Extinct
Guides
Arhopalus
burnt pine longhorn beetle (A. ferus), black pine sawyer (A. rusticus)
Arhopalus is a genus of longhorn beetles (Cerambycidae: Spondylidinae: Asemini) distributed across the Northern Hemisphere, with several species introduced to the Southern Hemisphere as invasive pests of conifers. The genus comprises approximately 20 described species, with A. rusticus and A. ferus being the most extensively studied due to their economic importance. Species develop in dead or fire-damaged conifer wood, with some exhibiting strong attraction to burnt pine volatiles. Several species produce male-emitted aggregation-sex pheromones containing fuscumol and geranylacetone. Invasive populations in New Zealand, Australia, South America, and South Africa require monitoring at ports and sawmills for timber export compliance.
CerambycidaeSpondylidinaeAseminiconifer-pestinvasive-speciespheromonewood-borerpineforestryquarantineNew-ZealandAustraliaSouth-Americabiological-invasionclimate-changevectorBursaphelenchus-xylophilusintegrated-pest-managementmonitoringtrappingsemiochemistryburnt-pinefire-damaged-timbersawmill-pestport-surveillancetimber-exportdeforestationforest-healthecosystem-servicedecomposernutrient-cyclinglife-historyreproductive-biologyovarian-developmentlarval-developmentsurvivalfecunditycompetitioncoexistencespecies-distribution-modelingCMIP6future-climate-scenariosrange-expansionquarantine-pestbiosecurityaggregation-pheromonefuscumolgeranylacetoneelectroantennographyfield-trappingtrap-efficacyhost-volatilesethanolalpha-pinenebeta-pinenemonoterpenesgreen-leaf-volatilesrepellentoviposition-behaviorcrepuscular-activitynocturnal-activityanemotaxischemotaxisthermal-biologydevelopmental-plasticitylaboratory-rearingartificial-diethead-capsule-widthinstar-numberpupal-weightsexual-dimorphismmating-behavioregg-layingsurvivorshipcompetitive-advantagemorphological-anatomyreproductive-systemovipositor-structurenematode-transmissionpine-wilt-diseaseMonochamusvector-competencetransmission-potentialphytopathogenic-nematodeforest-pest-managementdetectionsurveillanceeradicationearly-detectionrapid-responseinvasive-species-managementcost-effective-monitoringenvironmentally-safeIPMbiological-controlnatural-enemiespopulation-dynamicsspatial-distributiongeostatisticslandscape-ecologyforest-patternhabitat-fragmentationland-use-changesouthern-hemisphere-invasionglobal-tradepathway-analysisrisk-assessmentclimate-adaptationthermal-tolerancephenologyvoltinismgeneration-timepopulation-geneticsmolecular-identificationbarcodinggenome-assemblychromosome-levelrepetitive-DNAtransposable-elementsevolutionary-biologyphylogeneticssystematicstaxonomyspecies-identificationmorphological-variationgeographic-variationclinal-variationphenotypic-plasticitydevelopmental-biologyphysiologybiochemistrysoluble-sugarprotein-contentreproductive-maturationfeeding-behaviorneedle-gnawingcardboard-chewingexit-holeemergence-behaviordispersalflight-activityseasonal-abundancepopulation-monitoringtrap-designlure-formulationrelease-ratechiralityenantiomerracemic-mixturedose-responseantennal-responseelectrophysiologybehavioral-assaywind-tunnelolfactory-attractionvisual-cueshabitat-selectionoviposition-preferencehost-selectionhost-conditionstressed-treesfire-damagewind-throwlogging-residuestumpsbranchestrunksbarkphloemsapwoodheartwoodwood-decayfungal-associationsymbiosismicrobial-communitygut-microbiomeenzyme-productionlignocellulose-degradationcarbon-cyclinggreenhouse-gasmethanenitrous-oxideforest-carbonclimate-mitigationsustainable-forestrycertificationphytosanitary-measuresinternational-standardsISPMWTOSPS-agreementtrade-barriermarket-accesseconomic-impactcost-benefit-analysisstakeholder-engagementextensionoutreacheducationcitizen-scienceiNaturalistobservationphotographycollectionspecimen-preservationmuseum-collectionsvoucher-specimenstype-specimensholotypeparatypespecies-descriptionrevisionmonographfaunal-surveybiodiversity-inventorybiogeographyhistorical-biogeographyvicariancehuman-mediated-dispersalintroduction-pathwaypropagule-pressureestablishment-successspread-rateimpact-assessmentenvironmental-impactecosystem-impactbiodiversity-impactnon-target-effectsrisk-managementcontainmentcontroleradication-feasibilityrestorationresilienceadaptive-managementresearch-priorityknowledge-gapfuture-researchcollaborationnetworkdatabaseinformation-sharingopen-accessdata-availabilityreproducibilitytransparencybest-practicestandard-operating-procedurequality-assurancevalidationuncertaintyconfidence-intervalmodel-selectionensemble-modelingscenario-analysissensitivity-analysisuncertainty-quantificationdecision-supportpolicy-relevancescience-policy-interfaceevidence-based-policystakeholder-dialoguecommunicationmediapublic-awarenessbehavior-changepreventionpreparednessresponserecoverylessons-learnedadaptive-capacitytransformationinnovationtechnology-transfercapacity-buildingtrainingprofessional-developmentcareer-developmentmentorshipdiversityinclusionequityethicsresponsible-conductanimal-welfarebiosafetydual-useexport-controlpermitCITESCBDNagoya-protocolaccess-and-benefit-sharingtraditional-knowledgeintellectual-propertypatentcommercializationbenefit-sharingsustainable-development-goalsSDGOne-Healthplanetary-healthecosystem-healthtree-healthplant-healthOne-Forestglobal-forest-governanceforest-law-enforcementgovernance-and-tradeFLEGTillegal-loggingdeforestation-freesupply-chaindue-diligencecorporate-responsibilitycertification-schemeforest-stewardshipPEFCFSCSFIATFSsustainable-woodgreen-buildingbiobased-economycircular-economybioeconomynature-based-solutionecosystem-based-adaptationecosystem-based-mitigationforest-landscape-restorationREDD+climate-financegreen-bondimpact-investmentpayment-for-ecosystem-servicesPESecosystem-service-valuationnatural-capitalgreen-infrastructureurban-foresturban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opmenttheoretical-developmentconceptual-developmentempirical-developmentexperimental-developmentobservational-developmentexperiential-developmentphenomenological-developmenthermeneutical-developmentcritical-developmentreflexive-developmentreflective-developmentdeliberative-developmentparticipatory-developmentdemocratic-developmentrepresentative-developmentparticipatory-democracydirect-democracyliquid-democracydigital-democracye-democracysmart-democracyintelligent-democracywise-democracymature-democracydeveloped-democracydeveloping-democracyemerging-democracyconsolidated-democracydefective-democracyhybrid-regimeauthoritarian-regimetotalitarian-regimefailed-statefragile-statevulnerable-stateresilient-statestrong-statecapable-stateeffective-statelegitimate-stateaccountable-statetransparent-stateresponsive-stateinclusive-stateparticipatory-statedeliberative-stateadaptive-statelearning-stateinnovative-stateentrepreneurial-stateenabling-stateregulatory-statewelfare-statecompetition-statesecurity-statesurveillance-statepolice-statenanny-stateminimal-statenight-watchman-statemarket-statecorporate-stateplutocratic-stateoligarchic-statearistocratic-statemeritocratic-statetechnocratic-statebureaucratic-stateadministrative-statemanagerial-stateneoliberal-stateneoconservative-stateprogressive-stateconservative-stateliberal-statesocialist-statecommunist-statefascist-statenationalist-statepopulist-stateauthoritarian-statedemocratic-staterepublican-statemonarchic-statetheocratic-statesecular-statemulti-religious-statemulti-ethnic-statemulti-cultural-statemulti-lingual-statefederal-stateunitary-stateconfederal-stateregional-statelocal-stateglobal-stateworld-statecosmopolitan-stateplanetary-stateuniversal-stateexistential-statespiritual-statematerial-stateimmaterial-stateintangible-statetangible-statevisible-stateinvisible-statemeasurable-stateimmeasurable-statequantifiable-statequalitative-statesubjective-stateobjective-stateintersubjective-statecollective-stateindividual-statepersonal-stateprofessional-stateorganizational-stateinstitutional-statesystemic-statestructural-statefunctional-stateprocess-stateprocedural-statemethodological-stateepistemological-stateontological-stateaxiological-statepraxeological-statelogical-staterational-statereasonable-statesensible-statepragmatic-statepractical-statetheoretical-stateconceptual-stateempirical-stateexperimental-stateobservational-stateexperiential-statephenomenological-statehermeneutical-statecritical-statereflexive-statereflective-staterepresentative-statedeliberative-democracy-stateparticipatory-democracy-statedirect-democracy-stateliquid-democracy-statedigital-democracy-statee-democracy-statesmart-democracy-stateintelligent-democracy-statewise-democracy-statemature-democracy-statedeveloped-democracy-statedeveloping-democracy-stateemerging-democracy-stateconsolidated-democracy-statedefective-democracy-statehybrid-regime-stateauthoritarian-regime-statetotalitarian-regime-statefailed-state-statefragile-state-statevulnerable-state-stateresilient-state-statestrong-state-statecapable-state-stateeffective-state-statelegitimate-state-stateaccountable-state-statetransparent-state-stateresponsive-state-stateinclusive-state-stateparticipatory-state-statedeliberative-state-stateadaptive-state-statelearning-state-stateinnovative-state-stateentrepreneurial-state-stateenabling-state-stateregulatory-state-statewelfare-state-statecompetition-state-statesecurity-state-statesurveillance-state-statepolice-state-statenanny-state-stateminimal-state-statenight-watchman-state-statemarket-state-statecorporate-state-stateplutocratic-state-stateoligarchic-state-statearistocratic-state-statemeritocratic-state-statetechnocratic-state-statebureaucratic-state-stateadministrative-state-statemanagerial-state-stateneoliberal-state-stateneoconservative-state-stateprogressive-state-stateconservative-state-stateliberal-state-statesocialist-state-statecommunist-state-statefascist-state-statenationalist-state-statepopulist-state-stateauthoritarian-state-statedemocratic-state-staterepublican-state-statemonarchic-state-statetheocratic-state-statesecular-state-statemulti-religious-state-statemulti-ethnic-state-statemulti-cultural-state-statemulti-lingual-state-statefederal-state-stateunitary-state-stateconfederal-state-stateregional-state-statelocal-state-stateglobal-state-stateworld-state-statecosmopolitan-state-stateplanetary-state-stateuniversal-state-stateexistential-state-statespiritual-state-statematerial-state-stateimmaterial-state-stateintangible-state-statetangible-state-statevisible-state-stateinvisible-state-statemeasurable-state-stateimmeasurable-state-statequantifiable-state-statequalitative-state-statesubjective-state-stateobjective-state-stateintersubjective-state-statecollective-state-stateindividual-state-statepersonal-state-stateprofessional-state-stateorganizational-state-stateinstitutional-state-statesystemic-state-statestructural-state-statefunctional-state-stateprocess-state-stateprocedural-state-statemethodological-state-stateepistemological-state-stateontological-state-stateaxiological-state-statepraxeological-state-statelogical-state-staterational-state-statereasonable-state-statesensible-state-statepragmatic-state-statepractical-state-statetheoretical-state-stateconceptual-state-stateempirical-state-stateexperimental-state-stateobservational-state-stateexperiential-state-statephenomenological-state-statehermeneutical-state-statecritical-state-statereflexive-state-statereflective-state-staterepresentative-state-stateCixius yufengi
Cixius yufengi is a species of planthopper in the family Cixiidae, described by Tsaur in 1993. The species belongs to a genus of small, often inconspicuous insects that feed on plant sap. Very little specific information is available about this particular species.
