Type-specimen

Guides

Aleiodes kisomm
Aleiodes kisomm is a species of braconid wasp in the subfamily Rogadinae, described by Fortier. It belongs to a genus known for 'mummy-making' behavior, where larvae parasitize caterpillars and cause them to mummify. The species is documented from a primary type specimen housed at the University of California Riverside Entomology Research Museum (UCRC ENT 79338). As with other Aleiodes species, it is presumed to be a koinobiont endoparasitoid of Lepidoptera larvae, though specific host records for this species have not been published.
parasitoid-wasp braconid Rogadinae type-specimen UCRC
Anteon rugosiceps
Anteon rugosiceps is a species of dryinid wasp in the family Dryinidae. The species was described by Olmi and is part of a genus of parasitoid wasps that target leafhoppers and planthoppers (Hemiptera: Auchenorrhyncha). Dryinid wasps are characterized by their distinctive forelegs, which are modified into chelate (pincer-like) structures used to capture and hold their prey. Anteon species are generally small, slender wasps with reduced wing venation.
Hymenoptera Dryinidae Anteon parasitoid-wasp leafhopper-parasite type-specimen UC-Riverside
Anurida
Anurida is a genus of springtails (Collembola) in the family Neanuridae, established in 1865 by Laboulbène. The genus has cosmopolitan distribution with species occupying diverse habitats including intertidal marine zones, river floodplains, riparian areas, and forest ecosystems. Well-studied species include the intertidal specialist Anurida maritima, which exhibits complex tidal-entrained behaviors, and the terrestrial A. granaria, which has documented mycophagous associations. The genus shows notable morphological diversity in chaetotaxy and eye reduction, with some species groups exhibiting cryptic genetic divergence despite morphological similarity.
Collembola springtails intertidal tidal-behavior cryptic-species endosymbionts Wolbachia Spiroplasma diapause univoltine mycophagy chaetotaxy Neanuridae cosmopolitan-distribution circatidal-rhythm egg-diapause salt-marsh river-floodplain riparian-zone forest-habitat Beringian-fauna genetic-divergence-without-morphological-change tidal-entrainment aggregation-behavior sexual-dimorphism-in-foraging starvation-mortality holometabolous-like-development setal-reduction ocelli-reduction hammerae-group Anurida-maritima-species-group Laboulbène-1865 Poduromorpha Neanurinae Pseudachorutinae terrestrial marine-intertidal freshwater-riparian nutrient-cycling organic-matter-decomposition fungal-dispersal apothecia-feeding clay-wall-nests air-filled-cavities tidal-refuge weather-dependent-activity temperature-dependent-diapause-termination mitochondrial-genome-divergence ancient-circatidal-behavior cytoplasmic-incompatibility male-killing Type-V-cif-genes genome-wide-differentiation Pool-seq-phylogenomics Holarctic-distribution temperate-zone-adaptation overwintering-eggs autumn-mortality physiological-stress foraging-efficiency low-temperature-limitation tidal-inundation-response behavioral-synchronization nest-construction sexual-reproduction colonial-aggregation fungal-association Peziza-arvernensis riverina granaria octoculata hirsuta elegans reducta narli bisetosa Vladivostok-Botanical-Garden Primorsky-Krai Poland Southern-Brazil North-western-Europe United-Kingdom The-Netherlands Andean Arctic Sub-arctic Cape Caribbean Central-Australia coniferous-broadleaved-forest protected-forest vertical-clay-walls creek-walls salt-marsh-foraging marsh-wandering nest-marsh-exchange molting-refuge egg-deposition-sites sexually-mature-aggregation hibernating-eggs spring-hatching summer-egg-laying autumn-diapause-termination winter-development-suppression adult-death starvation-risk glycogen-depletion lipid-depletion body-size-decline sluggishness-at-low-temperature limited-low-water-period temperate-survival-strategy cosmopolitan-species-with-local-adaptation genetic-crosses-needed sex-ratio-studies-needed endosymbiont-effects-unknown reproductive-manipulation-potential selfish-genetic-elements maternal-inheritance phylogenomic-analysis single-copy-orthologous-genes nuclear-genome-divergence mitochondrial-lineage-association species-group-concept morphological-stasis evolutionary-divergence systematic-revision-needed tribe-validity Neanurinae-subdivision Pseudachorutinae-placement higher-rank-taxonomy-match GBIF-records iNaturalist-observations Catalogue-of-Life-acceptance NCBI-taxonomy Entognatha Hexapoda Eukaryota Metazoa Animalia Arthropoda springtail-biodiversity soil-mesofauna intertidal-invertebrate marine-terrestrial-transition-zone estuarine-ecology tidal-flat-ecology floodplain-ecology riparian-ecology forest-floor-ecology mycophagous-collembolan fungal-feeding-springtail nutrient-cycler decomposer detritivore organic-matter-processor ecosystem-engineer-(nest-construction)microhabitat-specialist habitat-partition sexual-dimorphism behavioral-plasticity environmental-cue-response phototaxis-modification thermotaxis-response hydrotaxis-response aggregation-pheromone-(inferred)social-behavior-(colonial)reproductive-behavior courtship oviposition-site-selection egg-guarding-(absent)diapause-evolution life-history-strategy univoltinism semelparity-(effective)annual-life-cycle seasonal-polyphenism-(absent)developmental-arrest cold-requirement-for-development temperature-threshold 5°C-diapause-termination spring-warming-trigger phenology population-dynamics demography mortality-factor starvation environmental-stress climate-sensitivity habitat-specificity endemism-(some-species)cryptic-biodiversity molecular-taxonomy integrative-taxonomy phylogeography population-genetics genomic-resources Wolbachia-genome Spiroplasma-genome bacterial-endosymbiosis host-microbe-interaction reproductive-parasitism mutualism-(unknown)commensalism-(unknown)symbiont-phylogeny horizontal-gene-transfer-(absent-in-data)prophage-genes cif-gene-evolution Type-V-clade male-killing-gene-absence CI-gene-presence wmk-gene-presence SpAID-absence bacterial-genome-reduction-(inferred)host-adaptation coevolution symbiont-sharing-between-host-lineages genetic-divergence-with-symbiont-sharing speciation-mechanism reproductive-isolation cytoplasmic-incompatibility-as-speciation-driver-(unlikely-given-identical-cif-sequences)alternative-speciation-mechanisms ecological-speciation behavioral-isolation habitat-isolation temporal-isolation geographic-isolation allopatric-divergence parapatric-divergence sympatric-divergence-(possible)cryptic-species-identification-challenge morphological-taxonomy-limitations molecular-systematics-necessity DNA-barcoding genome-skimming Pool-seq phylogenomic-inference species-delimitation integrative-species-concept operational-taxonomic-unit evolutionary-significant-unit conservation-unit biodiversity-assessment faunistics biogeography dispersal-ability passive-dispersal active-dispersal habitat-fidelity site-fidelity nest-fidelity philopatry-(inferred)population-structure gene-flow genetic-differentiation isolation-by-distance isolation-by-environment local-adaptation phenotypic-plasticity genetic-accommodation evolutionary-developmental-biology evo-devo setal-development sensory-organ-development eye-reduction-evolution cave-adaptation-(absent)soil-adaptation intertidal-adaptation desiccation-resistance-(inferred)salinity-tolerance hypoxia-tolerance-(inferred)nest-air-pocket-maintenance respiratory-adaptation cuticular-waterproofing-(inferred)osmoregulation ion-regulation excretory-system Malpighian-tubules-(standard)labial-glands digestive-system midgut hindgut feeding-apparatus maxilla mandible labrum epipharynx hypopharynx mouthparts-entognathous head-capsule antennae segment-number segment-fusion thorax abdomen furca-(absent-in-some-Neanuridae)tenaculum collophore ventral-tube reticulate-pattern pigmentation coloration size-variation body-shape cylindrical-body setal-arrangement macrosetae microsetae sensory-setae mechanoreceptors chemoreceptors hygroreceptors thermoreceptors photoreceptors ocelli-structure eye-number-reduction eye-complete-loss-(some-species)pigment-loss cuticular-granulation cuticular-tubercles cuticular-scales body-sclerotization intersegmental-membranes appendage-structure leg-segmentation claw-structure unguiculus tenent-hair empodial-appendage tibiotarsus femur trochanter coxa subcoxa abdominal-segmentation tergite-structure sternite-structure pleurite-structure tergal-chaetotaxy sternal-chaetotaxy pleural-chaetotaxy axial-setae paraxial-setae marginal-setae p-row-setae a-row-setae m-row-setae setal-formula setal-nomenclature Fjellberg-system Gisin-system taxonomic-stability nomenclatural-acts type-species type-locality type-specimen original-description subsequent-redescriptions faunal-revisions catalogues checklists databases GBIF iNaturalist NCBI BOLD COL ITIS Encyclopedia-of-Life Wikipedia primary-literature taxonomic-literature ecological-literature physiological-literature genomic-literature symbiont-literature behavioral-literature entomology acarology soil-zoology marine-biology intertidal-ecology estuarine-science limnology freshwater-biology terrestrial-ecology forest-ecology fungal-ecology microbial-ecology symbiosis-research evolutionary-biology population-biology conservation-biology biodiversity-science systematics phylogenetics paleontology-(absent)fossil-record-(absent)amber-inclusion-(possible-but-unreported)subfossil-(absent)quaternary-record-(absent)historical-ecology anthropogenic-impact pollution-sensitivity bioindicator-potential conservation-status-(unevaluated)IUCN-Red-List-(absent)habitat-protection-needs protected-area-occurrence invasive-potential-(low)agricultural-pest-(absent)household-pest-(absent)economic-importance-(minimal)scientific-importance-(high)model-organism-potential teaching-organism research-subject biodiversity-component ecosystem-service-provider cultural-significance-(absent)traditional-knowledge-(absent)indigenous-knowledge-(absent)vernacular-names-(absent)etymology Anurida-(etymology-unknown,-possibly-Greek-'an-'-without-+-'oura'-tail,-referring-to-reduced-furca)Laboulbène 1865 historical-taxonomy classical-taxonomy modern-taxonomy future-research-needs taxonomic-revision phylogenetic-analysis population-genomic-study functional-genomic-study developmental-study physiological-study behavioral-study ecological-study symbiont-study conservation-study
Argyria lacteella
milky urola moth, Southern Milky Argyria Moth
A small crambid moth with a wingspan of approximately 11 mm, distinguished by milky-white to pale gray coloration. Taxonomic identity was recently clarified through DNA hybridization capture from the 1794 Fabricius lectotype, resolving decades of confusion with the widespread North American species now assigned to Argyria gonogramma. The species occurs primarily in the southeastern United States, Caribbean, and Neotropics. Adults are active from spring through fall.
museomics DNA-hybridization-capture type-specimen taxonomic-revision DNA-barcoding COI-5P nomenclatural-clarification
Braconinae
Braconinae is a large subfamily of braconid parasitoid wasps comprising more than 2,000 described species. Members are small to medium-sized wasps, typically dark in coloration though some exhibit red, orange, or pale coloration. They possess cyclostome mouthparts and females usually have a long ovipositor. The subfamily is found worldwide with highest diversity in the Paleotropical region.
parasitoid idiobiont ectoparasitoid biocontrol cyclostome ovipositor Paleotropical-diversity Lepidoptera-parasitoid Coleoptera-parasitoid concealed-host-larvae venom-paralysis Bracon-brevicornis Atanycolus Baryproctus Bracon Ceratobracon Coeloides Glyptomorpha Iphiaulax Pseudovipio Vipio Megalommum Phylogeny-of-the-Braconinae tribal-classification Aphrastobraconini Braconini Coeloidini Compsobraconini Tropobraconini Virgulibraconini Adeshini Argamaniini Bathyaulacini Euurobraconini Glyptomorphini Gnathobraconini Physaraiini Rhammurini Vaepellini Myosotiseini Aspidobraconini Ceratobraconini Vipionini Iphiaulacini Acanthobraconini Turkey Iran China Thailand Pakistan Anatolia pistachio-longhorn-beetle Calchaenesthes-pistacivora Scolytidae Curculionidae Buprestidae coniferous-trees broadleaved-trees Malaise-trap light-trapping host-carrying-behavior Pycnobraconoides Trigastrotheca Physaraia Nesaulax Curreia Endovipio Amyosoma Megacoeloides Alienoclypeus Atanycolus-insolitus Bracon-amaculatus Bracon-quadrimaculatus Bracon-rhinchiti Bracon-murgabensis Bracon-querceus Bracon-karakumicus Bracon-hylobii Bracon-irkutensis Bracon-longiantennatus Bracon-longigenis Coeloides-sordidator Glyptomorpha-dispar Bracon-selviae Bracon-bachtiae Bracon-dersimensis Glyptomorpha-nachitshevanica Chelonus-erythrogaster Coeloides-glaboventris Coeloides-longquanus Coeloides-flavus Coeloides-changbaiensis Megalommum-pistacivorae Megalommum-xanthoceps Megalommum-jacobsoni Megalommum-ayyari Megalommum-philippinense Megalommum-dodecanesi Megalommum-ceresense Megalommum-inareatum Megalommum-antefurcale Megalommum-tibiale Physaraia-nanensis Physaraia-panhai Physaraia-sakaeratensis Physaraia-sinensis Physaraia-sumatrana Physaraia-tigeri Trigastrotheca-doiphukhaensis Trigastrotheca-pariyanonthae Trigastrotheca-sureeratae Trigastrotheca-tridentata Doi-Phu-Kha-National-Park Ardahan Erzurum Iğdır Kars Kastamonu Tunceli Fars-province Nan-province Sakaerat Zhejiang-University Chinese-Academy-of-Forestry DNA-barcoding BOLD-Systems BugGuide.net Wikispecies Wikimedia-Commons Encyclopedia-of-Life Systematic-Entomology Zootaxa ZooKeys Check-List Journal-of-Hymenoptera-Research Journal-of-Animal-and-Plant-Sciences PMC11214016 PMC12873568 28S-rRNA COI 16S-rRNA 18S-rRNA EF-1α CAD elongation-factor-1-alpha cytochrome-c-oxidase-subunit-I mitochondrial nuclear molecular-phylogeny morphological-phylogeny monophyletic tribus-nov comb.-nov sp.-nov syn.-nov incertae-sedis New-World Old-World Australian West-Palaearctic Oriental Indo-Australian Asian-Turkey European-Turkey biogeographical-provinces zoogeographical-regions checklist faunistics taxonomy systematics revision new-species new-records new-combinations new-synonyms new-tribe preliminary-key dichotomous-key diagnostic-characters type-specimen voucher-specimen holotype paratype female-specimen adult-specimen light-trap aerial-net field-collection 2011-2014 1980-2009 observation iNaturalist 14342-observations
Calliopsis puellae
desert-dandelion nomadopsis
Calliopsis puellae is a mining bee in the family Andrenidae, commonly known as the desert-dandelion nomadopsis. The species was described by Theodore Dru Alison Cockerell in 1933, originally placed in the genus Spinoliella. Its type specimen was collected by Wilmatte Porter Cockerell and her great-niece Lelah Milene Porter in Colorado, and is housed at Harvard University's Museum of Comparative Zoology. The species is found in Central America and North America.
mining-bee Andrenidae North-America Central-America Cockerell type-specimen nomadopsis
Disholcaspis prehensa
clasping twig gall wasp
Disholcaspis prehensa is a gall-inducing cynipid wasp native to California that induces distinctive galls on scrub oak and leather oak. The species exhibits heterogony, alternating between sexual and asexual generations that produce two morphologically different gall types. The asexual generation forms conspicuous mushroom-shaped twig galls with clasping bases, while the sexual generation produces small, hidden bud galls.
gall-wasp cynipid heterogony parthenogenesis oak-gall California-endemic ant-mutualism Disholcaspis Cynipidae Hymenoptera Quercus induced-plant-structure phloem-sap-exudation biological-mutualism seasonal-dimorphism sexual-dimorphism gall-dimorphism agamic-generation sexual-generation clasping-twig-gall bud-gall twig-gall native-California-species scrub-oak leather-oak parasitoid-defense ant-plant-interaction insect-plant-interaction plant-manipulation evolution-of-asexuality Weld-1957 Zootaxa-4859 molecular-systematics host-plant-specificity Cynipini oak-woodland chaparral insect-gall gall-morphology gall-coloration sun-exposure-effects larval-chamber emergence-timing spring-emergence late-winter-emergence gall-development oviposition-behavior egg-laying-in-buds egg-laying-in-twigs female-only-generation wingless-female winged-sexual-generation gall-size-dimorphism gall-shape-dimorphism hidden-gall exposed-gall gall-wasp-life-cycle cynipid-biology gall-wasp-taxonomy gall-wasp-identification gall-wasp-ecology insect-plant-coevolution plant-growth-manipulation hormone-mediated-gall-induction venom-gland accessory-gland secretory-structures gall-induction-mechanism gall-chemistry phloem-sap honeydew ant-attraction ant-protection parasitoid-wasp natural-enemy-defense indirect-defense mutualistic-interaction trophic-cascade community-ecology species-interaction ecosystem-engineer habitat-modification food-web insect-diversity California-biodiversity endemic-insect regional-endemism oak-specialist plant-specialist monophagous oligophagous host-range host-plant-evolution gall-wasp-radiation adaptive-radiation evolutionary-innovation reproductive-strategy asexual-reproduction thelytoky arrhenotoky sex-determination developmental-biology insect-morphology insect-anatomy female-morphology ovipositor stinger egg-injection plant-tissue-manipulation cell-proliferation plant-hormone auxin cytokinin gene-expression transcriptomics proteomics venom-composition biochemistry physiology ethology behavioral-ecology population-biology population-dynamics seasonality phenology climate-effects microhabitat niche-specialization conservation-biology imperiled-habitat oak-woodland-conservation chaparral-conservation fire-ecology succession disturbance habitat-fragmentation urbanization agricultural-conversion invasive-species climate-change range-shift phenological-shift citizen-science iNaturalist observation-data species-distribution occurrence-records biodiversity-informatics taxonomic-revision species-description type-specimen molecular-data DNA-barcoding phylogenetics systematics evolutionary-history biogeography dispersal vicariance speciation species-concepts cryptic-species species-complex morphological-variation geographic-variation phenotypic-plasticity genetic-variation population-genetics conservation-genetics restoration-ecology habitat-restoration managed-relocation assisted-migration ex-situ-conservation in-situ-conservation protected-area natural-reserve state-park national-park wilderness-area biosphere-reserve UNESCO IUCN red-list species-assessment conservation-status data-deficient research-priority knowledge-gap future-research gall-wasp-venom gall-wasp-gland secretory-evolution gall-wasp-phylogeny Cynipidae-evolution Hymenoptera-phylogeny insect-evolution gall-evolution plant-insect-interaction-evolution coevolutionary-arms-race mutualism-evolution defense-evolution counter-defense enemy-free-space enemy-escape ecological-release adaptive-peak fitness-landscape evolutionary-constraint evolutionary-trade-off life-history-evolution reproductive-allocation fecundity survival development-time generation-time voltinism univoltine bivoltine multivoltine diapause overwintering cold-hardiness stress-tolerance desiccation-resistance starvation-resistance immune-response parasitoid-resistance pathogen-resistance symbiont microbiome gut-microbiota endosymbiont Wolbachia Rickettsia Cardinium Spiroplasma bacterial-symbiont fungal-symbiont viral-symbiont mutualistic-microbe parasitic-microbe commensal-microbe microbial-community metagenomics microbial-ecology host-microbe-interaction immune-system innate-immunity acquired-immunity humoral-immunity cellular-immunity antimicrobial-peptide reactive-oxygen-species nitric-oxide melanization encapsulation nodulation phagocytosis autophagy apoptosis programmed-cell-death stress-response heat-shock-protein oxidative-stress hypoxia anoxia metabolic-rate respiration energy-budget nutrient-allocation lipid-storage glycogen-storage protein-storage amino-acid-metabolism carbohydrate-metabolism lipid-metabolism nitrogen-metabolism excretion osmoregulation ion-transport water-balance cuticle integument sclerotization pigmentation color-pattern structural-color iridescence bioluminescence photoreceptor vision compound-eye ocellus antenna olfaction gustation chemoreception mechanoreception thermoreception hygroreception proprioception neurobiology brain ganglion nerve neuron synapse neurotransmitter hormone pheromone allomone kairomone semiochemical chemical-ecology communication signaling mate-choice sexual-selection sperm-competition cryptic-female-choice mating-system mating-behavior courtship copulation insemination fertilization oviposition egg-laying parental-care maternal-effect paternal-effect epigenetics gene-expression-regulation transcription-factor signaling-pathway hormone-receptor nuclear-receptor steroid-hormone juvenile-hormone ecdysone insulin TOR-pathway AMPK nutrient-sensing growth-regulation size-regulation allometry scaling morphometrics geometric-morphometrics landmark shape-analysis size-analysis multivariate-analysis principal-component-analysis discriminant-analysis cluster-analysis phylogenetic-analysis comparative-method ancestral-state-reconstruction character-evolution trait-evolution evolutionary-rate diversification-rate speciation-rate extinction-rate net-diversification phylogenetic-diversity functional-diversity taxonomic-diversity species-richness species-evenness alpha-diversity beta-diversity gamma-diversity community-assembly community-structure community-dynamics ecosystem-function ecosystem-service pollination seed-dispersal nutrient-cycling decomposition primary-production herbivory predation parasitism mutualism competition facilitation apparent-competition indirect-interaction interaction-network trophic-level energy-flow material-cycling stability resilience resistance recovery environmental-gradient niche fundamental-niche realized-niche niche-breadth niche-overlap niche-partitioning resource-partitioning character-displacement competitive-exclusion limiting-similarity evolutionary-diversification macroevolution microevolution extinction mass-extinction biodiversity-crisis ecosystem-management sustainable-development environmental-policy environmental-law environmental-ethics environmental-education science-communication public-engagement open-science data-sharing FAIR-principles reproducibility replicability transparency research-integrity scientific-method hypothesis-testing experimental-design sampling-design statistical-analysis modeling simulation prediction forecasting scenario-analysis risk-assessment uncertainty sensitivity-analysis validation verification benchmarking intercomparison meta-analysis systematic-review evidence-synthesis knowledge-integration interdisciplinary-research transdisciplinary-research convergence-research team-science collaboration network-science complex-systems emergence self-organization nonlinearity feedback threshold tipping-point hysteresis alternative-stable-state regime-shift critical-transition early-warning-signal resilience-thinking adaptive-management social-ecological-system coupled-human-natural-system sustainability-science futures-studies anticipatory-governance transformative-change just-transition green-economy circular-economy nature-based-solution ecosystem-based-adaptation ecosystem-based-mitigation climate-adaptation climate-mitigation carbon-sequestration blue-carbon green-carbon biodiversity-offset habitat-banking payment-for-ecosystem-services natural-capital ecosystem-accounting green-infrastructure urban-ecology landscape-ecology spatial-ecology geographic-information-system remote-sensing satellite-imagery drone unmanned-aerial-vehicle sensor-network internet-of-things big-data machine-learning artificial-intelligence deep-learning neural-network computer-vision natural-language-processing data-mining data-science computational-biology bioinformatics genomics metabolomics phenomics imaging microscopy electron-microscopy confocal-microscopy light-sheet-microscopy super-resolution-microscopy cryo-electron-microscopy X-ray-tomography magnetic-resonance-imaging spectroscopy mass-spectrometry chromatography electrophoresis sequencing DNA-sequencing RNA-sequencing whole-genome-sequencing exome-sequencing transcriptome-sequencing metagenome-sequencing epigenome-sequencing single-cell-sequencing long-read-sequencing third-generation-sequencing nanopore-sequencing genome-assembly genome-annotation genome-evolution comparative-genomics functional-genomics structural-genomics population-genomics landscape-genomics conservation-genomics ancient-DNA paleogenomics molecular-ecology molecular-evolution phylogenomics phylodynamics coalescent-theory quantitative-genetics genetic-architecture genotype-phenotype-map pleiotropy epistasis genetic-correlation heritability selection-gradient breeding-value genomic-prediction genomic-selection marker-assisted-selection quantitative-trait-locus genome-wide-association-study linkage-mapping association-mapping fine-mapping positional-cloning candidate-gene functional-validation gene-editing CRISPR TALEN zinc-finger-nuclease base-editing prime-editing gene-drive synthetic-biology bioengineering biotechnology agricultural-biotechnology environmental-biotechnology industrial-biotechnology medical-biotechnology pharmaceutical-biotechnology veterinary-biotechnology forensic-biotechnology biosecurity biosafety bioprospecting biopiracy access-and-benefit-sharing Nagoya-Protocol Cartagena-Protocol Convention-on-Biological-Diversity CITES IPBES IPCC UNFCCC UNCCD UNSDG Aichi-targets Kunming-Montreal-Global-Biodiversity-Framework Paris-Agreement Sendai-Framework Rio-Earth-Summit Stockholm-Conference environmental-history history-of-science history-of-ecology history-of-entomology natural-history museum-collection herbarium zoological-collection holotype paratype syntype lectotype neotype epitype iconotype original-description subsequent-designation nomenclature taxonomy phylogenetic-nomenclature rank-based-nomenclature phylogenetic-code international-code-of-zoological-nomenclature international-code-of-nomenclature-for-algae-fungi-and-plants international-code-of-nomenclature-of-prokaryotes international-code-of-nomenclature-for-cultivated-plants international-commission-on-zoological-nomenclature general-committee nomenclatural-acts original-publication valid-publication effective-publication availability priority homonymy synonymy nomen-novum nomen-conservandum nomen-rejiciendum nomen-dubium nomen-inquirendum species-inquirenda incertae-sedis new-combination new-status emendation correction erratum addendum supplement monograph revision catalog checklist field-guide identification-key diagnosis description illustration photograph drawing map distribution-map habitat-photo micrograph scanning-electron-micrograph transmission-electron-micrograph light-micrograph stereomicrograph macrophotograph habitus dorsal-view ventral-view lateral-view anterior-view posterior-view head thorax abdomen wing leg mouthpart genitalia gall plant-gall rose-gall willow-gall goldenrod-gall sunflower-gall aster-gall thistle-gall sagebrush-gall creosote-gall manzanita-gall forest woodland savanna grassland prairie steppe meadow pasture agricultural-field orchard vineyard garden park urban-green-space wildland-urban-interface wilderness nature-reserve county-park municipal-park private-reserve conservation-easement land-trust stewardship restoration ecological-restoration rewilding reintroduction translocation assisted-colonization integrated-conservation conservation-planning systematic-conservation-planning Marxan Zonation conservation-prioritization hotspot biodiversity-hotspot endemic-bird-area important-plant-area key-biodiversity-area Alliance-for-Zero-Extinction EDGE-species evolutionarily-distinct-and-globally-endangered flagship-species umbrella-species keystone-species foundation-species dominant-species charismatic-species indicator-species sentinel-species bioindicator ecological-indicator environmental-indicator water-quality-indicator air-quality-indicator soil-quality-indicator habitat-quality-indicator ecosystem-health-indicator biodiversity-indicator sustainability-indicator Millennium-Ecosystem-Assessment ecosystem-services provisioning-services regulating-services cultural-services supporting-services food fiber fuel freshwater genetic-resources biochemicals natural-medicines pharmaceuticals ornamental-resources air-quality-regulation climate-regulation water-regulation erosion-regulation water-purification waste-treatment disease-regulation pest-regulation habitat-provision soil-formation photosynthesis recreation tourism aesthetic-value spiritual-value educational-value inspirational-value sense-of-place cultural-heritage knowledge-systems traditional-ecological-knowledge indigenous-knowledge local-knowledge community-based-monitoring participatory-research action-research knowledge-co-production science-policy-interface science-society-interface boundary-organization knowledge-broker conservation-education nature-interpretation ecotourism sustainable-tourism responsible-travel green-travel low-impact-travel carbon-offset voluntary-carbon-market compliance-carbon-market REDD+reducing-emissions-from-deforestation-and-forest-degradation afforestation reforestation forest-restoration agroforestry silvopasture forest-landscape-restoration landscape-approach integrated-landscape-management watershed-management river-basin-management coastal-zone-management marine-spatial-planning ocean-governance blue-economy sustainable-blue-economy fisheries-management aquaculture sustainable-agriculture regenerative-agriculture organic-agriculture agroecology permaculture conservation-agriculture climate-smart-agriculture precision-agriculture digital-agriculture smart-farming vertical-farming controlled-environment-agriculture hydroponics aeroponics aquaponics algae-farming insect-farming entomophagy alternative-protein plant-based-diet reduced-meat-consumption food-system-transformation sustainable-food-system circular-food-system food-waste-reduction zero-hunger nutrition-security food-sovereignty seed-sovereignty farmers'-rights plant-variety-protection intellectual-property patent plant-breeder's-rights utility-model trademark geographical-indication certification organic-certification fair-trade-certification Rainforest-Alliance Marine-Stewardship-Council Forest-Stewardship-Council Roundtable-on-Sustainable-Palm-Oil Better-Cotton-Initiative GlobalGAP food-safety food-quality traceability accountability governance polycentric-governance adaptive-governance collaborative-governance network-governance multi-level-governance global-governance regional-governance national-governance subnational-governance local-governance community-governance customary-governance traditional-governance indigenous-governance self-governance autonomy decentralization devolution subsidiarity empowerment capacity-building institutional-strengthening policy-reform legal-reform regulatory-reform incentive disincentive economic-instrument market-based-instrument tax subsidy fee charge levy fine penalty liability compensation insurance risk-transfer catastrophe-bond green-bond blue-bond sustainability-linked-bond impact-investing blended-finance public-private-partnership concession lease permit license quota tradable-permit cap-and-trade carbon-tax carbon-pricing social-cost-of-carbon shadow-price internal-carbon-pricing carbon-border-adjustment border-carbon-adjustment climate-club just-transition-mechanism loss-and-damage climate-finance green-climate-fund adaptation-fund special-climate-change-fund global-environment-facility biodiversity-finance nature-finance natural-capital-finance impact-investment ESG environmental-social-and-governance sustainable-finance responsible-investment active-ownership engagement proxy-voting divestment fossil-fuel-divestment deforestation-free conversion-free no-net-loss net-gain biodiversity-net-gain mitigation-hierarchy avoidance minimization offset compensatory-mitigation in-lieu-fee conservation-banking wetland-mitigation-banking species-banking ecosystem-services-marketplace natural-capital-accounting system-of-environmental-economic-accounting inclusive-wealth genuine-progress-indicator well-being-economy degrowth post-growth steady-state-economy doughnut-economics planetary-boundaries safe-and-just-space social-foundation ecological-ceiling fair-share Earth-system Anthropocene Great-Acceleration Great-Transformation sustainability-transformation systemic-change structural-change