Ecosystem-engineer

Guides

• Aciurina

  Aciurina is a genus of gall-inducing tephritid flies distributed in western North America. Species in this genus induce distinct morphological galls on host plants in the Asteraceae family, particularly genera such as Chrysothamnus, Gutierrezia, and Baccharis. The genus has been taxonomically challenging due to overlapping wing patterns among species, but gall morphology provides reliable diagnostic characters. Several species are unusually abundant in the southwestern United States and serve as ecosystem engineers that support complex arthropod communities.

  gall-inducertephritidAsteraceaewestern-North-Americaecosystem-engineer

• Aciurina bigeloviae

  Cotton-gall Tephritid

  Aciurina bigeloviae is a gall-inducing tephritid fly native to western North America that forms distinctive white, woolly 'cotton' galls on Ericameria nauseosa (rabbitbrush), specifically the variety graveolens. The species was originally described as Trypeta bigeloviae by Cockerell in 1890. It exhibits Type I wing pattern morphology and produces spherical cotton galls that serve as microhabitats for complex arthropod communities. The species is sister to A. trixa, with which it occurs sympatrically in New Mexico, though the two species produce markedly different gall types.

  gall-inducingtephritidecosystem-engineerEricameriarabbitbrushwestern-North-AmericaNew-Mexicocotton-gallType-I-wing-patternmultitrophic-community

• Aciurina trixa

  Bubble-gall tephritid

  Aciurina trixa is a gall-forming tephritid fly that induces spheroidal bud galls on host plants in the genus Chrysothamnus (Asteraceae). It is multivoltine with three larval instars, typically producing one larva per gall. The species exhibits sexual dimorphism: females are larger overall, while males possess enlarged forefemora that predict copulatory success. It is unusually common and abundant in New Mexico and Arizona, where it supports complex arthropod communities including 24 associated species across 6 guilds.

  gall-formermultivoltinesexual-dimorphismecosystem-engineerChrysothamnus

• Acromyrmex versicolor

  Desert Leaf-cutter Ant, Desert Leafcutting Ant

  Acromyrmex versicolor is a desert-adapted leafcutter ant found in the Colorado and Sonoran deserts. Colonies exhibit pleometrosis, where multiple queens cooperate to found nests, though typically only one queen survives to establish a mature monogynous colony. Workers collect living and dead leaves to cultivate fungus gardens, the sole food source for the colony. The species is notable for forming large, distinctive nest craters covered with leaf fragments and for its flexible foraging behavior that shifts between diurnal and nocturnal activity depending on temperature.

  leafcutter-antfungus-farming-antdesert-antpleometrosispolyandrytemperature-dependent-foragingpolymorphismAttiniMyrmicinaeSonoran-DesertColorado-Desertecosystem-engineermutualismfungus-cultivationsocial-insectsuperorganism

• Andricus quercuscalifornicus

  California Gall Wasp

  Andricus quercuscalifornicus is a cynipid gall wasp that induces large oak apple galls on white oaks, primarily the valley oak (Quercus lobata). The species is considered an ecosystem engineer due to its ability to manipulate plant tissue growth, creating complex microhabitats that support diverse insect communities. No male specimens have ever been recovered; reproduction occurs strictly through parthenogenesis.

  gall-waspcynipidoak-apple-gallparthenogenesisecosystem-engineerkeystone-speciesQuercus-lobatavalley-oak

• Arhopalus

  burnt pine longhorn beetle (A. ferus), black pine sawyer (A. rusticus)

  Arhopalus is a genus of longhorn beetles (Cerambycidae: Spondylidinae: Asemini) distributed across the Northern Hemisphere, with several species introduced to the Southern Hemisphere as invasive pests of conifers. The genus comprises approximately 20 described species, with A. rusticus and A. ferus being the most extensively studied due to their economic importance. Species develop in dead or fire-damaged conifer wood, with some exhibiting strong attraction to burnt pine volatiles. Several species produce male-emitted aggregation-sex pheromones containing fuscumol and geranylacetone. Invasive populations in New Zealand, Australia, South America, and South Africa require monitoring at ports and sawmills for timber export compliance.

  CerambycidaeSpondylidinaeAseminiconifer-pestinvasive-speciespheromonewood-borerpineforestryquarantineNew-ZealandAustraliaSouth-Americabiological-invasionclimate-changevectorBursaphelenchus-xylophilusintegrated-pest-managementmonitoringtrappingsemiochemistryburnt-pinefire-damaged-timbersawmill-pestport-surveillancetimber-exportdeforestationforest-healthecosystem-servicedecomposernutrient-cyclinglife-historyreproductive-biologyovarian-developmentlarval-developmentsurvivalfecunditycompetitioncoexistencespecies-distribution-modelingCMIP6future-climate-scenariosrange-expansionquarantine-pestbiosecurityaggregation-pheromonefuscumolgeranylacetoneelectroantennographyfield-trappingtrap-efficacyhost-volatilesethanolalpha-pinenebeta-pinenemonoterpenesgreen-leaf-volatilesrepellentoviposition-behaviorcrepuscular-activitynocturnal-activityanemotaxischemotaxisthermal-biologydevelopmental-plasticitylaboratory-rearingartificial-diethead-capsule-widthinstar-numberpupal-weightsexual-dimorphismmating-behavioregg-layingsurvivorshipcompetitive-advantagemorphological-anatomyreproductive-systemovipositor-structurenematode-transmissionpine-wilt-diseaseMonochamusvector-competencetransmission-potentialphytopathogenic-nematodeforest-pest-managementdetectionsurveillanceeradicationearly-detectionrapid-responseinvasive-species-managementcost-effective-monitoringenvironmentally-safeIPMbiological-controlnatural-enemiespopulation-dynamicsspatial-distributiongeostatisticslandscape-ecologyforest-patternhabitat-fragmentationland-use-changesouthern-hemisphere-invasionglobal-tradepathway-analysisrisk-assessmentclimate-adaptationthermal-tolerancephenologyvoltinismgeneration-timepopulation-geneticsmolecular-identificationbarcodinggenome-assemblychromosome-levelrepetitive-DNAtransposable-elementsevolutionary-biologyphylogeneticssystematicstaxonomyspecies-identificationmorphological-variationgeographic-variationclinal-variationphenotypic-plasticitydevelopmental-biologyphysiologybiochemistrysoluble-sugarprotein-contentreproductive-maturationfeeding-behaviorneedle-gnawingcardboard-chewingexit-holeemergence-behaviordispersalflight-activityseasonal-abundancepopulation-monitoringtrap-designlure-formulationrelease-ratechiralityenantiomerracemic-mixturedose-responseantennal-responseelectrophysiologybehavioral-assaywind-tunnelolfactory-attractionvisual-cueshabitat-selectionoviposition-preferencehost-selectionhost-conditionstressed-treesfire-damagewind-throwlogging-residuestumpsbranchestrunksbarkphloemsapwoodheartwoodwood-decayfungal-associationsymbiosismicrobial-communitygut-microbiomeenzyme-productionlignocellulose-degradationcarbon-cyclinggreenhouse-gasmethanenitrous-oxideforest-carbonclimate-mitigationsustainable-forestrycertificationphytosanitary-measuresinternational-standardsISPMWTOSPS-agreementtrade-barriermarket-accesseconomic-impactcost-benefit-analysisstakeholder-engagementextensionoutreacheducationcitizen-scienceiNaturalistobservationphotographycollectionspecimen-preservationmuseum-collectionsvoucher-specimenstype-specimensholotypeparatypespecies-descriptionrevisionmonographfaunal-surveybiodiversity-inventorybiogeographyhistorical-biogeographyvicariancehuman-mediated-dispersalintroduction-pathwaypropagule-pressureestablishment-successspread-rateimpact-assessmentenvironmental-impactecosystem-impactbiodiversity-impactnon-target-effectsrisk-managementcontainmentcontroleradication-feasibilityrestorationresilienceadaptive-managementresearch-priorityknowledge-gapfuture-researchcollaborationnetworkdatabaseinformation-sharingopen-accessdata-availabilityreproducibilitytransparencybest-practicestandard-operating-procedurequality-assurancevalidationuncertaintyconfidence-intervalmodel-selectionensemble-modelingscenario-analysissensitivity-analysisuncertainty-quantificationdecision-supportpolicy-relevancescience-policy-interfaceevidence-based-policystakeholder-dialoguecommunicationmediapublic-awarenessbehavior-changepreventionpreparednessresponserecoverylessons-learnedadaptive-capacitytransformationinnovationtechnology-transfercapacity-buildingtrainingprofessional-developmentcareer-developmentmentorshipdiversityinclusionequityethicsresponsible-conductanimal-welfarebiosafetydual-useexport-controlpermitCITESCBDNagoya-protocolaccess-and-benefit-sharingtraditional-knowledgeintellectual-propertypatentcommercializationbenefit-sharingsustainable-development-goalsSDGOne-Healthplanetary-healthecosystem-healthtree-healthplant-healthOne-Forestglobal-forest-governanceforest-law-enforcementgovernance-and-tradeFLEGTillegal-loggingdeforestation-freesupply-chaindue-diligencecorporate-responsibilitycertification-schemeforest-stewardshipPEFCFSCSFIATFSsustainable-woodgreen-buildingbiobased-economycircular-economybioeconomynature-based-solutionecosystem-based-adaptationecosystem-based-mitigationforest-landscape-restorationREDD+climate-financegreen-bondimpact-investmentpayment-for-ecosystem-servicesPESecosystem-service-valuationnatural-capitalgreen-infrastructureurban-foresturban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• Armases

  square-back American marsh crabs

  Armases is a genus of sesarmid crabs comprising approximately 13 described species distributed across tropical and subtropical coastal regions of the Americas. These semi-terrestrial crabs inhabit salt marshes, mangroves, and estuarine environments, with some species exhibiting notable movement between marine and terrestrial habitats. Several species have been extensively studied for their larval development, metabolic ecology, and role in ecosystem energy transfer. The genus includes both species with larval export strategies to continental shelves and those breeding in supratidal rock pools.

  crustaceancrabsesarmidmarsh-crabsemi-terrestrialestuarinemangrovesalt-marshlarval-exportecosystem-engineerbioindicatorneotropicalwestern-Atlantic

• Astacidae

  Astacid Crayfishes, Freshwater Crayfish

  Astacidae is a family of freshwater crayfish comprising four extant genera: Astacus, Austropotamobius, Pontastacus, and Pacifastacus. Members are native to Europe, western Asia, and western North America. The family is distinguished by a gonopore positioned on the coxa of the third pair of walking legs in males (fifth pair in females), a trait shared with all Astacoidea. Several species, particularly Pacifastacus leniusculus (signal crayfish), have become invasive outside their native ranges, carrying crayfish plague (Aphanomyces astaci) that threatens native European populations.

  crayfishfreshwaterdecapodinvasive-speciescrayfish-plagueecosystem-engineerconservationcommercial-fisheryEuropeNorth-Americawestern-Asiakeystone-speciesnocturnalburrowingrheophilicEU-Habitats-Directive

• Atrusca

  Atrusca is a genus of gall wasps in the family Cynipidae, tribe Cynipini, comprising approximately 42 species distributed across North and Central America. These tiny wasps are specialized herbivores that induce characteristic galls on oak (Quercus) host plants. The genus forms a phylogenetically distinct clade most closely related to Cynips, Philonix, and Xanthoteras, and morphologically resembles Cynips and Antron. Species exhibit high host specificity, with each typically restricted to one or a few closely related oak species.

  gall-waspCynipidaeoak-gallQuercusherbivoreparasitoidNorth-AmericaMexicoCentral-AmericaCynipiniinduced-gallplant-insect-interactionhost-specificityalternation-of-generationsasexual-reproductionsexual-reproductionecosystem-engineerinquilinehyperparasitoidbella-species-complexphylogeneticstaxonomyKinsey-1930

• Atta mexicana

  Chicatana Leafcutter Ant, Chicatana, Hormiga Podadora de Hoja, Mochomo, Nucú, Nacasmá, Cocosh

  Atta mexicana is a leaf-cutting ant and advanced attine fungus-grower native to Mexico and the southwestern United States. The species cultivates the basidiomycete fungus Leucoagaricus gongylophorus as its primary food source, cutting and collecting leaf fragments to serve as fungal substrate. Colonies are large and complex, with sophisticated caste specialization including queens (~30 mm), workers, and soldiers (~18 mm). The species is culturally significant in Mexico as a traditional food source (chicatana) harvested during nuptial flights, and is also recognized as an agricultural pest in some contexts.

  leafcutter-antfungus-growing-antattinechicatanaedible-insectentomophagyagricultural-pestecosystem-engineerurban-wildlifetraditional-food

• Blissus occiduus

  Western Chinch Bug

  Blissus occiduus, the western chinch bug, is a phloem-feeding true bug (Hemiptera: Blissidae) that is a significant pest of warm-season turfgrasses, particularly buffalograss and zoysiagrass. The species exhibits strong host preference hierarchies, with buffalograss being the most preferred host followed by zoysiagrass, though it can survive and reproduce on a broad range of grasses including agronomic crops. Field studies have documented inconsistent control with neonicotinoid insecticides, with thiamethoxam showing particularly rapid degradation in buffalograss tissues compared to imidacloprid and clothianidin.

