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Entedoninae
Entedoninae is a subfamily of chalcid wasps in the family Eulophidae comprising over 90 genera. Members are primarily parasitoids with documented associations across diverse host groups including gall-inducing insects, leafminers, sawflies, thrips, and ants. The subfamily exhibits considerable morphological and ecological diversity, with species found in tropical to temperate habitats worldwide.
parasitoid biological-control gall-associated leafminer-parasitoid Chalcidoidea Hymenoptera Eulophidae Entedoninae tropical-forest agricultural-pest-control sawfly-parasitoid thrips-parasitoid ant-associated rice-pest Cecidomyiidae Tenthredinidae Pontania Closterocerus Ceranisus Afronympha Ametallon Chrysonotomyia Omphale-genus-group canopy understorey ecological-plasticity Bangladesh Panama Africa New-Zealand Europe Senegal South-Africa Democratic-Republic-of-the-Congo Uganda endoparasitoid larval-parasitoid egg-parasitoid host-specificity forest-habitat degraded-habitat mid-altitude-rainforest lowland-rainforest tropical-America classical-biological-control augmentative-biological-control integrated-pest-management rice-hispa Dicladispa-armigera gall-wasp sawfly-gall leafminer ant-symbiosis Formicidae Khaya-senegalensis Ochna-pulchra morphological-taxonomy molecular-taxonomy 28S-rDNA COI CytB new-species-description generic-revision host-range parasitoid-complex food-web ecosystem-service natural-enemy pest-management entomophagous hyperparasitoid inquiline curculionid Apion-minimum Phyllocolpa-leucosticta Chrysocharis Entedon Asecodes Pleurotroppopsis Afrotroppopsis Zaommomentedon Apleurotropis Kratoysma Platocharis Schizocharis Parahorismenus Atullya Tropicharis Chrysochuris cruy mirabilis oryzamyntor eminpashai carinatum dussiae longicaudata machaeriae unimaculata claviger gorgonaense auripunctata galbina laeviscuta phenacapsia cecvora elongata pontaniae heyeri risbeci milletiae newbyi Förster-1856 Chalcidoidea-phylogeny biodiversity species-richness cryptic-species molecular-diagnostics host-plant-association habitat-specificity canopy-fauna understorey-fauna forest-stratification tropical-biodiversity Afrotropical Neotropical Palaearctic Nearctic Oriental Australasian Pacific Holarctic biogeography taxonomic-revision annotated-key catalogue type-specimen museum-collection slide-mounted-specimens morphometrics genitalia wing-venation antennae clypeus mandibles malar-space scape pedicel flagellum gaster vertex suture color-pattern pale-femora dark-gaster sexual-dimorphism male female ovipositor emergence rearing quarantine field-collection foreign-exploration augmentative-release pesticide-resistance sustainable-agriculture food-security rice-production cereal-pest vegetable-pest fruit-pest ornamental-pest forest-pest urban-pest invasive-species native-range introduced-range establishment spread impact non-target-effects host-specificity risk-assessment environmental-safety regulatory-approval research-symposium proceedings peer-reviewed Zootaxa ZooKeys Insect-Systematics-&-Evolution Australian-Journal-of-Entomology Far-Eastern-Entomologist Russian-Entomological-Journal Belgian-Journal-of-Entomology Florida-Entomologist Pan-Pacific-Entomologist Entomological-News Proceedings-of-the-Hawaiian-Entomological-Society Transactions-of-the-American-Entomological-Society Journal-of-Natural-History Zoological-Journal-of-the-Linnean-Society BioControl Journal-of-Economic-Entomology Entomologia-Experimentalis-et-Applicata Environmental-Entomology Oecologia Cladistics Molecular-Phylogenetics-and-Evolution Heredity Canadian-Entomologist Proceedings-of-the-Entomological-Society-of-Washington Annals-of-the-Entomological-Society-of-America Bulletin-of-Entomological-Research Neotropical-Entomology Journal-of-Asia-Pacific-Entomology Turkish-Journal-of-Zoology Polish-Journal-of-Entomology North-Western-Journal-of-Zoology Entomologica-Americana Entomologische-Berichten Sahlbergia Dugesiana Acta-Zoologica-Lilloana Folia-Entomologica-Mexicana Revista-de-la-Sociedad-Entomologica-Argentina Boletin-de-la-Asociacion-Espanola-de-Entomologia Entomologicheskoe-Obozrenie Entomological-Review Zoological-Institute-RAS Proceedings-of-the-Zoological-Institute-of-the-Russian-Academy-of-Sciences Taiwan-Agricultural-Research-Institute Council-of-Agriculture University-of-California Riverside 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sustainable-intensification agroecology conservation-biological-control habitat-management floral-resources refuge buffer-zone landscape-complexity connectivity spillover natural-enemy-diversity food-web-structure trophic-interactions multitrophic indirect-effects trait-mediated density-dependent inverse-density-dependent functional-response numerical-response searching-efficiency handling-time oviposition-behavior host-selection host-acceptance host-suitability host-quality host-stage-preference egg-load progeny-allocation sex-ratio local-mate-competition sib-mating inbreeding fitness longevity fecundity development-time thermal-requirements degree-days phenology synchrony asynchrony seasonal-adaptation diapause quiescence cold-hardiness heat-tolerance desiccation-resistance starvation-resistance dispersal flight walking foraging patch-use area-restricted-search oviposition-marking host-marking superparasitism multiparasitism intraguild-predation hyperparasitism autoparasitism heteronomous-hyperparasitism facultative-hyperparasitism obligate-hyperparasitism primary-parasitoid secondary-parasitoid tertiary-parasitoid mummy aphid-mummy scale-mummy pupal-parasitoid egg-larval-parasitoid larval-pupal-parasitoid idiobiont koinobiont ectoparasitoid solitary gregarious polyembryony thelytoky arrhenotoky diploid haplodiploid chromosome-number genome-size transcriptome proteome metabolome morphome phenome extended-phenotype niche-construction ecosystem-engineering keystone-species ecosystem-service-provider functional-group guild trophic-level energy-flow nutrient-cycling decomposition predation herbivory parasitism mutualism commensalism competition apparent-competition exploitative-competition interference-competition scramble-competition contest-competition resource-partitioning niche-differentiation character-displacement co-evolution arms-race Red-Queen geographic-mosaic local-adaptation phenotypic-plasticity genetic-adaptation epigenetics transgenerational-effects maternal-effects paternal-effects parental-care mating-system sexual-selection sperm-competition cryptic-female-choice mate-choice lek territory aggression alarm defense venom allomone kairomone synomone semiochemical pheromone trail-pheromone alarm-pheromone aggregation-pheromone sex-pheromone marking-pheromone oviposition-deterrent host-marking-pheromone plant-volatile induced-plant-defense systemic-acquired-resistance induced-systemic-resistance priming volatile-organic-compound green-leaf-volatile terpenoid phenylpropanoid jasmonic-acid salicylic-acid ethylene abscisic-acid gibberellin auxin cytokinin brassinosteroid strigolactone nitric-oxide reactive-oxygen-species hypersensitive-response programmed-cell-death autophagy apoptosis necrosis pathogen-associated-molecular-pattern damage-associated-molecular-pattern pattern-recognition-receptor effector-triggered-immunity gene-for-gene avirulence virulence effector toxin phytotoxin mycotoxin bacterial-toxin viral-toxin insecticidal-protein cry-protein vip-protein cyt-protein spore conidium appressorium haustorium mycelium sclerotium fruiting-body basidiocarp ascocarp perithecium cleistothecium pycnidium acervulus sporodochium symptom sign disease-triangle disease-cycle epidemic endemic pandemic outbreak eradication containment management control prevention forecasting warning-system decision-support-system expert-system model simulation economic-threshold economic-injury-level action-threshold treatment-threshold damage-threshold aesthetic-injury-level cultural-control physical-control mechanical-control chemical-control genetic-control sterile-insect-technique incompatible-insect-technique precision-agriculture site-specific-management variable-rate-application remote-sensing geographic-information-system global-positioning-system unmanned-aerial-vehicle drone sensor imaging spectroscopy machine-learning artificial-intelligence deep-learning neural-network computer-vision image-analysis pattern-recognition classification segmentation detection tracking counting sampling sampling-plan sequential-sampling binomial-sampling presence-absence-sampling fixed-precision-sampling variable-precision-sampling spatial-sampling temporal-sampling stratified-sampling cluster-sampling systematic-sampling random-sampling convenience-sampling judgment-sampling expert-sampling quota-sampling snowball-sampling respondent-driven-sampling capture-recapture mark-recapture mark-resight distance-sampling line-transect point-transect strip-transect plot quadrat pitfall-trap Malaise-trap pan-trap water-pan yellow-pan blue-pan white-pan flight-interception-trap window-trap canopy-fogging beating-tray sweep-net aerial-net ground-net aquatic-net D-vac vacuum-sampler aspirator mouth-aspirator mechanical-aspirator berlese-funnel Tullgren-funnel Winkler-extractor pitfall barber-trap pitfall-array drift-fence funnel-trap light-trap UV-light black-light mercury-vapor sodium-vapor LED actinic color-trap sticky-trap pheromone-trap synthetic-pheromone natural-pheromone kairomone-trap food-bait protein-bait sugar-bait fermentation-bait alcohol-bait vinegar-bait molasses-bait fruit-bait meat-bait fish-bait dung-bait carrion-bait decoy lure attractant repellent deterrent antifeedant oviposition-stimulant arrestant suppressant toxicant stomach-poison contact-poison fumigant systemic translaminar residual persistent non-persistent broad-spectrum narrow-spectrum selective non-selective target-specific host-specific mode-of-action nerve-poison acetylcholinesterase-inhibitor GABA-antagonist sodium-channel-modulator nicotinic-acetylcholine-receptor-agonist ryanodine-receptor-modulator chitin-synthesis-inhibitor juvenile-hormone-mimic ecdysone-agonist microbial-pesticide entomopathogenic-nematode entomopathogenic-fungus entomopathogenic-bacterium entomopathogenic-virus entomopathogenic-protozoa Bacillus-thuringiensis Bt Bacillus-sphaericus Paenibacillus-popilliae Serratia-entomophila Metarhizium-anisopliae Beauveria-bassiana Lecanicillium-lecanii Isaria-fumosorosea Nomuraea-rileyi Hirsutella-thompsonii Entomophthora Neozygites Nosema Vairimorpha Thelohania Anticarsia-gemmatalis-nucleopolyhedrovirus AgMNPV Cydia-pomonella-granulovirus CpGV Spodoptera-exigua-multiple-nucleopolyhedrovirus SeMNPV Autographa-californica-multiple-nucleopolyhedrovirus AcMNPV nucleopolyhedrovirus granulovirus cypovirus entomopoxvirus iridovirus parvovirus tetravirus nodavirus reovirus picornavirus iflavirus densovirus birnavirus totivirus partitivirus hypovirulence mycovirus bacteriophage phage transduction conjugation transformation horizontal-gene-transfer vertical-gene-transfer mobile-genetic-element transposon plasmid integron genomic-island pathogenicity-island symbiosis-island metabolic-island resistance-island virulence-factor antibiotic-resistance insecticide-resistance acaricide-resistance herbicide-resistance fungicide-resistance bactericide-resistance nematicide-resistance 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Eustala
humpbacked orb-weavers
Eustala is a genus of orb-weaver spiders in the family Araneidae, first described by Eugène Simon in 1895. As of November 2024, the genus contains 87 species distributed primarily across the Americas, from the United States through Central America to South America, with highest diversity in Brazil, Uruguay, and Argentina. Several species have been documented exhibiting specialized ecological relationships, including myrmecophily (living among ants) in Central American species that inhabit ant-defended acacia plants. The genus includes the well-known humpbacked orb-weaver, Eustala anastera, found across North and Central America.