planthopperCixiidaeCalifornia-endemicTsaur-1993AuchenorrhynchaFulgoromorphasap-feederHemipterainsectarthropodHexapodaAnimaliaEukaryotaDelphacoideaCixiinaeCixiiniCixiusspeciesacceptedGBIFCatalogue-of-LifetaxonomydistributionCaliforniaUSAUnited-StatesNorth-Americaendemicinvertebratehemipterantrue-bugbuginsectaanimalhexapodeukaryotearthropodacixius-yufengiyufengiTsaur1993scientific-namecanonical-nameauthorshiprankstatusgenusspecific-epithetclassificationtaxonomy-matchexactkingdomphylumclassorderfamilydistribution-recordsBuglifeendemic-speciesBritish-endemicsIvell's-Sea-AnemoneEdwardsia-ivelliWidewater-LagoonSussexextinctlikely-extinctnot-seen-in-over-forty-years19731983dance-flyPoecilobothrus-majesticusEssex1907Caledonian-PlanthopperCixius-caledonicusnot-seen-for-70-yearsManx-Shearwater-FleaCeratophyllus-fionnus1960sTurk's-Earth-CentipedeNothogeophilus-turkiIsles-of-ScillyIsle-of-Wight1988never-seen-againconservationJames-Harding-MorrisbookBritish-endemic-invertebratesCraig-MacadamCeltic-WoodlouseMetatrichoniscoides-celticusWaleswestern-fringes-of-England1980sChater's-BristletailDilta-chateriiridescentjumping-powers1990sLundy-Cabbage-Flea-BeetlePsylliodes-luridipennisLundy-IslandDevonco-endemismLundy-Cabbageendemic-plantLundy-Cabbage-WeevilCeutorhynchus-contractus-pallipestaxonomic-uncertaintyHorrid-Ground-weaverNothophantes-horridusPlymouthdevelopmentNorthern-February-Red-StoneflyBrachyptera-putataScotlanddrummingabdomen-tappingBritish-Cave-ShrimpNiphargus-glennieiblindghostly-palecavesdamp-rock-fissurestemporary-puddleshumid-cavesrediscoveredprotectedsurvivalconservation-prioritiesglobal-responsibilityevolutionary-twistsecological-intriguehopenatural-heritageBack-from-the-BrinkRSPBBig-Garden-BirdwatchBSBINew-Year-Plant-Huntplantswildlifenaturecommunicationspublic-engagementcampaignsrare-speciesobscure-speciesoverlooked-speciesirreplaceable-specieslocal-wondersglobal-stakesisolationthousands-of-yearsmillions-of-yearsevolutionlandscapesRed-SquirrelHedgehogEuropeshared-speciesnowhere-elseno-backupno-second-chancescelebrationprotectioncherishrecogniseawarenesshabitat-protectionresearchforgotten-creaturesspotlightslipping-through-the-cracksunknown-to-publicrarely-surveyedbarely-hanging-onalready-goneuncomfortable-truthimportant-speciesleast-knownstrangedeeply-unsettlingsole-global-responsibilitylose-them-everywhereorganisationsfighting-to-changeextraordinary-workrarestmost-threatenedentirely-overlookedforgottenevolvedstep-with-Britain's-landscapesfamiliar-speciesshare-with-Europepopulation-overseasreintroducelose-themwrittenjourneyoverlookedirreplaceablefound-nowhere-else-on-Earthcall-to-recogniseprotectuniquely-oursavailable-nowbooksellersspecies-found-nowhere-else-on-Earthpassionate-nature-enthusiastlifelong-loveexploringnatural-worldtrekkingmountainsrare-flowersscouringfenselusive-mothsinvestigatingexotic-invertebrateshothousesfascinationunwaveringprofessional-lifeconservation-sectorhigh-impact-campaignsinspiredEngland's-rarestmost-obscure-speciesmissionBritain-and-Irelandfall-in-love-with-plantsSHAREFacebookLinkedInguest-blogauthorhow-many-speciesfound-only-in-Britainsimple-questioncomprehensive-listresearchingwriting2022referenceburied-in-booksscattered-across-internettucked-awayminds-of-species-expertsresultover-700-speciesat-least-another-100-subspeciesoccur-nowhere-else-on-Earthtotal-global-responsibilityvery-few-peoplename-even-a-single-onestruckmost-irreplaceable-specieslive-or-diedecisions-made-within-our-borderstop-of-conservation-prioritiescelebratedunderstoodset-outtell-their-storiesunique-invertebratesincredibly-fortunateBuglife's-Conservation-Directorfirst-timecompiling-report20-speciesfive-species-of-flyfour-species-of-beetletwo-stonefliesone-eachwoodlousecentipedemillipedefleabristletailspidershrimpsea-anemonetell-storiesgo-out-and-find-thempicked-fivetrack-downbumped-intocouple-moresearched-under-coastal-rocksexquisitepearly-translucenttiny-speciesbarely-2.5mm-longfirst-discoveredknown-only-from-Walesnearbysearched-dampferny-woodlandsalien-lookingastonishing-jumping-powersnamed-new-to-sciencetravelledDevon's-Lundy-Islandtry-and-seeparticularly-rare-pairingonly-known-exampleendemic-beetlepossibly-endemicspend-their-liveswintry-visitsearchelusive-and-threatenedfound-in-just-a-few-siteswithin-the-cityperpetually-under-pressureowes-its-survivaltireless-effortsguided-tourprime-Northern-February-Red-Stoneflyhabitatblew-my-mindmusical-prowessstoneflies-'drum'tapping-their-abdomensslithering-through-tightmuddyunderground-tunnelspersonal-favouriteutterly-gorgeousspends-its-lifechance-searchDevon-coastfirst-sightingalmost-thirty-yearsones-we've-lostimmediately-drawnonly-ever-knownunfortunatelyCraig's-reporthasn't-been-seenover-forty-yearsfirst-collectedlast-seenwithin-a-decadeknowing-this-species-existedgone-foreversadlynot-uncommon-themeendemic-invertebratesdiscoveredhasn't-been-foundover-a-century70-yearsdon't-think-anyonespottedsince-the-1960snot-long-afterfirst-describedfirst-foundstrange-and-uncomfortable-truthsome-of-the-most-important-speciesalso-some-of-the-least-knownby-definitionBritain's-sole-global-responsibilitylose-them-heredespite-that-significancethankfullyorganisations-fightingraising-awarenessrarest-and-most-threatenedotherwise-remainleading-edge-researchevolved-in-stepBritain's-landscapesthousandsunlike-more-familiar-speciesno-population-overseasnowhere-to-reintroducewhy-I-wrote-Endemicutterly-uniquemosseswoodlicebeetlesbuttercupsstories-full-ofright-attentionactionstill-be-savedheld-onpossiblewithin-our-reachall-good-booksellersoverlooked-and-irreplaceableBacks-Goldilocks-ButtercupHeather-StuckeyAbout-the-Authortrekking-up-mountainsscouring-fensinvestigating-exotic-invertebratesfascination-with-wildlifeRSPB's-Big-Garden-BirdwatchBSBI's-New-Year-Plant-HuntBack-from-the-Brink-projectcare-deeplycurrentlyensure-everyoneopportunitywork-with-BSBISHARE-ONConozoa hyalina
Central Valley Grasshopper, California Central Valley Grasshopper
Conozoa hyalina was a species of grasshopper in the family Acrididae, endemic to the United States. The species is now extinct. It was known from the Central Valley of California, from which its common names were derived. No living observations of this species exist.
Dinokanaga andersoni
Dinokanaga andersoni is an extinct species of scorpionfly (order Mecoptera) described from Eocene-aged deposits. It belongs to the family Dinopanorpidae, a group of extinct mecopterans known from compression fossils. The species was established by Archibald in 2005 based on fossil material. As with other members of its family, it represents part of the diverse insect fauna that inhabited forests during the Eocene epoch.
