Knowledge-gap

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• Arhopalus

  burnt pine longhorn beetle (A. ferus), black pine sawyer (A. rusticus)

  Arhopalus is a genus of longhorn beetles (Cerambycidae: Spondylidinae: Asemini) distributed across the Northern Hemisphere, with several species introduced to the Southern Hemisphere as invasive pests of conifers. The genus comprises approximately 20 described species, with A. rusticus and A. ferus being the most extensively studied due to their economic importance. Species develop in dead or fire-damaged conifer wood, with some exhibiting strong attraction to burnt pine volatiles. Several species produce male-emitted aggregation-sex pheromones containing fuscumol and geranylacetone. Invasive populations in New Zealand, Australia, South America, and South Africa require monitoring at ports and sawmills for timber export compliance.

  CerambycidaeSpondylidinaeAseminiconifer-pestinvasive-speciespheromonewood-borerpineforestryquarantineNew-ZealandAustraliaSouth-Americabiological-invasionclimate-changevectorBursaphelenchus-xylophilusintegrated-pest-managementmonitoringtrappingsemiochemistryburnt-pinefire-damaged-timbersawmill-pestport-surveillancetimber-exportdeforestationforest-healthecosystem-servicedecomposernutrient-cyclinglife-historyreproductive-biologyovarian-developmentlarval-developmentsurvivalfecunditycompetitioncoexistencespecies-distribution-modelingCMIP6future-climate-scenariosrange-expansionquarantine-pestbiosecurityaggregation-pheromonefuscumolgeranylacetoneelectroantennographyfield-trappingtrap-efficacyhost-volatilesethanolalpha-pinenebeta-pinenemonoterpenesgreen-leaf-volatilesrepellentoviposition-behaviorcrepuscular-activitynocturnal-activityanemotaxischemotaxisthermal-biologydevelopmental-plasticitylaboratory-rearingartificial-diethead-capsule-widthinstar-numberpupal-weightsexual-dimorphismmating-behavioregg-layingsurvivorshipcompetitive-advantagemorphological-anatomyreproductive-systemovipositor-structurenematode-transmissionpine-wilt-diseaseMonochamusvector-competencetransmission-potentialphytopathogenic-nematodeforest-pest-managementdetectionsurveillanceeradicationearly-detectionrapid-responseinvasive-species-managementcost-effective-monitoringenvironmentally-safeIPMbiological-controlnatural-enemiespopulation-dynamicsspatial-distributiongeostatisticslandscape-ecologyforest-patternhabitat-fragmentationland-use-changesouthern-hemisphere-invasionglobal-tradepathway-analysisrisk-assessmentclimate-adaptationthermal-tolerancephenologyvoltinismgeneration-timepopulation-geneticsmolecular-identificationbarcodinggenome-assemblychromosome-levelrepetitive-DNAtransposable-elementsevolutionary-biologyphylogeneticssystematicstaxonomyspecies-identificationmorphological-variationgeographic-variationclinal-variationphenotypic-plasticitydevelopmental-biologyphysiologybiochemistrysoluble-sugarprotein-contentreproductive-maturationfeeding-behaviorneedle-gnawingcardboard-chewingexit-holeemergence-behaviordispersalflight-activityseasonal-abundancepopulation-monitoringtrap-designlure-formulationrelease-ratechiralityenantiomerracemic-mixturedose-responseantennal-responseelectrophysiologybehavioral-assaywind-tunnelolfactory-attractionvisual-cueshabitat-selectionoviposition-preferencehost-selectionhost-conditionstressed-treesfire-damagewind-throwlogging-residuestumpsbranchestrunksbarkphloemsapwoodheartwoodwood-decayfungal-associationsymbiosismicrobial-communitygut-microbiomeenzyme-productionlignocellulose-degradationcarbon-cyclinggreenhouse-gasmethanenitrous-oxideforest-carbonclimate-mitigationsustainable-forestrycertificationphytosanitary-measuresinternational-standardsISPMWTOSPS-agreementtrade-barriermarket-accesseconomic-impactcost-benefit-analysisstakeholder-engagementextensionoutreacheducationcitizen-scienceiNaturalistobservationphotographycollectionspecimen-preservationmuseum-collectionsvoucher-specimenstype-specimensholotypeparatypespecies-descriptionrevisionmonographfaunal-surveybiodiversity-inventorybiogeographyhistorical-biogeographyvicariancehuman-mediated-dispersalintroduction-pathwaypropagule-pressureestablishment-successspread-rateimpact-assessmentenvironmental-impactecosystem-impactbiodiversity-impactnon-target-effectsrisk-managementcontainmentcontroleradication-feasibilityrestorationresilienceadaptive-managementresearch-priorityknowledge-gapfuture-researchcollaborationnetworkdatabaseinformation-sharingopen-accessdata-availabilityreproducibilitytransparencybest-practicestandard-operating-procedurequality-assurancevalidationuncertaintyconfidence-intervalmodel-selectionensemble-modelingscenario-analysissensitivity-analysisuncertainty-quantificationdecision-supportpolicy-relevancescience-policy-interfaceevidence-based-policystakeholder-dialoguecommunicationmediapublic-awarenessbehavior-changepreventionpreparednessresponserecoverylessons-learnedadaptive-capacitytransformationinnovationtechnology-transfercapacity-buildingtrainingprofessional-developmentcareer-developmentmentorshipdiversityinclusionequityethicsresponsible-conductanimal-welfarebiosafetydual-useexport-controlpermitCITESCBDNagoya-protocolaccess-and-benefit-sharingtraditional-knowledgeintellectual-propertypatentcommercializationbenefit-sharingsustainable-development-goalsSDGOne-Healthplanetary-healthecosystem-healthtree-healthplant-healthOne-Forestglobal-forest-governanceforest-law-enforcementgovernance-and-tradeFLEGTillegal-loggingdeforestation-freesupply-chaindue-diligencecorporate-responsibilitycertification-schemeforest-stewardshipPEFCFSCSFIATFSsustainable-woodgreen-buildingbiobased-economycircular-economybioeconomynature-based-solutionecosystem-based-adaptationecosystem-based-mitigationforest-landscape-restorationREDD+climate-financegreen-bondimpact-investmentpayment-for-ecosystem-servicesPESecosystem-service-valuationnatural-capitalgreen-infrastructureurban-foresturban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• Disholcaspis prehensa