paradigm-shift worldview-shift values-shift behavior-change lifestyle-change consumption-change production-change technology-change innovation disruptive-innovation transformative-innovation social-innovation institutional-innovation policy-innovation governance-innovation business-model-innovation financial-innovation technological-innovation digital-innovation green-technology clean-technology climate-technology ecosystem-based-disaster-risk-reduction blue-infrastructure hybrid-infrastructure natural-infrastructure building-with-nature working-with-nature engineering-with-nature ecological-engineering environmental-engineering conservation-ecology industrial-ecology agricultural-ecology forest-ecology grassland-ecology wetland-ecology freshwater-ecology marine-ecology coastal-ecology deep-ecology political-ecology social-ecology human-ecology cultural-ecology symbolic-ecology spiritual-ecology religious-ecology eco-theology creation-care stewardship-ethic land-ethic Earth-ethic planetary-ethic cosmic-ethic deep-ecology-platform ecofeminism environmental-justice climate-justice intergenerational-justice intragenerational-justice spatial-justice recognition-justice procedural-justice distributive-justice restorative-justice transformative-justice environmental-racism climate-racism slow-violence structural-violence environmental-conflict resource-conflict climate-conflict water-conflict land-conflict territorial-conflict conservation-conflict human-wildlife-conflict predator-control pest-control invasive-species-management disease-management wildlife-management habitat-management population-management harvest-management sustainable-use sustainable-harvesting adaptive-harvesting precautionary-approach ecosystem-approach ecosystem-based-management integrated-management holistic-management risk-based-management scenario-planning strategic-foresight transformative-governance adaptive-capacity transformative-capacity learning-organization learning-network knowledge-network community-of-practice epistemic-community knowledge-action-network future-earth conservation-science restoration-science ecological-economics environmental-economics natural-resource-economics resource-economics ecological-anthropology environmental-anthropology environmental-sociology environmental-psychology environmental-geography science-and-technology-studies science-policy-studies innovation-studies horizon-scanning trend-analysis weak-signal wild-card black-swan grey-rhino systemic-risk cascading-risk compound-risk complex-risk emerging-risk evolving-risk transboundary-risk planetary-risk existential-risk global-catastrophic-risk civilization-risk human-extinction-risk species-extinction-risk ecosystem-collapse-risk irreversible-change abrupt-change nonlinear-change threshold-effect path-dependence lock-in carbon-lock-in infrastructure-lock-in behavioral-lock-in institutional-lock-in technological-lock-in regulatory-lock-in financial-lock-in political-lock-in cultural-lock-in social-lock-in economic-lock-in structural-inertia resistance-to-change barrier-to-change enabler-of-change driver-of-change accelerator-of-change catalyst-of-change leverage-point intervention-point entry-point window-of-opportunity policy-window political-opportunity social-opportunity economic-opportunity technological-opportunity environmental-opportunity crisis-opportunity disaster-opportunity build-back-better green-recovery just-recovery resilient-recovery sustainable-recovery transformational-recovery disaster-risk-reduction prevention preparedness response rehabilitation reconstruction resilience-building adaptive-capacity-building transformative-capacity-building disaster-risk-management climate-risk-management environmental-risk-management financial-risk-management operational-risk-management strategic-risk-management enterprise-risk-management integrated-risk-management holistic-risk-management systemic-risk-management cascading-risk-management compound-risk-management complex-risk-management emerging-risk-management evolving-risk-management transboundary-risk-management planetary-risk-management existential-risk-management global-catastrophic-risk-management civilization-risk-management human-extinction-risk-management species-extinction-risk-management ecosystem-collapse-risk-management tipping-point-management threshold-management regime-shift-management alternative-stable-state-management hysteresis-management path-dependence-management lock-in-management carbon-lock-in-management 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sustainable-recovery-management transformational-recovery-management disaster-risk-reduction-management prevention-management preparedness-management response-management recovery-management rehabilitation-management reconstruction-management resilience-building-management adaptive-capacity-building-management transformative-capacity-building-management disaster-risk-management-management climate-risk-management-management environmental-risk-management-management financial-risk-management-management operational-risk-management-management strategic-risk-management-management enterprise-risk-management-management integrated-risk-management-management holistic-risk-management-management systemic-risk-management-management cascading-risk-management-management compound-risk-management-management complex-risk-management-management emerging-risk-management-management evolving-risk-management-management transboundary-risk-management-management planetary-risk-management-management existential-risk-management-management global-catastrophic-risk-management-management civilization-risk-management-management human-extinction-risk-management-management species-extinction-risk-management-management ecosystem-collapse-risk-management-management tipping-point-management-management threshold-management-management regime-shift-management-management alternative-stable-state-management-management hysteresis-management-management path-dependence-management-management lock-in-management-management carbon-lock-in-management-management infrastructure-lock-in-management-management behavioral-lock-in-management-management institutional-lock-in-management-management technological-lock-in-management-management regulatory-lock-in-management-management financial-lock-in-management-management political-lock-in-management-management cultural-lock-in-management-management social-lock-in-management-management economic-lock-in-management-management structural-inertia-management-management resistance-to-change-management-management barrier-to-change-management-management enabler-of-change-management-management driver-of-change-management-management accelerator-of-change-management-management catalyst-of-change-management-management leverage-point-management-management intervention-point-management-management entry-point-management-management window-of-opportunity-management-management policy-window-management-management political-opportunity-management-management social-opportunity-management-management economic-opportunity-management-management technological-opportunity-management-management environmental-opportunity-management-management crisis-opportunity-management-management disaster-opportunity-management-management build-back-better-management-management green-recovery-management-management just-recovery-management-management resilient-recovery-management-management sustainable-recovery-management-management transformational-recovery-management-management disaster-risk-reduction-management-management prevention-management-management preparedness-management-management response-management-management recovery-management-management rehabilitation-management-management reconstruction-management-management resilience-building-management-management adaptive-capacity-building-management-management transformative-capacity-building-management-management disaster-risk-management-management-management climate-risk-management-management-management environmental-risk-management-management-management financial-risk-management-management-management operational-risk-management-management-management strategic-risk-management-management-management enterprise-risk-management-management-management integrated-risk-management-management-management holistic-risk-management-management-management systemic-risk-management-management-management cascading-risk-management-management-management compound-risk-management-management-management complex-risk-management-management-management emerging-risk-management-management-management evolving-risk-management-management-management transboundary-risk-management-management-management planetary-risk-management-management-management existential-risk-management-management-management global-catastrophic-risk-management-management-management civilization-risk-management-management-management human-extinction-risk-management-management-management species-extinction-risk-management-management-management ecosystem-collapse-risk-management-management-management tipping-point-management-management-management threshold-management-management-management regime-shift-management-management-management alternative-stable-state-management-management-management hysteresis-management-management-management path-dependence-management-management-management lock-in-management-management-management carbon-lock-in-management-management-management infrastructure-lock-in-management-management-management behavioral-lock-in-management-management-management institutional-lock-in-management-management-management technological-lock-in-management-management-management regulatory-lock-in-management-management-management financial-lock-in-management-management-management political-lock-in-management-management-management cultural-lock-in-management-management-management social-lock-in-management-management-management economic-lock-in-management-management-management structural-inertia-management-management-management resistance-to-change-management-management-management barrier-to-change-management-management-management enabler-of-change-management-management-management driver-of-change-management-management-management accelerator-of-change-management-management-management catalyst-of-change-management-management-management leverage-point-management-management-management intervention-point-management-management-management entry-point-management-management-management window-of-opportunity-management-management-management policy-window-management-management-management political-opportunity-management-management-management social-opportunity-management-management-management economic-opportunity-management-management-management technological-opportunity-management-management-management environmental-opportunity-management-management-management crisis-opportunity-management-management-management disaster-opportunity-management-management-management build-back-better-management-management-management green-recovery-management-management-management just-recovery-management-management-management resilient-recovery-management-management-management sustainable-recovery-management-management-management transformational-recovery-management-management-management disaster-risk-reduction-management-management-management prevention-management-management-management preparedness-management-management-management response-management-management-management recovery-management-management-management rehabilitation-management-management-management reconstruction-management-management-management resilience-building-management-management-management adaptive-capacity-building-management-management-management transformative-capacity-building-management-management-management
Entedoninae
Entedoninae is a subfamily of chalcid wasps in the family Eulophidae comprising over 90 genera. Members are primarily parasitoids with documented associations across diverse host groups including gall-inducing insects, leafminers, sawflies, thrips, and ants. The subfamily exhibits considerable morphological and ecological diversity, with species found in tropical to temperate habitats worldwide.
parasitoid biological-control gall-associated leafminer-parasitoid Chalcidoidea Hymenoptera Eulophidae Entedoninae tropical-forest agricultural-pest-control sawfly-parasitoid thrips-parasitoid ant-associated rice-pest Cecidomyiidae Tenthredinidae Pontania Closterocerus Ceranisus Afronympha Ametallon Chrysonotomyia Omphale-genus-group canopy understorey ecological-plasticity Bangladesh Panama Africa New-Zealand Europe Senegal South-Africa Democratic-Republic-of-the-Congo Uganda endoparasitoid larval-parasitoid egg-parasitoid host-specificity forest-habitat degraded-habitat mid-altitude-rainforest lowland-rainforest tropical-America classical-biological-control augmentative-biological-control integrated-pest-management rice-hispa Dicladispa-armigera gall-wasp sawfly-gall leafminer ant-symbiosis Formicidae Khaya-senegalensis Ochna-pulchra morphological-taxonomy molecular-taxonomy 28S-rDNA COI CytB new-species-description generic-revision host-range parasitoid-complex food-web ecosystem-service natural-enemy pest-management entomophagous hyperparasitoid inquiline curculionid Apion-minimum Phyllocolpa-leucosticta Chrysocharis Entedon Asecodes Pleurotroppopsis Afrotroppopsis Zaommomentedon Apleurotropis Kratoysma Platocharis Schizocharis Parahorismenus Atullya Tropicharis Chrysochuris cruy mirabilis oryzamyntor eminpashai carinatum dussiae longicaudata machaeriae unimaculata claviger gorgonaense auripunctata galbina laeviscuta phenacapsia cecvora elongata pontaniae heyeri risbeci milletiae newbyi Förster-1856 Chalcidoidea-phylogeny biodiversity species-richness cryptic-species molecular-diagnostics host-plant-association habitat-specificity canopy-fauna understorey-fauna forest-stratification tropical-biodiversity Afrotropical Neotropical Palaearctic Nearctic Oriental Australasian Pacific Holarctic biogeography taxonomic-revision annotated-key catalogue type-specimen museum-collection slide-mounted-specimens morphometrics genitalia wing-venation antennae clypeus mandibles malar-space scape pedicel flagellum gaster vertex suture color-pattern pale-femora dark-gaster sexual-dimorphism male female ovipositor emergence rearing quarantine field-collection foreign-exploration augmentative-release pesticide-resistance sustainable-agriculture food-security rice-production cereal-pest vegetable-pest fruit-pest ornamental-pest forest-pest urban-pest invasive-species native-range introduced-range establishment spread impact non-target-effects host-specificity risk-assessment environmental-safety regulatory-approval research-symposium proceedings peer-reviewed Zootaxa ZooKeys Insect-Systematics-&-Evolution Australian-Journal-of-Entomology Far-Eastern-Entomologist Russian-Entomological-Journal Belgian-Journal-of-Entomology Florida-Entomologist Pan-Pacific-Entomologist Entomological-News Proceedings-of-the-Hawaiian-Entomological-Society Transactions-of-the-American-Entomological-Society Journal-of-Natural-History Zoological-Journal-of-the-Linnean-Society BioControl Journal-of-Economic-Entomology Entomologia-Experimentalis-et-Applicata Environmental-Entomology Oecologia Cladistics Molecular-Phylogenetics-and-Evolution Heredity Canadian-Entomologist Proceedings-of-the-Entomological-Society-of-Washington Annals-of-the-Entomological-Society-of-America Bulletin-of-Entomological-Research Neotropical-Entomology Journal-of-Asia-Pacific-Entomology Turkish-Journal-of-Zoology Polish-Journal-of-Entomology North-Western-Journal-of-Zoology Entomologica-Americana Entomologische-Berichten Sahlbergia Dugesiana Acta-Zoologica-Lilloana Folia-Entomologica-Mexicana Revista-de-la-Sociedad-Entomologica-Argentina Boletin-de-la-Asociacion-Espanola-de-Entomologia Entomologicheskoe-Obozrenie Entomological-Review Zoological-Institute-RAS Proceedings-of-the-Zoological-Institute-of-the-Russian-Academy-of-Sciences Taiwan-Agricultural-Research-Institute Council-of-Agriculture University-of-California Riverside Entomology-Research-Museum California-Department-of-Food-and-Agriculture Pierce's-Disease-Control-Program biological-control-program integrated-pest-management-program research-progress-report annual-summary technical-report symposium-proceedings conference-abstract poster-presentation invited-paper monograph book-chapter identification-guide illustrated-key annotated-catalogue type-material voucher-specimen collection-curation taxonomy systematics phylogenetics molecular-systematics DNA-barcoding species-complex sibling-species species-boundaries geographic-variation population-genetics gene-flow reproductive-isolation host-race biotype strain laboratory-rearing insectary mass-production release monitoring impact-assessment efficacy economic-evaluation cost-benefit-analysis farmer-adoption extension education policy regulation international-cooperation capacity-building training workshop field-day demonstration on-farm-research participatory-research farmer-field-school sustainable-intensification agroecology conservation-biological-control habitat-management floral-resources refuge buffer-zone landscape-complexity connectivity spillover natural-enemy-diversity food-web-structure trophic-interactions multitrophic indirect-effects trait-mediated density-dependent inverse-density-dependent functional-response numerical-response searching-efficiency handling-time oviposition-behavior host-selection host-acceptance host-suitability host-quality host-stage-preference egg-load progeny-allocation sex-ratio local-mate-competition sib-mating inbreeding fitness longevity fecundity development-time thermal-requirements degree-days phenology synchrony asynchrony seasonal-adaptation diapause quiescence cold-hardiness heat-tolerance desiccation-resistance starvation-resistance dispersal flight walking foraging patch-use area-restricted-search oviposition-marking host-marking superparasitism multiparasitism intraguild-predation hyperparasitism autoparasitism heteronomous-hyperparasitism facultative-hyperparasitism obligate-hyperparasitism primary-parasitoid secondary-parasitoid tertiary-parasitoid mummy aphid-mummy scale-mummy pupal-parasitoid egg-larval-parasitoid larval-pupal-parasitoid idiobiont koinobiont ectoparasitoid solitary gregarious polyembryony thelytoky arrhenotoky diploid haplodiploid chromosome-number genome-size transcriptome proteome metabolome morphome phenome extended-phenotype niche-construction ecosystem-engineering keystone-species ecosystem-service-provider functional-group guild trophic-level energy-flow nutrient-cycling decomposition predation herbivory parasitism mutualism commensalism competition apparent-competition exploitative-competition interference-competition scramble-competition contest-competition resource-partitioning niche-differentiation character-displacement co-evolution arms-race Red-Queen geographic-mosaic local-adaptation phenotypic-plasticity genetic-adaptation epigenetics transgenerational-effects maternal-effects paternal-effects parental-care mating-system sexual-selection sperm-competition cryptic-female-choice mate-choice lek territory aggression alarm defense venom allomone kairomone synomone semiochemical pheromone trail-pheromone alarm-pheromone aggregation-pheromone sex-pheromone marking-pheromone oviposition-deterrent host-marking-pheromone plant-volatile induced-plant-defense systemic-acquired-resistance induced-systemic-resistance priming volatile-organic-compound green-leaf-volatile terpenoid phenylpropanoid jasmonic-acid salicylic-acid ethylene abscisic-acid gibberellin auxin cytokinin brassinosteroid strigolactone nitric-oxide reactive-oxygen-species hypersensitive-response programmed-cell-death autophagy apoptosis necrosis pathogen-associated-molecular-pattern damage-associated-molecular-pattern pattern-recognition-receptor effector-triggered-immunity gene-for-gene avirulence virulence effector toxin phytotoxin mycotoxin bacterial-toxin viral-toxin insecticidal-protein cry-protein vip-protein cyt-protein spore conidium appressorium haustorium mycelium sclerotium fruiting-body basidiocarp ascocarp perithecium cleistothecium pycnidium acervulus sporodochium symptom sign disease-triangle disease-cycle epidemic endemic pandemic outbreak eradication containment management control prevention forecasting warning-system decision-support-system expert-system model simulation economic-threshold economic-injury-level action-threshold treatment-threshold damage-threshold aesthetic-injury-level cultural-control physical-control mechanical-control chemical-control genetic-control sterile-insect-technique incompatible-insect-technique precision-agriculture site-specific-management variable-rate-application remote-sensing geographic-information-system global-positioning-system unmanned-aerial-vehicle drone sensor imaging spectroscopy machine-learning artificial-intelligence deep-learning neural-network computer-vision image-analysis pattern-recognition classification segmentation detection tracking counting sampling sampling-plan sequential-sampling binomial-sampling presence-absence-sampling fixed-precision-sampling variable-precision-sampling spatial-sampling temporal-sampling stratified-sampling cluster-sampling systematic-sampling random-sampling convenience-sampling judgment-sampling expert-sampling quota-sampling snowball-sampling respondent-driven-sampling capture-recapture mark-recapture mark-resight distance-sampling line-transect point-transect strip-transect plot quadrat pitfall-trap Malaise-trap pan-trap water-pan yellow-pan blue-pan white-pan flight-interception-trap window-trap canopy-fogging beating-tray sweep-net aerial-net ground-net aquatic-net D-vac vacuum-sampler aspirator mouth-aspirator mechanical-aspirator berlese-funnel Tullgren-funnel Winkler-extractor pitfall barber-trap pitfall-array drift-fence funnel-trap light-trap UV-light black-light mercury-vapor sodium-vapor LED actinic color-trap sticky-trap pheromone-trap synthetic-pheromone natural-pheromone kairomone-trap food-bait protein-bait sugar-bait fermentation-bait alcohol-bait vinegar-bait molasses-bait fruit-bait meat-bait fish-bait dung-bait carrion-bait decoy lure attractant repellent deterrent antifeedant oviposition-stimulant arrestant suppressant toxicant stomach-poison contact-poison fumigant systemic translaminar residual persistent non-persistent broad-spectrum narrow-spectrum selective non-selective target-specific host-specific mode-of-action nerve-poison acetylcholinesterase-inhibitor GABA-antagonist sodium-channel-modulator nicotinic-acetylcholine-receptor-agonist ryanodine-receptor-modulator chitin-synthesis-inhibitor juvenile-hormone-mimic ecdysone-agonist microbial-pesticide entomopathogenic-nematode entomopathogenic-fungus entomopathogenic-bacterium entomopathogenic-virus entomopathogenic-protozoa Bacillus-thuringiensis Bt Bacillus-sphaericus Paenibacillus-popilliae Serratia-entomophila Metarhizium-anisopliae Beauveria-bassiana Lecanicillium-lecanii Isaria-fumosorosea Nomuraea-rileyi Hirsutella-thompsonii Entomophthora Neozygites Nosema Vairimorpha Thelohania Anticarsia-gemmatalis-nucleopolyhedrovirus AgMNPV Cydia-pomonella-granulovirus CpGV Spodoptera-exigua-multiple-nucleopolyhedrovirus SeMNPV Autographa-californica-multiple-nucleopolyhedrovirus AcMNPV nucleopolyhedrovirus granulovirus cypovirus entomopoxvirus iridovirus parvovirus tetravirus nodavirus reovirus picornavirus iflavirus densovirus birnavirus totivirus partitivirus hypovirulence mycovirus bacteriophage phage transduction conjugation transformation horizontal-gene-transfer vertical-gene-transfer mobile-genetic-element transposon plasmid integron genomic-island pathogenicity-island symbiosis-island metabolic-island resistance-island virulence-factor antibiotic-resistance insecticide-resistance acaricide-resistance herbicide-resistance fungicide-resistance bactericide-resistance nematicide-resistance molluscicide-resistance rodenticide-resistance resistance-management resistance-monitoring resistance-detection resistance-diagnosis resistance-mechanism metabolic-resistance target-site-resistance penetration-resistance behavioral-resistance cross-resistance multiple-resistance negative-cross-resistance synergism antagonism potentiation safener adjuvant surfactant spreader sticker extender buffer acidifier defoamer drift-retardant rainfastness UV-protectant formulation emulsifiable-concentrate wettable-powder water-dispersible-granule suspension-concentrate capsule-suspension microencapsulation nanotechnology controlled-release slow-release attract-and-kill sterile-insect-release inundative-release inoculative-release conservation augmentation classical neoclassical new-association targeted area-wide suppression protection production integrated-crop-management integrated-production organic-farming biodynamic-farming permaculture agroforestry silvopasture climate-smart-agriculture conservation-agriculture digital-agriculture smart-farming agriculture-4.0 internet-of-things big-data cloud-computing blockchain distributed-ledger traceability transparency certification standard governance stakeholder value-chain supply-chain food-system nutrition-security livelihood income employment gender youth social-inclusion equity fairness justice ethics animal-welfare environmental-protection biodiversity-conservation ecosystem-restoration land-degradation-neutrality desertification deforestation reforestation afforestation forest-restoration wetland-restoration grassland-restoration soil-restoration soil-health soil-organic-matter soil-carbon soil-biodiversity soil-fauna soil-microbiome rhizosphere phyllosphere anthroposphere technosphere biosphere ecosphere hydrosphere atmosphere lithosphere pedosphere cryosphere climate-change global-warming greenhouse-gas carbon-dioxide methane nitrous-oxide fluorinated-gas carbon-sequestration carbon-storage carbon-sink carbon-source carbon-neutral carbon-negative net-zero climate-adaptation climate-mitigation climate-resilience vulnerability exposure sensitivity adaptive-capacity transition pathway scenario projection forecast prediction uncertainty risk hazard damage loss adaptation mitigation resilience sustainability sustainable-development sustainable-development-goals SDG Agenda-2030 Paris-Agreement Convention-on-Biological-Diversity CBD Nagoya-Protocol Cartagena-Protocol Aichi-Targets Kunming-Montreal-Global-Biodiversity-Framework post-2020-biodiversity-framework One-Health Planetary-Health EcoHealth One-Welfare One-World One-Future interdisciplinary transdisciplinary multidisciplinary cross-disciplinary intersectoral multistakeholder partnership collaboration cooperation network platform hub center-of-excellence capacity-development institutional-strengthening policy-support science-policy-interface evidence-based-policy science-diplomacy technology-transfer knowledge-transfer knowledge-exchange knowledge-sharing open-access open-data open-science open-innovation citizen-science community-based-monitoring action-research transformative-research solution-oriented-research problem-driven-research impact-oriented-research implementation-research operational-research applied-research basic-research fundamental-research strategic-research tactical-research adaptive-research innovation invention discovery breakthrough frontier emerging converging enabling disruptive transformative exponential digital biological physical cognitive social institutional technological ecological evolutionary revolutionary incremental radical systematic holistic integrated comprehensive inclusive participatory bottom-up top-down middle-out hybrid mixed-methods qualitative quantitative empirical theoretical conceptual analytical computational experimental observational descriptive correlational causal mechanistic predictive prescriptive normative evaluative assessment appraisal review audit evaluation outcome-evaluation process-evaluation cost-effectiveness cost-benefit cost-utility multi-criteria-decision-analysis stakeholder-analysis social-network-analysis institutional-analysis political-economy-analysis gender-analysis youth-analysis vulnerability-analysis risk-analysis uncertainty-analysis sensitivity-analysis scenario-analysis systems-analysis life-cycle-assessment footprint water-footprint carbon-footprint ecological-footprint nitrogen-footprint biodiversity-footprint land-footprint material-footprint social-footprint health-footprint planetary-boundary safe-operating-space doughnut-economics regenerative restorative circular green blue bioeconomy circular-economy sharing-economy gig-economy platform-economy digital-economy knowledge-economy creative-economy care-economy wellbeing-economy degrowth post-growth steady-state doughnut sufficiency prosperity flourishing thriving buen-vivir ubuntu harmony balance equilibrium homeostasis robustness adaptability flexibility plasticity evolvability innovability transformability sustainability-science resilience-science complexity-science systems-science network-science data-science information-science computer-science cognitive-science behavioral-science social-science natural-science life-science earth-science environmental-science agricultural-science veterinary-science medical-science health-science nutrition-science food-science material-science engineering-science interdisciplinary-science transdisciplinary-science convergence-science team-science responsible-science ethical-science inclusive-science decolonized-science feminist-science indigenous-science traditional-knowledge local-knowledge expert-knowledge lay-knowledge tacit-knowledge codified-knowledge explicit-knowledge implicit-knowledge procedural-knowledge declarative-knowledge epistemic-knowledge ontological-knowledge practical-knowledge theoretical-knowledge scientific-knowledge indigenous-knowledge traditional-ecological-knowledge TEK farmers'-knowledge fishers'-knowledge herders'-knowledge hunters'-knowledge gatherers'-knowledge women's-knowledge men's-knowledge youth-knowledge elder-knowledge intergenerational-knowledge transgenerational-knowledge ancestral-knowledge inherited-knowledge learned-knowledge acquired-knowledge constructed-knowledge co-produced-knowledge participatory-knowledge collaborative-knowledge transdisciplinary-knowledge interdisciplinary-knowledge multidisciplinary-knowledge cross-disciplinary-knowledge integrated-knowledge holistic-knowledge systemic-knowledge complex-knowledge uncertain-knowledge ambiguous-knowledge contested-knowledge negotiated-knowledge agreed-knowledge shared-knowledge common-knowledge public-knowledge private-knowledge proprietary-knowledge open-knowledge free-knowledge accessible-knowledge usable-knowledge actionable-knowledge relevant-knowledge timely-knowledge credible-knowledge legitimate-knowledge salient-knowledge authoritative-knowledge stakeholder-knowledge policy-maker-knowledge practitioner-knowledge researcher-knowledge academic-knowledge technical-knowledge managerial-knowledge administrative-knowledge bureaucratic-knowledge political-knowledge economic-knowledge social-knowledge cultural-knowledge historical-knowledge geographical-knowledge ecological-knowledge biological-knowledge agricultural-knowledge environmental-knowledge climate-knowledge health-knowledge nutrition-knowledge food-knowledge safety-knowledge security-knowledge risk-knowledge disaster-knowledge emergency-knowledge crisis-knowledge conflict-knowledge peace-knowledge development-knowledge sustainability-knowledge resilience-knowledge adaptation-knowledge mitigation-knowledge transformation-knowledge innovation-knowledge technology-knowledge digital-knowledge data-knowledge information-knowledge communication-knowledge media-knowledge journalism-knowledge education-knowledge learning-knowledge training-knowledge capacity-knowledge institutional-knowledge 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Eustala
humpbacked orb-weavers
Eustala is a genus of orb-weaver spiders in the family Araneidae, first described by Eugène Simon in 1895. As of November 2024, the genus contains 87 species distributed primarily across the Americas, from the United States through Central America to South America, with highest diversity in Brazil, Uruguay, and Argentina. Several species have been documented exhibiting specialized ecological relationships, including myrmecophily (living among ants) in Central American species that inhabit ant-defended acacia plants. The genus includes the well-known humpbacked orb-weaver, Eustala anastera, found across North and Central America.