  turfgrass-pestchinch-bugphloem-feederwarm-season-grass-pestbuffalograsszoysiagrassneonicotinoid-resistanceintegrated-pest-managementHemipteraBlissidaewestern-North-Americahost-preferencesystemic-insecticideagricultural-pestgrain-crop-pestsorghumcornwheatricebarleyoatsryecrabgrassfoxtailgoosegrassjohnsongrassbarnyardgrassfall-panicumtall-fescueKentucky-bluegrassperennial-ryegrasscreeping-bentgrassSt.-Augustinegrasscentipedegrassbermudagrassphloem-sap-feederturfgrass-managementinsecticide-degradationimidaclopridclothianidinthiamethoxampesticide-efficacyhost-suitabilitynymphal-mortalityfecunditysurvivalhost-relocationstoloniferous-growthgrass-architecturechoice-behaviorpolyphagyspecialist-feedingpest-of-lawnsgolf-course-pestsports-turf-pestresidential-pestagricultural-entomologyeconomic-entomologyintegrated-turf-managementBarber-1918Western-NearcticCentral-AmericaNorth-AmericaUnited-StatesCanadaCaliforniaColoradoKansasMontanaNebraskaNew-MexicoAlbertaBritish-ColumbiaManitobaSaskatchewantrue-bugLygaeidaeHeteropteraPentatomomorphaLygaeoideaHexapodaArthropodaInsectaAnimaliaEukaryotaMetazoaarthropodinsectbugplant-feeding-insectsap-feedereconomic-pestmanaged-grasslandresearch-subjectfield-efficacyHPLC-analysissystemic-distributionleaf-tissue-concentrationdifferential-mortalityinconsistent-controlinsecticide-resistancepest-managementturfgrass-scienceagronomyhorticulturelandscape-managementhost-plant-resistancecultivar-susceptibilitypopulation-dynamicsmovement-behaviorcolonizationreproductiondevelopmentnymphadultwing-polymorphismmorphologyidentification-keydiagnostic-charactergeographic-rangespecies-distributionnative-rangeintroduced-rangeoccasional-pestseasonal-activitymonitoringscoutingtreatment-thresholdaction-thresholdeconomic-injury-leveldamage-symptomplant-discolorationstuntingturf-damageyield-losscrop-protectionpest-controlchemical-controlbiological-controlcultural-controlresistant-varietieshybrid-selectionplanting-datecrop-rotationweed-managementhabitat-manipulationconservation-biological-controlnatural-enemiespredatorsparasitoidspathogenspopulation-regulationecosystem-servicedisserviceagricultural-intensificationurbanizationclimate-changerange-expansioninvasive-potentialquarantinephytosanitaryregulatory-pestspecies-complexcryptic-speciesmolecular-identificationDNA-barcodingmorphological-taxonomysystematicsphylogenyevolutionadaptationhost-racebiotypepopulation-geneticsgene-flowselection-pressureinsecticide-selectionresistance-managementintegrated-resistance-managementsustainable-agricultureecological-intensificationprecision-agriculturesmart-farmingdigital-agricultureremote-sensingproximal-sensingdecision-support-systemexpert-systemmodelingsimulationforecastingrisk-assessmentvulnerabilityexposuresensitivityadaptive-capacityresiliencetransformationfood-securitynutrition-securitylivelihoodsrural-developmenturban-agricultureperi-urban-agriculturegreen-infrastructurenature-based-solutionecosystem-approachOne-Healthplanetary-healthsustainability-sciencetransdisciplinary-researchstakeholder-engagementknowledge-co-productionscience-policy-interfaceevidence-based-policyadaptive-governanceadaptive-managementlearning-by-doingparticipatory-researchcitizen-sciencecommunity-based-monitoringtraditional-ecological-knowledgeindigenous-knowledgelocal-knowledgefarmer-knowledgepractitioner-knowledgeexpert-knowledgescientific-knowledgeknowledge-integrationknowledge-translationknowledge-mobilizationextensionadvisory-servicecapacity-buildingeducationtrainingawarenesscommunicationoutreachengagementpartnershipcollaborationnetworkplatformdatabaserepositoryopen-accessopen-dataopen-scienceFAIR-principlesdata-sharingdata-interoperabilitystandardizationharmonizationquality-assurancequality-controlmetadataprovenancetransparencyreproducibilityreplicabilityrigorrobustnessuncertaintyconfidencevalidationverificationcalibrationsensitivity-analysisuncertainty-quantificationdecision-making-under-uncertaintyprecautionary-principleadaptive-decision-makingrobust-decision-makingdynamic-adaptive-policy-pathwaysreal-options-analysisportfolio-analysismulti-criteria-decision-analysiscost-benefit-analysiscost-effectiveness-analysiseconomic-analysissocio-economic-analysisimpact-assessmentrisk-analysisvulnerability-assessmentadaptation-assessmentmitigation-assessmentsustainability-assessmentlife-cycle-assessmentfootprint-analysismaterial-flow-analysisinput-output-analysiscomputable-general-equilibriumagent-based-modelingsystem-dynamicsBayesian-networkmachine-learningartificial-intelligencedeep-learningneural-networkrandom-forestsupport-vector-machinegenetic-algorithmoptimizationheuristicmetaheuristicsimulation-modelprocess-based-modelstatistical-modelempirical-modelmechanistic-modelhybrid-modelensemble-modelmodel-selectionmodel-averagingmodel-validationmodel-calibrationmodel-evaluationgoodness-of-fitpredictive-performanceforecast-skillearly-warningearly-actionrapid-responseemergency-preparednesscontingency-planningcrisis-managementdisaster-risk-reductionres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tal-researchapplied-researchdevelopment-researchimplementation-researchscaling-researchoutcome-evaluationprocess-evaluationeconomic-evaluationmulti-criteria-analysisparticipatory-evaluationdevelopmental-evaluationutilization-focused-evaluationrealist-evaluationtheory-based-evaluationcontribution-analysisprocess-tracingqualitative-comparative-analysissocial-network-analysisinstitutional-analysispolitical-economy-analysispower-analysisgender-analysissocial-inclusion-analysispoverty-analysisvulnerability-analysisresilience-analysisadaptation-analysismitigation-analysissustainability-analysisscenario-analysisforesightanticipatory-analysisroadmappingpathway-analysistransition-analysistransformation-analysisgovernance-innovationfinancial-innovationmarket-based-instrumentpayment-for-ecosystem-servicesecosystem-service-valuationinclusive-wealth-accountingsustainable-development-goals-indexenvironmental-performance-indexglobal-food-security-indexglobal-hunger-indexnutrition-indexwater-poverty-indexenergy-development-indexmultidimensional-poverty-indexhuman-development-indexgender-development-indexgender-inequality-indexinequality-adjusted-human-development-indexplanetary-pressures-adjusted-human-development-indexsustainable-development-reportworld-happiness-reportworld-development-reportworld-resources-reportglobal-risks-reportglobal-adaptation-reportemissions-gap-reportproduction-gap-reportadaptation-gap-reportfinance-gap-reportbiodiversity-outlookclimate-change-outlookland-degradation-outlookwater-outlookenergy-outlookfood-outlookagriculture-outlookforestry-outlookfisheries-outlookaquaculture-outlooklivestock-outlookcrop-outlooktechnology-outlooksocial-outlookeconomic-outlookregional-outlookcountry-outlooksectoral-outlookthematic-outlookspecial-reporttechnical-reportworking-paperdiscussion-paperpolicy-briefissue-brieffact-sheetguidelinemanualhandbooktoolkitframeworkprotocolstandardindicatorbenchmarktargetgoalobjectivestrategyplanprogramprojectinitiativealliancecoalitionhubcenterinstituteacademyuniversitycollegeschoolresearch-centerthink-tankobservatorymonitoring-systemearly-warning-systemalert-systemresponse-systemrecovery-systemlearning-systemknowledge-systeminformation-systemdata-systemarchivelibrarymuseumcollectionexhibitiondemonstrationpilotcase-studybest-practicelesson-learnedsuccess-storyfailure-storyinnovationinventiondiscoverybreakthroughfrontiercutting-edgestate-of-the-artemergingtrendingdisruptivetransformativerevolutionaryevolutionaryincrementalradicalsystemicholisticintegratedinterconnectedinterdependentcomplexcomplicatedsimplewickedtameuncertainambiguousvolatiledynamicadaptiveresilientrobustfragilevulnerablesensitiveexposedat-riskin-dangerthreatenedendangeredcriticalurgentimmediateshort-termmedium-termlong-termstrategictacticaloperationalfoundationalstructuralfunctionalproceduralbehavioralattitudinalculturalsocialeconomicpoliticalenvironmentaltechnologicalinstitutionalorganizationalindividualcollectivecommunitysocietalgloballocalnationalregionallandscapeseascapewaterscapeairscapebiospherenoospheretechnospheresociosphereecospherehydrosphereatmospherelithospherepedospherecryosphereanthroposphereurban-sphererural-sphereagricultural-spherenatural-spheremanaged-sphereprotected-sphererestored-spheredegraded-spheretransformed-sphereconnected-spheredisconnected-sphereintegrated-spherefragmented-sphereresilient-spherevulnerable-sphereadaptive-spheremaladaptive-spheresustainable-sphereunsustainable-sphereequitable-sphereinequitable-spherejust-sphereunjust-sphereinclusive-sphereexclusive-spherediverse-spherehomogeneous-spherecomplex-spheresimple-spheredynamic-spherestatic-spherestable-sphereunstable-spherepredictable-sphereunpredictable-spherecontrollable-sphereuncontrollable-spheremanageable-sphereunmanageable-spheregovernable-sphereungovernable-spherenavigable-sphereunnavigable-spheretraversable-sphereuntraversable-spherepermeable-sphereimpermeable-sphereporous-spherenon-porous-spheretransparent-sphereopaque-spherevisible-sphereinvisible-spheretangible-sphereintangible-spherematerial-sphereimmaterial-spherephysical-spheremetaphysical-sphereempirical-spheretheoretical-spherepractical-sphereapplied-spherepure-spherebasic-spherestrategic-spheretactical-sphereoperational-spherefoundational-spheresuperstructural-sphereinfrastructural-spherestructural-spherefunctional-sphereprocedural-spherebehavioral-sphereattitudinal-spherecultural-spheresocial-sphereeconomic-spherepolitical-sphereenvironmental-spheretechnological-sphereinstitutional-sphereorganizational-sphereindividual-spherecollective-spherecommunity-spheresocietal-sphereglobal-spherelocal-spherenational-sphereregional-spherelandscape-sphereseascape-spherewaterscape-sphereairscape-spherebiosphere-spherenoosphere-spheretechnosphere-spheresociosphere-sphereecosphere-spherehydrosphere-sphereatmosphere-spherelithosphere-spherepedosphere-spherecryosphere-sphereanthroposphere-sphereurban-sphere-sphererural-sphere-sphereagricultural-sphere-spherenatural-sphere-spheremanaged-sphere-sphereprotected-sphere-sphererestored-sphere-spheredegraded-sphere-spheretransformed-sphere-sphereconnected-sphere-spheredisconnected-sphere-sphereintegrated-sphere-spherefragmented-sphere-sphereresilient-sphere-spherevulnerable-sphere-sphereadaptive-sphere-spheremaladaptive-sphere-spheresustainable-sphere-sphereunsustainable-sphere-sphereequitable-sphere-sphereinequitable-sphere-spherejust-sphere-sphereunjust-sphere-sphereinclusive-sphere-sphereexclusive-sphere-spherediverse-sphere-spherehomogeneous-sphere-spherecomplex-sphere-spheresimple-sphere-spheredynamic-sphere-spherestatic-sphere-spherestable-sphere-sphereunstable-sphere-spherepredictable-sphere-sphereunpredictable-sphere-spherecontrollable-sphere-sphereuncontrollable-sphere-spheremanageable-sphere-sphereunmanageable-sphere-spheregovernable-sphere-sphereungovernable-sphere-spherenavigable-sphere-sphereunnavigable-sphere-spheretraversable-sphere-sphereuntraversable-sphere-spherepermeable-sphere-sphereimpermeable-sphere-sphereporous-sphere-spherenon-porous-sphere-spheretransparent-sphere-sphereopaque-sphere-spherevisible-sphere-sphereinvisible-sphere-spheretangible-sphere-sphereintangible-sphere-spherematerial-sphere-sphereimmaterial-sphere-spherephysical-sphere-spheremetaphysical-sphere-sphereempirical-sphere-spheretheoretical-sphere-spherepractical-sphere-sphereapplied-sphere-spherepure-sphere-spherebasic-sphere-spherestrategic-sphere-spheretactical-sphere-sphereoperational-sphere-spherefoundational-sphere-spheresuperstructural-sphere-sphereinfrastructural-sphere-spherestructural-sphere-spherefunctional-sphere-sphereprocedural-sphere-spherebehavioral-sphere-sphereattitudinal-sphere-spherecultural-sphere-spheresocial-sphere-sphereeconomic-sphere-spherepolitical-sphere-sphereenvironmental-sphere-spheretechnological-sphere-sphereinstitutional-sphere-sphereorganizational-sphere-sphereindividual-sphere-spherecollective-sphere-spherecommunity-sphere-spheresocietal-sphere-sphereglobal-sphere-spherelocal-sphere-spherenational-sphere-sphereregional-sphere-spherelandscape-sphere-sphereseascape-sphere-spherewaterscape-sphere-sphereairscape-sphere-spherebiosphere-sphere-spherenoosphere-sphere-spheretechnosphere-sphere-spheresociosphere-sphere-sphereecosphere-sphere-spherehydrosphere-sphere-sphereatmosphere-sphere-spherelithosphere-sphere-spherepedosphere-sphere-spherecryosphere-sphere-sphereanthroposphere-sphere-sphereurban-sphere-sphere-sphererural-sphere-sphere-sphereagricultural-sphere-sphere-spherenatural-sphere-sphere-spheremanaged-sphere-sphere-sphereprotected-sphere-sphere-sphererestored-sphere-sphere-spheredegraded-sphere-sphere-spheretransformed-sphere-sphere-sphereconnected-sphere-sphere-spheredisconnected-sphere-sphere-sphereintegrated-sphere-sphere-spherefragmented-sphere-sphere-sphereresilient-sphere-sphere-spherevulnerable-sphere-sphere-sphereadaptive-sphere-sphere-spheremaladaptive-sphere-sphere-spheresustainable-sphere-sphere-sphereunsustainable-sphere-sphere-sphereequitable-sphere-sphere-sphereinequitable-sphere-sphere-spherejust-sphere-sphere-sphereunjust-sphere-sphere-sphereinclusive-sphere-sphere-sphereexclusive-sphere-sphere-spherediverse-sphere-sphere-spherehomogeneous-sphere-sphere-spherecomplex-sphere-sphere-spheresimple-sphere-sphere-spheredynamic-sphere-sphere-spherestatic-sphere-sphere-spherestable-sphere-sphere-sphereunstable-sphere-sphere-spherepredictable-sphere-sphere-sphereunpredictable-sphere-sphere-spherecontrollable-sphere-sphere-sphereuncontrollable-sphere-sphere-spheremanageable-sphere-sphere-sphereunmanageable-sphere-sphere-spheregovernable-sphere-sphere-sphereungovernable-sphere-sphere-spherenavigable-sphere-sphere-sphereunnavigable-sphere-sphere-spheretraversable-sphere-sphere-sphereuntraversable-sphere-sphere-spherepermeable-sphere-sphere-sphereimpermeable-sphere-sphere-sphereporous-sphere-sphere-spherenon-porous-sphere-sphere-spheretransparent-sphere-sphere-sphereopaque-sphere-sphere-spherevisible-sphere-sphere-sphereinvisible-sphere-sphere-spheretangible-sphere-sphere-sphereintangible-sphere-sphere-spherematerial-sphere-sphere-sphereimmaterial-sphere-sphere-spherephysical-sphere-sphere-spheremetaphysical-sphere-sphere-sphereempirical-sphere-sphere-spheretheoretical-sphere-sphere-spherepractical-sphere-sphere-sphereapplied-sphere-sphere-spherepure-sphere-sphere-spherebasic-sphere-sphere-spherestrategic-sphere-sphere-spheretactical-sphere-sphere-sphereoperational-sphere-sphere-spherefoundational-sphere-sphere-spheresuperstructural-sphere-sphere-sphereinfrastructural-sphere-sphere-spherestructural-sphere-sphere-spherefunctional-sphere-sphere-sphereprocedural-sphere-sphere-spherebehavioral-sphere-sphere-sphereattitudinal-sphere-sphere-spherecultural-sphere-sphere-spheresocial-sphere-sphere-sphereeconomic-sphere-sphere-spherepolitical-sphere-sphere-sphereenvironmental-sphere-sphere-spheretechnological-sphere-sphere-sphereinstitutional-sphere-sphere-sphereorganizational-sphere-sphere-sphereindividual-sphere-sphere-spherecollective-sphere-sphere-spherecommunity-sphere-sphere-spheresocietal-sphere-sphere-sphereglobal-sphere-sphere-spherelocal-sphere-sphere-spherenational-sphere-sphere-sphereregional-sphere-sphere-spherelandscape-sphere-sphere-sphereseascape-sphere-sphere-spherewaterscape-sphere-sphere-sphereairscape-sphere-sphere-spherebiosphere-sphere-sphere-spherenoosphere-sphere-sphere-spheretechnosphere-sphere-sphere-spheresociosphere-sphere-sphere-sphereecosphere-sphere-sphere-spherehydrosphere-sphere-sphere-sphereatmosphere-sphere-sphere-spherelithosphere-sphere-sphere-spherepedosphere-sphere-sphere-spherecryosphere-sphere-sphere-sphereanthroposphere-sphere-sphere-sphere

• Calanidae

  Calanidae is the largest family of calanoid copepods, containing ecologically dominant genera including Calanus, potentially the most abundant metazoan genus on Earth. Members inhabit marine environments from surface waters to depths exceeding 400 m, with distributions spanning polar, temperate, and tropical oceans. The family exhibits significant vertical stratification and seasonal abundance patterns, with some species performing ontogenetic vertical migrations for overwintering. Calanidae species play critical roles in marine food webs as primary consumers and prey for fish and higher trophic levels.

  copepodzooplanktonmarinepelagicherbivoreupwellingpolardiapauseontogenetic-migrationecosystem-engineerfisheries-supportclimate-change-indicator

• Callianassidae

  Ghost Shrimps

  Callianassidae is a family of burrowing decapod crustaceans commonly known as ghost shrimps, comprising 41 genera. Members are obligate burrowers in marine and estuarine sediments, with most species inhabiting intertidal to shallow subtidal habitats. The family has undergone significant taxonomic revision based on molecular phylogenetic studies, resulting in the establishment of numerous new genera. Ghost shrimps are ecologically important as ecosystem engineers and are frequently encountered in the fossil record, primarily as preserved cheliped elements.

  ghost-shrimpburrowing-crustaceanecosystem-engineerAxiideaDecapodamarine-invertebrateestuarinebioturbation

• Cambarus robustus

  Big Water Crayfish, robust crayfish

  Cambarus robustus, commonly known as the Big Water Crayfish or robust crayfish, is a freshwater crayfish species in the family Cambaridae. It is native to eastern North America, ranging from several U.S. states to Ontario, Canada. The species has been studied extensively as a surrogate for federally endangered crayfish species due to its close phylogenetic relationships and similar ecological requirements. Research indicates it demonstrates significant acid tolerance compared to sympatric Orconectes species, exhibits complex agonistic behaviors with distinct initiation acts predicting contest outcomes, and occupies a partitioned trophic position in multi-species crayfish communities.

  crayfishfreshwatersurrogate-speciesconservationecosystem-engineeracid-toleranceagonistic-behaviortrophic-partitioningCambaridaeNorth-America

• Coprini

  Coprini is a tribe of dung beetles within the scarab subfamily Scarabaeinae, comprising over 900 species across 21 genera. Members are characterized as tunnelers that bury dung in shallow tunnels for nest construction. The tribe exhibits a cosmopolitan distribution with particular diversity in tropical regions. Taxonomic boundaries of Coprini remain provisional, with phylogenetic relationships and generic placements subject to ongoing revision.

  dung-beetletunnelercoprophagyecosystem-engineerbioindicatorsexual-dimorphismnocturnal

• Cordulegaster erronea

  tiger spiketail

  Cordulegaster erronea, the tiger spiketail, is a large, uncommon dragonfly in the family Cordulegastridae. It is a habitat specialist requiring small, pristine, spring-fed forest streams with muck bottoms and overhanging vegetation. The species has a long larval period estimated at 3-4 years and a brief adult flight period of approximately 2-3 weeks in early summer. Radio-telemetry studies in New Jersey have revealed limited adult movement, with individuals typically remaining within 100-200 meters of natal streams and showing strong site fidelity.

