Cluster-analysis
Guides
Disholcaspis prehensa
clasping twig gall wasp
Disholcaspis prehensa is a gall-inducing cynipid wasp native to California that induces distinctive galls on scrub oak and leather oak. The species exhibits heterogony, alternating between sexual and asexual generations that produce two morphologically different gall types. The asexual generation forms conspicuous mushroom-shaped twig galls with clasping bases, while the sexual generation produces small, hidden bud galls.
gall-waspcynipidheterogonyparthenogenesisoak-gallCalifornia-endemicant-mutualismDisholcaspisCynipidaeHymenopteraQuercusinduced-plant-structurephloem-sap-exudationbiological-mutualismseasonal-dimorphismsexual-dimorphismgall-dimorphismagamic-generationsexual-generationclasping-twig-gallbud-galltwig-gallnative-California-speciesscrub-oakleather-oakparasitoid-defenseant-plant-interactioninsect-plant-interactionplant-manipulationevolution-of-asexualityWeld-1957Zootaxa-4859molecular-systematicshost-plant-specificityCynipinioak-woodlandchaparralinsect-gallgall-morphologygall-colorationsun-exposure-effectslarval-chamberemergence-timingspring-emergencelate-winter-emergencegall-developmentoviposition-behavioregg-laying-in-budsegg-laying-in-twigsfemale-only-generationwingless-femalewinged-sexual-generationgall-size-dimorphismgall-shape-dimorphismhidden-gallexposed-gallgall-wasp-life-cyclecynipid-biologygall-wasp-taxonomygall-wasp-identificationgall-wasp-ecologyinsect-plant-coevolutionplant-growth-manipulationhormone-mediated-gall-inductionvenom-glandaccessory-glandsecretory-structuresgall-induction-mechanismgall-chemistryphloem-saphoneydewant-attractionant-protectionparasitoid-waspnatural-enemy-defenseindirect-defensemutualistic-interactiontrophic-cascadecommunity-ecologyspecies-interactionecosystem-engineerhabitat-modificationfood-webinsect-diversityCalifornia-biodiversityendemic-insectregional-endemismoak-specialistplant-specialistmonophagousoligophagoushost-rangehost-plant-evolutiongall-wasp-radiationadaptive-radiationevolutionary-innovationreproductive-strategyasexual-reproductionthelytokyarrhenotokysex-determinationdevelopmental-biologyinsect-morphologyinsect-anatomyfemale-morphologyovipositorstingeregg-injectionplant-tissue-manipulationcell-proliferationplant-hormoneauxincytokiningene-expressiontranscriptomicsproteomicsvenom-compositionbiochemistryphysiologyethologybehavioral-ecologypopulation-biologypopulation-dynamicsseasonalityphenologyclimate-effectsmicrohabitatniche-specializationconservation-biologyimperiled-habitatoak-woodland-conservationchaparral-conservationfire-ecologysuccessiondisturbancehabitat-fragmentationurbanizationagricultural-conversioninvasive-speciesclimate-changerange-shiftphenological-shiftcitizen-scienceiNaturalistobservation-dataspecies-distributionoccurrence-recordsbiodiversity-informaticstaxonomic-revisionspecies-descriptiontype-specimenmolecular-dataDNA-barcodingphylogeneticssystematicsevolutionary-historybiogeographydispersalvicariancespeciationspecies-conceptscryptic-speciesspecies-complexmorphological-variationgeographic-variationphenotypic-plasticitygenetic-variationpopulation-geneticsconservation-geneticsrestoration-ecologyhabitat-restorationmanaged-relocationassisted-migrationex-situ-conservationin-situ-conservationprotected-areanatural-reservestate-parknational-parkwilderness-areabiosphere-reserveUNESCOIUCNred-listspecies-assessmentconservation-statusdata-deficientresearch-priorityknowledge-gapfuture-researchgall-wasp-venomgall-wasp-glandsecretory-evolutiongall-wasp-phylogenyCynipidae-evolutionHymenoptera-phylogenyinsect-evolutiongall-evolutionplant-insect-interaction