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Arhopalus
burnt pine longhorn beetle (A. ferus), black pine sawyer (A. rusticus)
Arhopalus is a genus of longhorn beetles (Cerambycidae: Spondylidinae: Asemini) distributed across the Northern Hemisphere, with several species introduced to the Southern Hemisphere as invasive pests of conifers. The genus comprises approximately 20 described species, with A. rusticus and A. ferus being the most extensively studied due to their economic importance. Species develop in dead or fire-damaged conifer wood, with some exhibiting strong attraction to burnt pine volatiles. Several species produce male-emitted aggregation-sex pheromones containing fuscumol and geranylacetone. Invasive populations in New Zealand, Australia, South America, and South Africa require monitoring at ports and sawmills for timber export compliance.
Cerambycidae Spondylidinae Asemini conifer-pest invasive-species pheromone wood-borer pine forestry quarantine New-Zealand Australia South-America biological-invasion climate-change vector Bursaphelenchus-xylophilus integrated-pest-management monitoring trapping semiochemistry burnt-pine fire-damaged-timber sawmill-pest port-surveillance timber-export deforestation forest-health ecosystem-service decomposer nutrient-cycling life-history reproductive-biology ovarian-development larval-development survival fecundity competition coexistence species-distribution-modeling CMIP6 future-climate-scenarios range-expansion quarantine-pest biosecurity aggregation-pheromone fuscumol geranylacetone electroantennography field-trapping trap-efficacy host-volatiles ethanol alpha-pinene beta-pinene monoterpenes green-leaf-volatiles repellent oviposition-behavior crepuscular-activity nocturnal-activity anemotaxis chemotaxis thermal-biology developmental-plasticity laboratory-rearing artificial-diet head-capsule-width instar-number pupal-weight sexual-dimorphism mating-behavior egg-laying survivorship competitive-advantage morphological-anatomy reproductive-system ovipositor-structure nematode-transmission pine-wilt-disease Monochamus vector-competence transmission-potential phytopathogenic-nematode forest-pest-management detection surveillance eradication early-detection rapid-response invasive-species-management cost-effective-monitoring environmentally-safe IPM biological-control natural-enemies population-dynamics spatial-distribution geostatistics landscape-ecology forest-pattern habitat-fragmentation land-use-change southern-hemisphere-invasion global-trade pathway-analysis risk-assessment climate-adaptation thermal-tolerance phenology voltinism generation-time population-genetics molecular-identification barcoding genome-assembly chromosome-level repetitive-DNA transposable-elements evolutionary-biology phylogenetics systematics taxonomy species-identification 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Blissus occiduus
Western Chinch Bug
Blissus occiduus, the western chinch bug, is a phloem-feeding true bug (Hemiptera: Blissidae) that is a significant pest of warm-season turfgrasses, particularly buffalograss and zoysiagrass. The species exhibits strong host preference hierarchies, with buffalograss being the most preferred host followed by zoysiagrass, though it can survive and reproduce on a broad range of grasses including agronomic crops. Field studies have documented inconsistent control with neonicotinoid insecticides, with thiamethoxam showing particularly rapid degradation in buffalograss tissues compared to imidacloprid and clothianidin.
turfgrass-pest chinch-bug phloem-feeder warm-season-grass-pest buffalograss zoysiagrass neonicotinoid-resistance integrated-pest-management Hemiptera Blissidae western-North-America host-preference systemic-insecticide agricultural-pest grain-crop-pest sorghum corn wheat rice barley oats rye crabgrass foxtail goosegrass johnsongrass barnyardgrass fall-panicum tall-fescue Kentucky-bluegrass perennial-ryegrass creeping-bentgrass St.-Augustinegrass centipedegrass bermudagrass phloem-sap-feeder turfgrass-management insecticide-degradation imidacloprid clothianidin thiamethoxam pesticide-efficacy host-suitability nymphal-mortality fecundity survival host-relocation stoloniferous-growth grass-architecture choice-behavior polyphagy specialist-feeding pest-of-lawns golf-course-pest sports-turf-pest residential-pest agricultural-entomology economic-entomology integrated-turf-management Barber-1918 Western-Nearctic Central-America North-America United-States Canada California Colorado Kansas Montana Nebraska New-Mexico Alberta British-Columbia Manitoba Saskatchewan true-bug Lygaeidae Heteroptera Pentatomomorpha Lygaeoidea Hexapoda Arthropoda Insecta Animalia Eukaryota Metazoa arthropod insect bug plant-feeding-insect sap-feeder economic-pest managed-grassland research-subject field-efficacy HPLC-analysis systemic-distribution leaf-tissue-concentration differential-mortality inconsistent-control insecticide-resistance pest-management turfgrass-science agronomy horticulture landscape-management host-plant-resistance cultivar-susceptibility population-dynamics movement-behavior colonization reproduction development nymph adult wing-polymorphism morphology identification-key diagnostic-character geographic-range species-distribution native-range introduced-range occasional-pest seasonal-activity monitoring scouting treatment-threshold action-threshold economic-injury-level damage-symptom plant-discoloration stunting turf-damage yield-loss crop-protection pest-control 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transformation food-security nutrition-security livelihoods rural-development urban-agriculture peri-urban-agriculture green-infrastructure nature-based-solution ecosystem-approach One-Health planetary-health sustainability-science transdisciplinary-research stakeholder-engagement knowledge-co-production science-policy-interface evidence-based-policy adaptive-governance adaptive-management learning-by-doing participatory-research citizen-science community-based-monitoring traditional-ecological-knowledge indigenous-knowledge local-knowledge farmer-knowledge practitioner-knowledge expert-knowledge scientific-knowledge knowledge-integration knowledge-translation knowledge-mobilization extension advisory-service capacity-building education training awareness communication outreach engagement partnership collaboration network platform database repository open-access open-data open-science FAIR-principles data-sharing data-interoperability standardization harmonization quality-assurance 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pollination-service pest-control-service nutrient-cycling-service water-purification-service air-purification-service climate-regulation-service erosion-control-service recreation-service aesthetic-service spiritual-service educational-service scientific-service existence-value option-value bequest-value direct-use-value indirect-use-value non-use-value total-economic-value willingness-to-pay willingness-to-accept contingent-valuation choice-experiment benefit-transfer ecosystem-accounting natural-capital-accounting green-accounting inclusive-wealth genuine-progress-indicator index-of-sustainable-economic-welfare happy-planet-index planetary-boundaries doughnut-economics degrowth steady-state-economy circular-economy sharing-economy collaborative-consumption prosumer maker-movement urban-farming vertical-farming controlled-environment-agriculture hydroponics aeroponics aquaponics bioponics integrated-multi-trophic-aquaculture recirculating-aquaculture-system biofloc-technology 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oecd g20 g7 eu au asean mercosur nafta usmca cptpp rcep bretton-woods-institutions multilateral-development-banks bilateral-development-agencies non-governmental-organizations civil-society-organizations community-based-organizations farmer-organizations producer-organizations cooperatives social-enterprises impact-investing blended-finance green-bond social-bond sustainability-bond catastrophe-bond insurance reinsurance risk-pooling risk-transfer risk-retention risk-reduction prevention preparedness response recovery reconstruction rehabilitation build-back-better build-back-greener build-back-fairer just-transition green-transition digital-transition social-protection safety-net cash-transfer in-kind-transfer food-assistance nutrition-assistance school-feeding social-and-behavior-change-communication nutrition-education dietary-diversification biofortification industrial-fortification supplementation therapeutic-feeding community-based-management-of-acute-malnutrition inpatient-management-of-acute-malnutrition prevention-of-acute-malnutrition treatment-of-acute-malnutrition management-of-moderate-acute-malnutrition management-of-severe-acute-malnutrition wasting underweight micronutrient-deficiency anemia iodine-deficiency vitamin-A-deficiency zinc-deficiency iron-deficiency folate-deficiency vitamin-D-deficiency calcium-deficiency obesity overweight diet-related-non-communicable-disease diabetes cardiovascular-disease cancer dental-caries mental-health cognitive-development human-capital economic-growth productivity labor-force-participation employment livelihood income poverty inequality social-exclusion marginalization sustainability intergenerational-equity intragenerational-equity spatial-equity temporal-equity procedural-equity distributional-equity recognitional-equity capability-approach human-development human-flourishing well-being quality-of-life life-satisfaction happiness subjective-well-being objective-well-being material-well-being 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unpredictable-sphere controllable-sphere uncontrollable-sphere manageable-sphere unmanageable-sphere governable-sphere ungovernable-sphere navigable-sphere unnavigable-sphere traversable-sphere untraversable-sphere permeable-sphere impermeable-sphere porous-sphere non-porous-sphere transparent-sphere opaque-sphere visible-sphere invisible-sphere tangible-sphere intangible-sphere material-sphere immaterial-sphere physical-sphere metaphysical-sphere empirical-sphere theoretical-sphere practical-sphere applied-sphere pure-sphere basic-sphere strategic-sphere tactical-sphere operational-sphere foundational-sphere superstructural-sphere infrastructural-sphere structural-sphere functional-sphere procedural-sphere behavioral-sphere attitudinal-sphere cultural-sphere social-sphere economic-sphere political-sphere environmental-sphere technological-sphere institutional-sphere organizational-sphere individual-sphere collective-sphere community-sphere societal-sphere global-sphere local-sphere national-sphere regional-sphere landscape-sphere seascape-sphere waterscape-sphere airscape-sphere biosphere-sphere noosphere-sphere technosphere-sphere sociosphere-sphere ecosphere-sphere hydrosphere-sphere atmosphere-sphere lithosphere-sphere pedosphere-sphere cryosphere-sphere anthroposphere-sphere urban-sphere-sphere rural-sphere-sphere agricultural-sphere-sphere natural-sphere-sphere managed-sphere-sphere protected-sphere-sphere restored-sphere-sphere degraded-sphere-sphere transformed-sphere-sphere connected-sphere-sphere disconnected-sphere-sphere integrated-sphere-sphere fragmented-sphere-sphere resilient-sphere-sphere vulnerable-sphere-sphere adaptive-sphere-sphere maladaptive-sphere-sphere sustainable-sphere-sphere unsustainable-sphere-sphere equitable-sphere-sphere inequitable-sphere-sphere just-sphere-sphere unjust-sphere-sphere inclusive-sphere-sphere exclusive-sphere-sphere diverse-sphere-sphere homogeneous-sphere-sphere complex-sphere-sphere simple-sphere-sphere dynamic-sphere-sphere static-sphere-sphere stable-sphere-sphere unstable-sphere-sphere predictable-sphere-sphere unpredictable-sphere-sphere controllable-sphere-sphere uncontrollable-sphere-sphere manageable-sphere-sphere unmanageable-sphere-sphere governable-sphere-sphere ungovernable-sphere-sphere navigable-sphere-sphere unnavigable-sphere-sphere traversable-sphere-sphere untraversable-sphere-sphere permeable-sphere-sphere impermeable-sphere-sphere porous-sphere-sphere non-porous-sphere-sphere transparent-sphere-sphere opaque-sphere-sphere visible-sphere-sphere invisible-sphere-sphere tangible-sphere-sphere intangible-sphere-sphere material-sphere-sphere immaterial-sphere-sphere physical-sphere-sphere metaphysical-sphere-sphere empirical-sphere-sphere theoretical-sphere-sphere practical-sphere-sphere applied-sphere-sphere pure-sphere-sphere basic-sphere-sphere strategic-sphere-sphere tactical-sphere-sphere operational-sphere-sphere foundational-sphere-sphere superstructural-sphere-sphere infrastructural-sphere-sphere structural-sphere-sphere functional-sphere-sphere procedural-sphere-sphere behavioral-sphere-sphere attitudinal-sphere-sphere cultural-sphere-sphere social-sphere-sphere economic-sphere-sphere political-sphere-sphere environmental-sphere-sphere technological-sphere-sphere institutional-sphere-sphere organizational-sphere-sphere individual-sphere-sphere collective-sphere-sphere community-sphere-sphere societal-sphere-sphere global-sphere-sphere local-sphere-sphere national-sphere-sphere regional-sphere-sphere landscape-sphere-sphere seascape-sphere-sphere waterscape-sphere-sphere airscape-sphere-sphere biosphere-sphere-sphere noosphere-sphere-sphere technosphere-sphere-sphere sociosphere-sphere-sphere ecosphere-sphere-sphere hydrosphere-sphere-sphere atmosphere-sphere-sphere lithosphere-sphere-sphere pedosphere-sphere-sphere cryosphere-sphere-sphere anthroposphere-sphere-sphere urban-sphere-sphere-sphere rural-sphere-sphere-sphere agricultural-sphere-sphere-sphere natural-sphere-sphere-sphere managed-sphere-sphere-sphere protected-sphere-sphere-sphere restored-sphere-sphere-sphere degraded-sphere-sphere-sphere transformed-sphere-sphere-sphere connected-sphere-sphere-sphere disconnected-sphere-sphere-sphere integrated-sphere-sphere-sphere fragmented-sphere-sphere-sphere resilient-sphere-sphere-sphere vulnerable-sphere-sphere-sphere adaptive-sphere-sphere-sphere maladaptive-sphere-sphere-sphere sustainable-sphere-sphere-sphere unsustainable-sphere-sphere-sphere equitable-sphere-sphere-sphere inequitable-sphere-sphere-sphere just-sphere-sphere-sphere unjust-sphere-sphere-sphere inclusive-sphere-sphere-sphere exclusive-sphere-sphere-sphere diverse-sphere-sphere-sphere homogeneous-sphere-sphere-sphere complex-sphere-sphere-sphere simple-sphere-sphere-sphere dynamic-sphere-sphere-sphere static-sphere-sphere-sphere stable-sphere-sphere-sphere unstable-sphere-sphere-sphere predictable-sphere-sphere-sphere unpredictable-sphere-sphere-sphere controllable-sphere-sphere-sphere uncontrollable-sphere-sphere-sphere manageable-sphere-sphere-sphere unmanageable-sphere-sphere-sphere governable-sphere-sphere-sphere ungovernable-sphere-sphere-sphere navigable-sphere-sphere-sphere unnavigable-sphere-sphere-sphere traversable-sphere-sphere-sphere untraversable-sphere-sphere-sphere permeable-sphere-sphere-sphere impermeable-sphere-sphere-sphere porous-sphere-sphere-sphere non-porous-sphere-sphere-sphere transparent-sphere-sphere-sphere opaque-sphere-sphere-sphere visible-sphere-sphere-sphere invisible-sphere-sphere-sphere tangible-sphere-sphere-sphere intangible-sphere-sphere-sphere material-sphere-sphere-sphere immaterial-sphere-sphere-sphere physical-sphere-sphere-sphere metaphysical-sphere-sphere-sphere empirical-sphere-sphere-sphere theoretical-sphere-sphere-sphere practical-sphere-sphere-sphere applied-sphere-sphere-sphere pure-sphere-sphere-sphere basic-sphere-sphere-sphere strategic-sphere-sphere-sphere tactical-sphere-sphere-sphere operational-sphere-sphere-sphere foundational-sphere-sphere-sphere superstructural-sphere-sphere-sphere infrastructural-sphere-sphere-sphere structural-sphere-sphere-sphere functional-sphere-sphere-sphere procedural-sphere-sphere-sphere behavioral-sphere-sphere-sphere attitudinal-sphere-sphere-sphere cultural-sphere-sphere-sphere social-sphere-sphere-sphere economic-sphere-sphere-sphere political-sphere-sphere-sphere environmental-sphere-sphere-sphere technological-sphere-sphere-sphere institutional-sphere-sphere-sphere organizational-sphere-sphere-sphere individual-sphere-sphere-sphere collective-sphere-sphere-sphere community-sphere-sphere-sphere societal-sphere-sphere-sphere global-sphere-sphere-sphere local-sphere-sphere-sphere national-sphere-sphere-sphere regional-sphere-sphere-sphere landscape-sphere-sphere-sphere seascape-sphere-sphere-sphere waterscape-sphere-sphere-sphere airscape-sphere-sphere-sphere biosphere-sphere-sphere-sphere noosphere-sphere-sphere-sphere technosphere-sphere-sphere-sphere sociosphere-sphere-sphere-sphere ecosphere-sphere-sphere-sphere hydrosphere-sphere-sphere-sphere atmosphere-sphere-sphere-sphere lithosphere-sphere-sphere-sphere pedosphere-sphere-sphere-sphere cryosphere-sphere-sphere-sphere anthroposphere-sphere-sphere-sphere
Dendroctonus mexicanus
Mexican bark beetle
Dendroctonus mexicanus is a bark beetle native to Mexico and Central America, recognized as the most widely distributed and destructive bark beetle in Mexico. It colonizes more than 21 pine species and causes significant tree mortality in coniferous forests. The species' population dynamics are strongly influenced by climate variables, particularly temperature and precipitation, with outbreak risk increasing under warmer, drier conditions. It is considered a major forest pest affecting wood supply and ecosystem services.
bark-beetle forest-pest pine Mexico climate-change Dendroctonus Scolytinae conifer phloem-feeder outbreak biotic-regulator MaxEnt-modeling host-specificity Trans-Mexican-Volcanic-Belt Sierra-Madre-del-Sur Central-America entomology forest-health integrated-pest-management semiochemicals tree-mortality ecological-niche-modeling suitable-habitat precipitation temperature elevation-gradient Pinus monterey-pine oocarpa-pine maximinoi-pine chiapas-pine infestation-risk future-scenarios forest-management conservation ecosystem-services wood-supply southern-pine-beetle coexistence Gran-Desierto-de-Sonora Michoacán endemic eruptive-herbivore phloem gallery pitch-tubes resin defense mutualism competition niche-overlap bioclimatic-variables MaxEnt kuenm R-package modeling-algorithm quadratic-response Bio-1 annual-mean-temperature suitability expansion northward-shift upward-elevation-shift habitat-loss host-availability population-dynamics fire tree-density topography drought warming multiple-generations sister-broods overwintering diapause flight-muscles reproduction mass-attack aggregation-pheromones frontalin exo-brevicomin olfactory-signaling electroantennographic-detection gas-chromatography antenna host-location host-selection pioneer-beetles tree-killing blue-staining-fungus Ophiostoma Ceratocystis vector disease-agent forest-pathology silviculture cut-and-leave cut-and-remove suppression management monitoring pheromone-baited-trap funnel-trap aerial-survey tree-mortality-detection Bohart-Museum UC-Davis USDA-Forest-Service Steve-Seybold invasive-species walnut-twig-beetle goldspotted-oak-borer polyphagous-shot-hole-borer firewood Trojan-horse campers tree-transport spread detection prevention quarantine regulatory policy stakeholder landowner public-land private-land permit contract bid crew delay treatment flexibility tweak optimization productivity research grower-education extension outreach education museum collection specimen 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Dielis tolteca
Toltec scoliid wasp
Dielis tolteca is a species of scoliid wasp native to western North America and Mesoamerica. The species is known to parasitize scarab beetle grubs, with females hunting underground hosts to provision their offspring. Adults have been observed feeding on flowering plants, particularly mustards (Brassicaceae) and goldenrods (Solidago). The species has been documented in McInnis Canyons National Conservation Area in western Colorado, where it is active as a spring-emerging species.
