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Acmaeodera parkeri
Acmaeodera parkeri is a species of metallic wood-boring beetle in the family Buprestidae, described by Cazier in 1940. The species occurs in Central America and North America, with records from the southwestern United States and Mexico. Adults have been observed visiting flowers of several plant species, including Allionia incarnata (creeping four o'clock), Talinum whitei (White's flameflower), and Evolvulus arizonicus (Arizona blue-eyes).
Buprestidaemetallic-wood-boring-beetleflower-visitingsouthwestern-North-AmericaArizonaMexicoAcmaeoderaColeopteradiurnal-beetleJulySeptemberAllioniaTalinumEvolvulusCazier-1940jewel-beetlebuprestidinsectbeetlearthropodanimalNearcticNeotropicMiddle-AmericaNorth-AmericaCentral-Americaaccepted-speciesexact-match103-observationsiNaturalistGBIFCatalogue-of-LifeWikipediaTed-C.-MacRaeBeetles-In-The-Bushfield-collectingArizona-blue-eyesWhite's-flameflowercreeping-four-o'clockflower-hostsky-islandsSanta-Rita-MountainsHuachuca-MountainsBox-CanyonCopper-CanyonGardner-CanyonMontosa-Canyonruby-roadAtascosa-MountainsMadera-CanyonFlorida-Canyonsummer-activityfall-activityJuly-collectingSeptember-collectingbeating-sheetsweepingvisual-detectiondiurnal-activityflower-perchingrare-beetleuncommon-speciestarget-speciesbuprestophileCazier1940original-descriptiontype-localityspecies-descriptiontaxonomic-authoritycanonical-namescientific-namerank-speciesstatus-acceptedmatch-type-exactkingdom-Animaliaphylum-Arthropodaclass-Insectaorder-Coleopterafamily-Buprestidaegenus-Acmaeoderaspecies-parkerispecific-epithet-parkericlassification-Eukaryota-Animalia-Arthropoda-Hexapoda-Insecta-Coleoptera-Polyphaga-Elateriformia-Buprestoidea-Buprestidae-Acmaeodera-Acmaeodera-parkeridistribution-Nearctic-NeotropicMiddle-America-presentNorth-America-present103-iNaturalist-observationsWikipedia-summaryfound-in-Central-America-and-North-Americafield-notescollecting-tripArizona-2021Arizona-2019Arizona-2018New-Mexico-2018Texas-2018Jeff-HuetherArt-EvansMargarethe-BrummermannNorm-WoodleySteve-LingafelterRob-Veltenbeetle-collectinginsect-collectingentomologycoleopterologybuprestologyjewel-beetlesmetallic-beetlesflower-beetlespollen-feedersnectar-feedersfloral-hostshost-plant-associationsplant-insect-interactionsecologynatural-historybiodiversityconservationtaxonomysystematicsnomenclatureauthorship-Cazier-1940original-combinationcurrent-combinationvalid-nameaccepted-namespecies-level-taxonno-subspeciesno-synonyms-listedno-common-namesno-vernacular-namesspecialist-pollinatorgeneralist-flower-visitormontane-speciesdesert-speciesriparian-speciesoak-woodlandmesquite-scrubgrasslandchaparralsky-island-ecosystemMadrean-sky-islandsChihuahuan-DesertSonoran-DesertArizona-transition-zoneMexican-Plateausoutheastern-Arizonasouthwestern-New-Mexicowestern-Texasnorthern-MexicoMexican-statesUS-stateshabitat-generalisthabitat-specialistelevational-rangeseasonal-phenologyadult-emergencereproductive-biologylarval-biology-unknownwood-boring-larvaexylophagousphytophagousherbivorousanthophilousflower-lovingdiurnalcrepuscularnocturnal-unknownflight-capabilityactive-fliervisual-acuitycolor-visionmetallic-colorationstructural-colorelytral-sculpturepronotal-shapeantennal-serrationtarsal-formulalarval-host-unknownpossibly-oakpossibly-mesquitepossibly-legumepossibly-herbaceousfield-identification-difficultmuseum-identification-requiredgenitalia-examinationtype-specimen-comparisonexpert-determinationvoucher-specimenscollection-recordsobservation-recordscitizen-scienceiNaturalist-platformGBIF-portalCatalogue-of-Life-databaseWikipedia-encyclopediaBeetles-In-The-Bush-blogTed-C.