Araneidae orb-weaver spider humpbacked myrmecophily ant-plant-mutualism Neotropical Nearctic Eustala-anastera Eustala-oblonga Eustala-illicita web-building predator acacia Pseudomyrmex Panama Brazil Galápagos Simon-1895 arachnology entomology tropical-ecology commensalism ontogeny ballooning overwintering juvenile spider-wasp-prey Caliadurgus-hyalinatus host-parasitoid-interactions third-party-exploitation mutualism-exploitation cryptic-coloration nocturnal-foraging diurnal-hiding-behavior thorn-defense orb-web-architecture sawtoothed-web population-ecology substrate-selection spatial-distribution species-diversity endemic-species taxonomic-revision nomen-dubium genitalic-morphology microscopy-identification temperate-North-America Central-America South-America Caribbean Pacific-islands conservation-biology climate-change-vulnerability phenology-disruption winter-activity spring-emergence forest-understory dry-forest moist-forest riparian grassland scrub edge-habitat woodland parkland urban-adapted generalist-predator specialist-predator ant-alate-predation herbivore-interception ecosystem-service biological-control natural-history field-identification cryptic-species-complex sexual-dimorphism wingless-female-moths ballooning-dispersal silk-production web-ontogeny instar-development overwintering-strategy cold-hardiness metabolic-suppression diapause voltinism univoltine bivoltine multivoltine seasonal-polyphenism color-morph pattern-polymorphism habitat-specificity host-plant-specificity ant-colony-integration alarm-pheromone-avoidance sting-avoidance predator-avoidance bird-predation lizard-predation wasp-predation assassin-bug-predation stink-bug-predation defensive-behavior thanatosis camouflage masquerade mimicry aggressive-mimicry foraging-ecology prey-capture web-maintenance web-repair silk-recycling energetics reproductive-output fecundity egg-sac-construction maternal-care male-searching-behavior mate-location pheromone-communication vibratory-communication visual-signaling territoriality web-site-fidelity web-site-tenacity dispersal-behavior colonization-ability range-expansion invasive-potential non-native-species introduced-species synanthropic agricultural-landscape orchard vineyard forest-plantation conservation-status data-deficient IUCN CITES protected-area national-park soberania chiribiquete galapagos floreana ecuador peru bolivia chile argentina uruguay paraguay venezuela colombia costa-rica nicaragua honduras guatemala belize mexico usa canada cuba hispaniola jamaica puerto-rico trinidad tobago barbados lesser-antilles greater-antilles west-indies caribbean-basin amazon-basin orinoco parana pantanal cerrado caatinga atlantic-forest pampas chaco yungas valdivian magellanic fynbos none research-model educational-resource citizen-science inaturalist gbif catalogue-of-life ncbi wikipedia zootaxa journal-of-natural-history psyche entomology-today bug-eric araneology chelicerate-biology arthropod-ecology tropical-biology neotropical-fauna nearctic-fauna holartic pantropical gondwanan biogeography phylogeography molecular-systematics morphological-taxonomy integrative-taxonomy species-concept cryptic-diversity undescribed-species new-species-description redescription neotype-designation lectotype-designation type-locality type-specimen museum-collection voucher-specimen dna-barcoding coi its phylogenomics transcriptomics proteomics metabolomics ecological-genomics adaptation local-adaptation phenotypic-plasticity canalization evolutionary-developmental-biology evo-devo behavioral-ecology sensory-ecology visual-ecology chemical-ecology mechanoreception proprioception neuroethology hunting-behavior web-building-behavior construction-behavior tool-use extended-phenotype niche-construction ecosystem-engineering trophic-cascade food-web energy-flow nutrient-cycling decomposition predator-prey-dynamics population-dynamics metapopulation source-sink-dynamics habitat-fragmentation edge-effects corridors connectivity landscape-ecology macroecology biogeochemistry climate-envelope species-distribution-modeling maxent bioclim worldclim remote-sensing gis spatial-analysis geostatistics abundance density biomass productivity diversity-index richness evenness dominance rarity endemism beta-diversity gamma-diversity alpha-diversity turnover nestedness community-assembly neutral-theory niche-theory competition facilitation apparent-competition indirect-effects interaction-networks mutualism parasitism predation herbivory detritivory scavenging cannibalism intraspecific-competition interspecific-competition character-displacement resource-partitioning niche-differentiation coexistence stability resilience resistance recovery succession disturbance perturbation extinction colonization invasion establishment spread impact risk-assessment management control eradication restoration reintroduction rewilding conservation-breeding ex-situ in-situ protected-area-network gap-analysis systematic-conservation-planning prioritization hotspot coldspot majority minority flagship umbrella keystone foundation ecosystem-engineer indicator sentinel bioindicator biomonitor ecosystem-health environmental-quality pollution pesticide heavy-metal contaminant bioaccumulation biomagnification toxicology risk hazard exposure dose-response no-observed-effect-concentration lowest-observed-effect-concentration median-lethal-concentration median-effective-concentration safety-factor uncertainty-factor probabilistic-risk-assessment life-cycle-assessment carbon-footprint water-footprint ecological-footprint sustainability sustainable-development green-economy circular-economy ecosystem-services natural-capital green-infrastructure nature-based-solutions ecological-restoration habitat-restoration species-recovery population-recovery genetic-rescue outbreeding-depression inbreeding-depression genetic-diversity heterozygosity effective-population-size bottleneck founder-effect genetic-drift 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emerging-infectious-disease pandemic epidemic endemic sporadic outbreak cluster index-case contact-tracing quarantine isolation social-distancing masking vaccination herd-immunity reproduction-number basic-reproduction-number effective-reproduction-number doubling-time generation-time serial-interval incubation-period latent-period infectious-period clinical-period pre-symptomatic asymptomatic pauci-symptomatic mild moderate severe critical fatal case-fatality-rate infection-fatality-rate attack-rate secondary-attack-rate excess-mortality years-of-life-lost disability-adjusted-life-year quality-adjusted-life-year burden-of-disease global-burden-of-disease one-health planetary-health ecohealth conservation-medicine veterinary-medicine human-medicine comparative-medicine translational-medicine precision-medicine personalized-medicine genomic-medicine regenerative-medicine stem-cell tissue-engineering organoid organ-on-chip microphysiological-system in-silico in-vitro ex-vivo in-vivo animal-model non-human-primate rodent mouse rat guinea-pig rabbit dog cat livestock poultry aquaculture fishery wild-capture mariculture algaculture seaweed-farming integrated-multi-trophic-aquaculture recirculating-aquaculture-system aquaponics hydroponics aeroponics vertical-farming controlled-environment-agriculture precision-agriculture smart-farming digital-agriculture agriculture-4.0 internet-of-things artificial-intelligence machine-learning deep-learning neural-network computer-vision drone unmanned-aerial-vehicle satellite gps gnss geographic-information-system decision-support-system expert-system knowledge-based-system ontology semantic-web linked-data open-data big-data data-mining data-science data-analytics predictive-analytics prescriptive-analytics descriptive-analytics diagnostic-analytics business-intelligence artificial-intelligence-ethics algorithmic-bias fairness accountability transparency explainability interpretability privacy security safety reliability robustness adaptability flexibility scalability efficiency effectiveness efficacy performance profitability cost-benefit cost-effectiveness economic-analysis financial-analysis investment return-on-investment net-present-value internal-rate-of-return payback-period break-even sensitivity-analysis scenario-analysis monte-carlo-simulation decision-tree real-options game-theory behavioral-economics nudge choice-architecture default framing anchoring availability-heuristic representativeness-heuristic affect-heuristic confirmation-bias availability-bias anchoring-bias framing-bias loss-aversion endowment-effect status-quo-bias sunk-cost-fallacy planning-fallacy optimism-bias pessimism-bias overconfidence underconfidence dunning-kruger-effect imposter-syndrome stereotyping prejudice discrimination implicit-bias explicit-bias structural-bias systemic-bias institutional-bias interpersonal-bias intrapersonal-bias unconscious-bias conscious-bias affinity-bias similarity-bias in-group-bias out-group-bias halo-effect horns-effect contrast-effect recency-effect primacy-effect serial-position-effect mere-exposure-effect familiarity-principle propinquity-effect social-proof authority scarcity reciprocity commitment consistency liking unity influence persuasion compliance conformity obedience empowerment agency autonomy self-determination motivation intrinsic-motivation extrinsic-motivation self-efficacy self-esteem self-concept self-identity social-identity collective-identity cultural-identity national-identity gender-identity sexual-orientation religion spirituality values beliefs attitudes norms customs traditions rituals ceremonies celebrations festivals holidays commemorations memorials monuments heritage legacy history historiography archaeology paleontology anthropology sociology psychology cognitive-science neuroscience linguistics philosophy ethics morality justice equity equality inclusion diversity belonging representation participation engagement 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eukaryome phytobiome rhizobiome endobiome ectobiome exo-biome astrobiome planetary-biome earth-system climate-system ocean-system terrestrial-system freshwater-system atmospheric-system cryospheric-system anthroposphere technosphere noosphere biosphere hydrosphere lithosphere atmosphere magnetosphere heliosphere cosmosphere universe multiverse dimension space time spacetime relativity quantum-mechanics string-theory m-theory loop-quantum-gravity cosmology astronomy astrophysics planetary-science earth-science geology geophysics geochemistry geomorphology soil-science pedology edaphology hydrology limnology oceanography meteorology climatology paleoclimatology stratigraphy sedimentology mineralogy petrology volcanology seismology tectonics plate-tectonics continental-drift orogeny epeirogeny isostasy erosion weathering deposition transport sedimentation lithification diagenesis metamorphism melting crystallization igneous sedimentary metamorphic rock-cycle water-cycle carbon-cycle nitrogen-cycle phosphorus-cycle sulfur-cycle oxygen-cycle hydrogen-cycle biogeochemical-cycle nutrient-cycle trophic-pyramid food-chain ecological-network interaction-web mutualistic-network antagonistic-network competition-network facilitation-network ecosystem-function ecosystem-process provisioning-service regulating-service cultural-service supporting-service habitat-service refuge nursery corridor stepping-stone buffer-zone core-area periphery matrix patch landscape seascape riverscape skyscape soundscape smellscape tastescape touchscape mindscape dreamscape memoryscape heritagescape futurescape possibility-space scenario-space decision-space policy-space governance-space institutional-space social-space cultural-space economic-space political-space legal-space ethical-space moral-space spiritual-space sacred-space profane-space liminal-space transitional-space threshold boundary edge frontier margin center core hinterland heartland homeland motherland fatherland native-land ancestral-land promised-land wilderness pioneer settler colonist immigrant emigrant refugee asylum-seeker displaced-person stateless-person nomad traveler tourist pilgrim explorer adventurer discoverer inventor innovator creator maker builder craftsman artisan artist designer architect engineer scientist scholar academic researcher investigator detective journalist reporter correspondent chronicler historian biographer memoirist autobiographer diarist letter-writer poet novelist playwright screenwriter essayist critic reviewer editor publisher librarian archivist curator conservator restorer preservationist conservationist environmentalist ecologist naturalist botanist zoologist entomologist arachnologist herpetologist ichthyologist ornithologist mammalogist primatologist anthropologist archaeologist paleontologist geologist geographer cartographer surveyor navigator expedition voyage journey travel tour trip visit stay residence habitation dwelling shelter home house building 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wellness fitness soundness integrity wholeness completeness perfection excellence quality standard criterion benchmark yardstick measure gauge sign signal symptom mark token symbol image icon emblem logo brand trademark copyright patent intellectual-property proprietary confidential secret classified restricted limited controlled regulated managed governed administered operated maintained sustained supported funded financed resourced staffed equipped furnished supplied provided delivered distributed allocated assigned designated appointed elected selected chosen picked preferred favored liked loved adored cherished treasured valued esteemed respected honored celebrated commemorated remembered recalled recollected reminisced reflected contemplated meditated pondered considered weighed evaluated judged assessed appraised rated ranked graded scored marked pointed noted observed noticed seen viewed looked watched gazed stared glanced peeked peered scanned surveyed examined inspected 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experiment investigation inquiry query question problem issue matter subject topic theme motif leitmotif thread strand line sequence series progression continuation extension expansion enlargement evolution change transformation conversion transition shift alteration modification adjustment accommodation reconciliation harmonization synchronization coordination integration unification combination merger fusion blend mixture compound composite alloy amalgam incorporation assimilation absorption digestion metabolism catabolism anabolism equilibrium balance steadiness constancy permanence endurance persistence continuity duration longevity lifespan lifetime existence being entity object thing item article piece part portion segment section division component element constituent ingredient factor aspect facet feature characteristic property attribute trait indication evidence proof verification validation confirmation corroboration substantiation authentication certification accreditation licensing permitting authorizing empowering enabling allowing letting tolerating accepting receiving taking getting obtaining acquiring gaining earning winning achieving attaining reaching arriving coming approaching nearing closing converging meeting joining connecting linking binding tying fastening attaching affixing appending adding supplementing complementing completing finishing perfecting consummating fulfilling satisfying gratifying pleasing delighting enjoying relishing savoring appreciating valuing prizing treasuring cherishing nurturing cultivating fostering promoting advancing furthering encouraging supporting backing assisting aiding helping serving attending tending caring nursing healing curing treating remedying correcting rectifying repairing fixing mending restoring renewing reviving resuscitating reanimating reawakening rekindling reigniting relaunching restarting rebooting refreshing rejuvenating reinvigorating reenergizing recharging refueling replenishing 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Lygus hesperus
Western Tarnished Plant Bug
Lygus hesperus is a significant agricultural pest in western North America, causing extensive damage to cotton, strawberries, alfalfa seed, and other crops. In California alone, annual losses exceed $30 million in cotton and $40 million in strawberries. Adults overwinter in reproductive diapause triggered by short day lengths, resuming activity when conditions improve. The species has been the subject of extensive research on sampling methods, biological control, and insecticide resistance.