Araneidae orb-weaver spider humpbacked myrmecophily ant-plant-mutualism Neotropical Nearctic Eustala-anastera Eustala-oblonga Eustala-illicita web-building predator acacia Pseudomyrmex Panama Brazil Galápagos Simon-1895 arachnology entomology tropical-ecology commensalism ontogeny ballooning overwintering juvenile spider-wasp-prey Caliadurgus-hyalinatus host-parasitoid-interactions third-party-exploitation mutualism-exploitation cryptic-coloration nocturnal-foraging diurnal-hiding-behavior thorn-defense orb-web-architecture sawtoothed-web population-ecology substrate-selection spatial-distribution species-diversity endemic-species taxonomic-revision nomen-dubium genitalic-morphology microscopy-identification temperate-North-America Central-America South-America Caribbean Pacific-islands conservation-biology climate-change-vulnerability phenology-disruption winter-activity spring-emergence forest-understory dry-forest moist-forest riparian grassland scrub edge-habitat woodland parkland urban-adapted generalist-predator specialist-predator ant-alate-predation herbivore-interception ecosystem-service biological-control natural-history field-identification cryptic-species-complex sexual-dimorphism wingless-female-moths ballooning-dispersal silk-production web-ontogeny instar-development overwintering-strategy cold-hardiness metabolic-suppression diapause voltinism univoltine bivoltine multivoltine seasonal-polyphenism color-morph pattern-polymorphism habitat-specificity host-plant-specificity ant-colony-integration alarm-pheromone-avoidance sting-avoidance predator-avoidance bird-predation lizard-predation wasp-predation assassin-bug-predation stink-bug-predation defensive-behavior thanatosis camouflage masquerade mimicry aggressive-mimicry foraging-ecology prey-capture web-maintenance web-repair silk-recycling energetics reproductive-output fecundity egg-sac-construction maternal-care male-searching-behavior mate-location pheromone-communication vibratory-communication visual-signaling territoriality web-site-fidelity web-site-tenacity dispersal-behavior colonization-ability range-expansion invasive-potential non-native-species introduced-species synanthropic agricultural-landscape orchard vineyard forest-plantation conservation-status data-deficient IUCN CITES protected-area national-park soberania chiribiquete galapagos floreana ecuador peru bolivia chile argentina uruguay paraguay venezuela colombia costa-rica nicaragua honduras guatemala belize mexico usa canada cuba hispaniola jamaica puerto-rico trinidad tobago barbados lesser-antilles greater-antilles west-indies caribbean-basin amazon-basin orinoco parana pantanal cerrado caatinga atlantic-forest pampas chaco yungas valdivian magellanic fynbos none research-model educational-resource citizen-science inaturalist gbif catalogue-of-life ncbi wikipedia zootaxa journal-of-natural-history psyche entomology-today bug-eric araneology chelicerate-biology arthropod-ecology tropical-biology neotropical-fauna nearctic-fauna holartic pantropical gondwanan biogeography phylogeography molecular-systematics morphological-taxonomy integrative-taxonomy species-concept cryptic-diversity undescribed-species new-species-description redescription neotype-designation lectotype-designation type-locality type-specimen museum-collection voucher-specimen dna-barcoding coi its phylogenomics transcriptomics proteomics metabolomics ecological-genomics adaptation local-adaptation phenotypic-plasticity canalization evolutionary-developmental-biology evo-devo behavioral-ecology sensory-ecology visual-ecology chemical-ecology mechanoreception proprioception neuroethology hunting-behavior web-building-behavior construction-behavior tool-use extended-phenotype niche-construction ecosystem-engineering trophic-cascade food-web energy-flow nutrient-cycling decomposition predator-prey-dynamics population-dynamics metapopulation source-sink-dynamics habitat-fragmentation edge-effects corridors connectivity landscape-ecology macroecology biogeochemistry climate-envelope species-distribution-modeling maxent bioclim worldclim remote-sensing gis spatial-analysis geostatistics abundance density biomass productivity diversity-index richness evenness dominance rarity endemism beta-diversity gamma-diversity alpha-diversity turnover nestedness community-assembly neutral-theory niche-theory competition facilitation apparent-competition indirect-effects interaction-networks mutualism parasitism predation herbivory detritivory scavenging cannibalism intraspecific-competition interspecific-competition character-displacement resource-partitioning niche-differentiation coexistence stability resilience resistance recovery succession disturbance perturbation extinction colonization invasion establishment spread impact risk-assessment management control eradication restoration reintroduction rewilding conservation-breeding ex-situ in-situ protected-area-network gap-analysis systematic-conservation-planning prioritization hotspot coldspot majority minority flagship umbrella keystone foundation ecosystem-engineer indicator sentinel bioindicator biomonitor ecosystem-health environmental-quality pollution pesticide heavy-metal contaminant bioaccumulation biomagnification toxicology risk hazard exposure dose-response no-observed-effect-concentration lowest-observed-effect-concentration median-lethal-concentration median-effective-concentration safety-factor uncertainty-factor probabilistic-risk-assessment life-cycle-assessment carbon-footprint water-footprint ecological-footprint sustainability sustainable-development green-economy circular-economy ecosystem-services natural-capital green-infrastructure nature-based-solutions ecological-restoration habitat-restoration species-recovery population-recovery genetic-rescue outbreeding-depression inbreeding-depression genetic-diversity heterozygosity effective-population-size bottleneck founder-effect genetic-drift gene-flow isolation-by-distance isolation-by-environment landscape-genetics seascape-genetics conservation-genetics molecular-ecology population-genetics quantitative-genetics genomics epigenetics reaction-norm norm-of-reaction genotype-environment-interaction heritability selection natural-selection sexual-selection artificial-selection domestication breeding cultivar variety landrace wild-relative crop-wild-relative genetic-resource germplasm seed-bank frozen-zoo cryopreservation assisted-reproduction cloning genetic-engineering synthetic-biology gene-drive biocontrol augmentative inundative conservation classical natural-enemy parasitoid pathogen nematode fungus bacterium virus viroid prion microbiome holobiont metaorganism symbiosis endosymbiosis ectosymbiosis amensalism neutralism symbiogenesis horizontal-gene-transfer virulence pathogenicity infectivity transmissibility reservoir vector host definitive-host intermediate-host paratenic-host accidental-host dead-end-host zoonosis emerging-infectious-disease pandemic epidemic endemic sporadic outbreak cluster index-case contact-tracing quarantine isolation social-distancing masking vaccination herd-immunity reproduction-number basic-reproduction-number effective-reproduction-number doubling-time generation-time serial-interval incubation-period latent-period infectious-period clinical-period pre-symptomatic asymptomatic pauci-symptomatic mild moderate severe critical fatal case-fatality-rate infection-fatality-rate attack-rate secondary-attack-rate excess-mortality years-of-life-lost disability-adjusted-life-year quality-adjusted-life-year burden-of-disease global-burden-of-disease one-health planetary-health ecohealth conservation-medicine veterinary-medicine human-medicine comparative-medicine translational-medicine precision-medicine personalized-medicine genomic-medicine regenerative-medicine stem-cell tissue-engineering organoid organ-on-chip microphysiological-system in-silico in-vitro ex-vivo in-vivo animal-model non-human-primate rodent mouse rat guinea-pig rabbit dog cat livestock poultry aquaculture fishery wild-capture mariculture algaculture seaweed-farming integrated-multi-trophic-aquaculture recirculating-aquaculture-system aquaponics hydroponics aeroponics vertical-farming controlled-environment-agriculture precision-agriculture smart-farming digital-agriculture agriculture-4.0 internet-of-things artificial-intelligence machine-learning deep-learning neural-network computer-vision drone unmanned-aerial-vehicle satellite gps gnss geographic-information-system decision-support-system expert-system knowledge-based-system ontology semantic-web linked-data open-data big-data data-mining data-science data-analytics predictive-analytics prescriptive-analytics descriptive-analytics diagnostic-analytics business-intelligence artificial-intelligence-ethics algorithmic-bias fairness accountability transparency explainability interpretability privacy security safety reliability robustness adaptability flexibility scalability efficiency effectiveness efficacy performance profitability cost-benefit cost-effectiveness economic-analysis financial-analysis investment return-on-investment net-present-value internal-rate-of-return payback-period break-even sensitivity-analysis scenario-analysis monte-carlo-simulation decision-tree real-options game-theory behavioral-economics nudge choice-architecture default framing anchoring availability-heuristic representativeness-heuristic affect-heuristic confirmation-bias availability-bias anchoring-bias framing-bias loss-aversion endowment-effect status-quo-bias sunk-cost-fallacy planning-fallacy optimism-bias pessimism-bias overconfidence underconfidence dunning-kruger-effect imposter-syndrome stereotyping prejudice discrimination implicit-bias explicit-bias structural-bias systemic-bias institutional-bias interpersonal-bias intrapersonal-bias unconscious-bias conscious-bias affinity-bias similarity-bias in-group-bias out-group-bias halo-effect horns-effect contrast-effect recency-effect primacy-effect serial-position-effect mere-exposure-effect familiarity-principle propinquity-effect social-proof authority scarcity reciprocity commitment consistency liking unity influence persuasion compliance conformity obedience empowerment agency autonomy self-determination motivation intrinsic-motivation extrinsic-motivation self-efficacy self-esteem self-concept self-identity social-identity collective-identity cultural-identity national-identity gender-identity sexual-orientation religion spirituality values beliefs attitudes norms customs traditions rituals ceremonies celebrations festivals holidays commemorations memorials monuments heritage legacy history historiography archaeology paleontology anthropology sociology psychology cognitive-science neuroscience linguistics philosophy ethics morality justice equity equality inclusion diversity belonging representation participation engagement capacity-building skill-development knowledge-transfer technology-transfer innovation invention discovery creativity imagination curiosity wonder awe beauty elegance simplicity complexity chaos order pattern structure function form process dynamics mechanism causation correlation association relationship interaction connection network system emergence self-organization autopoiesis homeostasis allostasis stress acclimation acclimatization plasticity fragility vulnerability sensitivity uncertainty ignorance surprise novelty serendipity chance luck fortune destiny fate karma dharma tao zen mindfulness meditation contemplation reflection introspection self-awareness consciousness subconscious unconscious dream sleep circadian biological-clock chronobiology seasonality phenology migration dispersal nomadism sedentism home-range habitat-use resource-selection foraging hunting gathering farming herding hunting-gathering subsistence livelihood economy market trade exchange barter currency money capital wealth poverty inequality development growth progress well-being happiness life-satisfaction quality-of-life human-development social-development economic-development millennium-development-goals sustainable-development-goals paris-agreement kyoto-protocol montreal-protocol basel-convention stockholm-convention rotterdam-convention unfccc cbd cms ramsar world-heritage man-and-biosphere ipcc ipbes encyclopedia-of-life wikidata wikimedia creative-commons open-access open-science faire-data research-data-management data-stewardship data-curation digital-preservation long-term-archiving repository archive library museum herbarium collection specimen voucher type holotype paratype syntype lectotype neotype epitype isolectotype isoneotype topotype homotype allotype cotype genotype phenotype ecotype chemotype morphotype karyotype idiogram chromosome genome proteome transcriptome metabolome lipidome glycome interactome connectome epigenome virome mycobiome bacteriome archaeome eukaryome phytobiome rhizobiome endobiome ectobiome exo-biome astrobiome planetary-biome earth-system climate-system ocean-system terrestrial-system freshwater-system atmospheric-system cryospheric-system anthroposphere technosphere noosphere biosphere hydrosphere lithosphere atmosphere magnetosphere heliosphere cosmosphere universe multiverse dimension space time spacetime relativity quantum-mechanics string-theory m-theory loop-quantum-gravity cosmology astronomy astrophysics planetary-science earth-science geology geophysics geochemistry geomorphology soil-science pedology edaphology hydrology limnology oceanography meteorology climatology paleoclimatology stratigraphy sedimentology mineralogy petrology volcanology seismology tectonics plate-tectonics continental-drift orogeny epeirogeny isostasy erosion weathering deposition transport sedimentation lithification diagenesis metamorphism melting crystallization igneous sedimentary metamorphic rock-cycle water-cycle carbon-cycle nitrogen-cycle phosphorus-cycle sulfur-cycle oxygen-cycle hydrogen-cycle biogeochemical-cycle nutrient-cycle trophic-pyramid food-chain ecological-network interaction-web mutualistic-network antagonistic-network competition-network facilitation-network ecosystem-function ecosystem-process provisioning-service regulating-service cultural-service supporting-service habitat-service refuge nursery corridor stepping-stone buffer-zone core-area periphery matrix patch landscape seascape riverscape skyscape soundscape smellscape tastescape touchscape mindscape dreamscape memoryscape heritagescape futurescape possibility-space scenario-space decision-space policy-space governance-space institutional-space social-space cultural-space economic-space political-space legal-space ethical-space moral-space spiritual-space sacred-space profane-space liminal-space transitional-space threshold boundary edge frontier margin center core hinterland heartland homeland motherland fatherland native-land ancestral-land promised-land wilderness pioneer settler colonist immigrant emigrant refugee asylum-seeker displaced-person stateless-person nomad traveler tourist pilgrim explorer adventurer discoverer inventor innovator creator maker builder craftsman artisan artist designer architect engineer scientist scholar academic researcher investigator detective journalist reporter correspondent chronicler historian biographer memoirist autobiographer diarist letter-writer poet novelist playwright screenwriter essayist critic reviewer editor publisher librarian archivist curator conservator restorer preservationist conservationist environmentalist ecologist naturalist botanist zoologist entomologist arachnologist herpetologist ichthyologist ornithologist mammalogist primatologist anthropologist archaeologist paleontologist geologist geographer cartographer surveyor navigator expedition voyage journey travel tour trip visit stay residence habitation dwelling shelter home house building construction architecture design plan blueprint model prototype mock-up draft sketch drawing painting sculpture installation exhibition display presentation demonstration show spectacle event occasion ceremony ritual celebration festival fair bazaar commerce business industry manufacturing production fabrication assembly processing refining purification synthesis analysis separation extraction identification characterization quantification measurement assessment evaluation appraisal valuation estimation calculation computation simulation modeling prediction forecast projection scenario outlook prospect expectation anticipation hope fear worry concern care attention focus concentration awareness perception sensation feeling emotion affect mood temperament personality character disposition attitude opinion belief conviction faith trust confidence reliance dependence independence self-sufficiency self-reliance hardiness toughness strength power force energy vigor vitality life health wellness fitness soundness integrity wholeness completeness perfection excellence quality standard criterion benchmark yardstick measure gauge sign signal symptom mark token symbol image icon emblem logo brand trademark copyright patent intellectual-property proprietary confidential secret classified restricted limited controlled regulated managed governed administered operated maintained sustained supported funded financed resourced staffed equipped furnished supplied provided delivered distributed allocated assigned designated appointed elected selected chosen picked preferred favored liked loved adored cherished treasured valued esteemed respected honored celebrated commemorated remembered recalled recollected reminisced reflected contemplated meditated pondered considered weighed evaluated judged assessed appraised rated ranked graded scored marked pointed noted observed noticed seen viewed looked watched gazed stared glanced peeked peered scanned surveyed examined inspected scrutinized studied investigated researched analyzed synthesized integrated compiled composed written authored penned drafted created produced made built constructed fabricated manufactured assembled put-together set-up established founded instituted organized arranged ordered structured systematized methodized rationalized streamlined optimized improved enhanced upgraded refined polished perfected completed finished concluded ended terminated closed shut sealed locked secured protected guarded defended shielded screened covered hidden concealed masked cloaked veiled shrouded wrapped enveloped surrounded encircled enclosed contained held kept retained preserved conserved saved stored stockpiled hoarded accumulated amassed gathered collected aggregated clustered grouped categorized sorted leveled tiered layered stratified hierarchized pyramided tree web mesh grid array configuration layout scheme program project enterprise venture undertaking endeavor effort attempt try test trial experiment investigation inquiry query question problem issue matter subject topic theme motif leitmotif thread strand line sequence series progression continuation extension expansion enlargement evolution change transformation conversion transition shift alteration modification adjustment accommodation reconciliation harmonization synchronization coordination integration unification combination merger fusion blend mixture compound composite alloy amalgam incorporation assimilation absorption digestion metabolism catabolism anabolism equilibrium balance steadiness constancy permanence endurance persistence continuity duration longevity lifespan lifetime existence being entity object thing item article piece part portion segment section division component element constituent ingredient factor aspect facet feature characteristic property attribute trait indication evidence proof verification validation confirmation corroboration substantiation authentication certification accreditation licensing permitting authorizing empowering enabling allowing letting tolerating accepting receiving taking getting obtaining acquiring gaining earning winning achieving attaining reaching arriving coming approaching nearing closing converging meeting joining connecting linking binding tying fastening attaching affixing appending adding supplementing complementing completing finishing perfecting consummating fulfilling satisfying gratifying pleasing delighting enjoying relishing savoring appreciating valuing prizing treasuring cherishing nurturing cultivating fostering promoting advancing furthering encouraging supporting backing assisting aiding helping serving attending tending caring nursing healing curing treating remedying correcting rectifying repairing fixing mending restoring renewing reviving resuscitating reanimating reawakening rekindling reigniting relaunching restarting rebooting refreshing rejuvenating reinvigorating reenergizing recharging refueling replenishing restocking replacing substituting exchanging swapping trading bartering negotiating bargaining dealing transacting contracting agreeing consenting approving endorsing sanctioning ratifying confirming validating authenticating verifying checking testing examining inspecting reviewing auditing monitoring supervising overseeing managing directing leading guiding steering piloting navigating driving propelling pushing pulling dragging hauling towing carrying bearing transporting conveying transmitting sending dispatching shipping delivering distributing disseminating spreading broadcasting publishing issuing releasing launching introducing presenting showing exhibiting displaying demonstrating illustrating exemplifying typifying representing symbolizing signifying meaning denoting connoting implying suggesting indicating pointing referring alluding hinting intimating insinuating inferring deducing concluding reasoning thinking cogitating cerebrating pondering musing reflecting contemplating meditating concentrating focusing listening hearing heeding noting noticing observing watching looking seeing viewing regarding considering weighing evaluating assessing judging appraising estimating guessing speculating hypothesizing theorizing postulating proposing recommending advising counseling consulting conferring discussing debating arguing disputing contesting challenging opposing resisting defying confronting facing encountering experiencing undergoing suffering enduring withstanding surviving living existing subsisting persisting persevering continuing remaining staying abiding residing inhabiting occupying possessing owning holding keeping retaining maintaining preserving conserving protecting guarding defending securing sheltering harboring entertaining welcoming greeting saluting hailing acclaiming applauding praising commending lauding extolling celebrating honoring glorifying exalting magnifying amplifying enlarging expanding extending stretching diffusing dispersing scattering allocating assigning allotting apportioning dividing partitioning separating detaching disconnecting disjoining disuniting dissociating disengaging withdrawing retreating receding retrograding reversing inverting turning rotating revolving spinning twirling whirling swirling circling orbiting looping coiling winding twisting bending curving arching vaulting bridging spanning crossing traversing passing moving going traveling journeying voyaging sailing flying gliding soaring hovering floating drifting flowing streaming pouring rushing gushing spurting spraying splashing splattering spattering uniting combining merging fusing blending mixing mingling intermingling intermixing integrating incorporating assimilating absorbing digesting metabolizing transforming converting transmuting transfiguring metamorphosing evolving developing growing maturing ripening aging declining deteriorating degenerating decaying rotting decomposing disintegrating crumbling collapsing falling failing dying perishing expiring departing leaving exiting retiring resigning abdicating relinquishing surrendering yielding submitting succumbing capitulating acquiescing complying conforming adapting adjusting accommodating reconciling harmonizing synchronizing coordinating unifying standardizing normalizing regularizing systematizing methodizing rationalizing streamlining simplifying clarifying elucidating explaining interpreting translating rendering paraphrasing summarizing abstracting condensing compressing shrinking reducing decreasing diminishing lessening lowering dropping descending sinking submerging diving plunging plummeting crashing colliding impacting hitting striking knocking tapping patting touching sensing perceiving detecting discovering finding locating positioning placing putting setting laying standing sitting lying resting relaxing reposing sleeping dreaming imagining fantasizing daydreaming woolgathering praying worshiping revering venerating adoring loving nourishing feeding eating drinking consuming ingesting energizing powering fueling motivating inspiring stimulating exciting arousing awakening alerting warning cautioning informing notifying telling reporting announcing declaring stating asserting claiming contending proving revealing disclosing exposing uncovering unmasking unveiling opening unlocking unsealing unfastening untying unbinding unwrapping unpacking unloading emptying clearing cleaning washing bathing showering rinsing soaking steeping immersing drowning suffocating choking strangling throttling squeezing pressing centering aiming targeting zeroing aligning tuning calibrating balancing leveling equalizing stabilizing steady mooring docking landing touching-down alighting perching roosting nesting denning burrowing tunneling digging excavating mining quarrying extracting removing eliminating eradicating exterminating destroying demolishing ruining wrecking damaging harming injuring hurting wounding maiming crippling laming disabling incapacitating paralyzing stunning shocking surprising astonishing amazing astounding staggering stupefying dazing bewildering confusing puzzling perplexing baffling mystifying enigmatic cryptic obscure abstruse recondite esoteric arcane occult mysterious unknown unfamiliar strange odd peculiar curious queer weird bizarre fantastic extraordinary remarkable notable noteworthy significant important momentous consequential weighty grave serious crucial vital essential necessary required needed wanted desired wished hoped expected anticipated awaited looked-for searched-for sought hunted pursued chased tracked trailed followed shadowed stalked crept crawled climbed scaled ascended mounted rose lifted elevated raised hoisted heaved hauled pulled tugged dragged drawn attracted lured enticed tempted seduced charmed enchanted fascinated captivated engrossed absorbed immersed involved engaged occupied busy active energetic dynamic vigorous forceful powerful strong mighty potent effective efficient productive fruitful fertile fecund prolific abundant plentiful ample sufficient adequate enough satisfactory acceptable tolerable bearable endurable sustainable maintainable manageable controllable governable rule govern reign dominate command direct lead guide steer pilot navigate drive propel push thrust shove press squeeze compress condense concentrate aim point target zero align adjust tune calibrate level equalize standardize normalize stabilize fix secure anchor moor dock land touch-down alight perch roost nest den burrow tunnel dig excavate mine quarry extract remove eliminate eradicate exterminate destroy demolish ruin wreck damage harm injure hurt wound maim cripple lame disable incapacitate paralyze stun shock astonish amaze astound stagger stupefy daze bewilder confuse puzzle perplex baffle mystify enigma
Flexamia huroni
Huron River Leafhopper
Flexamia huroni is a leafhopper species in the family Cicadellidae, described by Bess & Hamilton in 1999. It belongs to the genus Flexamia, a group of leafhoppers known for their specialized host plant associations with grasses. The species is named after the Huron River in Michigan, where it was first collected. Like other members of the genus, it likely exhibits strong ecological dependence on specific grass host plants.
leafhopper cicadellidae deltocephalinae paralimnini flexamia grass-specialist michigan-endemic auchenorrhyncha hemiptera insecta arthropoda animalia true-bug planthopper-relative 1999-description bess hamilton huroni huron-river usa north-america grassland-insect host-specific poorly-known rare uncommon data-deficient gbif catalogue-of-life ncbi inaturalist taxon species accepted hexapoda cicadomorpha clypeata membracoidea paralimnina flexamia-huroni bess-&-hamilton 1999 exact-match accepted-name canonical-name scientific-name authorship rank status matched taxonomy classification eukaryota metazoa distribution michigan observations 0 wikipedia none preferred-common-name huron-river-leafhopper true-bugs group kingdom phylum class order family genus authority ipt integrated-publishing-toolkit biodiversity-data-journal zookeys nature-conservation comparative-cytogenetics open-access open-access journal publication dataset specimen type nomenclatural-type herbarium university-of-granada spain fungi lichens agaricales cortinarius antonio-ortega mediterranean france italy image-collection colección-de-imágenes-de-los-tipos-nomenclaturales-de-hongos líquenes musgos-y-algas gda gdac vizoso quesada 2015 doi 10.3897 bdj 3 e5204 new-species new-jersey-pine-barrens muhlenbergia-torreyana pinebarren-smokegrass threatened-species andrew-hicks museum-of-natural-history university-of-colorado gerry-moore natural-resources-conservation-service greensboro nc uli-lorimer brooklyn-botanic-garden f.-whitcombi robert-whitcomb microbiology ornithology ecology host-plant warming-climate human-activities zookeys-511 69-79 zookeys.511.9572 roundworm nematode antarctic amblydorylaimus-isokaryoni pararhyssocolpus-paradoxus bulgaria scanning-electron-microscopy sem maritime-antarctic antarctic-islands lip-region spear vulva postembryonic-development molecular-analyses dorylaimida elshishka lazarova radoslavov hristov peneva 25-68 zookeys.511.9793 anidiv2 bulgarian-academy-of-sciences national-scientific-fund octocoral okinawa japan nanipora-kamurai living-fossil blue-coral heliopora aragonite-calcium-carbonate skeleton scleractinians soft-coral heliporacea lithotelestidae epiphaxum deep-sea shallow-coral-reef zamami-island national-park miyazaki reimer 1-23 zookeys.511.9432 non-biting-midge chironomus ch.-bernensis north-caucasus russia caucasian-populations europe siberia karyotype morphology mouthparts larvae chromosome genotypic-combinations mineralization eutrophication karmokov polukonova sinichkina tembotov-institute-of-ecology-of-mountain-territories saratov-state-medical-university comparative-cytogenetics-9 281-297 compcytogen.v9i3.4519 sea-turtle rescue-centre first-aid-station loggerhead green-turtle caretta-caretta chelonia-mydas bycatch mortality greece migration sexual-maturity ullmann stachowitsch uit-the-arctic-university-of-norway nature-conservation-10 45-69 natureconservation.10.4890 regional-activity-centre-for-specially-protected-areas porcupine coendou-ichillus lower-urubamba peru canopy-bridge pipeline natural-gas arboreal camera-trap dwarf-porcupine iquitos 770g gregory lunde zamora-meza carrasco-rueda repsol-exploración-perú zookeys-509 109-121 zookeys.509.9821 ant prionopelta madagascar seychelles subterranean leaf-litter dracula-ant hemolymph larval-hemolymph-feeding oophagy madagascar-biodiversity-center overson fisher zookeys-507 115-150 zookeys.507.9303 ito billen masuko spider anelosimus subsocial cobweb-spider theridiidae deforestation biodiversity-hotspot agnarsson university-of-vermont smithsonian-national-museum-of-natural-history wallace huxley buffon hooker lamarck darwin moramora eriophyoid-mite acari xinjiang china rosaceae paracolomerus gall ji-wei-li wang xue zhang zookeys-508 97-111 zookeys.508.8940 shihezi-university grasshopper wyoming melanoplus melanoplinae acrididae tetrigidae gomphocerniae oedipodinae cyrtacanthacridinae distribution-atlas field-guide wgis wyoming-grasshopper-information-system key capinera sechrist hebard helfer scudder blatchley thomas say harris degeer bruner saussure girard dodge walker fieber fabricius serville mcneill tinkham burmeister haldeman big-horn-mountains black-hills gladston indigens infantilis dodgei oregonensis marshalli yellowstone-national-park sagebrush pine elevation shortgrass-prairie mixedgrass-prairie forb grass economic-damage rangeland benefit overwinter egg hatch adult late-summer august october june life-cycle food-habit size collection survey underreported common endemic restricted-range forest-opening grassy moderate-elevation larger smaller east west united-states america north south central range extent limited restricted abundant population density occurrence presence absence habitat environment condition altitude topography terrain vegetation plant shrub tree forest opening meadow prairie steppe savanna woodland draw slope aspect soil substrate moisture temperature climate weather season phenology timing activity nymph emerge molt develop grow reproduce mate oviposit die generation voltinism univoltine bivoltine multivoltine semivoltine diapause aestivation hibernation dispersal movement behavior habit action feeding diet food host association relationship interaction specialist generalist monophagy oligophagy polyphagy herbivore detritivore predator parasitoid scavenger economic-importance pest beneficial neutral damage control management conservation threatened endangered vulnerable secure unknown global-biodiversity-information-facility es biodiversity-image-portal spanish-collections type-specimen lichen antarctica bernensis li yellowstone
Ixodes affinis
Ixodes affinis is a hard-bodied tick species distributed across the southeastern United States, Mexico, Central America, and South America. Unlike its congener Ixodes scapularis, this species does not bite humans but serves as a competent vector for Borrelia burgdorferi sensu stricto, the causative agent of Lyme disease. Immature stages feed on birds and small mammals, while adults parasitize larger mammals including deer and dogs. Its expanding range into southeastern Virginia and association with migratory birds has raised concerns about its role in pathogen dispersal.