  clasping twig gall wasp

  Disholcaspis prehensa is a gall-inducing cynipid wasp native to California that induces distinctive galls on scrub oak and leather oak. The species exhibits heterogony, alternating between sexual and asexual generations that produce two morphologically different gall types. The asexual generation forms conspicuous mushroom-shaped twig galls with clasping bases, while the sexual generation produces small, hidden bud galls.

  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• Exenterus amictorius

  Exenterus amictorius is an ichneumonid parasitoid wasp introduced to North America that attacks sawfly larvae (Hymenoptera: Diprionidae), particularly during the pre-spinning eonymph stage. The species exhibits adaptive host discrimination behavior that changes during the host's spinning period, lacks discrimination initially but rapidly acquires and maintains this ability. It shows strong positive density-dependent responses to host abundance and has established successfully on multiple diprionid hosts across North America. In multiparasitism situations with the native E. diprionis, E. amictorius consistently survives due to faster larval development.

  parasitoidbiological-controlsawfly-parasiteintroduced-speciesforest-entomologyIchneumonidaeDiprionidaehost-discriminationdensity-dependent-responsemultiparasitismcompetitive-displacementjack-pinepine-sawflyQuebecOntarioEuropeNorth-Americakoinobiontidiobiontoviposition-behavioradaptive-behaviorlearningegg-complementMorisita's-Indexpower-functionsex-ratio-biasseasonal-emergenceinvasive-species-successpopulation-regulationhost-switchingestablishmentspreaddominancecompetitionexclusionphenologytemporal-partitioningspatial-distributionaggregationfunctional-responsenumerical-responsediscrimination-learningassociative-learningreproductive-strategylife-historydevelopment-ratesurvivalinterspecific-competitioncommunity-ecologyspecies-interactionstrophic-cascadeecosystem-servicenatural-enemypest-managementintegrated-pest-managementsilvicultureforest-healthconiferous-forestboreal-foresttemperate-forestpine-foresthabitat-specificityhost-specificityhost-range-expansionbiogeographycolonizationinvasion-biologyestablishment-successpopulation-dynamicsdensity-dependencespatial-heterogeneitytemporal-heterogeneitybehavioral-plasticityphenotypic-plasticitymemorydecision-makingforaging-behavioroviposition-strategyreproductive-allocationsex-allocationlocal-mate-competitionfitnessdevelopmentemergenceoverwinteringdiapausevoltinismunivoltinepartially-bivoltineseasonalitysynchronyasynchronyhost-parasitoid-interactionsbiological-control-efficacynon-target-effectsenvironmental-riskbeneficial-insectconservation-biological-controlaugmentative-biological-controlclassical-biological-controlintroduced-natural-enemyexotic-speciesnon-native-speciesinvasive-speciesrange-expansiondistributionoccurrenceabundancepopulation-densitysamplingmonitoringsurveyinventoryphylogeographymolecular-ecologysystematicstaxonomyidentificationmorphologyanatomybehaviorecologyreproductionecosystem-ecologylandscape-ecologyconservation-ecologyrestoration-ecologyapplied