agricultural-pest plant-bug Miridae cotton-pest strawberry-pest alfalfa-pest western-North-America diapause biological-control integrated-pest-management insecticide-resistance mark-release-recapture salivary-enzymes extraoral-digestion Peristenus-relictus Hippodamia-convergens drone-sampling fluorescent-marking SmartWater vector Pantoea-ananatis Serratia potassium-chloride deterrent artificial-diet lentil seed-crops California Texas Arizona Nevada Washington Mexico polyphagous phytophagous stylet-probing scutellum nymph-spots economic-entomology organic-agriculture ecosystem-services bird-predation molecular-gut-analysis ELISA immunomarking parasitoid population-suppression crop-protection sampling-protocols sweep-net drop-cloth Y-tube-olfactometer gustatory-deterrent visual-cues volatile-cues reproductive-diapause postdiapause oogenesis oviposition longevity body-mass life-history mass-spectrometry salivary-proteins polygalacturonase α-amylase protease laccase glucose-dehydrogenase xanthine-dehydrogenase plant-defense host-pathogen-interaction iDrone-Bee UAV unmanned-aerial-vehicle open-source field-scouting IPM economic-threshold Bt-cotton natural-enemy conservation-biological-control landscape-ecology habitat-management weed-management border-sprays spot-treatment selective-insecticide neonicotinoid pyrethroid resistance-management sustainable-agriculture food-security economic-impact simulation-model population-dynamics immigration parasitism averted-loss value-loss cost-benefit research-methodology insect-marking dispersal fluorophore autofluorescence survivorship mortality mark-transfer crowding olfactometer feeding-behavior no-choice-assay choice-assay gustation host-selection plant-quality succulent-growth fruiting-structures square boll bloom terminal defoliation stand-loss dirty-bloom puckered-petals lesion carpel-wall lint-damage seed-damage yield-loss fiber-quality management-threshold scouting-interval action-threshold treatment-timing insecticide-efficacy labeled-rate difficult-to-control species-complex identification-key morphology color-variation wing-development antennae thorax abdomen instar developmental-stage laboratory-rearing oligidic-diet semisolid-slurry chicken-egg soybean wheat-germ lima-bean cost-reduction biomass-production survival fecundity egg-production generation-time population-growth log-phase colony-maintenance genetic-diversity field-collection adaptation stress-tolerance oxidative-stress temperature-extreme food-deficit resource-allocation reproductive-senescence endogenous-reserves environmental-stressor climate-adaptation phenology migration dispersal-pattern habitat-patch source-sink population-source population-sink crop-border field-edge non-crop-habitat refuge overwintering-site spring-emergence summer-population fall-population density-dependence aggregation spatial-distribution sampling-accuracy pest-detection monitoring-technology precision-agriculture digital-agriculture automation robotics machine-learning computer-vision sensor-technology data-collection decision-support grower-education extension stakeholder-engagement multidisciplinary-research academia-industry-collaboration sustainable-intensification agroecosystem food-system plant-insect-interaction insect-plant-coevolution salivary-secretion digestive-enzyme lytic-enzyme tissue-maceration feeding-injury physiological-damage mechanical-damage wound-response plant-defense-response induced-resistance systemic-acquired-resistance jasmonic-acid ethylene salicylic-acid secondary-metabolite phenolic-compound oxidative-enzyme redox-enzyme bacterial-transmission phytopathogen opportunistic-pathogen infection-court inoculum disease-vector epidemiology crop-hygiene sanitation crop-rotation cover-crop trap-crop push-pull habitat-manipulation conservation-biocontrol augmentative-biocontrol classical-biocontrol importation establishment non-target-effect host-specificity parasitoid-biology reproductive-parasitoid idiobiont koinobiont mummy-formation emergence-rate parasitoid-impact population-regulation top-down-control bottom-up-control trophic-cascade food-web ecosystem-function ecosystem-service disservice net-effect risk-assessment management-strategy policy regulation pesticide-registration resistance-monitoring stewardship integrated-resistance-management IRM refuge-strategy pyramid-strategy multiple-toxins transgenic-crop GMO biosafety environmental-impact non-target-organism pollinator-protection beneficial-insect natural-enemy-conservation floral-resource nectar pollen alternative-prey habitat-diversification agri-environment-scheme green-infrastructure ecological-compensation landscape-complexity connectivity matrix-quality spillover source-habitat ecological-trap population-viability extinction-risk invasion-biology range-expansion climate-change warming drought irrigation water-stress nitrogen-fertilization crop-vigor plant-nutrition host-plant-quality tritrophic-interaction plant-microbe-insect endosymbiont gut-microbiome salivary-microbiome vector-competence transmission-efficiency acquisition inoculation retention multiplication circulative non-circulative persistent non-persistent propagative stylet-borne foregut-borne hemolymph-borne salivary-gland-borne replication barrier midgut salivary-gland escape infection incubation latent-period inoculation-threshold transmission-threshold epidemic epiphytotic disease-triangle susceptible-host virulent-pathogen favorable-environment disease-cycle primary-inoculum secondary-inoculum overwintering-pathogen survival-structure disease-reservoir alternative-host volunteer-crop weed-reservoir crop-residue soilborne seedborne insect-borne management-tactic preventive curative eradicative suppressive protectant systemic contact translaminar fumigant biological-pesticide microbial-pesticide entomopathogenic-fungus entomopathogenic-nematode entomopathogenic-virus baculovirus granulovirus nucleopolyhedrovirus bacteria Bacillus-thuringiensis Bt-toxin crystal-protein cytolytic-protein vegetative-insecticidal-protein mode-of-action gut-paralysis septicemia toxemia antibiotic symbiont antibiotic-resistance fitness-cost resistance-allele resistance-mechanism metabolic-resistance target-site-resistance penetration-resistance behavioral-resistance cross-resistance multiple-resistance resistance-evolution selection-pressure dose-response lethal-concentration lethal-dose median-lethal-concentration median-lethal-dose confidence-interval probit-analysis logit-analysis dose-mortality time-mortality resistance-ratio resistance-factor diagnostic-dose discriminating-dose baseline-susceptibility susceptible-strain reference-strain field-population resistant-population homozygous-resistant heterozygous dominance recessive incomplete-dominance gene-frequency allele-frequency selection-coefficient fitness relative-fitness population-genetics quantitative-genetics heritability genetic-correlation breeding-value 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phytoalexin lignin suberin callose papilla cell-wall-reinforcement hypersensitive-cell-death induced-systemic-resistance priming defense-elicitor plant-activator biostimulant plant-strengthener organic-amendment compost manure biochar vermicompost compost-tea microbial-inoculant plant-growth-promoting-rhizobacteria mycorrhizal-fungus endophyte epiphyte phytosphere rhizosphere phyllosphere spermosphere anthroposphere ecosystem-process nutrient-cycling carbon-sequestration nitrogen-fixation phosphorus-solubilization potassium-mobilization micronutrient-availability soil-structure water-infiltration water-holding-capacity erosion-control pollination pest-suppression disease-suppression weed-suppression decomposition detritivory predation herbivory mutualism commensalism competition facilitation inhibition amensalism neutralism symbiosis antagonism trophic-interaction food-chain energy-flow material-cycling biomass-pyramid trophic-pyramid ecological-efficiency production-efficiency assimilation-efficiency ecological-footprint sustainability resilience resistance adaptability transformability socio-ecological-system adaptive-management participatory-research citizen-science knowledge-co-production transdisciplinary-research interdisciplinary-research disciplinary-research basic-research applied-research development-research action-research participatory-action-research farmer-participatory-research on-farm-research station-research controlled-experiment field-experiment observational-study correlational-study manipulative-experiment natural-experiment quasi-experiment longitudinal-study cross-sectional-study case-study comparative-study meta-analysis systematic-review scoping-review rapid-review living-review evidence-synthesis knowledge-synthesis research-synthesis systematic-map bibliometric-analysis citation-analysis co-citation-analysis bibliographic-coupling keyword-analysis topic-modeling science-mapping research-front emerging-topic hot-topic research-gap future-direction research-priority research-agenda strategic-research mission-oriented-research societal-challenge sustainable-development-goal nutrition-security climate-smart-agriculture climate-resilient-agriculture low-input-agriculture regenerative-agriculture agroecology integrated-farming mixed-farming diversified-farming specialized-farming intensive-farming extensive-farming conventional-farming alternative-farming urban-agriculture peri-urban-agriculture rural-agriculture smallholder-agriculture large-scale-agriculture family-farming corporate-farming contract-farming tenant-farming subsistence-farming commercial-farming export-oriented-farming import-substituting-farming local-food-system global-food-system food-value-chain supply-chain value-chain commodity-chain agribusiness agrifood-system food-processing food-distribution food-retail food-service food-consumption food-waste food-loss post-harvest-loss storage-loss processing-loss distribution-loss retail-loss consumer-waste plate-waste kitchen-waste food-rescue food-recovery food-redistribution food-banking composting anaerobic-digestion bioenergy biofuel bioproduct biomaterial biorefinery circular-economy circular-bioeconomy green-economy blue-economy bioeconomy nature-based-solution ecosystem-based-adaptation ecosystem-based-mitigation climate-mitigation disaster-risk-reduction extreme-event flood heat-wave cold-snap frost hail windstorm tornado hurricane typhoon cyclone storm-surge sea-level-rise salinization desertification land-degradation soil-degradation soil-erosion soil-compaction soil-sealing soil-contamination soil-pollution heavy-metal persistent-organic-pollutant pesticide-residue antibiotic-residue hormone-residue plastic-pollution microplastic nanoplastic particulate-matter air-pollution water-pollution groundwater-pollution surface-water-pollution eutrophication hypoxia dead-zone biodiversity-loss habitat-loss habitat-fragmentation habitat-degradation habitat-destruction invasive-species introduced-species alien-species non-native-species exotic-species naturalized-species casual-species escaped-species cultivated-species ornamental-species pet-species livestock poultry aquaculture fishery capture-fishery marine-fishery freshwater-fishery inland-fishery recreational-fishery subsistence-fishery artisanal-fishery industrial-fishery overfishing bycatch discards ghost-fishing fishing-gear fishing-method fishing-effort fishing-capacity fishing-pressure stock-assessment fishery-management marine-protected-area no-take-zone marine-reserve terrestrial-protected-area national-park nature-reserve wildlife-sanctuary biosphere-reserve world-heritage-site ramsar-site important-bird-area key-biodiversity-area ecological-network protected-area-network conservation-corridor wildlife-corridor habitat-corridor stepping-stone buffer-zone transition-zone core-zone management-zone sustainable-use-zone restoration-zone rehabilitation-zone reclamation-zone remediation-zone containment-zone eradication-zone surveillance-zone monitoring-zone quarantine-zone biosecurity phytosanitary veterinary animal-health plant-health one-health ecohealth planetary-health global-health public-health environmental-health occupational-health food-safety feed-safety water-safety air-quality environmental-quality environmental-monitoring environmental-assessment environmental-impact-assessment strategic-environmental-assessment cumulative-impact-assessment social-impact-assessment health-impact-assessment hazard-assessment exposure-assessment dose-response-assessment risk-characterization risk-management risk-communication risk-perception risk-governance precautionary-principle polluter-pays-principle user-pays-principle extended-producer-responsibility corporate-social-responsibility environmental-stewardship sustainable-livelihood sustainable-consumption sustainable-production green-procurement green-supply-chain green-logistics green-manufacturing green-chemistry green-technology clean-technology environmental-technology pollution-prevention waste-minimization resource-efficiency energy-efficiency water-efficiency material-efficiency land-use-efficiency nutrient-use-efficiency nitrogen-use-efficiency phosphorus-use-efficiency potassium-use-efficiency micronutrient-use-efficiency fertilizer-use-efficiency pesticide-use-efficiency water-productivity land-productivity labor-productivity capital-productivity total-factor-productivity