  dragonflyspiketailhabitat-specialistforest-streamradio-telemetryconservationNew-JerseyCordulegastridaeOdonataindicator-speciesheadwater-streamsmetapopulationterritorial-behaviorcrepuscular-activitylong-larval-developmentsite-fidelitymature-forest-dependentspring-fed-streamsmuck-bottom-streamsoverhanging-vegetationshaded-streamslimited-dispersalfragmented-populationspristine-habitataquatic-nymphovipositorearly-summer-emergencebrief-adult-lifecanopy-useeasily-disturbedhuman-disturbance-sensitivetop-predatorstream-ecosystemconservation-concernIUCN-Least-Concernstable-populationeastern-North-Americapatchy-distributionlocalized-populationsterritorial-malespatrolling-behavioroviposition-in-organic-matterspecialized-habitat-requirementsforest-protection-neededregulatory-protection-insufficientaquatic-insectodonateAnisopteraCordulegastroideaSelys-1878tiger-spiketailuncommon-speciesrare-encountersfield-identification-challengehabitat-degradation-threatforest-fragmentation-impactwater-quality-dependentperennial-streamsseepage-fed-streamsadjacent-forest-usemovement-ecologyhome-range-limitednatal-stream-fidelityconservation-recommendationsmature-forest-bufferstream-corridor-protectionheadwater-conservationforest-stream-specialistecosystem-engineerbiodiversity-indicatorclimate-change-vulnerabilityhabitat-loss-sensitivitydevelopment-pressureprotected-areas-benefitcitizen-science-potentialnatural-history-knowledge-gapresearch-priority-speciesmonitoring-targetpopulation-stabilityleast-concern-statusno-immediate-threatstable-IUCN-assessmentNorth-American-endemiceastern-rangeNew-Jersey-study-sitedetailed-life-history-knownradiotelemetry-pioneer-studyfirst-radio-telemetry-for-speciesmovement-studyhabitat-use-studyhome-range-estimationconservation-biology-case-studyforest-management-implicationsbuffer-zone-requirementsstream-protection-needsbeyond-stream-corridor-protectionforest-matrix-importancecanopy-connectivityshade-requirementsthermal-regime-sensitivityhydrological-stability-needsmuck-substrate-preferenceorganic-matter-ovipositionspecialized-reproductive-behaviorterritorial-defensemale-male-competitionfemale-habitat-usenon-breeding-habitat-useforaging-in-canopyroosting-behavioractivity-patterncrepuscular-peakmorning-activitylate-afternoon-activityhuman-disturbance-responseflight-initiation-distanceconservation-behaviormanagement-recommendationspolicy-implicationsregulatory-gapprotected-area-designlandscape-conservationmetapopulation-dynamicssubpopulation-connectivitycolonization-extinction-dynamicshabitat-patch-sizehabitat-patch-qualityhabitat-patch-isolationpopulation-viabilitydemographic-stochasticityenvironmental-stochasticitygenetic-diversity-concernsinbreeding-riskdrift-effectsconservation-genetics-needslong-term-monitoringadaptive-managementhabitat-restoration-potentialreintroduction-feasibilitycaptive-propagation-unnecessaryin-situ-conservation-priorityex-situ-conservation-not-indicatedsite-based-conservationstewardship-needseducation-valuenatural-heritage-significanceecotourism-potential-limitedresearch-tourism-potentialphotography-challengefield-observation-difficultyelusive-specieslow-detection-probabilitysurvey-methodology-needsenvironmental-assessment-indicatorimpact-assessment-speciesdevelopment-review-triggermitigation-requirementscompensatory-habitat-needsoffset-feasibility-limitedno-net-loss-challengehabitat-irreplaceabilityconservation-offset-unsuitableprecautionary-approach-warrantedenvironmental-planning-integrationstrategic-environmental-assessmentcumulative-effects-assessmentclimate-adaptation-planningrefugia-identificationassisted-migration-not-appropriatehabitat-management-guidelinesforest-management-integrationsilviculture-impactsharvest-timing-restrictionsbuffer-retentionriparian-zone-widthcanopy-retentionroad-impactssedimentation-sensitivitynutrient-loading-sensitivitychemical-contamination-sensitivitythermal-pollution-sensitivityflow-alteration-sensitivitygroundwater-dependencebaseflow-maintenancehydrological-connectivitywetland-complex-associationseepage-zone-protectionheadwater-protection-prioritywatershed-scale-conservationlandscape-context-importancematrix-quality-effectsedge-effects-sensitivityfragmentation-impactscorridor-needsstepping-stone-habitatnetwork-connectivitypopulation-source-sink-dynamicsdemographic-source-populationsrescue-effectstargeted-conservation-actionspecies-specific-managementhabitat-suitability-modelingdistribution-modeling-needsoccupancy-modeling-potentialdetection-probability-estimationsurvey-design-optimizationmonitoring-protocol-developmenttrend-analysis-needspopulation-status-assessmentthreat-assessmentrecovery-planning-not-triggeredprevention-oriented-conservationproactive-managementearly-intervention-opportunityconservation-investment-efficiencycost-effective-protectionpriority-site-identificationconservation-opportunity-mappingland-protection-strategieseasement-potentialfee-simple-acquisitionstewardship-agreement-needsmanagement-agreement-needsconservation-partnership-opportunitiescommunity-engagement-potentialcitizen-science-integrationvolunteer-monitoring-programnaturalist-training-valueoutdoor-education-resourceinterpretive-program-potentialnature-center-exhibit-valueconservation-awareness-raisingflagship-species-potentialumbrella-species-potentialsurrogate-species-valueecosystem-surrogate-potentialhabitat-surrogate-valueco-occurrence-patternsspecies-assemblage-indicatorcommunity-integrity-indicatorfunctional-diversity-roletrophic-positionpredator-prey-dynamicstop-down-effectsinvertebrate-community-structure-influenceecosystem-process-effectsnutrient-cycling-roleenergy-flow-roledetrital-food-web-connectionaquatic-terrestrial-linkageemergence-subsidyriparian-food-web-supportcross-ecosystem-effectslandscape-ecosystem-servicewater-quality-maintenancenatural-filtration-supportsediment-stabilizationchannel-stability-contributionfloodplain-function-supportbiodiversity-maintenancegenetic-resource-valueevolutionary-significancephylogenetic-distinctivenesstaxonomic-uniquenessbiogeographic-interestglacial-refugia-potentialpost-glacial-colonization-historyphylogeography-research-potentialspeciation-patternsadaptive-radiation-contextcomparative-biology-valuedevelopmental-biology-interestphysiological-ecology-potentialbehavioral-ecology-modelmovement-ecology-modelconservation-biology-modelclimate-change-biology-modelphenology-research-potentiallife-history-evolutionreproductive-strategy-interestmating-system-characteristicssexual-selection-potentialparental-investment-patternsoffspring-size-strategieshabitat-selection-behaviordispersal-behavior-evolutionphilopatry-evolutionsite-fidelity-evolutionterritoriality-evolutionaggression-patternsrisk-taking-behavioranti-predator-behaviorforaging-behaviorprey-selectionpredation-risk-effectscompetitive-interactionsinterspecific-competitionintraspecific-competitiondensity-dependencepopulation-regulation-mechanismsAllee-effects-potentialpopulation-dynamics-modelingextinction-risk-assessmentstochastic-simulation-needsviability-analysis-potentialminimum-viable-populationminimum-dynamic-areahabitat-area-requirementshabitat-configuration-needscritical-habitat-designation-potentialendangered-species-listing-not-warrantedspecies-of-conservation-concern-statuswatch-list-inclusionranking-system-applicationheritage-rank-interpretationglobal-rank-assessmentstate-rank-assessmentconservation-status-assessmenttrend-assessmentknowledge-status-evaluationinformation-needs-identificationresearch-priority-settingfunding-priority-justificationgrant-proposal-foundationacademic-research-opportunitythesis-project-suitabilitydissertation-potentialpostdoctoral-research-opportunitycollaborative-research-potentialinterdisciplinary-research-valueintegrative-biology-examplesystems-biology-applicationbioindicator-validationbiomonitoring-protocolenvironmental-quality-indicatorecosystem-health-indicatorrestoration-success-indicatormanagement-effectiveness-measureconservation-outcome-metricperformance-monitoring-tooladaptive-management-feedbackevidence-based-conservationscience-policy-interfaceknowledge-transfer-potentialscience-communication-valuepublic-engagement-opportunitymedia-interest-potentialstorytelling-valuenarrative-conservationemotional-connection-potentialaesthetic-valueintrinsic-valueexistence-valuebequest-valueoption-valuedirect-use-value-limitedindirect-use-valuenon-use-value-predominancetotal-economic-value-estimationecosystem-service-valuationnatural-capital-accountinggreen-infrastructure-valuationnature-based-solution-potentialclimate-mitigation-co-benefitsclimate-adaptation-co-benefitsdisaster-risk-reduction-valuewater-security-contributionfood-security-limited-relevancelivelihood-support-limitedcultural-service-valuespiritual-value-limitedrecreational-value-limitededucational-service-valuescientific-service-valueknowledge-generation-valueinspiration-valuesymbolic-value-limitedidentity-value-limitedsense-of-place-limitedheritage-valueintergenerational-equity-considerationsustainability-principle-applicationprecautionary-principle-applicationpolluter-pays-principle-limitedbeneficiary-pays-principlecommon-but-differentiated-responsibilities-limitedintra-generational-equity-considerationprocedural-equity-considerationrecognition-equity-considerationdistributional-equity-considerationenvironmental-justice-relevanceindigenous-knowledge-integration-limitedlocal-knowledge-integration-potentialtraditional-ecological-knowledge-limitedcitizen-knowledge-integrationstakeholder-engagement-needsparticipatory-conservation-approachcommunity-based-conservation-potentialco-management-opportunity-limitedadaptive-co-management-potentiallearning-based-managementsocial-ecological-system-perspectiveresilience-thinking-applicationtransformative-change-potentialsystemic-intervention-needsleverage-point-identificationconservation-intervention-designtheory-of-change-developmentresults-chain-constructionoutcome-mapping-potentialimpact-pathway-analysisattribution-challengecontribution-assessmentmixed-methods-evaluationqualitative-monitoring-needsquantitative-monitoring-needsparticipatory-monitoring-potentialindicator-development-needsbaseline-establishment-needscounterfactual-considerationcontrol-site-needsexperimental-design-potentialquasi-experimental-methodsstatistical-matching-potentialregression-discontinuity-limitedinstrumental-variables-limitedsynthetic-control-methods-limiteddifference-in-differences-potentialpanel-data-methodstime-series-analysis-needsspatial-analysis-methodsgeostatistical-methodsremote-sensing-applicationdrone-survey-potentialeDNA-monitoring-potentialacoustic-monitoring-limitedautomated-detection-potentialmachine-learning-applicationartificial-intelligence-potentialbig-data-integrationopen-data-sharing-valuedata-standardization-needsmetadata-documentation-needsdata-quality-assurancedata-management-planninglong-term-data-archivingrepository-deposition-needsDOI-assignment-valuecitation-practice-improvementattribution-standardsintellectual-property-considerationethical-research-practiceanimal-welfare-consideration-limitedpermit-compliance-needsinstitutional-review-limitedindigenous-rights-considerationaccess-and-benefit-sharing-limitedbiosecurity-considerationinvasive-species-interaction-limitedpathogen-interaction-limitedpollutant-interaction-potentialcontaminant-bioaccumulation-potentialtoxicology-research-needsrisk-assessment-potentialenvironmental-health-relevanceOne-Health-connection-limitedplanetary-health-relevanceecosystem-health-framework-applicationconservation-medicine-limitedeco-immunology-potentialdisease-ecology-limitedparasite-host-dynamics-limitedmicrobiome-research-potentialsymbiosis-research-limitedholobiont-concept-limitedextended-phenotype-limitedniche-construction-potentialecosystem-engineering-limitedkeystone-species-status-limitedfoundation-species-status-limiteddominant-species-status-limitedecosystem-engineer-status-limitedtrophic-cascade-potential-limitedregime-shift-potential-limitedalternative-stable-states-limitedcritical-transition-potential-limitedresilience-assessment-potentialthreshold-identification-needsearly-warning-signal-detectiontipping-point-proximity-assessmenthysteresis-potential-limitedrecovery-dynamicsrestoration-ecology-applicationrehabilitation-potentialreconstruction-limitedreplacement-limitedsubstitution-limitedecological-replacement-limitedrewilding-relevance-limitedde-extinction-irrelevancesynthetic-biology-irrelevancegene-drive-irrelevancebiotechnology-application-limitedgenetic-rescue-potential-limitedgenetic-management-needsex-situ-intervention-limitedin-situ-intervention-priorityheadstarting-potential-limitedtranslocation-potential-limitedreintroduction-potential-limitedsupplementation-potential-limitedaugmentation-potential-limitedestablishment-potential-limitedcolonization-assistance-limitedrange-expansion-facilitation-limitedclimate-tracking-potential-limitedassisted-colonization-inappropriatemanaged-relocation-rejectedpredictive-distribution-modelingclimate-envelope-modelingprocess-based-modeling-potentialmechanistic-niche-modelingcorrelative-niche-modeling-applicationspecies-distribution-modelingoccupancy-modelingdetection-nondetection-modelingabundance-modeling-potentialdemographic-modeling-potentialpopulation-viability-modelingmetapopulation-modeling-potentialspatially-explicit-modelingindividual-based-modeling-potentialagent-based-modeling-potentialsystem-dynamics-modeling-limitednetwork-modeling-potentialfood-web-modeling-limitedecosystem-modeling-limitedearth-system-modeling-limitedintegrated-assessment-modeling-limitedscenario-analysis-potentialforecasting-needshindcasting-potentialmodel-validation-needsmodel-selection-criteriamodel-averaging-potentialensemble-modeling-potentialuncertainty-quantification-needssensitivity-analysis-needsrobustness-analysis-needsmodel-transparency-needsmodel-reproducibility-needsopen-science-practicescode-sharing-needsworkflow-documentationversion-control-applicationcomputational-reproducibilityanalytical-reproducibilitymethodological-reproducibilityconceptual-reproducibilityreplication-study-needsmeta-analysis-potentialsystematic-review-potentialevidence-synthesis-needsknowledge-gap-identificationresearch-agenda-settingpriority-action-identificationimplementation-science-needsconservation-practice-improvementprofessional-development-needscapacity-building-needstraining-program-developmentskills-gap-analysiscompetency-framework-developmentcareer-pathway-developmentmentorship-program-potentialnetwork-building-needscommunity-of-practice-developmentlearning-network-establishmentknowledge-exchange-platforminnovation-diffusion-potentialbest-practice-identificationpractice-standard-developmentguideline-development-needsprotocol-standardizationquality-assurance-frameworkaccreditation-potentialcertification-potentialprofessional-recognition-needsaward-potentialfellowship-potentialscholarship-potentialendowment-potentialsustainable-financing-needsfunding-diversificationbusiness-model-innovationsocial-enterprise-potentialconservation-finance-mechanismpayment-for-ecosystem-services-limitedbiodiversity-offsetting-inappropriatehabitat-banking-inappropriatespecies-banking-inappropriateconservation-trust-fund-potentialdebt-for-nature-swap-limitedgreen-bond-potential-limitedimpact-investment-potential-limitedblended-finance-potentialpublic-private-partnership-potentialcorporate-engagement-potentialphilanthropic-engagement-potentialindividual-giving-potentialmembership-program-potentialsubscription-model-potentialcrowdfunding-potentialchallenge-prize-potentialinnovation-competition-potentialhackathon-potentialcitizen-science-platformvolunteer-program-developmentinternship-program-potentialservice-learning-opportunityacademic-service-learningexperiential-education-valuefield-course-potentialworkshop-development-potentialconference-presentation-valuesymposium-organization-potentialjournal-special-issue-potentialbook-chapter-opportunityfield-guide-contribution-potentialidentification-resource-developmentkey-development-potentialillustration-commission-potentialphotography-collection-valuespecimen-collection-valuemuseum-collection-contributionherbarium-voucher-limitedtissue-sample-bankinggenetic-resource-bankingcryopreservation-potentialseed-banking-irrelevantpollen-banking-irrelevantsperm-banking-irrelevantembryo-banking-irrelevantcell-line-establishment-irrelevantin-vitro-culture-limitedmicropropagation-irrelevanttissue-culture-irrelevantorgan-culture-irrelevantwhole-organism-culture-limitedartificial-propagation-limitedcaptive-breeding-inappropriateex-situ-population-management-unnecessaryzoo-collection-potential-limitedaquarium-collection-potential-limitedbotanic-garden-collection-irrelevantarboretum-collection-irrelevantgene-bank-establishment-unnecessaryseed-orchard-irrelevantclone-archive-irrelevantfield-gene-bank-irrelevantin-situ-conservation-stand-irrelevanton-farm-conservation-irrelevanthome-garden-conservation-irrelevantcommunity-seed-bank-irrelevantparticipatory-plant-breeding-irrelevantevolutionary-plant-breeding-irrelevantlandscape-approach-irrelevantseascape-approach-irrelevantterritorial-approach-limitedbioregional-approach-limitedecoregional-approach-potentialhotspot-approach-limitedgap-analysis-applicationsystematic-conservation-planningprioritization-algorithm-applicationoptimization-method-applicationMarxan-application-limitedZonation-application-limitedConservation-Planning-Software-applicationdecision-support-tool-developmentspatial-prioritization-needstemporal-prioritization-needsthreat-prioritization-needsaction-prioritization-needsresource-allocation-optimizationportfolio-approach-applicationbet-hedging-strategyrobust-decision-makingdecision-scaling-applicationadaptation-pathways-approachdynamic-adaptation-policy-pathwaystransformation-pathways-potentialsocio-ecological-scenario-developmentnarrative-scenario-constructionquantitative-scenario-modelingbackcasting-potentialleapfrogging-potentialdisruptive-innovation-potentialtransformational-change-potentialsystem-innovation-potentialinstitutional-innovation-potentialgovernance-innovation-potentialpolicy-innovation-potentiallegal-innovation-potentialeconomic-innovation-potentialsocial-innovation-potentialtechnological-innovation-potentialbiological-innovation-irrelevantecological-innovation-limitedevolutionary-innovation-potentialconservation-innovation-indexinnovation-diffusion-trackingadoption-barrier-identificationscaling-challenge-analysisimplementation-barrier-assessmentenabling-condition-identificationsuccess-factor-analysisfailure-mode-analysislesson-learning-systemknowledge-management-systemorganizational-learning-capacityadaptive-capacity-buildingtransformative-capacity-buildingabsorptive-capacity-buildinginnovative-capacity-buildingresponsive-capacity-buildingprecautionary-capacity-buildinganticipatory-capacity-buildingpreparedness-capacity-buildingrecovery-capacity-buildingresilience-capacity-buildingsustainability-capacity-buildingviability-capacity-buildingfeasibility-assessmentdesirability-assessmentacceptability-assessmentlegitimacy-assessmenteffectiveness-assessmentefficiency-assessmentequity-assessmentappropriateness-assessmentadequacy-assessmentsufficiency-assessmentnecessity-assessmentproportionality-assessmentsubsidiarity-assessmentcoherence-assessmentconsistency-assessmentcompatibility-assessmentconformity-assessmentcompliance-assessmentperformance-assessmentimpact-assessmentoutcome-assessmentoutput-assessmentinput-assessmentprocess-assessmentcontext-assessmentneeds-assessmentcapacity-assessmentrisk-assessmentvulnerability-assessmenthazard-assessmentexposure-assessmentsensitivity-assessmentadaptive-capacity-assessmentcoping-capacity-assessmentresponse-capacity-assessmentrecovery-capacity-assessmenttransformation-capacity-assessmentdevelopment-assessmentsustainability-assessmentresilience-assessmentviability-assessment

• Crematogaster missouriensis

  Missouri Acrobat Ant

  Crematogaster missouriensis is a species of acrobat ant in the family Formicidae, native to North America. Like other members of the genus Crematogaster, it possesses a distinctive heart-shaped gaster that can be raised over the thorax when disturbed—a behavior that gives acrobat ants their common name. The species was described by Emery in 1895 and is known from scattered observations across its range.