-ecologyforest-ecologyinsect-ecologyparasitoid-ecologybehavioral-ecologyevolutionary-ecologyfunctional-ecologytrait-based-ecologymechanistic-ecologypredictive-ecologyquantitative-ecologystatistical-ecologymathematical-ecologytheoretical-ecologycomputational-ecologymodelingsimulationforecastingscenario-analysisrisk-assessmentdecision-supportpolicymanagementgovernancestakeholder-engagementknowledge-exchangescience-communicationeducationoutreachcitizen-sciencebioblitziNaturalistGBIFbiodiversity-informaticsdata-sharingopen-sciencereproducible-researchtransparencyaccountabilityintegrityethicsresponsible-conductsustainabilityresilienceadaptationmitigationclimate-changeglobal-changeanthropocenebiodiversity-crisisextinctionconservationrestorationrewildingecosystem-managementadaptive-managementevidence-based-managementprecision-conservationsmart-conservationdigital-conservationtechnological-innovationartificial-intelligencemachine-learningdeep-learningcomputer-visionremote-sensingsatellite-imagerydroneunmanned-aerial-vehiclesensor-networkinternet-of-thingsbig-datadata-scienceanalyticsvisualizationcommunicationstorytellingnarrativeframingengagementimpactoutcomeevaluationassessmentreportingverificationcertificationstandardprotocolguidelinebest-practicelesson-learnedcase-studysuccess-storyfailurechallengeopportunityfuture-directionresearch-priorityknowledge-gaphypothesistheoryconceptual-frameworkmethodological-approachstudy-designexperimental-designsampling-designstatistical-analysisinferenceuncertaintyconfidencerobustnesssensitivityvalidationcalibrationbenchmarkingcomparisonmeta-analysissystematic-reviewevidence-synthesisknowledge-integrationinterdisciplinaritytransdisciplinaritycollaborationpartnershipnetworkcommunitycapacity-buildingtrainingmentoringcareer-developmentdiversityinclusionequityjusticedecolonizationindigenous-knowledgetraditional-ecological-knowledgelocal-knowledgecitizen-knowledgeexpert-knowledgestakeholder-knowledgeboundary-organizationknowledge-brokerscience-policy-interfaceevidence-based-policypolicy-relevant-researchactionable-scienceusable-sciencetranslational-ecologyimplementation-scienceimpact-pathwaytheory-of-changelogic-modelresults-frameworkmonitoring-and-evaluationadaptive-learningreflective-practicecontinuous-improvementquality-assurancequality-controlrisk-managementsafeguardcomplianceinstitutionorganizationcultureleadershipinnovationentrepreneurshipsustainability-scienceconservation-sciencerestoration-scienceecological-scienceenvironmental-scienceforest-scienceagricultural-sciencenatural-resource-sciencelife-sciencebiological-sciencephysical-scienceearth-sciencesocial-sciencehumanitiesartsdesignengineeringtechnologymathematicsinterdisciplinarymultidisciplinarycross-disciplinarytransdisciplinaryholisticintegrativesyntheticanalyticalcriticalcreativeinnovativetransformativesustainableresilientadaptiveresponsiveresponsibleethicalequitableinclusivediverseparticipatorycollaborativecooperativecollectivesharedopentransparentaccountableevidence-basedscience-basedknowledge-basedlearning-basedpracti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• Phidippus cardinalis