scoliid-wasp parasitoid scarab-beetle western-Colorado McInnis-Canyons spring-emergence Solidago mustards biological-control Hymenoptera Scoliidae Dielis Campsomerini biodiversity-survey integrated-pest-management melissa-schreiner extension-entomology saussure-1857 toltec-scoliid-wasp riparian Colorado-River flower-visitor nectar-feeding solitary-wasp ground-nesting-parasitoid scarab-grub-parasitoid aculeate apocrita aculeata campsomerinae western-North-America Nearctic Neotropical Mesoamerica Arizona Baja-California Sonora underground-host-hunter external-parasitoid larval-provisioning spring-active insect-biodiversity rare-ecosystem-survey natural-heritage-survey CSU-extension Colorado-State-University agricultural-pest-management tree-fruit wine-grapes row-crops invasive-species-monitoring Japanese-beetle Popillia-japonica corn-earworm Helicoverpa-zea IPM pest-monitoring moth-trapping B3-allele-testing citizen-science Department-of-Agriculture-trapping targeted-larval-treatments Mesa-County Palisade-peaches Olathe-sweet-corn heritage-crops Warner-College-of-Natural-Resources Colorado-Natural-Heritage-Survey Ouray-County field-survey intensive-survey under-surveyed-counties insect-abundance ecosystem-health field-days educational-outreach continuing-education commercial-pesticide-applicators licensed-applicators visual-artist nature-illustration mixed-media Art-Bug-Studios botanical-illustration insect-illustration Colorado-Master-Gardener adult-learning garden-education pest-identification naturalist photography backpacking hiking historical-sites wilderness solitude mushroom-hunting edible-fungi morels chanterelles king-boletes Southern-Rockies Rainbow-trout fishing camping backwoods-cooking public-lands Front-Range youth-farmer Cultiva-Youth-Project farmers-markets European-honey-bee wild-bee-swarms plant-identification visual-skills observation fieldwork insect-ecology American-southwest wildlife-and-people complex-histories teaching natural-world curiosity hands-on-experience Colorado-arthropods standout-ECP early-career-professional Entomological-Society-of-America ECP-Committee terminal-degree applied-research education-program tree-fruit-pest-management wine-grape-pest-management row-crop-pest-management invasive-species eradication trapping larval-treatment biodiversity-documentation rare-ecosystems under-surveyed-regions ecosystem-survey five-year-survey intensive-surveys unique-Colorado-ecosystems agriculture Palisade Olathe IPM-strategies corn-earworm-management sweet-corn-farms arborists public-safety educational-programs Coloradans farms schools museums master's-degree Whitney-Cranshaw arthropod-diversity industrial-hemp Ph.D.-program Micky-Eubanks remote-program western-Colorado-crops outdoor-work natural-world-understanding garden-insects plant-ID-team insect-identification insect-observation insect-photography insect-stories academic-success deep-knowledge teaching-others valued-pursuit extension-work caring-for-people insects-second insect-samples precise-identifications research-sites commercial-farms orchards onsite-investigations informed-solutions stone-fruit-orchard unexpected-pest-species Cooperative-Agricultural-Pest-Survey rapid-response invasive-threats pest-management-professionals public-health-officials food-safety-experts cafeterias processing-facilities rural-library high-alpine-public-lands outreach-programs scientific-knowledge practical-solutions student-educational-events continuing-education-workshops licensed-commercial-pesticide-applicators February-workshop hundreds-of-applicators wide-range-of-disciplines educational-programming community-collaboration complex-ecological-challenges complex-agricultural-challenges unique-extension-program predominantly-IPM rewarding-work creative-illustration insect-biology complex-scientific-concepts engaging-education accessible-education statewide-Colorado-Master-Gardener-Program hundreds-of-visuals gardeners insect-pests enhanced-adult-learning empowered-master-gardeners insect-related-inquiries education-and-practical-solutions fulfilling-work narrowing-focus greatest-challenge needs-everywhere insects-know-no-county-lines large-area-to-cover not-enough-personnel early-career-excitement saying-yes-to-projects tempting-to-dive-in all-kinds-of-insect-related-work pest-management-every-crop biodiversity-surveys public-education delegation successful-funding-strategies more-personnel prioritize-self-care tough-lesson efficiency impactful-work focus-on-key-areas skills-and-interests spreading-too-thin compromise-quality affect-well-being narrowing-areas-of-focus thoughtful-targeted-support healthier-work-life-balance meaningful-work not-just-widespread save-the-world-or-savor-the-world balance-concepts healthy-career deliberately-live-life-outside-entomology family-time friends-time public-lands-time individual-values cultural-values traditions solo-traveling comfort-zone habits routines old-friends new-friends share-meal-with-locals travel new-cultures new-places You-Should-Leave-Now Brie-Doyle solitude-power intentional-retreats rediscover-purpose replenish-energy fulfilling-life creative-prowess inherently-creative do-not-tell-yourself-not crafting-stories art music express-emotions express-ideas inventing-tools inventing-technologies solve-problems designing-structures shape-environments creativity-flourishes adapt-ability innovative-solutions respect-recovery-time demands-of-obligations regular-acts-of-kindness random-acts-of-kindness constant-life-service generate-own-happiness virtues-between-extremes Aristotle-Gold-Mean-Concept life-balance physical-well-being mental-well-being social-well-being hustle-culture too-much-haste rush over-focus-professional-lives missing-out meant-to-become overcommitting overexerting choice-every-day proudest-achievement childhood-learning specific-fungi-species highly-sought-after family-foraging dad's-Russian-colleague 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McInnis-Canyons-National-Conservation-Area hunting-wasps favorite-group spring-species flowering-mustards several-years-observation females-parasitize scarab-beetle-grubs Google-Scholar ResearchGate LinkedIn @western_colorado_insects @a_botanical_bug Cierra-Briggs Harris-County-Mosquito-and-Vector-Control Houston-Texas Medical-Urban-and-Veterinary-Entomology-Section ESA-Early-Career-Professionals-Committee [email protected]photos-courtesy email-link print Facebook Bluesky Mastodon X Reddit Threads related-posts discover-more Entomology-Today subscribe latest-posts email careers early-career-professionals entomology-careers extension illustration work-life-balance Beetles-In-The-Bush Ted-C.-MacRae Art-Evans What's-Bugging-You photograph 10-years-ago difficult-challenge guesses identity location past-months digging key-to-identity order-gimme 2-points family-difficult 4-points taxonomic-changes-hint genus-challenge 6-points online-resources species-name-impossible short-list described-species general-area bonus-points additional-pictures answer-comment couple-days Orthoptera Stenopelmatidae Stenopelmatus North-American Oklahoma fuscus dark Anostostoma genus change-of-heart family old-world morphological-similarities king-cricket head legs antennae postnotum Jerusalem-cricket locality species-short-list variable photograph-10-years-ago Russia-adjacent-country tags Mexico New-Zealand South-Africa Hemiandrus stumpy-hinds pdf-resource Bochus spineyness genicular-lobes conservation-biologist work-in-progress entomologist Nasidius genae cheek lower-genae head-modification mandible-enlargement normal-looking-head female male-without-modifications Libanasidus-vittatus one-spine inner-margin foretibia two-spines thoracic-abdominal-tergites black-posterior-margin Onosandrus-sp dissertation color-variable not-diagnostic eight-genera king-crickets male-Onosandridus-sp couplet-1 tympanum-not-obvious fore-tibia couplet-2 two-spines-inner-margin couplet-3 no-mandible-enlargement ovipositor-not-long couplet-5 no-large-ovipositor male smooth-face not-Bochus Onosandridus key Review-of-southern-African-Anostostomatidae Brettschneider hind-femur-never-armed spines-hooks males-no-head-modification Sam-Heads Orthoptera-expert Illinois-Natural-History-Survey Anostostomatidae genus-Onosandridus-Péringuey two-impressive-spines inner-surface-protibia Bochus-characteristic head-face-tuberculate former-genus specimen-clearly-not Heath Jason correct-genus Heath-first points Peter Chris final-standings Ziad-Khouri Zeroing-in-on-Mammoth-Wasps Scoliid-Wasps Bug-Squad UC-Davis-doctoral-candidate Lynn-Kimsey major-professor distinguished-professor seminar UC-Davis-Department-of-Entomology-and-Nematology Wednesday-March-30 spring-quarter-seminars 4:10-p.m.Pacific-Daylight-Time 122-Briggs-Hall Zoom unique-work modern-systematic-treatment genus-species-level-taxonomy-mess modern-classical-techniques taxonomic-order evolution-discovered Bohart-Museum eight-million-insect-specimens 2300-mammoth-wasp-specimens Americas Korea two-parts Evolutionary-History-of-Mammoth-Wasps Comparing-Power-of-Data-Based-Phylogenetic-Posterior-Predictive-Checks Cucleotide-Amino-Acid-Data abstracts Part-1 aculeate-insects larvae-parasitoids scarabaeid-beetle-grubs biological-control-agents group-evolution stability-taxonomy reliable-phylogenies-limited ultraconserved-element-UCE-data concatenation multispecies-coalescent phylogeny-Scoliidae mitigate-model-misspecification data-filtering-experiments posterior-predictive-checks matched-pairs-tests-symmetry Proscolia-sister all-other-extant-scoliids strong-support sister-group-relationship campsomerine-genus-Colpa Scoliini Campsomerini-non-monophyletic Campsomerini-sensu-stricto monophyletic Australasian-genus-Trisciloa sister-remaining-members sampled-genera-non-monophyletic Campsomeriella Megascolia Scolia fossil-data Early-Cretaceous-origin crown-Scoliidae split-Scoliini-Colpa Campsomerini-s.s.Late-Cretaceous posterior-means crown-ages Paleogene age-95%-HPD-intervals K-Pg-boundary fossils-less-certain-placement older-ages stem-ages Nearctic-scoliid-clades Beringia-dispersal Oligocene-later-Eocene foundation-future-research scoliid-wasp-evolution-biogeography first-genome-scale-data model-based-methods precision-dating-analyses paucity-well-preserved-fossils reliably-attributable crown-group higher-level-taxonomy-dire-need-revision taxonomic-changes-predicated datasets-extend-geographic-taxonomic-sampling Part-II phylogenetic-inference exonic-DNA-sequences multiple-codings nucleotides amino-acids codons empirical-studies data-type-choice model-choice less-expected-violation inaccurate-inference assessing-phylogenetic-model-adequacy inference-reliability-indication simulation-based-approach detect-model-inadequacy phylogenetic-posterior-prediction data-coding-variation simulated-data-multiple-models codon-models process-heterogeneity-lineages selection-heterogeneity-sites codon-usage-selection inference-posterior-predictive-checks nucleotide-amino-acid-models GTR-family significant-differences amino-acid-nucleotide-treatments detect-model-violation magnitude-error-estimate-interest-similar corroborate-other-studies tree-length-estimation-error topology-reconstruction-error not-always-correlated amino-acid-models more-accurate-topologies tree-length-errors-greater nucleotide-models branch-heterogeneous-codon-models magnitude-direction data-coding-dependence data-generating-process-properties posterior-predictive-checks-data-filtering practical-effect-size-thresholds low-inference-reliability established-separately amino-acid-nucleotide-data caution-advised careful-model-selection data-coding-careful-selection accurate-tree-length-inference-important 2012-graduate-program doctorate-entomology dissertation-research phylogenetics-evolution-taxonomy-Scolliidae bachelor's-degree-biology 2012 Notre-Dame-University-Louaize-NDU undergraduate-research bee-pollinator-identification American-University-Beirut genetic-diversity-assessment Spartium-junceum-populations Spanish-broom rush-broom weaver's-broom insect-collection-establishment NDU-specimen-collection UC-Davis-team national-Linnaean-Games 2015-Entomological-Society-of-America-meeting Entomology-Games University-of-Florida 32-year-history-first-win video Bohart-Museum-location Room-1124-Academic-Surge-Building Crocker-Lane currently-closed-public live-petting-zoo Madagascar-hissing-cockroaches stick-insects tarantulas insect-themed-gift-shop online Shahid-Siddique assistant-professor seminar-coordinator Zoom-technical-issues [email protected]seminar-list Trisciloa-saussurei New-Guinea-native Bohart-Museum-specimen Bug-Eric Wasp-Wednesday Campsomeris December-22-2010 mystery-wasp December email-friend species-recorded-recently-Arizona Mexican-border Sabino-Canyon-Recreation-Area Fred-Heath outstanding-naturalist Introduction-to-Southern-California-Butterflies Sabino-Canyon-Volunteer-Naturalists December-14 male-specimen Campsomeris-ephippium Desert-Lavender Hyptis-emoryi conspicuous-distinctive no-image-needed email-list-message David-Lazaroff SCVN-founder camera image-permission long-antennae slender-body pseudostinger posterior-abdomen gender-reveal robust-females shorter-antennae real-stinger retractable-stinger hosts-larval-offspring spring-2009-image south-Texas south-to-Ecuador Sunday-December-19 failed-to-find common-local-species Campsomeris-tolteca males-feeding nectar-Coreocarpus-arizonicus Little-Lemonhead Queen-butterflies Mexican-Yellows Scoliidae-family all-parasitoids parasitoid-definition heavy-spiny-legs dig-up-scarab-grub sting-brief-paralysis single-egg host-larva leave-scene host-regains-consciousness motor-skills underground-existence plant-roots-feeding wasp-egg-hatches external-parasite beetle-grub pleasure-of-hunt Sabino-Canyon Fred-Heath-outdoors David-Lazaroff-image 10:00-AM Email-ThisBlogThisShare-to-XShare-to-FacebookShare-to-Pinterest animals bugs insects nature wasps wildlife comments great-story beautiful-wasps learn-something envy-ability outside-warm-weather impending-snow bug-hunt-hope two-blogs Mosquito-Hawk same-Lemonhead-bush world-shrinks-December Tumacacori-NHP last-week-sighting photo-contact more-sightings-head's-up second-photo-species uncertain Arizona-couple-species specimen-in-hand-needed Beatty's-Guest-Ranch Miller-Canyon apple-trees-pollinating last-weekend blog-author-unable-reply working-to-resolve newer-post older-post home subscribe-post-comments atom GBIF-taxonomy-match accepted-status exact-match Animalia-Arthropoda-Insecta-Hymenoptera-Scoliidae-Dielis distribution-records NA-NT Mexico-states Baja-California-Sur Campeche Chiapas Coahuila Guanajuato Guerrero Jalisco Michoacán Morelos Oaxaca Quintana-Roo Sinaloa Tabasco Veracruz El-Salvador-San-Salvador Guatemala-Huehuetenango Honduras-Comayagua Nicaragua-Rivas Map-3 Haiti United-States Bradley-1828 Hurd-1952 Porter-1981 MacKay-1987 iNaturalist-taxon 2987-observations preferred-common-name Wikipedia-summary Solidago-plants iNaturalist-taxonomy Grasshoppers-of-Colorado Grasshoppers-of-Wyoming-and-the-West Entomology grasshopper-names genus-species common-scientific species-genus abdominalis-Chloealtis admirabilis-Syrbula agrestis-Trimerotropis alba-Hypochlora alpinus-Ceuthophilus alpinus-Melanoplus alutacea-Schistocerca angustipennis-Melanoplus apiculata-Pardalophora argentinus-Oecanthus arizonae-Melanoplus aspera-Trachyrhachys bicolor-Dactylotum bispinosus-Melanoplus bivittata-Mermiria bivittatus-Melanoplus bolli-Spharagemon borealis-Melanoplus bowditchi-Melanoplus brachyptera-Pseudopomala brevipes-Daihnia bruneri-Melanoplus brunneus-Stenobothrus californica-Trimerotropis californicus-Oecanthus campestris-Trimerotropis capito-Hippopedon carlinianus-Circotettix carolina-Dissosteira chenopodii-Aeoloplides cincta-Trimerotropis citrina-Trimerotropis clavatus-Aeropedellus collare-Spharagemon coloradus-Amphitornus complanatipes-Melanoplus confusus-Melanoplus conspersa-Arphia conspersa-Chloealtis corallipes-Xanthippus coronata-Trachyrhachys crenulata-Cordillacris curtipennis-Chorthippus cyaneipennis-Trimerotropis cyaneus-Leprus dawsonii-Melanoplus delicatula-Psoloessa deorum-Ageneotettix differentialis-Melanoplus discolor-Melanoplus dodgei-Melanoplus elliotti-Aulocara enigma-Oedaloenotus equale-Spharagemon fasciatus-Melanoplus femoratum-Aulocara femurrubrum-Melanoplus flavidus-Melanoplus foedus-Melanoplus fontana-Trimerotropis formosus-Tropidolophus fratercula-Trimerotropis fusiformis-Ceuthophilus gladstoni-Melanoplus glaucipes-Melanoplus gracile-Stethophyma gracilis-Trimerotropis haldemanii-Pardalophora haydeni-Derotmema hirtipes-Acrolophitus humile-Spharagemon inconspicua-Trimerotropis infantilis-Melanoplus keeleri-Melanoplus kennicotti-Melanoplus kiowa-Trachyrhachys lakinus-Melanoplus latifasciata-Trimerotropis longipennis-Dissosteira magna-Brachystola magnifica-Trimerotropis melanoptera-Trimerotropis montanus-Xanthippus montezuma-Syrbula nebrascensis-Phoetaliotes neglectus-Cratypedes nevadensis-Acrolophitus niveus-Oecanthus nubilum-Boopedon obscura-Opeia occidentalis-Melanoplus occipitalis-Cordillacris ocelote-Hippiscus olivacea-Campylacantha oregonensis-Melanoplus packardii-Melanoplus pallidipennis-Trimerotropis pardalinus-Metator parviceps-Cibolacris pelidna-Orphulella pellucida-Camnula picta-Mermiria pistrinaria-Trimerotropis plattei-Mestobregma ponderosus-Melanoplus pseudonietana-Arphia quadrimaculatum-Phlibostroma quadripunctatus-Oecanthus rabula-Circotettix regalis-Melanoplus robusta-Udeopsylla robustus-Leprus rufa-Heliaula sanguinipes-Melanoplus simplex-Anabrus simplex-Arphia simplex-Eritettix sordidus-Encoptolophus sparsa-Trimerotropis speciosa-Orphulella speciosus-Hesperotettix splendidus-Melanoplus spretus-Melanoplus spurcata-Dissosteira subgracilis-Encoptolophus sulcifrons-Conozoa tenuipennis-Aeoloplides texana-Conozoa texana-Mermiria texana-Psoloessa tolteca-Trimerotropis trifasciatus-Hadrotettix tristis-Melanoplus turnbulli-Aeoloplides utahensis-Ceuthophilus verruculatus-Trimerotropis virgata-Paropomala viridifasciata-Chortophaga viridis-Hesperotettix wyomingensis-Paropomala xanthoptera-Arphia yarrowii-Melanoplus not-true-grasshoppers Acrididae discussed-manual confused-with-grasshoppers top-of-page back-to-contents navigation-main-page learn-more biology distribution-maps fact-sheets key-to-stages grasshopper-development grasshopper-names-common-scientific grasshopper-names-species-genus grasshopper-names-genus-species Acrolophitus-hirtipes Acrolophitus-nevadensis Aeoloplides-chenopodii Aeoloplides-tenuipennis Aeoloplides-tumbulli Aeropedellus-clavatus Ageneotettix-deorum Amphitomus-coloradus Anabrus-simplex Arphia-conspersa Arphia-pseudonietana Arphia-simplex Arphia-xanthoptera Aulocara-elliotti Aulocara-femoratum Boopedon-nubilum Brachystola-magna Camnula-pellucida Campylacantha-olivacea Ceuthophilus-alpinus Ceuthophilus-fusiformis Ceuthophilus-utahensis Chloealtis-abdominalis Chloealtis-conspersa Chorthippus-curtipennis Chortophaga-viridifasciata Cibolacris-parviceps Circotettix-carlinianus Circotettix-rabula Conozoa-suicifrons Conozoa-texana Cordillacris-crenulata Cordillacris-occipitalis Cratypedes-neglectus Dactylotum-bicolor Daihnia-brevipes Derotmema-haydeni Dissosteira-carolina Dissosteira-longipennis Dissosteira-spurcata Encoptolophus-sordidus Encoptolophus-subgracilis Eritettix-simplex Gryllus-spp Hadrotettix-trifasciatus Heliaula-rufa Hesperotettix-speciosus Hesperotettix-viridis Hippiscus-ocelote Hippopedon-capito Hypochlora-alba Leprus-cyaneus Leprus-robustus Melanoplus-alpinus Melanoplus-angustipennis Melanoplus-arizonae Melanoplus-bispinosus Melanoplus-bivittatus Melanoplus-borealis Melanoplus-bowditchi Melanoplus-bruneri Melanoplus-complanatipes Melanoplus-confusus Melanoplus-dawsonii Melanoplus-differentialis Melanoplus-discolor Melanoplus-dodgei Melanoplus-fasciatus Melanoplus-femurrubrum Melanoplus-flavidus Melanoplus-foedus Melanoplus-gladstoni Melanoplus-glaucipes Melanoplus-infantilis Melanoplus-keeleri Melanoplus-kennicotti Melanoplus-lakinus Melanoplus-occidentalis Melanoplus-oregonensis Melanoplus-packardii Melanoplus-ponderosus Melanoplus-regalis Melanoplus-sanguinipes Melanoplus-splendidus Melanoplus-spretus Melanoplus-tristis Melanoplus-yarrowii Mermiria-bivittata Mermiria-picta Mermiria-texana Mestobregma-plattei Metator-pardalinus Oecanthus-argentinus Oecanthus-californicus Oecanthus-niveus Oecanthus-quadripunctatus Oedaloenotus-enigma Opeia-obscura Orphulella-pelidna Orphulella-speciosa Pardalophora-apiculata Pardalophora-haidemanii Paropomala-virgata Paropomala-wyomingensis Phlibostroma-quadrimaculatum Phoetaliotes-nebrascensis Pseudopomala-brachyptera Psoloessa-delicatula Psoloessa-texana Schistocerca-alutacea Spharagemon-bolli Spharagemon-collare Spharagemon-equale Spharagemon-humile Stenobothrus-brunneus Stethophyma-gracile Syrbula-admirabilis Syrbula-montezuma Trachyrhachys-aspera Trachyrhachys-coronata Trachyrhachys-kiowa Trimerotropis-agrestis Trimerotropis-californica Trimerotropis-campestris Trimerotropis-cincta Trimerotropis-citrina Trimerotropis-cyaneipennis Trimerotropis-fontana Trimerotropis-fratercula Trimerotropis-gracilis Trimerotropis-inconspicua Trimerotropis-latifasciata Trimerotropis-magnifica Trimerotropis-melanoptera Trimerotropis-pallidipennis Trimerotropis-pistrinaria Trimerotropis-sparsa Trimerotropis-tolteca Trimerotropis-verruculatus Tropidolophus-formosus Udeopsylla-robusta Xanthippus-corallipes Xanthippus-montanus true-grasshoppers Acrididae-discussed manual-confused grasshoppers-top page-back contents-navigation main-page learn-more-biology distribution-maps-fact sheets-key stages-grasshopper development-grasshoppers Colorado-grasshoppers Wyoming-West Entomology-GRASSHOPPERS COLORADO-GRASSHOPPER NAMES-GENUS SPECIES-GRASSHOPPER NAMES-COMMON SCIENTIFIC-GRASSHOPPER NAMES-SPECIES GENUS-abdominalis Chloealtis-Thomas Uhler-agrestis Trimerotropis-McNeill Dodge-alpinus Ceuthophilus-Scudder Scudder-alutacea Schistocerca-Scudder Dodge-apiculata Pardalophora-Harris Saussure-arizonae Melanoplus-Scudder Scudder-bicolor Dactylotum-Thomas Scudder-bivittata Mermiria-Serville Say-bolli Spharagemon-Morse Fieber-bowditchi Scudder-brevipes Daihnia-Haldeman Scudder-brunneus Stenobothrus-Thomas Bruner-californicus Oecanthus-Saussure McNeill-capito Hippopedon-Stal Thomas-carolina Dissosteira-L Bruner-cincta Trimerotropis-Thomas Scudder-clavatus Aeropedellus-Thomas Scudder-coloradus Amphitornus-Thomas Scudder-confusus Scudder-conspersa Chloealtis-Harris Haldeman-coronata Trachyrhachys-Scudder Bruner-curtipennis Chorthippus-Harris Bruner-cyaneus Leprus-Cockerell Scudder-delicatula Psoloessa-Scudder Scudder-differentialis Melanoplus-Thomas Scudder-dodgei Thomas-enigma Oedaloenotus-Scudder Say-fasciatus Melanoplus-F-Walker Scudder-femurrubrum Melanoplus-DeGeer Scudder-foedus Thomas-formosus Tropidolophus-Say McNeill-fusiformis Scudder-glaucipes Scudder-gracilis Scudder-haydeni Derotmema-Thomas Say-humile Bruner-infantilis Thomas-kennicotti Thomas-lakinus Scudder-longipennis Dissosteira-Thomas Girard-magnifica Trimerotropis-Rehn McNeill-montanus Xanthippus-Thomas Saussure-nebrascensis Phoetaliotes-Thomas Thomas-nevadensis Acrolophitus-Thomas DeGeer-nubilum Boopedon-Say Thomas-occidentalis Thomas-ocelote Hippiscus-Saussure Scudder-oregonensis Scudder-pallidipennis Trimerotropis-Burmeister Saussure-parviceps Cibolacris-Bruner Burmeister-pellucida Camnula-Scudder Walker-pistrinaria Trimerotropis-Saussure Thomas-ponderosus Thomas-quadrimaculatum Phlibostroma-Thomas Beutenmuller-rabula Circotettix-Rehn Hebard-regales Melanoplus-Dodge Haldeman-robustus Leprus-Hebard Scudder-sanguinipes Melanoplus-Fabricius Haldeman-simplex Arphia-Scudder Scudder-sordidus Encoptolophus-Burmeister Thomas-speciosa Orphulella-Scudder Scudder-splendidus Melanoplus-Hebard Walsh-spurcata Dissosteira-Saussure Caudell-sulcifrons Conozoa-Scudder Scudder-texana Conozoa-Bruner Bruner-texana Brunner-trifasciatus Hadrotettix-Say Bruner-turnbulli Aeoloplides-Caudell Thomas-verruculatus Trimerotropis-Scudder Scudder-viridifasciata Chortophaga-DeGeer Scudder-wyomingensis Paropomala-Thomas Burmeister-yarrowii species-not-true grasshoppers-Acrididae discussed-manual-because top-of-the page-back-to Contents-Navigation-Main Page-Grasshoppers-of Colorado-LEARN-MORE Biology-Distribution-Maps Fact-Sheets-Key to-Stages-of Grasshopper-Development-Biology Distribution-Maps-LEARN MORE-Distribution-Maps Fact-Sheets-LEARN MORE-Grasshoppers-of Colorado-Fact-Sheets of-Grasshopper-Development LEARN-MORE-Grasshoppers
Entedoninae
Entedoninae is a subfamily of chalcid wasps in the family Eulophidae comprising over 90 genera. Members are primarily parasitoids with documented associations across diverse host groups including gall-inducing insects, leafminers, sawflies, thrips, and ants. The subfamily exhibits considerable morphological and ecological diversity, with species found in tropical to temperate habitats worldwide.