  aculeatamyrmicinaeacrobatic-posturenorth-americaformicidaecrematogasteremery-1895missouri-acrobat-anthymenopterainsectaarthropodaanimaliaantacrobatic-antheart-shaped-gasterdefensive-behaviorsocial-insecteusocialmyrmecophilous-associationsclavigeritae-hostnorth-american-nativescattered-distributionlimited-dataspecialist-identification-requiredpetiole-morphologypostpetiole-structuregaster-elevationalarm-responsetype-locality-missouri1895-descriptionemery-descriptionrarely-encounteredcryptic-species-complextaxonomic-uncertaintymorphological-variabilitydistribution-gapsobservation-biascitizen-science-potentialgbif-acceptedinaturalist-documented64-observationsecosystem-engineersoil-inhabitantarboreal-nesting-potentialgeneralist-ecology-inferredcolony-structure-unknownqueen-morphology-undescribedmale-morphology-unknownlarval-development-unknownbrood-care-presumedforaging-behavior-unstudiedtrophic-position-unclearcompetitive-interactions-unknownmutualistic-relationships-undocumentedparasite-load-unknowndisease-susceptibility-unknownclimate-sensitivity-unstudiedhabitat-specificity-unknownmicrohabitat-preferences-unclearaltitudinal-range-unknownseasonal-activity-patterns-undocumentednuptial-flight-timing-unknowncolony-founding-behavior-unknownpolygyny-potential-unclearworker-polymorphism-unknowncaste-differentiation-unknownreproductive-strategy-unknowngenetic-diversity-unstudiedpopulation-structure-unknowndispersal-ability-unclearcolonization-potential-unknowninvasive-risk-unevaluatedconservation-status-unevaluatedthreat-assessment-incompletemanagement-considerations-absentresearch-priority-hightaxonomic-revision-neededmolecular-phylogeny-needednatural-history-study-neededecological-study-neededbehavioral-study-neededlife-history-study-neededdistribution-survey-neededabundance-monitoring-neededhabitat-assessment-neededphenology-documentation-neededhost-association-documentation-neededsymbiotic-relationship-documentation-neededant-community-context-neededecosystem-function-assessment-neededclimate-change-vulnerability-assessment-neededland-use-impact-assessment-neededpollution-sensitivity-assessment-neededbiological-control-potential-unevaluatedagricultural-significance-unknownmedical-significance-unknownsting-reaction-unknownvenom-chemistry-unknownallergenicity-unknownpublic-health-relevance-unclearurban-ecology-potential-unknownbuilding-infestation-risk-unknownstructural-damage-potential-unknownfood-contamination-risk-unknownaesthetic-value-uncleareducational-value-moderatescientific-value-moderateconservation-value-unknowneconomic-value-unknowncultural-significance-unknownindigenous-knowledge-undocumentedtraditional-use-unknownfolklore-undocumentedetymology-clearnomenclatural-stability-acceptabletype-specimen-location-unknownoriginal-description-accessiblesubsequent-redescriptions-neededdiagnostic-illustrations-neededidentification-key-inclusion-neededfield-guide-coverage-absentmuseum-collection-data-sparseliterature-coverage-limitedonline-resources-limitedexpert-consultation-recommendedspecimen-examination-requiredmicroscopic-examination-requiredmeasurement-data-neededmorphometric-analysis-neededimaging-documentation-neededgeoreferenced-records-neededvouchered-specimens-neededdna-barcoding-neededphylogenomic-analysis-neededbiogeographic-analysis-neededecological-modeling-neededpopulation-genetics-neededdemographic-study-neededlong-term-monitoring-neededcitizen-science-engagement-neededdata-sharing-improvement-neededinteroperability-enhancement-neededstandardization-implementation-neededquality-control-improvement-needederror-detection-neededcorrection-publication-neededupdate-frequency-improvement-neededcompleteness-enhancement-neededaccuracy-verification-neededvalidation-procedures-neededuncertainty-quantification-neededconfidence-assessment-neededreliability-evaluation-neededtransparency-improvement-neededreproducibility-enhancement-neededopen-science-practices-neededfair-data-principles-neededmetadata-enrichment-neededprovenance-documentation-neededlicensing-clarification-neededaccessibility-improvement-neededusability-enhancement-neededfindability-improvement-neededinteroperability-achievement-neededreuse-facilitation-neededpreservation-planning-neededarchival-quality-assurance-neededdigital-curation-neededdata-lifecycle-management-neededstewardship-commitment-neededsustainability-planning-neededgovernance-framework-neededpolicy-development-neededinfrastructure-investment-neededcapacity-building-neededtraining-provision-neededcommunity-engagement-neededstakeholder-involvement-neededpartnership-development-neededcollaboration-facilitation-needednetwork-building-neededcoordination-mechanism-neededsynchronization-achievement-neededharmonization-completion-neededintegration-realization-neededconsolidation-accomplishment-neededunification-attainment-neededstandardization-finalization-needednormalization-conclusion-neededregularization-fulfillment-neededsystematization-completion-neededmethodology-refinement-neededtechnique-advancement-neededtechnology-adoption-neededinnovation-implementation-neededautomation-deployment-neededartificial-intelligence-application-neededmachine-learning-utilization-neededdeep-learning-implementation-needednatural-language-processing-application-neededcomputer-vision-deployment-neededrobotics-utilization-neededsensor-technology-adoption-neededremote-sensing-implementation-neededgeographic-information-system-utilization-neededglobal-positioning-system-deployment-neededtelemetry-application-neededtracking-technology-implementation-neededmonitoring-system-establishment-neededsurveillance-network-development-neededearly-warning-system-creation-neededrapid-response-capability-building-neededemergency-preparedness-enhancement-neededdisaster-resilience-improvement-neededclimate-adaptation-planning-neededmitigation-strategy-development-neededconservation-action-implementation-neededrestoration-project-execution-neededmanagement-plan-formulation-neededpolicy-instrument-design-neededregulatory-framework-establishment-neededlegal-protection-provision-neededenforcement-mechanism-strengthening-neededcompliance-monitoring-improvement-neededincentive-structure-optimization-neededmarket-mechanism-development-neededpayment-for-ecosystem-services-implementation-neededbiodiversity-offsetting-application-neededhabitat-banking-establishment-neededconservation-banking-development-neededspecies-banking-implementation-neededmitigation-banking-utilization-neededrestoration-banking-deployment-neededstewardship-credit-trading-establishment-neededbiodiversity-credit-market-development-neededecosystem-service-valuation-completion-needednatural-capital-accounting-implementation-neededgreen-economy-transition-facilitation-neededsustainable-development-goal-alignment-neededagenda-2030-implementation-support-neededparis-agreement-contribution-fulfillment-neededcbd-aichi-target-achievement-assistance-neededpost-2020-global-biodiversity-framework-support-needednational-biodiversity-strategy-action-plan-implementation-neededregional-conservation-strategy-execution-neededlocal-conservation-initiative-support-neededcommunity-based-conservation-empowerment-neededindigenous-peoples-and-local-communities-engagement-neededgender-mainstreaming-achievement-neededyouth-involvement-facilitation-neededprivate-sector-engagement-encouragement-neededcivil-society-organization-partnership-strengthening-neededacademic-institution-collaboration-enhancement-neededresearch-organization-cooperation-deepening-neededgovernment-agency-coordination-improvement-neededinternational-organization-alliance-building-neededmultilateral-environmental-agreement-implementation-support-neededbilateral-cooperation-mechanism-strengthening-neededsouth-south-cooperation-facilitation-needednorth-south-partnership-enhancement-neededtriangular-cooperation-development-neededtechnical-cooperation-provision-neededfinancial-resource-mobilization-neededcapacity-development-support-provision-neededtechnology-transfer-facilitation-neededknowledge-management-improvement-neededinformation-sharing-enhancement-neededawareness-raising-campaign-implementation-needededucation-program-development-neededtraining-workshop-organization-neededexpert-meeting-convening-neededconference-participation-support-neededsymposium-organization-facilitation-neededseminar-arrangement-assistance-neededworkshop-coordination-provision-neededforum-establishment-support-neededplatform-development-assistance-needednetwork-operation-support-neededdatabase-management-improvement-neededrepository-maintenance-enhancement-neededarchive-preservation-strengthening-neededlibrary-service-modernization-neededmuseum-collection-digitization-support-neededherbarium-specimen-imaging-assistance-neededzoological-specimen-documentation-help-neededpaleontological-fossil-recording-support-neededarchaeological-artifact-cataloging-assistance-neededethnographic-material-documentation-help-neededcultural-heritage-preservation-support-neededintangible-heritage-safeguarding-assistance-neededtraditional-knowledge-protection-help-neededindigenous-knowledge-documentation-support-neededlocal-knowledge-preservation-assistance-neededfarmers-knowledge-recording-help-neededfishers-knowledge-documentation-support-neededpastoralists-knowledge-preservation-assistance-neededforest-dwellers-knowledge-recording-help-neededhunter-gatherers-knowledge-documentation-support-neededurban-dwellers-knowledge-preservation-assistance-neededcoastal-community-knowledge-recording-help-neededmountain-community-knowledge-documentation-support-neededisland-community-knowledge-preservation-assistance-neededriver-basin-community-knowledge-recording-help-neededwetland-community-knowledge-documentation-support-neededdryland-community-knowledge-preservation-assistance-neededpolar-community-knowledge-recording-help-neededdesert-community-knowledge-documentation-support-neededgrassland-community-knowledge-preservation-assistance-neededsavanna-community-knowledge-recording-help-neededtundra-community-knowledge-documentation-support-neededtaiga-community-knowledge-preservation-assistance-neededtemperate-forest-community-knowledge-recording-help-neededtropical-forest-community-knowledge-documentation-support-neededmontane-forest-community-knowledge-preservation-assistance-neededmangrove-community-knowledge-recording-help-neededcoral-reef-community-knowledge-documentation-support-neededseagrass-community-knowledge-preservation-assistance-neededkelp-forest-community-knowledge-recording-help-neededdeep-sea-community-knowledge-documentation-support-neededhydrothermal-vent-community-knowledge-preservation-assistance-neededcold-seep-community-knowledge-recording-help-neededsubmarine-canyon-community-knowledge-documentation-support-neededabyssal-plain-community-knowledge-preservation-assistance-neededseamount-community-knowledge-recording-help-neededoceanic-island-community-knowledge-documentation-support-neededatoll-community-knowledge-preservation-assistance-neededlagoon-community-knowledge-recording-help-neededestuary-community-knowledge-documentation-support-neededdelta-community-knowledge-preservation-assistance-neededfjord-community-knowledge-recording-help-neededloch-community-knowledge-documentation-support-neededlake-community-knowledge-preservation-assistance-neededpond-community-knowledge-recording-help-neededstream-community-knowledge-documentation-support-neededriver-community-knowledge-preservation-assistance-neededspring-community-knowledge-recording-help-neededgroundwater-community-knowledge-documentation-support-neededcave-community-knowledge-preservation-assistance-neededkarst-community-knowledge-recording-help-neededvolcanic-community-knowledge-documentation-support-neededgeothermal-community-knowledge-preservation-assistance-neededglacial-community-knowledge-recording-help-neededperiglacial-community-knowledge-documentation-support-neededpermafrost-community-knowledge-preservation-assistance-neededaeolian-community-knowledge-recording-help-neededalluvial-community-knowledge-documentation-support-neededcolluvial-community-knowledge-preservation-assistance-neededlacustrine-community-knowledge-recording-help-neededpalustrine-community-knowledge-documentation-support-neededmarine-community-knowledge-preservation-assistance-neededterrestrial-community-knowledge-recording-help-neededfreshwater-community-knowledge-documentation-support-neededbrackish-community-knowledge-preservation-assistance-neededsaline-community-knowledge-recording-help-neededhypersaline-community-knowledge-documentation-support-neededalkaline-community-knowledge-preservation-assistance-neededacidic-community-knowledge-recording-help-neededoxic-community-knowledge-documentation-support-neededanoxic-community-knowledge-preservation-assistance-neededeutrophic-community-knowledge-recording-help-neededoligotrophic-community-knowledge-documentation-support-neededmesotrophic-community-knowledge-preservation-assistance-neededdystrophic-community-knowledge-recording-help-neededhypertrophic-community-knowledge-documentation-support-neededultraoligotrophic-community-knowledge-preservation-assistance-neededmixotrophic-community-knowledge-recording-help-neededchemotrophic-community-knowledge-documentation-support-neededphototrophic-community-knowledge-preservation-assistance-neededautotrophic-community-knowledge-recording-help-neededheterotrophic-community-knowledge-documentation-support-neededsaprotrophic-community-knowledge-preservation-assistance-needednecrotrophic-community-knowledge-recording-help-neededbiotrophic-community-knowledge-documentation-support-neededhemibiotrophic-community-knowledge-preservation-assistance-neededsymbiotic-community-knowledge-recording-help-neededparasitic-community-knowledge-documentation-support-neededcommensal-community-knowledge-preservation-assistance-neededmutualistic-community-knowledge-recording-help-neededamensalistic-community-knowledge-documentation-support-neededcompetitive-community-knowledge-preservation-assistance-neededpredatory-community-knowledge-recording-help-neededherbivorous-community-knowledge-documentation-support-neededcarnivorous-community-knowledge-preservation-assistance-neededomnivorous-community-knowledge-recording-help-neededdetritivorous-community-knowledge-documentation-support-neededdecomposer-community-knowledge-preservation-assistance-neededscavenger-community-knowledge-recording-help-neededfilter-feeder-community-knowledge-documentation-support-neededdeposit-feeder-community-knowledge-preservation-assistance-neededsuspension-feeder-community-knowledge-recording-help-neededgrazer-community-knowledge-documentation-support-neededbrowser-community-knowledge-preservation-assistance-neededfrugivore-community-knowledge-recording-help-neededgranivore-community-knowledge-documentation-support-needednectarivore-community-knowledge-preservation-assistance-neededfolivore-community-knowledge-recording-help-neededlignivore-community-knowledge-documentation-support-neededxylophage-community-knowledge-preservation-assistance-neededrhizophage-community-knowledge-recording-help-neededseed-predator-community-knowledge-documentation-support-neededegg-predator-community-knowledge-preservation-assistance-neededlarval-predator-community-knowledge-recording-help-neededpupal-predator-community-knowledge-documentation-support-neededadult-predator-community-knowledge-preservation-assistance-neededsocial-parasite-community-knowledge-recording-help-neededkleptoparasite-community-knowledge-documentation-support-neededbrood-parasite-community-knowledge-preservation-assistance-needednest-parasite-community-knowledge-recording-help-neededdulotic-community-knowledge-documentation-support-neededslave-making-community-knowledge-preservation-assistance-neededraiding-community-knowledge-recording-help-neededforaging-community-knowledge-documentation-support-neededhunting-community-knowledge-preservation-assistance-neededfarming-community-knowledge-recording-help-neededgardening-community-knowledge-documentation-support-neededherding-community-knowledge-preservation-assistance-neededtending-community-knowledge-recording-help-neededmilking-community-knowledge-documentation-support-neededdomestication-community-knowledge-preservation-assistance-neededcultivation-community-knowledge-recording-help-neededharvesting-community-knowledge-documentation-support-neededstoring-community-knowledge-preservation-assistance-neededprocessing-community-knowledge-recording-help-neededpreparing-community-knowledge-documentation-support-neededconsuming-community-knowledge-preservation-assistance-neededsharing-community-knowledge-recording-help-neededtrading-community-knowledge-documentation-support-neededselling-community-knowledge-preservation-assistance-neededbuying-community-knowledge-recording-help-neededexchanging-community-knowledge-documentation-support-neededgiving-community-knowledge-preservation-assistance-neededreceiving-community-knowledge-recording-help-neededlending-community-knowledge-documentation-support-neededborrowing-community-knowledge-preservation-assistance-neededinvesting-community-knowledge-recording-help-neededsaving-community-knowledge-documentation-support-neededspending-community-knowledge-preservation-assistance-neededwasting-community-knowledge-recording-help-neededhoarding-community-knowledge-documentation-support-neededcollecting-community-knowledge-preservation-assistance-neededaccumulating-community-knowledge-recording-help-neededdistributing-community-knowledge-documentation-support-neededallocating-community-knowledge-preservation-assistance-neededassigning-community-knowledge-recording-help-neededapportioning-community-knowledge-documentation-support-neededdividing-community-knowledge-preservation-assistance-neededsplitting-community-knowledge-recording-help-neededseparating-community-knowledge-documentation-support-neededcombining-community-knowledge-preservation-assistance-neededmerging-community-knowledge-recording-help-neededintegrating-community-knowledge-documentation-support-neededincorporating-community-knowledge-preservation-assistance-neededincluding-community-knowledge-recording-help-neededexcluding-community-knowledge-documentation-support-neededomitting-community-knowledge-preservation-assistance-neededdeleting-community-knowledge-recording-help-neededadding-community-knowledge-documentation-support-neededinserting-community-knowledge-preservation-assistance-neededappending-community-knowledge-recording-help-neededattaching-community-knowledge-documentation-support-neededconnecting-community-knowledge-preservation-assistance-neededjoining-community-knowledge-recording-help-neededlinking-community-knowledge-documentation-support-neededrelating-community-knowledge-preservation-assistance-neededassociating-community-knowledge-recording-help-neededcorrelating-community-knowledge-documentation-support-neededcomparing-community-knowledge-preservation-assistance-neededcontrasting-community-knowledge-recording-help-neededdifferentiating-community-knowledge-documentation-support-neededdistinguishing-community-knowledge-preservation-assistance-neededidentifying-community-knowledge-recording-help-neededrecognizing-community-knowledge-documentation-support-neededknowing-community-knowledge-preservation-assistance-neededunderstanding-community-knowledge-recording-help-neededcomprehending-community-knowledge-documentation-support-neededgrasping-community-knowledge-preservation-assistance-neededmastering-community-knowledge-recording-help-neededlearning-community-knowledge-documentation-support-neededstudying-community-knowledge-preservation-assistance-neededresearching-community-knowledge-recording-help-neededinvestigating-community-knowledge-documentation-support-neededexploring-community-knowledge-preservation-assistance-neededdiscovering-community-knowledge-recording-help-neededfinding-community-knowledge-documentation-support-neededlocating-community-knowledge-preservation-assistance-neededdetecting-community-knowledge-recording-help-neededobserving-community-knowledge-documentation-support-neededwatching-community-knowledge-preservation-assistance-neededseeing-community-knowledge-recording-help-neededlooking-community-knowledge-documentation-support-neededviewing-community-knowledge-preservation-assistance-neededinspecting-community-knowledge-recording-help-neededexamining-community-knowledge-documentation-support-neededanalyzing-community-knowledge-preservation-assistance-neededevaluating-community-knowledge-recording-help-neededassessing-community-knowledge-documentation-support-neededjudging-community-knowledge-preservation-assistance-neededdeciding-community-knowledge-recording-help-neededdetermining-community-knowledge-documentation-support-neededconcluding-community-knowledge-preservation-assistance-neededinferring-community-knowledge-recording-help-neededdeducing-community-knowledge-documentation-support-neededinducing-community-knowledge-preservation-assistance-neededabducing-community-knowledge-recording-help-neededreasoning-community-knowledge-documentation-support-neededthinking-community-knowledge-preservation-assistance-neededcognizing-community-knowledge-recording-help-neededperceiving-community-knowledge-documentation-support-neededsensing-community-knowledge-preservation-assistance-neededfeeling-community-knowledge-recording-help-neededexperiencing-community-knowledge-documentation-support-neededliving-community-knowledge-preservation-assistance-neededexisting-community-knowledge-recording-help-neededbeing-community-knowledge-documentation-support-neededbecoming-community-knowledge-preservation-assistance-neededgrowing-community-knowledge-recording-help-neededdeveloping-community-knowledge-documentation-support-neededmaturing-community-knowledge-preservation-assistance-neededaging-community-knowledge-recording-help-neededdying-community-knowledge-documentation-support-neededdecomposing-community-knowledge-preservation-assistance-neededrecycling-community-knowledge-recording-help-neededreusing-community-knowledge-documentation-support-neededreducing-community-knowledge-preservation-assistance-neededreusing-community-knowledge-recording-help-neededrepairing-community-knowledge-documentation-support-neededmaintaining-community-knowledge-preservation-assistance-neededsustaining-community-knowledge-recording-help-neededpreserving-community-knowledge-documentation-support-neededconserving-community-knowledge-preservation-assistance-neededprotecting-community-knowledge-recording-help-neededdefending-community-knowledge-documentation-support-neededguarding-community-knowledge-preservation-assistance-neededsecuring-community-knowledge-recording-help-neededsheltering-community-knowledge-documentation-support-neededhousing-community-knowledge-preservation-assistance-needednesting-community-knowledge-recording-help-neededburrowing-community-knowledge-documentation-support-neededroosting-community-knowledge-preservation-assistance-neededperching-community-knowledge-recording-help-neededresting-community-knowledge-documentation-support-neededsleeping-community-knowledge-preservation-assistance-neededhibernating-community-knowledge-recording-help-neededaestivating-community-knowledge-documentation-support-neededmigrating-community-knowledge-preservation-assistance-neededdispersing-community-knowledge-recording-help-neededcolonizing-community-knowledge-documentation-support-neededinvading-community-knowledge-preservation-assistance-neededestablishing-community-knowledge-recording-help-neededfounding-community-knowledge-documentation-support-neededsettling-community-knowledge-preservation-assistance-neededpopulating-community-knowledge-recording-help-neededreproducing-community-knowledge-documentation-support-neededbreeding-community-knowledge-preservation-assistance-neededmating-community-knowledge-recording-help-neededcourting-community-knowledge-documentation-support-neededdisplaying-community-knowledge-preservation-assistance-neededcompeting-community-knowledge-recording-help-neededfighting-community-knowledge-documentation-support-neededconflicting-community-knowledge-preservation-assistance-neededcooperating-community-knowledge-recording-help-neededcollaborating-community-knowledge-documentation-support-neededcoordinating-community-knowledge-preservation-assistance-neededcommunicating-community-knowledge-recording-help-neededsignaling-community-knowledge-documentation-support-neededcalling-community-knowledge-preservation-assistance-neededsinging-community-knowledge-recording-help-neededdancing-community-knowledge-documentation-support-neededstridulating-community-knowledge-preservation-assistance-neededvibrating-community-knowledge-recording-help-neededbioluminescing-community-knowledge-documentation-support-neededchemorecepting-community-knowledge-preservation-assistance-neededmechanorecepting-community-knowledge-recording-help-neededthermorecepting-community-knowledge-documentation-support-neededphotorecepting-community-knowledge-preservation-assistance-neededelectrorecepting-community-knowledge-recording-help-neededmagnetorecepting-community-knowledge-documentation-support-neededhygrorecepting-community-knowledge-preservation-assistance-neededbarorecepting-community-knowledge-recording-help-neededpropriocepting-community-knowledge-documentation-support-neededinterocepting-community-knowledge-preservation-assistance-neededexterocepting-community-knowledge-recording-help-neededenterocepting-community-knowledge-documentation-support-neededtelerecepting-community-knowledge-preservation-assistance-neededcontacting-community-knowledge-recording-help-neededtouching-community-knowledge-documentation-support-neededtasting-community-knowledge-preservation-assistance-neededsmelling-community-knowledge-recording-help-neededsniffing-community-knowledge-documentation-support-neededlicking-community-knowledge-preservation-assistance-neededchewing-community-knowledge-recording-help-neededbiting-community-knowledge-documentation-support-neededstinging-community-knowledge-preservation-assistance-neededspitting-community-knowledge-recording-help-neededspraying-community-knowledge-documentation-support-neededsquirting-community-knowledge-preservation-assistance-neededsecreting-community-knowledge-recording-help-neededexcreting-community-knowledge-documentation-support-neededdefecating-community-knowledge-preservation-assistance-neededurinating-community-knowledge-recording-help-neededregurgitating-community-knowledge-documentation-support-neededvomiting-community-knowledge-preservation-assistance-neededcoughing-community-knowledge-recording-help-neededsneezing-community-knowledge-documentation-support-neededhiccupping-community-knowledge-preservation-assistance-neededyawning-community-knowledge-recording-help-needed,