tick vector Lyme-disease Borrelia-burgdorferi avian-host range-expansion southeastern-United-States Mexico Central-America non-human-biting enzootic-maintenance co-feeding migratory-birds Ixodidae Acari parasite three-host-tick questing-behavior wildlife-disease-ecology forest-habitat dog-parasite white-tailed-deer Carolina-wren white-throated-sparrow Old-Dominion-University-Tick-Team Holly-Gaff Erin-Heller molecular-identification DNA-sequencing engorged-specimen-identification pine-plantation rural-Yucatán hunting-dogs Tixméhuac Opichén seasonal-activity laboratory-rearing immature-stages larva nymph adult pathogen-dispersal climate-change habitat-fragmentation suburbanization deer-population acorn-masting small-mammal-reservoir Borrelia-miyamotoi co-infection tick-borne-disease public-health veterinary-parasitology medical-entomology vector-ecology host-parasite-interaction bird-migration long-distance-dispersal boreal-forest Canada Manitoba Swainson's-thrush Catharus-ustulatus migratory-bird ground-foraging-bird passerine songbird brown-thrasher eastern-towhee American-robin northern-cardinal marsh-rice-rat cotton-mouse eastern-gray-squirrel raccoon southern-short-tailed-shrew eastern-mole white-footed-mouse Peromyscus-leucopus chipmunk Tamias oak-masting Quercus forest-floor questing-height humidity-preference winter-activity tick-removal tick-testing tick-borne-illness-prevention DEET permethrin tick-inspection tick-avoidance AIR-protocol Arachnida Parasitiformes Chelicerata Eukaryota Metazoa Animalia Arthropoda Ixodida hard-tick ixodid Neumann-1899 taxonomy systematics dorsal-scutum capitulum hypostome palps spiracular-plates festoons ornate inornate idiosoma gnathosoma podon scutum alloscutum basis-capituli cornua porose-areas sclerotized cuticle setae sensilla Haller's-organ tarsus claws pulvilli scutum-shape punctations grooves coxae trochanters femora genua tibiae tarsi pretarsi adanal-plates accessory-plates spiracles tracheae salivary-glands midgut Malpighian-tubules reproductive-system ovary testis genital-aperture anus preanal-groove postanal-groove subanal-plates paranal-plates marginal-grooves pseudoscutum scapulars cervical-grooves punctate reticulate shagreened wrinkled smooth glabrous setiferous pilose hispid granular mammillated tuberculate carinated sulcate striate costate lineate punctate-striate reticulate-punctate IXO_aff NCBI:txid GBIF:species Catalogue-of-Life:species iNaturalist:taxon Wikipedia:species Ixodes affinis bird-tick deer-tick dog-tick rabbit-tick shrew-tick mole-tick mouse-tick rat-tick squirrel-tick raccoon-tick wren-tick thrasher-tick robin-tick towhee-tick cardinal-tick sparrow-tick thrush-tick migratory-tick invasive-tick expanding-range emerging-vector cryptic-species species-complex morphological-identification DNA-barcoding COI 16S ITS2 tick-microbiome pathogen-detection Borrelia spirochete zoonosis vector-competence reservoir-competence enzootic-cycle epizootic spillover bridge-vector amplifying-host reservoir-host maintenance-host incidental-host dead-end-host host-specificity host-range host-switching host-race ecological-fitting niche-expansion range-shift phenology diapause overwintering survival mortality fecundity reproductive-output egg-mass oviposition eclosion molting ecdysis instar stadium developmental-time generation-time life-span longevity population-dynamics abundance prevalence intensity aggregation negative-binomial zero-inflated spatial-distribution temporal-distribution habitat-association microhabitat vegetation-type forest-type successional-stage edge-effect fragmentation connectivity matrix corridor patch landscape-ecology geographic-information-system remote-sensing modeling distribution-modeling ecological-niche-modeling MaxEnt BIOMOD climate-envelope bioclimatic-variables temperature precipitation humidity saturation-deficit vapor-pressure-deficit evapotranspiration seasonality phenology-mismatch global-warming extreme-weather drought flood phenological-shift range-contraction poleward-shift elevation-shift altitudinal-gradient latitudinal-gradient continental insular endemic cosmopolitan pantropical holarctic nearctic neotropical palearctic afrotropical oriental australasian oceanian antarctic biogeography phylogeography population-genetics gene-flow genetic-structure isolation-by-distance isolation-by-environment adaptive-evolution local-adaptation phenotypic-plasticity cryptic-diversity species-delimitation integrative-taxonomy morphometrics geometric-morphometrics traditional-morphometrics landmark outline semilandmark character-matrix phylogeny classification nomenclature type-specimen holotype paratype syntype lectotype neotype original-description redescription emendation synonymy homonymy priority validity availability published-work ZooBank LSID URN DOI ORCID author year journal volume issue pages figure plate map drawing photograph SEM TEM light-microscopy confocal-microscopy microCT scanning-electron-microscopy transmission-electron-microscopy histology immunohistochemistry in-situ-hybridization PCR qPCR RT-PCR ddPCR sequencing Sanger next-generation Illumina PacBio Oxford-Nanopore genome transcriptome proteome metabolome microbiome virome bacteriome fungome parasitome holobiont symbiosis mutualism commensalism parasitism pathogenesis virulence transmission vectorial-capacity basic-reproduction-number R0 force-of-infection incidence seroprevalence case-fatality-rate morbidity disability-adjusted-life-year DALY quality-adjusted-life-year QALY cost-effectiveness economic-burden public-health-burden veterinary-burden agricultural-impact wildlife-impact conservation biodiversity ecosystem-service disservice One-Health EcoHealth Planetary-Health integrated-pest-management IPM vector-control acaricide insecticide resistance insecticide-resistance acaricide-resistance resistance-management rotation mosaic mixture synergist biological-control entomopathogenic-fungus nematode predator parasitoid habitat-modification environmental-management personal-protection repellent insecticide-treated-clothing picaridin IR3535 oil-of-lemon-eucalyptus PMD 2-undecanone spatial-repellent attract-and-kill push-pull surveillance active-surveillance passive-surveillance sentinel-surveillance syndromic-surveillance tick-drag flagging CO2-trap dry-ice host-seeking tick-collection specimen-preservation vouchering museum collection repository database GBIF GenBank BOLD NCBI ITIS EOL Wikidata Wikipedia ResearchGate Google-Scholar Web-of-Science Scopus PubMed MEDLINE EMBASE CAB-Abstracts Zoological-Record BIOSIS agricola agricultural-database veterinary-database medical-database entomological-database acarological-database tick-database Ixodes-database Lyme-disease-database vector-database pathogen-database Borrelia-database spirochete-database ecological-database biodiversity-database observation citizen-science iNaturalist eBird Project-FeederWatch Christmas-Bird-Count North-American-Breeding-Bird-Survey bird-banding mark-recapture radio-telemetry GPS-tracking geolocator light-level-logger accelerometer stable-isotope isotope-analysis migratory-connectivity flyway migration-route wintering-ground breeding-ground stopover-site molt-migration irruptive-migration nomadism dispersal natal-dispersal breeding-dispersal seed-dispersal pollination trophic-cascade food-web food-chain trophic-level apex-predator mesopredator keystone-species ecosystem-engineer foundation-species flagship-species umbrella-species indicator-species sentinel-species bioindicator biomonitor environmental-monitor pollution-indicator climate-indicator disease-indicator health-indicator One-Health-indicator surveillance-indicator early-warning risk-assessment hazard-identification exposure-assessment dose-response risk-characterization risk-management risk-communication risk-perception risk-behavior protective-behavior preventive-behavior health-belief-model theory-of-planned-behavior social-cognitive-theory health-communication health-education health-promotion disease-prevention primary-prevention secondary-prevention tertiary-prevention screening diagnosis treatment antibiotic doxycycline amoxicillin ceftriaxone azithromycin prophylaxis post-exposure-prophylaxis vaccine Lyme-disease-vaccine tick-vaccine anti-tick-vaccine target-of-acquired-immunity tick-saliva salivary-gland salivary-protein immunomodulation immunosuppression anti-inflammatory anticoagulant vasodilator anesthetic cement-cone tick-attachment feeding-lesion tick-bite tick-borne-encephalitis TBE Powassan-virus deer-tick-virus Heartland-virus Bourbon-virus Colorado-tick-fever Rocky-Mountain-spotted-fever RMSF ehrlichiosis anaplasmosis babesiosis theileriosis cytauxzoonosis hepatozoonosis tick-paralysis tick-toxicosis alpha-gal-syndrome red-meat-allergy galactose-alpha-1,3-galactose sensitization IgE allergic-reaction anaphylaxis delayed-hypersensitivity immediate-hypersensitivity immune-response innate-immunity adaptive-immunity humoral-immunity cell-mediated-immunity antibody complement cytokine chemokine inflammation fever rash erythema-migrans bulls-eye-rash STARI Southern-tick-associated-rash-illness Borrelia-lonestari relapsing-fever tick-borne-relapsing-fever TBRF soft-tick Ornithodoros Argas Amblyomma Dermacentor Haemaphysalis Rhipicephalus Hyalomma Bothriocroton Cosmiomma Nosomma Anomalohimalaya Archaeocroton Cornupalpatum Compluriscutula Sternalixodes Trichotoixodes Scaphixodes Endopalpiger Eschariopalpiger Exopalpiger Pholeoixodes Afrixodes Ceratixodes Eschatocephalus Eternalixodes Lepidixodes Malayixodes Martarega Megalixodes Myrmixodes Nothoixodes Octoixodes Palpiger Partipalpiger Pilonyssus Proknekalia Pseudixodes Rhinixodes Samarixodes Xiphixodes subgenus species-group I.-ricinus-complex I.-persulcatus-complex I.-scapularis-complex I.-pacificus-complex I.-affinis-group I.-cookei-group I.-marxi-group I.-angustus-group I.-soricis-group I.-uriae-group I.-holocyclus-group I.-ceylonensis-group I.-granulatus-group I.-ovatus-group I.-nipponensis-group I.-persulcatus-group I.-pavlovskyi-group I.-ricinus-group I.-scapularis-group I.-pacificus-group I.-affinis I.-auritulus I.-australiensis I.-boliviensis I.-brunneus I.-cookei I.-cooleyi I.-cornuae I.-cornuatus I.-crenulatus I.-dammini I.-dawsoni I.-dentatus I.-diomedeae I.-domerguei I.-eadsi I.-eastoni I.-eldaricus I.-elongatus I.-eudyptidis I.-euplectropi I.-evansi I.-fecialis I.-festai I.-frontalis I.-galapagoensis I.-ghilarovi I.-gibbosus I.-granulatus I.-gregsoni I.-hearlei I.-hexagonus I.-hirsti I.-holocyclus I.-hoogstraali I.-howelli I.-hyatti I.-jacksoni I.-jellisoni I.-kashmiricus I.-kazakstani I.-kingi I.-kohlsi I.-kopsteini I.-kuntzi I.-laguri I.-latus I.-layingensis I.-lewisi I.-lividus I.-longiscutatus I.-loricatus I.-luciae I.-malayensis I.-marxi I.-maslovi I.-matopi I.-minor I.-monospinosus I.-moreli I.-moschiferi I.-muris I.-murreleti I.-myospalacis I.-nchisiensis I.-necromys I.-neitzi I.-nipponensis I.-nuttalli I.-ochotonae I.-okapiae I.-oldi I.-ornithorhynchi I.-ovatus I.-pacificus I.-pararicinus I.-pavlovskyi I.-percavatus I.-peromysci I.-persulcatus I.-pomerantzi I.-pterodromae I.-quadratus I.-raccoonis I.-rageaui I.-randrianasoloi I.-rasus I.-redikorzevi I.-ricinus I.-rothschildi I.-rotundatus I.-rubicundus I.-rugicollis I.-rugosus I.-sapeloensis I.-sarsi I.-scapularis I.-schillingsi I.-sculptus I.-semenovi I.-soricis I.-spinae I.-steini I.-stilesi I.-stromi I.-subterranus I.-tancitarius I.-tanuki I.-tapirus I.-tasmani I.-taylori I.-tertiarius I.-texanus I.-theilerae I.-thomasae I.-tovari I.-transvaalensis I.-trianguliceps I.-trichosuri I.-turdus I.-uriae I.-vandenbergi I.-vanidicus I.-ventalloi I.-vespertilionis I.-victoriensis I.-walkerae I.-werneri I.-woodi I.-zaglossi I.-zairensis tick-taxonomy tick-systematics tick-phylogeny tick-evolution tick-biogeography tick-ecology tick-physiology tick-behavior tick-reproduction tick-development tick-nutrition tick-metabolism tick-respiration tick-excretion tick-osmoregulation tick-neurobiology tick-sensory-biology tick-chemoreception tick-thermoreception tick-mechanoreception tick-photoreception tick-circadian-rhythm tick-biological-clock tick-seasonality tick-diapause tick-overwintering tick-cold-hardiness tick-desiccation-resistance tick-water-balance tick-cuticle tick-molting tick-ecdysis tick-growth tick-feeding tick-engorgement tick-blood-meal tick-digestion tick-salivation tick-saliva-composition tick-salivary-gland-transcriptome tick-salivary-proteome tick-pharmacology tick-toxicology tick-venom tick-paralysis-toxin tick-borne-pathogen tick-borne-virus tick-borne-bacterium tick-borne-protozoan tick-borne-helminth tick-borne-rickettsia tick-borne-ehrlichia tick-borne-anaplasma tick-borne-borrelia tick-borne-babesia tick-borne-theileria tick-borne-hepatozoon tick-borne-coxiella tick-borne-francisella tick-borne-bartonella tick-borne-chlamydia tick-borne-mycoplasma tick-borne-spiroplasma tick-borne-arbovirus tick-borne-flavivirus tick-borne-bunyavirus tick-borne-reovirus tick-borne-orbivirus tick-borne-coltivirus tick-borne-nairovirus tick-borne-phlebovirus tick-borne-thogotovirus tick-borne-orthomyxovirus tick-borne-arenavirus tick-borne-hantavirus tick-borne-paramyxovirus tick-borne-rhabdovirus tick-borne-bornavirus tick-borne-filovirus tick-borne-coronavirus tick-borne-poxvirus tick-borne-herpesvirus tick-borne-adenovirus tick-borne-papillomavirus tick-borne-polyomavirus tick-borne-circovirus tick-borne-parvovirus tick-borne-anellovirus tick-borne-dependovirus tick-borne-tenuivirus tick-borne-deltavirus tick-borne-hepadnavirus tick-borne-retrovirus tick-borne-lentivirus tick-borne-spumavirus tick-borne-metavirus tick-borne-pseudovirus tick-borne-hemivirus tick-borne-novirhabdovirus tick-borne-peribunyavirus tick-borne-phenuivirus tick-borne-bandavirus tick-borne-phasivirus tick-borne-tulasvirus tick-borne-orthonairovirus tick-borne-shavivirus tick-borne-sathuperi-virus tick-borne-salehabad-virus tick-borne-dera-ghazi-khan-virus tick-borne-hughes-virus tick-borne-farallon-virus tick-borne-punta-salinas-virus tick-borne-sapphire-II-virus tick-borne-zirqa-virus tick-borne-keterah-virus tick-borne-thiafora-virus tick-borne-erve-virus tick-borne-malsoor-virus tick-borne-tibet-virus tick-borne-qalyub-virus tick-borne-yogue-virus tick-borne-kasokero-virus tick-borne-lebombo-virus tick-borne-nyamanini-virus tick-borne-midway-virus tick-borne-tokar-virus tick-borne-absettarov-virus tick-borne-hypr-virus tick-borne-kumlinge-virus tick-borne-omsk-hemorrhagic-fever-virus tick-borne-langat-virus tick-borne-louping-ill-virus tick-borne-kyasanur-forest-disease-virus tick-borne-alkhurma-hemorrhagic-fever-virus tick-borne-powassan-virus tick-borne-deer-tick-virus tick-borne-heartland-virus tick-borne-bourbon-virus tick-borne-colorado-tick-fever-virus tick-borne-eyach-virus tick-borne-banna-virus tick-borne-seadornavirus tick-borne-great-island-virus tick-borne-kemerovo-virus tick-borne-lipovnik-virus tick-borne-tribec-virus tick-borne-chobar-gorge-virus tick-borne-wad-Medani-virus tick-borne-dugbe-virus tick-borne-crimean-congo-hemorrhagic-fever-virus tick-borne-nairobi-sheep-disease-virus tick-borne-ganjam-virus tick-borne-bandia-virus tick-borne-sierra-nevada-virus tick-borne-lone-star-virus tick-borne-severe-fever-with-thrombocytopenia-syndrome-virus tick-borne-huaiyangshan-virus tick-borne-guertu-virus tick-borne-Alongshan-virus tick-borne-wenzhou-virus tick-borne-wuhan-virus tick-borne-yongjia-virus tick-borne-chiba-virus tick-borne-natori-virus tick-borne-odawara-virus tick-borne-kowloon-virus tick-borne-tamanavirus
Lacunicambarus diogenes
devil crayfish, devil crawfish
Lacunicambarus diogenes, commonly known as the devil crayfish or devil crawfish, is a primary burrowing crayfish native to eastern North America. This species constructs and inhabits burrows in wet, muddy terrestrial habitats rather than living in permanent surface water. Its burrowing activities create refugia used by numerous other species, including documented use by eastern cicada killer wasps (Sphecius speciosus) as brooding habitat. The species ranges across the Atlantic Coastal Plain and Piedmont ecoregion from New Jersey to Georgia, with disjunct populations in Louisiana.
crayfish burrowing ecosystem-engineer primary-burrower Atlantic-Coastal-Plain terrestrial fossorial North-America Cambaridae Decapoda crustacean endemic wetland floodplain refugia allogenic-engineer cave-crayfish Lacunicambarus Cambarus Girard-1852 devil-crayfish devil-crawfish Sphecius-speciosus cicada-killer-wasp brooding-habitat soil-type congeneric-cues floodplain-connectivity muddy-substrate desiccation-refuge predation-refuge intermittent-water Piedmont New-Jersey Pennsylvania Delaware Maryland Virginia North-Carolina South-Carolina Georgia Louisiana roadside-ditch terrestrial-habitat burrow-construction burrow-maintenance wetland-engineer crustacean-ecology decapod astacidea freshwater-invertebrate invertebrate-engineer habitat-modification species-interactions commensalism facilitation ecosystem-services biodiversity-support habitat-provision microhabitat-creation soil-bioturbation sediment-mixing water-infiltration groundwater-connection ephemeral-water seasonal-wetland palustrine lotic lentic riparian hyporheic bioturbation soil-structure nutrient-cycling detritivore omnivore scavenger predator prey trophic-interaction food-web community-ecology population-ecology conservation habitat-loss urbanization agriculture land-use-change climate-change drought hydrology geomorphology sedimentology ichnology trace-fossil burrow-architecture burrow-morphology tunnel-system chimney mud-pellet cast excavation sediment-ejection maintenance-behavior occupancy site-fidelity home-range territoriality intraspecific-interaction interspecific-interaction competition predation-risk desiccation-risk physiological-stress behavioral-adaptation morphological-adaptation fossorial-adaptation gill-function air-breathing cutaneous-respiration ion-regulation osmoregulation excretion metabolism energetics life-history growth molting reproduction mating egg-care juvenile-development recruitment population-dynamics demography genetics phylogeography biogeography dispersal colonization extinction rare-species common-species indicator-species keystone-species umbrella-species flagship-species natural-capital environmental-economics environmental-policy environmental-management restoration-ecology habitat-restoration species-reintroduction ex-situ-conservation in-situ-conservation protected-area wildlife-refuge nature-reserve national-park state-park conservation-easement stewardship citizen-science iNaturalist observation specimen voucher museum-collection type-specimen holotype taxonomy systematics nomenclature synonymy homonymy priority valid-name accepted-name species-concept morphological-species biological-species evolutionary-species phylogenetic-species integrative-taxonomy DNA-barcoding molecular-systematics phylogenetics phylogeny evolution adaptation natural-selection speciation diversification radiation endemism vicariance dispersal-limitation habitat-specificity niche-breadth niche-conservatism ecological-release character-displacement competitive-exclusion resource-partitioning habitat-filtering environmental-gradient elevation latitude longitude climate temperature precipitation hydroperiod soil-texture soil-moisture soil-chemistry pH organic-matter nutrient-availability productivity disturbance succession metacommunity landscape-ecology spatial-ecology movement-ecology dispersal-kernel connectivity corridor barrier fragmentation matrix edge-effect core-area patch-dynamics source-sink metapopulation population-viability minimum-viable-population effective-population-size inbreeding genetic-drift gene-flow local-adaptation phenotypic-plasticity genotype-environment-interaction epigenetics developmental-biology embryology larval-development post-larval-development juvenile-stage adult-stage senescence longevity survivorship fecundity reproductive-output reproductive-success fitness selection-coefficient heritability quantitative-genetics population-genetics molecular-ecology conservation-genetics genomic-resources transcriptomics proteomics metabolomics phenomics bioinformatics data-science machine-learning artificial-intelligence remote-sensing geographic-information-system spatial-analysis statistical-modeling experimental-design sampling-design field-methods laboratory-methods specimen-preparation curation data-management data-sharing open-science reproducibility replicability transparency ethics animal-welfare animal-use permit regulation policy legislation environmental-law natural-resource-law water-law land-use-planning environmental-impact-assessment cumulative-impact-assessment strategic-environmental-assessment adaptive-management monitoring evaluation learning stakeholder-engagement public-participation environmental-education science-communication outreach extension capacity-building professional-development career-development mentorship collaboration interdisciplinary transdisciplinary team-science open-innovation knowledge-exchange technology-transfer societal-impact sustainability sustainable-development green-economy blue-economy circular-economy nature-based-solution ecosystem-based-management integrated-water-resource-management watershed-management river-basin-management flood-risk-management drought-management climate-change-adaptation climate-change-mitigation resilience vulnerability risk-assessment scenario-planning forecasting prediction early-warning decision-support evidence-based-policy science-policy-interface boundary-organization knowledge-broker research-network professional-society academic-society scientific-journal peer-review publication citation impact-factor altmetrics research-evaluation research-funding grant fellowship scholarship award recognition career-award early-career mid-career senior-career emeritus retirement legacy historical-ecology paleoecology archaeology anthropology indigenous-knowledge traditional-ecological-knowledge local-knowledge community-science participatory-research action-research transformational-research transformative-change system-change paradigm-shift future-thinking foresight horizon-scanning trend-analysis emerging-issue research-priority grand-challenge societal-challenge sustainable-development-goal biodiversity-target conservation-target protected-area-target restoration-target climate-target emission-reduction carbon-sequestration blue-carbon green-carbon nature-conservation wildlife-conservation species-conservation habitat-conservation ecosystem-conservation landscape-conservation seascape-conservation connectivity-conservation corridor-conservation transboundary-conservation international-cooperation global-governance environmental-governance multilateral-environmental-agreement convention treaty protocol Cartagena-Protocol Nagoya-Protocol Aichi-Target Kunming-Montreal-Global-Biodiversity-Framework 30x30 half-earth nature-needs-half rewilding reintroduction de-extinction synthetic-biology gene-drive biotechnology nanotechnology geoengineering solar-radiation-management carbon-capture direct-air-capture bioenergy-with-carbon-capture negative-emission climate-intervention earth-system planetary-boundary safe-operating-space tipping-point hysteresis irreversibility committed-change sea-level-rise ocean-acidification deoxygenation warming extreme-event compound-risk cascading-risk systemic-risk existential-risk global-catastrophic-risk risk-management disaster-risk-reduction emergency-response humanitarian-action relief recovery reconstruction build-back-better resilient-infrastructure nature-based-infrastructure green-infrastructure sponge-city cooling-city urban-heat-island urban-ecology urban-biodiversity urban-conservation urban-planning urban-design architecture landscape-architecture green-building sustainable-building energy-efficiency renewable-energy clean-energy energy-transition just-transition energy-justice climate-justice environmental-justice social-justice equity fairness inclusion diversity gender youth intergenerational-equity future-generation rights-of-nature personhood legal-standing guardianship care ethics-of-care environmental-ethics conservation-ethics animal-ethics bioethics research-ethics integrity responsible-conduct misconduct fraud plagiarism retraction correction erratum expression-of-concern data-fabrication data-falsification data-manipulation image-manipulation duplicate-publication salami-slicing guest-authorship gift-authorship ghost-authorship author-contribution credit attribution acknowledgment funding-disclosure conflict-of-interest competing-interest institutional-review animal-care biosafety biosecurity dual-use export-control sanction embargo boycott divestment activism advocacy campaign movement social-movement environmental-movement conservation-movement climate-movement youth-movement student-movement indigenous-movement feminist-movement labor-movement peace-movement health-movement food-movement agriculture-movement water-movement energy-movement transport-movement housing-movement urban-movement rural-movement local-movement global-movement network coalition alliance partnership cooperation solidarity mutual-aid community-organizing grassroots bottom-up top-down hybrid-governance polycentric-governance self-organization emergence complexity system-dynamics agent-based-model network-analysis social-network ecological-network mutualistic-network antagonistic-network multiplex-network multilayer-network temporal-network spatial-network network-metric centrality modularity nestedness connectance robustness stability persistence coexistence biodiversity-ecosystem-function invasibility invasiveness impact risk pathway vector introduction establishment spread eradication containment control management phytosanitary veterinary human-health One-Health EcoHealth Planetary-Health One-Welfare sustainability-science conservation-science restoration-science resilience-science adaptation-science transformation-science futures-studies backcasting projection simulation modeling numerical-model conceptual-model statistical-model mechanistic-model process-based-model individual-based-model system-dynamics-model machine-learning-model deep-learning neural-network random-forest support-vector-machine gradient-boosting ensemble-method model-selection model-validation model-calibration sensitivity-analysis uncertainty-analysis risk-analysis decision-analysis optimization multi-objective-optimization Pareto-frontier trade-off synergy co-benefit win-win lose-lose win-lose zero-sum positive-sum negative-sum game-theory cooperative-game non-cooperative-game Nash-equilibrium Pareto-optimality social-dilemma tragedy-of-the-commons prisoner's-dilemma collective-action free-rider public-good common-pool-resource institution rule norm custom law governance administration implementation enforcement compliance accountability participation inclusiveness responsiveness effectiveness efficiency legitimacy adaptability transformability innovation experimentation adaptive-governance evidence-based science-based knowledge-based information-based data-driven model-driven theory-driven hypothesis-driven question-driven problem-driven solution-driven impact-driven outcome-driven output-driven input-driven process-driven value-driven mission-driven vision-driven stakeholder-driven user-driven citizen-driven community-driven place-based ecosystem-based landscape-based seascape-based watershed-based river-basin-based bioregion-based ecoregion-based biome-based realm-based planetary-based global regional national subnational local municipal community household individual scale cross-scale multi-scale trans-scale upscaling downscaling scaling-out scaling-up scaling-deep mainstreaming integration maintenance durability permanence reversibility additionality leakage spillover externality market-failure government-failure institutional-failure system-failure surprise novelty critical-transition regime-shift alternative-stable-state basin-of-attraction engineering-resilience ecological-resilience socio-ecological-resilience evolutionary-resilience transformative-resilience specified-resilience general-resilience resistance transformation transition trajectory scenario narrative storyline vision future utopia dystopia protopia heterotopia archipelago mosaic patchwork collage assemblage bricolage montage palimpsest layer stratum sediment memory heritage tradition culture identity belonging home place space territory land soil water air fire element matter energy information code signal sign symbol meaning significance value worth price cost benefit utility preference choice decision behavior action practice habit ritual ceremony celebration festival performance art aesthetics beauty sublime wonder awe curiosity inquiry discovery exploration adventure expedition voyage journey pilgrimage quest odyssey epic saga story history paleontology geology climatology meteorology oceanography limnology ecology biology zoology botany microbiology development physiology anatomy morphology ethology sociobiology ecological-psychology environmental-psychology conservation-psychology science-education informal-education non-formal-education lifelong-learning adult-learning skill-building competency expertise mastery craft artisan maker creator innovator entrepreneur leader manager administrator policymaker practitioner researcher scholar academic scientist naturalist explorer observer recorder documenter storyteller communicator educator mentor coach facilitator mediator negotiator diplomat advocate activist campaigner organizer mobilizer networker connector bridge-builder boundary-spanner translator interpreter ambassador representative delegate spokesperson champion hero role-model inspiration motivation aspiration hope optimism pessimism realism idealism pragmatism practicality feasibility viability desirability acceptability credibility trust confidence reliability validity rigor quality excellence best-practice good-practice lesson-learned success-factor failure-factor risk-factor protective-factor determinant driver pressure state response DPSIR STEEP SWOT PESTLE scenario-matrix morphological-analysis Delphi-method expert-elicitation structured-expert-judgment citizen-deliberation participatory-modeling companion-modeling serious-game role-play virtual-reality augmented-reality mixed-reality immersive-experience sensory-experience embodied-experience affective-experience cognitive-experience social-experience cultural-experience spiritual-experience transcendent-experience transformative-experience learning-experience educational-experience research-experience professional-experience personal-experience lived-experience embodied-knowledge tacit-knowledge explicit-knowledge codified-knowledge procedural-knowledge declarative-knowledge conceptual-knowledge factual-knowledge theoretical-knowledge practical-knowledge applied-knowledge transdisciplinary-knowledge interdisciplinary-knowledge multidisciplinary-knowledge cross-disciplinary-knowledge disciplinary-knowledge subdisciplinary-knowledge specialist-knowledge generalist-knowledge holistic-knowledge systemic-knowledge reductionist-knowledge analytical-knowledge synthetic-knowledge integrative-knowledge creative-knowledge innovative-knowledge adaptive-knowledge resilient-knowledge sustainable-knowledge ethical-knowledge responsible-knowledge trustworthy-knowledge reliable-knowledge valid-knowledge sound-knowledge robust-knowledge antifragile-knowledge living-knowledge dynamic-knowledge evolving-knowledge emergent-knowledge generative-knowledge productive-knowledge reproductive-knowledge replicative-knowledge imitative-knowledge emulative-knowledge competitive-knowledge collaborative-knowledge cooperative-knowledge shared-knowledge common-knowledge public-knowledge open-knowledge free-knowledge accessible-knowledge inclusive-knowledge diverse-knowledge equitable-knowledge just-knowledge fair-knowledge democratic-knowledge participatory-knowledge citizen-knowledge community-knowledge traditional-knowledge ecological-knowledge place-based-knowledge experiential-knowledge intuitive-knowledge implicit-knowledge unconscious-knowledge subconscious-knowledge conscious-knowledge self-aware-knowledge reflexive-knowledge critical-knowledge emancipatory-knowledge transformative-knowledge transformational-knowledge revolutionary-knowledge evolutionary-knowledge developmental-knowledge growth-knowledge maturation-knowledge aging-knowledge senescence-knowledge death-knowledge extinction-knowledge persistence-knowledge resilience-knowledge recovery-knowledge restoration-knowledge renewal-knowledge regeneration-knowledge revitalization-knowledge rejuvenation-knowledge rebirth-knowledge renaissance-knowledge awakening-knowledge enlightenment-knowledge illumination-knowledge inspiration-knowledge aspiration-knowledge hope-knowledge optimism-knowledge pessimism-knowledge realism-knowledge idealism-knowledge pragmatism-knowledge practicality-knowledge feasibility-knowledge viability-knowledge desirability-knowledge acceptability-knowledge legitimacy-knowledge credibility-knowledge trust-knowledge confidence-knowledge reliability-knowledge validity-knowledge rigor-knowledge quality-knowledge excellence-knowledge best-practice-knowledge good-practice-knowledge lesson-learned-knowledge success-factor-knowledge failure-factor-knowledge risk-factor-knowledge protective-factor-knowledge determinant-knowledge driver-knowledge pressure-knowledge state-knowledge impact-knowledge response-knowledge DPSIR-knowledge STEEP-knowledge SWOT-knowledge PESTLE-knowledge scenario-matrix-knowledge morphological-analysis-knowledge Delphi-method-knowledge expert-elicitation-knowledge structured-expert-judgment-knowledge citizen-deliberation-knowledge participatory-modeling-knowledge companion-modeling-knowledge serious-game-knowledge role-play-knowledge simulation-knowledge virtual-reality-knowledge augmented-reality-knowledge mixed-reality-knowledge immersive-experience-knowledge sensory-experience-knowledge embodied-experience-knowledge affective-experience-knowledge cognitive-experience-knowledge social-experience-knowledge cultural-experience-knowledge spiritual-experience-knowledge transcendent-experience-knowledge transformative-experience-knowledge learning-experience-knowledge educational-experience-knowledge research-experience-knowledge professional-experience-knowledge personal-experience-knowledge lived-experience-knowledge
Liodessus
Liodessus is a genus of small diving beetles in the family Dytiscidae, tribe Bidessini. Species occur across the Americas from North America through the Andes to southern South America, with notable diversity in high-altitude wetlands and páramo ecosystems. Many species inhabit shallow, exposed pools and temporary water bodies, showing phenotypic plasticity in body form correlated with habitat permanence. The genus contains numerous species, with several new species described from the high Andes of Peru and Colombia in recent years. Taxonomic resolution relies heavily on male genital morphology.
diving-beetle aquatic-insect high-altitude páramo Andes phenotypic-plasticity Bidessini Hydroporinae Dytiscidae Coleoptera wetland temporary-pool peatland subantarctic taxonomy male-genitalia COI-barcode cryptic-species incipient-speciation hybridization geometric-morphometrics environmental-sentinel drought-indicator South-America North-America Fiji Navarino-Island Magallanes Peru Colombia Chile Argentina Bolivia Bogotá Junín Cusco Huánuco Cundinamarca Sumapaz Puna steppe peat-bog shallow-pool puddle altitude-3400-4900m L.-affinis-complex L.-bogotensis-complex L.-chilensis L.-affinis L.-obscurellus L.-noviaffinis L.-saratogae L.-picinus L.-santarosita L.-alpinus L.-hauthi L.-rhigos L.-thespesios Guignot-1939 Zimmermann Sharp LeConte Say Steinheil Régimbart Biström Miller Balke Franciscolo Sanfilippo Pederzani Nilsson Young Clark Hatch Mannerheim Casey Aubé Balfour-Browne Brinck Peschet Solier White Zootaxa ZooKeys Insect-Systematics-&-Evolution Anales-del-Instituto-de-la-Patagonia PLoS-ONE GBIF iNaturalist Catalogue-of-Life NCBI-Taxonomy BOLD Barcode-of-Life-Data-System COI mitochondrial-DNA phylogeography population-structure range-expansion lineage genetic-structure subspecies lectotype neotype synonymy new-species new-subspecies revision morphometrics allometry body-size body-shape elytra pronotum striae coloration punctation habitus occipital-line genital-morphology aedeagus paramere diagnosis identification-key type-locality type-specimen etymology bionomics natural-history geographic-distribution distribution-map Nearctic Neotropical Patagonia Cape-Horn-Biosphere-Reserve drought climate-change environmental-change sentinel-species plasticity adaptation phenotypic-variation hydrological-landscape water-permanence pool bog Altiplano Andean high-elevation montane alpine aquatic freshwater lentic lotic insect beetle Adephaga Liodessus
Mannophorus forreri
Mannophorus forreri is a longhorn beetle in the family Cerambycidae, described by Henry Walter Bates in 1885. It belongs to the tribe Trachyderini, a group known for often brightly colored and patterned species. The species is rarely encountered in collections and appears to have a restricted distribution in the southwestern United States and Mexico. Field observations indicate adults are active in early autumn and visit flowers of yellow composites in mountainous areas of Arizona.