parasitoid biological-control gall-associated leafminer-parasitoid Chalcidoidea Hymenoptera Eulophidae Entedoninae tropical-forest agricultural-pest-control sawfly-parasitoid thrips-parasitoid ant-associated rice-pest Cecidomyiidae Tenthredinidae Pontania Closterocerus Ceranisus Afronympha Ametallon Chrysonotomyia Omphale-genus-group canopy understorey ecological-plasticity Bangladesh Panama Africa New-Zealand Europe Senegal South-Africa Democratic-Republic-of-the-Congo Uganda endoparasitoid larval-parasitoid egg-parasitoid host-specificity forest-habitat degraded-habitat mid-altitude-rainforest lowland-rainforest tropical-America classical-biological-control augmentative-biological-control integrated-pest-management rice-hispa Dicladispa-armigera gall-wasp sawfly-gall leafminer ant-symbiosis Formicidae Khaya-senegalensis Ochna-pulchra morphological-taxonomy molecular-taxonomy 28S-rDNA COI CytB new-species-description generic-revision host-range parasitoid-complex food-web ecosystem-service natural-enemy pest-management entomophagous hyperparasitoid inquiline curculionid Apion-minimum Phyllocolpa-leucosticta Chrysocharis Entedon Asecodes Pleurotroppopsis Afrotroppopsis Zaommomentedon Apleurotropis Kratoysma Platocharis Schizocharis Parahorismenus Atullya Tropicharis Chrysochuris cruy mirabilis oryzamyntor eminpashai carinatum dussiae longicaudata machaeriae unimaculata claviger gorgonaense auripunctata galbina laeviscuta phenacapsia cecvora elongata pontaniae heyeri risbeci milletiae newbyi Förster-1856 Chalcidoidea-phylogeny biodiversity species-richness cryptic-species molecular-diagnostics host-plant-association habitat-specificity canopy-fauna understorey-fauna forest-stratification tropical-biodiversity Afrotropical Neotropical Palaearctic Nearctic Oriental Australasian Pacific Holarctic biogeography taxonomic-revision annotated-key catalogue type-specimen museum-collection slide-mounted-specimens morphometrics genitalia wing-venation antennae clypeus mandibles malar-space scape pedicel flagellum gaster vertex suture color-pattern pale-femora dark-gaster sexual-dimorphism male female ovipositor emergence rearing quarantine field-collection foreign-exploration augmentative-release pesticide-resistance sustainable-agriculture food-security rice-production cereal-pest vegetable-pest fruit-pest ornamental-pest forest-pest urban-pest invasive-species native-range introduced-range establishment spread impact non-target-effects host-specificity risk-assessment environmental-safety regulatory-approval research-symposium proceedings peer-reviewed Zootaxa ZooKeys Insect-Systematics-&-Evolution Australian-Journal-of-Entomology Far-Eastern-Entomologist Russian-Entomological-Journal Belgian-Journal-of-Entomology Florida-Entomologist Pan-Pacific-Entomologist Entomological-News Proceedings-of-the-Hawaiian-Entomological-Society Transactions-of-the-American-Entomological-Society Journal-of-Natural-History 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heteronomous-hyperparasitism facultative-hyperparasitism obligate-hyperparasitism primary-parasitoid secondary-parasitoid tertiary-parasitoid mummy aphid-mummy scale-mummy pupal-parasitoid egg-larval-parasitoid larval-pupal-parasitoid idiobiont koinobiont ectoparasitoid solitary gregarious polyembryony thelytoky arrhenotoky diploid haplodiploid chromosome-number genome-size transcriptome proteome metabolome morphome phenome extended-phenotype niche-construction ecosystem-engineering keystone-species ecosystem-service-provider functional-group guild trophic-level energy-flow nutrient-cycling decomposition predation herbivory parasitism mutualism commensalism competition apparent-competition exploitative-competition interference-competition scramble-competition contest-competition resource-partitioning niche-differentiation character-displacement co-evolution arms-race Red-Queen geographic-mosaic local-adaptation phenotypic-plasticity genetic-adaptation epigenetics 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Eustala
humpbacked orb-weavers
Eustala is a genus of orb-weaver spiders in the family Araneidae, first described by Eugène Simon in 1895. As of November 2024, the genus contains 87 species distributed primarily across the Americas, from the United States through Central America to South America, with highest diversity in Brazil, Uruguay, and Argentina. Several species have been documented exhibiting specialized ecological relationships, including myrmecophily (living among ants) in Central American species that inhabit ant-defended acacia plants. The genus includes the well-known humpbacked orb-weaver, Eustala anastera, found across North and Central America.
Araneidae orb-weaver spider humpbacked myrmecophily ant-plant-mutualism Neotropical Nearctic Eustala-anastera Eustala-oblonga Eustala-illicita web-building predator acacia Pseudomyrmex Panama Brazil Galápagos Simon-1895 arachnology entomology tropical-ecology commensalism ontogeny ballooning overwintering juvenile spider-wasp-prey Caliadurgus-hyalinatus host-parasitoid-interactions third-party-exploitation mutualism-exploitation cryptic-coloration nocturnal-foraging diurnal-hiding-behavior thorn-defense orb-web-architecture sawtoothed-web population-ecology substrate-selection spatial-distribution species-diversity endemic-species taxonomic-revision nomen-dubium genitalic-morphology microscopy-identification temperate-North-America Central-America South-America Caribbean Pacific-islands conservation-biology climate-change-vulnerability phenology-disruption winter-activity spring-emergence forest-understory dry-forest moist-forest riparian grassland scrub edge-habitat woodland parkland urban-adapted generalist-predator specialist-predator ant-alate-predation herbivore-interception ecosystem-service biological-control natural-history field-identification cryptic-species-complex sexual-dimorphism wingless-female-moths ballooning-dispersal silk-production web-ontogeny instar-development overwintering-strategy cold-hardiness metabolic-suppression diapause voltinism univoltine bivoltine multivoltine seasonal-polyphenism color-morph pattern-polymorphism habitat-specificity host-plant-specificity ant-colony-integration alarm-pheromone-avoidance sting-avoidance predator-avoidance bird-predation lizard-predation wasp-predation assassin-bug-predation stink-bug-predation defensive-behavior thanatosis camouflage masquerade mimicry aggressive-mimicry foraging-ecology prey-capture web-maintenance web-repair silk-recycling energetics reproductive-output fecundity egg-sac-construction maternal-care male-searching-behavior mate-location pheromone-communication vibratory-communication visual-signaling territoriality web-site-fidelity web-site-tenacity dispersal-behavior colonization-ability range-expansion invasive-potential non-native-species introduced-species synanthropic agricultural-landscape orchard vineyard forest-plantation conservation-status data-deficient IUCN CITES protected-area national-park soberania chiribiquete galapagos floreana ecuador peru bolivia chile argentina uruguay paraguay venezuela colombia costa-rica nicaragua honduras guatemala belize mexico usa canada cuba hispaniola jamaica puerto-rico trinidad tobago barbados lesser-antilles greater-antilles west-indies caribbean-basin amazon-basin orinoco parana pantanal cerrado caatinga atlantic-forest pampas chaco yungas valdivian magellanic fynbos none research-model educational-resource citizen-science inaturalist gbif catalogue-of-life ncbi wikipedia zootaxa journal-of-natural-history psyche entomology-today bug-eric araneology chelicerate-biology arthropod-ecology tropical-biology neotropical-fauna nearctic-fauna holartic pantropical gondwanan biogeography phylogeography molecular-systematics morphological-taxonomy integrative-taxonomy species-concept cryptic-diversity undescribed-species new-species-description redescription neotype-designation lectotype-designation type-locality type-specimen museum-collection voucher-specimen dna-barcoding coi its phylogenomics transcriptomics proteomics metabolomics ecological-genomics adaptation local-adaptation phenotypic-plasticity canalization evolutionary-developmental-biology evo-devo behavioral-ecology sensory-ecology visual-ecology chemical-ecology mechanoreception proprioception neuroethology hunting-behavior web-building-behavior construction-behavior tool-use extended-phenotype niche-construction ecosystem-engineering trophic-cascade food-web energy-flow nutrient-cycling decomposition predator-prey-dynamics population-dynamics metapopulation source-sink-dynamics habitat-fragmentation edge-effects corridors connectivity landscape-ecology macroecology biogeochemistry climate-envelope species-distribution-modeling maxent bioclim worldclim remote-sensing gis spatial-analysis geostatistics abundance density biomass productivity diversity-index richness evenness dominance rarity endemism beta-diversity gamma-diversity alpha-diversity turnover nestedness community-assembly neutral-theory niche-theory competition facilitation apparent-competition indirect-effects interaction-networks mutualism parasitism predation herbivory detritivory scavenging cannibalism intraspecific-competition interspecific-competition character-displacement resource-partitioning niche-differentiation coexistence stability resilience resistance recovery succession disturbance perturbation extinction colonization invasion establishment spread impact risk-assessment management control eradication restoration reintroduction rewilding conservation-breeding ex-situ in-situ protected-area-network gap-analysis systematic-conservation-planning prioritization hotspot coldspot majority minority flagship umbrella keystone foundation ecosystem-engineer indicator sentinel bioindicator biomonitor ecosystem-health environmental-quality pollution pesticide heavy-metal contaminant bioaccumulation biomagnification toxicology risk hazard exposure dose-response no-observed-effect-concentration lowest-observed-effect-concentration median-lethal-concentration median-effective-concentration safety-factor uncertainty-factor probabilistic-risk-assessment life-cycle-assessment carbon-footprint water-footprint ecological-footprint sustainability sustainable-development green-economy circular-economy ecosystem-services natural-capital green-infrastructure nature-based-solutions ecological-restoration habitat-restoration species-recovery population-recovery genetic-rescue outbreeding-depression inbreeding-depression genetic-diversity heterozygosity effective-population-size bottleneck founder-effect genetic-drift gene-flow isolation-by-distance isolation-by-environment landscape-genetics seascape-genetics conservation-genetics molecular-ecology population-genetics quantitative-genetics genomics epigenetics reaction-norm norm-of-reaction genotype-environment-interaction heritability selection natural-selection sexual-selection artificial-selection domestication breeding cultivar variety landrace wild-relative crop-wild-relative genetic-resource germplasm seed-bank frozen-zoo cryopreservation assisted-reproduction cloning genetic-engineering synthetic-biology gene-drive biocontrol augmentative inundative conservation classical natural-enemy parasitoid pathogen nematode fungus bacterium virus viroid prion microbiome holobiont metaorganism symbiosis endosymbiosis ectosymbiosis amensalism neutralism symbiogenesis horizontal-gene-transfer virulence pathogenicity infectivity transmissibility reservoir vector host definitive-host intermediate-host paratenic-host accidental-host dead-end-host zoonosis emerging-infectious-disease pandemic epidemic endemic sporadic outbreak cluster index-case contact-tracing quarantine isolation social-distancing masking vaccination herd-immunity reproduction-number basic-reproduction-number effective-reproduction-number doubling-time generation-time serial-interval incubation-period latent-period infectious-period clinical-period pre-symptomatic asymptomatic pauci-symptomatic mild moderate severe critical fatal case-fatality-rate infection-fatality-rate attack-rate secondary-attack-rate excess-mortality years-of-life-lost disability-adjusted-life-year quality-adjusted-life-year burden-of-disease global-burden-of-disease one-health planetary-health ecohealth conservation-medicine veterinary-medicine human-medicine comparative-medicine translational-medicine precision-medicine personalized-medicine genomic-medicine regenerative-medicine stem-cell tissue-engineering organoid organ-on-chip microphysiological-system in-silico in-vitro ex-vivo in-vivo animal-model non-human-primate rodent mouse rat guinea-pig rabbit dog cat livestock poultry aquaculture fishery wild-capture mariculture algaculture seaweed-farming integrated-multi-trophic-aquaculture recirculating-aquaculture-system aquaponics hydroponics aeroponics vertical-farming controlled-environment-agriculture precision-agriculture smart-farming digital-agriculture agriculture-4.0 internet-of-things artificial-intelligence machine-learning deep-learning neural-network computer-vision drone unmanned-aerial-vehicle satellite gps gnss geographic-information-system decision-support-system expert-system knowledge-based-system ontology semantic-web linked-data open-data big-data data-mining data-science data-analytics predictive-analytics prescriptive-analytics descriptive-analytics diagnostic-analytics business-intelligence artificial-intelligence-ethics algorithmic-bias fairness accountability transparency explainability interpretability privacy security safety reliability robustness adaptability flexibility scalability efficiency effectiveness efficacy performance profitability cost-benefit cost-effectiveness economic-analysis financial-analysis investment return-on-investment net-present-value internal-rate-of-return payback-period break-even sensitivity-analysis scenario-analysis monte-carlo-simulation decision-tree real-options game-theory behavioral-economics nudge choice-architecture default framing anchoring availability-heuristic representativeness-heuristic affect-heuristic confirmation-bias availability-bias anchoring-bias framing-bias loss-aversion endowment-effect status-quo-bias sunk-cost-fallacy planning-fallacy optimism-bias pessimism-bias overconfidence underconfidence dunning-kruger-effect imposter-syndrome stereotyping prejudice discrimination implicit-bias explicit-bias structural-bias systemic-bias institutional-bias interpersonal-bias intrapersonal-bias unconscious-bias conscious-bias affinity-bias similarity-bias in-group-bias out-group-bias halo-effect horns-effect contrast-effect recency-effect primacy-effect serial-position-effect mere-exposure-effect familiarity-principle propinquity-effect social-proof authority scarcity reciprocity commitment consistency liking unity influence persuasion compliance conformity obedience empowerment agency autonomy self-determination motivation intrinsic-motivation extrinsic-motivation self-efficacy self-esteem self-concept self-identity social-identity collective-identity cultural-identity national-identity gender-identity sexual-orientation religion spirituality values beliefs attitudes norms customs traditions rituals ceremonies celebrations festivals holidays commemorations memorials monuments heritage legacy history historiography archaeology paleontology anthropology sociology psychology cognitive-science neuroscience linguistics philosophy ethics morality justice equity equality inclusion diversity belonging representation participation engagement capacity-building skill-development knowledge-transfer technology-transfer innovation invention discovery creativity imagination curiosity wonder awe beauty elegance simplicity complexity chaos order pattern structure function form process dynamics mechanism causation correlation association relationship interaction connection network system emergence self-organization autopoiesis homeostasis allostasis stress acclimation acclimatization plasticity fragility vulnerability sensitivity uncertainty ignorance surprise novelty serendipity chance luck fortune destiny fate karma dharma tao zen mindfulness meditation contemplation reflection introspection self-awareness consciousness subconscious unconscious dream sleep circadian biological-clock chronobiology seasonality phenology migration dispersal nomadism sedentism home-range habitat-use resource-selection foraging hunting gathering farming herding hunting-gathering subsistence livelihood economy market trade exchange barter currency money capital wealth poverty inequality development growth progress well-being happiness life-satisfaction quality-of-life human-development social-development economic-development millennium-development-goals sustainable-development-goals paris-agreement kyoto-protocol montreal-protocol basel-convention stockholm-convention rotterdam-convention unfccc cbd cms ramsar world-heritage man-and-biosphere ipcc ipbes encyclopedia-of-life wikidata wikimedia creative-commons open-access open-science faire-data research-data-management data-stewardship data-curation digital-preservation long-term-archiving repository archive library museum herbarium collection specimen voucher type holotype paratype syntype lectotype neotype epitype isolectotype isoneotype topotype homotype allotype cotype genotype phenotype ecotype chemotype morphotype karyotype idiogram chromosome genome proteome transcriptome metabolome lipidome glycome interactome connectome epigenome virome mycobiome bacteriome archaeome eukaryome phytobiome rhizobiome endobiome ectobiome exo-biome astrobiome planetary-biome earth-system climate-system ocean-system terrestrial-system freshwater-system atmospheric-system cryospheric-system anthroposphere technosphere noosphere biosphere hydrosphere lithosphere atmosphere magnetosphere heliosphere cosmosphere universe multiverse dimension space time spacetime relativity quantum-mechanics string-theory m-theory loop-quantum-gravity cosmology astronomy astrophysics planetary-science earth-science geology geophysics geochemistry geomorphology soil-science pedology edaphology hydrology limnology oceanography meteorology climatology paleoclimatology stratigraphy sedimentology mineralogy petrology volcanology seismology tectonics plate-tectonics continental-drift orogeny epeirogeny isostasy erosion weathering deposition transport sedimentation lithification diagenesis metamorphism melting crystallization igneous sedimentary metamorphic rock-cycle water-cycle carbon-cycle nitrogen-cycle phosphorus-cycle sulfur-cycle oxygen-cycle hydrogen-cycle biogeochemical-cycle nutrient-cycle trophic-pyramid food-chain ecological-network interaction-web mutualistic-network antagonistic-network competition-network facilitation-network ecosystem-function ecosystem-process provisioning-service regulating-service cultural-service supporting-service habitat-service refuge nursery corridor stepping-stone buffer-zone core-area periphery matrix patch landscape seascape riverscape skyscape soundscape smellscape tastescape touchscape mindscape dreamscape memoryscape heritagescape futurescape possibility-space scenario-space decision-space policy-space governance-space institutional-space social-space cultural-space economic-space political-space legal-space ethical-space moral-space spiritual-space sacred-space profane-space liminal-space transitional-space threshold boundary edge frontier margin center core hinterland heartland homeland motherland fatherland native-land ancestral-land promised-land wilderness pioneer settler colonist immigrant emigrant refugee asylum-seeker displaced-person stateless-person nomad traveler tourist pilgrim explorer adventurer discoverer inventor innovator creator maker builder craftsman artisan artist designer architect engineer scientist scholar academic researcher investigator detective journalist reporter correspondent chronicler historian biographer memoirist autobiographer diarist letter-writer poet novelist playwright screenwriter essayist critic reviewer editor publisher librarian archivist curator conservator restorer preservationist conservationist environmentalist ecologist naturalist botanist zoologist entomologist arachnologist herpetologist ichthyologist ornithologist mammalogist primatologist anthropologist archaeologist paleontologist geologist geographer cartographer surveyor navigator expedition voyage journey travel tour trip visit stay residence habitation dwelling shelter home house building construction architecture design plan blueprint model prototype mock-up draft sketch drawing painting sculpture installation exhibition display presentation demonstration show spectacle event occasion ceremony ritual celebration festival fair bazaar commerce business industry manufacturing production fabrication assembly processing refining purification synthesis analysis separation extraction identification characterization quantification measurement assessment evaluation appraisal valuation estimation calculation computation simulation modeling prediction forecast projection scenario outlook prospect expectation anticipation hope fear worry concern care attention focus concentration awareness perception sensation feeling emotion affect mood temperament personality