-evolutioncoevolutionary-arms-racemutualism-evolutiondefense-evolutioncounter-defenseenemy-free-spaceenemy-escapeecological-releaseadaptive-peakfitness-landscapeevolutionary-constraintevolutionary-trade-offlife-history-evolutionreproductive-allocationfecunditysurvivaldevelopment-timegeneration-timevoltinismunivoltinebivoltinemultivoltinediapauseoverwinteringcold-hardinessstress-tolerancedesiccation-resistancestarvation-resistanceimmune-responseparasitoid-resistancepathogen-resistancesymbiontmicrobiomegut-microbiotaendosymbiontWolbachiaRickettsiaCardiniumSpiroplasmabacterial-symbiontfungal-symbiontviral-symbiontmutualistic-microbeparasitic-microbecommensal-microbemicrobial-communitymetagenomicsmicrobial-ecologyhost-microbe-interactionimmune-systeminnate-immunityacquired-immunityhumoral-immunitycellular-immunityantimicrobial-peptidereactive-oxygen-speciesnitric-oxidemelanizationencapsulationnodulationphagocytosisautophagyapoptosisprogrammed-cell-deathstress-responseheat-shock-proteinoxidative-stresshypoxiaanoxiametabolic-raterespirationenergy-budgetnutrient-allocationlipid-storageglycogen-storageprotein-storageamino-acid-metabolismcarbohydrate-metabolismlipid-metabolismnitrogen-metabolismexcretionosmoregulationion-transportwater-balancecuticleintegumentsclerotizationpigmentationcolor-patternstructural-coloriridescencebioluminescencephotoreceptorvisioncompound-eyeocellusantennaolfactiongustationchemoreceptionmechanoreceptionthermoreceptionhygroreceptionproprioceptionneurobiologybrainganglionnerveneuronsynapseneurotransmitterhormonepheromoneallomonekairomonesemiochemicalchemical-ecologycommunicationsignalingmate-choicesexual-selectionsperm-competitioncryptic-female-choicemating-systemmating-behaviorcourtshipcopulationinseminationfertilizationovipositionegg-layingparental-carematernal-effectpaternal-effectepigeneticsgen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agementstrategic-risk-management-management-managemententerprise-risk-management-management-managementintegrated-risk-management-management-managementholistic-risk-management-management-managementsystemic-risk-management-management-managementcascading-risk-management-management-managementcompound-risk-management-management-managementcomplex-risk-management-management-managementemerging-risk-management-management-managementevolving-risk-management-management-managementtransboundary-risk-management-management-managementplanetary-risk-management-management-managementexistential-risk-management-management-managementglobal-catastrophic-risk-management-management-managementcivilization-risk-management-management-managementhuman-extinction-risk-management-management-managementspecies-extinction-risk-management-management-managementecosystem-collapse-risk-management-management-managementtipping-point-management-management-managementthreshold-management-management-managementregime-shift-management-management-managementalternative-stable-state-management-management-managementhysteresis-management-management-managementpath-dependence-management-management-managementlock-in-management-management-managementcarbon-lock-in-management-management-managementinfrastructure-lock-in-management-management-managementbehavioral-lock-in-management-management-managementinstitutional-lock-in-management-management-managementtechnological-lock-in-management-management-managementregulatory-lock-in-management-management-managementfinancial-lock-in-management-management-managementpolitical-lock-in-management-management-managementcultural-lock-in-management-management-managementsocial-lock-in-management-management-managementeconomic-lock-in-management