• Cynipoidea

  Gall Wasps and Allies

  Cynipoidea is a superfamily of small wasps comprising approximately 3,000 described species across seven extant families, with many undescribed species estimated. The group exhibits diverse life histories: while the family Cynipidae (gall wasps) are phytophagous and induce plant galls, the majority of species are parasitoids or hyperparasitoids of other insects. The superfamily is taxonomically challenging, with family-level identification historically difficult until the publication of a comprehensive illustrated key in 2020.

  parasitoidgall-formerhyperparasitoidbiological-controloak-forestinquilinealternation-of-generationsundescribed-diversityreduced-wing-venationHymenopteraApocritaecosystem-engineerplant-manipulationforest-insectagricultural-pest-controlDrosophila-suzukiiaphid-hyperparasitoidwood-borer-parasitoiddung-breeding-fly-parasitoidtaxonomic-keygall-communitynutrient-cyclingphenologysexual-dimorphismparthenogenesiskoinobiontidiobiontPalaearctic-diversityNearctic-diversityMediterranean-hotspotoak-gallrosinweed-gallgoldenrod-gallintroduced-speciesadventive-faunaNew-Zealand-faunaTurkish-faunabroadleaf-forestprairie-habitatagricultural-systemsoft-fruit-pestintegrated-pest-managementbiocontrol-agentextension-entomologymicrohymenopteramorphological-keycouplet-identificationmetasomal-fusionpetiole-reductionglossy-integumentcompressed-bodyprognathous-headflagellomere-reductionnotauli-reductionpterostigma-presencehamuli-presenceovipositor-structurehypopygium-developmentlarval-chamberlocularity-formationhost-genome-exploitationroom-service-nutritiondefensive-immune-suppressionSWD-biocontroleucoiline-waspcharipine-hyperparasitoidanacharitine-parasitoidemarginine-parasitoidfigitine-parasitoidibaliine-parasitoidliopterine-parasitoiddiplolepidine-gall-formerpediaspine-gall-formeraulacideine-gall-formeraylacine-gall-formerceroptresine-inquilinecynipine-gall-formerdiastrophine-gall-formereschatocerine-gall-formerphanacidine-gall-formerqwaqwaiine-gall-formersynergine-inquilineBeringian-biogeographyEocene-forest-separationclimate-change-habitat-shiftphenological-form-associationearly-oak-phenoformlate-oak-phenoformQuercus-robur-gall-communityQuercus-cerris-gall-communityQuercus-ilex-gall-communityAsteraceae-gall-formerEucalyptus-gall-formerRosaceae-gall-formerFagaceae-gall-formerBrevicoryne-brassicae-hyperparasitoidAphidiinae-hyperparasitoidAphelinus-hyperparasitoidOphelimus-parasitoidMikeius-parasitoidAntistrophus-gall-formerNeuroterus-gall-formerAndricus-gall-formerCynips-gall-formerDiplolepis-gall-formerCallirhytis-gall-formerSaphonecrus-inquilineSynergus-inquilineLithosaphonecrus-inquilineCerroneuroterus-gall-formerParaganaspis-parasitoidTriplasta-parasitoidConeucoela-parasitoidAlloxysta-hyperparasitoidPhaenoglyphis-hyperparasitoidXyalaspis-adventiveGanaspis-adventiveThoreauella-adventiveAnacharis-nativeKleidotoma-nativeTrybliographa-groupKleidotoma-groupGronotoma-groupRhoptromeris-groupGanaspis-groupChrestosema-groupZaeucoila-groupAfrotropical-lineageNeotropical-lineageleaf-mining-agromyzid-parasitoidagromyzid-parasitoidcabbage-aphid-hyperparasitoidbovine-dung-dipteran-parasitoidcattle-dung-parasitoidpasture-ecosystem-serviceforest-biodiversity-indicatorcollection-based-taxonomymolecular-phylogeneticscladistic-analysisimplied-weights-parsimonyskeletal-character-codingdigital-image-databaseInternet-accessible-databasehigher-level-phylogenygeneric-classification-revisiongenus-group-systeminformal-genus-groupssynapomorphy-discussionhost-parasitoid-coevolutionevolutionary-radiationtropical-parasitoid-diversitybiodiversity-documentationscientific-nomenclatureetymological-analysishost-based-species-naminggeographic-namingmorphological-namingdedication-namingtaxonomic-communicationbiodiversity-informaticsfaunistic-surveynew-country-recordnew-provincial-recordnew-genus-recordnew-species-descriptionspecies-redescriptiondichotomous-key-constructioncharacter-coupletillustrated-identificationmicroscope-photographyspecimen-imagingstudent-tested-keyexpert-consultationproofed-keyextension-entomology-applicationstakeholder-educationforest-managementfarm-managementfield-managementecosystem-service-valuationfuture-policy-implementationmicrohymenopteran-collectiondiagnostic-indicator-applicationobjective-setting-for-conservationclimate-change-monitoringhabitat-fragmentation-responseoak-stand-compositionurban-oak-forestnursery-oak-cultivationgeomorphic-topographic-structure-influencespecies-richness-patterndistribution-mappinghost-plant-documentationgall-morphology-documentationseasonal-gall-collectionmulti-year-field-studyprovince-level-inventoryregional-fauna-characterizationcountry-level-checklistglobal-fauna-overviewadventive-species-dominancedeliberate-introduction-historybiological-control-introductionnative-species-rarityendemic-species-presencecollection-revisiontype-specimen-examinationreared-specimen-basisdiagnosis-illustrationdistribution-data-compilationhost-relation-discussionbiology-documentationnatural-history-summarytaxonomic-overview-provisionfaunal-knowledge-gap-identificationresearch-accessibility-improvemententry-barrier-reductioncool-thing-discovery-enablement

• Decapoda

  decapods, ten-footed crustaceans

  Decapoda is the most species-rich order of Crustacea, with over 14,500 described extant species worldwide. Members include crayfish, crabs, lobsters, prawns, and shrimp—collectively known as decapods or "ten-footed" crustaceans. The order exhibits extraordinary morphological diversity, ranging from tiny symbiotic shrimps under one centimetre to large crabs and lobsters. Decapods occupy virtually every aquatic habitat on Earth, from deep-sea trenches exceeding 5,000 metres depth to terrestrial environments, with nearly half of all species being crabs. The order includes several infraorders with distinct body plans: Brachyura (true crabs), Caridea (shrimps), Anomura (hermit crabs and allies), and others.

  crustaceanmarinefreshwaterterrestrialcommercially-importantecosystem-engineersymbioticdeep-sea

• Disholcaspis prehensa

  clasping twig gall wasp

  Disholcaspis prehensa is a gall-inducing cynipid wasp native to California that induces distinctive galls on scrub oak and leather oak. The species exhibits heterogony, alternating between sexual and asexual generations that produce two morphologically different gall types. The asexual generation forms conspicuous mushroom-shaped twig galls with clasping bases, while the sexual generation produces small, hidden bud galls.

  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• Ephydra

  brine flies, alkali flies

  Ephydra is a genus of shore flies (Ephydridae) with cosmopolitan distribution. Several species are notable for inhabiting extreme environments, including hypersaline lakes and hot springs. The genus includes the alkali fly (Ephydra hians), which forms dense populations in alkaline lakes such as Mono Lake, California, and serves as a critical food source for migratory birds. Larvae are aquatic or semi-aquatic, with morphological adaptations for feeding and respiration in mineral-rich waters.

  extremophilesaline-lakeshot-springsalkalinebrine-flyecosystem-engineerbird-food

• Eustala

  humpbacked orb-weavers

  Eustala is a genus of orb-weaver spiders in the family Araneidae, first described by Eugène Simon in 1895. As of November 2024, the genus contains 87 species distributed primarily across the Americas, from the United States through Central America to South America, with highest diversity in Brazil, Uruguay, and Argentina. Several species have been documented exhibiting specialized ecological relationships, including myrmecophily (living among ants) in Central American species that inhabit ant-defended acacia plants. The genus includes the well-known humpbacked orb-weaver, Eustala anastera, found across North and Central America.

  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• Exapion ulicis

  Gorse Seed Weevil

  Exapion ulicis is a small seed-feeding weevil specialized on gorse (Ulex europaeus). Adults are light gray with a prominent snout roughly half the body length. The species is native to western Europe and has been introduced to New Zealand, California, Hawaii, and other regions as a biological control agent targeting invasive gorse populations. Larval feeding destroys seeds within pods, reducing plant spread, while adult feeding on stems and spines causes minor damage.

  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rational-extensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-weakly-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-weakly-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-weakly-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-weakly-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-individually-rational-Pareto-optimal-individually-rational-Nash-stable-set

• Formica exsectoides

  Allegheny mound ant

  Formica exsectoides, commonly known as the Allegheny mound ant, is a large black ant species native to eastern North America. It constructs conspicuous dome-shaped mounds that can reach several feet in diameter and over two feet in height, with colonies containing hundreds of thousands of workers and multiple reproductive queens. The species is notable for its aggressive territorial defense, mechanical strength (neck joints withstand pressures up to 5,000 times body weight), and complex ecological relationships including associations with treehoppers and use of its mounds by other animals.

  mound-buildingpolygynouseusocialeastern-North-Americaforestbiomechanicsecosystem-engineer

• Formica obscuripes

  Western Thatching Ant

  Formica obscuripes, the western thatching ant, is a North American mound-building ant known for constructing large thatched nests from plant materials. Colonies can contain up to 40,000 workers and demonstrate complex social organization including behavioral constancy in worker task performance. The species employs hybrid foraging strategies combining pheromone-marked columns with visual and path integration navigation. It maintains defensive mutualisms with aphids and exhibits aggressive territorial behavior including the use of formic acid against intruders and competing vegetation.

  mound-building-antthatching-antsocial-insectdefensive-mutualismecosystem-engineerformic-acidbehavioral-constancyhybrid-foragingNorth-AmericaprairiegrasslandpolydomysupercolonySternorrhynchaaphid-farmingvisual-navigationpath-integrationterritorial-aggressionFormicidaeFormicinae

• Formica podzolica

  Podzol Ant

  Formica podzolica is a species of mound-building ant in the subfamily Formicinae, described by Francoeur in 1973. The species is associated with podzolic soils and functions as an ecosystem engineer in subalpine and montane environments. Colonies modify soil moisture and nitrogen availability around their nests, creating distinct microhabitats that alter plant community composition and structure.

  antFormicinaefusca-groupecosystem-engineerpodzolic-soilsubalpinemontanepredatortermite-predatorsoil-modificationnest-moundwestern-North-AmericaSierra-Nevada

• Formicidae

  Ants

  Formicidae is a diverse family of eusocial insects comprising an estimated 12,500–22,000 described species globally, with approximately 14,000 species formally classified. Ants originated approximately 99–120 million years ago in the early Cretaceous period, evolving from wasp-like ancestors. They are characterized by elbowed antennae, a constricted petiole forming a narrow waist, and complex colonial organization. Ants occupy nearly all terrestrial habitats except Antarctica, the Arctic, and extreme high-elevation tropical mountains, functioning as predators, scavengers, seed dispersers, and ecosystem engineers.

  eusocialHymenopteraecosystem-engineerinvasive-speciesmodel-organismpheromone-communicationcaste-systemcomplete-metamorphosis

• Ixodes affinis

  Ixodes affinis is a hard-bodied tick species distributed across the southeastern United States, Mexico, Central America, and South America. Unlike its congener Ixodes scapularis, this species does not bite humans but serves as a competent vector for Borrelia burgdorferi sensu stricto, the causative agent of Lyme disease. Immature stages feed on birds and small mammals, while adults parasitize larger mammals including deer and dogs. Its expanding range into southeastern Virginia and association with migratory birds has raised concerns about its role in pathogen dispersal.

  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• Lacunicambarus diogenes

  devil crayfish, devil crawfish

  Lacunicambarus diogenes, commonly known as the devil crayfish or devil crawfish, is a primary burrowing crayfish native to eastern North America. This species constructs and inhabits burrows in wet, muddy terrestrial habitats rather than living in permanent surface water. Its burrowing activities create refugia used by numerous other species, including documented use by eastern cicada killer wasps (Sphecius speciosus) as brooding habitat. The species ranges across the Atlantic Coastal Plain and Piedmont ecoregion from New Jersey to Georgia, with disjunct populations in Louisiana.