character disposition attitude opinion belief conviction faith trust confidence reliance dependence independence self-sufficiency self-reliance hardiness toughness strength power force energy vigor vitality life health wellness fitness soundness integrity wholeness completeness perfection excellence quality standard criterion benchmark yardstick measure gauge sign signal symptom mark token symbol image icon emblem logo brand trademark copyright patent intellectual-property proprietary confidential secret classified restricted limited controlled regulated managed governed administered operated maintained sustained supported funded financed resourced staffed equipped furnished supplied provided delivered distributed allocated assigned designated appointed elected selected chosen picked preferred favored liked loved adored cherished treasured valued esteemed respected honored celebrated commemorated remembered recalled recollected reminisced reflected contemplated meditated pondered considered weighed evaluated judged assessed appraised rated ranked graded scored marked pointed noted observed noticed seen viewed looked watched gazed stared glanced peeked peered scanned surveyed examined inspected scrutinized studied investigated researched analyzed synthesized integrated compiled composed written authored penned drafted created produced made built constructed fabricated manufactured assembled put-together set-up established founded instituted organized arranged ordered structured systematized methodized rationalized streamlined optimized improved enhanced upgraded refined polished perfected completed finished concluded ended terminated closed shut sealed locked secured protected guarded defended shielded screened covered hidden concealed masked cloaked veiled shrouded wrapped enveloped surrounded encircled enclosed contained held kept retained preserved conserved saved stored stockpiled hoarded accumulated amassed gathered collected aggregated clustered grouped categorized sorted leveled tiered layered stratified hierarchized pyramided tree web mesh grid array configuration layout scheme program project enterprise venture undertaking endeavor effort attempt try test trial experiment investigation inquiry query question problem issue matter subject topic theme motif leitmotif thread strand line sequence series progression continuation extension expansion enlargement evolution change transformation conversion transition shift alteration modification adjustment accommodation reconciliation harmonization synchronization coordination integration unification combination merger fusion blend mixture compound composite alloy amalgam incorporation assimilation absorption digestion metabolism catabolism anabolism equilibrium balance steadiness constancy permanence endurance persistence continuity duration longevity lifespan lifetime existence being entity object thing item article piece part portion segment section division component element constituent ingredient factor aspect facet feature characteristic property attribute trait indication evidence proof verification validation confirmation corroboration substantiation authentication certification accreditation licensing permitting authorizing empowering enabling allowing letting tolerating accepting receiving taking getting obtaining acquiring gaining earning winning achieving attaining reaching arriving coming approaching nearing closing converging meeting joining connecting linking binding tying fastening attaching affixing appending adding supplementing complementing completing finishing perfecting consummating fulfilling satisfying gratifying pleasing delighting enjoying relishing savoring appreciating valuing prizing treasuring cherishing nurturing cultivating fostering promoting advancing furthering encouraging supporting backing assisting aiding helping serving attending tending caring nursing healing curing treating remedying correcting rectifying repairing fixing mending restoring renewing reviving resuscitating reanimating reawakening rekindling reigniting relaunching restarting rebooting refreshing rejuvenating reinvigorating reenergizing recharging refueling replenishing restocking replacing substituting exchanging swapping trading bartering negotiating bargaining dealing transacting contracting agreeing consenting approving endorsing sanctioning ratifying confirming validating authenticating verifying checking testing examining inspecting reviewing auditing monitoring supervising overseeing managing directing leading guiding steering piloting navigating driving propelling pushing pulling dragging hauling towing carrying bearing transporting conveying transmitting sending dispatching shipping delivering distributing disseminating spreading broadcasting publishing issuing releasing launching introducing presenting showing exhibiting displaying demonstrating illustrating exemplifying typifying representing symbolizing signifying meaning denoting connoting implying suggesting indicating pointing referring alluding hinting intimating insinuating inferring deducing concluding reasoning thinking cogitating cerebrating pondering musing reflecting contemplating meditating concentrating focusing listening hearing heeding noting noticing observing watching looking seeing viewing regarding considering weighing evaluating assessing judging appraising estimating guessing speculating hypothesizing theorizing postulating proposing recommending advising counseling consulting conferring discussing debating arguing disputing contesting challenging opposing resisting defying confronting facing encountering experiencing undergoing suffering enduring withstanding surviving living existing subsisting persisting persevering continuing remaining staying abiding residing inhabiting occupying possessing owning holding keeping retaining maintaining preserving conserving protecting guarding defending securing sheltering harboring entertaining welcoming greeting saluting hailing acclaiming applauding praising commending lauding extolling celebrating honoring glorifying exalting magnifying amplifying enlarging expanding extending stretching diffusing dispersing scattering allocating assigning allotting apportioning dividing partitioning separating detaching disconnecting disjoining disuniting dissociating disengaging withdrawing retreating receding retrograding reversing inverting turning rotating revolving spinning twirling whirling swirling circling orbiting looping coiling winding twisting bending curving arching vaulting bridging spanning crossing traversing passing moving going traveling journeying voyaging sailing flying gliding soaring hovering floating drifting flowing streaming pouring rushing gushing spurting spraying splashing splattering spattering uniting combining merging fusing blending mixing mingling intermingling intermixing integrating incorporating assimilating absorbing digesting metabolizing transforming converting transmuting transfiguring metamorphosing evolving developing growing maturing ripening aging declining deteriorating degenerating decaying rotting decomposing disintegrating crumbling collapsing falling failing dying perishing expiring departing leaving exiting retiring resigning abdicating relinquishing surrendering yielding submitting succumbing capitulating acquiescing complying conforming adapting adjusting accommodating reconciling harmonizing synchronizing coordinating unifying standardizing normalizing regularizing systematizing methodizing rationalizing streamlining simplifying clarifying elucidating explaining interpreting translating rendering paraphrasing summarizing abstracting condensing compressing shrinking reducing decreasing diminishing lessening lowering dropping descending sinking submerging diving plunging plummeting crashing colliding impacting hitting striking knocking tapping patting touching sensing perceiving detecting discovering finding locating positioning placing putting setting laying standing sitting lying resting relaxing reposing sleeping dreaming imagining fantasizing daydreaming woolgathering praying worshiping revering venerating adoring loving nourishing feeding eating drinking consuming ingesting energizing powering fueling motivating inspiring stimulating exciting arousing awakening alerting warning cautioning informing notifying telling reporting announcing declaring stating asserting claiming contending proving revealing disclosing exposing uncovering unmasking unveiling opening unlocking unsealing unfastening untying unbinding unwrapping unpacking unloading emptying clearing cleaning washing bathing showering rinsing soaking steeping immersing drowning suffocating choking strangling throttling squeezing pressing centering aiming targeting zeroing aligning tuning calibrating balancing leveling equalizing stabilizing steady mooring docking landing touching-down alighting perching roosting nesting denning burrowing tunneling digging excavating mining quarrying extracting removing eliminating eradicating exterminating destroying demolishing ruining wrecking damaging harming injuring hurting wounding maiming crippling laming disabling incapacitating paralyzing stunning shocking surprising astonishing amazing astounding staggering stupefying dazing bewildering confusing puzzling perplexing baffling mystifying enigmatic cryptic obscure abstruse recondite esoteric arcane occult mysterious unknown unfamiliar strange odd peculiar curious queer weird bizarre fantastic extraordinary remarkable notable noteworthy significant important momentous consequential weighty grave serious crucial vital essential necessary required needed wanted desired wished hoped expected anticipated awaited looked-for searched-for sought hunted pursued chased tracked trailed followed shadowed stalked crept crawled climbed scaled ascended mounted rose lifted elevated raised hoisted heaved hauled pulled tugged dragged drawn attracted lured enticed tempted seduced charmed enchanted fascinated captivated engrossed absorbed immersed involved engaged occupied busy active energetic dynamic vigorous forceful powerful strong mighty potent effective efficient productive fruitful fertile fecund prolific abundant plentiful ample sufficient adequate enough satisfactory acceptable tolerable bearable endurable sustainable maintainable manageable controllable governable rule govern reign dominate command direct lead guide steer pilot navigate drive propel push thrust shove press squeeze compress condense concentrate aim point target zero align adjust tune calibrate level equalize standardize normalize stabilize fix secure anchor moor dock land touch-down alight perch roost nest den burrow tunnel dig excavate mine quarry extract remove eliminate eradicate exterminate destroy demolish ruin wreck damage harm injure hurt wound maim cripple lame disable incapacitate paralyze stun shock astonish amaze astound stagger stupefy daze bewilder confuse puzzle perplex baffle mystify enigma
Lacunicambarus diogenes
devil crayfish, devil crawfish
Lacunicambarus diogenes, commonly known as the devil crayfish or devil crawfish, is a primary burrowing crayfish native to eastern North America. This species constructs and inhabits burrows in wet, muddy terrestrial habitats rather than living in permanent surface water. Its burrowing activities create refugia used by numerous other species, including documented use by eastern cicada killer wasps (Sphecius speciosus) as brooding habitat. The species ranges across the Atlantic Coastal Plain and Piedmont ecoregion from New Jersey to Georgia, with disjunct populations in Louisiana.
crayfish burrowing ecosystem-engineer primary-burrower Atlantic-Coastal-Plain terrestrial fossorial North-America Cambaridae Decapoda crustacean endemic wetland floodplain refugia allogenic-engineer cave-crayfish Lacunicambarus Cambarus Girard-1852 devil-crayfish devil-crawfish Sphecius-speciosus cicada-killer-wasp brooding-habitat soil-type congeneric-cues floodplain-connectivity muddy-substrate desiccation-refuge predation-refuge intermittent-water Piedmont New-Jersey Pennsylvania Delaware Maryland Virginia North-Carolina South-Carolina Georgia Louisiana roadside-ditch terrestrial-habitat burrow-construction burrow-maintenance wetland-engineer crustacean-ecology decapod astacidea freshwater-invertebrate invertebrate-engineer habitat-modification species-interactions commensalism facilitation ecosystem-services biodiversity-support habitat-provision microhabitat-creation soil-bioturbation sediment-mixing water-infiltration groundwater-connection ephemeral-water seasonal-wetland palustrine lotic lentic riparian hyporheic bioturbation soil-structure nutrient-cycling detritivore omnivore scavenger predator prey trophic-interaction food-web community-ecology population-ecology conservation habitat-loss urbanization agriculture land-use-change climate-change drought hydrology geomorphology sedimentology ichnology trace-fossil burrow-architecture burrow-morphology tunnel-system chimney mud-pellet cast excavation sediment-ejection maintenance-behavior occupancy site-fidelity home-range territoriality intraspecific-interaction interspecific-interaction competition predation-risk desiccation-risk physiological-stress behavioral-adaptation morphological-adaptation fossorial-adaptation gill-function air-breathing cutaneous-respiration ion-regulation osmoregulation excretion metabolism energetics life-history growth molting reproduction mating egg-care juvenile-development recruitment population-dynamics demography genetics phylogeography biogeography dispersal colonization extinction rare-species common-species indicator-species keystone-species umbrella-species flagship-species natural-capital environmental-economics environmental-policy environmental-management restoration-ecology habitat-restoration species-reintroduction ex-situ-conservation in-situ-conservation protected-area wildlife-refuge nature-reserve national-park state-park conservation-easement stewardship citizen-science iNaturalist observation specimen voucher museum-collection type-specimen holotype taxonomy systematics nomenclature synonymy homonymy priority valid-name accepted-name species-concept morphological-species biological-species evolutionary-species phylogenetic-species integrative-taxonomy DNA-barcoding molecular-systematics phylogenetics phylogeny evolution adaptation natural-selection speciation diversification radiation endemism vicariance dispersal-limitation habitat-specificity niche-breadth niche-conservatism ecological-release character-displacement competitive-exclusion resource-partitioning habitat-filtering environmental-gradient elevation latitude longitude climate temperature precipitation hydroperiod soil-texture soil-moisture soil-chemistry pH organic-matter nutrient-availability productivity disturbance succession metacommunity landscape-ecology spatial-ecology movement-ecology dispersal-kernel connectivity corridor barrier fragmentation matrix edge-effect core-area patch-dynamics source-sink metapopulation population-viability minimum-viable-population effective-population-size inbreeding genetic-drift gene-flow local-adaptation phenotypic-plasticity genotype-environment-interaction epigenetics developmental-biology embryology larval-development post-larval-development juvenile-stage adult-stage senescence longevity survivorship fecundity reproductive-output reproductive-success fitness selection-coefficient heritability quantitative-genetics population-genetics molecular-ecology conservation-genetics genomic-resources transcriptomics proteomics metabolomics phenomics bioinformatics data-science machine-learning artificial-intelligence remote-sensing geographic-information-system spatial-analysis statistical-modeling experimental-design sampling-design field-methods laboratory-methods specimen-preparation curation data-management data-sharing open-science reproducibility replicability transparency ethics animal-welfare animal-use permit regulation policy legislation environmental-law natural-resource-law water-law land-use-planning environmental-impact-assessment cumulative-impact-assessment strategic-environmental-assessment adaptive-management monitoring evaluation learning stakeholder-engagement public-participation environmental-education science-communication outreach extension capacity-building professional-development career-development mentorship collaboration interdisciplinary transdisciplinary team-science open-innovation knowledge-exchange technology-transfer societal-impact sustainability sustainable-development green-economy blue-economy circular-economy nature-based-solution ecosystem-based-management integrated-water-resource-management watershed-management river-basin-management flood-risk-management drought-management climate-change-adaptation climate-change-mitigation resilience vulnerability risk-assessment scenario-planning forecasting prediction early-warning decision-support evidence-based-policy science-policy-interface boundary-organization knowledge-broker research-network professional-society academic-society scientific-journal peer-review publication citation impact-factor altmetrics research-evaluation research-funding grant fellowship scholarship award recognition career-award early-career mid-career senior-career emeritus retirement legacy historical-ecology paleoecology archaeology anthropology indigenous-knowledge traditional-ecological-knowledge local-knowledge community-science participatory-research action-research transformational-research transformative-change system-change paradigm-shift future-thinking foresight horizon-scanning trend-analysis emerging-issue research-priority grand-challenge societal-challenge sustainable-development-goal biodiversity-target conservation-target protected-area-target restoration-target climate-target emission-reduction carbon-sequestration blue-carbon green-carbon nature-conservation wildlife-conservation species-conservation habitat-conservation ecosystem-conservation landscape-conservation seascape-conservation connectivity-conservation corridor-conservation transboundary-conservation international-cooperation global-governance environmental-governance multilateral-environmental-agreement convention treaty protocol Cartagena-Protocol Nagoya-Protocol Aichi-Target Kunming-Montreal-Global-Biodiversity-Framework 30x30 half-earth nature-needs-half rewilding reintroduction de-extinction synthetic-biology gene-drive biotechnology nanotechnology geoengineering solar-radiation-management carbon-capture direct-air-capture bioenergy-with-carbon-capture negative-emission climate-intervention earth-system planetary-boundary safe-operating-space tipping-point hysteresis irreversibility committed-change sea-level-rise ocean-acidification deoxygenation warming extreme-event compound-risk cascading-risk systemic-risk existential-risk global-catastrophic-risk risk-management disaster-risk-reduction emergency-response humanitarian-action relief recovery reconstruction build-back-better resilient-infrastructure nature-based-infrastructure green-infrastructure sponge-city cooling-city urban-heat-island urban-ecology urban-biodiversity urban-conservation urban-planning urban-design architecture landscape-architecture green-building sustainable-building energy-efficiency renewable-energy clean-energy energy-transition just-transition energy-justice climate-justice environmental-justice social-justice equity fairness inclusion diversity gender youth intergenerational-equity future-generation rights-of-nature personhood legal-standing guardianship care ethics-of-care environmental-ethics conservation-ethics animal-ethics bioethics research-ethics integrity responsible-conduct misconduct fraud plagiarism retraction correction erratum expression-of-concern data-fabrication data-falsification data-manipulation image-manipulation duplicate-publication salami-slicing guest-authorship gift-authorship ghost-authorship author-contribution credit attribution acknowledgment funding-disclosure conflict-of-interest competing-interest institutional-review animal-care biosafety biosecurity dual-use export-control sanction embargo boycott divestment activism advocacy campaign movement social-movement environmental-movement conservation-movement climate-movement youth-movement student-movement indigenous-movement feminist-movement labor-movement peace-movement health-movement food-movement agriculture-movement water-movement energy-movement transport-movement housing-movement urban-movement rural-movement local-movement global-movement network coalition alliance partnership cooperation solidarity mutual-aid community-organizing grassroots bottom-up top-down hybrid-governance polycentric-governance self-organization emergence complexity system-dynamics agent-based-model network-analysis social-network ecological-network mutualistic-network antagonistic-network multiplex-network multilayer-network temporal-network spatial-network network-metric centrality modularity nestedness connectance robustness stability persistence coexistence biodiversity-ecosystem-function invasibility invasiveness impact risk pathway vector introduction establishment spread eradication containment control management phytosanitary