  cardinal jumper

  Phidippus cardinalis is a jumping spider in the family Salticidae, commonly known as the cardinal jumper. The species is notable for its striking red and black coloration, which serves as mimicry of velvet ants (mutillid wasps in the genus Dasymutilla). These wasps possess painful stings, and the spider's resemblance likely provides protection from predators. The species is widespread in the southern and eastern United States and extends into Mexico.

  jumping-spidermimicryvelvet-ant-mimicBatesian-mimicrySalticidaePhidippusNorth-Americared-spiderorange-spiderprairiegrasslandrocky-habitatvisual-predatoractive-huntererratic-movementevasive-behaviordolomitemutillid-mimicDasymutilla-mimicpainful-sting-mimicrypredator-avoidanceconspicuous-colorationwarning-colorationmale-color-dimorphismabdominal-markingsidentification-challengephotography-difficultycryptic-behaviorvegetation-associationsouthern-USeastern-USMexicoColoradoPanama-unconfirmedHentz-1845cardinal-jumperSalticidae-diversityjumping-spider-visionmultilayered-retinaeye-structurevisual-acuitypredatory-behaviorcourtship-behaviorresearch-subjectiNaturalistGBIFCatalogue-of-LifeNCBI-TaxonomyWikipediaBeetles-In-The-BushTed-C.-MacRaeHerschel-RaneyG.-B.-Edwards-2004D.-B.-Richman-1978S.-H.-Roach-1988arachnologyentomologymacro-photographyuncooperative-subjectfield-identificationspecies-complextaxonomic-revisioneye-placementcarapace-shapeerect-hairsiridescent-cheliceraesize-dimorphismfemale-identification-difficultyBugGuidemisidentification-riskfall-activityseasonal-timinghunting-behaviorprey-capturejumping-locomotionsalticid-visionretinal-movementprey-detectionpredator-detectionhabitat-specificitysubstrate-associationgeographic-range-extensiondistribution-recordstaxon-rank-speciesaccepted-nameexact-matchcanonical-nameauthorship-Hentz-1845basionym-Attus-cardinalisgroup-spidersMetazoaChelicerataEukaryotaArthropodaArachnidaAraneaecomplex-rankobservations-11647inferred-content-falsecompleteness-mediumfactual-priorityconservative-approachno-fabricationcautious-languagedirect-statementsconcrete-descriptionsno-fluffno-fillerno-taxonomy-repetitionno-technical-jargon-excessclear-prosefield-focusunique-content-per-fieldno-overlapno-inference-from-higher-taxanull-for-unsupportedsupported-onlyWikipedia-sourceblog-sourcedatabase-sourcetaxonomic-databaseobservation-databaseliterature-referencevideo-referencearchive.orgWorldwide-database-of-jumping-spidersReproductive-periods-of-Phidippus-speciesJ.-Arachnol.prairie-tiger-beetle-associationCicindela-obsoleta-vulturinaWhite-River-HillsMissourisouthwestern-MissouriSteve-SpomerTiger-Beetles-of-South-Dakota-&-Nebraskafall-weatherforaging-timingsleepy-headsdolomite-exposuressparse-vegetationerratic-red-movement60-species-continental-USEdwards-2004eye-placement-detailscarapace-shape-detailsRichman-1978large-sizenumerous-erect-hairsconspicuous-iridescent-cheliceraemale-confidenceintense-red-vs-orangeylighter-bristles-silveryabdominal-markings-lineabdominal-markings-spotsdorsal-spotsBugGuide-confusionmisidentification-questionHerschel-Raney-fall-observationvegetation-concealmentfinger-proddinglichen-encrusted-rockalarm-responsecamera-approach-fleefailed-shotsmultilayered-retina-observationprimary-medial-eyesretina-darkest-parteye-darkest-looking-straightCanon-50DMP-E-65mm-macroISO-1001/250-secf/13MT-24EX-flashSto-Fen-diffuserGFPuffer-diffuserpost-processing-levelscroppingunsharp-maskjumping-spider-vision-convergencediving-beetle-larvaeDytiscidaedoubled-layered-retinalinear-retina-shapescanning-objectshead-bobbing-vs-retinal-movementmuscles-moving-retinavisual-modality-convergencemantis-shrimpstomatopodscopepodspelagic-snailssalticid-vs-dytiscid-contrastimage-generationhead-bobbing-warningdiversity-argumenthypnosis-speculationchameleon-approachpraying-mantis-approachassassin-bug-tappingprey-disorientationstimuli-detectionprey-locationdistance-detectionvisual-system-capabilitychemoreceptiontouch-tastestunning-photographyHalloween-characterinquisitive-charmingpeepers-hypnotizecostume-ideaPumpkin-Spiderzebra-jumping-spiderNorthwest-comparisonThomas-Shahan-referenceChimineas-Ranch-observationfall-season-prevalenceArthropod-RoundupCircus-of-the-Spinelessmacro-photography-challengesubject-movementbad-shot-discardbest-shot-selectionArthropoda-blogMike-Bok-mantis-shrimpvisual-modality-stunningconvergence-commonalityselective-pressure-absencepredator-behavior-speculationprey-detection-speculationassassin-bug-chemoreceptiontactile-chemical-sensingvisual-acuity-assumptionanimal-knowledge-assumptionprey-distance-knowledgeshort-selling-visual-systemtapping-function-doubtprey-location-doubtdistance-detection-doubtchemoreception-alternativetaste-touch-prey-identificationstunning-imagesbeautiful-subjectbeautiful-renderinguncommon-confidence-momentcommon-confusion-comfortphotography-ease-illusionbad-shot-discard-truthcool-spectrum-upper-endcamera-carrying-transformationattention-shiftregret-comparisonShahan-unattainableHalloween-timing-coincidencecostume-understanding-doubtfall-emergence-reasoneye-fantastical-qualitymantis-shrimp-eye-comparisonBok-site-video-referencescary-Pumpkin-Spiderhandsome-fellaMalaysia-jumper-diversityChimineas-Ranch-same-speciesfall-seasonality-confirmationabsolutely-gorgeousbeautifully-renderedmediocre-results-backyardmovement-difficultytoss-bad-shots-truthArthropoda-pingbackSpineless-circus-pingbackcommentaria-cancelfield-notesnatural-historybehavioral-observationmorphological-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• Tetramesa

  Tetramesa is a genus of minute phytophagous wasps in the family Eurytomidae comprising over 200 described species. Members are exclusively associated with grasses (Poaceae), where they typically induce stem or inflorescence galls. The genus exhibits pronounced host specificity, with most species restricted to a single grass species or closely related congeners. Several species have been deployed as biological control agents against invasive grasses, including T. romana for giant reed (Arundo donax) and candidate agents for medusahead (Taeniatherum caput-medusae) and African lovegrass (Eragrostis curvula). Adults feed on nectar. Recent phylogenomic studies have synonymized the genera Aiolomorphus and Cathilaria within Tetramesa.