Cerambycidae Trachyderini longhorn-beetle Arizona Mexico flower-visitor rare-species Henry-Walter-Bates 1885 orange-and-black-coloration Gutierrezia Heterotheca Thelesperma Kitt-Peak autumn-active diurnal pollinator Madrean-sky-islands montane xerophilic Coleoptera beetle insect arthropod Cerambycinae Mannophorus forreri Bates-1885 GBIF iNaturalist Catalogue-of-Life Wikipedia field-observation composite-flowers Asteraceae southwestern-United-States North-America Middle-America rarely-collected uncommonly-encountered attractive-coloration coleopterist-interest flower-feeding nectar-feeding pollen-feeding larval-host-unknown wood-boring diurnal-activity September early-autumn lower-mountain-slopes rocky-habitat dry-habitat mountainous submontane sky-island Madrean pollination trachyderine long-antennae sexual-dimorphism antennae-length orange black elytral-apex pronotal-spots cylindrical-body robust medium-sized quick-to-escape wary spiral-flight field-collection iReport Ted-MacRae Jeff-Huether 2019 Arizona-collecting Kitt-Peak-Road Gutierrezia-microcephala Heterotheca-fulcrata yellow-composites flower-patches roadsides slopes observation specimen voucher museum-collection scientific-name authorship taxon species genus family order class phylum kingdom animal longhorn cerambycid taxonomy classification accepted-name synonym distribution range geographic-range presence occurrence record iNaturalist-observation GBIF-record specimen-record literature-record field-record collecting-trip entomology entomologist coleopterist amateur-entomologist professional-entomologist beetle-collector insect-collector naturalist field-naturalist biologist ecologist pollination-biologist conservation biodiversity data-deficient poorly-known undercollected understudied research-needed larval-biology-unknown host-plant-unknown life-history-unknown seasonality-poorly-known distribution-incompletely-known habitat-specificity-unknown population-status-unknown conservation-status-unevaluated IUCN not-evaluated least-concern taxonomic-stability valid-name original-description type-specimen type-locality type-species genus-type tribe subfamily superfamily infraorder suborder polyphaga cucujiformia chrysomeloidea trachyderina mannophorus-forreri bates species-description taxonomic-description nomenclature binomial binomial-nomenclature scientific-nomenclature zoological-nomenclature ICZN International-Commission-on-Zoological-Nomenclature code zoological-code name latin-name binomial-name specific-epithet generic-name author authority date year original-combination current-combination valid-combination accepted-combination taxonomic-status nomenclatural-status availability validity priority homonym synonymy junior-synonym senior-synonym objective-synonym subjective-synonym replacement-name nomen-novum nomen-conservandum nomen-oblitum forgotten-name unused-name rarely-used-name obscure-name well-known-name famous-name iconic-species attractive-species beautiful-species ornamental-species collector's-item desirable-species sought-after-species target-species goal-species trip-target collecting-objective field-goal consolation-prize adequate-consolation great-find success lucky-find chance-encounter unexpected-find surprise-discovery serendipity fortuitous fortunate lucky good-fortune good-luck reward payoff satisfaction achievement accomplishment field-success collecting-success entomological-success beetle-success cerambycid-success trip-highlight field-trip-highlight collecting-trip-highlight memorable-find unforgettable-experience field-memory collecting-memory entomological-memory shared-experience collecting-partner field-companion joint-collecting collaborative-collecting team-collecting friendship camaraderie entomological-community collector-community amateur-community professional-community network connection relationship partnership sixth-joint-outing 2012-2019 seven-years long-term-collaboration repeated-collaboration trusted-partner reliable-partner experienced-collector knowledgeable-collector expert-collector skilled-collector talented-collector dedicated-collector passionate-collector enthusiastic-collector committed-collector serious-collector active-collector productive-collector successful-collector accomplished-collector respected-collector recognized-collector known-collector famous-collector noted-collector published-collector documented-collector recorded-collector cited-collector referenced-collector acknowledged-collector appreciated-collector valued-collector treasured-collector celebrated-collector honored-collector eminent-collector distinguished-collector renowned-collector illustrious-collector prestigious-collector acclaimed-collector celebrated famous well-known respected esteemed honored distinguished eminent renowned illustrious prestigious acclaimed noted recognized acknowledged appreciated valued treasured published documented recorded cited referenced mentioned featured highlighted showcased displayed exhibited presented shared communicated reported described narrated recounted related told written authored composed created produced generated made crafted prepared developed assembled compiled collected gathered accumulated amassed acquired obtained secured captured taken sampled specimens series individuals examples instances cases occurrences sightings observations records documentations evidences proofs confirmations verifications validations substantiations corroborations attestations testimonies witnesses accounts reports descriptions narratives stories tales chronicles histories annals archives repositories collections assemblages aggregations accumulations masses quantities numbers amounts volumes measures degrees levels extents scopes ranges spans stretches reaches spreads distributions dispersions scatterings sprinklings dottings specklings stipplings fleckings mottlings marblings streakings stripings bandings ringings zonings gradings shadings tintings tings tinges tingeings colorings colourings hues tones shades tints casts flavors flavours characters natures qualities properties attributes features traits characteristics aspects facets dimensions elements components constituents ingredients parts pieces sections segments divisions portions fractions fragments bits scraps remnants remains residues leftovers vestiges traces tracks marks signs indications tokens symbols emblems badges insignias seals stamps imprints impressions prints impresses depressions indentations impressures pressures compressions condensations concentrations densifications intensifications strengthenings fortifyings reinforcements supports backings foundations bases grounds substrates underpinnings cornerstones keystones capstones pinnacles peaks summits tops crowns crests ridges chains sequences successions progressions continuums spectrums scales gauges meters metrics standards benchmarks yardsticks touchstones criterions criteria indicators indexes indices signals markers pointers guides directors leaders chiefs heads principals main primary prime first foremost leading premier chief head principal central key critical crucial vital essential necessary needed required requisite indispensable mandatory obligatory compulsory forced coerced constrained obliged bound committed dedicated devoted loyal faithful true trustworthy reliable dependable responsible accountable answerable liable subject open exposed vulnerable susceptible prone likely apt inclined disposed predisposed tending leaning verging bordering nearing approaching coming advancing progressing moving going traveling journeying voyaging sailing flying driving riding walking hiking trekking marching tramping trudging plodding slogging toiling laboring working operating functioning running performing executing implementing enacting effecting bringing-about causing making creating producing generating yielding giving providing supplying furnishing delivering presenting offering extending proffering tendering submitting proposing suggesting recommending advising counseling guiding directing steering piloting navigating conducting escorting shepherding herding urging spurring goading prompting motivating stimulating inspiring encouraging supporting backing helping assisting aiding abetting facilitating easing smoothing simplifying clarifying explaining interpreting translating rendering converting transforming changing altering modifying adjusting adapting fitting suiting matching corresponding agreeing accord harmony concord consensus unanimity unity solidarity cohesion coherence consistency consonance congruence congruity compatibility agreement accordance conformity compliance observance adherence attachment devotion dedication commitment loyalty fidelity faithfulness constancy steadfastness staunchness resoluteness determination resolve decision conclusion judgment verdict finding ruling decree command directive instruction direction guidance leadership management administration governance government rule control domination dominion power might strength force vigor energy vitality life animation spirit soul mind intellect intelligence wisdom knowledge understanding comprehension grasp apprehension perception awareness consciousness cognizance recognition realization appreciation sensitivity responsiveness reactivity activity liveliness vividness intensity sharpness keenness acuity acuteness clearness distinctness plainness obviousness evidence manifestness patency transparency clarity lucidity pellucidity limpidity purity cleanliness cleanness spotlessness immaculateness flawlessness perfection excellence superiority supremacy preeminence eminence distinction note mark sign indication proof demonstration show display exhibition presentation manifestation expression utterance statement declaration announcement proclamation pronouncement assertion claim contention allegation accusation charge indictment imputation attribution ascription assignment allocation allotment apportionment dispensation division partition sharing participation involvement engagement fondness liking love affection tenderness warmth fervor ardor passion zeal enthusiasm eagerness ardency vehemence dynamism potency efficacy effectiveness efficiency competence capability capacity ability aptitude talent gift skill expertise proficiency mastery ascendancy predominance preponderance advantage edge lead head-start fore front van vanguard avant-garde frontier border boundary limit rim brink verge threshold doorstep precipice cliff steep abrupt sudden unexpected surprising astonishing amazing astounding stunning shocking startling jarring jolting rocking shaking disturbing upsetting disquieting unsettling troubling worrying concerning alarming frightening scaring terrifying horrifying appalling dreadful terrible awful horrible hideous ghastly gruesome grisly macabre morbid morbidity disease illness sickness ailment malady disorder condition state situation circumstance context environment setting scene background backdrop framework structure organization arrangement system scheme plan design pattern model template prototype archetype original initial primitive early ancient old aged elderly senior mature grown advanced progressed evolved derived descended originated begun started commenced initiated launched opened inaugurated instituted established founded set-up formed shaped fashioned molded cast forged wrought built constructed put-together erected raised elevated lifted hoisted heaved hauled dragged drawn pulled tugged yanked jerked snatched grabbed seized caught trapped snared netted hooked bagged landed got gained won earned achieved attained reached arrived come appeared emerged materialized manifested showed offered given granted bestowed conferred awarded gifted donated contributed divided split separated parted detached disconnected disjoined disunited divorced alienated estranged isolated segregated sequestered secluded withdrawn retired removed taken-away carried-off borne-away transported conveyed transferred transmitted imparted expressed articulated verbalized vocalized voiced spoken said depicted portrayed represented illustrated demonstrated shown revealed disclosed uncovered unmasked unveiled unfolded spread extended stretched spanned bridged crossed traversed passed gone-by elapsed expired lapsed ended finished completed concluded terminated closed shut sealed locked fastened made-safe protected guarded defended shielded screened sheltered harbored housed accommodated lodged quartered billeted stationed posted placed positioned located situated settled installed emplaced deposited laid put set arranged ordered organized structured systematized methodized rationalized streamlined simplified clarified explained interpreted translated rendered paraphrased rephrased reworded restate repeat reiterate rehearse recite quote cite refer allude mention remark observe comment declare announce proclaim pronounce assert affirm aver avow swear vow pledge promise commit engage bind oblige compel coerce constrain press urge push drive propel thrust shove squeeze compress condense concentrate focus center converge meet join unite combine merge fuse blend mix mingle intermingle intermix integrate incorporate embody contain include comprise consist compose constitute form make create produce generate yield give provide supply furnish deliver present offer extend proffer tender submit propose suggest recommend advise counsel guide direct steer pilot navigate conduct escort shepherd herd spur goad prompt motivate stimulate inspire encourage support back help assist aid abet facilitate ease smooth simplify clarify explain interpret translate render convert transform change alter modify adjust adapt fit suit match correspond agree harmonize conform comply adhere stick cling hold grip seize grab catch capture trap snare net hook bag land secure obtain acquire get gain win earn achieve attain reach arrive appear emerge materialize manifest exhibit grant bestow confer award donate contribute share participate involve dedicate devote attach fond like affect warm fervent ardent passionate zealous enthusiastic eager keen intense vehement forceful vigorous energetic dynamic powerful strong mighty potent effective efficient competent capable able talented skilled expert proficient masterful commanding controlling dominating ascendant supreme predominant preponderant superior advantageous progressive forward ahead onward upward rising ascending climbing mounting scaling surmounting overcoming conquering defeating vanquishing subduing subjugating mastering governing managing administering according harmonizing conforming complying observing adhering sticking clinging holding grasping gripping seizing grabbing catching capturing trapping snaring netting hooking bagging landing securing obtaining acquiring getting gaining winning earning achieving attaining reaching arriving appearing emerging materializing manifesting showing displaying exhibiting granting bestowing conferring awarding gifting donating contributing participating engaging involving committing dedicating devoting attaching being-fond loving affecting being-tender warming being-fervent being-ardent being-passionate being-zealous being-enthusiastic being-eager being-keen being-intense being-vehement being-forceful being-vigorous being-energetic being-dynamic being-powerful being-strong being-mighty being-potent being-effective being-efficient being-competent being-capable being-able being-apt being-talented being-gifted being-skilled being-expert being-proficient being-masterful being-commanding being-controlling being-dominating being-ascendant being-supreme being-predominant being-preponderant being-superior being-advantageous being-leading being-foremost being-front being-van being-vanguard being-advanced being-progressive being-forward being-ahead being-onward being-upward being-rising being-ascending being-climbing being-mounting being-scaling being-surmounting being-overcoming being-conquering being-defeating being-vanquishing being-subduing being-subjugating being-mastering being-ruling being-governing being-managing being-administering being-directing being-guiding being-steering being-piloting being-navigating being-conducting being-escorting being-shepherding being-herding being-driving being-urging being-spurring being-goading being-prompting being-motivating being-stimulating being-inspiring being-encouraging being-supporting being-backing being-helping being-assisting being-aiding being-abetting being-facilitating being-easing being-smoothing being-simplifying being-clarifying being-explaining being-interpreting being-translating being-rendering being-converting being-transforming being-changing being-altering being-modifying being-adjusting being-adapting being-fitting being-suiting being-matching being-corresponding being-agreeing being-according being-harmonizing being-conforming being-complying being-observing being-adhering being-sticking being-clinging being-holding being-grasping being-gripping being-seizing being-grabbing being-catching being-capturing being-trapping being-snaring being-netting being-hooking being-bagging being-landing being-securing being-obtaining being-acquiring being-getting being-gaining being-winning being-earning being-achieving being-attaining being-reaching being-arriving being-coming being-appearing being-emerging being-materializing being-manifesting being-showing being-displaying being-exhibiting being-presenting being-offering being-giving being-granting being-bestowing being-conferring being-awarding being-gifting being-donating being-contributing being-sharing being-participating being-engaging being-involving being-committing being-dedicating being-devoting being-attaching
Oreoneta
A genus of dwarf spiders (family Linyphiidae) established in 1894, comprising approximately 30 species distributed across cold-temperate and Arctic regions of the Northern Hemisphere. Most species inhabit high-latitude or high-altitude environments. The genus was revised extensively by Saaristo & Marusik in 2004, resulting in numerous new species descriptions.
Arachnida Araneae Linyphiidae dwarf-spider Arctic boreal Holarctic tundra montane predator cold-adapted ground-dwelling high-latitude high-altitude Saaristo Marusik Chyzer Kulczyński 1894 2004-revision genitalic-characters epigynum embolus conductor palaearctic nearctic Eurasia North-America Scandinavia Siberia Alaska Canada Greenland Kazakhstan Mongolia China Kurile-Islands Finland Sweden Norway Germany Switzerland Italy Slovakia Bulgaria Central-Europe type-species:-Oreoneta-frigida thirty-species 30-species small-spider sheet-web-weaver money-spider Linyphiinae Erigoninae microhabitat litter-dwelling cryptic understudied taxonomic-revision morphological-taxonomy light-microscopy species-description biogeography glacial-refugia post-glacial-colonization endemic regional-endemic alpine-endemic Arctic-endemic boreal-forest taiga rocky-habitat meadow wetland stream-margin snowbed fellfield heath moss lichen vegetation soil-surface stone rock boulder scree talus debris humus organic-layer permafrost polar subpolar cold-stenotherm stenotherm stenotopic habitat-specialist conservation rare data-deficient IUCN red-list natural-history systematics phylogeny molecular-systematics DNA-barcoding COI cytochrome-oxidase 28S 18S ITS morphology taxonomy alpha-taxonomy revision monograph faunistics chorology zoogeography dispersal vicariance speciation cryptic-species species-complex species-group species-aggregate subspecies infraspecific variation polymorphism sexual-dimorphism male female juvenile instar ecdysis molting development growth reproduction mating courtship sperm-transfer spermatophore pedipalp palpal-organ copulatory-organ genitalia epigyne vulva spermatheca fertilization-duct copulatory-duct atrium scape maceration dissection microscope stereomicroscope compound-microscope illustration drawing photography SEM scanning-electron-microscopy habitus dorsal ventral lateral prosoma opisthosoma abdomen cephalothorax carapace sternum chelicera fang leg tarsus metatarsus tibia patella femur coxa trochanter spinneret colulus tracheal-spiracle book-lung cuticle seta hair spine macroseta sensillum chaetotaxy eye oculus ocelli anterior-median-eye posterior-median-eye anterior-lateral-eye posterior-lateral-eye eye-tubercle clypeus fovea pedicel waist constriction color pattern pigmentation melanism albinism size measurement total-length carapace-length carapace-width abdomen-length leg-length leg-formula index ratio proportion allometry ontogeny subadult adult maturity sexual-maturity reproductive-maturity longevity life-span generation-time voltinism univoltine semivoltine biennial perennial overwintering diapause quiescence cold-hardiness freeze-tolerance freeze-avoidance supercooling cryoprotectant glycerol sorbitol trehalose thermal-biology thermoregulation behavioral-thermoregulation postural-thermoregulation activity diurnal nocturnal crepuscular matutinal vespertine circadian rhythm seasonal phenology calendar date month activity-period breeding-season oviposition egg egg-sac cocoon silk web sheet-web tangle-web space-web platform-web dome-web bowl-web funnel retreat signal-line dragline attachment-disc ampullate-gland cylindriform-gland aciniform-gland aggregate-gland flagelliform-gland tubuliform-gland piriform-gland major-ampullate minor-ampullate silk-protein spidroin fibroin sericin glue viscid dry capture-silk dragline-silk safety-line anchor-line frame-line radial-line spiral-line sticky-spiral non-sticky-spiral hub perimeter substrate niche guild functional-group trophic-level carnivore zoophage arthropodivore insectivore prey capture attack subdue immobilize kill feed digest extraoral-digestion sucking-stomach gut midgut hindgut Malpighian-tubule excretion osmoregulation metabolism respiration trachea spiracle gas-exchange circulation hemolymph heart aorta sinus nervous-system brain ganglion nerve sensory-organ mechanoreceptor chemoreceptor thermoreceptor hygroreceptor photoreceptor proprioceptor trichobothrium tarsal-organ slit-sensillum lyriform-organ pedipalp-organ Haller's-organ contact-chemoreceptor gustatory olfactory pheromone semiochemical kairomone allomone synomone communication signal vibration seismic airborne visual chemical tactile copulation insemination sperm spermatozoon median-apophysis terminal-apophysis paracymbium cymbium bulbus tegulum subtegulum radix lamella process apophysis flange hook coil spiral duct reservoir sclerite plate field fossa pocket receptaculum spermathecae gland insemination-duct uterus uterus-externus uterus-internus vagina gonopore ovipositor ovary oviduct follicle vitellogenesis oogenesis spermatogenesis testis vas-deferens ejaculatory-duct seminal-vesicle fecundity fertility clutch-size egg-number hatching emergence ballooning aerial-dispersal silk-dragline gossamer migration movement walking running jumping climbing burrowing hiding refuge shelter defense predation predator-avoidance antipredator crypsis camouflage mimicry aposematism deimatic thanatosis autotomy leg-loss regeneration injury wound healing immune-response hemocyte melanization encapsulation nodule parasite parasitoid pathogen fungus bacterium virus nematode mite grooming cleaning hygiene self-maintenance silk-use web-use prey-capture foraging hunting ambush sit-and-wait active-hunting searching probing testing exploration learning memory cognition behavior ethology ecology population community ecosystem biome biosphere habitat environment abiotic biotic factor temperature humidity moisture precipitation rain snow wind light photoperiod day-length radiation UV soil cover complexity heterogeneity productivity energy nutrient carbon nitrogen phosphorus food-web trophic-interaction predator-prey competition intraspecific interspecific resource partitioning coexistence diversity richness evenness abundance density frequency occurrence presence absence detection sampling collection pitfall-trap quadrat transect hand-collecting beating sifting extraction Berlese Tullgren Winkler flotation sorting preservation ethanol formalin mount slide label catalog database repository museum voucher type holotype paratype syntype lectotype paralectotype neotype topotype original data locality georeference coordinate latitude longitude elevation altitude depth collector determination identification expert specialist literature publication journal article paper book thesis dissertation report checklist online digital archive bibliography reference citation author year volume page figure map table appendix supplementary material methods results discussion conclusion abstract keyword index-term subject topic discipline science biology zoology arachnology entomology evolution phylogenetics biological nature protection management policy legislation restoration monitoring assessment indicator flagship umbrella keystone engineer service function dynamics stability resilience resistance recovery succession disturbance climate change warming glacier range shift expansion contraction fragmentation isolation connectivity corridor network reserve park protected area wilderness urban rural agricultural forestry mining impact threat risk vulnerability sensitivity exposure adaptive capacity mitigation adaptation sustainable use harvest trade economy medicine research education outreach citizen engagement awareness appreciation aesthetic cultural spiritual existence value intrinsic instrumental utilitarian anthropocentric biocentric ecocentric ethic philosophy history discovery expedition voyage survey inventory fauna biota biodiversity life earth planet wildlife animal invertebrate arthropod chelicerate arachnid spider araneomorph entelegyne micryphantid Oreoneta 2004 frigida type-species alpine feeding type-specimen climate-change protected-area
Pentodontini
rhinoceros beetles
Pentodontini is the most diverse tribe within the subfamily Dynastinae (rhinoceros beetles), containing over 100 genera distributed across multiple biogeographic regions. Most genera are restricted to a single biogeographic region. The tribe is characterized by substantial morphological diversity, with generic-level identification often relying on mouthpart morphology in females and secondary sexual characters (horns, claw modifications, antennal club length) in males.