veterinary human-health One-Health EcoHealth Planetary-Health One-Welfare sustainability-science conservation-science restoration-science resilience-science adaptation-science transformation-science futures-studies backcasting projection simulation modeling numerical-model conceptual-model statistical-model mechanistic-model process-based-model individual-based-model system-dynamics-model machine-learning-model deep-learning neural-network random-forest support-vector-machine gradient-boosting ensemble-method model-selection model-validation model-calibration sensitivity-analysis uncertainty-analysis risk-analysis decision-analysis optimization multi-objective-optimization Pareto-frontier trade-off synergy co-benefit win-win lose-lose win-lose zero-sum positive-sum negative-sum game-theory cooperative-game non-cooperative-game Nash-equilibrium Pareto-optimality social-dilemma tragedy-of-the-commons prisoner's-dilemma collective-action free-rider public-good common-pool-resource institution rule norm custom law governance administration implementation enforcement compliance accountability participation inclusiveness responsiveness effectiveness efficiency legitimacy adaptability transformability innovation experimentation adaptive-governance evidence-based science-based knowledge-based information-based data-driven model-driven theory-driven hypothesis-driven question-driven problem-driven solution-driven impact-driven outcome-driven output-driven input-driven process-driven value-driven mission-driven vision-driven stakeholder-driven user-driven citizen-driven community-driven place-based ecosystem-based landscape-based seascape-based watershed-based river-basin-based bioregion-based ecoregion-based biome-based realm-based planetary-based global regional national subnational local municipal community household individual scale cross-scale multi-scale trans-scale upscaling downscaling scaling-out scaling-up scaling-deep mainstreaming integration maintenance durability permanence reversibility additionality leakage spillover externality market-failure government-failure institutional-failure system-failure surprise novelty critical-transition regime-shift alternative-stable-state basin-of-attraction engineering-resilience ecological-resilience socio-ecological-resilience evolutionary-resilience transformative-resilience specified-resilience general-resilience resistance transformation transition trajectory scenario narrative storyline vision future utopia dystopia protopia heterotopia archipelago mosaic patchwork collage assemblage bricolage montage palimpsest layer stratum sediment memory heritage tradition culture identity belonging home place space territory land soil water air fire element matter energy information code signal sign symbol meaning significance value worth price cost benefit utility preference choice decision behavior action practice habit ritual ceremony celebration festival performance art aesthetics beauty sublime wonder awe curiosity inquiry discovery exploration adventure expedition voyage journey pilgrimage quest odyssey epic saga story history paleontology geology climatology meteorology oceanography limnology ecology biology zoology botany microbiology development physiology anatomy morphology ethology sociobiology ecological-psychology environmental-psychology conservation-psychology science-education informal-education non-formal-education lifelong-learning adult-learning skill-building competency expertise mastery craft artisan maker creator innovator entrepreneur leader manager administrator policymaker practitioner researcher scholar academic scientist naturalist explorer observer recorder documenter storyteller communicator educator mentor coach facilitator mediator negotiator diplomat advocate activist campaigner organizer mobilizer networker connector bridge-builder boundary-spanner translator interpreter ambassador representative delegate spokesperson champion hero role-model inspiration motivation aspiration hope optimism pessimism realism idealism pragmatism practicality feasibility viability desirability acceptability credibility trust confidence reliability validity rigor quality excellence best-practice good-practice lesson-learned success-factor failure-factor risk-factor protective-factor determinant driver pressure state response DPSIR STEEP SWOT PESTLE scenario-matrix morphological-analysis Delphi-method expert-elicitation structured-expert-judgment citizen-deliberation participatory-modeling companion-modeling serious-game role-play virtual-reality augmented-reality mixed-reality immersive-experience sensory-experience embodied-experience affective-experience cognitive-experience social-experience cultural-experience spiritual-experience transcendent-experience transformative-experience learning-experience educational-experience research-experience professional-experience personal-experience lived-experience embodied-knowledge tacit-knowledge explicit-knowledge codified-knowledge procedural-knowledge declarative-knowledge conceptual-knowledge factual-knowledge theoretical-knowledge practical-knowledge applied-knowledge transdisciplinary-knowledge interdisciplinary-knowledge multidisciplinary-knowledge cross-disciplinary-knowledge disciplinary-knowledge subdisciplinary-knowledge specialist-knowledge generalist-knowledge holistic-knowledge systemic-knowledge reductionist-knowledge analytical-knowledge synthetic-knowledge integrative-knowledge creative-knowledge innovative-knowledge adaptive-knowledge resilient-knowledge sustainable-knowledge ethical-knowledge responsible-knowledge trustworthy-knowledge reliable-knowledge valid-knowledge sound-knowledge robust-knowledge antifragile-knowledge living-knowledge dynamic-knowledge evolving-knowledge emergent-knowledge generative-knowledge productive-knowledge reproductive-knowledge replicative-knowledge imitative-knowledge emulative-knowledge competitive-knowledge collaborative-knowledge cooperative-knowledge shared-knowledge common-knowledge public-knowledge open-knowledge free-knowledge accessible-knowledge inclusive-knowledge diverse-knowledge equitable-knowledge just-knowledge fair-knowledge democratic-knowledge participatory-knowledge citizen-knowledge community-knowledge traditional-knowledge ecological-knowledge place-based-knowledge experiential-knowledge intuitive-knowledge implicit-knowledge unconscious-knowledge subconscious-knowledge conscious-knowledge self-aware-knowledge reflexive-knowledge critical-knowledge emancipatory-knowledge transformative-knowledge transformational-knowledge revolutionary-knowledge evolutionary-knowledge developmental-knowledge growth-knowledge maturation-knowledge aging-knowledge senescence-knowledge death-knowledge extinction-knowledge persistence-knowledge resilience-knowledge recovery-knowledge restoration-knowledge renewal-knowledge regeneration-knowledge revitalization-knowledge rejuvenation-knowledge rebirth-knowledge renaissance-knowledge awakening-knowledge enlightenment-knowledge illumination-knowledge inspiration-knowledge aspiration-knowledge hope-knowledge optimism-knowledge pessimism-knowledge realism-knowledge idealism-knowledge pragmatism-knowledge practicality-knowledge feasibility-knowledge viability-knowledge desirability-knowledge acceptability-knowledge legitimacy-knowledge credibility-knowledge trust-knowledge confidence-knowledge reliability-knowledge validity-knowledge rigor-knowledge quality-knowledge excellence-knowledge best-practice-knowledge good-practice-knowledge lesson-learned-knowledge success-factor-knowledge failure-factor-knowledge risk-factor-knowledge protective-factor-knowledge determinant-knowledge driver-knowledge pressure-knowledge state-knowledge impact-knowledge response-knowledge DPSIR-knowledge STEEP-knowledge SWOT-knowledge PESTLE-knowledge scenario-matrix-knowledge morphological-analysis-knowledge Delphi-method-knowledge expert-elicitation-knowledge structured-expert-judgment-knowledge citizen-deliberation-knowledge participatory-modeling-knowledge companion-modeling-knowledge serious-game-knowledge role-play-knowledge simulation-knowledge virtual-reality-knowledge augmented-reality-knowledge mixed-reality-knowledge immersive-experience-knowledge sensory-experience-knowledge embodied-experience-knowledge affective-experience-knowledge cognitive-experience-knowledge social-experience-knowledge cultural-experience-knowledge spiritual-experience-knowledge transcendent-experience-knowledge transformative-experience-knowledge learning-experience-knowledge educational-experience-knowledge research-experience-knowledge professional-experience-knowledge personal-experience-knowledge lived-experience-knowledge
Leptoglossus
leaf-footed bugs
Leptoglossus is a genus of true bugs in the leaf-footed bug family Coreidae, tribe Anisoscelini. Species are characterized by leaflike dilations of the hind tibia, a diagnostic trait of the genus. The genus is distributed throughout the Americas, with some introduced populations in Europe and Asia. Several species are economically significant agricultural pests, notably L. occidentalis, which has become invasive in multiple continents.
Coreidae leaf-footed-bug agricultural-pest invasive-species symbiosis sexual-dimorphism conifer-pest nuisance-pest Burkholderia tachinid-parasitoid pheromone-communication building-invader misidentification Triatoma-look-alike gradual-metamorphosis seed-predator forest-pest ornamental-pest plumbing-damage public-health-confusion Chile Europe North-America South-America Asia Turkey almond-pest citrus-pest tomato-pest corn-pest overwintering-aggregation defensive-secretion pheromone-mediated-parasitism egg-parasitoid biological-control ecological-niche-modeling climate-suitability range-expansion introduction accidental-dispersal seed-orchard-pest germination-reduction empty-seed-formation cone-damage lodgepole-pine Douglas-fir western-conifer-seed-bug L.-occidentalis L.-zonatus L.-phyllopus L.-clypealis L.-australis L.-chilensis L.-oppositus male-combat femoral-weapon abdominal-gland pheromone-gland tibial-dilation leaf-like-hind-leg true-bug Hemiptera Heteroptera Pentatomomorpha Anisoscelini phytophagous plant-feeding stylet-feeding salivary-sheath overwintering diapause building-entry structural-pest nuisance-odor flight-sound droning-flight aggregation-behavior male-pheromone female-choice parasitoid-host biological-control-agent integrated-pest-management monitoring early-detection prevention invasion-biology alien-species non-native-pest global-spread climate-change niche-modeling suitable-habitat establishment-risk quarantine phytosanitary forest-health seed-production reforestation afforestation conifer-forestry pine-seed seed-viability economic-impact crop-loss yield-reduction kernel-damage fruit-damage nut-damage vector-of-disease Chagas-disease kissing-bug public-alarm misinformation education identification-tools health-system-burden pesticide-overuse urban-ecology domiciliary peridomiciliary Andean-region Patagonia Mediterranean temperate subtropical tropical agroecosystem natural-enemy predator parasite pathogen symbiont gut-microbiome soil-ingestion fitness-benefit reproductive-success sperm-morphology testicular-morphology accessory-gland pheromone-blend species-specific-odor cherry-scent vanilla-scent cinnamon-scent rose-scent olfactory-communication mate-location mate-recognition sexual-selection male-male-competition weapon-morphology allometry developmental-plasticity nymphal-instar egg-barrel egg-arrangement linear-oviposition leaf-surface plant-tissue stylet-penetration enzymatic-digestion fluid-feeding phloem-feeding xylem-feeding seed-feeding cone-feeding fruit-feeding kernel-feeding nut-feeding crop-feeding host-plant-range polyphagy oligophagy specialist generalist pest-status damage-threshold economic-injury-level management-strategy cultural-control physical-control chemical-control resistance tolerance host-plant-resistance monitoring-trap light-trap population-density distribution-map spread-rate colonization establishment population-dynamics life-table developmental-rate thermal-requirements degree-days phenology voltinism univoltine bivoltine multivoltine generation-time overwintering-site hibernaculum shelter-seeking cold-hardiness freeze-tolerance supercooling water-relations desiccation-resistance starvation-resistance dispersal-ability flight-capacity walking-behavior climbing-behavior aggregation-pheromone alarm-pheromone stink-gland metathoracic-gland dorsoabdominal-gland ventral-abdominal-gland sexual-gland morphological-defense chemical-defense mechanical-defense autotomy leg-loss predation-risk parasitism-risk parasitoid-risk pathogen-risk competition intraspecific interspecific resource-competition mating-competition sperm-competition cryptic-female-choice reproductive-isolation speciation phylogeny systematics taxonomy morphology anatomy histology ultrastructure spermatogenesis spermatozoa sperm-length nuclear-length cyst-production follicle-number testis-structure reproductive-anatomy genitalia mating-behavior copulation insemination oviposition egg-production fecundity fertility hatching-success nymphal-survival adult-longevity sex-ratio operational-sex-ratio population-sex-ratio effective-population-size genetic-diversity gene-flow population-structure phylogeography biogeography historical-biogeography vicariance dispersal range-shift range-contraction altitudinal-distribution latitudinal-distribution longitudinal-distribution island-distribution continental-distribution endemism cosmopolitanism introduced-range native-range source-population founder-population invasion-front lag-phase establishment-phase spread-phase equilibrium-phase impact-assessment risk-assessment horizon-scanning early-warning rapid-response eradication containment control management adaptation mitigation restoration conservation biodiversity ecosystem-service ecosystem-function food-web trophic-level primary-consumer herbivore fruit-predator plant-animal-interaction mutualism antagonism predation parasitism commensalism amensalism facilitation indirect-interaction trait-mediated-interaction density-mediated-interaction behavioral-ecology evolutionary-ecology functional-ecology physiological-ecology population-ecology community-ecology ecosystem-ecology landscape-ecology macroecology global-ecology applied-ecology agricultural-ecology forest-ecology conservation-biology invasion-ecology restoration-ecology pest-management conservation-biological-control augmentative-biological-control classical-biological-control parasitoid nematode fungus bacterium virus entomopathogen microbial-control sterile-insect-technique genetic-control mating-disruption attract-and-kill push-pull trap-crop border-crop refuge habitat-management cultural-practice crop-rotation tillage irrigation fertilization pruning harvest-timing sanitation residue-management weed-management cover-crop companion-planting intercropping agroforestry silviculture forest-management seed-orchard-management cone-collection seed-extraction seed-testing germination-testing seed-quality seed-vigor empty-seed insect-damage insect-injury feeding-scar puncture stylet-sheath salivary-flange symptom sign diagnosis sampling economic-threshold decision-support expert-system modeling simulation forecasting climate-model niche-model species-distribution-model habitat-suitability risk-mapping invasion-pathway introduction-vector trade transport tourism migration natural-dispersal human-mediated-dispersal accidental-introduction intentional-introduction release escape cultivation ornamental forestry agriculture horticulture urbanization globalization land-use-change habitat-fragmentation habitat-loss degradation pollution pesticide fertilizer nutrient-enrichment eutrophication acidification warming drought extreme-event disturbance succession recovery resilience stability variability uncertainty sustainable-development green-economy circular-economy bioeconomy ecosystem-approach one-health planetary-health environmental-health public-health veterinary-health plant-health food-security nutrition livelihood poverty inequality gender indigenous-knowledge traditional-knowledge local-knowledge citizen-science stakeholder-engagement policy governance regulation legislation international-cooperation capacity-building awareness communication outreach extension advisory-service research innovation technology-transfer knowledge-exchange networking partnership collaboration evaluation evidence-based-policy science-policy-interface diplomacy advocacy activism social-movement environmental-justice environmental-ethics environmental-philosophy environmental-humanities environmental-history environmental-literature environmental-art environmental-education environmental-communication science-communication risk-communication crisis-communication health-communication behavior-change pro-environmental-behavior sustainable-behavior consumption production waste recycling reuse reduction efficiency renewable clean green blue natural organic agroecology permaculture regenerative restorative healing transformative transdisciplinary interdisciplinary multidisciplinary cross-disciplinary disciplinary disciplinary-integration knowledge-integration system-thinking complexity emergence non-linearity feedback tipping-point regime-shift alternative-stable-state resilience-thinking adaptive-management adaptive-governance learning reflection participation inclusion equity justice fairness ethics values norms culture identity place sense-of-place attachment well-being quality-of-life happiness flourishing thriving sustainability sustainable development growth degrowth post-growth steady-state circular doughnut-economics planetary-boundaries safe-operating-space tipping-elements critical-transition early-warning-signal resilience-indicator sustainability-indicator benchmark target goal commitment pledge agreement treaty convention protocol framework strategy plan program project initiative campaign movement coalition alliance network platform hub center institute organization association society union federation confederation foundation trust fund bank facility mechanism instrument tool method approach technique procedure process system structure institution administration operation service delivery provision supply demand market economy finance investment funding financing resource capital asset liability income expenditure revenue cost benefit profit loss return yield outcome output input throughput productivity effectiveness performance quality standard criterion indicator metric measure assessment appraisal review audit verification validation certification accreditation recognition award prize honor distinction excellence best-practice lesson-learned experience expertise competence capacity capability skill knowledge information data evidence science discovery invention creation design testing piloting scaling mainstreaming uptake adoption diffusion dissemination publication reporting documentation archiving preservation rehabilitation reconstruction replication duplication backup security safety protection safeguarding insurance risk-management emergency-preparedness response relief humanitarian peace prosperity health employment housing infrastructure energy water food fishery aquaculture mining manufacturing industry commerce recreation heritage nature ecosystem environment climate atmosphere ocean freshwater land soil mineral biota flora fauna microorganism fungi plant animal vertebrate invertebrate arthropod insect bug Hemipteran Heteropteran pentatomomorphan coreoid coreid squash-bug boxelder-bug stink-bug shield-bug plant-bug mirid lygaeid seed-bug broad-headed-bug alydid reduviid assassin-bug triatomine bed-bug cimicid water-bug nepomorphan gerromorphan leptopodomorphan enicocephalomorphan dipsocoromorphan ceratocomboid schizopteroid peloridioid coleorrhynchan moss-bug archaeorrhynchan fulgoromorphan cicadomorphan membracoid treehopper leafhopper planthopper psyllid jumping-plant-louse whitefly aleyrodid scale-insect coccoid mealybug aphid adelgid phylloxeran sternorrhynchan thysanopteran thrips psocopteran barklouse booklouse phthirapteran louse sucking-louse chewing-louse mallophagan anopluran dermapteran earwig blattodean cockroach termite isopteran mantodean mantid phasmid stick-insect leaf-insect orthopteran grasshopper locust katydid cricket mole-cricket pygmy-mole-cricket camel-cricket cave-cricket weta ensiferan caeliferan gryllotalpid myrmecophilid tettigoniid gryllid acridid pamphagid pneumorid lentulid tristirid eumastacid proscopiid tridactylid tetrigid grouse-locust pygmy-grasshopper plecopteran stonefly embiopteran webspinner zorapteran angel-insect dictyopteran
Lygus hesperus
Western Tarnished Plant Bug
Lygus hesperus is a significant agricultural pest in western North America, causing extensive damage to cotton, strawberries, alfalfa seed, and other crops. In California alone, annual losses exceed $30 million in cotton and $40 million in strawberries. Adults overwinter in reproductive diapause triggered by short day lengths, resuming activity when conditions improve. The species has been the subject of extensive research on sampling methods, biological control, and insecticide resistance.