partial-factor-productivity agricultural-productivity crop-productivity livestock-productivity fishery-productivity forestry-productivity economic-productivity social-productivity environmental-productivity ecological-productivity biological-productivity primary-productivity secondary-productivity net-primary-productivity gross-primary-productivity ecosystem-productivity landscape-productivity regional-productivity global-productivity carbon-productivity nitrogen-productivity water-footprint carbon-footprint nitrogen-footprint phosphorus-footprint biodiversity-footprint material-footprint land-footprint energy-footprint emission-factor intensity-factor decoupling absolute-decoupling relative-decoupling resource-decoupling impact-decoupling dematerialization rematerialization transmaterialization substitution complementarity trade-off synergy co-benefit win-win lose-lose win-lose zero-sum positive-sum negative-sum game-theory prisoner's-dilemma tragedy-of-the-commons collective-action common-pool-resource public-good private-good club-good common-good global-common global-public-good tipping-point critical-transition regime-shift hysteresis irreversibility path-dependence lock-in technological-lock-in institutional-lock-in behavioral-lock-in social-ecological-trap poverty-trap degradation-trap development-trap complex-adaptive-system social-ecological-system coupled-human-natural-system human-environment-system adaptive-capacity transformative-capacity learning-capacity innovation-capacity governance-capacity institutional-capacity organizational-capacity individual-capacity community-capacity social-capital human-capital natural-capital physical-capital financial-capital built-capital cultural-capital political-capital intellectual-capital knowledge-capital information-capital data-capital digital-capital technological-capital infrastructural-capital institutional-capital organizational-capital network-capital relationship-capital reputational-capital brand-capital symbolic-capital existential-capital spiritual-capital religious-capital moral-capital ethical-capital trust-capital reciprocity-capital solidarity-capital cooperation-capital coordination-capital collaboration-capital communication-capital information-sharing knowledge-sharing learning innovation experimentation transformation transition change evolution development growth progress improvement enhancement optimization maximization minimization satisfaction sufficiency abundance scarcity security insecurity vulnerability robustness stability instability equilibrium disequilibrium homeostasis allostasis stress distress eustress coping coping-capacity coping-strategy coping-mechanism coping-resource coping-support social-support emotional-support instrumental-support informational-support appraisal-support network-support community-support institutional-support policy-support financial-support technical-support technological-support infrastructural-support capacity-building capacity-development capacity-strengthening empowerment enablement mobilization activation engagement participation involvement inclusion exclusion marginalization discrimination inequity inequality disparity gap divide digital-divide knowledge-divide information-divide science-divide research-divide innovation-divide technology-divide development-divide north-south-divide east-west-divide urban-rural-divide gender-divide age-divide class-divide income-divide wealth-divide asset-divide opportunity-divide outcome-divide wellbeing-divide happiness-divide life-satisfaction-divide health-divide education-divide skill-divide competency-divide capability-divide functioning-divide agency-divide freedom-divide choice-divide autonomy-divide self-determination-divide dignity-divide respect-divide recognition-divide status-divide power-divide influence-divide voice-divide representation-divide participation-divide engagement-divide citizenship-divide belonging-divide identity-divide culture-divide value-divide belief-divide norm-divide practice-divide behavior-divide habit-divide custom-divide tradition-divide modernity-divide globalization-divide localization-divide universalism-divide particularism-divide cosmopolitanism-divide nationalism-divide patriotism-divide populism-divide elitism-divide democracy-divide authoritarianism-divide control-divide security-divide liberty-divide privacy-divide surveillance-divide transparency-divide accountability-divide responsibility-divide liability-divide guilt-divide blame-divide punishment-divide reward-divide incentive-divide motivation-divide intention-divide attention-divide perception-divide cognition-divide emotion-divide affect-divide attitude-divide preference-divide interest-divide need-divide want-divide desire-divide demand-divide supply-divide market-divide exchange-divide transaction-divide contract-divide agreement-divide cooperation-divide competition-divide conflict-divide coercion-divide force-divide violence-divide peace-divide harmony-divide coexistence-divide tolerance-divide acceptance-divide rejection-divide opposition-divide resistance-divide conformity-divide deviance-divide normality-divide abnormality-divide illness-divide disease-divide disorder-divide disability-divide impairment-divide handicap-divide capacity-divide potential-divide performance-divide achievement-divide attainment-divide accomplishment-divide success-divide failure-divide excellence-divide mediocrity-divide average-divide standard-divide benchmark-divide threshold-divide ceiling-divide floor-divide limit-divide boundary-divide border-divide frontier-divide horizon-divide perspective-divide viewpoint-divide standpoint-divide position-divide stance-divide approach-divide method-divide technique-divide tool-divide instrument-divide device-divide apparatus-divide equipment-divide machine-divide machinery-divide mechanism-divide system-divide structure-divide organization-divide institution-divide regime-divide order-divide arrangement-divide configuration-divide pattern-divide design-divide plan-divide scheme-divide program-divide project-divide initiative-divide intervention-divide action-divide activity-divide operation-divide process-divide procedure-divide protocol-divide guideline-divide recommendation-divide requirement-divide regulation-divide rule-divide law-divide policy-divide strategy-divide tactic-divide objective-divide goal-divide target-divide aim-divide purpose-divide mission-divide vision-divide principle-divide ethic-divide moral-divide criterion-divide indicator-divide measure-divide metric-divide index-divide scale-divide score-divide rating-divide ranking-divide grade-divide level-divide stage-divide phase-divide step-divide progression-divide sequence-divide series-divide succession-divide chain-divide cycle-divide loop-divide spiral-divide wave-divide pulse-divide rhythm-divide tempo-divide pace-divide speed-divide velocity-divide acceleration-divide deceleration-divide momentum-divide inertia-divide energy-divide work-divide effort-divide strain-divide stress-divide pressure-divide tension-divide compression-divide expansion-divide contraction-divide deformation-divide distortion-divide transformation-divide conversion-divide transduction-divide transmission-divide transfer-divide transport-divide movement-divide motion-divide flow-divide flux-divide current-divide stream-divide river-divide tide-divide wind-divide storm-divide weather-divide climate-divide season-divide year-divide decade-divide century-divide millennium-divide epoch-divide era-divide period-divide time-divide duration-divide interval-divide span-divide stretch-divide spell-divide bout-divide episode-divide event-divide occurrence-divide incident-divide accident-divide coincidence-divide chance-divide luck-divide fortune-divide destiny-divide fate-divide karma-divide dharma-divide tao-divide way-divide path-divide road-divide route-divide course-divide direction-divide orientation-divide alignment-divide positioning-divide placement-divide location-divide site-divide place-divide spot-divide point-divide situation-divide context-divide environment-divide setting-divide scene-divide platform-divide base-divide foundation-divide ground-divide soil-divide earth-divide land-divide terrain-divide territory-divide domain-divide realm-divide sphere-divide world-divide globe-divide planet-divide universe-divide cosmos-divide space-divide void-divide emptiness-divide fullness-divide plenum-divide vacuum-divide atmosphere-divide air-divide gas-divide vapor-divide mist-divide fog-divide cloud-divide rain-divide snow-divide ice-divide frost-divide dew-divide humidity-divide moisture-divide wetness-divide dryness-divide aridity-divide drought-divide flood-divide deluge-divide inundation-divide submersion-divide immersion-divide saturation-divide supersaturation-divide condensation-divide evaporation-divide transpiration-divide sublimation-divide deposition-divide precipitation-divide infiltration-divide percolation-divide seepage-divide leakage-divide discharge-divide runoff-divide erosion-divide sedimentation-divide accretion-divide aggregation-divide coagulation-divide flocculation-divide crystallization-divide solidification-divide liquefaction-divide vaporization-divide boiling-divide melting-divide freezing-divide thawing-divide cooling-divide heating-divide warming-divide thermal-divide temperature-divide heat-divide cold-divide cool-divide warm-divide hot-divide thermodynamic-divide enthalpy-divide entropy-divide free-energy-divide Gibbs-free-energy-divide Helmholtz-free-energy-divide chemical-potential-divide fugacity-divide partial-pressure-divide concentration-divide dilution-divide solution-divide mixture-divide compound-divide element-divide atom-divide molecule-divide ion-divide electron-divide proton-divide neutron-divide nucleus-divide particle-divide quark-divide lepton-divide boson-divide fermion-divide hadron-divide baryon-divide meson-divide photon-divide gluon-divide weak-boson-divide Higgs-boson-divide string-divide brane-divide dimension-divide space-time-divide continuum-divide discrete-divide quantum-divide classical-divide relativistic-divide Newtonian-divide Einsteinian-divide Planckian-divide cosmological-divide astronomical-divide microscopic-divide macroscopic-divide mesoscopic-divide nanoscopic-divide atomic-divide molecular-divide cellular-divide tissue-divide organ-divide organism-divide population-divide community-divide ecosystem-divide biome-divide biosphere-divide ecosphere-divide noosphere-divide technosphere-divide anthroposphere-divide sociosphere-divide politosphere-divide econosphere-divide culturosphere-divide religiosphere-divide ideosphere-divide memesphere-divide infosphere-divide cybersphere-divide digisphere-divide virtual-divide analog-divide real-divide actual-divide possible-divide impossible-divide necessary-divide contingent-divide probable-divide improbable-divide certain-divide uncertain-divide determinate-divide indeterminate-divide definite-divide indefinite-divide specific-divide general-divide particular-divide universal-divide singular-divide plural-divide individual-divide collective-divide social-divide cultural-divide historical-divide political-divide economic-divide religious-divide philosophical-divide scientific-divide technological-divide artistic-divide literary-divide musical-divide visual-divide performing-divide cinematic-divide photographic-divide architectural-divide fashion-divide culinary-divide gastronomic-divide olfactory-divide gustatory-divide tactile-divide kinesthetic-divide proprioceptive-divide vestibular-divide auditory-divide somatosensory-divide interoceptive-divide exteroceptive-divide multisensory-divide crossmodal-divide synesthetic-divide anesthetic-divide esthetic-divide aesthetic-divide poetic-divide rhetorical-divide logical-divide mathematical-divide statistical-divide computational-divide algorithmic-divide heuristic-divide systematic-divide methodical-divide empirical-divide theoretical-divide conceptual-divide abstract-divide concrete-divide practical-divide applied-divide pure-divide fundamental-divide basic-divide strategic-divide tactical-divide operational-divide administrative-divide managerial-divide executive-divide leadership-divide followership-divide membership-divide partnership-divide ownership-divide stewardship-divide guardianship-divide trusteeship-divide custodianship-divide wardship-divide tutelage-divide mentorship-divide sponsorship-divide championship-divide friendship-divide relationship-divide kinship-divide companionship-divide fellowship-divide scholarship-divide apprenticeship-divide internship-divide residency-divide assistantship-divide associateship-divide authorship-divide editorship-divide curatorship-divide directorship-divide governorship-divide presidency-divide chancellorship-divide deanship-divide headship-divide chairmanship-divide
Pentodontini
rhinoceros beetles
Pentodontini is the most diverse tribe within the subfamily Dynastinae (rhinoceros beetles), containing over 100 genera distributed across multiple biogeographic regions. Most genera are restricted to a single biogeographic region. The tribe is characterized by substantial morphological diversity, with generic-level identification often relying on mouthpart morphology in females and secondary sexual characters (horns, claw modifications, antennal club length) in males.