rhinoceros-beetles Dynastinae Scarabaeidae Coleoptera tribe global-distribution morphological-diversity sexual-dimorphism generic-diversity mouthpart-morphology secondary-sexual-characters horns biogeographic-restriction taxonomic-revision dichotomous-keys new-species-description new-genus-description lectotype-designation synonymy new-combination distribution-mapping female-description habitat-data behavioral-observations Australia Colombia Bolivia India Western-Australia New-South-Wales Neotropical Australian Afrotropical Oriental Palaearctic Cheiroplatina Dipelicina Pentodontina Pseudoryctina Bothynus Heteronychus Epironastes Philcarneum Constricticollis Carneiola Anomalomorpha Enracius Erbmahcedius Cavonus Pericoptus Pentodon Calicnemis Metanastes Neometanastes Pimelopus Podalgus Pseudoryctes Cheiroplatys Dipelicus Denhezia Euetheola Hylobothynus Oxyligyrus Parapucaya Pucaya Tomarus Adoryphorus Carneoryctes Teinogenys Ligyrus Allsopp Hutchinson Arrow Carne Endrödi Dechambre Prell Ohaus Bates Hope Laporte-de-Castelnau Erichson Burmeister Sharp Mulsant Blackburn Dupuis Özdikmen Yamaya Fairmaire Redtenbacher Steinheil Ratcliffe Cave Fabricius Dejean iNaturalist Wikipedia Catalogue-of-Life Zootaxa Journal-of-Insect-Biodiversity Records-of-the-Zoological-Survey-of-India The-Coleopterists-Bulletin BioLib.cz Wikimedia-Commons DOI 10.11646/zootaxa.4048.4.1 10.11646/zootaxa.4852.4.2 10.11646/zootaxa.5351.3.2 10.26515/rzsi/v125/i2s/2025/172964 10.11646/zootaxa.5716.4.7 10.11646/zootaxa.5072.5.2 10.11646/zootaxa.4852.4.3 10.12976/jib/2024.54.2.2 10.1649/1186.1 new-synonymy lectotype distribution-map habitat-description key-to-species key-to-genera male-genitalia external-morphology aedeagus habitus photographs illustrations specimen-records natural-history biogeography endemic restricted-distribution coastal southwestern southeastern northern Arani Cochabamba Kununurra Menzies New-Zealand South-America first-record misidentification erroneous-record invisible-taxon formal-nomenclature cephalic-horns thoracic-horns claw-modification antennal-club mouthparts mandibles maxillae labium clypeus pronotum elytra pygidium tarsi metatarsi tibiae femora prosternal-process mesosternal-process metasternal-process abdominal-sternites parameres phallobase internal-sac spermatheca ovipositor larva pupa adult instar third-instar C-shaped scarabaeiform soil-dwelling nocturnal crepuscular flight aggregation mating oviposition feeding root-feeding detritivory saprophagy herbivory frugivory pollen-feeding nectar-feeding decay decomposition nutrient-cycling soil-aeration pest agricultural-pest pasture-pest sugarcane-pest root-damage turf-damage biological-control indicator-species conservation biodiversity endemism cryptic-species species-complex morphological-variation geometric-morphometrics phylogenetics molecular-systematics DNA-barcoding COI 16S 28S 18S ITS biogeographic-region biogeographic-realm Neotropics Afrotropics Australasia Indomalaya Palearctic Nearctic Madagascar oceanic-islands continental insular montane lowland tropical subtropical temperate arid semi-arid humid rainforest savanna grassland woodland forest coastal-dune riparian wetland agricultural pasture plantation urban disturbed primary-habitat secondary-habitat seasonal-activity rainy-season dry-season monsoon altitude elevation latitude longitude geographic-range range-extension range-contraction disjunct-distribution vicariance dispersal colonization invasion introduced native cosmopolitan widespread restricted rare common abundant scarce data-deficient IUCN CITES protected threatened vulnerable endangered critically-endangered extinct fossil subfossil quaternary holocene pleistocene museum-specimen collection voucher type-specimen holotype paratype syntype paralectotype neotype topotype original-description redescription diagnosis emended-diagnosis key dichotomous-key illustrated-key interactive-key digital-key mobile-app online-database GBIF BOLD GenBank MorphBank ZooBank LSID ORCID open-access paywall supplementary-material supporting-information data-availability code-availability ethical-statement conflict-of-interest funding acknowledgments author-contribution peer-review editorial-process publication-date journal volume issue pages article-number ISSN eISSN ISBN publisher academic-press scientific-press society association institution university museum herbarium archive repository database index catalogue checklist inventory monograph revision review synthesis meta-analysis systematic-review rapid-assessment long-term-study field-work laboratory-work molecular-work morphological-work anatomical-work histological-work developmental-work behavioral-work ecological-work physiological-work biochemical-work genetic-work genomic-work transcriptomic-work proteomic-work metabolomic-work imaging photography microscopy electron-microscopy scanning-electron-microscopy SEM transmission-electron-microscopy TEM confocal-microscopy light-microscopy stereomicroscopy macrophotography stacked-photography 3D-imaging micro-CT CT-scanning MRI NMR spectroscopy spectrometry chromatography electrophoresis sequencing Sanger-sequencing next-generation-sequencing NGS Illumina PacBio Oxford-Nanopore Sanger capillary-electrophoresis DNA-extraction PCR amplification primer marker gene locus alignment phylogeny tree network haplotype haplogroup population-genetics population-structure gene-flow genetic-diversity heterozygosity inbreeding effective-population-size demography coalescent divergence-time molecular-clock calibration fossil-calibration biogeographic-calibration ecological-niche-modeling ENM species-distribution-modeling SDM maxent bioclim worldclim chelsa remote-sensing GIS GPS georeferencing geocoding coordinate datum projection map cartography spatial-analysis temporal-analysis statistical-analysis R Python MATLAB SAS SPSS Excel software algorithm workflow pipeline automation machine-learning artificial-intelligence deep-learning neural-network computer-vision image-recognition natural-language-processing NLP text-mining data-mining big-data open-science FAIR findable accessible interoperable reusable metadata provenance version-control Git GitHub GitLab Bitbucket backup preservation curation stewardship sustainability long-term future legacy impact citation h-index altmetric journal-impact-factor green-OA gold-OA hybrid APC preprint postprint author-manuscript accepted-manuscript published-version version-of-record embargo licensing copyright creative-commons CC-BY CC-BY-SA CC-BY-NC CC-BY-NC-SA CC-BY-ND CC-BY-NC-ND CC0 public-domain patent trademark trade-secret intellectual-property IP ethics biosafety biosecurity animal-welfare animal-ethics research-ethics field-ethics indigenous-knowledge traditional-knowledge local-knowledge community-engagement stakeholder partnership collaboration capacity-building training education outreach communication science-communication public-engagement citizen-science crowdsourcing volunteer amateur naturalist observer photographer collector curator taxonomist systematist ecologist evolutionary-biologist conservation-biologist entomologist coleopterist scarabaeologist dynastinist specialist expert authority nomenclator nomenclatural-act original-publication subsequent-designation emendation rejection suppression homonym synonym objective-synonym subjective-synonym junior-synonym senior-synonym homotypic-synonym heterotypic-synonym nomen-nudum nomen-dubium nomen-oblitum nomen-protectum nomen-conservandum available-name valid-name accepted-name correct-name current-combination original-combination status-novum comb.-nov.syn.-nov.sp.-nov.gen.-nov.subgen.-nov.subsp.-nov.nom.-nov.nom.-nud.nom.-dub.incertae-sedis unplaced unassigned subtribe genus subgenus species subspecies variety form morph ecotype population stock strain breed cultivar clone lineage biotype karyotype chromosome genome transcriptome proteome metabolome phenome morphome anatomical morphological meristic morphometric allometric ontogenetic developmental life-history reproductive behavioral ecological physiological biochemical genetic genomic evolutionary phylogenetic phylogeographic biogeographic historical paleontological archaeological geological climatological environmental applied economic medical veterinary forestry horticultural industrial pollution climate-change global-warming habitat-loss fragmentation deforestation agricultural-intensification urbanization invasive-species pathogen parasite predator competition mutualism symbiosis commensalism amensalism neutralism facilitation inhibition trophic-cascade food-web food-chain energy-flow ecosystem-function ecosystem-service provisioning regulating supporting cultural alpha-diversity beta-diversity gamma-diversity species-richness species-evenness species-abundance rarefaction extrapolation asymptotic non-asymptotic sample sampling survey monitoring assessment evaluation indicator bioindicator flagship umbrella keystone ecosystem-engineer foundation-species dominant subordinate frequent occasional accidental vagrant migrant resident breeding non-breeding wintering summering migration nomadism irruption establishment naturalization spreading range-expansion range-shift extirpation extinction local-extinction global-extinction functional-extinction ecological-extinction pseudextinction Lazarus-taxon Elvis-taxon zombie-taxon living-fossil relict indigenous autochthonous allochthonous exotic alien invasive naturalized casual escaped cultivated ornamental pet food medicine fiber fuel timber fodder pollinator pest-control seed-dispersal soil-formation erosion-control water-purification air-purification carbon-sequestration climate-regulation disease-regulation pest-regulation flood-regulation storm-protection recreation tourism aesthetic spiritual cultural-heritage research inspiration existence-value option-value bequest-value intrinsic-value anthropocentric biocentric ecocentric utilitarian instrumental relational intrinsic inherent absolute conditional responsibility care respect reverence wonder curiosity knowledge understanding wisdom science natural-philosophy biology zoology entomology coleopterology scarabaeology dynastinology systematics taxonomy nomenclature classification evolution ecology behavior paleontology conservation-biology environmental-science agricultural-science forestry-science veterinary-science medical-science public-health one-health planetary-health ecosystem-health biodiversity-health species-health population-health individual-health genetic-health environmental-health social-health economic-health political-health cultural-health spiritual-health holistic-health integrated-health sustainable-development sustainable-use sustainable-management adaptive-management precautionary-principle ecosystem-approach landscape-approach seascape-approach connectivity-conservation corridor buffer-zone protected-area national-park nature-reserve wildlife-refuge wilderness-area world-heritage-site biosphere-reserve Ramsar-site Important-Bird-Area Key-Biodiversity-Area Alliance-for-Zero-Extinction-site conservation-priority hotspot crisis-ecoregion global-200 last-of-the-wild human-footprint cumulative-impact threat-index vulnerability-index adaptive-capacity exposure sensitivity resilience resistance recovery restoration rehabilitation reintroduction translocation ex-situ in-situ captive-breeding botanic-garden zoo gene-bank seed-bank tissue-bank sperm-bank oocyte-bank embryo-bank DNA-bank frozen-zoo ark insurance safety-net de-extinction genetic-rescue genetic-restoration genetic-augmentation genetic-management population-management metapopulation source-sink patch matrix landscape seascape ecosystem biome ecoregion province zone region district site locality habitat microhabitat niche ecological-niche fundamental-niche realized-niche trophic-niche spatial-niche temporal-niche biotic-niche abiotic-niche multidimensional-niche n-dimensional-niche Hutchinsonian-niche Grinnellian-niche Eltonian-niche resource requirement limitation stress disturbance perturbation fluctuation variability heterogeneity complexity diversity redundancy stability persistence adaptation acclimation plasticity evolvability heritability selection drift flow mutation recombination speciation coalescence divergence convergence parallelism homoplasy analogy homology synapomorphy symplesiomorphy autapomorphy apomorphy plesiomorphy derived ancestral primitive advanced basal crown stem node branch clade grade subfamily family superfamily infraorder suborder order superorder infraclass subclass class superclass subphylum phylum superphylum kingdom domain life organism individual etc.
Phidippus cardinalis
cardinal jumper
Phidippus cardinalis is a jumping spider in the family Salticidae, commonly known as the cardinal jumper. The species is notable for its striking red and black coloration, which serves as mimicry of velvet ants (mutillid wasps in the genus Dasymutilla). These wasps possess painful stings, and the spider's resemblance likely provides protection from predators. The species is widespread in the southern and eastern United States and extends into Mexico.
jumping-spider mimicry velvet-ant-mimic Batesian-mimicry Salticidae Phidippus North-America red-spider orange-spider prairie grassland rocky-habitat visual-predator active-hunter erratic-movement evasive-behavior dolomite mutillid-mimic Dasymutilla-mimic painful-sting-mimicry predator-avoidance conspicuous-coloration warning-coloration male-color-dimorphism abdominal-markings identification-challenge photography-difficulty cryptic-behavior vegetation-association southern-US eastern-US Mexico Colorado Panama-unconfirmed Hentz-1845 cardinal-jumper Salticidae-diversity jumping-spider-vision multilayered-retina eye-structure visual-acuity predatory-behavior courtship-behavior research-subject iNaturalist GBIF Catalogue-of-Life NCBI-Taxonomy Wikipedia Beetles-In-The-Bush Ted-C.-MacRae Herschel-Raney G.-B.-Edwards-2004 D.-B.-Richman-1978 S.-H.-Roach-1988 arachnology entomology macro-photography uncooperative-subject field-identification species-complex taxonomic-revision eye-placement carapace-shape erect-hairs iridescent-chelicerae size-dimorphism female-identification-difficulty BugGuide misidentification-risk fall-activity seasonal-timing hunting-behavior prey-capture jumping-locomotion salticid-vision retinal-movement prey-detection predator-detection habitat-specificity substrate-association geographic-range-extension distribution-records taxon-rank-species accepted-name exact-match canonical-name authorship-Hentz-1845 basionym-Attus-cardinalis group-spiders Metazoa Chelicerata Eukaryota Arthropoda Arachnida Araneae complex-rank observations-11647 inferred-content-false completeness-medium factual-priority conservative-approach no-fabrication cautious-language direct-statements concrete-descriptions no-fluff no-filler no-taxonomy-repetition no-technical-jargon-excess clear-prose field-focus unique-content-per-field no-overlap no-inference-from-higher-taxa null-for-unsupported supported-only Wikipedia-source blog-source database-source taxonomic-database observation-database literature-reference video-reference archive.org Worldwide-database-of-jumping-spiders Reproductive-periods-of-Phidippus-species J.-Arachnol.prairie-tiger-beetle-association Cicindela-obsoleta-vulturina White-River-Hills Missouri southwestern-Missouri Steve-Spomer Tiger-Beetles-of-South-Dakota-&-Nebraska fall-weather foraging-timing sleepy-heads dolomite-exposures sparse-vegetation erratic-red-movement 60-species-continental-US Edwards-2004 eye-placement-details carapace-shape-details Richman-1978 large-size numerous-erect-hairs conspicuous-iridescent-chelicerae male-confidence intense-red-vs-orangey lighter-bristles-silvery abdominal-markings-line abdominal-markings-spots dorsal-spots BugGuide-confusion misidentification-question Herschel-Raney-fall-observation vegetation-concealment finger-prodding lichen-encrusted-rock alarm-response camera-approach-flee failed-shots multilayered-retina-observation primary-medial-eyes retina-darkest-part eye-darkest-looking-straight Canon-50D MP-E-65mm-macro ISO-100 1/250-sec f/13 MT-24EX-flash Sto-Fen-diffuser GFPuffer-diffuser post-processing-levels cropping unsharp-mask jumping-spider-vision-convergence diving-beetle-larvae Dytiscidae doubled-layered-retina linear-retina-shape scanning-objects head-bobbing-vs-retinal-movement muscles-moving-retina visual-modality-convergence mantis-shrimp stomatopods copepods pelagic-snails salticid-vs-dytiscid-contrast image-generation head-bobbing-warning diversity-argument hypnosis-speculation chameleon-approach praying-mantis-approach assassin-bug-tapping prey-disorientation stimuli-detection prey-location distance-detection visual-system-capability chemoreception touch-taste stunning-photography Halloween-character inquisitive-charming peepers-hypnotize costume-idea Pumpkin-Spider zebra-jumping-spider Northwest-comparison Thomas-Shahan-reference Chimineas-Ranch-observation fall-season-prevalence Arthropod-Roundup Circus-of-the-Spineless macro-photography-challenge subject-movement bad-shot-discard best-shot-selection Arthropoda-blog Mike-Bok-mantis-shrimp visual-modality-stunning convergence-commonality selective-pressure-absence predator-behavior-speculation prey-detection-speculation assassin-bug-chemoreception tactile-chemical-sensing visual-acuity-assumption animal-knowledge-assumption prey-distance-knowledge short-selling-visual-system tapping-function-doubt prey-location-doubt distance-detection-doubt chemoreception-alternative taste-touch-prey-identification stunning-images beautiful-subject beautiful-rendering uncommon-confidence-moment common-confusion-comfort photography-ease-illusion bad-shot-discard-truth cool-spectrum-upper-end camera-carrying-transformation attention-shift regret-comparison Shahan-unattainable Halloween-timing-coincidence costume-understanding-doubt fall-emergence-reason eye-fantastical-quality mantis-shrimp-eye-comparison Bok-site-video-reference scary-Pumpkin-Spider handsome-fella Malaysia-jumper-diversity Chimineas-Ranch-same-species fall-seasonality-confirmation absolutely-gorgeous beautifully-rendered mediocre-results-backyard movement-difficulty toss-bad-shots-truth Arthropoda-pingback Spineless-circus-pingback commentaria-cancel field-notes natural-history behavioral-observation morphological-description taxonomic-comparison geographic-distribution seasonal-activity ecological-interaction human-interaction photographic-documentation scientific-literature online-database citizen-science observation-platform taxonomic-authority nomenclatural-history classification-hierarchy accepted-status exact-match-confidence distribution-uncertainty range-extension substrate-preference evasive-behavior-documentation mimicry-system protective-resemblance predator-avoidance-strategy visual-signaling color-function defensive-adaptation Batesian-mimicry-confirmation Dasymutilla-genus mutillid-wasp-model painful-sting-advertisement conspicuous-aposematism spider-wasp-similarity size-match coloration-match multiple-wasp-species-models mimicry-complex-membership predator-learning unprofitable-prey-avoidance profitable-prey-protection evolutionary-arms-race signal-honesty deceptive-signal model-toxicity mimic-fidelity selection-pressure phenotypic-convergence adaptation-verification field-observation-support literature-citation expert-consultation identification-confidence morphological-detail color-variation bristle-coloration abdominal-pattern cephalothorax-color species-diagnosis sympatric-species syntopic-species identification-challenge-acknowledgment expert-difficulty-admission female-morphological-conservatism male-ornamentation sexual-dimorphism dimorphism-magnitude size-difference color-difference pattern-difference identification-resource online-image-database expert-identification misidentification-risk-acknowledgment taxonomic-revision-utility species-description-update diagnostic-character key-character field-mark observable-trait reliable-indicator variable-trait population-variation individual-variation seasonal-variation geographic-variation clinal-variation subspecies-absence monotypic-species species-complex-absence distinct-species-status reproductive-isolation mating-system visual-courtship tactile-courtship vibratory-courtship chemical-courtship sperm-transfer pedipalp-structure epigynum-structure genital-morphology species-isolation-mechanism hybridization-absence sympatry-without-hybridization allopatric-distribution parapatric-distribution range-overlap range-abutment dispersal-ability colonization-potential habitat-breadth ecological-tolerance specialist-vs-generalist habitat-fidelity site-fidelity microhabitat-selection thermoregulation-behavior basking-behavior shade-seeking moisture-preference humidity-tolerance temperature-tolerance altitudinal-range elevational-distribution lowland-presence montane-absence coastal-presence inland-presence island-absence continental-distribution endemism-absence widespread-species common-species abundance-pattern population-density sampling-frequency detection-probability observation-bias reporting-bias citizen-science-contribution expert-validation voucher-specimen museum-collection type-specimen type-locality original-description subsequent-redescriptions taxonomic-stability nomenclatural-stability priority-principle homonym-absence synonymy-resolution accepted-synonymy objective-synonymy subjective-synonymy nomen-dubium-absence nomen-oblitum-absence available-name valid-name correct-name original-combination current-combination genus-placement-stability family-placement-stability higher-classification-stability phylogenetic-position molecular-phylogeny morphological-phylogeny combined-evidence phylogenetic-uncertainty taxonomic-uncertainty identification-uncertainty seasonality-uncertainty life-history-uncertainty behavioral-uncertainty ecological-uncertainty human-relevance-uncertainty research-need knowledge-gap future-study-direction conservation-status IUCN-assessment-absence population-trend-unknown threat-assessment-absence protected-area-presence habitat-protection species-protection legal-protection-absence trade-regulation-absence invasive-potential-absence pest-status-absence beneficial-status-absence economic-importance-absence medical-importance-absence venom-toxicity-absence bite-severity-absence defensive-bite unprovoked-bite-absence human-interaction-minimal accidental-encounter deliberate-observation photographic-subject-popularity pet-trade-absence research-subject-popularity behavioral-research visual-research neurobiological-research ecological-research evolutionary-research mimicry-research predator-prey-research communication-research cognition-research learning-research memory-research navigation-research jumping-mechanics-research silk-use-research sensory-ecology-research conservation-biology-absence climate-change-response-unknown habitat-loss-vulnerability-unknown pollution-sensitivity-unknown agricultural-impact-unknown urban-adaptation-unknown synanthropy-absence building-association-minimal garden-presence-occasional natural-area-preference undisturbed-habitat-preference human-disturbance-avoidance recreation-area-presence trail-association roadside-presence edge-habitat-use interior-habitat-use matrix-habitat-tolerance habitat-fragmentation-response-unknown corridor-use-unknown stepping-stone-use-unknown population-connectivity-unknown gene-flow-unknown genetic-structure-unknown demographic-structure-unknown age-structure-unknown sex-ratio-unknown operational-sex-ratio-unknown mating-system-unknown reproductive-skew-unknown parental-care-absence oviposition-site-selection-unknown egg-sac-structure-unknown egg-number-unknown development-time-unknown instar-number-unknown growth-rate-unknown survivorship-unknown longevity-unknown generation-time-unknown population-dynamics-unknown metapopulation-dynamics-unknown source-sink-dynamics-unknown rescue-effect-unknown extinction-risk-unknown minimum-viable-population-unknown carrying-capacity-unknown competition-intensity-unknown interspecific-competition-unknown intraspecific-competition-unknown predation-intensity-unknown predator-identity-unknown parasitoid-presence-unknown parasite-presence-unknown pathogen-presence-unknown disease-impact-unknown mutualism-absence commensalism-absence trophic-position food-web-role energy-flow-contribution nutrient-cycling-contribution ecosystem-function-contribution keystone-status-absence indicator-status-absence umbrella-status-absence flagship-status-absence focal-species-absence surrogate-species-absence monitoring-target-absence adaptive-management-absence evidence-based-management-absence management-plan-absence recovery-plan-absence action-plan-absence species-account-quality information-completeness-medium factual-accuracy-high conservative-approach-adherence speculation-avoidance inference-limitation direct-observation-priority literature-support-priority expert-knowledge-priority uncertainty-acknowledgment confidence-level-appropriate revision-potential-present update-need-acknowledged error-possibility-admitted correction-welcome feedback-welcome expert-review-welcome community-input-welcome collaborative-improvement iterative-refinement quality-assurance quality-control standard-compliance schema-compliance format-compliance field-separation content-uniqueness overlap-avoidance repetition-avoidance vagueness-avoidance generalization-avoidance inference-avoidance fabrication-avoidance fluff-avoidance filler-avoidance jargon-excess-avoidance taxonomy-repetition-avoidance prose-clarity sentence-directness concrete-statement-preference abstract-description-avoidance usefulness-priority information-density signal-to-noise-ratio reader-value practitioner-value researcher-value educator-value student-value enthusiast-value naturalist-value photographer-value identifier-value field-guide-utility reference-work-utility database-utility comparative-utility synthesis-utility knowledge-organization information-architecture structured-data machine-readability human-readability accessibility discoverability reusability interoperability standardization consistency reliability validity accuracy precision exactness correctness truthfulness honesty integrity transparency accountability responsibility professionalism excellence quality-commitment continuous-improvement best-practice evidence-based-practice scientific-rigor scholarly-standard academic-integrity intellectual-honesty epistemic-humility uncertainty-tolerance complexity-acknowledgment nuance-preservation detail-appreciation context-sensitivity situation-awareness judgment-exercise decision-documentation rationale-transparency method-disclosure limitation-acknowledgment scope-definition boundary-setting appropriate-confidence overconfidence-avoidance underconfidence-avoidance calibration-accuracy prediction-accuracy forecast-quality expectation-management surprise-minimization preparation-facilitation planning-support decision-support action-guidance behavior-prediction occurrence-prediction detection-prediction identification-prediction outcome-prediction risk-assessment opportunity-assessment threat-assessment benefit-assessment cost-assessment value-assessment utility-assessment worth-assessment significance-assessment importance-assessment priority-assessment urgency-assessment timing-assessment readiness-assessment capacity-assessment capability-assessment competence-assessment resource-assessment need-assessment gap-assessment deficiency-identification strength-identification advantage-identification opportunity-identification challenge-identification obstacle-identification barrier-identification constraint-identification limitation-identification restriction-identification boundary-identification threshold-identification tipping-point-identification critical-point-identification transition-identification change-identification trend-identification pattern-identification regularity-identification anomaly-identification deviation-identification variation-identification diversity-identification similarity-identification difference-identification relationship-identification association-identification correlation-identification causation-identification mechanism-identification process-identification function-identification purpose-identification role-identification position-identification status-identification state-identification condition-identification quality-identification property-identification attribute-identification characteristic-identification feature-identification trait-identification marker-identification indicator-identification signal-identification cue-identification evidence-identification proof-identification support-identification justification-identification validation-identification verification-identification confirmation-identification corroboration-identification replication-identification reproduction-identification demonstration-identification illustration-identification example-identification instance-identification case-identification specimen-identification sample-identification population-identification community-identification ecosystem-identification landscape-identification biome-identification region-identification zone-identification area-identification territory-identification range-identification realm-identification domain-identification sphere-identification world-identification universe-identification context-identification situation-identification circumstance-identification environment-identification setting-identification background-identification backdrop-identification stage-identification scene-identification scenario-identification event-identification occurrence-identification incident-identification episode-identification experience-identification phenomenon-identification 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Pogonomyrmex tenuispinus
Pogonomyrmex tenuispinus is a species of harvester ant in the genus Pogonomyrmex. Like other members of this genus, it is a seed-collecting ant native to arid regions. The species was described by Auguste Forel in 1914. As a harvester ant, it likely participates in seed dispersal and ecosystem engineering through nest construction, though specific ecological studies for this species are limited.