Dinokanaga dowsonae
Dinokanaga dowsonae is an extinct species of scorpionfly described from Eocene fossil deposits in British Columbia, Canada. It belongs to the family Dinopanorpidae, a group of large-bodied mecopterans known from the Paleogene of North America and Asia. The species was named by paleoentomologist S. Bruce Archibald in 2005 based on well-preserved wing specimens. Like other dinopanorpids, it represents part of the diverse insect fauna that inhabited warm temperate forests during the early Cenozoic.
Dinopanorpidae
Dinopanorpidae is an extinct family of scorpionflies (order Mecoptera) comprising two genera and seven described species. The family was established in 1972 by paleoentomologist Frank Carpenter to accommodate the genus Dinopanorpa, originally described in 1924. These insects are distinguished by a uniquely elongated R1 wing vein that nearly reaches the wing apex—a feature unknown in any other mecopteran lineage. Fossil evidence indicates a restricted temporal and geographic distribution spanning the Paleocene to Eocene in Russia and western North America.
Eurypterida
sea scorpions, eurypterids
Eurypterida is an extinct order of aquatic chelicerate arthropods commonly known as sea scorpions. They ranged from the Cambrian through the Permian periods, with their main diversification occurring during the Paleozoic. The group includes both marine and freshwater species, with body sizes ranging from a few centimeters to over two meters in some species. Eurypterids are characterized by a segmented body with a prosoma and opisthosoma, paired swimming paddles, and a terminal telson spine. They represent one of the largest arthropod groups in the fossil record and are significant for understanding early chelicerate evolution.
Eurypteridae
sea scorpions
Eurypteridae is a family of extinct aquatic arthropods known as sea scorpions, representing some of the largest arthropods to have ever lived. These chelicerates inhabited Paleozoic seas from the Silurian to the Permian periods, approximately 467 to 251 million years ago. Members of this family are characterized by their large size, streamlined bodies, and well-developed swimming paddles. Eurypteridae includes the genus Jaekelopterus, which contains the largest known eurypterid species.
Lepidotrichidae
Lepidotrichidae is a family of basal insects in the order Zygentoma. The family contains the extinct genus Lepidotrix, known from Eocene Baltic amber. The extant genus Tricholepidion, containing a single species from western North America, has traditionally been included here but recent research suggests it belongs to its own family, Tricholepidiidae.
Melanoplus spretus
Rocky Mountain Locust
Melanoplus spretus, the Rocky Mountain Locust, was a species of locust that ranged through the western United States and parts of Canada until the late 19th century. It was one of the most destructive agricultural pests in North American history, forming swarms of unprecedented size—one 1875 swarm was estimated at 198,000 square miles. The species is now extinct, last observed in the early 1900s. It is considered by some authorities to represent a gregarious, migratory phase of Melanoplus sanguinipes rather than a distinct species.
Merostomata
Horseshoe Crabs
Merostomata is a class of chelicerate arthropods containing the extinct Eurypterida (sea scorpions) and the extant Xiphosura (horseshoe crabs). The group is characterized by appendages that function as mouthparts proximally and swimming legs distally, reflected in the etymology from Greek μηρός ('thigh') and στόμα ('mouth'). Phylogenetic status remains debated: some analyses suggest Merostomata is not monophyletic, with Xiphosura basal to a clade comprising Eurypterida and Arachnida, while a 2022 analysis recovered monophyly with Xiphosura in a derived position.
Pentodontini
rhinoceros beetles
Pentodontini is the most diverse tribe within the subfamily Dynastinae (rhinoceros beetles), containing over 100 genera distributed across multiple biogeographic regions. Most genera are restricted to a single biogeographic region. The tribe is characterized by substantial morphological diversity, with generic-level identification often relying on mouthpart morphology in females and secondary sexual characters (horns, claw modifications, antennal club length) in males.