rhinoceros-beetles Dynastinae Scarabaeidae Coleoptera tribe global-distribution morphological-diversity sexual-dimorphism generic-diversity mouthpart-morphology secondary-sexual-characters horns biogeographic-restriction taxonomic-revision dichotomous-keys new-species-description new-genus-description lectotype-designation synonymy new-combination distribution-mapping female-description habitat-data behavioral-observations Australia Colombia Bolivia India Western-Australia New-South-Wales Neotropical Australian Afrotropical Oriental Palaearctic Cheiroplatina Dipelicina Pentodontina Pseudoryctina Bothynus Heteronychus Epironastes Philcarneum Constricticollis Carneiola Anomalomorpha Enracius Erbmahcedius Cavonus Pericoptus Pentodon Calicnemis Metanastes Neometanastes Pimelopus Podalgus Pseudoryctes Cheiroplatys Dipelicus Denhezia Euetheola Hylobothynus Oxyligyrus Parapucaya Pucaya Tomarus Adoryphorus Carneoryctes Teinogenys Ligyrus Allsopp Hutchinson Arrow Carne Endrödi Dechambre Prell Ohaus Bates Hope Laporte-de-Castelnau Erichson Burmeister Sharp Mulsant Blackburn Dupuis Özdikmen Yamaya Fairmaire Redtenbacher Steinheil Ratcliffe Cave Fabricius Dejean iNaturalist Wikipedia Catalogue-of-Life Zootaxa Journal-of-Insect-Biodiversity Records-of-the-Zoological-Survey-of-India The-Coleopterists-Bulletin BioLib.cz Wikimedia-Commons DOI 10.11646/zootaxa.4048.4.1 10.11646/zootaxa.4852.4.2 10.11646/zootaxa.5351.3.2 10.26515/rzsi/v125/i2s/2025/172964 10.11646/zootaxa.5716.4.7 10.11646/zootaxa.5072.5.2 10.11646/zootaxa.4852.4.3 10.12976/jib/2024.54.2.2 10.1649/1186.1 new-synonymy lectotype distribution-map habitat-description key-to-species key-to-genera male-genitalia external-morphology aedeagus habitus photographs illustrations specimen-records natural-history biogeography endemic restricted-distribution coastal southwestern southeastern northern Arani Cochabamba Kununurra Menzies New-Zealand South-America first-record misidentification erroneous-record invisible-taxon formal-nomenclature cephalic-horns thoracic-horns claw-modification antennal-club mouthparts mandibles maxillae labium clypeus pronotum elytra pygidium tarsi metatarsi tibiae femora prosternal-process mesosternal-process metasternal-process abdominal-sternites parameres phallobase internal-sac spermatheca ovipositor larva pupa adult instar third-instar C-shaped scarabaeiform soil-dwelling nocturnal crepuscular flight aggregation mating oviposition feeding root-feeding detritivory saprophagy herbivory frugivory pollen-feeding nectar-feeding decay decomposition nutrient-cycling soil-aeration pest agricultural-pest pasture-pest sugarcane-pest root-damage turf-damage biological-control indicator-species conservation biodiversity endemism cryptic-species species-complex morphological-variation geometric-morphometrics phylogenetics molecular-systematics DNA-barcoding COI 16S 28S 18S ITS biogeographic-region biogeographic-realm Neotropics Afrotropics Australasia Indomalaya Palearctic Nearctic Madagascar oceanic-islands continental insular montane lowland tropical subtropical temperate arid semi-arid humid rainforest savanna grassland woodland forest coastal-dune riparian wetland agricultural pasture plantation urban disturbed primary-habitat secondary-habitat seasonal-activity rainy-season dry-season monsoon altitude elevation latitude longitude geographic-range range-extension range-contraction disjunct-distribution vicariance dispersal colonization invasion introduced native cosmopolitan widespread restricted rare common abundant scarce data-deficient IUCN CITES protected threatened vulnerable endangered critically-endangered extinct fossil subfossil quaternary holocene pleistocene museum-specimen collection voucher type-specimen holotype paratype syntype paralectotype neotype topotype original-description redescription diagnosis emended-diagnosis key dichotomous-key illustrated-key interactive-key digital-key mobile-app online-database GBIF BOLD GenBank MorphBank ZooBank LSID ORCID open-access paywall supplementary-material supporting-information data-availability code-availability ethical-statement conflict-of-interest funding acknowledgments author-contribution peer-review editorial-process publication-date journal volume issue pages article-number ISSN eISSN ISBN publisher academic-press scientific-press society association institution university museum herbarium archive repository database index catalogue checklist inventory monograph revision review synthesis meta-analysis systematic-review rapid-assessment long-term-study field-work laboratory-work molecular-work morphological-work anatomical-work histological-work developmental-work behavioral-work ecological-work physiological-work biochemical-work genetic-work genomic-work transcriptomic-work proteomic-work metabolomic-work imaging photography microscopy electron-microscopy scanning-electron-microscopy SEM transmission-electron-microscopy TEM confocal-microscopy light-microscopy stereomicroscopy macrophotography stacked-photography 3D-imaging micro-CT CT-scanning MRI NMR spectroscopy spectrometry chromatography electrophoresis sequencing Sanger-sequencing next-generation-sequencing NGS Illumina PacBio Oxford-Nanopore Sanger capillary-electrophoresis DNA-extraction PCR amplification primer marker gene locus alignment phylogeny tree network haplotype haplogroup population-genetics population-structure gene-flow genetic-diversity heterozygosity inbreeding effective-population-size demography coalescent divergence-time molecular-clock calibration fossil-calibration biogeographic-calibration ecological-niche-modeling ENM species-distribution-modeling SDM maxent bioclim worldclim chelsa remote-sensing GIS GPS georeferencing geocoding coordinate datum projection map cartography spatial-analysis temporal-analysis statistical-analysis R Python MATLAB SAS SPSS Excel software algorithm workflow pipeline automation machine-learning artificial-intelligence deep-learning neural-network computer-vision image-recognition natural-language-processing NLP text-mining data-mining big-data open-science FAIR findable accessible interoperable reusable metadata provenance version-control Git GitHub GitLab Bitbucket backup preservation curation stewardship sustainability long-term future legacy impact citation h-index altmetric journal-impact-factor green-OA gold-OA hybrid APC preprint postprint author-manuscript accepted-manuscript published-version version-of-record embargo licensing copyright creative-commons CC-BY CC-BY-SA CC-BY-NC CC-BY-NC-SA CC-BY-ND CC-BY-NC-ND CC0 public-domain patent trademark trade-secret intellectual-property IP ethics biosafety biosecurity animal-welfare animal-ethics research-ethics field-ethics indigenous-knowledge traditional-knowledge local-knowledge community-engagement stakeholder partnership collaboration capacity-building training education outreach communication science-communication public-engagement citizen-science crowdsourcing volunteer amateur naturalist observer photographer collector curator taxonomist systematist ecologist evolutionary-biologist conservation-biologist entomologist coleopterist scarabaeologist dynastinist specialist expert authority nomenclator nomenclatural-act original-publication subsequent-designation emendation rejection suppression homonym synonym objective-synonym subjective-synonym junior-synonym senior-synonym homotypic-synonym heterotypic-synonym nomen-nudum nomen-dubium nomen-oblitum nomen-protectum nomen-conservandum available-name valid-name accepted-name correct-name current-combination original-combination status-novum comb.-nov.syn.-nov.sp.-nov.gen.-nov.subgen.-nov.subsp.-nov.nom.-nov.nom.-nud.nom.-dub.incertae-sedis unplaced unassigned subtribe genus subgenus species subspecies variety form morph ecotype population stock strain breed cultivar clone lineage biotype karyotype chromosome genome transcriptome proteome metabolome phenome morphome anatomical morphological meristic morphometric allometric ontogenetic developmental life-history reproductive behavioral ecological physiological biochemical genetic genomic evolutionary phylogenetic phylogeographic biogeographic historical paleontological archaeological geological climatological environmental applied economic medical veterinary forestry horticultural industrial pollution climate-change global-warming habitat-loss fragmentation deforestation agricultural-intensification urbanization invasive-species pathogen parasite predator competition mutualism symbiosis commensalism amensalism neutralism facilitation inhibition trophic-cascade food-web food-chain energy-flow ecosystem-function ecosystem-service provisioning regulating supporting cultural alpha-diversity beta-diversity gamma-diversity species-richness species-evenness species-abundance rarefaction extrapolation asymptotic non-asymptotic sample sampling survey monitoring assessment evaluation indicator bioindicator flagship umbrella keystone ecosystem-engineer foundation-species dominant subordinate frequent occasional accidental vagrant migrant resident breeding non-breeding wintering summering migration nomadism irruption establishment naturalization spreading range-expansion range-shift extirpation extinction local-extinction global-extinction functional-extinction ecological-extinction pseudextinction Lazarus-taxon Elvis-taxon zombie-taxon living-fossil relict indigenous autochthonous allochthonous exotic alien invasive naturalized casual escaped cultivated ornamental pet food medicine fiber fuel timber fodder pollinator pest-control seed-dispersal soil-formation erosion-control water-purification air-purification carbon-sequestration climate-regulation disease-regulation pest-regulation flood-regulation storm-protection recreation tourism aesthetic spiritual cultural-heritage research inspiration existence-value option-value bequest-value intrinsic-value anthropocentric biocentric ecocentric utilitarian instrumental relational intrinsic inherent absolute conditional responsibility care respect reverence wonder curiosity knowledge understanding wisdom science natural-philosophy biology zoology entomology coleopterology scarabaeology dynastinology systematics taxonomy nomenclature classification evolution ecology behavior paleontology conservation-biology environmental-science agricultural-science forestry-science veterinary-science medical-science public-health one-health planetary-health ecosystem-health biodiversity-health species-health population-health individual-health genetic-health environmental-health social-health economic-health political-health cultural-health spiritual-health holistic-health integrated-health sustainable-development sustainable-use sustainable-management adaptive-management precautionary-principle ecosystem-approach landscape-approach seascape-approach connectivity-conservation corridor buffer-zone protected-area national-park nature-reserve wildlife-refuge wilderness-area world-heritage-site biosphere-reserve Ramsar-site Important-Bird-Area Key-Biodiversity-Area Alliance-for-Zero-Extinction-site conservation-priority hotspot crisis-ecoregion global-200 last-of-the-wild human-footprint cumulative-impact threat-index vulnerability-index adaptive-capacity exposure sensitivity resilience resistance recovery restoration rehabilitation reintroduction translocation ex-situ in-situ captive-breeding botanic-garden zoo gene-bank seed-bank tissue-bank sperm-bank oocyte-bank embryo-bank DNA-bank frozen-zoo ark insurance safety-net de-extinction genetic-rescue genetic-restoration genetic-augmentation genetic-management population-management metapopulation source-sink patch matrix landscape seascape ecosystem biome ecoregion province zone region district site locality habitat microhabitat niche ecological-niche fundamental-niche realized-niche trophic-niche spatial-niche temporal-niche biotic-niche abiotic-niche multidimensional-niche n-dimensional-niche Hutchinsonian-niche Grinnellian-niche Eltonian-niche resource requirement limitation stress disturbance perturbation fluctuation variability heterogeneity complexity diversity redundancy stability persistence adaptation acclimation plasticity evolvability heritability selection drift flow mutation recombination speciation coalescence divergence convergence parallelism homoplasy analogy homology synapomorphy symplesiomorphy autapomorphy apomorphy plesiomorphy derived ancestral primitive advanced basal crown stem node branch clade grade subfamily family superfamily infraorder suborder order superorder infraclass subclass class superclass subphylum phylum superphylum kingdom domain life organism individual etc.
Rothschildia lebeau
Lebeau's Silk Moth, Forbes Moth
Rothschildia lebeau is a giant silkmoth in the family Saturniidae, distributed from the Lower Rio Grande Valley through eastern Mexico to Brazil. Adults are short-lived and do not feed, relying on fat reserves accumulated during larval development. The species exhibits a life history tradeoff between larval survival and adult body size based on host plant quality, with larger males showing higher mating success.