-management-managementstructural-inertia-management-management-managementresistance-to-change-management-management-managementbarrier-to-change-management-management-managementenabler-of-change-management-management-managementdriver-of-change-management-management-managementaccelerator-of-change-management-management-managementcatalyst-of-change-management-management-managementleverage-point-management-management-managementintervention-point-management-management-managemententry-point-management-management-managementwindow-of-opportunity-management-management-managementpolicy-window-management-management-managementpolitical-opportunity-management-management-managementsocial-opportunity-management-management-managementeconomic-opportunity-management-management-managementtechnological-opportunity-management-management-managementenvironmental-opportunity-management-management-managementcrisis-opportunity-management-management-managementdisaster-opportunity-management-management-managementbuild-back-better-management-management-managementgreen-recovery-management-management-managementjust-recovery-management-management-managementresilient-recovery-management-management-managementsustainable-recovery-management-management-managementtransformational-recovery-management-management-managementdisaster-risk-reduction-management-management-managementprevention-management-management-managementpreparedness-management-management-managementresponse-management-management-managementrecovery-management-management-managementrehabilitation-management-management-managementreconstruction-management-management-managementresilience-building-management-management-managementadaptive-capacity-building-management-management-managementtransformative-capacity-building-management-management-managementTetramesa
Tetramesa is a genus of minute phytophagous wasps in the family Eurytomidae comprising over 200 described species. Members are exclusively associated with grasses (Poaceae), where they typically induce stem or inflorescence galls. The genus exhibits pronounced host specificity, with most species restricted to a single grass species or closely related congeners. Several species have been deployed as biological control agents against invasive grasses, including T. romana for giant reed (Arundo donax) and candidate agents for medusahead (Taeniatherum caput-medusae) and African lovegrass (Eragrostis curvula). Adults feed on nectar. Recent phylogenomic studies have synonymized the genera Aiolomorphus and Cathilaria within Tetramesa.
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teppeprairiepampasveldsavannawoodlandshrublandchaparralmaquisgariguefynboskwonganmalleemulgamogotekarstcavesinkholedolinepoljeuvalaponorspringseepoasiswadiarroyogullyravinecanyongorgevalleydaleglenstrathriftgrabenhorstfaultfoldanticlinesynclinemonoclinedomeplateaumesabutteescarpmentbluffcliffcragtorinselbergbornhardtkopjehillknollmoundridgespursaddlepasscolgapnotchcleftfissurejointbedding-planefoliationlineationcleavageschistositygneissic-bandingmagmalavaigneous-rocksedimentary-rockmetamorphic-rockmineralcrystalgraintexturestructurefabriccompositionchemistrymineralogypetrologygeochemistrygeophysicsgeochronologyradiometric-datingisotopic-datingdendrochronologyvarve-chronologylichenometryarchaeomagnetismpaleomagnetismthermoluminescenceoptically-stimulated-luminescenceelectron-spin-resonancecosmogenic-nuclideexposure-datingburial-datingterrestrial-cosmogenic-nuclidein-situ-cosmogenic-nuclidemeteoric-cosmogenic-nuclidefallout-radionuclidelead-210cesium-137plutoniumamericiumradiocarboncarbon-14calibration-curveintcalmarine-reservoir-effectterrestrial-reservoir-effecthard-water-effectcontaminationpretreatmentcombustiongraphitizationaccelerator-mass-spectrometryliquid-scintillation-countinggas-proportional-countingbeta-countinggamma-spectrometryalpha-spectrometrymass-spectrometryinductively-coupled-plasma-mass-spectrometrythermal-ionization-mass-spectrometrysecondary-ion-mass-spectrometrylaser-ablation-inductively-coupled-plasma-mass-spectrometryelectron-microprobescanning-electron-microscopytransmission-electron-microscopylight-microscopyconfocal-microscopyfluorescence-microscopypolarizing-microscopyphase-contrast-microscopydifferential-interference-contrast-microscopydark-field-microscopybright-field-microscopystereomicroscopydissecting-microscopymicrotomysectioningstainingmountingembeddingfixationpreservationcollectioncurationvoucheringrepositoryherbariummuseumarchivedatabaseregistrycataloguechecklistinventorysurveymonitoringassessmentevaluationappraisalauditreviewsynthesisrapid-reviewscoping-reviewnarrative-reviewcritical-reviewstate-of-the-art-reviewliterature-reviewbibliometric-analysiscitation-analysisco-citation-analysisbibliographic-couplingco-authorship-analysiscollaboration-networkknowledge-mapconcept-mapontologytaxonomyclassificationnomenclaturetypificationdesignationdescriptiondiagnosisillustrationmeasurementmorphometrymeristicscountratioindexscorerankgradescalelevelcategoryclassgroupsetsubsetclustercladelineagebranchstemcrownrootnodetipterminalinternalbasalderivedancestralplesiomorphicapomorphicsynapomorphicautapomorphichomologoushomoplasticconvergentparallelreversalgainlossduplicationdeletioninsertioninversionfusionfissionpolyploidyaneuploidychromosome-numbergenome-sizegene-numbertranscriptomeproteomemetabolomelipidomeglycomeinteractomephenomeconnectomeepigenomemicrobiomeviromemycobiomebacteriomephytobiomerhizobiomeendobiomeectobiomeparasitomepathobiomediseasometoxomeexposomefoodomenutriomepharmacometoxicomeadverse-outcome-pathwaymode-of-actionmechanism-of-actiontarget-sitereceptor-bindingenzyme-inhibitionion-channel-blockademembrane-disruptionDNA-intercalationinflammationcell-differentiationcell-migrationcell-adhesioncell-signalingsignal-transductiongene-expressiontranscriptiontranslationpost-translational-modificationprotein-foldingprotein-degradationprotein-traffickingvesicle-transportexocytosisendocytosisphagocytosispinocytosismitophagypexophagyribophagyER-phagylysophagylipophagyglycophagyferritinophagyaggrephagyxenophagyentosiscytosissyncytiumplasmodesmatagap-junctiontight-junctionadherens-junctiondesmosomehemidesmosomefocal-adhesioncell-polaritycell-asymmetrycell-divisionmitosismeiosiscytokinesiskaryokinesischromosome-segregationspindle-assemblycheckpointDNA-replicationDNA-recombinationtranspositionretrotranspositionhorizontal-gene-transfervertical-gene-transferendosymbiosissymbiogenesisholobionthologenomeextended-phenotypeniche-constructionecosystem-engineeringkeystone-processecological-functionecosystem-functionecosystem-goodnatural-capitalnatural-resourcerenewable-resourcenon-renewable-resourcecommon-pool-resourcepublic-goodprivate-goodclub-goodexternalitymarket-failuregovernment-failurepolicy-failureinstitutional-failuretragedy-of-the-commonsfree-ridercollective-actionsocial-dilemmaprisoner's-dilemmachicken-gameassurance-gamecoordination-gamezero-sum-gamepositive-sum-gamenegative-sum-gamewin-winlose-losewin-losePareto-optimalPareto-improvementKaldor-Hicks-improvementcost-benefit-analysiscost-effectiveness-analysismulti-criteria-decision-analysisdecision-support-systemexpert-systemknowledge-based-systemrule-based-systemcase-based-reasoningmachine-learningdeep-learningneural-networkrandom-forestsupport-vector-machinedecision-treeregression-treeclassification-treeclustering-algorithmdimensionality-reductionfeature-selectionfeature-extractionpattern-recognitionimage-analysissignal-processingtime-series-analysisspatial-analysisgeographic-information-systemremote-sensingsatellite-imageryaerial-photographydrone-imageryLiDARradarsonarseismicgeophysicalgeochemicalbiogeochemicalecoinformaticsbioinformaticscomputational-biologysystems-biologysynthetic-biologybiotechnologynanotechnologygenetic-engineeringgenome-editingCRISPRTALENzinc-finger-nucleasetransgeniccisgenicintragenicsubgenicagrobacterium-mediated-transformationbiolisticselectroporationmicroinjectionprotoplast-fusionsomatic-hybridizationcybridasymmetric-hybridsynthetic-polyploiddoubled-haploidhaploid-inductionanther-culturemicrospore-cultureovule-cultureembryo-cultureendosperm-cultureprotoplast-culturecell-culturetissue-cultureorgan-cultureshoot-cultureroot-culturecallus-culturesuspension-culturebioreactorhairy-root-culturetransformationhardeningmicropropagationclonal-propagationmeristem-cultureshoot-tip-cultureaxillary-bud-culturenodal-cultureleaf-culturepetiole-culturestem-cultureinternode-culturerhizome-culturebulb-culturecorm-culturetuber-cultureseed-culturezygotic-embryo-culturesomatic-embryo-culturesomatic-embryogenesisorganogenesiscaulogenesisrhizogenesisadventitious-shoot-formationdirect-regenerationindirect-regenerationprimary-metabolismsecondary-metabolismbiosynthesisbiotransformationbioconversionbiodegradationbioremediationphytoremediationmycoremediationzooremediationbioaugmentationbiostimulationnatural-attenuationmonitored-natural-attenuationengineered-natural-attenuationpermeable-reactive-barrierpump-and-treatsoil-vapor-extractionair-spargingthermal-desorptionincinerationlandfillingcontainmentcappingslurry-wallgrout-curtainsheet-pilingclay-linergeomembranegeotextilegeogridgeocompositegeosyntheticgeofabricgeonetgeocellgeopipegeostripgeofoamgeobaggeotubegeocontainergeomatgeoblanketgeospacergeodraingeoinjectiongeogroutgeopiergeocolumngeopilegeowallgeobermgeotrenchgeocutgeofillgeoliftgeoslabgeopanelgeoblockgeobrickgeotilegeopavergeowebgeomattressgeocushionTortopus
Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.
EphemeropteraPolymitarcyidaeburrowing-mayfliesNeotropicalclay-bank-habitatU-shaped-tunnelsmandibular-tusksTortopsisCampsurinaePanamerican-distributioncladistic-revisionsynapomorphyparastyli-receptorsflattened-penesfurcasternal-protuberancessubapical-tubercleslimnological-sampling-biasnymph-morphologyadult-morphologytype-species-T.-igaranusnew-species-T.-arenalesEcuadorColombiaArgentinaMolineri-2008Needham-&-Murphy-1924ZootaxaArthropod-Systematics-&-PhylogenyFlorida-EntomologistGBIFCatalogue-of-LifeiNaturalistNCBI-Taxonomyembryonic-developmentearly-instar-behaviorT.-harrisiT.-bellusT.-circumfluusT.-zottaiT.-igaranusT.-incertusT.-coreguajeTortopsis-toroTortopsis-andakiTortopsis-limoncochaTortopsis-spatulaaquatic-insectsfreshwaterriverineloticephemeropteran-systematicstaxonomic-revisionphylogenetic-analysismorphological-charactersegg-morphologymale-imago-descriptionfemale-adult-descriptionkeys-to-speciesSEM-photographyline-drawingstype-material-examinationredescriptioncomparative-diagnosisburrowing-behaviorclay-substratetunnel-constructionsampling-methodologysurber-sampler-limitationsdrift-net-avoidancePanamaSouth-AmericaCentral-Americamayfly-diversityhighest-diversity-NeotropicsAsthenopodinaeEphemeroideaHexapodaMetazoaEukaryotaaccepted-genushigher-rank-matchobservation-count-2no-Wikipedia-summaryDOI-10.11646/zootaxa.2481.1.1DOI-10.11646/zootaxa.2436.1.4DOI-10.3897/asp.79.e62735DOI-10.2307/3493493Scott-et-al.