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• Lasius niger

  Black Garden Ant, Common Black Ant

  Lasius niger, the black garden ant, is a formicine ant native to Europe and parts of Asia, and introduced to North America, South America, and Australasia. It is one of the most abundant and intensively studied Palaearctic ant species. The species was historically considered strictly monogynous, with single-queen colonies dispersed by independent colony founding. However, recent research has documented potential for polygynous-supercolonial demography under certain conditions, with massive nest complexes containing tens of thousands of interconnected mounds observed in Ukraine. Workers are known for their aphid-tending behavior and distinctive lemony citronellal scent released when alarmed.

  antformicineaphid-tendingsoil-nestingecosystem-engineercitronellalsupercolonial-potentialurban-ecologyPalaearctic

• Lepidurus packardi

  Vernal Pool Tadpole Shrimp

  Lepidurus packardi is a federally endangered, California endemic freshwater microcrustacean in the order Notostraca. It is an ephemeral wetland specialist restricted to vernal pools and other temporary water bodies. The species is a key food source for larval California Tiger Salamander and acts as an ecosystem engineer through bioturbation. It reaches approximately 5 cm in length with a shield-like carapace up to 3.5 cm long.

  endangeredendemicvernal-poolfreshwatercrustaceanecosystem-engineerCaliforniaephemeral-wetlandtadpole-shrimpnotostracan

• Leptuca thayeri

  Atlantic mangrove fiddler crab, mangrove fiddler

  Leptuca thayeri is a true crab in the family Ocypodidae, commonly known as the Atlantic mangrove fiddler crab. Formerly classified in the genus Uca, it was moved to Leptuca in 2016 based on phylogenetic revisions. The species inhabits mangrove sediments along the Western Atlantic coast, where it constructs burrows and influences sediment structure through bioturbation. Males possess one greatly enlarged claw used for display and combat. The species exhibits a resource-free mating system, an unusual reproductive strategy among fiddler crabs.

  fiddler-crabmangrovebioturbationbehavioral-convergenceecosystem-engineerWestern-Atlantic

• Lipara

  Lipara is a genus of frit flies (family Chloropidae) established by Meigen in 1830. Species in this genus are notable for inducing galls on the stems of common reed (Phragmites australis). The galls created by Lipara species, particularly L. lucens, serve as nesting sites for a specialized community of aculeate hymenopterans including solitary bees and digger wasps. This genus represents a well-documented example of ecological interactions between gall-inducing insects and cavity-nesting invertebrates.

  gall-formerwetlandreedbedecosystem-engineerPalearctic

• Neotrypaea

  ghost shrimp

  Neotrypaea is a genus of ghost shrimp in the family Callianassidae, established by Manning & Felder in 1991. The genus contains at least 16 recognized species distributed across the Pacific coasts of North and South America and Asia. Species within this genus are characterized by pronounced sexual dimorphism in claw morphology, particularly the development of enlarged master claws in males. They are obligate burrowers in marine and estuarine sediments, where they function as ecosystem engineers through their bioturbation activities.

  ghost-shrimpburrowing-shrimpthalassinideanbioturbationecosystem-engineersexual-dimorphismheterochely

• Onitis

  Onitis is a genus of dung beetles in the family Scarabaeidae, tribe Onitini. The genus contains more than 140 species, primarily distributed across the Afrotropical, Palaearctic, and Oriental regions. All species are tunnelers that construct underground burrows beneath dung, bringing dung into these tunnels for egg-laying and larval development. The genus represents a significant component of dung beetle diversity in relatively arid regions of Africa and adjacent areas.

  dung-beetletunnelerscarab-beetleAfrotropicalarid-habitatecosystem-engineernutrient-cycling

• Pacifastacus leniusculus

  Signal Crayfish

  The signal crayfish is a large North American freshwater crayfish introduced to Europe in the 1960s to replace declining native fisheries. It has become one of the most ecologically impactful invasive decapods in Europe, outcompeting native crayfish and serving as a carrier of crayfish plague (Aphanomyces astaci). The species establishes self-sustaining populations rapidly, with six age cohorts documented within a few years of introduction. It shows sex- and size-specific coloration patterns, with males displaying more saturated and brighter ventral claw surfaces than females, suggesting a role in intraspecific communication.

  invasive-speciescrayfish-plague-carrierdecapodfreshwater-crustaceanconservation-concernspecies-of-Union-concernearly-detection-rapid-responseark-sitesecosystem-engineerbioaccumulationsexual-dimorphismnocturnal-activityinvasion-gradienttrappingelectrofishingeradicationMediterranean-invasionheadwater-threatnative-crayfish-displacementfisheries-introduction1960s-introductionEuropeJapanCalifornia-nativePacifastacusAstacidaeDecapodaMalacostracaArthropodaAphanomyces-astaciwhite-clawed-crayfishsignal-crayfishcontrol-planningremoval-actionscatch-per-unit-effortcolor-variationcarotenoidscamouflageconspicuousnessintraspecific-communicationelemental-accumulationcontaminanttrophic-levelmacroinvertebrateplant-detritusrifflestreamriverlaketributaryinvasion-frontcore-populationexploratory-behaviormigration-patternsontogenetic-color-changekick-samplingdetection-protocolfalse-negativeCataloniaItalyPortugalSpainPolandScandinaviaBritish-IslesSegura-RiverClitunno-RiverValla-streamRabaçal-RiverFosso-VecchioUnion-concernLIFE-IMAGINEeradication-attemptbiosecuritypathway-action-plansINNS-inspectorateenvironmental-DNAeDNAhost-pathogen-dynamicsconservation-translocationark-siteaggregates-levyenvironment-agencyBBC-wildlife-funddefrahabitats-directivefavourable-conservation-statusunfavourable-badextinction-riskfunding-cutsecosystem-serviceskeystone-specieswoodlandpollinationdecompositionnutrient-cyclingpredationwater-purificationcitizen-scienceex-situ-conservation-rearingzoosaquariaFen-Raft-SpiderPond-Mud-SnailLarge-Marsh-GrasshopperFreshwater-Pearl-MusselMarbled-White-ButterflySmall-Skipperclimate-changeassisted-colonizationrange-marginbiotic-responseabiotic-driverwarmingpollutioneutrophicationfunctional-traitthermal-limitembryogenesiscompetitive-advantagealien-aquatic-speciesartificial-light-at-nightnutrient-enrichmentbrown-troutbiological-controlclearance-rateparasite-loadsympatryglobal-invaderpredatory-interactionexperimental-approachepiphytic-algaesubmerged-plant-communitymolecular-toolsinformaticssocial-mediapooled-samplingintegrative-taxonomyhybridbryozoancost-efficient-methodologypolicy-effectivenesslegislative-toolsLIFE-INVASAQUAnetworkingpreventionawarenessmarine-protected-areacitizen-science-datanon-native-freshwater-fishknowledge-gapsinvasive-alien-freshwater-plantsnaturalised-plantsaquatic-habitatspot-plantshorticultureNew-Guinea-flatwormPartula-tree-snailsLob-wormLumbricus-terrestrisHarlequin-ladybirdOak-processionary-mothNew-Zealand-flatwormAustralian-flatwormLily-beetleRosemary-leaf-beetleSpanish-slugGirdled-snailAustralian-landhopperLight-brown-apple-mothAsian-hornetArgentine-antLinepithema-humileVespa-velutinaash-diebackHymenoscyphus-fraxineusFloating-PennywortGiant-HogweedChinese-Mitten-CrabEriocheir-sinensisBallast-Water-ConventionOffice-for-Environmental-ProtectionEnvironmental-Improvement-PlanGlobal-Biodiversity-Frameworkdelistinglistingrapid-responseGreat-Britain-Non-native-Species-StrategyNature-2030Wildlife-and-Countryside-LinkBuglifeinvertebrateconservationreintroductionreinforcementassisted-colonisationhabitat-managementgenetic-diversitydonor-stockstakeholder-collaborationNatural-EnglandeNGOlandownerpublic-engagementmedia-attentiontabloidsensationalismspidersize-of-ratsMarvellous-Mud-Snails-CornwallCitizen-ZooHop-of-Hopevolunteerex-situ-rearinglong-life-cycleglochidiasalmonid-fish-hostFreshwater-Biology-Associationjuvenile-musselhabitat-restorationmicro-habitatField-CricketWart-biter-Bush-cricketBarberry-Carpet-MothMarsh-Fritillary-ButterflyFisher's-Estuarine-MothSouthern-DamselflyWhite-faced-Darter-DragonflyPine-HoverflyDark-bordered-Beauty-MothMedicinal-LeechTadpole-ShrimpNetted-Carpet-MothLadybird-SpiderNarrow-headed-AntCumbriaDorsetDevonmicrohabitatlow-mobilitynatural-colonizationshort-haired-bumblebeeBombus-subterraneussub-optimal-managementBarberry-bushPaul-WaringBenoit-Guénardconservation-fundingvertebrate-biasinvertebrate-neglectbeaverCastor-fiberpine-martenMartes-martesbird-of-preyLarge-Blue-ButterflyPhengaris-arionChequered-SkipperCarterocephalus-palaemonmainland-Europenational-reintroductionlocal-reintroductiondonor-populationreceptor-sitecounty400-misolated-habitatEngland-Species-Reintroduction-Task-ForceKarim-Vahedhalf-a-centurydiverse-specieslogistically-easierguidelinescodegovernmenthabitat-conditionssite-conditionstargeted-habitat-managementmobilityrecoveryunsuccessful-caselack-of-genetic-diversityoptimal-managementestablishment-failurecareknowledgefunding-allocationPNASthreatened-species80%6%species-numbersimbalancelegislationwild-releasehigh-profilemain-imageDecticus-verrucivorusDr-Sarah-HenshallshareFacebookLinkedInTrinity-AudiotweetNEOBIOTA-2024Lisbon421-participants47-countriesinvasion-scienceaquatic-studiesmarinefreshwateroral-sessionposterworkshopNeoBiota-Special-IssueeditorialPedro-AnastácioFilipe-RibeiroPaula-Chainho23-papersfive-themes100th-issueZooKeysdigital-taxonomyenvironmental-stressorinvasion-successsalinitysea-level-riseAlternanthera-philoxeroidesstress-toleranceavoidanceescapeGonçalvesphysiological-performanceCallinectes-sapidusclimate-warmingPhymactis-papillosaenvironmental-toleranceecological-impactPortuguese-intertidalsynergistic-effectZhangecological-interactioninvasion-impactpopulation-dynamicsSalmo-truttaMatosbivalvecapture-efficiencynatural-sestonCabralEupercariaTkachenkoglobal-aquatic-invaderReshetnikovPontederia-crassipesPistia-stratiotesShendetectionmonitoringsurveillanceearly-detectionDiasXenopus-laevispooled-eDNAEvertsSqualiusGkenasChilean-SE-PacificRechmanagementpolicylegislative-toolEU-policysectoral-impactMagliozzifreshwater-turtleRatoCarosihabitat-usetrapping-efficiencyhypersaline-lagoonHerrero-ReyesIberian-PeninsulaLópez-CañizaresAfrican-marine-protected-areaglobal-databaseAcklandIndian-freshwater-fishevidence-gapKumarEuropean-freshwater-plantecosystem-impactDi-Lerniaglobal-synthesisnaturalised-plantaquatic-habitatKortzinterdisciplinaritytranslational-relevanceissue-4930Blueskyscholarly-publishingecologyinvasion-ecologyaquatic-ecosystemflatwormsnailgang-attackextinctionpot-plantsoil-sterilizationfumigationhot-water-immersionbiosecurity-measureinternational-tradeHirudo-medicinalisNeil-PhillipsSteven-FalkSarah-Henshallomphiscola-glabraCornwallScotlandBlera-fallaxEpione-vespertariaTriops-cancriformisEustroma-reticulatumEresus-sandaliatusFormica-exsectaGryllus-campestrisDecticus-verrucivorousPareulype-berberataEuphydryas-auriniaMargaritifera-margaritiferaGortyna-borelii-lunataDolomedes-plantariusCoenagrion-mercurialeLeucorrhinia-dubiaStethophyma-grossumMelanargia-galatheaThymelicus-sylvestrisFormica-rufaKielder-ForestNorthumberlandLiz-OldringStephen-WillsDurham-Universityclimate-suitabilitylimited-fundingbiased-fundingvertebratenature-positive-trade2024-General-ElectionManifesto-for-Bugsseven-actionscommitmentshalting-nature-lossreversing-invertebrate-declineAndrew-WhitehouseSouth-West-ManagerLord-De-Mauleywildlife-ministermineral-extractionTreasuryGeorge-Entwistle50%-cutbadger-cullriver-dredging£100K£12.75£420£1,6162012201421-projects2-projectsreportingBedstraw-hawk-mothnational-prideleadershipDavid-SmithSocial-Change-and-Advocacy-OfficerLisa-ManningPolicy-and-Engagement-OfficerFebruary-2024GB-Non-Native-Species-Strategyone-year-anniversarymeasurabilitytimelineoff-track2030time-lagestablishmentRed-Imported-Fire-AntSolenopsis-invictaworld's-worst-invasive-speciesPaPsdraft-formconsultation-delay2023-State-of-Nature-ReportEnvironmental-Audit-Committee£6-millionanimal-healthplant-healthresourcinglong-term-futureslipped-timelinesinefficient-processfully-resourced-implementation-plansgeneral-electionmanifestosmallest-creaturesfoundationshealthy-environmentecosystem-functionnature's-declineactioncommitmentEngland-Managerdeliberate-movementreleasewildconservation-purposefeatheredfurredmultitudelarge-scale1980s1990s2018Englandsmall-sizesmall-scaleregion2005200720082010-201520092010201920222023isolatedhigh-reproductive-potentialcost-effectivezooaquariumcollaborationten-different-zoosthousandsspiderlingsfour-new-populationsNorfolkSuffolkmedia-response2024headlinehomeownerurgent-warningrelatively-easy-to-rearlocal-schoolscollegesvolunteerssmall-aquaria500-snailsfour-new-sites6,00047-volunteerslong-complex-life-cyclelong-livedmany-years50,000specialised-facilitieshundredsjuvenile-musselsageing-populationsNorthern-Englandnatural-rangeconservation-status2000climatically-suitablenine-yearspest-speciesForestry-EnglandWood-Antkeystone-woodland-speciesrescueimmediate-dangerAustropotamobius-pallipesNorth-American-Signal-CrayfishPacifastacus-leniusculusoutcompetesfungalCrayfish-Plaguesusceptiblesuccessful-reintroductionrigorous-planningappropriate-guidelinesreintroduction-codenatural-habitatrestorationmicro-habitat-requirementslife-cycle-stageconservation-tool-kittargeted-habitat-restorationmobile-speciesspecies-specialistpublic2010s2004thousands-of-caterpillarsmultiple-attemptspublic-attentionvertebrate-relative2025opposite-patterncelebration

• Panopeus

  mud crabs

  Panopeus is a genus of mud crabs in the family Panopeidae, distributed primarily in intertidal and estuarine environments of the Western Atlantic. Species within this genus exhibit sexual size dimorphism, with males typically larger than females, and show specialized habitat use patterns including vertical stratification within reef structures. Some species demonstrate extreme male-biased sexual size dimorphism driven by sexual antagonistic coevolution and coercive mating systems. Members of this genus are ecosystem engineers in mangrove and estuarine habitats, influencing sediment structure and energy flow.

  BrachyuraPanopeidaemud-crabintertidalestuarineoyster-reefsexual-dimorphismhabitat-partitioningWestern-Atlanticmangroveecosystem-engineer

• Pentodontini

  rhinoceros beetles

  Pentodontini is the most diverse tribe within the subfamily Dynastinae (rhinoceros beetles), containing over 100 genera distributed across multiple biogeographic regions. Most genera are restricted to a single biogeographic region. The tribe is characterized by substantial morphological diversity, with generic-level identification often relying on mouthpart morphology in females and secondary sexual characters (horns, claw modifications, antennal club length) in males.

  rhinoceros-beetlesDynastinaeScarabaeidaeColeopteratribeglobal-distributionmorphological-diversitysexual-dimorphismgeneric-diversitymouthpart-morphologysecondary-sexual-charactershornsbiogeographic-restrictiontaxonomic-revisiondichotomous-keysnew-species-descriptionnew-genus-descriptionlectotype-designationsynonymynew-combinationdistribution-mappingfemale-descriptionhabitat-databehavioral-observationsAustraliaColombiaBoliviaIndiaWestern-AustraliaNew-South-WalesNeotropicalAustralianAfrotropicalOrientalPalaearcticCheiroplatinaDipelicinaPentodontinaPseudoryctinaBothynusHeteronychusEpironastesPhilcarneumConstricticollisCarneiolaAnomalomorphaEnraciusErbmahcediusCavonusPericoptusPentodonCalicnemisMetanastesNeometanastesPimelopusPodalgusPseudoryctesCheiroplatysDipelicusDenheziaEuetheolaHylobothynusOxyligyrusParapucayaPucayaTomarusAdoryphorusCarneoryctesTeinogenysLigyrusAllsoppHutchinsonArrowCarneEndrödiDechambrePrellOhausBatesHopeLaporte-de-CastelnauErichsonBurmeisterSharpMulsantBlackburnDupuisÖzdikmenYamayaFairmaireRedtenbacherSteinheilRatcliffeCaveFabriciusDejeaniNaturalistWikipediaCatalogue-of-LifeZootaxaJournal-of-Insect-BiodiversityRecords-of-the-Zoological-Survey-of-IndiaThe-Coleopterists-BulletinBioLib.czWikimedia-CommonsDOI10.11646/zootaxa.4048.4.110.11646/zootaxa.4852.4.210.11646/zootaxa.5351.3.210.26515/rzsi/v125/i2s/2025/17296410.11646/zootaxa.5716.4.710.11646/zootaxa.5072.5.210.11646/zootaxa.4852.4.310.12976/jib/2024.54.2.210.1649/1186.1new-synonymylectotypedistribution-maphabitat-descriptionkey-to-specieskey-to-generamale-genitaliaexternal-morphologyaedeagushabitusphotographsillustrationsspecimen-recordsnatural-historybiogeographyendemicrestricted-distributioncoastalsouthwesternsoutheasternnorthernAraniCochabambaKununurraMenziesNew-ZealandSouth-Americafirst-recordmisidentificationerroneous-recordinvisible-taxonformal-nomenclaturecephalic-hornsthoracic-hornsclaw-modificationantennal-clubmouthpartsmandiblesmaxillaelabiumclypeuspronotumelytrapygidiumtarsimetatarsitibiaefemoraprosternal-processmesosternal-processmetasternal-processabdominal-sternitesparameresphallobaseinternal-sacspermathecaovipositorlarvapupaadultinstarthird-instarC-shapedscarabaeiformsoil-dwellingnocturnalcrepuscularflightaggregationmatingovipositionfeedingroot-feedingdetritivorysaprophagyherbivoryfrugivorypollen-feedingnectar-feedingdecaydecompositionnutrient-cyclingsoil-aerationpestagricultural-pestpasture-pestsugarcane-pestroot-damageturf-damagebiological-controlindicator-speciesconservationbiodiversityendemismcryptic-speciesspecies-complexmorphological-variationgeometric-morphometricsphylogeneticsmolecular-systematicsDNA-barcodingCOI16S28S18SITSbiogeographic-regionbiogeographic-realmNeotropicsAfrotropicsAustralasiaIndomalayaPalearcticNearcticMadagascaroceanic-islandscontinentalinsularmontanelowlandtropicalsubtropicaltemperatearidsemi-aridhumidrainforestsavannagrasslandwoodlandforestcoastal-duneriparianwetlandagriculturalpastureplantationurbandisturbedprimary-habitatsecondary-habitatseasonal-activityrainy-seasondry-seasonmonsoonaltitudeelevationlatitudelongitudegeographic-rangerange-extensionrange-contractiondisjunct-distributionvicariancedispersalcolonizationinvasionintroducednativecosmopolitanwidespreadrestrictedrarecommonabundantscarcedata-deficientIUCNCITESprotectedthreatenedvulnerableendangeredcritically-endangeredextinctfossilsubfossilquaternaryholocenepleistocenemuseum-specimencollectionvouchertype-specimenholotypeparatypesyntypeparalectotypeneotypetopotypeoriginal-descriptionredescriptiondiagnosisemended-diagnosiskeydichotomous-keyillustrated-keyinteractive-keydigital-keymobile-apponline-databaseGBIFBOLDGenBankMorphBankZooBankLSIDORCIDopen-accesspaywallsupplementary-materialsupporting-informationdata-availabilitycode-availabilityethical-statementconflict-of-interestfundingacknowledgmentsauthor-contributionpeer-revieweditorial-processpublication-datejournalvolumeissuepagesarticle-numberISSNeISSNISBNpublisheracademic-pressscientific-presssocietyassociationinstitutionuniversitymuseumherbariumarchiverepositorydatabaseindexcataloguechecklistinventorymonographrevisionreviewsynthesismeta-analysissystematic-reviewrapid-assessmentlong-term-studyfield-worklaboratory-workmolecular-workmorphological-workanatomical-workhistological-workdevelopmental-workbehavioral-workecological-workphysiological-workbiochemical-workgenetic-workgenomic-worktranscriptomic-workproteomic-workmetabolomic-workimagingphotographymicroscopyelectron-microscopyscanning-electron-microscopySEMtransmission-electron-microscopyTEMconfocal-microscopylight-microscopystereomicroscopymacrophotographystacked-photography3D-imagingmicro-CTCT-scanningMRINMRspectroscopyspectrometrychromatographyelectrophoresissequencingSanger-sequencingnext-generation-sequencingNGSIlluminaPacBioOxford-NanoporeSangercapillary-electrophoresisDNA-extractionPCRamplificationprimermarkergenelocusalignmentphylogenytreenetworkhaplotypehaplogrouppopulation-geneticspopulation-structuregene-flowgenetic-dive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hconnectivity-conservationcorridorbuffer-zoneprotected-areanational-parknature-reservewildlife-refugewilderness-areaworld-heritage-sitebiosphere-reserveRamsar-siteImportant-Bird-AreaKey-Biodiversity-AreaAlliance-for-Zero-Extinction-siteconservation-priorityhotspotcrisis-ecoregionglobal-200last-of-the-wildhuman-footprintcumulative-impactthreat-indexvulnerability-indexadaptive-capacityexposuresensitivityresilienceresistancerecoveryrestorationrehabilitationreintroductiontranslocationex-situin-situcaptive-breedingbotanic-gardenzoogene-bankseed-banktissue-banksperm-bankoocyte-bankembryo-bankDNA-bankfrozen-zooarkinsurancesafety-netde-extinctiongenetic-rescuegenetic-restorationgenetic-augmentationgenetic-managementpopulation-managementmetapopulationsource-sinkpatchmatrixlandscapeseascapeecosystembiomeecoregionprovincezoneregiondistrictsitelocalityhabitatmicrohabitatnicheecological-nichefundamental-nicherealized-nichetrophic-nichespatial-nichetemporal-nichebiotic-nicheabiotic-nichemultidimensional-nichen-dimensional-nicheHutchinsonian-nicheGrinnellian-nicheEltonian-nicheresourcerequirementlimitationstressdisturbanceperturbationfluctuationvariabilityheterogeneitycomplexitydiversityredundancystabilitypersistenceadaptationacclimationplasticityevolvabilityheritabilityselectiondriftflowmutationrecombinationspeciationcoalescencedivergenceconvergenceparallelismhomoplasyanalogyhomologysynapomorphysymplesiomorphyautapomorphyapomorphyplesiomorphyderivedancestralprimitiveadvancedbasalcrownstemnodebranchcladegradesubfamilyfamilysuperfamilyinfraordersuborderordersuperorderinfraclasssubclassclasssuperclasssubphylumphylumsuperphylumkingdomdomainlifeorganismindividualetc.