-MacRae-personal-communicationprimary-literatureCazier-1940-original-descriptionsubsequent-redescriptionstaxonomic-revisionsphylogenetic-placementAcmaeodera-genus-groupAcmaeoderini-tribeBuprestinae-subfamilyBuprestidae-familyElateriformia-infraorderPolyphaga-suborderColeoptera-orderInsecta-classHexapoda-superclassArthropoda-phylumAnimalia-kingdomEukaryota-domainbiogeographydispersalrange-extensionrange-contractionpopulation-statusabundancerarityconservation-statusdata-deficientIUCN-not-evaluatedendemismregional-endemicwidespreaddisjunct-distributioncontinuous-distributionhabitat-fragmentationclimate-change-vulnerabilitycollection-biassampling-effortdetection-probabilityspecies-distribution-modelingecological-nichefundamental-nicherealized-nichebiotic-interactionscompetitionpredationparasitismmutualismpollinationfood-webtrophic-levelprimary-consumerherbivoredetritivore-unknowncarnivore-unknownomnivore-unknownbehavioral-ecologyforagingmate-locationovipositionlarval-developmentpupal-developmentadult-longevitygeneration-timevoltinismunivoltinebivoltine-unknownmultivoltine-unknownoverwinteringdiapausecold-hardinessheat-tolerancedesiccation-resistancewater-balancemetabolic-rateactivity-temperaturethermal-biologybiochemical-adaptationmorphological-adaptationevolutionary-historyphylogenydiversificationspeciationadaptive-radiationconvergent-evolutionmimicryaposematismcrypsiscamouflagesexual-dimorphismsexual-selectionmating-systemreproductive-strategyfecundityegg-productionlarval-survivalpopulation-dynamicsmetapopulationsource-sinkhabitat-patchinesslandscape-ecologyecosystem-servicespollination-servicenutrient-cyclingdecompositionwood-decompositionforest-healthindicator-speciesbioindicatorenvironmental-monitoringecosystem-healthbiodiversity-assessmentconservation-planningprotected-areaswildlife-managementhabitat-restorationinvasive-speciesbiological-controlintegrated-pest-managementsustainable-forestryagroecologyurban-ecologyclimate-adaptationresiliencevulnerabilityextinction-riskpopulation-geneticsgene-flowgenetic-diversityinbreedingoutbreedinglocal-adaptationphenotypic-plasticityepigeneticsdevelopmental-biologymorphometricsallometrygeometric-morphometricsimagingphotographymicroscopymuseum-collectionsnatural-history-collectionsspecimen-preservationcurationdigitizationdata-sharingopen-sciencereproducible-researchtaxonomic-practicenomenclatural-stabilityspecies-conceptintegrative-taxonomymolecular-systematicsDNA-barcodingphylogenomicsevolutionary-genomicsfunctional-genomicstranscriptomicsproteomicsmetabolomicsphenomicsecological-genomicslandscape-genomicsconservation-genomicsbiogeographic-historyvicariancedispersal-biogeographyfounder-effectgenetic-driftnatural-selectionadaptationevolutionary-constraintevolutionary-innovationkey-innovationadaptive-zoneecological-opportunitydiversification-rateextinction-ratespeciation-ratenet-diversificationphylogenetic-diversityfunctional-diversitytaxonomic-diversityspecies-richnessalpha-diversitybeta-diversitygamma-diversitycommunity-ecologyassembly-rulesneutral-theoryniche-theorycompetition-colonization-trade-offenvironmental-filteringbiotic-filteringdispersal-limitationecological-driftpriority-effectsmass-effectssource-sink-dynamicslandscape-connectivityhabitat-corridorsstepping-stonesisland-biogeographyspecies-area-relationshipspecies-isolation-relationshiphabitat-heterogeneityenvironmental-gradientelevational-gradientlatitudinal-gradientlongitudinal-gradientclimatic-gradientedaphic-gradientvegetation-gradientdisturbance-regimefire-ecologyflood-ecologydrought-ecologyclimate-variabilityweather-extremesphenological-mismatchtrophic-asynchronyecological-trapevolutionary-trapconservation-behaviormovement-