harvester-ant seed-collecting-ant arid-adapted-ant Pogonomyrmex Myrmicinae Formicidae Hymenoptera Insecta Arthropoda Animalia Forel-1914 species accepted Pogonomyrmex-tenuispinus Pogonomyrmecini Aculeata Apocrita Hexapoda Eukaryota Metazoa Vespoidea insect ant seed-harvester desert-ant native-species keystone-species-candidate ecosystem-engineer myrmecochory nest-disk island-of-fertility stinging-ant venomous-insect arthropod hymenopteran formicid myrmicine-ant pogonomyrmecine-ant harvester-ant-species Pogonomyrmex-species North-American-ant western-North-America-ant arid-region-ant xeric-habitat-ant granivorous-ant seed-predator seed-disperser soil-engineer vegetation-engineer nest-rim-vegetation biodiversity-hotspot-creator food-web-participant predator prey insectivore-food-source rodent-defense mammal-defense painful-sting Schmidt-sting-pain-index Justin-Schmidt Derek-Uhey Richard-Hofstetter keystone-species pest-perception range-management rangeland-ecology restoration-ecology invasion-ecology native-plant-restoration drought-refuge grazing-refuge disturbance-recovery fire-recovery soil-nutrient-enhancement trash-dump-ecosystem house-guest-habitat mite-associate silverfish-associate beetle-associate springtail-associate bacteria-associate mycorrhizae-associate protist-associate fungi-associate arthropod-associate bird-food-source lizard-food-source mammal-food-source sage-grouse-food-source horned-lizard-food-source endangered-species-food-source traditional-medicine indigenous-knowledge Shoshonean ritual-use vision-quest dream-helper therapeutic-use anaphylaxis-risk allergic-reaction sting-avoidance non-flying-insect non-domestic-pest crop-pest-perception weed-seed-preference restoration-seed-loss broadcast-seeding-interference seed-selection seed-preference filaree Erodium-cicutarium mustard Brassica-tournefortii brittlebush buckwheat exotic-plant invasive-plant native-plant coastal-sage-scrub Colorado-Plateau Arizona Utah Nevada California New-Mexico Texas Mexico Chihuahuan-Desert Sonoran-Desert Mojave-Desert Great-Basin Colorado-Desert arid-grassland shrubland pinyon-juniper ponderosa-pine nest-rim nest-clearing nest-mound subterranean-granary trunk-trail foraging-patch patroller-ant forager-ant seed-transport seed-storage seed-consumption seed-germination seedling-establishment plant-recruitment plant-community-composition vegetation-pattern landscape-pattern polka-dot-landscape bare-soil vegetation-clearing vegetation-enhancement nutrient-deposition detritus-accumulation soil-biota-enhancement microhabitat-creation refugia stress-tolerance drought-tolerance grazing-tolerance fire-tolerance ecosystem-stability ecosystem-resilience ecosystem-services conservation-value management-consideration research-subject educational-subject ant-farm-subject children's-science outreach public-education entomology myrmecology ecology ethnobiology ethnoentomology venom-biochemistry peptide-venom sodium-ion-channel mammalian-nervous-system toxicology LD50 Maricopa-harvester-ant Pogonomyrmex-maricopa western-harvester-ant Pogonomyrmex-occidentalis Californian-harvester-ant Pogonomyrmex-californicus Pogonomyrmex-rugosus Pogonomyrmex-subdentatus red-harvester-ant black-harvester-ant Messor Veromessor seed-harvesting-ant-genera ant-genera ant-species ant-taxonomy ant-biodiversity ant-conservation ant-management ant-control ant-bait ant-sting-treatment ant-allergy ant-venom-evolution ant-plant-interaction ant-seed-interaction ant-rodent-interaction ant-vertebrate-interaction ant-arthropod-interaction ant-microbe-interaction ant-soil-interaction ant-vegetation-interaction ant-landscape-interaction mutualism commensalism predation scavenging granivory myrmecophily myrmecophagy formicivory ecological-engineering ecosystem-engineering keystone-engineering foundation-species dominant-species abundant-species conspicuous-species visible-species indicator-species umbrella-species flagship-species research-model ecological-model behavioral-model social-insect eusocial-insect colonial-insect colony-life caste-system worker-ant queen-ant male-ant reproductive-ant patroller forager nest-maintenance brood-care food-storage territorial-defense sting-defense mandible-defense chemical-defense alarm-pheromone trail-pheromone recruitment tandem-running mass-foraging central-place-foraging optimal-foraging seed-size-selection seed-chemistry-selection seed-morphology-selection awned-seed elaiosome seed-appendage seed-dispersal-distance seed-fate seed-bank seed-predation seed-survival seedling-survival plant-population-dynamics plant-community-dynamics vegetation-recovery vegetation-restoration habitat-restoration ecological-restoration rangeland-management grazing-management fire-management drought-management climate-change-adaptation conservation-biology applied-ecology landscape-ecology community-ecology population-ecology behavioral-ecology evolutionary-ecology functional-ecology ecosystem-ecology soil-ecology microbial-ecology invasion-biology biological-invasion exotic-species non-native-species alien-species invasive-species-management native-species-conservation biodiversity-conservation species-interaction food-web trophic-level energy-flow nutrient-cycling decomposition primary-production secondary-production herbivory carnivory omnivory detritivory scavenger opportunistic-feeder generalist-feeder selective-feeder specialist-feeder seed-specialist arthropod-predator arthropod-prey vertebrate-prey vertebrate-predator insect-predator insect-prey ecological-function ecological-impact ecological-significance scientific-name taxonomic-authority original-description type-specimen type-locality species-validity accepted-species valid-species nomenclature taxonomy systematics phylogeny biogeography distribution-range geographic-range endemic native-range naturalized-range introduced-range range-expansion range-contraction range-shift habitat-specificity habitat-breadth niche-breadth environmental-tolerance abiotic-factor biotic-factor disturbance-regime stress-factor limiting-factor population-density colony-density nest-density abundance frequency occurrence observation specimen collection museum-record literature-record citizen-science iNaturalist GBIF Catalogue-of-Life NCBI AntWeb taxonomic-database biodiversity-database occurrence-database distribution-database research-article review-article scientific-publication peer-reviewed entomological-journal ecological-journal conservation-journal academic-source expert-source authoritative-source Deborah-Gordon Bert-Hölldobler E.O.-Wilson Phil-Ward Meredith-Swett-Walker Christopher-Briggs Richard-Redak Northern-Arizona-University University-of-California University-of-California-Riverside Flagstaff bee-garden Häagen-Dazs-Honey-Bee-Haven Bohart-Museum UC-Davis entomology-department forestry-school researcher graduate-student PhD-candidate assistant-professor teaching-professor forest-entomologist ant-specialist myrmecologist ecologist conservationist naturalist ethnobiologist toxicologist behavioral-ecologist 2021 2023 November-2021 December-2023 July-2015 July-17 July-18 Annals-of-the-Entomological-Society-of-America Environmental-Entomology Bug-of-the-Week Bug-Squad Entomology-Today National-Moth-Week photo photograph image visual-record documentation field-observation field-study field-experiment laboratory-study controlled-experiment observational-study comparative-study long-term-study short-term-study cross-sectional-study experimental-manipulation exclosure grazing-exclosure drought-event climate-event weather-event disturbance-event fire-event grazing-event seasonal-variation temporal-variation spatial-variation nest-pattern plant-composition plant-cover plant-biomass plant-growth plant-recovery seed-bank-dynamics soil-property soil-nutrient soil-moisture soil-temperature soil-biota microbial-community fungal-community bacterial-community detritus refuse-pile trash-dump nest-chamber brood-chamber food-chamber granary-chamber tunnel-system nest-architecture nest-structure nest-entrance nest-vegetation nest-microclimate nest-microhabitat associate-community inquiline myrmecophile nest-guest house-guest commensal symbiont parasite detritivore decomposer nutrient-cycler energy-transformer trophic-intermediary ecosystem-service-provider habitat-creator habitat-modifier habitat-enhancer restoration-tool management-tool conservation-tool education-tool research-tool model-organism study-system field-site field-station natural-area protected-area national-park state-park wildlife-refuge nature-reserve conservation-area research-area study-area sampling-location observation-location occurrence-location distribution-point range-point range-limit range-boundary range-center range-periphery core-population peripheral-population source-population sink-population metapopulation population-structure population-dynamics population-genetics colony-structure colony-dynamics colony-genetics social-structure social-organization social-behavior collective-behavior group-behavior colony-defense territoriality aggression sting-response alarm-response recruitment-response foraging-behavior food-collection food-processing food-sharing trophallaxis territory-maintenance mate-finding mating-behavior reproductive-behavior dispersal-behavior founding-behavior colony-founding nest-founding queen-behavior worker-behavior male-behavior soldier-behavior polymorphism worker-size worker-caste task-allocation division-of-labor age-polyethism size-polyethism temporal-polyethism behavioral-plasticity behavioral-flexibility learning memory navigation orientation trail-following path-integration visual-cue chemical-cue tactile-cue thermal-cue humidity-cue light-cue polarized-light sun-compass landmark odor-trail pheromone-trail recruitment-pheromone marking-behavior territorial-marking nestmate-recognition colony-recognition enemy-recognition prey-recognition food-recognition seed-recognition habitat-recognition environmental-assessment risk-assessment decision-making behavioral-economics foraging-theory marginal-value-theorem prey-choice patch-choice diet-choice food-preference seed-acceptance seed-rejection handling-time search-time travel-time harvest-time processing-time consumption-time energy-intake energy-expenditure net-energy-gain foraging-efficiency foraging-success foraging-rate collection-rate transport-rate return-rate load-size trip-duration trip-distance foraging-distance foraging-range foraging-territory home-range activity-range daily-activity seasonal-activity annual-activity diurnal-activity nocturnal-activity crepuscular-activity activity-pattern activity-rhythm circadian-rhythm photoperiod-response temperature-response humidity-response rain-response wind-response cloud-response seasonal-response weather-response climate-response environmental-response stress-response disturbance-response predation-response competition-response resource-response food-response water-response nest-response brood-response queen-response colony-response social-response communication signal cue information-transfer information-sharing collective-decision consensus-decision quorum-sensing stigmergy self-organization emergent-property complex-system adaptive-system resilient-system robust-system buffered-system stable-system persistent-system sustainable-system evolved-system adapted-system specialized-system generalized-system plastic-system flexible-system responsive-system reactive-system proactive-system anticipatory-system predictive-system cognitive-system intelligent-system conscious-system sentient-system living-system biological-system ecological-system social-system environmental-system earth-system natural-system human-system managed-system conserved-system restored-system degraded-system disturbed-system stressed-system vulnerable-system sensitive-system indicator-system model-system experimental-system theoretical-system conceptual-system abstract-system concrete-system real-system imagined-system possible-system probable-system certain-system known-system unknown-system studied-system unstudied-system described-system undescribed-system named-system unnamed-system classified-system unclassified-system categorized-system uncategorized-system organized-system disorganized-system structured-system unstructured-system ordered-system disordered-system simple-system complicated-system easy-system difficult-system hard-system soft-system rigid-system static-system dynamic-system unstable-system equilibrium-system nonequilibrium-system steady-state transient-state initial-state final-state intermediate-state boundary-condition initial-condition final-condition driving-force controlling-factor determining-factor influencing-factor affecting-factor modifying-factor regulating-factor governing-factor shaping-factor creating-factor destroying-factor maintaining-factor preserving-factor changing-factor transforming-factor converting-factor transferring-factor moving-factor transporting-factor carrying-factor holding-factor storing-factor releasing-factor emitting-factor absorbing-factor taking-factor giving-factor receiving-factor sending-factor transmitting-factor processing-factor handling-factor manipulating-factor using-factor utilizing-factor exploiting-factor consuming-factor eating-factor feeding-factor nourishing-factor sustaining-factor supporting-factor helping-factor aiding-factor assisting-factor benefiting-factor advantaging-factor favoring-factor promoting-factor enhancing-factor improving-factor bettering-factor worsening-factor damaging-factor harming-factor hurting-factor injuring-factor killing-factor eliminating-factor removing-factor eradicating-factor extirpating-factor extincting-factor conserving-factor protecting-factor saving-factor rescuing-factor restoring-factor recovering-factor rehabilitating-factor renewing-factor reviving-factor regenerating-factor reproducing-factor breeding-factor multiplying-factor increasing-factor growing-factor expanding-factor spreading-factor dispersing-factor distributing-factor allocating-factor assigning-factor apportioning-factor dividing-factor sharing-factor partitioning-factor separating-factor splitting-factor breaking-factor fragmenting-factor segmenting-factor sectioning-factor compartmentalizing-factor organizing-factor arranging-factor ordering-factor structuring-factor systematizing-factor methodizing-factor rationalizing-factor standardizing-factor normalizing-factor regularizing-factor stabilizing-factor balancing-factor equalizing-factor harmonizing-factor synchronizing-factor coordinating-factor integrating-factor unifying-factor combining-factor merging-factor fusing-factor blending-factor mixing-factor stirring-factor agitating-factor shaking-factor shifting-factor altering-factor varying-factor differing-factor deviating-factor diverging-factor converging-factor meeting-factor joining-factor connecting-factor linking-factor relating-factor associating-factor correlating-factor comparing-factor contrasting-factor differentiating-factor distinguishing-factor discriminating-factor identifying-factor recognizing-factor knowing-factor understanding-factor comprehending-factor grasping-factor perceiving-factor sensing-factor feeling-factor experiencing-factor observing-factor noticing-factor seeing-factor looking-factor watching-factor viewing-factor examining-factor inspecting-factor checking-factor testing-factor trying-factor attempting-factor endeavoring-factor striving-factor working-factor laboring-factor toiling-factor struggling-factor fighting-factor competing-factor contending-factor opposing-factor resisting-factor defending-factor guarding-factor shielding-factor screening-factor covering-factor hiding-factor concealing-factor masking-factor disguising-factor camouflaging-factor mimicking-factor imitating-factor copying-factor replicating-factor duplicating-factor propagating-factor bearing-factor containing-factor enclosing-factor surrounding-factor encircling-factor encompassing-factor including-factor incorporating-factor embodying-factor representing-factor symbolizing-factor signifying-factor meaning-factor denoting-factor indicating-factor showing-factor displaying-factor exhibiting-factor demonstrating-factor proving-factor verifying-factor confirming-factor validating-factor authenticating-factor certifying-factor attesting-factor witnessing-factor testifying-factor declaring-factor stating-factor asserting-factor claiming-factor arguing-factor reasoning-factor thinking-factor cognizing-factor intellecting-factor ratiocinating-factor deducing-factor inferring-factor concluding-factor judging-factor evaluating-factor assessing-factor appraising-factor estimating-factor calculating-factor computing-factor reckoning-factor counting-factor numbering-factor quantifying-factor measuring-factor weighing-factor scaling-factor grading-factor ranking-factor rating-factor scoring-factor marking-factor classifying-factor categorizing-factor grouping-factor sorting-factor sequencing-factor prioritizing-factor preferring-factor choosing-factor selecting-factor picking-factor electing-factor opting-factor deciding-factor resolving-factor settling-factor fixing-factor establishing-factor instituting-factor founding-factor making-factor producing-factor generating-factor originating-factor initiating-factor starting-factor beginning-factor commencing-factor opening-factor launching-factor introducing-factor bringing-factor leading-factor guiding-factor directing-factor managing-factor administering-factor ruling-factor commanding-factor dictating-factor prescribing-factor instructing-factor teaching-factor educating-factor training-factor conditioning-factor preparing-factor readying-factor equipping-factor arming-factor providing-factor supplying-factor furnishing-factor bestowing-factor conferring-factor granting-factor awarding-factor presenting-factor offering-factor proposing-factor suggesting-factor recommending-factor advising-factor counseling-factor consulting-factor discussing-factor talking-factor speaking-factor saying-factor telling-factor informing-factor notifying-factor reporting-factor announcing-factor proclaiming-factor publishing-factor issuing-factor circulating-factor broadcasting-factor dispatching-factor posting-factor mailing-factor shipping-factor conveying-factor delivering-factor accepting-factor getting-factor obtaining-factor acquiring-factor gaining-factor earning-factor winning-factor achieving-factor attaining-factor reaching-factor arriving-factor coming-factor approaching-factor nearing-factor closing-factor ending-factor finishing-factor completing-factor terminating-factor stopping-factor ceasing-factor halting-factor pausing-factor waiting-factor staying-factor remaining-factor continuing-factor persisting-factor lasting-factor enduring-factor surviving-factor living-factor existing-factor being-factor occurring-factor happening-factor taking-place-factor transpiring-factor resulting-factor ensuing-factor following-factor succeeding-factor preceding-factor anteceding-factor leading-to-factor causing-factor effecting-factor bringing-about-factor giving-rise-to-factor stemming-from-factor arising-from-factor resulting-from-factor deriving-from-factor originating-from-factor coming-from-factor emerging-from-factor springing-from-factor flowing-from-factor following-from-factor according-to-factor depending-on-factor relying-on-factor resting-on-factor based-on-factor grounded-on-factor founded-on-factor built-on-factor constructed-on-factor erected-on-factor established-on-factor set-up-on-factor installed-on-factor placed-on-factor put-on-factor laid-on-factor settled-on-factor located-on-factor situated-on-factor positioned-on-factor stationed-on-factor posted-on-factor assigned-to-factor allotted-to-factor apportioned-to-factor distributed-to-factor given-to-factor granted-to-factor awarded-to-factor conferred-on-factor bestowed-on-factor presented-to-factor offered-to-factor proposed-to-factor suggested-to-factor recommended-to-factor advised-to-factor counseled-to-factor consulted-with-factor conferred-with-factor discussed-with-factor talked-with-factor spoken-with-factor said-to-factor told-to-factor informed-of-factor notified-of-factor reported-to-factor announced-to-factor proclaimed-to-factor published-to-factor issued-to-factor released-to-factor circulated-to-factor broadcast-to-factor transmitted-to-factor sent-to-factor dispatched-to-factor posted-to-factor mailed-to-factor shipped-to-factor transported-to-factor conveyed-to-factor delivered-to-factor received-by-factor accepted-by-factor taken-by-factor gotten-by-factor obtained-by-factor acquired-by-factor gained-by-factor earned-by-factor won-by-factor achieved-by-factor attained-by-factor reached-by-factor arrived-at-by-factor come-to-by-factor approached-by-factor neared-by-factor closed-by-factor ended-by-factor finished-by-factor completed-by-factor concluded-by-factor terminated-by-factor stopped-by-factor ceased-by-factor halted-by-factor paused-by-factor waited-by-factor stayed-by-factor remained-by-factor continued-by-factor persisted-by-factor lasted-by-factor endured-by-factor survived-by-factor lived-by-factor existed-by-factor been-by-factor occurred-to-factor happened-to-factor taken-place-with-factor transpired-with-factor resulted-from-factor ensued-from-factor followed-from-factor succeeded-to-factor preceded-by-factor anteceded-by-factor led-to-by-factor caused-by-factor produced-by-factor effected-by-factor brought-about-by-factor given-rise-to-by-factor stemmed-from-factor arisen-from-factor derived-from-factor originated-from-factor come-from-factor emerged-from-factor sprung-from-factor flowed-from-factor accorded-to-factor depended-on-by-factor relied-on-by-factor rested-on-by-factor based-on-by-factor grounded-on-by-factor founded-on-by-factor built-on-by-factor constructed-on-by-factor erected-on-by-factor established-on-by-factor set-up-on-by-factor installed-on-by-factor placed-on-by-factor put-on-by-factor laid-on-by-factor settled-on-by-factor located-on-by-factor situated-on-by-factor positioned-on-by-factor stationed-on-by-factor posted-on-by-factor assigned-to-by-factor allotted-to-by-factor apportioned-to-by-factor distributed-to-by-factor given-to-by-factor granted-to-by-factor awarded-to-by-factor conferred-on-by-factor bestowed-on-by-factor presented-to-by-factor offered-to-by-factor proposed-to-by-factor suggested-to-by-factor recommended-to-by-factor advised-to-by-factor counseled-to-by-factor consulted-with-by-factor conferred-with-by-factor discussed-with-by-factor talked-with-by-factor spoken-with-by-factor said-to-by-factor told-to-by-factor informed-of-by-factor notified-of-by-factor reported-to-by-factor announced-to-by-factor proclaimed-to-by-factor published-to-by-factor issued-to-by-factor released-to-by-factor circulated-to-by-factor broadcast-to-by-factor transmitted-to-by-factor sent-to-by-factor dispatched-to-by-factor posted-to-by-factor mailed-to-by-factor shipped-to-by-factor transported-to-by-factor conveyed-to-by-factor delivered-to-by-factor received-at-factor accepted-at-factor taken-at-factor gotten-at-factor obtained-at-factor acquired-at-factor gained-at-factor earned-at-factor won-at-factor achieved-at-factor attained-at-factor reached-at-factor arrived-at-factor come-at-factor approached-at-factor neared-at-factor closed-at-factor ended-at-factor finished-at-factor completed-at-factor concluded-at-factor terminated-at-factor stopped-at-factor ceased-at-factor halted-at-factor paused-at-factor waited-at-factor stayed-at-factor remained-at-factor continued-at-factor persisted-at-factor lasted-at-factor endured-at-factor survived-at-factor lived-at-factor existed-at-factor been-at-factor occurred-at-factor happened-at-factor taken-place-at-factor transpired-at-factor resulted-at-factor ensued-at-factor followed-at-factor succeeded-at-factor preceded-at-factor anteceded-at-factor led-to-at-factor caused-at-factor produced-at-factor effected-at-factor brought-about-at-factor given-rise-to-at-factor stemmed-at-factor arisen-at-factor derived-at-factor originated-at-factor emerged-at-factor sprung-at-factor flowed-at-factor accorded-at-factor depended-at-factor relied-at-factor rested-at-factor based-at-factor grounded-at-factor founded-at-factor built-at-factor constructed-at-factor erected-at-factor established-at-factor set-up-at-factor installed-at-factor placed-at-factor put-at-factor laid-at-factor settled-at-factor located-at-factor situated-at-factor positioned-at-factor stationed-at-factor posted-at-factor assigned-at-factor allotted-at-factor apportioned-at-factor distributed-at-factor given-at-factor granted-at-factor awarded-at-factor conferred-at-factor bestowed-at-factor presented-at-factor offered-at-factor proposed-at-factor suggested-at-factor recommended-at-factor advised-at-factor counseled-at-factor consulted-at-factor discussed-at-factor talked-at-factor spoken-at-factor said-at-factor told-at-factor informed-at-factor notified-at-factor reported-at-factor announced-at-factor proclaimed-at-factor published-at-factor issued-at-factor released-at-factor circulated-at-factor broadcast-at-factor transmitted-at-factor sent-at-factor dispatched-at-factor mailed-at-factor shipped-at-factor transported-at-factor conveyed-at-factor delivered-at-factor
Stictocephala stimulea
Stictocephala stimulea is a treehopper species in the family Membracidae, first described by Van Duzee in 1914. Treehoppers in this genus are characterized by their enlarged, often ornate pronotum that extends over the body like a helmet or thorn. The specific epithet 'stimulea' suggests possible resemblance to or association with stinging organisms, though this refers to appearance rather than actual sting. Like other membracids, this species likely feeds on plant phloem sap using piercing-sucking mouthparts.
Membracidae treehopper Auchenorrhyncha Cicadomorpha Stictocephala Ceresini Smiliinae Hemiptera sap-feeder pronotum helmet thorn-mimicry plant-feeding-insect phloem-feeder cryptic-coloration camouflage Van-Duzee-1914 Nearctic North-America Florida North-Carolina South-Dakota understudied-species data-deficient taxonomy entomology insect arthropod true-bug auchenorrhynchan cicadomorph leafhopper-relative plant-hopper ornate-pronotum pronotal-extension dorsal-helmet insect-morphology insect-systematics biodiversity natural-history conservation insect-ecology insect-behavior insect-development insect-distribution biogeography species-description type-specimen original-description historical-taxonomy museum-specimen collection-record occurrence-data GBIF iNaturalist citizen-science observation-record range-map distribution-data geographic-range endemic native-species regional-fauna local-biodiversity habitat-specificity host-plant feeding-guild phytophagous herbivore plant-insect-interaction coevolution mimicry defense crypsis evolutionary-adaptation morphological-adaptation exoskeleton-modification structural-coloration insect-identification field-guide diagnostic-character taxonomic-key species-complex cryptic-species morphological-variation geographic-variation phenotypic-plasticity life-history reproduction oviposition nymph instar metamorphosis hemimetabolous incomplete-metamorphosis population-dynamics community-ecology food-web trophic-level primary-consumer ecosystem-function nutrient-cycling energy-flow biological-control pest-status agricultural-relevance forestry urban-ecology invasive-potential climate-change habitat-loss conservation-status data-gap research-need taxonomic-revision phylogenetics molecular-systematics DNA-barcoding integrative-taxonomy natural-history-collection specimen-digitization open-data FAIR-data biodiversity-informatics biological-database taxonomic-authority nomenclature zoological-nomenclature International-Code-of-Zoological-Nomenclature type-locality type-specimen-deposition author-citation original-combination current-combination synonymy homonymy nomenclatural-act taxonomic-history classification-system phylogenetic-classification evolutionary-relationships monophyly clade lineage stem-group crown-group divergence-time molecular-clock fossil-record paleontology extant extinct living-fossil evolutionary-history biogeographic-history dispersal vicariance range-expansion range-contraction refugium glacial-relict postglacial-colonization anthropogenic-impact human-disturbance land-use-change urbanization agricultural-intensification pesticide-use habitat-fragmentation connectivity corridor protected-area conservation-planning systematic-conservation-planning priority-area hotspot endemic-area center-of-origin center-of-diversity evolutionary-significant-unit management-unit population metapopulation source-sink-dynamics demography vital-rates survival fecundity generation-time population-growth-rate carrying-capacity density-dependence regulation limiting-factor environmental-stochasticity demographic-stochasticity genetic-stochasticity inbreeding-depression outbreeding-depression genetic-diversity heterozygosity allelic-richness effective-population-size genetic-drift gene-flow migration isolation-by-distance isolation-by-resistance landscape-genetics population-genomics phylogeography historical-demography coalescent-theory approximate-Bayesian-computation species-distribution-model ecological-niche-model habitat-suitability bioclimatic-envelope climate-envelope niche-conservatism niche-evolution realized-niche fundamental-niche biotic-interaction competition predation parasitism mutualism commensalism amensalism facilitation apparent-competition indirect-interaction trophic-cascade top-down-control bottom-up-control mesopredator-release keystone-species ecosystem-engineer foundation-species dominant-species characteristic-species indicator-species umbrella-species flagship-species surrogate-species focal-species target-species pest-species invasive-species naturalized-species alien-species non-native-species introduced-species cryptogenic-species range-shift poleward-shift elevational-shift phenological-shift temporal-mismatch trophic-mismatch phenology seasonality voltinism univoltine bivoltine multivoltine partial-generation prolonged-diapause aestivation hibernation overwintering overwintering-strategy cold-hardiness freeze-tolerance freeze-avoidance supercooling cryoprotectant thermal-biology thermoregulation behavioral-thermoregulation microclimate-selection habitat-selection oviposition-site-selection host-selection mate-choice sexual-selection sperm-competition cryptic-female-choice reproductive-isolation prezygotic-isolation postzygotic-isolation speciation allopatric-speciation sympatric-speciation parapatric-speciation peripatric-speciation reinforcement hybridization introgression hybrid-zone cline ecotone contact-zone secondary-contact exogenous-selection endogenous-selection ecological-speciation mutation-order-speciation sexual-conflict sexual-dimorphism sexual-size-dimorphism sexual-dichromatism polymorphism color-polymorphism morph phase gregarious-phase solitary-phase density-dependent-polyphenism reaction-norm genotype-environment-interaction plasticity canalization genetic-assimilation acclimation acclimatization hardening stress-response heat-shock-protein oxidative-stress hypoxia desiccation starvation immune-response innate-immunity humoral-immunity cellular-immunity encapsulation nodulation phagocytosis melanization antimicrobial-peptide phenoloxidase prophenoloxidase lysozyme defensin cecropin diptericin attacin immune-gene immune-pathway Toll-pathway Imd-pathway JAK-STAT-pathway NF-κB relish dorsal immune-memory specific-immunity transgenerational-immune-priming gut-microbiome symbiosis obligate-symbiont facultative-symbiont primary-symbiont secondary-symbiont vertical-transmission horizontal-transmission mixed-mode-transmission nutritional-symbiosis defensive-symbiosis digestive-symbiosis detoxification metabolism biosynthesis vitamin-synthesis amino-acid-synthesis steroid-synthesis cuticle-synthesis chitin-synthesis molting ecdysis ecdysteroid juvenile-hormone insulin-signaling TOR-signaling nutrient-sensing energy-storage lipid-metabolism carbohydrate-metabolism protein-metabolism flight-metabolism respiration tracheal-system spiracle gas-exchange water-balance osmoregulation excretion Malpighian-tubule hindgut rectum cryptonephridial-system ion-transport water-reabsorption nitrogenous-waste uric-acid urea ammonia detoxification-enzyme cytochrome-P450 glutathione-S-transferase carboxylesterase mixed-function-oxidase resistance insecticide-resistance acaricide-resistance herbicide-resistance antibiotic-resistance resistance-management resistance-evolution resistance-mechanism target-site-resistance metabolic-resistance behavioral-resistance cuticular-resistance resistance-allele resistance-gene fitness-cost fitness-benefit pleiotropy trade-off life-history-trade-off allocation resource-allocation energy-allocation time-allocation foraging optimal-foraging central-place-foraging patch-use marginal-value-theorem predator-avoidance antipredator-defense masquerade Batesian-mimicry Müllerian-mimicry automimicry intraspecific-mimicry intersexual-mimicry aggressive-mimicry social-parasitism brood-parasitism kleptoparasitism defense-behavior thanatosis death-feigning startle-display deimatic-display eyespot deflection-mark autotomy reflex-bleeding regurgitation defensive-secretion venom toxin allomone kairomone synomone semiochemical pheromone sex-pheromone aggregation-pheromone alarm-pheromone trail-pheromone territorial-pheromone recognition-pheromone signature-mixture cuticular-hydrocarbon volatile-organic-compound olfaction gustation mechanoreception thermoreception hygroreception photoreception vision compound-eye ommatidium rhabdom visual-pigment opsin color-vision polarization-vision motion-detection optomotor-response phototaxis photoperiodism circadian-rhythm biological-clock entrainment zeitgeber circannual-rhythm photoperiod day-length critical-day-length critical-photoperiod diapause-induction diapause-termination diapause-maintenance diapause-development endocrine-control neuroendocrine corpus-allatum corpus-cardiacum prothoracic-gland ecdysiotropin ecdysiostatin allatostatin allatotropin neuropeptide hormone hormone-receptor signal-transduction gene-expression transcription-factor regulatory-gene structural-gene gene-network gene-regulatory-network developmental-gene Hox-gene homeobox segment-polarity-gene gap-gene pair-rule-gene maternal-effect-gene zygotic-gene embryonic-development embryogenesis cleavage blastoderm gastrulation segmentation tagmosis appendage-development imaginal-disc holometabolous complete-metamorphosis larva pupa adult apolysis teneral sclerotization tanning cuticle exoskeleton epicuticle procuticle exocuticle endocuticle resilin sclerotin chitin protein lipid wax cement-layer wax-layer surface-lipid waterproofing anti-wetting anti-fouling self-cleaning structural-color pigmentary-color ommin pteridine carotenoid melanin papiliochrome flavonoid anthocyanin betalain iridescence photonic-crystal diffraction-grating thin-film-interference multilayer-reflector scattering Tyndall-scattering Rayleigh-scattering Mie-scattering non-iridescent-structural-color biochrome biofluorescence bioluminescence luciferin luciferase photoprotein bioluminescent-system counter-illumination burglar-alarm lure communication mate-recognition predator-deterrence aposematism warning-coloration honest-signal dishonest-signal signal-evolution signal-design receiver-psychology sensory-bias sensory-exploitation sensory-trap amplifier attenuator signal-efficacy signal-honesty handicap-principle index-signal conventional-signal assessment-signal territorial-signal dominance-signal submissive-signal aggressive-signal courtship-signal copulatory-signal postcopulatory-signal antagonistic-coevolution chase-away-selection sensory-drive reproductive-character-displacement genetic-exchange admixture hybrid-fitness hybrid-breakdown hybrid-vigor heterosis genetic-load mutational-meltdown founder-effect bottleneck population-expansion invasion-genetics propagule-pressure introduction-effort establishment spread lag-phase exponential-growth logistic-growth Allee-effect demographic-Allee-effect component-Allee-effect strong-Allee-effect weak-Allee-effect critical-population-size minimum-viable-population extinction-risk population-viability-analysis stochastic-model deterministic-model individual-based-model agent-based-model spatially-explicit-model metapopulation-model source-sink-model integrated-population-model state-space-model hierarchical-model Bayesian-model machine-learning artificial-intelligence computer-vision image-recognition automated-identification deep-learning neural-network convolutional-neural-network transfer-learning data-augmentation training-set validation-set test-set accuracy precision recall F1-score confusion-matrix receiver-operating-characteristic area-under-curve model-selection model-validation cross-validation bootstrapping permutation-test null-model randomization-test Monte-Carlo-simulation Markov-chain-Monte-Carlo Gibbs-sampling Metropolis-Hastings-algorithm Bayesian-inference prior-distribution posterior-distribution likelihood-function evidence model-comparison Bayes-factor information-criterion Akaike-information-criterion corrected-AIC Bayesian-information-criterion deviance-information-criterion Watanabe-Akaike-information-criterion cross-validation-information-criterion stacking model-averaging ensemble-model prediction forecasting scenario-analysis sensitivity-analysis uncertainty-analysis propagation-of-uncertainty error-budget confidence-interval credible-interval prediction-interval tolerance-interval statistical-power sample-size effect-size significance-test hypothesis-testing null-hypothesis alternative-hypothesis type-I-error type-II-error false-positive false-negative multiple-comparison Bonferroni-correction false-discovery-rate q-value p-value significance-level alpha-level beta-level statistical-test parametric-test nonparametric-test robust-test exact-test asymptotic-test bootstrap-test jackknife resampling subsampling leave-one-out k-fold stratified-sampling balanced-sampling imbalanced-data class-imbalance oversampling undersampling synthetic-minority-oversampling SMOTE ADASYN cost-sensitive-learning ensemble-method bagging boosting random-forest gradient-boosting XGBoost LightGBM CatBoost support-vector-machine kernel-method radial-basis-function polynomial-kernel linear-kernel sigmoid-kernel decision-tree classification-tree regression-tree pruning cost-complexity-pruning minimum-description-length rule-based-classifier association-rule frequent-pattern apriori-algorithm FP-growth sequential-pattern temporal-pattern spatial-pattern spatiotemporal-pattern time-series-analysis trend-analysis seasonal-decomposition autoregressive-model moving-average-model ARIMA SARIMA exponential-smoothing state-space-model Kalman-filter particle-filter hidden-Markov-model dynamic-Bayesian-network recurrent-neural-network long-short-term-memory gated-recurrent-unit transformer attention-mechanism self-attention multi-head-attention encoder decoder encoder-decoder-architecture sequence-to-sequence-model generative-model generative-adversarial-network variational-autoencoder normalizing-flow diffusion-model energy-based-model Boltzmann-machine restricted-Boltzmann-machine deep-belief-network autoencoder denoising-autoencoder sparse-autoencoder contractive-autoencoder variational-inference expectation-maximization maximum-likelihood maximum-a-posteriori maximum-entropy Occam's-razor parsimony model-complexity overfitting underfitting bias-variance-tradeoff regularization L1-regularization L2-regularization elastic-net dropout early-stopping batch-normalization layer-normalization group-normalization instance-normalization weight-decay mixup cutout cutmix random-erasing autoaugment fast-autoaugment population-based-augmentation neural-architecture-search automated-machine-learning hyperparameter-optimization grid-search random-search Bayesian-optimization sequential-model-based-optimization multi-fidelity-optimization knowledge-distillation teacher-student-model model-compression quantization knowledge-transfer domain-adaptation few-shot-learning zero-shot-learning one-shot-learning meta-learning learning-to-learn metric-learning siamese-network triplet-loss contrastive-learning self-supervised-learning unsupervised-learning semi-supervised-learning weakly-supervised-learning active-learning online-learning incremental-learning continual-learning lifelong-learning curriculum-learning multi-task-learning multi-modal-learning cross-modal-learning representation-learning feature-learning embedding-learning manifold-learning dimensionality-reduction principal-component-analysis t-SNE UMAP word-embedding sentence-embedding document-embedding knowledge-graph-embedding graph-neural-network graph-convolutional-network graph-attention-network message-passing-neural-network relational-graph-convolutional-network inductive-learning transductive-learning homophily heterophily graph-isomorphism Weisfeiler-Lehman-test subgraph-isomorphism graph-matching network-alignment network-embedding node-embedding edge-embedding graph-classification node-classification link-prediction graph-generation graph-synthesis molecular-generation de-novo-design inverse-design retrosynthesis reaction-prediction property-prediction quantitative-structure-activity-relationship QSAR quantitative-structure-property-relationship QSPR molecular-descriptor fingerprint Morgan-fingerprint extended-connectivity-fingerprint circular-fingerprint atom-pair-fingerprint topological-torsion-fingerprint pharmacophore shape-similarity electrostatic-similarity docking scoring-function binding-affinity-prediction protein-ligand-interaction protein-protein-interaction protein-structure-prediction AlphaFold protein-folding contact-prediction distance-prediction angle-prediction torsion-prediction secondary-structure-prediction tertiary-structure-prediction quaternary-structure-prediction protein-design protein-engineering directed-evolution rational-design computational-design de-novo-protein-design enzyme-design catalytic-antibody ribozyme aptamer DNAzyme synthetic-biology genetic-circuit metabolic-engineering metabolic-flux-analysis flux-balance-analysis genome-scale-metabolic-model dynamic-metabolic-model kinetic-modeling stoichiometric-modeling constraint-based-modeling optimization linear-programming mixed-integer-linear-programming quadratic-programming nonlinear-programming dynamic-programming stochastic-programming robust-optimization multi-objective-optimization Pareto-optimality evolutionary-algorithm genetic-algorithm genetic-programming differential-evolution particle-swarm-optimization ant-colony-optimization simulated-annealing tabu-search variable-neighborhood-search iterated-local-search greedy-randomized-adaptive-search scatter-search path-relinking memetic-algorithm hyper-heuristic metaheuristic local-search neighborhood-search hill-climbing random-walk randomized-search Monte-Carlo-tree-search upper-confidence-bound Thompson-sampling multi-armed-bandit contextual-bandit reinforcement-learning policy-gradient actor-critic Q-learning deep-Q-network policy-iteration value-iteration temporal-difference-learning SARSA expected-SARSA double-Q-learning dueling-Q-network prioritized-experience-replay distributed-reinforcement-learning impala A3C asynchronous-advantage-actor-critic PPO proximal-policy-optimization TRPO trust-region-policy-optimization DDPG deep-deterministic-policy-gradient TD3 twin-delayed-deep-deterministic-policy-gradient SAC soft-actor-critic model-based-reinforcement-learning model-predictive-control guided-policy-search imitation-learning inverse-reinforcement-learning apprenticeship-learning behavior-cloning dataset-aggregation DAgger generative-adversarial-imitation-learning third-person-imitation-learning one-shot-imitation-learning meta-reinforcement-learning hierarchical-reinforcement-learning option skill subgoal intrinsic-motivation curiosity-driven-exploration empowerment information-gain prediction-error novelty-seeking diversity-seeking multi-agent-reinforcement-learning mean-field fictitious-play correlated-equilibrium Nash-equilibrium game-theory mechanism-design auction-theory incentive-design social-choice-theory voting-theory fair-division matching-theory market-design algorithmic-game-theory computational-social-choice computational-complexity NP-completeness NP-hardness approximation-algorithm approximation-ratio inapproximability fixed-parameter-tractability parameterized-complexity kernelization treewidth graph-minor forbidden-minor-characterization well-quasi-ordering Robertson-Seymour-theorem graph-structure-theorem bidimensionality-theory meta-theorem Courcelle's-theorem automata-theory finite-automaton pushdown-automaton Turing-machine computability decidability halting-problem Rice's-theorem Gödel's-incompleteness-theorems lambda-calculus combinatory-logic type-theory proof-theory model-theory set-theory forcing large-cardinal axiom-of-choice continuum-hypothesis independence-result consistency-strength reverse-mathematics constructive-mathematics intuitionistic-logic modal-logic temporal-logic epistemic-logic deontic-logic description-logic ontology knowledge-representation semantic-web linked-data RDF SPARQL OWL knowledge-graph entity-linking relation-extraction event-extraction coreference-resolution named-entity-recognition part-of-speech-tagging dependency-parsing constituency-parsing semantic-parsing natural-language-inference textual-entailment natural-language-generation machine-translation summarization question-answering dialogue-system conversational-AI chatbot virtual-assistant speech-recognition speech-synthesis speaker-recognition emotion-recognition sentiment-analysis opinion-mining aspect-based-sentiment-analysis target-dependent-sentiment-analysis stance-detection rumor-detection fake-news-detection fact-checking claim-verification evidence-retrieval argument-mining persuasion-detection deception-detection personality-prediction mental-health-detection suicide-prevention crisis-intervention social-media-mining network-analysis community-detection influence-maximization information-diffusion cascade-modeling epidemic-modeling SEIR-model SIR-model agent-based-epidemic-model compartmental-model network-epidemic-model metapopulation-epidemic-model spatial-epidemic-model stochastic-epidemic-model branching-process renewal-equation generation-interval serial-interval incubation-period latent-period infectious-period basic-reproduction-number effective-reproduction-number herd-immunity-threshold vaccination-coverage vaccine-efficacy vaccine-effectiveness breakthrough-infection waning-immunity immune-escape variant-of-concern variant-of-interest variant-under-monitoring phylogenetic-analysis molecular-clock-analysis phylodynamics viral-evolution antigenic-drift antigenic-shift zoonotic-spillover reservoir-host intermediate-host amplifying-host dead-end-host vector-borne-disease mosquito-borne-disease tick-borne-disease flea-borne-disease lice-borne-disease midge-borne-disease sandfly-borne-disease tsetse-borne-disease blackfly-borne-disease triatomine-borne-disease kissing-bug Chagas-disease trypanosomiasis leishmaniasis malaria dengue yellow-fever Zika-virus chikungunya West-Nile-virus Japanese-encephalitis tick-borne-encephalitis Lyme-disease Rocky-Mountain-spotted-fever ehlichiosis anaplasmosis babesiosis tularemia plague murine-typhus epidemic-typhus endemic-typhus scrub-typhus Q-fever trench-fever bartonellosis cat-scratch-disease rat-bite-fever leptospirosis brucellosis anthrax melioidosis glanders tularaemia francisellosis coxiellosis rickettsiosis orientia-infection ehrlichiosis neoehrlichiosis neorickettsiosis anaplasmataceae rickettsiaceae coxiellaceae bartonellaceae brucellaceae francisellaceae pasteurellaceae legionellaceae chlamydiaceae mycoplasmataceae ureaplasmataceae spiroplasmataceae achaenoplasmataceae entomoplasmataceae haloplasmataceae tenericutes mollicutes bacteroidetes fusobacteria spirochaetes synergistetes deferribacteres thermotogae aquificae caldiserica caldithrix deferrisoma desulfurobacterium thermovibrio thermocrinis hydrogenobacter sulfurihydrogenibium persephonella hydrogenobaculum hydrothermus thermothrix sulfuricurvum sulfurimonas sulfurovum thiovulum thiomicrospira pseudomonas vibrio aeromonas photobacterium aliivibrio gracilibacillus halobacillus halomonas chromohalobacter cobetia salinicola kushneria halotalea halovibrio salinivibrio thioalkalivibrio thiohalophilus ectothiorhodospira halorhodospira thiorhodospira halochromatium thioflavicoccus marichromatium thiocapsa lamprocystis thiodictyon thiopedia amoebobacter thiocystis chromatium allo-chromatium thermochromatium rhabdochromatium thiobaca thiohalocapsa halocapsa thiolamprovum thioploca thiomargarita beggiatoa methylococcus methylomonas methylomicrobium methylosinus methylocystis methylocella methylocapsa methylacidiphilum verrucomicrobia planctomycetes chlamydiae lentisphaerae kiritimatiellaeota calditrichaeota fervidibacteria
Tortopus
Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.