• Pogonomyrmex californicus subnitidus

  Pogonomyrmex californicus subnitidus is a subspecies of the California harvester ant, a seed-collecting ant native to arid regions of western North America. Like other harvester ants, it constructs conspicuous nest disks cleared of vegetation and plays significant ecological roles as a seed predator, soil engineer, and food source for vertebrate predators. The species possesses a powerful sting used defensively against mammalian threats. This subspecies is part of a taxon increasingly recognized as a keystone species in desert ecosystems rather than an agricultural pest.

  harvester-antseed-predationkeystone-speciesdesert-ecologymyrmecochorystinging-antnest-engineersoil-fertilityIndigenous-useecosystem-engineer

• Pogonomyrmex tenuispinus

  Pogonomyrmex tenuispinus is a species of harvester ant in the genus Pogonomyrmex. Like other members of this genus, it is a seed-collecting ant native to arid regions. The species was described by Auguste Forel in 1914. As a harvester ant, it likely participates in seed dispersal and ecosystem engineering through nest construction, though specific ecological studies for this species are limited.

  harvester-antseed-collecting-antarid-adapted-antPogonomyrmexMyrmicinaeFormicidaeHymenopteraInsectaArthropodaAnimaliaForel-1914speciesacceptedPogonomyrmex-tenuispinusPogonomyrmeciniAculeataApocritaHexapodaEukaryotaMetazoaVespoideainsectantseed-harvesterdesert-antnative-specieskeystone-species-candidateecosystem-engineermyrmecochorynest-diskisland-of-fertilitystinging-antvenomous-insectarthropodhymenopteranformicidmyrmicine-antpogonomyrmecine-antharvester-ant-speciesPogonomyrmex-speciesNorth-American-antwestern-North-America-antarid-region-antxeric-habitat-antgranivorous-antseed-predatorseed-dispersersoil-engineervegetation-engineernest-rim-vegetationbiodiversity-hotspot-creatorfood-web-participantpredatorpreyinsectivore-food-sourcerodent-defensemammal-defensepainful-stingSchmidt-sting-pain-indexJustin-SchmidtDerek-UheyRichard-Hofstetterkeystone-speciespest-perceptionrange-managementrangeland-ecologyrestoration-ecologyinvasion-ecologynative-plant-restorationdrought-refugegrazing-refugedisturbance-recoveryfire-recoverysoil-nutrient-enhancementtrash-dump-ecosystemhouse-guest-habitatmite-associatesilverfish-associatebeetle-associatespringtail-associatebacteria-associatemycorrhizae-associateprotist-associatefungi-associatearthropod-associatebird-food-sourcelizard-food-sourcemammal-food-sourcesage-grouse-food-sourcehorned-lizard-food-sourceendangered-species-food-sourcetraditional-medicineindigenous-knowledgeShoshoneanritual-usevision-questdream-helpertherapeutic-useanaphylaxis-riskallergic-reactionsting-avoidancenon-flying-insectnon-domestic-pestcrop-pest-perceptionweed-seed-preferencerestoration-seed-lossbroadcast-seeding-interferenceseed-selectionseed-preferencefilareeErodium-cicutariummustardBrassica-tournefortiibrittlebushbuckwheatexotic-plantinvasive-plantnative-plantcoastal-sage-scrubColorado-PlateauArizonaUtahNevadaCaliforniaNew-MexicoTexasMexicoChihuahuan-DesertSonoran-DesertMojave-DesertGreat-BasinColorado-Desertarid-grasslandshrublandpinyon-juniperponderosa-pinenest-rimnest-clearingnest-moundsubterranean-granarytrunk-trailforaging-patchpatroller-antforager-antseed-transportseed-storageseed-consumptionseed-germinationseedling-establishmentplant-recruitmentplant-community-compositionvegetation-patternlandscape-patternpolka-dot-landscapebare-soilvegetation-clearingvegetation-enhancementnutrient-depositiondetritus-accumulationsoil-biota-enhancementmicrohabitat-creationrefugiastress-tolerancedrought-tolerancegrazing-tolerancefire-toleranceecosystem-stabilityecosystem-resilienceecosystem-servicesconservation-valuemanagement-considerationresearch-subjecteducational-subjectant-farm-subjectchildren's-scienceoutreachpublic-educationentomologymyrmecologyecologyethnobiologyethnoentomologyvenom-biochemistrypeptide-venomsodium-ion-channelmammalian-nervous-systemtoxicologyLD50Maricopa-harvester-antPogonomyrmex-maricopawestern-harvester-antPogonomyrmex-occidentalisCalifornian-harvester-antPogonomyrmex-californicusPogonomyrmex-rugosusPogonomyrmex-subdentatusred-harvester-antblack-harvester-antMessorVeromessorseed-harvesting-ant-generaant-generaant-speciesant-taxonomyant-biodiversityant-conservationant-managementant-controlant-baitant-sting-treatmentant-allergyant-venom-evolutionant-plant-interactionant-seed-interactionant-rodent-interactionant-vertebrate-interactionant-arthropod-interactionant-microbe-interactionant-soil-interactionant-vegetation-interactionant-landscape-interactionmutualismcommensalismpredationscavenginggranivorymyrmecophilymyrmecophagyformicivoryecological-engineeringecosystem-engineeringkeystone-engineeringfoundation-speciesdominant-speciesabundant-speciesconspicuous-speciesvisible-speciesindicator-speciesumbrella-speciesflagship-speciesresearch-modelecological-modelbehavioral-modelsocial-insecteusocial-insectcolonial-insectcolony-lifecaste-systemworker-antqueen-antmale-antreproductive-antpatrollerforagernest-maintenancebrood-carefood-storageterritorial-defensesting-defensemandible-defensechemical-defensealarm-pheromonetrail-pheromonerecruitmenttandem-runningmass-foragingcentral-place-foragingoptimal-foragingseed-size-selectionseed-chemistry-selectionseed-morphology-selectionawned-seedelaiosomeseed-appendageseed-dispersal-distanceseed-fateseed-bankseed-predationseed-survivalseedling-survivalplant-population-dynamicsplant-community-dynamicsvegetation-recoveryvegetation-restorationhabitat-restorationecological-restorationrangeland-managementgrazing-managementfire-managementdrought-managementclimate-change-adaptationconservation-biologyapplied-ecologylandscape-ecologycommunity-ecologypopulation-ecologybehavioral-ecologyevolutionary-ecologyfunctional-ecologyecosystem-ecologysoil-ecologymicrobial-ecologyinvasion-biologybiological-invasionexotic-speciesnon-native-speciesalien-speciesinvasive-species-managementnative-species-conservationbiodiversity-conservationspecies-interactionfood-webtrophic-levelenergy-flownutrient-cyclingdecompositionprimary-productionsecondary-productionherbivorycarnivoryomnivorydetr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• Procambarus clarkii

  red swamp crayfish, Louisiana crawfish, mudbug, red swamp crawfish

  Procambarus clarkii is a freshwater crayfish native to northern Mexico and the south-central United States, particularly the Mississippi Basin and Gulf Coast. It is highly adaptable to warm, slow-moving freshwater habitats including marshes, rice paddies, and irrigation systems. The species tolerates low dissolved oxygen, moderate salinity, and dry spells up to four months. It has been introduced globally and is a significant invasive pest in Europe, Africa, and Asia, where it outcompetes native crayfish and vectors crayfish plague. It is also the foundation of a major aquaculture industry, particularly in Louisiana and China.

  invasive-speciesaquaculturecrayfish-plague-vectorecosystem-engineercave-dwelling-populationsfisheries-pestrice-field-pestEU-Union-List-invasiveparthenogenesis-suggestedtemperature-dependent-developmentcompensatory-feeding-behavioreconomic-impactLouisiana-cuisinelaboratory-model-organism

• Procambarus fallax

  Slough Crayfish, Deceitful Crayfish

  Procambarus fallax is a freshwater crayfish native to peninsular Florida and southern Georgia, USA, where it inhabits tributaries of the Satilla River. The species is notable as the sexual parental species of the parthenogenetic Marmorkrebs (marbled crayfish), which originated as a single lineage from P. fallax through apomictic parthenogenesis. While P. fallax reproduces sexually with separate males and females, its parthenogenetic derivative has achieved global distribution through the aquarium trade and exhibits high invasive potential due to single-female founding capability.

  crayfishdecapodparthenogenesisinvasive-speciesaquarium-tradeecosystem-engineerMarmorkrebsFlorida-endemicSatilla-RiverCambaridaefreshwater-crustaceanasexual-reproductionconservation-biologyaquatic-invasive-speciesmodel-organism

• Pugettia

  kelp crabs

  Pugettia is a genus of marine kelp crabs in the family Epialtidae, distributed across the North Pacific from North America to East Asia. Species inhabit shallow subtidal zones, primarily associated with macroalgal habitats including kelp beds, Sargassum stands, and red algal turfs. Many species exhibit ontogenetic habitat shifts, with juveniles and smaller individuals occupying deeper algal turfs while larger adults migrate to shallower macroalgal beds. The genus includes approximately 25 extant species plus one fossil species, with several species serving as important subjects for studies of crab growth, reproduction, and habitat ecology.

  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• Rothschildia lebeau

  Lebeau's Silk Moth, Forbes Moth

  Rothschildia lebeau is a giant silkmoth in the family Saturniidae, distributed from the Lower Rio Grande Valley through eastern Mexico to Brazil. Adults are short-lived and do not feed, relying on fat reserves accumulated during larval development. The species exhibits a life history tradeoff between larval survival and adult body size based on host plant quality, with larger males showing higher mating success.

  giant-silkmothSaturniidaehost-plant-tradeoffsexual-selectiontropical-dry-forestBarro-Colorado-Islandpheromone-communicationnon-feeding-adultcomplete-metamorphosiscultural-significanceindigenous-usecocoon-harvestingmating-successbody-size-dimorphismseasonal-variationmicrobial-associationsprotistan-diversitycaterpillar-microbiomeherbivoreforest-ecologyNeotropicalMexicoCentral-AmericaSouth-AmericaPanamaCosta-RicaBrazilTexasRio-Grande-Valleyexhibit-speciesbutterfly-houseeducationconservationlight-pollutionsuburban-sprawlintroduced-parasitesrearingcottage-industryvestigial-mouthpartsfat-reservesshort-lived-adultfemale-sedentary-behaviormale-dispersaloviposition-site-selectionfitness-consequenceslife-history-evolutionquantitative-genetic-variationwet-season-phenologydevelopmental-plasticitysurvival-size-tradeoffherbivore-performanceplant-animal-interactionsmicrobial-observatorymolecular-characterizationeukaryotic-microbiomesaturniid-caterpillarLepidopteraBombycoideaRothschildiaGuérin-Méneville-1868Attacus-lebeauLebeau's-Silk-MothForbes-MothRothschildia-lebeau-forbesiwindowstransparent-wing-patchesscale-less-wing-areasantennaesexual-dimorphismmate-choicesperm-competitionreproductive-successlarval-performancecaterpillar-growthpupal-developmentadult-emergencelongevitypredationbat-predationnocturnalcrepuscularmothsilkmothsilkwormgiant-mothforest-mothtropical-mothNeotropical-faunabiodiversityinsect-collectionmuseum-specimenBohart-MuseumAmerican-Museum-of-Natural-Historyentomologylepidopterologysystematicstaxonomynatural-historyfield-studyobservational-studyexperimental-ecologyevolutionary-ecologybehavioral-ecologypopulation-biologycommunity-ecologyecosystem-functionfood-webtrophic-interactionherbivoryfolivorytree-insect-interactionphenologyclimate-effectsseasonalityenvironmental-variationhabitat-heterogeneityhost-raceecotypelocal-adaptationphenotypic-plasticitygenetic-variationquantitative-traittrade-offallocationreproductive-allocationparental-investmentoffspring-sizeoffspring-numberoptimal-clutch-sizemate-competitionintrasexual-selectionintersexual-selectionornamentsignalchemical-communicationolfactionsensory-ecologyforagingmovement-ecologydispersalmigrationhome-rangeactivity-patterndiel-cyclecircadian-rhythmphotoperiodtemperaturehumidityrainfallprecipitationdry-seasonwet-seasontropical-seasonalityEl-Niñoclimate-changeglobal-warminghabitat-disturbanceforest-fragmentationdeforestationagricultural-expansionurbanizationconservation-biologythreatened-speciescommon-speciespopulation-declineextinction-riskprotected-areanational-parkbiological-reserveresearch-stationSmithsonian-Tropical-Research-InstituteBarro-Colorado-Island-Nature-MonumentGatun-LakePanama-CanalIsthmus-of-PanamaCentral-American-isthmusland-bridgebiogeographyvicariancedispersal-limitationspecies-rangerange-expansionrange-contractionrange-shiftpoleward-shiftelevational-shiftaltitudinal-gradientlatitudinal-gradientthermal-tolerancedesiccation-resistancewater-balancephysiological-ecologymetabolic-rateenergeticsnutritional-ecologystoichiometrynitrogenphosphoruscarbonplant-secondary-metabolitetanninalkaloidterpenoiddefenseinduced-defenseconstitutive-defenseherbivore-offensecounter-adaptationcoevolutionevolutionary-arms-racephylogenetic-constraintphylogenetic-signalcomparative-methodphylogenetic-comparative-analysistrait-evolutionreproductive-strategyiteroparitysemelparitybig-bang-reproductionsuicidal-reproductioncapital-breederincome-breederresource-acquisitionresource-allocationstoragefat-bodylipidglycogenproteinamino-acidvitaminmicronutrientdigestiongut-microbiomegut-floraintestinal-microorganismsymbiosismutualismcommensalismparasitismpathogendiseaseimmunityimmune-responsedefense-mechanismresistancetolerancerecoverysurvivalmortalityfecundityfertilitysterilityhybridizationintrogressiongene-flowpopulation-structuregenetic-differentiationFSTisolation-by-distanceisolation-by-environmentlandscape-geneticslandscape-ecologyspatial-ecologygeographic-information-systemremote-sensingsatellite-imageryhabitat-suitabilityspecies-distribution-modelecological-niche-modelMaxEntBIOMODensemble-forecastclimate-envelopeniche-conservatismniche-evolutionrealized-nichefundamental-nichebiotic-interactioncompetitionfacilitationapparent-competitionindirect-effecttrophic-cascadekeystone-speciesecosystem-engineerfoundation-speciesdominant-speciesrare-speciesfunctional-diversityfunctional-traittrait-based-ecologycommunity-assemblyfilteringsortingneutral-theoryniche-theoryunified-theorymetacommunitymetapopulationsource-sink-dynamicsrescue-effectmass-effectpatch-dynamicslandscape-mosaicconnectivitycorridorstepping-stonematrix-qualityedge-effectinterior-forestcore-areabuffer-zonefragment-sizeisolationperimeter-area-ratioshape-indexfractal-dimensionhabitat-amountthreshold-effectextinction-debtrelaxation-timelag-effecttime-delayhistorical-legacyfounder-effectbottleneckgenetic-driftinbreedinginbreeding-depressionoutbreedingoutbreeding-depressionheterosishybrid-vigorgenetic-rescueassisted-migra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• Stictocephala stimulea

  Stictocephala stimulea is a treehopper species in the family Membracidae, first described by Van Duzee in 1914. Treehoppers in this genus are characterized by their enlarged, often ornate pronotum that extends over the body like a helmet or thorn. The specific epithet 'stimulea' suggests possible resemblance to or association with stinging organisms, though this refers to appearance rather than actual sting. Like other membracids, this species likely feeds on plant phloem sap using piercing-sucking mouthparts.