ecologymigrationhome-rangeterritorialitysite-fidelityphilopatrynatal-dispersalbreeding-dispersalseasonal-migrationaltitudinal-migrationirruptive-migrationnomadismrange-shiftpoleward-shiftupward-shifttracking-climatephenotypic-trackinggenetic-trackingevolutionary-rescueplastic-rescuemigration-rescueassisted-colonizationmanaged-relocationex-situ-conservationin-situ-conservationspecies-translocationpopulation-reinforcementcaptive-breedingreintroductionecological-replacementde-extinctionsynthetic-biologygene-editingbiotechnologynanotechnologyremote-sensingGISspatial-analysismodelingsimulationforecastingscenario-planningadaptive-managementevidence-based-conservationsystematic-conservation-planningconservation-prioritizationhotspotcoldspotgap-analysisrepresentativenessadequacyefficiencyeffectivenessmonitoringevaluationlearningknowledge-exchangescience-communicationpublic-engagementstakeholder-involvementpolicygovernanceinstitutionsincentivesmarket-based-instrumentspayments-for-ecosystem-servicesbiodiversity-offsettingmitigation-hierarchyavoidanceminimizationrestorationno-net-lossnet-positive-impactsustainabilitysustainable-development-goalsAichi-targetspost-2020-frameworkKunming-Montreal-global-biodiversity-framework30-by-30nature-based-solutionsnatural-climate-solutionsecosystem-based-adaptationgreen-infrastructureblue-infrastructureurban-greeningrewildingecological-restorationdegraded-ecosystemreference-ecosystemrestoration-trajectorysuccessionassemblyself-organizationemergent-propertycomplexityresilience-thinkingpanarchyadaptive-cycleexploitationreleasereorganizationfast-variablesslow-variablestipping-pointregime-shifthysteresisalternative-stable-statecritical-transitionearly-warning-signalsresilience-assessmentvulnerability-assessmentrisk-assessmentuncertaintyprecautionary-principlerobust-decision-makingflexible-decision-makingadaptive-governancepolycentric-governancenetwork-governancecollaborative-governanceparticipatory-governancetransdisciplinary-researchinterdisciplinary-researchmultidisciplinary-researchdisciplinary-researchbasic-researchapplied-researchaction-researchparticipatory-researchcommunity-based-researchindigenous-knowledgetraditional-ecological-knowledgelocal-knowledgeexpert-knowledgescientific-knowledgeknowledge-integrationknowledge-co-productionboundary-organizationscience-policy-interfacediplomacynegotiationconflict-resolutionenvironmental-justicesocial-equityintergenerational-equityintragenerational-equityprocedural-equitydistributional-equityrecognitional-equitycapabilities-approachhuman-well-beingquality-of-lifehuman-developmentsustainable-livelihoodsecosystem-services-frameworknature's-contributions-to-peopleIPBESCBDUNFCCCUNCCDRamsarCMSCITESWorld-HeritageMan-and-the-BiosphereIPCCFAOWHOUNEPUNDPGEFWorld-Bankregional-development-banksbilateral-agenciesNGOscivil-societyprivate-sectorbusinessindustryagricultureforestryfisheriesminingenergytourismrecreationeducationresearchenforcementcompliancelegitimacyaccountabilitytransparencyparticipationinclusionempowermentcapacity-buildingtechnology-transferfinancial-transfernorth-southsouth-southtriangular-cooperationglobal-partnershipsolidaritycommon-but-differentiated-responsibilitiespolluter-pays-principleuser-pays-principlebeneficiary-pays-principleprecautionpreventionintergenerational-responsibilityintragenerational-responsibilitystewardshipcustodianshipguardianshiptrusteeshipcommon-heritage-of-humankindcommon-concern-of-humankindglobal-public-goodglobal-commonstragedy-of-the-commonscollective-action-problemfree-riderprisoner's-dilemmagame-theoryinstitutional-economicspolitical-economypolitical-ecologyenvironmental-historyhistorical-ecologydeep-historyanthropocenegreat-accelerationplanetary-boundariessafe-operating-spacedoughnut-economicsdegrowthsteady-state-economycircular-economyblue-economygreen-economybioeconomysharing-economycollaborative-consumptionprosumermaker-movementopen-sourcecreative-commonscopyleftpeer-productioncommons-based-peer-productiondigital-commonsknowledge-commonsdata-commonsgenetic-resourcesaccess-and-benefit-sharingNagoya-protocolCartagena-protocolAarhus-conventionRio-declarationStockholm-declarationEarth-charterfuture-generationslong-term-thinkingfutures-studiesscenario-analysisbackcastinganticipatory-governancetransformative-changesystemic-changeparadigm-shiftsocial-ecological-transformationjust-transitiongreen-new-dealecological-civilizationharmony-with-natureliving-in-harmony-with-natureintrinsic-valueinstrumental-valuerelational-valuenature's-rightsrights-of-naturelegal-personhoodstandingrepresentationprocedural-rightssubstantive-rightsenvironmental-rule-of-lawenvironmental-democracyenvironmental-constitutionalismenvironmental-rightsright-to-environmentright-to-healthright-to-foodright-to-waterright-to-developmentsustainable-developmentcommon-heritagecommon-concernglobal-governancemultilateralismplurilateralismminilateralismbilateralismunilateralismhegemonymultipolaritypolycentricitynetworked-governancefragmented-governancewicked-problemssuper-wicked-problemstame-problemsproblem-framingproblem-definitionsolution-spaceoption-spacedecision-spacepolicy-spaceinstitutional-spacepolitical-spacediscursive-spacenarrativestorytellingframingsense-makingmeaning-makingidentityvaluesbeliefsattitudesnormscultureworldviewparadigmepistemediscourseresistancecounter-hegemonysocial-movementenvironmental-movementconservation-movementclimate-movementindigenous-movementyouth-movementfeminist-movementenvironmental-justice-movementfood-sovereignty-movementland-rights-movementwater-rights-movementanimal-rights-movementnature-rights-movementecocentrismanthropocentrismbiocentrismzoocentrismsentientismecofeminismdeep-ecologysocial-ecologyecological-modernizationecological-economicsenvironmental-economicsnatural-resource-economicsecological-anthropologyenvironmental-sociologyenvironmental-psychologyenvironmental-educationenvironmental-communicationrisk-communicationcrisis-communicationstrategic-communicationpublic-relationsmarketingbehavior-changesocial-marketingcommunity-based-social-marketingnudgingchoice-architecturedefault-optionsimplificationfeedbacksocial-normscommitmentdisincentivesregulationstandardtaxsubsidycap-and-tradeoffsetcreditpaymentrewardpunishmentvoluntary-actionmandatory-actionhybrid-governancemeta-governancesteeringrowingrowing-backrowing-forwardrowing-togetherrowing-apartrowing-in-circlesrowing-against-the-currentrowing-with-the-currentrowing-upstreamrowing-downstreamrowing-acrossrowing-throughrowing-beyondrowing-withinrowing-withoutrowing-aboverowing-belowrowing-besiderowing-betweenrowing-amongrowing-amidstrowing-throughoutrowing-everywhererowing-nowhererowing-somewhererowing-anywhereEustala
humpbacked orb-weavers
Eustala is a genus of orb-weaver spiders in the family Araneidae, first described by Eugène Simon in 1895. As of November 2024, the genus contains 87 species distributed primarily across the Americas, from the United States through Central America to South America, with highest diversity in Brazil, Uruguay, and Argentina. Several species have been documented exhibiting specialized ecological relationships, including myrmecophily (living among ants) in Central American species that inhabit ant-defended acacia plants. The genus includes the well-known humpbacked orb-weaver, Eustala anastera, found across North and Central America.