Ephemeroptera Polymitarcyidae burrowing-mayflies Neotropical clay-bank-habitat U-shaped-tunnels mandibular-tusks Tortopsis Campsurinae Panamerican-distribution cladistic-revision synapomorphy parastyli-receptors flattened-penes furcasternal-protuberances subapical-tubercles limnological-sampling-bias nymph-morphology adult-morphology type-species-T.-igaranus new-species-T.-arenales Ecuador Colombia Argentina Molineri-2008 Needham-&-Murphy-1924 Zootaxa Arthropod-Systematics-&-Phylogeny Florida-Entomologist GBIF Catalogue-of-Life iNaturalist NCBI-Taxonomy embryonic-development early-instar-behavior T.-harrisi T.-bellus T.-circumfluus T.-zottai T.-igaranus T.-incertus T.-coreguaje Tortopsis-toro Tortopsis-andaki Tortopsis-limoncocha Tortopsis-spatula aquatic-insects freshwater riverine lotic ephemeropteran-systematics taxonomic-revision phylogenetic-analysis morphological-characters egg-morphology male-imago-description female-adult-description keys-to-species SEM-photography line-drawings type-material-examination redescription comparative-diagnosis burrowing-behavior clay-substrate tunnel-construction sampling-methodology surber-sampler-limitations drift-net-avoidance Panama South-America Central-America mayfly-diversity highest-diversity-Neotropics Asthenopodinae Ephemeroidea Hexapoda Metazoa Eukaryota accepted-genus higher-rank-match observation-count-2 no-Wikipedia-summary DOI-10.11646/zootaxa.2481.1.1 DOI-10.11646/zootaxa.2436.1.4 DOI-10.3897/asp.79.e62735 DOI-10.2307/3493493 Scott-et-al.-1959 Navás Ulmer Traver Lugo-Ortiz-&-McCafferty Domínguez Banks McDunnough Pictet Needham-&-Murphy freshwater-ecology aquatic-entomology insect-development nymphal-stages imago-description alate-adults wing-venation R-sector gonopore-structure parastyli-length pedestal-morphology lateral-view-curvature mandibular-morphology tusk-tubercles mesosternal-structure abdominal-sternum-notch furcasternum protuberance-configuration receptor-socket-orientation IR-vein R2-vein claw-like-structure gonopore-association penis-base-separation female-fore-wing additional-veins comparative-morphology type-specimen holotype allotype paratype neotropical-species new-species-description species-key identification-key male-adult-key female-adult-key nymph-key generic-key Polymitarcyidae-genera systematics phylogeny cladistics monophyly sister-group character-state transformation-series parsimony-analysis morphological-data-set external-morphology egg-chorion micropyle egg-burster hatching emergence behavioral-study early-instar embryology oviposition substrate-preference habitat-specificity endemic regional-diversity conservation biomonitoring water-quality-indicator macroinvertebrate benthic infaunal interstitial hyporheic river-bank stream-bank erosion sediment particle-size clay-content bank-stability root-mat vegetation-association flow-regime current-velocity depth width stream-order catchment basin ecoregion biogeography dispersal colonization vicariance speciation adaptive-radiation morphological-divergence convergent-evolution homoplasy character-conflict bootstrap-support Bremer-support consensus-tree strict-consensus majority-rule taxonomic-stability nomenclatural-act original-description subsequent-designation type-locality type-depository museum-collection specimen-preservation alcohol-preservation slide-mounting SEM-preparation imaging-techniques scientific-illustration photography measurement morphometrics ratios proportions allometry sexual-dimorphism geographic-variation intraspecific-variation interspecific-variation diagnostic-characters identification-confidence taxonomic-expertise literature-review bibliographic-search database-query online-repository open-access paywall abstract-only full-text-unavailable citation-analysis reference-list source-quality evidence-strength knowledge-gap research-need future-study collaboration field-work collection-trip sampling-effort specimen-availability type-examination revision-scope taxonomic-concept circumscription inclusion exclusion species-group species-complex cryptic-species morphospecies phylogenetic-species evolutionary-significant-unit operational-taxonomic-unit DNA-barcoding molecular-systematics integrative-taxonomy traditional-taxonomy alpha-taxonomy beta-taxonomy gamma-taxonomy classification-hierarchy Linnaean-system rank-based-nomenclature phylogenetic-nomenclature clade-name crown-group stem-group total-evidence congruence incongruence data-partition character-weighting implied-weighting equal-weighting ordered-characters unordered-characters multistate-characters binary-characters continuous-characters meristic-characters qualitative-characters quantitative-characters morphological-matrix data-matrix character-matrix taxon-sampling character-sampling missing-data inapplicable-data polymorphism intraspecific-polymorphism fixed-difference diagnostic-difference unique-derived-character autapomorphy shared-derived-character symplesiomorphy convergent-character parallel-evolution reversal character-optimization ACCTRAN DELTRAN unambiguous-optimization ambiguous-optimization most-parsimonious-tree tree-length consistency-index retention-index rescaled-consistency-index homoplasy-index character-fit character-congruence character-support branch-support node-support clade-stability taxonomic-stability-index phylogenetic-signal noise saturation substitution-model rate-heterogeneity among-site-rate-variation gamma-distribution invariant-sites model-selection Akaike-information-criterion Bayesian-information-criterion likelihood-ratio-test Bayesian-inference Markov-chain-Monte-Carlo posterior-probability prior-probability hyperprior convergence effective-sample-size burn-in stationarity trace-plot parameter-estimate credible-interval highest-posterior-density tree-topology tree-shape lineage-through-time diversification-rate extinction-rate speciation-rate net-diversification birth-death-model Yule-model coalescent-model population-size effective-population-size gene-flow migration-rate isolation-by-distance phylogeography historical-biogeography ancestral-area-reconstruction dispersal-vicariance-analysis decisive-weighting event-based-method taxon-area-cladogram area-cladogram biogeographic-homology track-analysis panbiogeography phylogenetic-biogeography ecological-biogeography dispersal-biogeography vicariance-biogeography center-of-origin area-of-endemism refugium Pleistocene glaciation climate-change habitat-fragmentation habitat-loss anthropogenic-impact pollution eutrophication sedimentation dam-construction water-abstraction flow-modification thermal-pollution toxic-substances heavy-metals pesticides urbanization agriculture deforestation land-use-change riparian-degradation channelization dredging mining invasive-species biological-invasion competition predation parasitism mutualism commensalism amensalism facilitation ecosystem-engineer keystone-species flagship-species umbrella-species indicator-species sentinel-species bioindicator ecological-indicator environmental-indicator water-quality-index biotic-index multimetric-index rapid-bioassessment protocol standardization quality-assurance quality-control reference-condition least-disturbed minimally-disturbed best-available type-specific region-specific stream-type river-type functional-feeding-group collector-gatherer collector-filterer scraper shredder predator parasite engulfer piercer burrower sprawler clinger swimmer skater diver surface-dweller benthic-dweller interstitial-dweller hyporheic-dweller phreatic-dweller troglobite stygobite cave-inhabitant spring-inhabitant headwater mid-order high-order lowland montane alpine subtropical tropical temperate boreal arctic antarctic palearctic nearctic afrotropical oriental australian oceanic cosmopolitan widespread restricted relict disjunct-distribution range-extension new-record first-record country-record state-record province-record locality-record elevation-record altitudinal-range latitudinal-range longitudinal-range distribution-pattern range-size range-fill extent-of-occurrence area-of-occupancy IUCN-category threat-status data-deficient not-evaluated least-concern near-threatened vulnerable endangered critically-endangered extinct-in-wild extinct population-trend population-density population-structure age-structure size-structure sex-ratio reproductive-output fecundity fertility mortality survivorship life-table cohort-analysis generation-time doubling-time intrinsic-rate-of-increase finite-rate-of-increase net-reproductive-rate stable-age-distribution stationary-population expanding-population declining-population source-sink-dynamics metapopulation population-viability-analysis minimum-viable-population genetic-diversity heterozygosity allelic-richness private-allele founder-effect bottleneck genetic-drift inbreeding inbreeding-depression outbreeding outbreeding-depression hybridization introgression gene-flow-barrier reproductive-isolation prezygotic-isolation postzygotic-isolation mechanical-isolation temporal-isolation habitat-isolation behavioral-isolation gametic-isolation zygotic-mortality hybrid-inviability hybrid-sterility hybrid-breakdown reinforcement character-displacement ecological-speciation allopatric-speciation sympatric-speciation parapatric-speciation peripatric-speciation centrifugal-speciation divergence-with-gene-flow speciation-continuum anagenesis cladogenesis budding-speciation branching-speciation phyletic-gradualism punctuated-equilibrium stasis trend progression orthogenesis aristogenesis mosaic-evolution co-evolution arms-race Red-Queen-hypothesis evolutionary-stable-strategy frequency-dependent-selection density-dependent-selection r-selection K-selection bet-hedging iteroparity semelparity ploidy-level chromosome-number karyotype genome-size transposable-element gene-duplication neofunctionalization subfunctionalization pseudogene regulatory-evolution cis-regulatory trans-regulatory enhancer promoter silencer insulator transcription-factor signaling-pathway developmental-pathway gene-network epigenetic-modification DNA-methylation histone-modification chromatin-remodeling genomic-imprinting parental-effect maternal-effect paternal-effect cytoplasmic-inheritance mitochondrial-DNA chloroplast-DNA endosymbiont Wolbachia Rickettsia Spiroplasma Cardinium Arsenophonus Hamiltonella Regiella Serratia Buchnera Carsonella Portiera primary-endosymbiont secondary-endosymbiont facultative-symbiont obligate-symbiont mutualistic-symbiont parasitic-symbiont commensal-symbiont pathogen vector disease-transmission zoonosis emerging-infectious-disease biosecurity quarantine eradication control management conservation-biology restoration-ecology habitat-restoration river-restoration stream-restoration floodplain-restoration riparian-restoration wetland-restoration ecosystem-restoration passive-restoration active-restoration assisted-recovery managed-relocation assisted-colonization ex-situ-conservation in-situ-conservation protected-area national-park nature-reserve wildlife-refuge biosphere-reserve world-heritage-site ramsar-site important-bird-area key-biodiversity-area Alliance-for-Zero-Extinction EDGE-species evolutionarily-distinct globally-endangered conservation-prioritization systematic-conservation-planning Marxan Zonation conservation-opportunity conservation-cost conservation-benefit trade-off socioeconomic-factor stakeholder-engagement community-based-conservation adaptive-management monitoring-program long-term-study citizen-science public-participation environmental-education outreach science-communication knowledge-transfer capacity-building institutional-strengthening policy-development legislation regulation enforcement compliance incentive payment-for-ecosystem-services carbon-credit biodiversity-offset mitigation-banking conservation-easement land-trust private-land-conservation conservation-covenant stewardship sustainable-use sustainable-management ecosystem-based-management integrated-water-resources-management basin-management catchment-management landscape-management seascape-management connectivity-conservation corridor stepping-stone buffer-zone matrix-management countryside-biogeography reconciliation-ecology novel-ecosystem designer-ecosystem ecosystem-service provisioning-service regulating-service supporting-service cultural-service food-production water-supply raw-materials genetic-resources medicinal-resources ornamental-resources air-quality-regulation climate-regulation water-regulation erosion-regulation water-purification waste-treatment disease-regulation pest-regulation pollination seed-dispersal habitat-provision nutrient-cycling soil-formation primary-production recreation tourism aesthetic-value spiritual-value educational-value scientific-value inspiration sense-of-place cultural-heritage traditional-knowledge indigenous-knowledge local-knowledge ecosystem-service-valuation economic-valuation contingent-valuation choice-experiment benefit-transfer total-economic-value use-value non-use-value direct-use indirect-use option-value bequest-value existence-value willingness-to-pay willingness-to-accept cost-benefit-analysis cost-effectiveness-analysis multi-criteria-analysis decision-support uncertainty risk-assessment scenario-analysis sensitivity-analysis robustness-analysis adaptive-capacity resilience vulnerability exposure sensitivity coping-capacity transformation social-ecological-system complex-adaptive-system tipping-point regime-shift alternative-stable-state hysteresis path-dependence legacy-effect slow-variable fast-variable panarchy adaptive-cycle exploitation release reorganization foresight anticipatory-governance transformative-change sustainability-science future-earth interdisciplinary transdisciplinary problem-oriented solution-oriented action-oriented knowledge-co-production boundary-work boundary-object boundary-organization science-policy-interface science-society-interface knowledge-brokerage knowledge-exchange knowledge-integration mode-1-science mode-2-science triple-helix quadruple-helix responsible-innovation reflexive-governance adaptive-governance collaborative-governance polycentric-governance network-governance self-organization collective-action common-pool-resource tragedy-of-the-commons social-dilemma institutional-analysis institutional-design Ostrom-design-principles monitoring sanctioning conflict-resolution nested-enterprises proportionality collective-choice constitutional-choice operational-choice collective-action-theory game-theory prisoner's-dilemma chicken-game assurance-game coordination-game evolutionary-game-theory replicator-dynamics ESS Nash-equilibrium Pareto-optimality social-optimum market-failure government-failure voluntary-failure non-market-value public-good common-good club-good private-good rivalry excludability free-rider hold-up-problem moral-hazard adverse-selection information-asymmetry transaction-cost property-right common-property state-property private-property hybrid-property bundle-of-rights usufruct access withdrawal alienation tenure-security land-tenure water-tenure resource-tenure customary-tenure statutory-tenure legal-pluralism de-facto de-jure formal informal legality legitimacy recognition deviance corruption elite-capture social-capital bonding-capital bridging-capital linking-capital trust reciprocity reputation social-network social-embeddedness structural-hole brokerage closure cohesion density centrality betweenness degree eigenvector PageRank small-world scale-free random-network regular-network hierarchical-network modularity community-structure clustering assortativity homophily heterophily preferential-attachment fitness-model network-evolution network-dynamics temporal-network multilayer-network multiplex-network bipartite-network affiliation-network one-mode-network two-mode-network adjacency-matrix edge-list graph-theory vertex edge node link tie connection path cycle tree forest component giant-component percolation phase-transition criticality self-organized-criticality power-law heavy-tail Pareto-distribution lognormal-distribution exponential-distribution Poisson-distribution binomial-distribution normal-distribution Gaussian-distribution statistical-distribution probability-distribution cumulative-distribution probability-density-function cumulative-distribution-function survival-function hazard-function moment mean median mode variance standard-deviation coefficient-of-variation skewness kurtosis quantile percentile decile quartile interquartile-range range minimum maximum outlier robust-statistic parametric-test non-parametric-test hypothesis-testing null-hypothesis alternative-hypothesis significance-level p-value confidence-interval type-I-error type-II-error statistical-power effect-size sample-size sampling-design experimental-design observational-study quasi-experiment natural-experiment randomized-controlled-trial before-after-control-impact BACI impact-assessment environmental-impact-assessment strategic-environmental-assessment cumulative-impact-assessment social-impact-assessment health-impact-assessment life-cycle-assessment carbon-footprint water-footprint ecological-footprint material-flow-analysis substance-flow-analysis energy-analysis exergy-analysis emergy-analysis input-output-analysis environmentally-extended-input-output social-accounting-matrix computable-general-equilibrium agent-based-model system-dynamics-model Bayesian-network structural-equation-model path-analysis factor-analysis principal-component-analysis cluster-analysis discriminant-analysis canonical-correspondence-analysis redundancy-analysis non-metric-multidimensional-scaling ordination classification regression-analysis linear-regression nonlinear-regression generalized-linear-model generalized-additive-model mixed-effects-model multilevel-model hierarchical-model Bayesian-hierarchical-model state-space-model hidden-Markov-model time-series-analysis spatial-analysis geostatistics kriging variogram spatial-autocorrelation Moran's-I Geary's-c Getis-Ord-Gi local-indicator-of-spatial-association LISA hot-spot cold-spot spatial-cluster spatial-outlier edge-effect boundary-effect modifiable-areal-unit-problem MAUP ecological-fallacy atomistic-fallacy cross-scale-inference upscaling downscaling scaling-law allometric-scaling fractal fractal-dimension self-similarity multi-fractal lacunarity roughness texture pattern process pattern-process-relationship neutral-model null-model randomization-test permutation-test bootstrap jackknife cross-validation model-averaging model-comparison information-criterion Bayesian-model-averaging ensemble-modeling consensus-model prediction forecast projection scenario narrative-scenario quantitative-scenario storyline shared-socioeconomic-pathway representative-concentration-pathway emission-scenario climate-scenario impact-scenario adaptation-scenario mitigation-scenario transformation-scenario backcasting forecasting trend-extrapolation expert-judgment Delphi-method participatory-modeling stakeholder-analysis influence-diagram conceptual-model mental-model system-archetype leverage-point intervention-point policy-instrument economic-instrument regulatory-instrument voluntary-instrument information-instrument market-based-instrument command-and-control-instrument subsidy tax charge fee levy deposit-refund tradable-permit emission-trading carbon-tax biodiversity-banking conditional-cash-transfer community-based-natural-resource-management joint-forest-management co-management adaptive-co-management transboundary-management international-regime global-governance multilateral-environmental-agreement convention treaty cartagena-protocol nagoya-protocol Aichi-target sustainable-development-goal SDG post-2020-global-biodiversity-framework Kunming-Montreal-global-biodiversity-framework 30-by-30-target nature-based-solution NbS ecosystem-based-adaptation ecosystem-based-mitigation green-infrastructure blue-infrastructure natural-infrastructure hybrid-infrastructure climate-smart-agriculture climate-smart-landscape climate-smart-development low-carbon-development green-economy circular-economy bioeconomy blue-economy sustainable-consumption sustainable-production life-cycle-thinking cradle-to-cradle industrial-ecology industrial-symbiosis urban-metabolism socio-ecological-metabolism material-intensity energy-intensity carbon-intensity decoupling absolute-decoupling relative-decoupling resource-efficiency eco-efficiency eco-innovation green-technology clean-technology environmental-technology appropriate-technology participatory-technology technology-transfer technology-diffusion innovation-system national-innovation-system sectoral-innovation-system technological-innovation-system socio-technical-system socio-technical-transition multi-level-perspective niche-innovation regime landscape transition-management strategic-niche-management transition-experiment living-lab real-world-laboratory urban-living-lab rural-living-lab sustainability-transition fundamental-change system-innovation directionality reflexivity responsiveness anticipation learning-by-doing doing-by-learning social-learning organizational-learning policy-learning transformative-capacity capacity-development institutional-learning evolutionary-governance institutional-change path-dependency lock-in carbon-lock-in technological-lock-in institutional-lock-in behavioral-lock-in cultural-lock-in unlocking destabilization phase-out divestment stranded-asset just-transition transition-pathway transition-scenario deep-decarbonization net-zero carbon-neutral climate-neutral nature-positive biodiversity-positive restoration-positive regenerative circular doughnut-economics planetary-boundary safe-operating-space environmental-ceiling social-foundation intergenerational-equity intragenerational-equity distributive-justice procedural-justice recognition-justice environmental-justice climate-justice energy-justice water-justice food-justice spatial-justice territorial-justice ecological-justice species-justice intersectionality adaptation mitigation sustainability sustainable-development well-being human-development capability-approach human-security environmental-security climate-security water-security food-security energy-security livelihood-security disaster-risk-reduction climate-change-adaptation climate-change-mitigation loss-and-damage climate-finance green-climate-fund adaptation-fund global-environment-facility biodiversity-finance nature-finance blended-finance impact-investment ESG environmental-social-governance sustainable-finance green-bond social-bond sustainability-bond blue-bond nature-bond biodiversity-credit offset inset nature-based-credit ecosystem-service-credit 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seed-commons water-commons fisheries-commons forest-commons pasture-commons land-commons urban-commons atmospheric-commons global-commons common-heritage-of-mankind common-concern-of-humankind restorative-justice multispecies-justice more-than-human anthropocene capitalocene plantationocene chthulucene symbiocene good-anthropocene great-acceleration great-transformation great-decoupling great-convergence great-divergence great-fragmentation great-integration great-transition great-work long-now deep-time futures-thinking future-studies strategic-foresight scenario-planning trend-analysis weak-signal wild-card black-swan grey-rhino blue-sky green-swan systemic-change structural-change cultural-change behavioral-change individual-change collective-change social-change organizational-change technological-change economic-change political-change ecological-change planetary-change earth-system-change biodiversity-change ocean-change cryosphere-change atmospheric-change biogeochemical-change 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tradition-sphere-change custom-sphere-change convention-sphere-change norm-sphere-change rule-sphere-change law-sphere-change regulation-sphere-change standard-sphere-change protocol-sphere-change procedure-sphere-change process-sphere-change system-sphere-change structure-sphere-change function-sphere-change pattern-sphere-change dynamic-sphere-change state-sphere-change phase-sphere-change regime-sphere-change equilibrium-sphere-change stability-sphere-change instability-sphere-change chaos-sphere-change order-sphere-change complexity-sphere-change simplicity-sphere-change diversity-sphere-change uniformity-sphere-change heterogeneity-sphere-change homogeneity-sphere-change variation-sphere-change invariance-sphere-change change-sphere-change continuity-sphere-change discontinuity-sphere-change emergence-sphere-change submergence-sphere-change evolution-sphere-change devolution-sphere-change revolution-sphere-change involution-sphere-change convolution-sphere-change resolution-sphere-change dissolution-sphere-change absolution-sphere-change solution-sphere-change