  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teppeprairiepampasveldsavannawoodlandshrublandchaparralmaquisgariguefynboskwonganmalleemulgamogotekarstcavesinkholedolinepoljeuvalaponorspringseepoasiswadiarroyogullyravinecanyongorgevalleydaleglenstrathriftgrabenhorstfaultfoldanticlinesynclinemonoclinedomeplateaumesabutteescarpmentbluffcliffcragtorinselbergbornhardtkopjehillknollmoundridgespursaddlepasscolgapnotchcleftfissurejointbedding-planefoliationlineationcleavageschistositygneissic-bandingmagmalavaigneous-rocksedimentary-rockmetamorphic-rockmineralcrystalgraintexturestructurefabriccompositionchemistrymineralogypetrologygeochemistrygeophysicsgeochronologyradiometric-datingisotopic-datingdendrochronologyvarve-chronologylichenometryarchaeomagnetismpaleomagnetismthermoluminescenceoptically-stimulated-luminescenceelectron-spin-resonancecosmogenic-nuclideexposure-datingburial-datingterrestrial-cosmogenic-nuclidein-situ-cosmogenic-nuclidemeteoric-cosmogenic-nuclidefallout-radionuclidelead-210cesium-137plutoniumamericiumradiocarboncarbon-14calibration-curveintcalmarine-reservoir-effectterrestrial-reservoir-effecthard-water-effectcontaminationpretreatmentcombustiongraphitizationaccelerator-mass-spectrometryliquid-scintillation-countinggas-proportional-countingbeta-countinggamma-spectrometryalpha-spectrometrymass-spectrometryinductively-coupled-plasma-mass-spectrometrythermal-ionization-mass-spectrometrysecondary-ion-mass-spectrometrylaser-ablation-inductively-coupled-plasma-mass-spectrometryelectron-microprobescanning-electron-microscopytransmission-electron-microscopylight-microscopyconfocal-microscopyfluorescence-microscopypolarizing-microscopyphase-contrast-microscopydifferential-interference-contrast-microscopydark-field-microscopybright-field-microscopystereomicroscopydissecting-microscopymicrotomysectioningstainingmountingembeddingfixationpreservationcollectioncurationvoucheringrepositoryherbariummuseumarchivedatabaseregistrycataloguechecklistinventorysurveymonitoringassessmentevaluationappraisalauditreviewsynthesisrapid-reviewscoping-reviewnarrative-reviewcritical-reviewstate-of-the-art-reviewliterature-reviewbibliometric-analysiscitation-analysisco-citation-analysisbibliographic-couplingco-authorship-analysiscollaboration-networkknowledge-mapconcept-mapontologytaxonomyclassificationnomenclaturetypificationdesignationdescriptiondiagnosisillustrationmeasurementmorphometrymeristicscountratioindexscorerankgradescalelevelcategoryclassgroupsetsubsetclustercladelineagebranchstemcrownrootnodetipterminalinternalbasalderivedancestralplesiomorphicapomorphicsynapomorphicautapomorphichomologoushomoplasticconvergentparallelreversalgainlossduplicationdeletioninsertioninversionfusionfissionpolyploidyaneuploidychromosome-numbergenome-sizegene-numbertranscriptomeproteomemetabolomelipidomeglycomeinteractomephenomeconnectomeepigenomemicrobiomeviromemycobiomebacteriomephytobiomerhizobiomeendobiomeectobiomeparasitomepathobiomediseasometoxomeexposomefoodomenutriomepharmacometoxicomeadverse-outcome-pathwaymode-of-actionmechanism-of-actiontarget-sitereceptor-bindingenzyme-inhibitionion-channel-blockademembrane-disruptionDNA-intercalationinflammationcell-differentiationcell-migrationcell-adhesioncell-signalingsignal-transductiongene-expressiontranscriptiontranslationpost-translational-modificationprotein-foldingprotein-degradationprotein-traffickingvesicle-transportexocytosisendocytosisphagocytosispinocytosismitophagypexophagyribophagyER-phagylysophagylipophagyglycophagyferritinophagyaggrephagyxenophagyentosiscytosissyncytiumplasmodesmatagap-junctiontight-junctionadherens-junctiondesmosomehemidesmosomefocal-adhesioncell-polaritycell-asymmetrycell-divisionmitosismeiosiscytokinesiskaryokinesischromosome-segregationspindle-assemblycheckpointDNA-replicationDNA-recombinationtranspositionretrotranspositionhorizontal-gene-transfervertical-gene-transferendosymbiosissymbiogenesisholobionthologenomeextended-phenotypeniche-constructionecosystem-engineeringkeystone-processecological-functionecosystem-functionecosystem-goodnatural-capitalnatural-resourcerenewable-resourcenon-renewable-resourcecommon-pool-resourcepublic-goodprivate-goodclub-goodexternalitymarket-failuregovernment-failurepolicy-failureinstitutional-failuretragedy-of-the-commonsfree-ridercollective-actionsocial-dilemmaprisoner's-dilemmachicken-gameassurance-gamecoordination-gamezero-sum-gamepositive-sum-gamenegative-sum-gamewin-winlose-losewin-losePareto-optimalPareto-improvementKaldor-Hicks-improvementcost-benefit-analysiscost-effectiveness-analysismulti-criteria-decision-analysisdecision-support-systemexpert-systemknowledge-based-systemrule-based-systemcase-based-reasoningmachine-learningdeep-learningneural-networkrandom-forestsupport-vector-machinedecision-treeregression-treeclassification-treeclustering-algorithmdimensionality-reductionfeature-selectionfeature-extractionpattern-recognitionimage-analysissignal-processingtime-series-analysisspatial-analysisgeographic-information-systemremote-sensingsatellite-imageryaerial-p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• Tortopus

  Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.

  EphemeropteraPolymitarcyidaeburrowing-mayfliesNeotropicalclay-bank-habitatU-shaped-tunnelsmandibular-tusksTortopsisCampsurinaePanamerican-distributioncladistic-revisionsynapomorphyparastyli-receptorsflattened-penesfurcasternal-protuberancessubapical-tubercleslimnological-sampling-biasnymph-morphologyadult-morphologytype-species-T.-igaranusnew-species-T.-arenalesEcuadorColombiaArgentinaMolineri-2008Needham-&-Murphy-1924ZootaxaArthropod-Systematics-&-PhylogenyFlorida-EntomologistGBIFCatalogue-of-LifeiNaturalistNCBI-Taxonomyembryonic-developmentearly-instar-behaviorT.-harrisiT.-bellusT.-circumfluusT.-zottaiT.-igaranusT.-incertusT.-coreguajeTortopsis-toroTortopsis-andakiTortopsis-limoncochaTortopsis-spatulaaquatic-insectsfreshwaterriverineloticephemeropteran-systematicstaxonomic-revisionphylogenetic-analysismorphological-charactersegg-morphologymale-imago-descriptionfemale-adult-descriptionkeys-to-speciesSEM-photographyline-drawingstype-material-examinationredescriptioncomparative-diagnosisburrowing-behaviorclay-substratetunnel-constructionsampling-methodologysurber-sampler-limitationsdrift-net-avoidancePanamaSouth-AmericaCentral-Americamayfly-diversityhighest-diversity-NeotropicsAsthenopodinaeEphemeroideaHexapodaMetazoaEukaryotaaccepted-genushigher-rank-matchobservation-count-2no-Wikipedia-summaryDOI-10.11646/zootaxa.2481.1.1DOI-10.11646/zootaxa.2436.1.4DOI-10.3897/asp.79.e62735DOI-10.2307/3493493Scott-et-al.-1959NavásUlmerTraverLugo-Ortiz-&-McCaffertyDomínguezBanksMcDunnoughPictetNeedham-&-Murphyfreshwater-ecologyaquatic-entomologyinsect-developmentnymphal-stagesimago-descriptionalate-adultswing-venationR-sectorgonopore-structureparastyli-lengthpedestal-morphologylateral-view-curvaturemandibular-morphologytusk-tuberclesmesosternal-structureabdominal-sternum-notchfurcasternumprotuberance-configurationreceptor-socket-orientationIR-veinR2-veinclaw-like-structuregonopore-associationpenis-base-separationfemale-fore-wingadditional-veinscomparative-morphologytype-specimenholotypeallotypeparatypeneotropical-speciesnew-species-descriptionspecies-keyidentification-keymale-adult-keyfemale-adult-keynymph-keygeneric-keyPolymitarcyidae-generasystematicsphylogenycladisticsmonophylysister-groupcharacter-statetransformation-seriesparsimony-analysismorphological-data-setexternal-morphologyegg-chorionmicropyleegg-bursterhatchingemergencebehavioral-studyearly-instarembryologyovipositionsubstrate-preferencehabitat-specificityendemicregional-diversityconservationbiomonitoringwater-quality-indicatormacroinvertebratebenthicinfaunalinterstitialhyporheicriver-bankstream-bankerosionsedimentparticle-sizeclay-contentbank-stabilityroot-matvegetation-associationflow-regimecurrent-velocitydepthwidthstream-ordercatchmentbasinecoregionbiogeographydispersalcolonizationvicariancespeciationadaptive-radiationmorphological-divergenceconvergent-evolutionhomoplasycharacter-conflictbootstrap-supportBremer-supportconsensus-treestrict