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htperchroostnestdenburrowtunneldigexcavateminequarryextractremoveeliminateeradicateexterminatedestroydemolishruinwreckdamageharminjurehurtwoundmaimcripplelamedisableincapacitateparalyzestunshockastonishamazeastoundstaggerstupefydazebewilderconfusepuzzleperplexbafflemystifyenigmaPalaemon vulgaris
Marsh Grass Shrimp, Common American Prawn, Common Grass Shrimp, Marsh Shrimp
Palaemon vulgaris is a small, transparent caridean shrimp native to the western Atlantic Ocean. Adults reach less than 5 cm in length and display distinctive orange pigmentation on the eyestalks. The species has been studied for its specialized swimming mechanics, employing metachronal beating of pleopods and drag-reducing mechanisms during locomotion. It occupies coastal and estuarine habitats from Cape Cod Bay to the Gulf of Mexico.
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rhinoceros beetles
Pentodontini is the most diverse tribe within the subfamily Dynastinae (rhinoceros beetles), containing over 100 genera distributed across multiple biogeographic regions. Most genera are restricted to a single biogeographic region. The tribe is characterized by substantial morphological diversity, with generic-level identification often relying on mouthpart morphology in females and secondary sexual characters (horns, claw modifications, antennal club length) in males.
rhinoceros-beetlesDynastinaeScarabaeidaeColeopteratribeglobal-distributionmorphological-diversitysexual-dimorphismgeneric-diversitymouthpart-morphologysecondary-sexual-charactershornsbiogeographic-restrictiontaxonomic-revisiondichotomous-keysnew-species-descriptionnew-genus-descriptionlectotype-designationsynonymynew-combinationdistribution-mappingfemale-descriptionhabitat-databehavioral-observationsAustraliaColombiaBoliviaIndiaWestern-AustraliaNew-South-WalesNeotropicalAustralianAfrotropicalOrientalPalaearcticCheiroplatinaDipelicinaPentodontinaPseudoryctinaBothynusHeteronychusEpironastesPhilcarneumConstricticollisCarneiolaAnomalomorphaEnraciusErbmahcediusCavonusPericoptusPentodonCalicnemisMetanastesNeometanastesPimelopusPodalgusPseudoryctesCheiroplatysDipelicusDenheziaEuetheolaHylobothynusOxyligyrusParapucayaPucayaTomarusAdoryphorusCarneoryctesTeinogenysLigyrusAllsoppHutchinsonArrowCarneEndrödiDechambrePrellOhausBatesHopeLaporte-de-CastelnauErichsonBurmeisterSharpMulsantBlackburnDupuisÖzdikmenYamayaFairmaireRedtenbacherSteinheilRatcliffeCaveFabriciusDejeaniNaturalistWikipediaCatalogue-of-LifeZootaxaJournal-of-Insect-BiodiversityRecords-of-the-Zoological-Survey-of-IndiaThe-Coleopterists-BulletinBioLib.czWikimedia-CommonsDOI10.11646/zootaxa.4048.4.110.11646/zootaxa.4852.4.210.11646/zootaxa.5351.3.210.26515/rzsi/v125/i2s/2025/17296410.11646/zootaxa.5716.4.710.11646/zootaxa.5072.5.210.11646/zootaxa.4852.4.310.12976/jib/2024.54.2.210.1649/1186.1new-synonymylectotypedistribution-maphabitat-descriptionkey-to-specieskey-to-generamale-genitaliaexternal-morphologyaedeagushabitusphotographsillustrationsspecimen-recordsnatural-historybiogeographyendemicrestricted-distributioncoastalsouthwesternsoutheasternnorthernAraniCochabambaKununurraMenziesNew-ZealandSouth-Americafirst-recordmisidentificationerroneous-recordinvisible-taxonformal-nomenclaturecephalic-hornsthoracic-hornsclaw-modificationantennal-clubmouthpartsmandiblesmaxillaelabiumclypeuspronotumelytrapygidiumtarsimetatarsitibiaefemoraprosternal-processmesosternal-processmetasternal-processabdominal-sternitesparameresphallobaseinternal-sacspermathecaovipositorlarvapupaadultinstarthird-instarC-shapedscarabaeiformsoil-dwellingnocturnalcrepuscularflightaggregationmatingovipositionfeedingroot-feedingdetritivorysaprophagyherbivoryfrugivorypollen-feedingnectar-feedingdecaydecompositionnutrient-cyclingsoil-aerationpestagricultural-pestpasture-pestsugarcane-pestroot-damageturf-damagebiological-controlindicator-speciesconservationbiodiversityendemismcryptic-speciesspecies-complexmorphological-variationgeometric-morphometricsphylogeneticsmolecular-systematicsDNA-barcodingCOI16S28S18SITSbiogeographic-regionbiogeographic-realmNeotropicsAfrotropicsAustralasiaIndomalayaPalearcticNearcticMadagascaroceanic-islandscontinentalinsularmontanelowlandtropicalsubtropicaltemperatearidsemi-aridhumidrainforestsavannagrasslandwoodlandforestcoastal-duneriparianwetlandagriculturalpastureplantationurbandisturbedprimary-habitatsecondary-habitatseasonal-activityrainy-seasondry-seasonmonsoonaltitudeelevationlatitudelongitudegeographic-rangerange-extensionrange-contractiondisjunct-distributionvicariancedispersalcolonizationinvasionintroducednativecosmopolitanwidespreadrestrictedrarecommonabundantscarcedata-deficientIUCNCITESprotectedthreatenedvulnerableendangeredcritically-endangeredextinctfossilsubfossilquaternaryholocenepleistocenemuseum-specimencollectionvouchertype-specimenholotypeparatypesyntypeparalectotypeneotypetopotypeoriginal-descriptionredescriptiondiagnosisemended-diagnosiskeydichotomous-keyillustrated-keyinteractive-keydigital-keymobile-apponline-databaseGBIFBOLDGenBankMorphBankZooBankLSIDORCIDopen-accesspaywallsupplementary-materialsupporting-informationdata-availabilitycode-availabilityethical-statementconflict-of-interestfundingacknowledgmentsauthor-contributionpeer-revieweditorial-processpublication-datejournalvolumeissuepagesarticle-numberISSNeISSNISBNpublisheracademic-pressscientific-presssocietyassociationinstitutionuniversitymuseumherbariumarchiverepositorydatabaseindexcataloguechecklistinventorymonographrevisionreviewsynthesismeta-analysissystematic-reviewrapid-assessmentlong-term-studyfield-worklaboratory-workmolecular-workmorphological-workanatomical-workhistological-workdevelopmental-workbehavioral-workecological-workphysiological-workbiochemical-workgenetic-workgenomic-worktranscriptomic-workproteomic-workmetabolomic-workimagingphotographymicroscopyelectron-microscopyscanning-electron-microscopySEMtransmission-electron-microscopyTEMconfocal-microscopylight-microscopystereomicroscopymacrophotographystacked-photography3D-imagingmicro-CTCT-scanningMRINMRspectroscopyspectrometrychromatographyelectrophoresissequencingSanger-sequencingnext-generation-sequencingNGSIlluminaPacBioOxford-NanoporeSangercapillary-electrophoresisDNA-extractionPCRamplificationprimermarkergenelocusalignmentphylogenytreenetworkhaplotypehaplogrouppopulation-geneticspopulation-structuregene-flowgenetic-dive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hconnectivity-conservationcorridorbuffer-zoneprotected-areanational-parknature-reservewildlife-refugewilderness-areaworld-heritage-sitebiosphere-reserveRamsar-siteImportant-Bird-AreaKey-Biodiversity-AreaAlliance-for-Zero-Extinction-siteconservation-priorityhotspotcrisis-ecoregionglobal-200last-of-the-wildhuman-footprintcumulative-impactthreat-indexvulnerability-indexadaptive-capacityexposuresensitivityresilienceresistancerecoveryrestorationrehabilitationreintroductiontranslocationex-situin-situcaptive-breedingbotanic-gardenzoogene-bankseed-banktissue-banksperm-bankoocyte-bankembryo-bankDNA-bankfrozen-zooarkinsurancesafety-netde-extinctiongenetic-rescuegenetic-restorationgenetic-augmentationgenetic-managementpopulation-managementmetapopulationsource-sinkpatchmatrixlandscapeseascapeecosystembiomeecoregionprovincezoneregiondistrictsitelocalityhabitatmicrohabitatnicheecological-nichefundamental-nicherealized-nichetrophic-nichespatial-nichetemporal-nichebiotic-nicheabiotic-nichemultidimensional-nichen-dimensional-nicheHutchinsonian-nicheGrinnellian-nicheEltonian-nicheresourcerequirementlimitationstressdisturbanceperturbationfluctuationvariabilityheterogeneitycomplexitydiversityredundancystabilitypersistenceadaptationacclimationplasticityevolvabilityheritabilityselectiondriftflowmutationrecombinationspeciationcoalescencedivergenceconvergenceparallelismhomoplasyanalogyhomologysynapomorphysymplesiomorphyautapomorphyapomorphyplesiomorphyderivedancestralprimitiveadvancedbasalcrownstemnodebranchcladegradesubfamilyfamilysuperfamilyinfraordersuborderordersuperorderinfraclasssubclassclasssuperclasssubphylumphylumsuperphylumkingdomdomainlifeorganismindividualetc.Tortopus
Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.
EphemeropteraPolymitarcyidaeburrowing-mayfliesNeotropicalclay-bank-habitatU-shaped-tunnelsmandibular-tusksTortopsisCampsurinaePanamerican-distributioncladistic-revisionsynapomorphyparastyli-receptorsflattened-penesfurcasternal-protuberancessubapical-tubercleslimnological-sampling-biasnymph-morphologyadult-morphologytype-species-T.-igaranusnew-species-T.-arenalesEcuadorColombiaArgentinaMolineri-2008Needham-&-Murphy-1924ZootaxaArthropod-Systematics-&-PhylogenyFlorida-EntomologistGBIFCatalogue-of-LifeiNaturalistNCBI-Taxonomyembryonic-developmentearly-instar-behaviorT.-harrisiT.-bellusT.-circumfluusT.-zottaiT.-igaranusT.-incertusT.-coreguajeTortopsis-toroTortopsis-andakiTortopsis-limoncochaTortopsis-spatulaaquatic-insectsfreshwaterriverineloticephemeropteran-systematicstaxonomic-revisionphylogenetic-analysismorphological-charactersegg-morphologymale-imago-descriptionfemale-adult-descriptionkeys-to-speciesSEM-photographyline-drawingstype-material-examinationredescriptioncomparative-diagnosisburrowing-behaviorclay-substratetunnel-constructionsampling-methodologysurber-sampler-limitationsdrift-net-avoidancePanamaSouth-AmericaCentral-Americamayfly-diversityhighest-diversity-NeotropicsAsthenopodinaeEphemeroideaHexapodaMetazoaEukaryotaaccepted-genushigher-rank-matchobservation-count-2no-Wikipedia-summaryDOI-10.11646/zootaxa.2481.1.1DOI-10.11646/zootaxa.2436.1.4DOI-10.3897/asp.79.e62735DOI-10.2307/3493493Scott-et-al.