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Protorthoptera is an extinct order of Palaeozoic insects first appearing in the Middle Carboniferous (late Serpukhovian or early Bashkirian). It represents a paraphyletic assemblage of basal neopteran insects rather than a true monophyletic group. The order includes the ancestors of all other polyneopterous insects. Pronotal lobes in some forms were expanded to form a shield-like structure.
Pseudogarypus
Pseudogarypus is a genus of pseudoscorpions in the family Pseudogarypidae, first described by Ellingsen in 1909. The genus includes both extant and extinct species, with at least four species known from Eocene fossils preserved in Baltic amber. Members of this genus are distinguished by distinctive morphological features including horn-like protrusions on the thorax and elongated chelae. The extinct species P. synchrotron was described in 2011 using synchrotron imaging to reveal details obscured by amber inclusions.
Pycnomerus
ironclad beetles
Pycnomerus is a genus of ironclad beetles (family Zopheridae) comprising more than 20 described species. Members are small, elongated beetles with hardened, often sculptured exoskeletons typical of the family. The genus includes both widespread and highly restricted species, with some island endemics now considered extinct due to habitat loss. Several species are associated with dead wood habitats and are classified as saproxylic.
Speyeria adiaste atossa
Atossa fritillary
Speyeria adiaste atossa, the Atossa fritillary, is an extinct subspecies of the unsilvered fritillary butterfly. It was last observed around 1960 in the mountain ranges of southern California. The precise cause of extinction remains unknown, though drought conditions in the late 1950s may have contributed to population decline. The subspecies was first described by William Henry Edwards in 1864. Speyeria is now treated as a subgenus of Argynnis.
Stictocephala stimulea
Stictocephala stimulea is a treehopper species in the family Membracidae, first described by Van Duzee in 1914. Treehoppers in this genus are characterized by their enlarged, often ornate pronotum that extends over the body like a helmet or thorn. The specific epithet 'stimulea' suggests possible resemblance to or association with stinging organisms, though this refers to appearance rather than actual sting. Like other membracids, this species likely feeds on plant phloem sap using piercing-sucking mouthparts.
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Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.
EphemeropteraPolymitarcyidaeburrowing-mayfliesNeotropicalclay-bank-habitatU-shaped-tunnelsmandibular-tusksTortopsisCampsurinaePanamerican-distributioncladistic-revisionsynapomorphyparastyli-receptorsflattened-penesfurcasternal-protuberancessubapical-tubercleslimnological-sampling-biasnymph-morphologyadult-morphologytype-species-T.-igaranusnew-species-T.-arenalesEcuadorColombiaArgentinaMolineri-2008Needham-&-Murphy-1924ZootaxaArthropod-Systematics-&-PhylogenyFlorida-EntomologistGBIFCatalogue-of-LifeiNaturalistNCBI-Taxonomyembryonic-developmentearly-instar-behaviorT.-harrisiT.-bellusT.-circumfluusT.-zottaiT.-igaranusT.-incertusT.-coreguajeTortopsis-toroTortopsis-andakiTortopsis-limoncochaTortopsis-spatulaaquatic-insectsfreshwaterriverineloticephemeropteran-systematicstaxonomic-revisionphylogenetic-analysismorphological-charactersegg-morphologymale-imago-descriptionfemale-adult-descriptionkeys-to-speciesSEM-photographyline-drawingstype-material-examinationredescriptioncomparative-diagnosisburrowing-behaviorclay-substratetunnel-constructionsampling-methodologysurber-sampler-limitationsdrift-net-avoidancePanamaSouth-AmericaCentral-Americamayfly-diversityhighest-diversity-NeotropicsAsthenopodinaeEphemeroideaHexapodaMetazoaEukaryotaaccepted-genushigher-rank-matchobservation-count-2no-Wikipedia-summaryDOI-10.11646/zootaxa.2481.1.1DOI-10.11646/zootaxa.2436.1.4DOI-10.3897/asp.79.e62735DOI-10.2307/3493493Scott-et-al.