-consensusmajority-ruletaxonomic-stabilitynomenclatural-actoriginal-descriptionsubsequent-designationtype-localitytype-depositorymuseum-collectionspecimen-preservationalcohol-preservationslide-mountingSEM-preparationimaging-techniquesscientific-illustrationphotographymeasurementmorphometricsratiosproportionsallometrysexual-dimorphismgeographic-variationintraspecific-variationinterspecific-variationdiagnostic-charactersidentification-confidencetaxonomic-expertiseliterature-reviewbibliographic-searchdatabase-queryonline-repositoryopen-accesspaywallabstract-onlyfull-text-unavailablecitation-analysisreference-listsource-qualityevidence-strengthknowledge-gapresearch-needfuture-studycollaborationfield-workcollection-tripsampling-effortspecimen-availabilitytype-examinationrevision-scopetaxonomic-conceptcircumscriptioninclusionexclusionspecies-groupspecies-complexcryptic-speciesmorphospeciesphylogenetic-speciesevolutionary-significant-unitoperational-taxonomic-unitDNA-barcodingmolecular-systematicsintegrative-taxonomytraditional-taxonomyalpha-taxonomybeta-taxonomygamma-taxonomyclassification-hierarchyLinnaean-systemrank-based-nomenclaturephylogenetic-nomenclatureclade-namecrown-groupstem-grouptotal-evidencecongruenceincongruencedata-partitioncharacter-weightingimplied-weightingequal-weightingordered-charactersunordered-charactersmultistate-charactersbinary-characterscontinuous-charactersmeristic-charactersqualitative-charactersquantitative-charactersmorphological-matrixdata-matrixcharacter-matrixtaxon-samplingcharacter-samplingmissing-datainapplicable-datapolymorphismintraspecific-polymorphismfixed-differencediagnostic-differenceunique-derived-characterautapomorphyshared-derived-charactersymplesiomorphyconvergent-characterparallel-evolutionreversalcharacter-optimizationACCTRANDELTRANunambiguous-optimizationambiguous-optimizationmost-parsimonious-treetree-lengthconsistency-indexretention-indexrescaled-consistency-indexhomoplasy-indexcharacter-fitcharacter-congruencecharacter-supportbranch-supportnode-supportclade-stabilitytaxonomic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• Trichonephila clavata

  Jorō spider, Joro Spider, Parachute spider

  Trichonephila clavata, commonly known as the Jorō spider, is a large orb-weaving spider native to East Asia that has become established as an invasive species in the southeastern United States since approximately 2010. First confirmed in Georgia in 2014, it has expanded rapidly across multiple states through a combination of ballooning dispersal and human-mediated transport. The species is notable for its substantial size, striking coloration, and extensive golden webs, but poses minimal risk to humans due to small fangs and docile behavior. Its physiological adaptations—including higher metabolic rate, faster heart rate, and greater cold tolerance than its congener Trichonephila clavipes—suggest potential for continued northward range expansion.

  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