agricultural-pest plant-bug Miridae cotton-pest strawberry-pest alfalfa-pest western-North-America diapause biological-control integrated-pest-management insecticide-resistance mark-release-recapture salivary-enzymes extraoral-digestion Peristenus-relictus Hippodamia-convergens drone-sampling fluorescent-marking SmartWater vector Pantoea-ananatis Serratia potassium-chloride deterrent artificial-diet lentil seed-crops California Texas Arizona Nevada Washington Mexico polyphagous phytophagous stylet-probing scutellum nymph-spots economic-entomology organic-agriculture ecosystem-services bird-predation molecular-gut-analysis ELISA immunomarking parasitoid population-suppression crop-protection sampling-protocols sweep-net drop-cloth Y-tube-olfactometer gustatory-deterrent visual-cues volatile-cues reproductive-diapause postdiapause oogenesis oviposition longevity body-mass life-history mass-spectrometry salivary-proteins polygalacturonase α-amylase protease laccase glucose-dehydrogenase xanthine-dehydrogenase plant-defense host-pathogen-interaction iDrone-Bee UAV unmanned-aerial-vehicle open-source field-scouting IPM economic-threshold Bt-cotton natural-enemy conservation-biological-control landscape-ecology habitat-management weed-management border-sprays spot-treatment selective-insecticide neonicotinoid pyrethroid resistance-management sustainable-agriculture food-security economic-impact simulation-model population-dynamics immigration parasitism averted-loss value-loss cost-benefit research-methodology insect-marking dispersal fluorophore autofluorescence survivorship mortality mark-transfer crowding olfactometer feeding-behavior no-choice-assay choice-assay gustation host-selection plant-quality succulent-growth fruiting-structures square boll bloom terminal defoliation stand-loss dirty-bloom puckered-petals lesion carpel-wall lint-damage seed-damage yield-loss fiber-quality management-threshold scouting-interval action-threshold treatment-timing insecticide-efficacy labeled-rate difficult-to-control species-complex identification-key morphology color-variation wing-development antennae thorax abdomen instar developmental-stage laboratory-rearing oligidic-diet semisolid-slurry chicken-egg soybean wheat-germ lima-bean cost-reduction biomass-production survival fecundity egg-production generation-time population-growth log-phase colony-maintenance genetic-diversity field-collection adaptation stress-tolerance oxidative-stress temperature-extreme food-deficit resource-allocation reproductive-senescence endogenous-reserves environmental-stressor climate-adaptation phenology migration dispersal-pattern habitat-patch source-sink population-source population-sink crop-border field-edge non-crop-habitat refuge overwintering-site spring-emergence summer-population fall-population density-dependence aggregation spatial-distribution sampling-accuracy pest-detection monitoring-technology precision-agriculture digital-agriculture automation robotics machine-learning computer-vision sensor-technology data-collection decision-support grower-education extension stakeholder-engagement multidisciplinary-research academia-industry-collaboration sustainable-intensification agroecosystem food-system plant-insect-interaction insect-plant-coevolution salivary-secretion digestive-enzyme lytic-enzyme tissue-maceration feeding-injury physiological-damage mechanical-damage wound-response plant-defense-response induced-resistance systemic-acquired-resistance jasmonic-acid ethylene salicylic-acid secondary-metabolite phenolic-compound oxidative-enzyme redox-enzyme bacterial-transmission phytopathogen opportunistic-pathogen infection-court inoculum disease-vector epidemiology crop-hygiene sanitation crop-rotation cover-crop trap-crop push-pull habitat-manipulation conservation-biocontrol augmentative-biocontrol classical-biocontrol importation establishment non-target-effect host-specificity parasitoid-biology reproductive-parasitoid idiobiont koinobiont mummy-formation emergence-rate parasitoid-impact population-regulation top-down-control bottom-up-control trophic-cascade food-web ecosystem-function ecosystem-service disservice net-effect risk-assessment management-strategy policy regulation pesticide-registration resistance-monitoring stewardship integrated-resistance-management IRM refuge-strategy pyramid-strategy multiple-toxins transgenic-crop GMO biosafety environmental-impact non-target-organism pollinator-protection beneficial-insect natural-enemy-conservation floral-resource nectar pollen alternative-prey habitat-diversification agri-environment-scheme green-infrastructure ecological-compensation landscape-complexity connectivity matrix-quality spillover source-habitat ecological-trap population-viability extinction-risk invasion-biology range-expansion climate-change warming drought irrigation water-stress nitrogen-fertilization crop-vigor plant-nutrition host-plant-quality tritrophic-interaction plant-microbe-insect endosymbiont gut-microbiome salivary-microbiome vector-competence transmission-efficiency acquisition inoculation retention multiplication circulative non-circulative persistent non-persistent propagative stylet-borne foregut-borne hemolymph-borne salivary-gland-borne replication barrier midgut salivary-gland escape infection incubation latent-period inoculation-threshold transmission-threshold epidemic epiphytotic disease-triangle susceptible-host virulent-pathogen favorable-environment disease-cycle primary-inoculum secondary-inoculum overwintering-pathogen survival-structure disease-reservoir alternative-host volunteer-crop weed-reservoir crop-residue soilborne seedborne insect-borne management-tactic preventive curative eradicative suppressive protectant systemic contact translaminar fumigant biological-pesticide microbial-pesticide entomopathogenic-fungus entomopathogenic-nematode entomopathogenic-virus baculovirus granulovirus nucleopolyhedrovirus bacteria Bacillus-thuringiensis Bt-toxin crystal-protein cytolytic-protein vegetative-insecticidal-protein mode-of-action gut-paralysis septicemia toxemia antibiotic symbiont antibiotic-resistance fitness-cost resistance-allele resistance-mechanism metabolic-resistance target-site-resistance penetration-resistance behavioral-resistance cross-resistance multiple-resistance resistance-evolution selection-pressure dose-response lethal-concentration lethal-dose median-lethal-concentration median-lethal-dose confidence-interval probit-analysis logit-analysis dose-mortality time-mortality resistance-ratio resistance-factor diagnostic-dose discriminating-dose baseline-susceptibility susceptible-strain reference-strain field-population resistant-population homozygous-resistant heterozygous dominance recessive incomplete-dominance gene-frequency allele-frequency selection-coefficient fitness relative-fitness population-genetics quantitative-genetics heritability genetic-correlation breeding-value genetic-gain marker-assisted-selection genomic-selection transcriptomics proteomics metabolomics genomics functional-genomics comparative-genomics evolutionary-genomics population-genomics phylogenomics bioinformatics computational-biology systems-biology network-analysis pathway-analysis gene-ontology protein-protein-interaction signal-transduction hormone juvenile-hormone ecdysone insulin-like-peptide neuropeptide pheromone sex-pheromone aggregation-pheromone alarm-pheromone trail-pheromone kairomone allomone synomone semiochemical olfaction mechanoreception vision phototaxis chemotaxis anemotaxis thermotaxis hygrotaxis geotaxis kinesis taxis orthokinesis klinokinesis rheotaxis thigmotaxis contact-chemoreception olfactory-receptor gustatory-receptor ionotropic-receptor odorant-binding-protein chemosensory-protein sensillum sensory-neuron antennal-lobe mushroom-body lateral-horn central-complex optical-lobe compound-eye ocellus simple-eye photoreceptor rhabdom ommatidium facet visual-acuity motion-detection polarized-light-detection color-vision ultraviolet-vision infrared-detection thermoreception hygroreception mechanoreceptor chordotonal-organ campaniform-sensillum trichoid-sensillum basiconic-sensillum coeloconic-sensillum placoid-sensillum sensory-ecology foraging-behavior host-location host-acceptance host-suitability oviposition-preference oviposition-deterrence antixenosis antibiosis tolerance resistance-category gall-midge hessian-fly gene-for-gene avirulence-gene resistance-gene R-gene effector-triggered-immunity pattern-triggered-immunity pathogen-associated-molecular-pattern damage-associated-molecular-pattern microbe-associated-molecular-pattern recognition perception defense-response hypersensitive-response cell-death programmed-cell-death autophagy necrosis oxidative-burst reactive-oxygen-species nitric-oxide calcium-signaling mitogen-activated-protein-kinase transcription-factor defense-gene pathogenesis-related-protein antimicrobial-peptide phytoalexin lignin suberin callose papilla cell-wall-reinforcement hypersensitive-cell-death induced-systemic-resistance priming defense-elicitor plant-activator biostimulant plant-strengthener organic-amendment compost manure biochar vermicompost compost-tea microbial-inoculant plant-growth-promoting-rhizobacteria mycorrhizal-fungus endophyte epiphyte phytosphere rhizosphere phyllosphere spermosphere anthroposphere ecosystem-process nutrient-cycling carbon-sequestration nitrogen-fixation phosphorus-solubilization potassium-mobilization micronutrient-availability soil-structure water-infiltration water-holding-capacity erosion-control pollination pest-suppression disease-suppression weed-suppression decomposition detritivory predation herbivory mutualism commensalism competition facilitation inhibition amensalism neutralism symbiosis antagonism trophic-interaction food-chain energy-flow material-cycling biomass-pyramid trophic-pyramid ecological-efficiency production-efficiency assimilation-efficiency ecological-footprint sustainability resilience resistance adaptability transformability socio-ecological-system adaptive-management participatory-research citizen-science knowledge-co-production transdisciplinary-research interdisciplinary-research disciplinary-research basic-research applied-research development-research action-research participatory-action-research farmer-participatory-research on-farm-research station-research controlled-experiment field-experiment observational-study correlational-study manipulative-experiment natural-experiment quasi-experiment longitudinal-study cross-sectional-study case-study comparative-study meta-analysis systematic-review scoping-review rapid-review living-review evidence-synthesis knowledge-synthesis research-synthesis systematic-map bibliometric-analysis citation-analysis co-citation-analysis bibliographic-coupling keyword-analysis topic-modeling science-mapping research-front emerging-topic hot-topic research-gap future-direction research-priority research-agenda strategic-research mission-oriented-research societal-challenge sustainable-development-goal nutrition-security climate-smart-agriculture climate-resilient-agriculture low-input-agriculture regenerative-agriculture agroecology integrated-farming mixed-farming diversified-farming specialized-farming intensive-farming extensive-farming conventional-farming alternative-farming urban-agriculture peri-urban-agriculture rural-agriculture smallholder-agriculture large-scale-agriculture family-farming corporate-farming contract-farming tenant-farming subsistence-farming commercial-farming export-oriented-farming import-substituting-farming local-food-system global-food-system food-value-chain supply-chain value-chain commodity-chain agribusiness agrifood-system food-processing food-distribution food-retail food-service food-consumption food-waste food-loss post-harvest-loss storage-loss processing-loss distribution-loss retail-loss consumer-waste plate-waste kitchen-waste food-rescue food-recovery food-redistribution food-banking composting anaerobic-digestion bioenergy biofuel bioproduct biomaterial biorefinery circular-economy circular-bioeconomy green-economy blue-economy bioeconomy nature-based-solution ecosystem-based-adaptation ecosystem-based-mitigation climate-mitigation disaster-risk-reduction extreme-event flood heat-wave cold-snap frost hail windstorm tornado hurricane typhoon cyclone storm-surge sea-level-rise salinization desertification land-degradation soil-degradation soil-erosion soil-compaction soil-sealing soil-contamination soil-pollution heavy-metal persistent-organic-pollutant pesticide-residue antibiotic-residue hormone-residue plastic-pollution microplastic nanoplastic particulate-matter air-pollution water-pollution groundwater-pollution surface-water-pollution eutrophication hypoxia dead-zone biodiversity-loss habitat-loss habitat-fragmentation habitat-degradation habitat-destruction invasive-species introduced-species alien-species non-native-species exotic-species naturalized-species casual-species escaped-species cultivated-species ornamental-species pet-species livestock poultry aquaculture fishery capture-fishery marine-fishery freshwater-fishery inland-fishery recreational-fishery subsistence-fishery artisanal-fishery industrial-fishery overfishing bycatch discards ghost-fishing fishing-gear fishing-method fishing-effort fishing-capacity fishing-pressure stock-assessment fishery-management marine-protected-area no-take-zone marine-reserve terrestrial-protected-area national-park nature-reserve wildlife-sanctuary biosphere-reserve world-heritage-site ramsar-site important-bird-area key-biodiversity-area ecological-network protected-area-network conservation-corridor wildlife-corridor habitat-corridor stepping-stone buffer-zone transition-zone core-zone management-zone sustainable-use-zone restoration-zone rehabilitation-zone reclamation-zone remediation-zone containment-zone eradication-zone surveillance-zone monitoring-zone quarantine-zone biosecurity phytosanitary veterinary animal-health plant-health one-health ecohealth planetary-health global-health public-health environmental-health occupational-health food-safety feed-safety water-safety air-quality environmental-quality environmental-monitoring environmental-assessment environmental-impact-assessment strategic-environmental-assessment cumulative-impact-assessment social-impact-assessment health-impact-assessment hazard-assessment exposure-assessment dose-response-assessment risk-characterization risk-management risk-communication risk-perception risk-governance precautionary-principle polluter-pays-principle user-pays-principle extended-producer-responsibility corporate-social-responsibility environmental-stewardship sustainable-livelihood sustainable-consumption sustainable-production green-procurement green-supply-chain green-logistics green-manufacturing green-chemistry green-technology clean-technology environmental-technology pollution-prevention waste-minimization resource-efficiency energy-efficiency water-efficiency material-efficiency land-use-efficiency nutrient-use-efficiency nitrogen-use-efficiency phosphorus-use-efficiency potassium-use-efficiency micronutrient-use-efficiency fertilizer-use-efficiency pesticide-use-efficiency water-productivity land-productivity labor-productivity capital-productivity total-factor-productivity partial-factor-productivity agricultural-productivity crop-productivity livestock-productivity fishery-productivity forestry-productivity economic-productivity social-productivity environmental-productivity ecological-productivity biological-productivity primary-productivity secondary-productivity net-primary-productivity gross-primary-productivity ecosystem-productivity landscape-productivity regional-productivity global-productivity carbon-productivity nitrogen-productivity water-footprint carbon-footprint nitrogen-footprint phosphorus-footprint biodiversity-footprint material-footprint land-footprint energy-footprint emission-factor intensity-factor decoupling absolute-decoupling relative-decoupling resource-decoupling impact-decoupling dematerialization rematerialization transmaterialization substitution complementarity trade-off synergy co-benefit win-win lose-lose win-lose zero-sum positive-sum negative-sum game-theory prisoner's-dilemma tragedy-of-the-commons collective-action common-pool-resource public-good private-good club-good common-good global-common global-public-good tipping-point critical-transition regime-shift hysteresis irreversibility path-dependence lock-in technological-lock-in institutional-lock-in behavioral-lock-in social-ecological-trap poverty-trap degradation-trap development-trap complex-adaptive-system social-ecological-system coupled-human-natural-system human-environment-system adaptive-capacity transformative-capacity learning-capacity innovation-capacity governance-capacity institutional-capacity organizational-capacity individual-capacity community-capacity social-capital human-capital natural-capital physical-capital financial-capital built-capital cultural-capital political-capital intellectual-capital knowledge-capital information-capital data-capital digital-capital technological-capital infrastructural-capital institutional-capital organizational-capital network-capital relationship-capital reputational-capital brand-capital symbolic-capital existential-capital spiritual-capital religious-capital moral-capital ethical-capital trust-capital reciprocity-capital solidarity-capital cooperation-capital coordination-capital collaboration-capital communication-capital information-sharing knowledge-sharing learning innovation experimentation transformation transition change evolution development growth progress improvement enhancement optimization maximization minimization satisfaction sufficiency abundance scarcity security insecurity vulnerability robustness stability instability equilibrium disequilibrium homeostasis allostasis stress distress eustress coping coping-capacity coping-strategy coping-mechanism coping-resource coping-support social-support emotional-support instrumental-support informational-support appraisal-support network-support community-support institutional-support policy-support financial-support technical-support technological-support infrastructural-support capacity-building capacity-development capacity-strengthening empowerment enablement mobilization activation engagement participation involvement inclusion exclusion marginalization discrimination inequity inequality disparity gap divide digital-divide knowledge-divide information-divide science-divide research-divide innovation-divide technology-divide development-divide north-south-divide east-west-divide urban-rural-divide gender-divide age-divide class-divide income-divide wealth-divide asset-divide opportunity-divide outcome-divide wellbeing-divide happiness-divide life-satisfaction-divide health-divide education-divide skill-divide competency-divide capability-divide functioning-divide agency-divide freedom-divide choice-divide autonomy-divide self-determination-divide dignity-divide respect-divide recognition-divide status-divide power-divide influence-divide voice-divide representation-divide participation-divide engagement-divide citizenship-divide belonging-divide identity-divide culture-divide value-divide belief-divide norm-divide practice-divide behavior-divide habit-divide custom-divide tradition-divide modernity-divide globalization-divide localization-divide universalism-divide particularism-divide cosmopolitanism-divide nationalism-divide patriotism-divide populism-divide elitism-divide democracy-divide authoritarianism-divide control-divide security-divide liberty-divide privacy-divide surveillance-divide transparency-divide accountability-divide responsibility-divide liability-divide guilt-divide blame-divide punishment-divide reward-divide incentive-divide motivation-divide intention-divide attention-divide perception-divide cognition-divide emotion-divide affect-divide attitude-divide preference-divide interest-divide need-divide want-divide desire-divide demand-divide supply-divide market-divide exchange-divide transaction-divide contract-divide agreement-divide cooperation-divide competition-divide conflict-divide coercion-divide force-divide violence-divide peace-divide harmony-divide coexistence-divide tolerance-divide acceptance-divide rejection-divide opposition-divide resistance-divide conformity-divide deviance-divide normality-divide abnormality-divide illness-divide disease-divide disorder-divide disability-divide impairment-divide handicap-divide capacity-divide potential-divide performance-divide achievement-divide attainment-divide accomplishment-divide success-divide failure-divide excellence-divide mediocrity-divide average-divide standard-divide benchmark-divide threshold-divide ceiling-divide floor-divide limit-divide boundary-divide border-divide frontier-divide horizon-divide perspective-divide viewpoint-divide standpoint-divide position-divide stance-divide approach-divide method-divide technique-divide tool-divide instrument-divide device-divide apparatus-divide equipment-divide machine-divide machinery-divide mechanism-divide system-divide structure-divide organization-divide institution-divide regime-divide order-divide arrangement-divide configuration-divide pattern-divide design-divide plan-divide scheme-divide program-divide project-divide initiative-divide intervention-divide action-divide activity-divide operation-divide process-divide procedure-divide protocol-divide guideline-divide recommendation-divide requirement-divide regulation-divide rule-divide law-divide policy-divide strategy-divide tactic-divide objective-divide goal-divide target-divide aim-divide purpose-divide mission-divide vision-divide principle-divide ethic-divide moral-divide criterion-divide indicator-divide measure-divide metric-divide index-divide scale-divide score-divide rating-divide ranking-divide grade-divide level-divide stage-divide phase-divide step-divide progression-divide sequence-divide series-divide succession-divide chain-divide cycle-divide loop-divide spiral-divide wave-divide pulse-divide rhythm-divide tempo-divide pace-divide speed-divide velocity-divide acceleration-divide deceleration-divide momentum-divide inertia-divide energy-divide work-divide effort-divide strain-divide stress-divide pressure-divide tension-divide compression-divide expansion-divide contraction-divide deformation-divide distortion-divide transformation-divide conversion-divide transduction-divide transmission-divide transfer-divide transport-divide movement-divide motion-divide flow-divide flux-divide current-divide stream-divide river-divide tide-divide wind-divide storm-divide weather-divide climate-divide season-divide year-divide decade-divide century-divide millennium-divide epoch-divide era-divide period-divide time-divide duration-divide interval-divide span-divide stretch-divide spell-divide bout-divide episode-divide event-divide occurrence-divide incident-divide accident-divide coincidence-divide chance-divide luck-divide fortune-divide destiny-divide fate-divide karma-divide dharma-divide tao-divide way-divide path-divide road-divide route-divide course-divide direction-divide orientation-divide alignment-divide positioning-divide placement-divide location-divide site-divide place-divide spot-divide point-divide situation-divide context-divide environment-divide setting-divide scene-divide platform-divide base-divide foundation-divide ground-divide soil-divide earth-divide land-divide terrain-divide territory-divide domain-divide realm-divide sphere-divide world-divide globe-divide planet-divide universe-divide cosmos-divide space-divide void-divide emptiness-divide fullness-divide plenum-divide vacuum-divide atmosphere-divide air-divide gas-divide vapor-divide mist-divide fog-divide cloud-divide rain-divide snow-divide ice-divide frost-divide dew-divide humidity-divide moisture-divide wetness-divide dryness-divide aridity-divide drought-divide flood-divide deluge-divide inundation-divide submersion-divide immersion-divide saturation-divide supersaturation-divide condensation-divide evaporation-divide transpiration-divide sublimation-divide deposition-divide precipitation-divide infiltration-divide percolation-divide seepage-divide leakage-divide discharge-divide runoff-divide erosion-divide sedimentation-divide accretion-divide aggregation-divide coagulation-divide flocculation-divide crystallization-divide solidification-divide liquefaction-divide vaporization-divide boiling-divide melting-divide freezing-divide thawing-divide cooling-divide heating-divide warming-divide thermal-divide temperature-divide heat-divide cold-divide cool-divide warm-divide hot-divide thermodynamic-divide enthalpy-divide entropy-divide free-energy-divide Gibbs-free-energy-divide Helmholtz-free-energy-divide chemical-potential-divide fugacity-divide partial-pressure-divide concentration-divide dilution-divide solution-divide mixture-divide compound-divide element-divide atom-divide molecule-divide ion-divide electron-divide proton-divide neutron-divide nucleus-divide particle-divide quark-divide lepton-divide boson-divide fermion-divide hadron-divide baryon-divide meson-divide photon-divide gluon-divide weak-boson-divide Higgs-boson-divide string-divide brane-divide dimension-divide space-time-divide continuum-divide discrete-divide quantum-divide classical-divide relativistic-divide Newtonian-divide Einsteinian-divide Planckian-divide cosmological-divide astronomical-divide microscopic-divide macroscopic-divide mesoscopic-divide nanoscopic-divide atomic-divide molecular-divide cellular-divide tissue-divide organ-divide organism-divide population-divide community-divide ecosystem-divide biome-divide biosphere-divide ecosphere-divide noosphere-divide technosphere-divide anthroposphere-divide sociosphere-divide politosphere-divide econosphere-divide culturosphere-divide religiosphere-divide ideosphere-divide memesphere-divide infosphere-divide cybersphere-divide digisphere-divide virtual-divide analog-divide real-divide actual-divide possible-divide impossible-divide necessary-divide contingent-divide probable-divide improbable-divide certain-divide uncertain-divide determinate-divide indeterminate-divide definite-divide indefinite-divide specific-divide general-divide particular-divide universal-divide singular-divide plural-divide individual-divide collective-divide social-divide cultural-divide historical-divide political-divide economic-divide religious-divide philosophical-divide scientific-divide technological-divide artistic-divide literary-divide musical-divide visual-divide performing-divide cinematic-divide photographic-divide architectural-divide fashion-divide culinary-divide gastronomic-divide olfactory-divide gustatory-divide tactile-divide kinesthetic-divide proprioceptive-divide vestibular-divide auditory-divide somatosensory-divide interoceptive-divide exteroceptive-divide multisensory-divide crossmodal-divide synesthetic-divide anesthetic-divide esthetic-divide aesthetic-divide poetic-divide rhetorical-divide logical-divide mathematical-divide statistical-divide computational-divide algorithmic-divide heuristic-divide systematic-divide methodical-divide empirical-divide theoretical-divide conceptual-divide abstract-divide concrete-divide practical-divide applied-divide pure-divide fundamental-divide basic-divide strategic-divide tactical-divide operational-divide administrative-divide managerial-divide executive-divide leadership-divide followership-divide membership-divide partnership-divide ownership-divide stewardship-divide guardianship-divide trusteeship-divide custodianship-divide wardship-divide tutelage-divide mentorship-divide sponsorship-divide championship-divide friendship-divide relationship-divide kinship-divide companionship-divide fellowship-divide scholarship-divide apprenticeship-divide internship-divide residency-divide assistantship-divide associateship-divide authorship-divide editorship-divide curatorship-divide directorship-divide governorship-divide presidency-divide chancellorship-divide deanship-divide headship-divide chairmanship-divide
Nomotettix cristatus
crested pygmy grasshopper, crested grouse locust, northern crested grouse locust
Nomotettix cristatus is a small pygmy grasshopper in the family Tetrigidae, commonly known as the crested pygmy grasshopper or crested grouse locust. It is one of approximately 35 Nearctic species of Tetrigidae. The species exhibits three recognized subspecies with distinct geographic distributions across North America. Like other members of its family, it is characterized by an elongated pronotum that extends over the abdomen, a trait distinguishing pygmy grasshoppers from typical grasshoppers in Acrididae.
pygmy-grasshopper groundhopper Tetrigidae Nearctic moist-habitat pronotum cristate North-America leaf-litter aquatic-margin Orthoptera Caelifera jumping-insect small-grasshopper crested-pronotum subspecies-variation Alberta Connecticut Florida Georgia Illinois lake-shore pond-edge stream-margin ground-dwelling Scudder-1862 Batrachidea-cristata Nomotettix-cristatus-cristatus Nomotettix-cristatus-compressus Nomotettix-cristatus-floridanus Tetriginae Acrididea Tetrigoidea Hexapoda Pterygota insect arthropod animal eukaryote metazoa animalia arthropoda insecta nomotettix cristatus crested pygmy grasshopper grouse-locust northern-crested-grouse-locust crested-grouse-locust crested-pygmy-grasshopper iNaturalist GBIF Catalogue-of-Life NCBI-Taxonomy Wikipedia Skejo Tumbrinck Devriese Hochkirch Morse-1895 Hancock-1902 Scudder-1863 Batrachidea cristata compressus floridanus Nearctic-region 35-species 2000-species 230-million-years dinosaur-contemporaries water-dependent-lifecycle Peruća-lake Croatia Balkans Europe systematics taxonomy faunistics natural-history biodiversity conservation endangered imperiled ZooKeys research amateur-naturalist citizen-science observation 53-observations rare neglected-diversity tropics Madagascar Australia New-Guinea Borneo Brazil Atlantic-Forest leaf-like-morphology spines warts undulations horns bizarre grotesque Hancock-1907 Genera-Insectorum Acididae pronotal-projection Holocerus Notocerus Holocerus-lucifer Holocerus-devriesei Notocerus-formidabilis Metopomystrum-muriciense Ophiotettix-pulcherrima Paraselina-brunneri Selivinga-tribulata Criotettix-bispinosus Tetrix-subulata Tettigidea-lateralis Tetrix-undulata Tetrix-tenuicornis Paratettix-meridionalis Paratettix-mexicanus Tetrix-depressa Tetrix-arenosa Tetrix-bipunctata Tetrix-japonica Paratettix-aztecus Hyperyboella-orphania Scelimena-producta Eurymorphopus-bolivariensis Discotettix-belzebuth Tetrix-ceperoi Tetrix-transsylvanica Arulenus Rhacocleis-buchichii Barbitistes-kaltenbachi Adžić Deranja Pavlović Fran-Rebrina Ivan-Budinski Vrlika HPM Hrvatski-Prirodoslovni-Muzej Zagreb Kitonić Miko Franjević Medak Mathieu Silva Pereira De-Domenico Sperber Connors Hendriksen Lambert Chong McMaster Monaghan Rentz Richter Rose Telnov Barclay Pauwels Entomological-Society-of-Latvia Riga Latvia Wallacea Volume-III biogeography nature-conservation Trinity-Audio guest-blog Josip-Skejo Orthoptera-of-Croatia Balkan-endemic endemic-species natural-history-museum museum-collections field-research career-determination serendipitous-discovery new-species systematic-research understudied-group regular-publishing 2021 2017 2020 2014 2016 2013 2012 2011 primary-school high-school biology-student snakes Uropeltidae Scolecophidia shield-tailed-snakes blind-snakes grasshoppers crickets Croatian-endemic first-encounter water-dependency lifecycle leading-European-orthopterists rare-species new-Arulenus-species understudied publishing Tetrigidae-research encyclopedia Elsevier digital-taxonomy open-access innovations journal growth trending social-media Flickr unidentified-rare-pygmy-grasshoppers students amateurs new-records not-rare Tribulation-helmed-groundhopper Kuranda Tully-Range Redlynch Kingfisher-park Speewah selected-awesome-places selected-amazing-taxa rainforests humid-forests extremely-rare weird-looking largest most-colourful ZooKeys-studies photographs identified-to-species-level variability pronotal-projection-morphology holotype Maroantsentra Antongil-Bay Tamatave Andasibe Vohimana Ranomafana Analamazaotra Traveler's-Palm Ravenala-madagascariensis natural-habitat skilful-flier flight large-back-spines Formidable-Pygmy-Grasshopper Sava-region true-colours small interesting Pygmy-unicorns Southern-America Murici description male-holotype head sternum frontal-view dorsal-view lateral-view sternomentum scale-bars Giraffehoppers unique antennal-shape face-coloration visual-animals courtship mating-pair Yapen-Island Cenderawasih-Bay W-New-Guinea young-entomologists decision study-pygmy-grasshoppers direction primary-and-high-school compare-data first-idea familiar-animals field order-Orthoptera friend fellow-student first-systematic-research two-Croatian-endemic-species first-years 2011-2012 never-saw single-pygmy-grasshopper BIOM Sinj Tetrigidae-location around-water first-time-ever lifecycle-water-dependent researching contacting Hendrik-Devriese Axel-Hochkirch Josef-Tumbrinck Croatian-Natural-History-Museum Skejo-et-al.-2014 Skejo-&-Caballero-2016 regularly-publishing Adžić-et-al.-2021 Endangered-Pygmy-Grasshoppers Encyclopaedia-of-Conservation Mathieu-et-al.-2021 ZooKeys-1042 Silva-et-al.-2017 comments recent-changes Metopomystrum Cleostratini Miriatrini ZooKeys-702 Skejo-et-al.-2020 online-social-media Anaselina Paraselina Selivinga rare-Australian-pygmy-grasshoppers ZooKeys-948 Malagasy-devils north south ZooKeys-957 Tumbrinck-&-Skejo-2017 taxonomic-and-biogeographic-revision New-Guinean-genus Ophiotettix-Walker-1871 Metrodorinae Ophiotettigini-trib.-nov.33-new-species nature's-creations encouraged U.S.-species spectacular closed Halloween orange-and-black Argus-tortoise-beetle Chelymorpha-cassidea oleander-caterpillar Syntomeida-epilais wheel-bug Arilus-cristatus morning-glory Convolvulaceae sweet-potato toxic-compounds alkaloids nerve-poisons monarch-butterfly milkweed-leaf-beetle oleander-aphid predator-protection polka-dot-wasp-moth vermillion-wings iridescent-blue-body white-spots wasp-mimicry ultrasonic-song mate-attraction egg-deposition oleander-leaf caterpillars cluster-feeding cardiac-glycosides heart-poisons dogbane-beetles natural-enemies barrel-shaped-eggs pale-orange exoskeleton-hardening orange-black-orange antennae-pale-orange body-and-legs-jet-black rear-end-brilliant-orange stout-beak proboscis impalement digestive-enzymes liquefaction medieval-torture-wheel Hemiptera true-bugs sucking-mouthparts incomplete-metamorphosis egg-nymph-adult plant-feeders harlequin-bugs squash-bugs stink-bugs fierce-predators pest-control Reduviidae assassin-bugs Zelus-longipes milkweed-assassin-bug sticky-forelegs prey-snaring proteolytic-enzymes predigestion muscular-pump black-wing-buds goldenrods meadow-plants Pselliopus-barberi orange-assassin-bug black-stripes stealth leafhopper brown-marmorated-stink-bug Halyomorpha-halys biological-control-agent native-protein-sources autumn-egg-laying spring-hatching May-June red-abdomens orange-antennae sawfly-larvae beetles other-bugs painful-poke caution handling pet-keeping Mad-Magazine Spy-vs-Spy pointy-nosed-secret-agents entomological-equivalent Bug-vs-Bug Heteroptera clan awesome-predator native-to-North-America record-numbers nursery function-of-wheel speculation cautious-approach embrace liquefied-meal protein-for-growth egg-production cluster-size tree-bark magnificent-creatures wide-variety scores stealthy-stalking assassination complementary-outbreak benefit naturally-occurring-predators parasites stem-onslaught no-interest-in-humans retreat memorable-painful-poke firsthand-experience clever-assassin USDA-NIFA-SCRI-Award research-support action-video website predator-prey-relationship natural-enemy ultimate-comeuppance Chinese-praying-mantis yellow-and-black-garden-spider arachnid close-encounter invasive-pest Asia Allentown-PA 1990s millions-of-dollars-damage crops home-invasion 33-states winter-refuge rising-crescendo concern farm-fields landscapes creature's-bane bounty awesome-six-legged-predator medieval-torture-device source-of-speculation little-fact tasty-morsel mark hapless-victim slurp protein-for-development autumn well-fed-female several-to-more-than-one-hundred sycamore-tree attendance irony nocent triggered-outbreak remain-to-be-seen word-of-caution Assasi pet laboratory learned-firsthand try-not-to-handle victim Shrewsbury Martinson Sargent fascination inspiration archive creatures trees UF extension integrated-pest-management NC-State AG295 html tortoise-beetles IFAS orn ole-cpillar UKY Agriculture CritterFiles casefile insects bugs assassin suggested-common-name scientific-name Linnaeus Megha-Kalsi Dakshina-R.-Seal publication preparation happy-safe-Halloween eat-stink-bug part-3 meets-wheel-bug mid-Atlantic-region inimitable-pest apples vegetables soybeans homes discovery damage distress homeowners wicked-winter previous-episodes give-stink-bugs Baby-boomers outwit undo sucking-insects Mother-Nature liquefied-tissues egg-conversion clusters spring bright-red-abdomens lucky half-dozen stealthily-stalked assassinating stinky-marmorated-cousins no-particular-interest stalking-humans too-large taste-bad firsthand beware handling-directly Ashley Nancy Chris Ryan Erik Caroline wrangling death-of-stink-bug Part-2 vandalizing-vegetables sullying-soybeans invading-homes wide-variety-crops curious-reunion arachnid-kind stalking capturing assassinating-stink-bugs game gorgeous-wheel-bug-nymphs cluster-near-eggs venturing-off first-victim true-bug-clan bed-bugs leaf-footed-bugs nefarious-brown-marmorated-stink-bug predicted struggle protect-crops prevent-stinky-invaders overlook eating-mechanism business-end powerful-beak front-legs cautiously-approaches embraces impales pumps-strong-digestive-enzymes liquefy-body-tissues slurps larvae tens-to-more-than-one-hundred plant-hoppers aphids increase-in-populations stink-bug-laden-nursery curious-irony outbreak beneficial-wheel-bugs iNaturalist-taxon rank preferred-common-name observations-count 399 Wikipedia-summary kingdom phylum class order family genus GBIF-taxonomy-match matched-scientific-name canonical-name status accepted match-type exact distribution-records authorship specific-epithet classification eukaryota nomotettix-cristatus authority basionym group grasshoppers-crickets-katydids species instructions fill-all-fields available-knowledge field-cannot-be-supported return-null NOT-repeat-information each-section-focused useful-detail schema taxon-record entomology-guide accurate conservative informative factual-correctness completeness clarity verbosity usefulness CRITICAL-RULES information-not-clearly-supported NOT-infer-species-level-traits higher-taxa explicitly-justified NOT-repeat-same-information multiple-fields unique-non-overlapping-content vague-generalizations like-most-insects typically-feeds-on-plants cautious-language has-been-observed is-known-to NOT-fabricate behaviors diet life-cycle-details host-relationships FIELD-INTENT summary high-level-overview 3-5-sentences appearance physical-description-only identification distinguish-from-similar-taxa habitat environment-conditions distribution geographic-range-only seasonality timing-of-activity feeding-habits null-if-unknown developmental-stages behavior notable-actions-or-habits ecologicalRole role-in-ecosystem humanRelevance interaction-with-humans similarTaxa must-include-reason misconceptions only-if-meaningful extraDetails important-additional-context STYLE-RULES clear-direct-sentences avoid-fluff-filler-language repeating-taxonomy-in-prose overly-technical-jargon concrete-statements abstract-descriptions QUALITY-RULES completeness-high most-fields-well-supported completeness-medium partial-but-reliable completeness-low sparse-data hasInferredContent true-ONLY-if-generalization-used otherwise-false OUTPUT-FORMAT strictly-match-JSON-schema no-extra-fields no-commentary-outside-JSON water-associated three-subspecies N.-c.-cristatus N.-c.-compressus N.-c.-floridanus small-size 399-observations exact-match medium-completeness no-inferred-content factual-correctness-prioritized conservative-approach informative-content no-fluff no-vague-generalizations cautious-language-where-needed no-fabrication unique-field-content focused-sections JSON-schema-compliance no-commentary
Tetropium parvulum
Northern Spruce Borer
Tetropium parvulum, the northern spruce borer, is a small cerambycid beetle specialized on spruce hosts. It was elevated from synonymy with T. cinnamopterum based on distinct morphological differences in larvae and adults, and exhibits strict host specificity to Picea compared to the conifer generalist T. cinnamopterum. The species has a one-year life cycle and is an economically important borer of white and Engelmann spruce logs in western Canada.