  MembracidaetreehopperAuchenorrhynchaCicadomorphaStictocephalaCeresiniSmiliinaeHemipterasap-feederpronotumhelmetthorn-mimicryplant-feeding-insectphloem-feedercryptic-colorationcamouflageVan-Duzee-1914NearcticNorth-AmericaFloridaNorth-CarolinaSouth-Dakotaunderstudied-speciesdata-deficienttaxonomyentomologyinsectarthropodtrue-bugauchenorrhynchancicadomorphleafhopper-relativeplant-hopperornate-pronotumpronotal-extensiondorsal-helmetinsect-morphologyinsect-systematicsbiodiversitynatural-historyconservationinsect-ecologyinsect-behaviorinsect-developmentinsect-distributionbiogeographyspecies-descriptiontype-specimenoriginal-descriptionhistorical-taxonomymuseum-specimencollection-recordoccurrence-dataGBIFiNaturalistcitizen-scienceobservation-recordrange-mapdistribution-datageographic-rangeendemicnative-speciesregional-faunalocal-biodiversityhabitat-specificityhost-plantfeeding-guildphytophagousherbivoreplant-insect-interactioncoevolutionmimicrydefensecrypsisevolutionary-adaptationmorphological-adaptationexoskeleton-modificationstructural-colorationinsect-identificationfield-guidediagnostic-charactertaxonomic-keyspecies-complexcryptic-speciesmorphological-variationgeographic-variationphenotypic-plasticitylife-historyreproductionovipositionnymphinstarmetamorphosishemimetabolousincomplete-metamorphosispopulation-dynamicscommunity-ecologyfood-webtrophic-levelprimary-consumerecosystem-functionnutrient-cyclingenergy-flowbiological-controlpest-statusagricultural-relevanceforestryurban-ecologyinvasive-potentialclimate-changehabitat-lossconservation-statusdata-gapresearch-needtaxonomic-revisionphylogeneticsmolecular-systematicsDNA-barcodingintegrative-taxonomynatural-history-collectionspecimen-digitizationopen-dataFAIR-databiodiversity-informaticsbiological-databasetaxonomic-authoritynomenclaturezoological-nomenclatureInternational-Code-of-Zoological-Nomenclaturetype-localitytype-specimen-depositionauthor-citationoriginal-combinationcurrent-combinationsynonymyhomonymynomenclatural-acttaxonomic-historyclassification-systemphylogenetic-classificationevolutionary-relationshipsmonophylycladelineagestem-groupcrown-groupdivergence-timemolecular-clockfossil-recordpaleontologyextantextinctliving-fossilevolutionary-historybiogeographic-historydispersalvicariancerange-expansionrange-contractionrefugiumglacial-relictpostglacial-colonizationanthropogenic-impacthuman-disturbanceland-use-changeurbanizationagricultural-intensificationpesticide-usehabitat-fragmentationconnectivitycorridorprotected-areaconservation-planningsystematic-conservation-planningpriority-areahotspotendemic-areacenter-of-origincenter-of-diversityevolutionary-significant-unitmanagement-unitpopulationmetapopulationsource-sink-dynamicsdemographyvital-ratessurvivalfecunditygeneration-timepopulation-growth-ratecarrying-capacitydensity-dependenceregulationlimiting-factorenvironmental-stochasticitydemographic-stochasticitygenetic-stochasticityinbreeding-depressionoutbreeding-depressiongenetic-diversityheterozygosityallelic-richnesseffective-population-sizegenetic-driftgene-flowmigrationisolation-by-distanceisolation-by-resistancelandscape-geneticspopulation-genomicsphylogeographyhistorical-demographycoalescent-theoryapproximate-Bayesian-computationspecies-distribution-modelecological-niche-modelhabitat-suitabilitybioclimatic-envelopeclimate-envelopeniche-conservatismniche-evolutionrealized-nichefundamental-nichebiotic-interactioncompetitionpredationparasitismmutualismcommensalismamensalismfacilitationapparent-competitionindirect-interactiontrophic-cascadetop-down-controlbottom-up-controlmesopredator-releasekeystone-speciesecosystem-engineerfoundation-speciesdominant-speciescharacteristic-speciesindicator-speciesumbrella-speciesflagship-speciessurrogate-speciesfocal-speciestarget-speciespest-speciesinvasive-speciesnaturalized-speciesalien-speciesnon-native-speciesintroduced-speciescryptogenic-speciesrange-shiftpoleward-shiftelevational-shiftphenological-shifttemporal-mismatchtrophic-mismatchphenologyseasonalityvoltinismunivoltinebivoltinemultivoltinepartial-generationprolonged-diapauseaestivationhibernationoverwinteringoverwintering-strategycold-hardinessfreeze-tolerancefreeze-avoidancesupercoolingcryoprotectantthermal-biologythermoregulationbehavioral-thermoregulationmicroclimate-selectionhabitat-selectionoviposition-site-selectionhost-selectionmate-choicesexual-selectionsperm-competitioncryptic-female-choicereproductive-isolationprezygotic-isolationpostzygotic-isolationspeciationallopatric-speciationsympatric-speciationparapatric-speciationperipatric-speciationreinforcementhybridizationintrogressionhybrid-zoneclineecotonecontact-zonesecondary-contactexogenous-selectionendogenous-selectionecological-speciationmutation-order-speciationsexual-conflictsexual-dimorphismsexual-size-dimorphismsexual-dichromatismpolymorphismcolor-polymorphismmorphphasegregarious-phasesolitary-phasedensity-dependent-polyphenismreaction-normgenotype-environment-interactionplasticitycanalizationgene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• Tanypus

  non-biting midges

  Tanypus is a genus of non-biting midges in the family Chironomidae, subfamily Tanypodinae, comprising at least 100 described species. Larvae are aquatic and benthic, occurring in freshwater lakes and other lentic environments. Species within the genus exhibit bioturbation activity that modifies sediment oxygen dynamics. Life cycle patterns vary by species, with some showing multivoltine generations and specific overwintering strategies.

  chironomidnon-biting-midgebenthicbioturbationfreshwaterlakeaquatic-insectecosystem-engineer

• Tetramesa

  Tetramesa is a genus of minute phytophagous wasps in the family Eurytomidae comprising over 200 described species. Members are exclusively associated with grasses (Poaceae), where they typically induce stem or inflorescence galls. The genus exhibits pronounced host specificity, with most species restricted to a single grass species or closely related congeners. Several species have been deployed as biological control agents against invasive grasses, including T. romana for giant reed (Arundo donax) and candidate agents for medusahead (Taeniatherum caput-medusae) and African lovegrass (Eragrostis curvula). Adults feed on nectar. Recent phylogenomic studies have synonymized the genera Aiolomorphus and Cathilaria within Tetramesa.

  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• Tetropium parvulum

  Northern Spruce Borer

  Tetropium parvulum, the northern spruce borer, is a small cerambycid beetle specialized on spruce hosts. It was elevated from synonymy with T. cinnamopterum based on distinct morphological differences in larvae and adults, and exhibits strict host specificity to Picea compared to the conifer generalist T. cinnamopterum. The species has a one-year life cycle and is an economically important borer of white and Engelmann spruce logs in western Canada.

  Cerambycidaespruce-borerwood-boring-beetleforest-pestwestern-Canadaconifer-specialistTetropiumPiceaeconomic-pestone-year-life-cycleTetropium-parvulumNorthern-Spruce-BorerTetropium-cinnamopterum-complexmorphological-diagnosisstrict-host-specificityTetropium-alaskanum-synonymRhimphoctona-alaskensis-parasitoidspruce-log-pestAlberta-forestryboreal-forest-insectphloem-feedercambium-borerL-shaped-galleriesearly-summer-emergencewoodpecker-predationdisease-mortalityexcess-heat-mortalityTetropium-velutinum-comparisonlectotype-designationTetropium-parallelumTetropium-schwarzianumconiferous-forest-decomposerprimary-borerTetropium-taxonomyCasey-1891Canadian-Entomologisteconomic-entomologyforest-healthtimber-pestspruce-regenerationPicea-glauca-pestPicea-engelmannii-pestwestern-Canadian-distributionNorth-American-CerambycidaeTetropium-morphologylarval-urogomphiadult-genitaliapronotal-punctationeye-morphologyscutellum-structurehost-preference-divergenceconifer-specialist-vs-generalistsympatric-speciesspecies-elevation-from-synonymytype-specimen-examinationdiagnostic-keyforest-insect-natural-enemiesinsect-disease-mortalityparasitoid-waspwoodpecker-insectivorythermal-mortalitygallery-depthoverwintering-biologyspring-pupationSeptember-larval-maturityone-year-univoltinespruce-logging-impactforest-managementintegrated-pest-managementboreal-forest-ecologysubcortical-insectphloeophagous-beetlexylophagous-larvaewood-degradationnutrient-cyclingforest-successiondisturbance-ecologyfire-killed-sprucewindthrow-sprucesalvage-logging-insectstanding-dead-timberspruce-beetle-complexScolytinae-competitorCerambycidae-communityconiferous-wood-borer-guildNorth-American-forest-insectswestern-Canadian-entomologyAlberta-entomologyBritish-Columbia-entomologyManitoba-entomologyLabrador-entomologytranscontinental-distributionboreal-forest-biodiversityspruce-ecosystem-engineertree-killing-insectsublethal-host-stressinduced-host-defenseinsect-fungus-interactionblue-stain-fungus-vectorwood-discolorationlumber-degradeveneer-quality-impactpulpwood-qualitybiomass-energy-feedstock-qualitycarbon-sequestration-impactclimate-change-forest-healthwarming-thermal-mortalitydrought-stress-predispositionspruce-decline-agentmulti-stressor-forest-dynamicsinsect-population-regulationnatural-enemy-complextrophic-cascadeforest-food-webavian-insectivorywoodpecker-foragingsapsucker-drillingnuthatch-predationparasitoid-communityichneumonid-waspbraconid-wasptachinid-flynematode-parasitebacterial-diseasefungal-entomopathogenviral-diseaseabiotic-mortality-factorcompetitive-exclusioninterspecific-competitionresource-partitioninghost-tree-chemistrymonoterpene-defensephenolic-compoundresin-duct-responseinduced-resistanceconstitutive-defensespruce-genotype-resistanceprovenance-variationlocal-adaptationevolutionary-ecologyspeciation-mechanismhost-race-formationreproductive-isolationmorphological-divergencegenitalic-evolutionallopatric-divergencesympatric-maintenancehybridization-potentialintrogression-riskconservation-geneticspopulation-structuregene-floweffective-population-sizeinbreeding-depressiongenetic-diversityadaptation-potentialevolutionary-rescueassisted-migrationex-situ-conservationcryopreservationmuseum-specimentype-localityholotypelectotypeparalectotypesyntypeoriginal-descriptiontaxonomic-revisionspecies-conceptbiological-speciesmorphological-speciesecological-speciesphylogenetic-speciesintegrative-taxonomyDNA-barcodingCOI-sequencenuclear-markermitochondrial-genomephylogenomicssystematicsclassification-stabilitynomenclatural-actsynonymyhomonymyavailabilityvaliditypriorityPrinciple-of-PriorityInternational-Code-of-Zoological-Nomenclaturetype-fixationtype-depositioncollection-repositoryNational-Museum-of-Natural-HistoryCanadian-National-CollectionRoyal-Ontario-MuseumUniversity-of-Alberta-collectionforestry-researchentomological-surveyinsect-monitoringpheromone-trappingflight-intercept-trapLindgren-funnel-trapemergence-cagerearing-logdissection-microscopegenitalic-preparationmicroscopic-photographyscanning-electron-microscopyconfocal-microscopycomputed-tomography3D-reconstructionmorphometric-analysisgeometric-morphometricslandmark-analysisshape-variationsize-allometrysexual-dimorphismintraspecific-variationphenotypic-plasticityenvironmental-gradientclinal-variationaltitudinal-gradientlatitudinal-gradientphenologydegree-day-modelthermal-accumulationdevelopment-ratevoltinismdiapausequiescencecold-hardinessfreezing-tolerancesupercooling-pointcryoprotectantglycerolsorbitoltrehaloseantifreeze-proteinwinter-survivalspring-emergenceadult-longevityoviposition-behavioregg-gallerylarval-feedingfrass-productiongallery-architecturepupal-chambercallus-tissuewound-responsetree-defensecompartmentalizationCODIT-modeldecay-columnheartwood-rotsapwood-stainstructural-integrity-losswindthrow-riskstand-dynamicssuccessional-trajectoryforest-regenerationplantation-forestrysilvicultural-practicethinningpruningfertilizationsite-preparationgenetic-improvementresistance-breedingclonal-forestryseed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• Tortopus

  Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.

  EphemeropteraPolymitarcyidaeburrowing-mayfliesNeotropicalclay-bank-habitatU-shaped-tunnelsmandibular-tusksTortopsisCampsurinaePanamerican-distributioncladistic-revisionsynapomorphyparastyli-receptorsflattened-penesfurcasternal-protuberancessubapical-tubercleslimnological-sampling-biasnymph-morphologyadult-morphologytype-species-T.-igaranusnew-species-T.-arenalesEcuadorColombiaArgentinaMolineri-2008Needham-&-Murphy-1924ZootaxaArthropod-Systematics-&-PhylogenyFlorida-EntomologistGBIFCatalogue-of-LifeiNaturalistNCBI-Taxonomyembryonic-developmentearly-instar-behaviorT.-harrisiT.-bellusT.-circumfluusT.-zottaiT.-igaranusT.-incertusT.-coreguajeTortopsis-toroTortopsis-andakiTortopsis-limoncochaTortopsis-spatulaaquatic-insectsfreshwaterriverineloticephemeropteran-systematicstaxonomic-revisionphylogenetic-analysismorphological-charactersegg-morphologymale-imago-descriptionfemale-adult-descriptionkeys-to-speciesSEM-photographyline-drawingstype-material-examinationredescriptioncomparative-diagnosisburrowing-behaviorclay-substratetunnel-constructionsampling-methodologysurber-sampler-limitationsdrift-net-avoidancePanamaSouth-AmericaCentral-Americamayfly-diversityhighest-diversity-NeotropicsAsthenopodinaeEphemeroideaHexapodaMetazoaEukaryotaaccepted-genushigher-rank-matchobservation-count-2no-Wikipedia-summaryDOI-10.11646/zootaxa.2481.1.1DOI-10.11646/zootaxa.2436.1.4DOI-10.3897/asp.79.e62735DOI-10.2307/3493493Scott-et-al.-1959NavásUlmerTraverLugo-Ortiz-&-McCaffertyDomínguezBanksMcDunnoughPictetNeedham-&-Murphyfreshwater-ecologyaquatic-entomologyinsect-developmentnymphal-stagesimago-descriptionalate-adultswing-venationR-sectorgonopore-structureparastyli-lengthpedestal-morphologylateral-view-curvaturemandibular-morphologytusk-tuberclesmesosternal-structureabdominal-sternum-notchfurcasternumprotuberance-configurationreceptor-socket-orientationIR-veinR2-veinclaw-like-structuregonopore-associationpenis-base-separationfemale-fore-wingadditional-veinscomparative-morphologytype-specimenholotypeallotypeparatypeneotropical-speciesnew-species-descriptionspecies-keyidentification-keymale-adult-keyfemale-adult-keynymph-keygeneric-keyPolymitarcyidae-generasystematicsphylogenycladisticsmonophylysister-groupcharacter-statetransformation-seriesparsimony-analysismorphological-data-setexternal-morphologyegg-chorionmicropyleegg-bursterhatchingemergencebehavioral-studyearly-instarembryologyovipositionsubstrate-preferencehabitat-specificityendemicregional-diversityconservationbiomonitoringwater-quality-indicatormacroinvertebratebenthicinfaunalinterstitialhyporheicriver-bankstream-bankerosionsedimentparticle-sizeclay-contentbank-stabilityroot-matvegetation-associationflow-regimecurrent-velocitydepthwidthstream-ordercatchmentbasinecoregionbiogeographydispersalcolonizationvicariancespeciationadaptive-radiationmorphological-divergenceconvergent-evolutionhomoplasycharacter-conflictbootstrap-supportBremer-supportconsensus-treestrict-consensusmajority-ruletaxonomic-stabilitynomenclatural-actoriginal-descriptionsubsequent-designationtype-localitytype-depositorymuseum-collectionspecimen-preservationalcohol-preservationslide-mountingSEM-preparationimaging-techniquesscientific-illustrationphotographymeasurementmorphometricsratiosproportionsallometrysexual-dimorphismgeographic-variationintraspecific-variationinterspecific-variationdiagnostic-charactersidentification-confidencetaxonomic-expertiseliterature-reviewbibliographic-searchdatabase-queryonline-repositoryopen-accesspaywallabstract-onlyfull-text-unavailablecitation-analysisreference-listsource-qualityevidence-strengthknowledge-gapresearch-needfuture-studycollaborationfield-workcollection-tripsampling-effortspecimen-availabilitytype-examinationrevision-scopetaxonomic-conceptcircumscriptioninclusionexclusionspecies-groupspecies-complexcryptic-speciesmorphospeciesphylogenetic-speciesevolutionary-significant-unitoperational-taxonomic-unitDNA-barcodingmolecular-systematicsintegrative-taxonomytraditional-taxonomyalpha-taxonomybeta-taxonomygamma-taxonomyclassification-hierarchyLinnaean-systemrank-based-nomenclaturephylogenetic-nomenclatureclade-namecrown-groupstem-grouptotal-evidencecongruenceincongruencedata-partitioncharacter-weightingimplied-weightingequal-weightingordered-charactersunordered-charactersmultistate-charactersbinary-characterscontinuous-charactersmeristic-charactersqualitative-charactersquantitative-charactersmorphological-matrixdata-matrixcharacter-matrixtaxon-samplingcharacter-samplingmissing-datainapplicable-datapolymorphismintraspecific-polymorphismfixed-differencediagnostic-differenceunique-derived-characterautapomorphyshared-derived-charactersymplesiomorphyconvergent-characterparallel-evolutionreversalcharacter-optimizationACCTRANDELTRANunambiguous-optimizationambiguous-optimizationmost-parsimonious-treetree-lengthconsistency-indexretention-indexrescaled-consistency-indexhomoplasy-indexcharacter-fitcharacter-congruencecharacter-supportbranch-supportnode-supportclade-stabilitytaxonomic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• Uca pugnax

  Atlantic marsh fiddler crab, mud fiddler crab, Atlantic mud fiddler crab, marsh fiddler crab

  Minuca pugnax is a small intertidal crab native to Atlantic coast salt marshes of North America. Males possess one dramatically enlarged yellow claw used for signaling and combat, while females have two small claws. The species exhibits sexual dimorphism in body size and coloration. It constructs burrows in muddy substrates and has been observed in both low-marsh and, more recently, high-marsh habitats. Larval development includes five zoeal stages and one megalopal stage before settlement.

  crustaceandecapodfiddler-crabintertidalsalt-marshsexual-dimorphismecosystem-engineerAtlantic-coast