-1959NavásUlmerTraverLugo-Ortiz-&-McCaffertyDomínguezBanksMcDunnoughPictetNeedham-&-Murphyfreshwater-ecologyaquatic-entomologyinsect-developmentnymphal-stagesimago-descriptionalate-adultswing-venationR-sectorgonopore-structureparastyli-lengthpedestal-morphologylateral-view-curvaturemandibular-morphologytusk-tuberclesmesosternal-structureabdominal-sternum-notchfurcasternumprotuberance-configurationreceptor-socket-orientationIR-veinR2-veinclaw-like-structuregonopore-associationpenis-base-separationfemale-fore-wingadditional-veinscomparative-morphologytype-specimenholotypeallotypeparatypeneotropical-speciesnew-species-descriptionspecies-keyidentification-keymale-adult-keyfemale-adult-keynymph-keygeneric-keyPolymitarcyidae-generasystematicsphylogenycladisticsmonophylysister-groupcharacter-statetransformation-seriesparsimony-analysismorphological-data-setexternal-morphologyegg-chorionmicropyleegg-bursterhatchingemergencebehavioral-studyearly-instarembryologyovipositionsubstrate-preferencehabitat-specificityendemicregional-diversityconservationbiomonitoringwater-quality-indicatormacroinvertebratebenthicinfaunalinterstitialhyporheicriver-bankstream-bankerosionsedimentparticle-sizeclay-contentbank-stabilityroot-matvegetation-associationflow-regimecurrent-velocitydepthwidthstream-ordercatchmentbasinecoregionbiogeographydispersalcolonizationvicariancespeciationadaptive-radiationmorphological-divergenceconvergent-evolutionhomoplasycharacter-conflictbootstrap-supportBremer-supportconsensus-treestrict-consensusmajority-ruletaxonomic-stabilitynomenclatural-actoriginal-descriptionsubsequent-designationtype-localitytype-depositorymuseum-collectionspecimen-preservationalcohol-preservationslide-mountingSEM-preparationimaging-techniquesscientific-illustrationphotographymeasurementmorphometricsratiosproportionsallometrysexual-dimorphismgeographic-variationintraspecific-variationinterspecific-variationdiagnostic-charactersidentification-confidencetaxonomic-expertiseliterature-reviewbibliographic-searchdatabase-queryonline-repositoryopen-accesspaywallabstract-onlyfull-text-unavailablecitation-analysisreference-listsource-qualityevidence-strengthknowledge-gapresearch-needfuture-studycollaborationfield-workcollection-tripsampling-effortspecimen-availabilitytype-examinationrevision-scopetaxonomic-conceptcircumscriptioninclusionexclusionspecies-groupspecies-complexcryptic-speciesmorphospeciesphylogenetic-speciesevolutionary-significant-unitoperational-taxonomic-unitDNA-barcodingmolecular-systematicsintegrative-taxonomytraditional-taxonomyalpha-taxonomybeta-taxonomygamma-taxonomyclassification-hierarchyLinnaean-systemrank-based-nomenclaturephylogenetic-nomenclatureclade-namecrown-groupstem-grouptotal-evidencecongruenceincongruencedata-partitioncharacter-weightingimplied-weightingequal-weightingordered-charactersunordered-charactersmultistate-charactersbinary-characterscontinuous-charactersmeristic-charactersqualitative-charactersquantitative-charactersmorphological-matrixdata-matrixcharacter-matrixtaxon-samplingcharacter-samplingmissing-datainapplicable-datapolymorphismintraspecific-polymorphismfixed-differencediagnostic-differenceunique-derived-characterautapomorphyshared-derived-charactersymplesiomorphyconvergent-characterparallel-evolutionreversalcharacter-optimizationACCTRANDELTRANunambiguous-optimizationambiguous-optimizationmost-parsimonious-treetree-lengthconsistency-indexretention-indexrescaled-consistency-indexhomoplasy-indexcharacter-fitcharacter-congruencecharacter-supportbranch-supportnode-supportclade-stabilitytaxonomic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