giant-silkmoth Saturniidae host-plant-tradeoff sexual-selection tropical-dry-forest Barro-Colorado-Island pheromone-communication non-feeding-adult complete-metamorphosis cultural-significance indigenous-use cocoon-harvesting mating-success body-size-dimorphism seasonal-variation microbial-associations protistan-diversity caterpillar-microbiome herbivore forest-ecology Neotropical Mexico Central-America South-America Panama Costa-Rica Brazil Texas Rio-Grande-Valley exhibit-species butterfly-house education conservation light-pollution suburban-sprawl introduced-parasites rearing cottage-industry vestigial-mouthparts fat-reserves short-lived-adult female-sedentary-behavior male-dispersal oviposition-site-selection fitness-consequences life-history-evolution quantitative-genetic-variation wet-season-phenology developmental-plasticity survival-size-tradeoff herbivore-performance plant-animal-interactions microbial-observatory molecular-characterization eukaryotic-microbiome saturniid-caterpillar Lepidoptera Bombycoidea Rothschildia Guérin-Méneville-1868 Attacus-lebeau Lebeau's-Silk-Moth Forbes-Moth Rothschildia-lebeau-forbesi windows transparent-wing-patches scale-less-wing-areas antennae sexual-dimorphism mate-choice sperm-competition reproductive-success larval-performance caterpillar-growth pupal-development adult-emergence longevity predation bat-predation nocturnal crepuscular moth silkmoth silkworm giant-moth forest-moth tropical-moth Neotropical-fauna biodiversity insect-collection museum-specimen Bohart-Museum American-Museum-of-Natural-History entomology lepidopterology systematics taxonomy natural-history field-study observational-study experimental-ecology evolutionary-ecology behavioral-ecology population-biology community-ecology ecosystem-function food-web trophic-interaction herbivory folivory tree-insect-interaction phenology climate-effects seasonality environmental-variation habitat-heterogeneity host-race ecotype local-adaptation phenotypic-plasticity genetic-variation quantitative-trait trade-off allocation reproductive-allocation parental-investment offspring-size offspring-number optimal-clutch-size mate-competition intrasexual-selection intersexual-selection ornament signal chemical-communication olfaction sensory-ecology foraging movement-ecology dispersal migration home-range activity-pattern diel-cycle circadian-rhythm photoperiod temperature humidity rainfall precipitation dry-season wet-season tropical-seasonality El-Niño climate-change global-warming habitat-disturbance forest-fragmentation deforestation agricultural-expansion urbanization conservation-biology threatened-species common-species population-decline extinction-risk protected-area national-park biological-reserve research-station Smithsonian-Tropical-Research-Institute Barro-Colorado-Island-Nature-Monument Gatun-Lake Panama-Canal Isthmus-of-Panama Central-American-isthmus land-bridge biogeography vicariance dispersal-limitation species-range range-expansion range-contraction range-shift poleward-shift elevational-shift altitudinal-gradient latitudinal-gradient thermal-tolerance desiccation-resistance water-balance physiological-ecology metabolic-rate energetics nutritional-ecology stoichiometry nitrogen phosphorus carbon plant-secondary-metabolite tannin alkaloid terpenoid defense induced-defense constitutive-defense herbivore-offense counter-adaptation coevolution evolutionary-arms-race phylogenetic-constraint phylogenetic-signal comparative-method phylogenetic-comparative-analysis trait-evolution reproductive-strategy iteroparity semelparity big-bang-reproduction suicidal-reproduction capital-breeder income-breeder resource-acquisition resource-allocation storage fat-body lipid glycogen protein amino-acid vitamin micronutrient digestion gut-microbiome gut-flora intestinal-microorganism symbiosis mutualism commensalism parasitism pathogen disease immunity immune-response defense-mechanism resistance tolerance recovery survival mortality fecundity fertility sterility hybridization introgression gene-flow population-structure genetic-differentiation FST isolation-by-distance isolation-by-environment landscape-genetics landscape-ecology spatial-ecology geographic-information-system remote-sensing satellite-imagery habitat-suitability species-distribution-model ecological-niche-model MaxEnt BIOMOD ensemble-forecast climate-envelope niche-conservatism niche-evolution realized-niche fundamental-niche biotic-interaction competition facilitation apparent-competition indirect-effect trophic-cascade keystone-species ecosystem-engineer foundation-species dominant-species rare-species functional-diversity functional-trait trait-based-ecology community-assembly filtering sorting neutral-theory niche-theory unified-theory metacommunity metapopulation source-sink-dynamics rescue-effect mass-effect patch-dynamics landscape-mosaic connectivity corridor stepping-stone matrix-quality edge-effect interior-forest core-area buffer-zone fragment-size isolation perimeter-area-ratio shape-index fractal-dimension habitat-amount threshold-effect extinction-debt relaxation-time lag-effect time-delay historical-legacy founder-effect bottleneck genetic-drift inbreeding inbreeding-depression outbreeding outbreeding-depression heterosis hybrid-vigor genetic-rescue assisted-migration managed-relocation translocation reintroduction restoration-ecology habitat-restoration ecological-restoration rewilding de-extinction synthetic-biology conservation-genetics conservation-genomics population-genomics landscape-genomics environmental-DNA eDNA metabarcoding metagenomics transcriptomics proteomics metabolomics phenomics imaging computer-vision machine-learning artificial-intelligence deep-learning neural-network convolutional-neural-network recurrent-neural-network generative-adversarial-network transfer-learning data-augmentation training-set validation-set test-set cross-validation bootstrap jackknife permutation randomization Monte-Carlo Markov-chain Bayesian-inference posterior-probability prior-probability likelihood model-selection information-criterion AIC BIC DIC WAIC LOO model-averaging multimodel-inference structural-equation-model path-analysis causal-inference directed-acyclic-graph potential-outcomes propensity-score instrumental-variable regression-discontinuity difference-in-differences synthetic-control panel-data longitudinal-data time-series cross-sectional-data spatial-data spatiotemporal-data point-pattern areal-data geostatistics kriging variogram spatial-autocorrelation Moran's-I Geary's-c Getis-Ord-Gi LISA hot-spot cold-spot cluster-detection scan-statistic network-analysis graph-theory small-world scale-free-network centrality betweenness closeness eigenvector PageRank community-detection modularity clustering-coefficient path-length diameter radius percolation robustness resilience vulnerability adaptive-capacity transformability social-ecological-system complex-adaptive-system emergent-property self-organization nonlinearity threshold tipping-point hysteresis alternative-stable-state regime-shift critical-transition early-warning-signal resilience-indicator panarchy adaptive-cycle exploitation release reorganization fast-variable slow-variable controlling-variable driver pressure state impact response DPSIR ecosystem-service provisioning-service regulating-service cultural-service supporting-service natural-capital green-infrastructure nature-based-solution ecosystem-based-adaptation ecosystem-based-mitigation payment-for-ecosystem-service REDD+biodiversity-offset habitat-banking mitigation-hierarchy avoidance minimization restoration compensation no-net-loss net-gain biodiversity-metric condition-metric landscape-metric connectivity-metric genetic-metric functional-metric irreplaceability threat opportunity conservation-planning systematic-conservation-planning Marxan Zonation prioritization efficiency effectiveness equity stakeholder governance institution policy law regulation incentive market-based-instrument command-and-control voluntary-agreement certification eco-label sustainable-use sustainable-management adaptive-management learning-by-doing monitoring evaluation indicator target goal vision scenario projection forecast uncertainty risk precautionary-principle safe-minimum-standard environmental-impact-assessment strategic-environmental-assessment cumulative-impact-assessment social-impact-assessment health-impact-assessment life-cycle-assessment carbon-footprint water-footprint ecological-footprint material-flow-analysis substance-flow-analysis input-output-analysis environmentally-extended-input-output-analysis multi-regional-input-output-analysis computable-general-equilibrium integrated-assessment-model earth-system-model global-circulation-model regional-climate-model downscaling bias-correction ensemble-simulation perturbed-physics emulator surrogate-model reduced-complexity-model simple-climate-model integrated-earth-system-model socioeconomic-scenario shared-socioeconomic-pathway representative-concentration-pathway climate-scenario emission-scenario mitigation-scenario adaptation-scenario impact-scenario vulnerability-scenario risk-scenario resilience-scenario transformation-scenario sustainability-scenario narrative storyline pathway trajectory transition transformation decarbonization electrification renewable-energy energy-efficiency circular-economy green-economy blue-economy bioeconomy nature-economy natural-capital-accounting ecosystem-accounting environmental-accounting green-accounting inclusive-wealth genuine-progress-indicator happy-planet-index human-development-index sustainable-development-goals Agenda-2030 Paris-Agreement Convention-on-Biological-Diversity Aichi-targets post-2020-global-biodiversity-framework Kunming-Montreal-Global-Biodiversity-Framework 30-by-30 half-earth nature-needs-half rewilding-Europe rewilding-Britain rewilding-North-America Pleistocene-rewilding trophic-rewilding ecological-replacement taxon-substitution revival-biology molecular-de-extinction proxy-species gene-drive CRISPR genome-editing genetic-modification biotechnology nanotechnology information-technology global-positioning-system unmanned-aerial-vehicle drone satellite sensor camera-trap acoustic-monitoring citizen-science community-science participatory-monitoring traditional-ecological-knowledge indigenous-knowledge local-knowledge farmer-knowledge fishers-knowledge co-production-of-knowledge transdisciplinarity interdisciplinarity multidisciplinarity post-normal-science mode-2-science triple-helix quadruple-helix quintuple-helix open-science open-data open-access open-source FAIR-data findable accessible interoperable reusable data-management-plan metadata provenance quality-assurance quality-control validation verification uncertainty-quantification error-propagation sensitivity-analysis scenario-analysis Monte-Carlo-simulation Bayesian-belief-network influence-diagram decision-tree multi-criteria-decision-analysis cost-benefit-analysis cost-effectiveness-analysis multi-attribute-utility-theory game-theory mechanism-design incentive-compatibility revelation-principle social-choice-theory voting-theory fair-division institutional-design policy-design governance-design architecture polycentric-governance adaptive-governance collaborative-governance network-governance hierarchical-governance market-governance community-governance hybrid-governance meta-governance steering rowing regulatory-state enabling-state partner-state entrepreneurial-state innovation-state strategic-state developmental-state welfare-state green-state sustainable-state resilient-state smart-state digital-state platform-state algorithmic-state surveillance-state authoritarian-state democratic-state deliberative-democracy participatory-democracy direct-democracy representative-democracy liberal-democracy social-democracy green-democracy ecological-democracy biocracy ecofascism deep-ecology social-ecology ecofeminism environmental-justice climate-justice intergenerational-justice intragenerational-justice spatial-justice territorial-justice recognition-justice procedural-justice distributive-justice restorative-justice transformative-justice reparative-justice transitional-justice ecological-justice species-justice multispecies-justice more-than-human-justice anthropocene capitalocene plantationocene chthulucene symbiocene good-anthropocene bad-anthropocene great-acceleration great-divergence great-convergence great-transformation great-decoupling great-disruption great-moderation great-recession great-depression great-war great-famine great-migration great-replacement great-reset great-refusal great-enrichment great-leveling great-compression
Tetramesa
Tetramesa is a genus of minute phytophagous wasps in the family Eurytomidae comprising over 200 described species. Members are exclusively associated with grasses (Poaceae), where they typically induce stem or inflorescence galls. The genus exhibits pronounced host specificity, with most species restricted to a single grass species or closely related congeners. Several species have been deployed as biological control agents against invasive grasses, including T. romana for giant reed (Arundo donax) and candidate agents for medusahead (Taeniatherum caput-medusae) and African lovegrass (Eragrostis curvula). Adults feed on nectar. Recent phylogenomic studies have synonymized the genera Aiolomorphus and Cathilaria within Tetramesa.
biological-control gall-wasp grassland invasive-species-management host-specificity cryptic-species phylogenomics Eurytomidae Chalcidoidea Hymenoptera phytophagy Poaceae stem-gall inflorescence-gall integrated-pest-management riparian-restoration molecular-systematics species-delimitation GMYC COI 28S biocontrol-agent Arundo-donax giant-reed medusahead African-lovegrass Eragrostis Sporobolus barley-jointworm integrated-taxonomy Southern-Hemisphere-diversity grassland-expansion Miocene-diversification monophagy host-range-testing non-target-effects tick-habitat cattle-fever-tick Rio-Grande Big-Bend-National-Park greenhouse-rearing oviposition-behavior water-deficit-stress generation-time parasitoid Eurytoma-amicophaga Eurytominae Walker-1848 Terebrantes Apocrita microhymenoptera minute-wasp reduced-wing-venation antennal-segmentation mesosoma-sculpture ovipositor molecular-operational-taxonomic-units MOTU species-boundaries cryptic-diversity sampling-effects SPEDE-sampler host-genetic-proxies field-host-range no-choice-testing native-Australian-species non-target-risk host-phylogeny target-weed suppression topping mechanical-control integrated-weed-management native-vegetation-recovery light-penetration canopy-structure dispersal establishment post-release-evaluation adventive-population Austin-Texas Idaho England Wales Orissa Canary-Islands Canary-Is.Mediterranean South-Africa southern-Africa Northern-Hemisphere Australia North-America Texas California Greece central-Asia Iran Ukraine Moldova Russia Israel Tajikistan Japan Eragrostis-curvula Sporobolus-pyramidalis Sporobolus-natalensis Sporobolus-africanus Hyparrhenia-hirta Andropogon-gayanus Eragrostis-plana Eragrostis-planiculmis Eragrostis-biflora Eragrostis-capensis Eragrostis-trichophora Deschampsia-caespitosa Deschampsia-setacea Taeniatherum-caput-medusae Hordeum barley Phragmites common-reed Dactylis-glomerata orchardgrass Poales grass graminoid cereal-crop pasture rangeland invasive-weed noxious-weed habitat-structure ecosystem-functioning naturalised-species alien-grass donor-country receiver-country biocontrol-practitioner host-plant-resistance internode-infestation oviposition-preference behavioral-categories antennae-attitude laboratory-artifact polyphagy specialist generalist climate-fluctuation molecular-clock dated-analysis phylogenetic-clade intrapopulation-variation geographic-sub-structuring singleton-effect overestimation consensus-delimitation operational-taxonomic-unit integrative-taxonomy evidence-line caveat putative-species novel-taxon undescribed-species biodiversity-assessment faunal-survey field-collection genetic-sequence mitochondrial-DNA nuclear-DNA cytochrome-c-oxidase-I ribosomal-DNA internal-transcribed-spacer chloroplast-DNA rps16 rpl32 trnK trnL ITS plant-insect-interaction herbivore phytophage primary-consumer trophic-level food-web community-ecology restoration-ecology conservation-biological-control classical-biological-control augmentative-biological-control inundative-release establishment-rate population-increase downstream-dispersal river-mile biomass-reduction plant-community tick-feeding-ant tick-feeding-beetle habitat-modification law-enforcement international-border flood-control irrigation-channel stream-clogging river-bank-weakening native-vegetation-stifling wildlife-habitat-reduction cattle-inspection deer-inspection Rhipicephalus Boophilus tick-population vegetation-transition shading-effect long-term-suppression high-suppression cane-thinning mechanical-cutting growth-suppression susceptibility-to-attack combination-treatment synergistic-effect research-demonstration USDA Agricultural-Research-Service entomologist field-measurement post-release-monitoring impact-assessment agent-efficacy non-target-damage crop-safety agricultural-importance host-specific-herbivore sustainable-method cost-effective-method weed-control invasive-plant-management habitat-restoration ecosystem-service provisioning-service regulating-service supporting-service cultural-service biodiversity-conservation threatened-species endangered-species habitat-loss climate-change land-use-change anthropogenic-disturbance urbanization agricultural-intensification grazing-pressure fire-regime nitrogen-deposition carbon-sequestration soil-stabilization water-quality air-quality pollinator nectar-feeding adult-diet larval-diet tissue-feeding gall-induction plant-manipulation phytohormone cytokinin auxin cell-proliferation tissue-differentiation nutrient-sink sink-strength source-sink-dynamics plant-physiology reproductive-output seed-production seed-weight flowering-head inflorescence stem-elongation tiller-production biomass-allocation growth-rate relative-growth-rate net-primary-productivity ecosystem-productivity carbon-cycling nitrogen-cycling phosphorus-cycling decomposition litter-quality herbivory-impact top-down-control bottom-up-control trophic-cascade indirect-effect direct-effect density-dependence density-independence population-regulation population-dynamics life-table survivorship fecundity reproductive-rate intrinsic-rate-of-increase doubling-time carrying-capacity population-growth exponential-growth logistic-growth Allee-effect stochasticity environmental-variation demographic-variation genetic-variation phenotypic-plasticity local-adaptation evolutionary-response coevolution arms-race escape-and-radiate diversification-rate speciation extinction phylogenetic-diversity functional-diversity taxonomic-diversity alpha-diversity beta-diversity gamma-diversity species-richness species-evenness species-abundance-distribution rank-abundance-curve diversity-index Shannon-index Simpson-index evenness-index dominance-index similarity-index dissimilarity-index cluster-analysis ordination principal-component-analysis principal-coordinate-analysis non-metric-multidimensional-scaling canonical-correspondence-analysis redundancy-analysis variation-partitioning structural-equation-modeling path-analysis meta-analysis systematic-review evidence-synthesis research-priority knowledge-gap data-deficiency IUCN-Red-List conservation-status protected-area management-plan monitoring-protocol adaptive-management stakeholder-engagement policy-recommendation legislation regulation incentive market-based-instrument payment-for-ecosystem-services carbon-credit biodiversity-offset habitat-banking conservation-easement land-trust stewardship sustainable-agriculture organic-farming agroecology precision-agriculture conservation-tillage cover-cropping crop-rotation intercropping agroforestry silvopasture riparian-buffer filter-strip grassed-waterway wetland-restoration prairie-restoration savanna-restoration forest-restoration degraded-land abandoned-land marginal-land set-aside wildlife-corridor habitat-connectivity landscape-matrix patch-dynamics edge-effect interior-habitat core-area buffer-zone permeability resistance least-cost-path circuit-theory isolation-by-distance isolation-by-resistance gene-flow genetic-drift inbreeding outbreeding hybridization introgression admixture population-structure genetic-differentiation FST GST D allele-frequency haplotype genotype phenotype quantitative-trait morphological-trait behavioral-trait life-history-trait reproductive-trait survival-trait developmental-trait physiological-trait metabolic-trait stress-response immune-response defense-trait resistance-trait tolerance-trait avoidance-trait antipredator-defense chemical-defense mechanical-defense crypsis mimicry aposematism startle-display thanatosis autotomy regeneration compensatory-growth induced-defense constitutive-defense direct-defense indirect-defense volatile-organic-compound extrafloral-nectar domatia mutualism ant-plant-mutualism pollination-mutualism seed-dispersal-mutualism protection-mutualism nutrition-mutualism transport-mutualism cleaning-mutualism agricultural-mutualism domestication crop-wild-relative landrace cultivar variety breeding selection genetic-improvement yield quality stress-tolerance nutrient-use-efficiency water-use-efficiency photosynthetic-efficiency harvest-index grain-filling source-limitation sink-limitation assimilate-transport phloem-loading phloem-unloading remobilization senescence stay-green antioxidant reactive-oxygen-species oxidative-stress heat-stress cold-stress drought-stress salinity-stress metal-stress nutrient-deficiency nutrient-toxicity soil-pH soil-texture soil-structure soil-organic-matter soil-microbial-community rhizosphere mycorrhiza nitrogen-fixation phosphorus-solubilization potassium-mobilization micronutrient-uptake allelopathy competition facilitation niche-differentiation resource-partitioning temporal-partitioning spatial-partitioning morphological-partitioning physiological-partitioning phenological-partitioning competitive-exclusion competitive-displacement priority-effect founder-effect mass-effect species-sorting neutral-theory ecological-drift dispersal-limitation environmental-filtering biotic-filtering abiotic-filtering trait-based-approach functional-trait response-trait effect-trait community-assembly community-disassembly community-reassembly succession primary-succession secondary-succession pioneer-species climax-species keystone-species ecosystem-engineer foundation-species dominant-species subordinate-species transient-species resident-species migrant-species nomadic-species irruptive-species invasive-species naturalized-species casual-species established-species spreading-species widespread-species restricted-species endemic-species cosmopolitan-species disjunct-distribution relict-distribution refugium glacial-refugium interglacial-refugium postglacial-colonization range-shift range-expansion range-contraction range-fragmentation source-population sink-population metapopulation population-viability minimum-viable-population effective-population-size genetic-rescue demographic-rescue assisted-migration managed-relocation ex-situ-conservation in-situ-conservation cryopreservation seed-bank gene-bank botanic-garden arboretum zoo aquarium microbial-culture-collection DNA-bank tissue-bank sperm-bank oocyte-bank embryo-bank reintroduction translocation reinforcement supplementation headstarting captive-breeding propagation nucleus-population release-program soft-release hard-release acclimatization habitat-preparation predator-control competitor-control disease-control parasite-control health-screening quarantine veterinary-care nutrition-management environmental-enrichment behavioral-training imprinting fostering cross-fostering hand-rearing parent-rearing semi-natural-enclosure pre-release-conditioning survival-rate reproduction-rate dispersal-rate site-fidelity homeward-orientation migration navigation orientation magnetic-sense olfactory-sense visual-sense auditory-sense mechanosensory-sense thermosensory-sense hygrosensory-sense electrosensory-sense polarized-light celestial-cues landmark-cues odor-plume wind-direction sun-compass star-compass moon-compass magnetic-compass inertial-navigation path-integration cognitive-map spatial-memory temporal-memory episodic-memory procedural-memory associative-learning operant-conditioning classical-conditioning habituation sensitization song-learning tool-use problem-solving innovation social-learning cultural-transmission teaching cooperation altruism kin-selection reciprocal-altruism group-selection multilevel-selection selfish-gene inclusive-fitness reproductive-skew dominance-hierarchy territoriality home-range activity-center den-site nest-site roost-site singing-post display-site lek arena scrape burrow cavity crevice hole tunnel gallery chamber cell comb paper-nest mud-nest silk-nest spun-cocoon pupal-case ootheca egg-mass clutch brood litter covey pack herd flock colony society superorganism eusociality cooperative-breeding communal-breeding polygyny polyandry promiscuity monogamy serial-monogamy genetic-monogamy social-monogamy extra-pair-copulation sperm-competition cryptic-female-choice sexual-selection intersexual-selection intrasexual-selection mate-choice mate-guarding mate-switching divorce widowhood mate-replacement breeding-season breeding-cycle breeding-attempt breeding-success fledging-success recruitment juvenile-survival adult-survival longevity life-expectancy maximum-lifespan aging telomere oxidative-damage mitochondrial-dysfunction cellular-senescence apoptosis necrosis autophagy proteostasis DNA-repair epigenetic-modification transgenerational-effect maternal-effect paternal-effect parental-care maternal-care paternal-care biparental-care alloparental-care brood-care egg-care larval-care pupal-care food-provisioning nest-defense territory-defense predator-defense disease-defense parasite-defense hygiene-behavior grooming nest-cleaning cannibalism siblicide infanticide filial-cannibalism sexual-cannibalism cannibalistic-oophagy trophic-egg regurgitation trophallaxis food-sharing information-sharing alarm-call mobbing distraction-display broken-wing-display tailing persecution attack escape flight fight freeze faint death-feigning tonic-immobility startle-response withdrawal avoidance dishabituation latent-inhibition blocking overshadowing relative-validity conditioned-inhibition spontaneous-recovery renewal reinstatement rapid-reacquisition latent-learning insight-learning observational-learning vicarious-learning emulation imitation social-facilitation social-inhibition audience-effect bystander-effect conformity obedience compliance commitment consistency foot-in-the-door door-in-the-face low-ball that's-not-all sunk-cost endowment-effect loss-aversion risk-aversion risk-seeking probability-weighting hyperbolic-discounting temporal-discounting intertemporal-choice optimal-foraging marginal-value-theorem prey-choice patch-choice diet-breadth central-place-foraging load-size travel-time handling-time search-time encounter-rate prey-density prey-profitability prey-handling prey-detection prey-pursuit prey-capture prey-consumption prey-rejection ingestion-rate digestion-rate assimilation-efficiency production-efficiency ecological-efficiency trophic-transfer-efficiency Lindeman-efficiency ten-percent-rule pyramid-of-numbers pyramid-of-biomass pyramid-of-energy food-chain trophic-network energy-flow material-cycling biogeochemical-cycle carbon-cycle nitrogen-cycle phosphorus-cycle sulfur-cycle water-cycle hydrological-cycle atmospheric-circulation ocean-circulation thermohaline-circulation El-Niño La-Niña ENSO NAO AO PDO AMO climate-oscillation climate-anomaly weather-extreme drought flood heat-wave cold-snap storm hurricane typhoon cyclone tornado thunderstorm lightning wildfire volcanic-eruption earthquake tsunami landslide avalanche meteor-impact mass-extinction background-extinction speciation-rate extinction-rate net-diversification turnover standing-diversity fossil-record taphonomy preservation-bias sampling-bias collection-bias publication-bias citation-bias language-bias geographic-bias temporal-bias taxonomic-bias size-bias abundance-bias habitat-bias facies-bias Lagerstätte Konservat-Lagerstätte Konzentrat-Lagerstätte trace-fossil body-fossil chemical-fossil molecular-fossil biomarker isotope-signature paleoclimate-proxy paleoecology paleobiogeography paleoecosystem paleocommunity paleoenvironment paleogeography paleolatitude paleolongitude continental-drift plate-tectonics seafloor-spreading subduction orogeny mountain-building basin-formation sedimentation erosion weathering soil-formation pedogenesis horizon-development profile-development chronosequence toposequence biosequence climosequence landform geomorphology topography relief elevation slope aspect curvature roughness drainage watershed catchment basin subbasin reach segment zone riffle pool run glide cascade waterfall wetland marsh swamp bog fen mire peatland tundra taiga boreal-forest temperate-forest temperate-rainforest temperate-grassland tropical-forest tropical-rainforest tropical-dry-forest tropical-savanna tropical-grassland desert hot-desert cold-desert semiarid steppe prairie pampas veld savanna woodland shrubland chaparral maquis garigue fynbos kwongan mallee mulga mogote karst cave sinkhole doline polje uvala ponor spring seep oasis wadi arroyo gully ravine canyon gorge valley dale glen strath rift graben horst fault fold anticline syncline monocline dome plateau mesa butte escarpment bluff cliff crag tor inselberg bornhardt kopje hill knoll mound ridge spur saddle pass col gap notch cleft fissure joint bedding-plane foliation lineation cleavage schistosity gneissic-banding magma lava igneous-rock sedimentary-rock metamorphic-rock mineral crystal grain texture structure fabric composition chemistry mineralogy petrology geochemistry geophysics geochronology radiometric-dating isotopic-dating dendrochronology varve-chronology lichenometry archaeomagnetism paleomagnetism thermoluminescence optically-stimulated-luminescence electron-spin-resonance cosmogenic-nuclide exposure-dating burial-dating terrestrial-cosmogenic-nuclide in-situ-cosmogenic-nuclide meteoric-cosmogenic-nuclide fallout-radionuclide lead-210 cesium-137 plutonium americium radiocarbon carbon-14 calibration-curve