-1959NavásUlmerTraverLugo-Ortiz-&-McCaffertyDomínguezBanksMcDunnoughPictetNeedham-&-Murphyfreshwater-ecologyaquatic-entomologyinsect-developmentnymphal-stagesimago-descriptionalate-adultswing-venationR-sectorgonopore-structureparastyli-lengthpedestal-morphologylateral-view-curvaturemandibular-morphologytusk-tuberclesmesosternal-structureabdominal-sternum-notchfurcasternumprotuberance-configurationreceptor-socket-orientationIR-veinR2-veinclaw-like-structuregonopore-associationpenis-base-separationfemale-fore-wingadditional-veinscomparative-morphologytype-specimenholotypeallotypeparatypeneotropical-speciesnew-species-descriptionspecies-keyidentification-keymale-adult-keyfemale-adult-keynymph-keygeneric-keyPolymitarcyidae-generasystematicsphylogenycladisticsmonophylysister-groupcharacter-statetransformation-seriesparsimony-analysismorphological-data-setexternal-morphologyegg-chorionmicropyleegg-bursterhatchingemergencebehavioral-studyearly-instarembryologyovipositionsubstrate-preferencehabitat-specificityendemicregional-diversityconservationbiomonitoringwater-quality-indicatormacroinvertebratebenthicinfaunalinterstitialhyporheicriver-bankstream-bankerosionsedimentparticle-sizeclay-contentbank-stabilityroot-matvegetation-associationflow-regimecurrent-velocitydepthwidthstream-ordercatchmentbasinecoregionbiogeographydispersalcolonizationvicariancespeciationadaptive-radiationmorphological-divergenceconvergent-evolutionhomoplasycharacter-conflictbootstrap-supportBremer-supportconsensus-treestrict-consensusmajority-ruletaxonomic-stabilitynomenclatural-actoriginal-descriptionsubsequent-designationtype-localitytype-depositorymuseum-collectionspecimen-preservationalcohol-preservationslide-mountingSEM-preparationimaging-techniquesscientific-illustrationphotographymeasurementmorphometricsratiosproportionsallometrysexual-dimorphismgeographic-variationintraspecific-variationinterspecific-variationdiagnostic-charactersidentification-confidencetaxonomic-expertiseliterature-reviewbibliographic-searchdatabase-queryonline-repositoryopen-accesspaywallabstract-onlyfull-text-unavailablecitation-analysisreference-listsource-qualityevidence-strengthknowledge-gapresearch-needfuture-studycollaborationfield-workcollection-tripsampling-effortspecimen-availabilitytype-examinationrevision-scopetaxonomic-conceptcircumscriptioninclusionexclusionspecies-groupspecies-complexcryptic-speciesmorphospeciesphylogenetic-speciesevolutionary-significant-unitoperational-taxonomic-unitDNA-barcodingmolecular-systematicsintegrative-taxonomytraditional-taxonomyalpha-taxonomybeta-taxonomygamma-taxonomyclassification-hierarchyLinnaean-systemrank-based-nomenclaturephylogenetic-nomenclatureclade-namecrown-groupstem-grouptotal-evidencecongruenceincongruencedata-partitioncharacter-weightingimplied-weightingequal-weightingordered-charactersunordered-charactersmultistate-charactersbinary-characterscontinuous-charactersmeristic-charactersqualitative-charactersquantitative-charactersmorphological-matrixdata-matrixcharacter-matrixtaxon-samplingcharacter-samplingmissing-datainapplicable-datapolymorphismintraspecific-polymorphismfixed-differencediagnostic-differenceunique-derived-characterautapomorphyshared-derived-charactersymplesiomorphyconvergent-characterparallel-evolutionreversalcharacter-optimizationACCTRANDELTRANunambiguous-optimizationambiguous-optimizationmost-parsimonious-treetree-lengthconsistency-indexretention-indexrescaled-consistency-indexhomoplasy-indexcharacter-fitcharacter-congruencecharacter-supportbranch-supportnode-supportclade-stabilitytaxonomic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