Cerambycidae spruce-borer wood-boring-beetle forest-pest western-Canada conifer-specialist Tetropium Picea economic-pest one-year-life-cycle Tetropium-parvulum Northern-Spruce-Borer Tetropium-cinnamopterum-complex morphological-diagnosis strict-host-specificity Tetropium-alaskanum-synonym Rhimphoctona-alaskensis-parasitoid spruce-log-pest Alberta-forestry boreal-forest-insect phloem-feeder cambium-borer L-shaped-galleries early-summer-emergence woodpecker-predation disease-mortality excess-heat-mortality Tetropium-velutinum-comparison lectotype-designation Tetropium-parallelum Tetropium-schwarzianum coniferous-forest-decomposer primary-borer Tetropium-taxonomy Casey-1891 Canadian-Entomologist economic-entomology forest-health timber-pest spruce-regeneration Picea-glauca-pest Picea-engelmannii-pest western-Canadian-distribution North-American-Cerambycidae Tetropium-morphology larval-urogomphi adult-genitalia pronotal-punctation eye-morphology scutellum-structure host-preference-divergence conifer-specialist-vs-generalist sympatric-species species-elevation-from-synonymy type-specimen-examination diagnostic-key forest-insect-natural-enemies insect-disease-mortality parasitoid-wasp woodpecker-insectivory thermal-mortality gallery-depth overwintering-biology spring-pupation September-larval-maturity one-year-univoltine spruce-logging-impact forest-management integrated-pest-management boreal-forest-ecology subcortical-insect phloeophagous-beetle xylophagous-larvae wood-degradation nutrient-cycling forest-succession disturbance-ecology fire-killed-spruce windthrow-spruce salvage-logging-insect standing-dead-timber spruce-beetle-complex Scolytinae-competitor Cerambycidae-community coniferous-wood-borer-guild North-American-forest-insects western-Canadian-entomology Alberta-entomology British-Columbia-entomology Manitoba-entomology Labrador-entomology transcontinental-distribution boreal-forest-biodiversity spruce-ecosystem-engineer tree-killing-insect sublethal-host-stress induced-host-defense insect-fungus-interaction blue-stain-fungus-vector wood-discoloration lumber-degrade veneer-quality-impact pulpwood-quality biomass-energy-feedstock-quality carbon-sequestration-impact climate-change-forest-health warming-thermal-mortality drought-stress-predisposition spruce-decline-agent multi-stressor-forest-dynamics insect-population-regulation natural-enemy-complex trophic-cascade forest-food-web avian-insectivory woodpecker-foraging sapsucker-drilling nuthatch-predation parasitoid-community ichneumonid-wasp braconid-wasp tachinid-fly nematode-parasite bacterial-disease fungal-entomopathogen viral-disease abiotic-mortality-factor competitive-exclusion interspecific-competition resource-partitioning host-tree-chemistry monoterpene-defense phenolic-compound resin-duct-response induced-resistance constitutive-defense spruce-genotype-resistance provenance-variation local-adaptation evolutionary-ecology speciation-mechanism host-race-formation reproductive-isolation morphological-divergence genitalic-evolution allopatric-divergence sympatric-maintenance hybridization-potential introgression-risk conservation-genetics population-structure gene-flow effective-population-size inbreeding-depression genetic-diversity adaptation-potential evolutionary-rescue assisted-migration ex-situ-conservation cryopreservation museum-specimen type-locality holotype lectotype paralectotype syntype original-description taxonomic-revision species-concept biological-species morphological-species ecological-species phylogenetic-species integrative-taxonomy DNA-barcoding COI-sequence nuclear-marker mitochondrial-genome phylogenomics systematics classification-stability nomenclatural-act synonymy homonymy availability validity priority Principle-of-Priority International-Code-of-Zoological-Nomenclature type-fixation type-deposition collection-repository National-Museum-of-Natural-History Canadian-National-Collection Royal-Ontario-Museum University-of-Alberta-collection forestry-research entomological-survey insect-monitoring pheromone-trapping flight-intercept-trap Lindgren-funnel-trap emergence-cage rearing-log dissection-microscope genitalic-preparation microscopic-photography scanning-electron-microscopy confocal-microscopy computed-tomography 3D-reconstruction morphometric-analysis geometric-morphometrics landmark-analysis shape-variation size-allometry sexual-dimorphism intraspecific-variation phenotypic-plasticity environmental-gradient clinal-variation altitudinal-gradient latitudinal-gradient phenology degree-day-model thermal-accumulation development-rate voltinism diapause quiescence cold-hardiness freezing-tolerance supercooling-point cryoprotectant glycerol sorbitol trehalose antifreeze-protein winter-survival spring-emergence adult-longevity oviposition-behavior egg-gallery larval-feeding frass-production gallery-architecture pupal-chamber callus-tissue wound-response tree-defense compartmentalization CODIT-model decay-column heartwood-rot sapwood-stain structural-integrity-loss windthrow-risk stand-dynamics successional-trajectory forest-regeneration plantation-forestry silvicultural-practice thinning pruning fertilization site-preparation genetic-improvement resistance-breeding clonal-forestry seed-orchard provenance-trial common-garden quantitative-genetics heritability selection-differential response-to-selection genetic-gain deployment-strategy risk-spreading portfolio-effect climate-smart-forestry adaptive-silviculture resilience-management transformational-adaptation ecosystem-based-management sustainable-forest-management certification-standard FSC PEFC SFI forest-health-monitoring aerial-survey ground-survey remote-sensing satellite-imagery hyperspectral-imaging LiDAR drone-survey machine-learning species-distribution-model habitat-suitability climate-envelope bioclimatic-variable future-projection emission-scenario RCP SSP uncertainty-quantification ensemble-modeling model-validation independent-data expert-knowledge stakeholder-engagement decision-support management-recommendation policy-implication regulatory-framework phytosanitary-measure quarantine-regulation invasive-species-risk pathway-analysis horizon-scanning early-detection rapid-response eradication-feasibility containment-strategy population-suppression biological-control classical-biological-control augmentative-biological-control conservation-biological-control natural-enemy-importation host-specificity-testing non-target-risk environmental-safety regulatory-approval release-protocol establishment-monitoring spread-tracking population-impact non-target-impact ecosystem-service pollination biological-control-service decomposition habitat-provision food-web-support biodiversity-indicator flagship-species umbrella-species keystone-species ecosystem-engineer foundation-species functional-diversity trait-based-ecology community-assembly environmental-filtering biotic-interaction competition facilitation apparent-competition indirect-effect network-ecology food-web-topology trophic-level energy-flow biomass-pyramid productivity ecosystem-function ecosystem-service-valuation natural-capital green-accounting sustainable-development-goal Aichi-target Kunming-Montreal-framework 30x30-target protected-area OECM conservation-effectiveness management-effectiveness governance indigenous-knowledge local-knowledge citizen-science iNaturalist community-monitoring participatory-research knowledge-co-production transdisciplinary-research science-policy-interface IPBES IPCC FAO ITTO IUFRO CABI CFS USDA USFS NRCan provincial-forestry industrial-forestry smallholder-forestry community-forestry urban-forestry agroforestry social-forestry forest-restoration reforestation afforestation natural-regeneration assisted-natural-regeneration enrichment-planting direct-seeding nursery-practice seedling-quality outplanting-survival competition-control vegetation-management herbicide mechanical-site-preparation prescribed-fire wildfire fire-management fire-ecology pyrodiversity serotiny seed-bank post-fire-regeneration salvage-logging sanitation-cutting insecticide fumigant contact-insecticide systemic-insecticide insect-growth-regulator chitin-synthesis-inhibitor juvenile-hormone-analog ecdysone-agonist semiochemical aggregation-pheromone anti-aggregation-pheromone sex-pheromone kairomone allomone synomone push-pull-strategy mating-disruption mass-trapping attract-and-kill sterile-insect-technique incompatible-insect-technique gene-drive RNA-interference precision-pest-management site-specific-management variable-rate-application unmanned-aerial-vehicle precision-forestry digital-forestry smart-forestry Industry-4.0 Internet-of-Things sensor-network real-time-monitoring big-data artificial-intelligence deep-learning computer-vision automated-identification taxonomic-impediment taxonomic-capacity parataxonomist DNA-taxonomy metabarcoding environmental-DNA biosurveillance biosecurity One-Health planetary-health ecohealth sustainability-science resilience-science complexity-science systems-thinking adaptive-management learning-organization knowledge-management institutional-memory capacity-building education extension outreach science-communication public-engagement media-relations social-media open-access data-sharing FAIR-principles reproducible-research open-science research-integrity research-ethics animal-welfare 3Rs wildlife-research-ethics indigenous-data-sovereignty benefit-sharing access-and-benefit-sharing Nagoya-Protocol Convention-on-Biological-Diversity CITES IUCN Red-List-assessment conservation-status population-trend threat-assessment climate-change-vulnerability adaptive-capacity evolutionary-potential extinction-risk minimum-viable-population effective-conservation systematic-conservation-planning prioritization gap-analysis representativeness complementarity irreplaceability vulnerability connectivity corridor stepping-stone habitat-network landscape-matrix permeability resistance-surface least-cost-path circuit-theory isolation-by-distance isolation-by-resistance population-genetics landscape-genetics seascape-genetics riverscape-genetics conservation-genomics population-genomics landscape-genomics environmental-association genotype-environment-association outlier-detection selective-sweep genomic-offset genomic-vulnerability genetic-rescue genetic-restoration genetic-management pedigree-analysis parentage-analysis relatedness-inference kinship inbreeding-coefficient effective-migration-rate divergence-time coalescent-theory approximate-Bayesian-computation machine-learning-genomics deep-learning-genomics structural-variant copy-number-variant single-nucleotide-polymorphism microsatellite mitochondrial-DNA chloroplast-DNA whole-genome-sequencing transcriptomics proteomics metabolomics phenomics remote-phenotyping high-throughput-phenotyping phenotyping-platform 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Trichonephila clavata
Jorō spider, Joro Spider, Parachute spider
Trichonephila clavata, commonly known as the Jorō spider, is a large orb-weaving spider native to East Asia that has become established as an invasive species in the southeastern United States since approximately 2010. First confirmed in Georgia in 2014, it has expanded rapidly across multiple states through a combination of ballooning dispersal and human-mediated transport. The species is notable for its substantial size, striking coloration, and extensive golden webs, but poses minimal risk to humans due to small fangs and docile behavior. Its physiological adaptations—including higher metabolic rate, faster heart rate, and greater cold tolerance than its congener Trichonephila clavipes—suggest potential for continued northward range expansion.
invasive-species orb-weaver ballooning urban-wildlife biological-control cold-tolerance startle-response cannibalism kleptoparasitism Japanese-folklore Nephilidae Trichonephila spider-silk range-expansion physiological-adaptation urban-ecology agricultural-pest-predator iNaturalist Georgia Maryland East-Asia United-States spider-bite venom docile non-aggressive web-architecture egg-sac overwintering sexual-dimorphism metabolic-rate heart-rate freeze-tolerance road-ecology prey-capture competition native-spider-impacts public-perception media-sensationalism management pesticide-alternatives broom-removal ornamental structural-pest ornithology cardinal web-strength trophic-interactions DNA-metabarcoding diet-analysis species-distribution-model physiological-entomology behavioral-ecology arachnology ballooning-dispersal human-mediated-dispersal hitchhiker cargo-stowaway port-of-entry climate-change warming northward-expansion DMV Howard-County Elkridge Ilchester West-Elkridge beech-forest secondary-forest suburban roadside telephone-pole streetlamp forest-edge disturbance-tolerance prey-detection vibrational-noise auditory-disturbance motionless freezing shyness Hannibal-Lecter stink-bug lanternfly evolutionary-history ancient-association ecosystem-service biocontrol natural-enemy invasive-pest brown-marmorated-stink-bug spotted-lanternfly Halyomorpha-halys Lycorma-delicatula competitive-exclusion dominant-orb-weaver abundant population-growth rapid-spread range-span 120,000-km2 physiological-plasticity metabolism twice-as-high 77%-higher-heart-rate 74%-freeze-survival development-rate seasonal-window life-cycle-completion thermal-limitation congener-comparison natural-dispersal human-transport iNaturalist-record citizen-science confirmed-sighting breeding-population male-and-female gravid-female egg-case stowaway cargo-container port-introduction local-port vehicle-transport hitchhiking accelerated-arrival several-years natural-means prognostication prediction model suitable-habitat abundant-suitable-habitat eastern-North-America western-North-America potential-range invasive-range potential-impact environmental-impact ecological-impact ecosystem-impact monitor cautious evidence-based reasonable-journalism sensationalism exaggeration uncritical-acceptance harmless carefully-monitored next-steps research-opportunity call-for-research knowledge-gap little-known novel-ecosystem spreading rapidly-expanding invasive-predator invasive-orb-weaver non-native introduced established colonized thriving doing-fine happy merely-20-minutes-away road-trip visit access-spread beech-leaf-disease BLD state-forest remarkable-colony amongst-beech-trees confirmed-prognostication Davis-and-Frick escape-relative-warmth expand-range-northward eastern-seaboard merely-20-minutes-away-from-home resist-opportunity amazing-predators tales-of-arrival previous-episodes 2022 2024 reduce-angst large-non-native-spider establishing-in-DMV facts-presented past-episode venomous-terrible-painful Nah expert-Rick-Hoebeke risks-small puny-fangs unlikely-pierce-skin video completely-non-aggressive eastern-Howard-County haphazard-webs littered-with-remains former-victims leaves shed-exoskeletons much-larger-female dwarfs-mate positioned-just-above strand-of-silk spinnerets underside-of-abdomen near-red-mound relative-to-hand large-and-docile wait-and-see what-Jorō-means-to-ecosystems help-other-spiders put-a-beat-down-on-invasive-pests stink-bugs spotted-lanternflies passive-hunters enormous-webs larger-than-a-meter-in-diameter capture-prey-snared-in-silk arachnophobes-scary arachnophiles-beautiful important-ecosystem-services crop-pests native-range Asia having-an-old-friend-over-for-dinner reunite first-time large-spiders juicy-prey-items feathered-and-non-feathered-reptiles poses-no-known-threat difficult-to-predict-impact non-native-species experts-suggest beyond-scary-mien give-indigenous-large-orb-weavers-run-for-their-money black-and-yellow-garden-spider marbled-orb-weaver spotted-orb-weaver other-parts-of-world most-abundant-and-dominant-orb-weaver what-will-it-mean only-time-will-tell final-tidbit Jorō-shapeshifter Jorō-gumo beautiful-woman seduces-men binds-with-silk devours bad-date acknowledgements Rick-Hoebeke identifying-Jorō insights David-Coyle Bob-Bellinger great-images knowledge studies Veni-vidi-vici Nelsen-et-al.Physiological-evaluation Nephila-clavata-newly-recorded Hoebeke-et-al.Life-Cycle-Habitat-and-Variation Moore Startle-responses Davis-and-Anerao How-Urban-Tolerant-Are-They Davis-et-al.inspiration details stars Dave-Clement Miri-Talabac Maddie-Potter hooked-up-with-colony differentiate resources.ipmcenters.org link Jorō-guru David-Coyle's-take YouTube calm-your-fears Journal-of-Economic-Entomology Journal-of-Medical-Entomology chemical-management-strategies effective-management-methods spider-control-products safe-and-effective unlikely-to-bite generally-harmless big-spiders colorful-spiders invasive-spiders in-your-face-spiders webs-on-homes landscape-plants structures middle-of-where-people-live-work-and-play northern-Georgia-2014 presumed-arrived-few-years-prior 15-years increased-dramatically much-of-Southeast disjunct-populations northeastern-U.S.firestorm-of-interest rapidly-expanding-range southeastern-states many-counties Carolinas Tennessee Oklahoma more-than-a-century parts-of-Florida rarely-Pennsylvania tropics thermal-limitations higher-metabolism faster-heart-rate better-ability-to-tolerate-freezing complete-life-cycle-rapidly chilly-temperatures suite-of-adaptations expand-northward warm-loving-cousin findings-support-potential much-yet-to-be-learned successfully-survive-northern-winters nation-and-world-warm southern-species-expand higher-latitudes-and-altitudes rapidly-range-expansion-will-occur typical-mode-of-dispersal aerial-dispersal ballooning-on-strands-of-silk Charlotte's-babies Charlotte's-Web spectacular-moniker parachute-spider rain-down-from-airplanes long-distance-transit human-assistance good-hitchhikers inseminated-gravid-female egg-case-stowaway arrival-in-DMV several-years-by-natural-means new-introduction-at-local-port human-assist-in-vehicles accelerate-arrival terrible-and-painful small-fangs visited-cousin golden-silk-spider Costa-Rica exceeding-meter-diameter indigenous-large-orb-weavers run-for-their-money minimal-direct-impact firestorm Andrew-Davis Benjamin-Frick University-of-Georgia past-week native-to-eastern-Asia Japan China Korea Taiwan known-in-Georgia-since-2013 spreading-rapidly joined-its-cousin Trichonephila-clavipes established-over-160-years better-tolerate-freezing survive-northern-winters rapidly-range-expansion typical-mode long-distance new-introduction human-assist arachnophobes arachnophiles brown-marmorated-stink-bugs difficult-to-predict scary-mien indigenous-orb-weavers Mary-Nouri Sarah-Morgan WAMU relatively-harmless spider-season fall leaves-changing-color cool-crisp-autumn-air pumpkin-spice extension-entomologist Southeast not-just-spiders 20-feet-wide homes people-live-work-and-play 15-years-or-so influx-of-calls public entomology-extension-programs media-frenzy two-questions get-rid-of-Jorō-spiders are-Jorō-spiders-dangerous Clemson-University Southern-Adventist-University Washington-College Union-College two-new-studies test-methods-for-eliminating how-likely-to-bite harm scoured-internet recommended-treatment-options spider-specific-management-recommendations medically-relevant widow-spiders brown-recluses do-not-enter-homes relatively-new recommended-products labelled-spider-control-products tested-products several-products-eliminate not-legal-pesticides machine-lubricant not-recommended commercially-available strongly-encourage labelled-products easiest-and-cheapest without-chemicals broom-or-stick walked-into-spider-web rode-bike took-spider-to-face not-great-feeling wonder-if-bitey give-bitten-superpowers unique-studies evaluated-behaviors react-to-people rather-drop-off-web hang-around-and-bite really-provoke volunteered-to-get-bitten yes-you-read-correctly akin-to-mosquito-bite little-redness little-swelling largely-gone-next-day no-acquired-superpowers take-home-message twofold simplest-and-most-effective without-pesticides feel-you-need-pesticides product-labeled-for-spider-control try-not-to-freak-out doesn't-want-to-be-on-you odds-of-getting-bitten-extremely-low enjoy-glorious-fall-weather drink-pumpkin-spiced-beverage watch-where-you-walk Keep-calm-and-carry-on bites-unlikely cause-minimal-discomfort associate-professor Department-of-Forestry-and-Environmental-Conservation socials email extension featured pest-management Entomology-Today latest-posts email-subscription March-2022 wondered make-its-way-to-DMV September-2022 two-observations three-years 40-sightings happy-and-doing-just-fine several-Howard-County-locations week-or-so-ago team-of-scientists University mission killing-ultra-valuable-beech-trees thriving-amongst-beech-trees visit-amazing-predators mature-females both-Trichonephila-species three-locally-common-orb-weaving-species time-spent-immobile mild-disturbance brief-puff-of-air air-puff-response-data five-other-North-American-species published-literature 453-observations freezing-behavior 10-spider-species remained-immobile-under-a-minute both-Trichonephila-spiders over-an-hour unprecedented shyest-ever-documented tolerate-urban-environments remaining-motionless fleeing nonsexual-cannibalism mating-activity females-cannibalize-males food-availability territorial-aggression Southeastern-United-States expanding-range shy behavioral-reactions descriptive-observations photo-documentation anecdotal-observations controlled-pairings lab natural-webs field one-female-biting-and-killing-another short-fight similar-size container 25-trials fights-ensued-40%different-sizes 27-trials fights-happened-18%larger-females-not-always-aggressor 52-lab-trials six-bouts 9%direct-killing field-trials empty-web 14-trials one-fight 7%aggressor-killing-and-wrapping silk away-from-any-web not-necessarily-territoriality intraspecific-aggression provoked-or-stressed deserves-more-study newly-invasive biology physiology new-range closely-related-species successfully-established 160-years investigation compare online-records iNaturalist.org seasonal-distributions timing females physiological-traits environmental-tolerance metabolic-rates cardiac-functioning cold-exposure survival brief-below-freezing-temperature shorter-season complete-lifecycle narrow-period suitable-weather inherently-higher-metabolism low-temperature survive-better 74%-compared-to-50%brief-freeze colder-climatic-region southeastern-USA useful-information planning urban-landscapes busy-roads frequent-disturbances auditory-and-vibrational physiological-or-behavioral-changes northeast-Georgia varying-levels-of-road-traffic prey-capture-behavior simulated-prey tuning-fork 128-hz-frequency touched-to-web attacked 357-total-trials 20-different-roads attacked-59%high-variability some-roadsides-over-80%others-less-than-30%small-but-significant-negative-correlation daily-road-traffic spider-attack-rates moderate--to-heavy-traffic-roads slightly-less-likely-to-attack 51%-vs-65%able-to-live-near-roads cost-in-terms-of-prey-capture did-not-weigh-less compensate-for-disturbance accumulating-evidence human-dominated-landscapes aid-spread introduced-range Eriostethus-rufus polysphinctine-ectoparasitoid araneid-spiders Neoscona-spp.endemic-to-Japan Yamagata-Prefecture 38º46'-N northern-Japan northernmost-record genus identification-confirmed morphology DNA-barcoding cocoon large-modified-web unique inverted-triangle over-50-cm hanging-from-ill-defined-part no-organized-structure host-spider circumstantial-evidences structure-of-modified-web pre-existing-web-partly-reused orb-web-completely-removed sustaining-threads-newly-moored parasitises-host-spiders two-subfamilies unusual-for-group invasion-potential known-invasive-range review dispersal-abilities potential-impacts management-strategies October-2022 at-least-120,000-km2 South-Carolina North-Carolina Alabama West-Virginia pattern-of-spread natural-dispersal-mechanisms human-mediated-transport large-bodied-orb-weavers flying-insects small-animals thirteen-co-occurring-spider-species monitored-for-competition resources web-building-sites natural-and-urban-habitats management-options-limited advise-journalists-and-experts against-exaggerating-potential-environmental-impact ecologically-harmless rapid-spread-carefully-monitored cautious-evidence-based-approach determining-next-steps three-material-types web-capture introduced-species alter-established-trophic-interactions molecular-analysis resolve-changes community-structure foraging-links east-coast fecal-samples prey-remains-from-webs dissected-guts compare-diet-composition arthropod-targeted-COI-primers Illumina-MiSeq highest-diversity highest-richness highest-proportion-prey-reads recovery-of-prey-reads dissected-gut-content low overwhelmed-by-Jorō-spider-DNA high-proportions-Jorō-spider-reads higher-prey-diversity-and-richness detected-prey-DNA several-days-after-capture initial-gut-retention-time-estimates first-glimpse complexity-of-trophic-associations introduced-web-building-spider viable-material source-of-prey-DNA estimates-of-biodiversity Jammu-and-Kashmir northernmost-shifting document-presence behavior habitat-adaptation colder-temperature-environment Goha-Tehsil Doda-district 1556.74-meters road-bridge nearby-vegetation brightly-colored-abdomens black green yellow red long-legs yellow-and-black-bandings predominantly-seen smaller fewer unique-behaviors remaining-motionless-after-disturbances 5-10-meters-of-roads adaptation-to-human-settlements field-observations image-documentation expert-confirmation thrive-in-colder-conditions 15-20-degrees-Celsius October-November no-spider-found higher-elevations significantly-lower-temperatures adaptation-limited-to-milder-cold-regions new-insight adaptation need-for-further-research non-native-habitats eggs overwinter harsh-weather-conditions no-protection compact-silk-case milky-coating chorionic-microspheres CM fine-structural-characteristics ecological-importance uneven-size-distribution outer-eggmass evenly-covered single-layer diameter-2.3-µm short-papillary-projections segmental-adhesion mucous-components insoluble-in-water partially-soluble-in-absolute-ethanol spherical-structure HFIP strong-organic-solvent not-derived-from-vitellogenic-or-choriogenetic-processes CM-adhesive-coatings ovipositional-process equivalent-to-cocoon-silk protective-functions silken-eggcase Northern-Cardinal Cardinalis-cardinalis perches steals-food southeast-of-the-United-States questions native-fauna-affected consuming-prey-items example native-species-deriving-benefit manner-of-kleptoparasitism perched-directly-on-web supported-its-weight 42-48-g first-documented-case spider-web-supporting-perching-bird measurements force-gauge typical-webs support-masses-up-to-70-g small-but-growing-body-of-knowledge non-native-spider species-distribution-models ecological-niche-modeling predict-areas establish-if-introduced three-distinct-genetic-lineages mitochondrial-COI genome-wide-SNPs Korean-Peninsula