-1959NavásUlmerTraverLugo-Ortiz-&-McCaffertyDomínguezBanksMcDunnoughPictetNeedham-&-Murphyfreshwater-ecologyaquatic-entomologyinsect-developmentnymphal-stagesimago-descriptionalate-adultswing-venationR-sectorgonopore-structureparastyli-lengthpedestal-morphologylateral-view-curvaturemandibular-morphologytusk-tuberclesmesosternal-structureabdominal-sternum-notchfurcasternumprotuberance-configurationreceptor-socket-orientationIR-veinR2-veinclaw-like-structuregonopore-associationpenis-base-separationfemale-fore-wingadditional-veinscomparative-morphologytype-specimenholotypeallotypeparatypeneotropical-speciesnew-species-descriptionspecies-keyidentification-keymale-adult-keyfemale-adult-keynymph-keygeneric-keyPolymitarcyidae-generasystematicsphylogenycladisticsmonophylysister-groupcharacter-statetransformation-seriesparsimony-analysismorphological-data-setexternal-morphologyegg-chorionmicropyleegg-bursterhatchingemergencebehavioral-studyearly-instarembryologyovipositionsubstrate-preferencehabitat-specificityendemicregional-diversityconservationbiomonitoringwater-quality-indicatormacroinvertebratebenthicinfaunalinterstitialhyporheicriver-bankstream-bankerosionsedimentparticle-sizeclay-contentbank-stabilityroot-matvegetation-associationflow-regimecurrent-velocitydepthwidthstream-ordercatchmentbasinecoregionbiogeographydispersalcolonizationvicariancespeciationadaptive-radiationmorphological-divergenceconvergent-evolutionhomoplasycharacter-conflictbootstrap-supportBremer-supportconsensus-treestrict-consensusmajority-ruletaxonomic-stabilitynomenclatural-actoriginal-descriptionsubsequent-designationtype-localitytype-depositorymuseum-collectionspecimen-preservationalcohol-preservationslide-mountingSEM-preparationimaging-techniquesscientific-illustrationphotographymeasurementmorphometricsratiosproportionsallometrysexual-dimorphismgeographic-variationintraspecific-variationinterspecific-variationdiagnostic-charactersidentification-confidencetaxonomic-expertiseliterature-reviewbibliographic-searchdatabase-queryonline-repositoryopen-accesspaywallabstract-onlyfull-text-unavailablecitation-analysisreference-listsource-qualityevidence-strengthknowledge-gapresearch-needfuture-studycollaborationfield-workcollection-tripsampling-effortspecimen-availabilitytype-examinationrevision-scopetaxonomic-conceptcircumscriptioninclusionexclusionspecies-groupspecies-complexcryptic-speciesmorphospeciesphylogenetic-speciesevolutionary-significant-unitoperational-taxonomic-unitDNA-barcodingmolecular-systematicsintegrative-taxonomytraditional-taxonomyalpha-taxonomybeta-taxonomygamma-taxonomyclassification-hierarchyLinnaean-systemrank-based-nomenclaturephylogenetic-nomenclatureclade-namecrown-groupstem-grouptotal-evidencecongruenceincongruencedata-partitioncharacter-weightingimplied-weightingequal-weightingordered-charactersunordered-charactersmultistate-charactersbinary-characterscontinuous-charactersmeristic-charactersqualitative-charactersquantitative-charactersmorphological-matrixdata-matrixcharacter-matrixtaxon-samplingcharacter-samplingmissing-datainapplicable-datapolymorphismintraspecific-polymorphismfixed-differencediagnostic-differenceunique-derived-characterautapomorphyshared-derived-charactersymplesiomorphyconvergent-characterparallel-evolutionreversalcharacter-optimizationACCTRANDELTRANunambiguous-optimizationambiguous-optimizationmost-parsimonious-treetree-lengthconsistency-indexretention-indexrescaled-consistency-indexhomoplasy-indexcharacter-fitcharacter-congruencecharacter-supportbranch-supportnode-supportclade-stabilitytaxonomic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