intcal marine-reservoir-effect terrestrial-reservoir-effect hard-water-effect contamination pretreatment combustion graphitization accelerator-mass-spectrometry liquid-scintillation-counting gas-proportional-counting beta-counting gamma-spectrometry alpha-spectrometry mass-spectrometry inductively-coupled-plasma-mass-spectrometry thermal-ionization-mass-spectrometry secondary-ion-mass-spectrometry laser-ablation-inductively-coupled-plasma-mass-spectrometry electron-microprobe scanning-electron-microscopy transmission-electron-microscopy light-microscopy confocal-microscopy fluorescence-microscopy polarizing-microscopy phase-contrast-microscopy differential-interference-contrast-microscopy dark-field-microscopy bright-field-microscopy stereomicroscopy dissecting-microscopy microtomy sectioning staining mounting embedding fixation preservation collection curation vouchering repository herbarium museum archive database registry catalogue checklist inventory survey monitoring assessment evaluation appraisal audit review synthesis rapid-review scoping-review narrative-review critical-review state-of-the-art-review literature-review bibliometric-analysis citation-analysis co-citation-analysis bibliographic-coupling co-authorship-analysis collaboration-network knowledge-map concept-map ontology taxonomy classification nomenclature typification designation description diagnosis illustration measurement morphometry meristics count ratio index score rank grade scale level category class group set subset cluster clade lineage branch stem crown root node tip terminal internal basal derived ancestral plesiomorphic apomorphic synapomorphic autapomorphic homologous homoplastic convergent parallel reversal gain loss duplication deletion insertion inversion fusion fission polyploidy aneuploidy chromosome-number genome-size gene-number transcriptome proteome metabolome lipidome glycome interactome phenome connectome epigenome microbiome virome mycobiome bacteriome phytobiome rhizobiome endobiome ectobiome parasitome pathobiome diseasome toxome exposome foodome nutriome pharmacome toxicome adverse-outcome-pathway mode-of-action mechanism-of-action target-site receptor-binding enzyme-inhibition ion-channel-blockade membrane-disruption DNA-intercalation inflammation cell-differentiation cell-migration cell-adhesion cell-signaling signal-transduction gene-expression transcription translation post-translational-modification protein-folding protein-degradation protein-trafficking vesicle-transport exocytosis endocytosis phagocytosis pinocytosis mitophagy pexophagy ribophagy ER-phagy lysophagy lipophagy glycophagy ferritinophagy aggrephagy xenophagy entosis cytosis syncytium plasmodesmata gap-junction tight-junction adherens-junction desmosome hemidesmosome focal-adhesion cell-polarity cell-asymmetry cell-division mitosis meiosis cytokinesis karyokinesis chromosome-segregation spindle-assembly checkpoint DNA-replication DNA-recombination transposition retrotransposition horizontal-gene-transfer vertical-gene-transfer endosymbiosis symbiogenesis holobiont hologenome extended-phenotype niche-construction ecosystem-engineering keystone-process ecological-function ecosystem-function ecosystem-good natural-capital natural-resource renewable-resource non-renewable-resource common-pool-resource public-good private-good club-good externality market-failure government-failure policy-failure institutional-failure tragedy-of-the-commons free-rider collective-action social-dilemma prisoner's-dilemma chicken-game assurance-game coordination-game zero-sum-game positive-sum-game negative-sum-game win-win lose-lose win-lose Pareto-optimal Pareto-improvement Kaldor-Hicks-improvement cost-benefit-analysis cost-effectiveness-analysis multi-criteria-decision-analysis decision-support-system expert-system knowledge-based-system rule-based-system case-based-reasoning machine-learning deep-learning neural-network random-forest support-vector-machine decision-tree regression-tree classification-tree clustering-algorithm dimensionality-reduction feature-selection feature-extraction pattern-recognition image-analysis signal-processing time-series-analysis spatial-analysis geographic-information-system remote-sensing satellite-imagery aerial-photography drone-imagery LiDAR radar sonar seismic geophysical geochemical biogeochemical ecoinformatics bioinformatics computational-biology systems-biology synthetic-biology biotechnology nanotechnology genetic-engineering genome-editing CRISPR TALEN zinc-finger-nuclease transgenic cisgenic intragenic subgenic agrobacterium-mediated-transformation biolistics electroporation microinjection protoplast-fusion somatic-hybridization cybrid asymmetric-hybrid synthetic-polyploid doubled-haploid haploid-induction anther-culture microspore-culture ovule-culture embryo-culture endosperm-culture protoplast-culture cell-culture tissue-culture organ-culture shoot-culture root-culture callus-culture suspension-culture bioreactor hairy-root-culture transformation hardening micropropagation clonal-propagation meristem-culture shoot-tip-culture axillary-bud-culture nodal-culture leaf-culture petiole-culture stem-culture internode-culture rhizome-culture bulb-culture corm-culture tuber-culture seed-culture zygotic-embryo-culture somatic-embryo-culture somatic-embryogenesis organogenesis caulogenesis rhizogenesis adventitious-shoot-formation direct-regeneration indirect-regeneration primary-metabolism secondary-metabolism biosynthesis biotransformation bioconversion biodegradation bioremediation phytoremediation mycoremediation zooremediation bioaugmentation biostimulation natural-attenuation monitored-natural-attenuation engineered-natural-attenuation permeable-reactive-barrier pump-and-treat soil-vapor-extraction air-sparging thermal-desorption incineration landfilling containment capping slurry-wall grout-curtain sheet-piling clay-liner geomembrane geotextile geogrid geocomposite geosynthetic geofabric geonet geocell geopipe geostrip geofoam geobag geotube geocontainer geomat geoblanket geospacer geodrain geoinjection geogrout geopier geocolumn geopile geowall geoberm geotrench geocut geofill geolift geoslab geopanel geoblock geobrick geotile geopaver geoweb geomattress geocushion
Tortopus
Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.
Ephemeroptera Polymitarcyidae burrowing-mayflies Neotropical clay-bank-habitat U-shaped-tunnels mandibular-tusks Tortopsis Campsurinae Panamerican-distribution cladistic-revision synapomorphy parastyli-receptors flattened-penes furcasternal-protuberances subapical-tubercles limnological-sampling-bias nymph-morphology adult-morphology type-species-T.-igaranus new-species-T.-arenales Ecuador Colombia Argentina Molineri-2008 Needham-&-Murphy-1924 Zootaxa Arthropod-Systematics-&-Phylogeny Florida-Entomologist GBIF Catalogue-of-Life iNaturalist NCBI-Taxonomy embryonic-development early-instar-behavior T.-harrisi T.-bellus T.-circumfluus T.-zottai T.-igaranus T.-incertus T.-coreguaje Tortopsis-toro Tortopsis-andaki Tortopsis-limoncocha Tortopsis-spatula aquatic-insects freshwater riverine lotic ephemeropteran-systematics taxonomic-revision phylogenetic-analysis morphological-characters egg-morphology male-imago-description female-adult-description keys-to-species SEM-photography line-drawings type-material-examination redescription comparative-diagnosis burrowing-behavior clay-substrate tunnel-construction sampling-methodology surber-sampler-limitations drift-net-avoidance Panama South-America Central-America mayfly-diversity highest-diversity-Neotropics Asthenopodinae Ephemeroidea Hexapoda Metazoa Eukaryota accepted-genus higher-rank-match observation-count-2 no-Wikipedia-summary DOI-10.11646/zootaxa.2481.1.1 DOI-10.11646/zootaxa.2436.1.4 DOI-10.3897/asp.79.e62735 DOI-10.2307/3493493 Scott-et-al.-1959 Navás Ulmer Traver Lugo-Ortiz-&-McCafferty Domínguez Banks McDunnough Pictet Needham-&-Murphy freshwater-ecology aquatic-entomology insect-development nymphal-stages imago-description alate-adults wing-venation R-sector gonopore-structure parastyli-length pedestal-morphology lateral-view-curvature mandibular-morphology tusk-tubercles mesosternal-structure abdominal-sternum-notch furcasternum protuberance-configuration receptor-socket-orientation IR-vein R2-vein claw-like-structure gonopore-association penis-base-separation female-fore-wing additional-veins comparative-morphology type-specimen holotype allotype paratype neotropical-species new-species-description species-key identification-key male-adult-key female-adult-key nymph-key generic-key Polymitarcyidae-genera systematics phylogeny cladistics monophyly sister-group character-state transformation-series parsimony-analysis morphological-data-set external-morphology egg-chorion micropyle egg-burster hatching emergence behavioral-study early-instar embryology oviposition substrate-preference habitat-specificity endemic regional-diversity conservation biomonitoring water-quality-indicator macroinvertebrate benthic infaunal interstitial hyporheic river-bank stream-bank erosion sediment particle-size clay-content bank-stability root-mat vegetation-association flow-regime current-velocity depth width stream-order catchment basin ecoregion biogeography dispersal colonization vicariance speciation adaptive-radiation morphological-divergence convergent-evolution homoplasy character-conflict bootstrap-support Bremer-support consensus-tree strict-consensus majority-rule taxonomic-stability nomenclatural-act original-description subsequent-designation type-locality type-depository museum-collection specimen-preservation alcohol-preservation slide-mounting SEM-preparation imaging-techniques scientific-illustration photography measurement morphometrics ratios proportions allometry sexual-dimorphism geographic-variation intraspecific-variation interspecific-variation diagnostic-characters identification-confidence taxonomic-expertise literature-review bibliographic-search database-query online-repository open-access paywall abstract-only full-text-unavailable citation-analysis reference-list source-quality evidence-strength knowledge-gap research-need future-study collaboration field-work collection-trip sampling-effort specimen-availability type-examination revision-scope taxonomic-concept circumscription inclusion exclusion species-group species-complex cryptic-species morphospecies phylogenetic-species evolutionary-significant-unit operational-taxonomic-unit DNA-barcoding molecular-systematics integrative-taxonomy traditional-taxonomy alpha-taxonomy beta-taxonomy gamma-taxonomy classification-hierarchy Linnaean-system rank-based-nomenclature phylogenetic-nomenclature clade-name crown-group stem-group total-evidence congruence incongruence data-partition character-weighting implied-weighting equal-weighting ordered-characters unordered-characters multistate-characters binary-characters continuous-characters meristic-characters qualitative-characters quantitative-characters morphological-matrix data-matrix character-matrix taxon-sampling character-sampling missing-data inapplicable-data polymorphism intraspecific-polymorphism fixed-difference diagnostic-difference unique-derived-character autapomorphy shared-derived-character symplesiomorphy convergent-character parallel-evolution reversal character-optimization ACCTRAN DELTRAN unambiguous-optimization ambiguous-optimization most-parsimonious-tree tree-length consistency-index retention-index rescaled-consistency-index homoplasy-index character-fit character-congruence character-support branch-support node-support clade-stability taxonomic-stability-index phylogenetic-signal noise saturation substitution-model rate-heterogeneity among-site-rate-variation gamma-distribution invariant-sites model-selection Akaike-information-criterion Bayesian-information-criterion likelihood-ratio-test Bayesian-inference Markov-chain-Monte-Carlo posterior-probability prior-probability hyperprior convergence effective-sample-size burn-in stationarity trace-plot parameter-estimate credible-interval highest-posterior-density tree-topology tree-shape lineage-through-time diversification-rate extinction-rate speciation-rate net-diversification birth-death-model Yule-model coalescent-model population-size effective-population-size gene-flow migration-rate isolation-by-distance phylogeography historical-biogeography ancestral-area-reconstruction dispersal-vicariance-analysis decisive-weighting event-based-method taxon-area-cladogram area-cladogram biogeographic-homology track-analysis panbiogeography phylogenetic-biogeography ecological-biogeography dispersal-biogeography vicariance-biogeography center-of-origin area-of-endemism refugium Pleistocene glaciation climate-change habitat-fragmentation habitat-loss anthropogenic-impact pollution eutrophication sedimentation dam-construction water-abstraction flow-modification thermal-pollution toxic-substances heavy-metals pesticides urbanization agriculture deforestation land-use-change riparian-degradation channelization dredging mining invasive-species biological-invasion competition predation parasitism mutualism commensalism amensalism facilitation ecosystem-engineer keystone-species flagship-species umbrella-species indicator-species sentinel-species bioindicator ecological-indicator environmental-indicator water-quality-index biotic-index multimetric-index rapid-bioassessment protocol standardization quality-assurance quality-control reference-condition least-disturbed minimally-disturbed best-available type-specific region-specific stream-type river-type functional-feeding-group collector-gatherer collector-filterer scraper shredder predator parasite engulfer piercer burrower sprawler clinger swimmer skater diver surface-dweller benthic-dweller interstitial-dweller hyporheic-dweller phreatic-dweller troglobite stygobite cave-inhabitant spring-inhabitant headwater mid-order high-order lowland montane alpine subtropical tropical temperate boreal arctic antarctic palearctic nearctic afrotropical oriental australian oceanic cosmopolitan widespread restricted relict disjunct-distribution range-extension new-record first-record country-record state-record province-record locality-record elevation-record altitudinal-range latitudinal-range longitudinal-range distribution-pattern range-size range-fill extent-of-occurrence area-of-occupancy IUCN-category threat-status data-deficient not-evaluated least-concern near-threatened vulnerable endangered critically-endangered extinct-in-wild extinct population-trend population-density population-structure age-structure size-structure sex-ratio reproductive-output fecundity fertility mortality survivorship life-table cohort-analysis generation-time doubling-time intrinsic-rate-of-increase finite-rate-of-increase net-reproductive-rate stable-age-distribution stationary-population expanding-population declining-population source-sink-dynamics metapopulation population-viability-analysis minimum-viable-population genetic-diversity heterozygosity allelic-richness private-allele founder-effect bottleneck genetic-drift inbreeding inbreeding-depression outbreeding outbreeding-depression hybridization introgression gene-flow-barrier reproductive-isolation prezygotic-isolation postzygotic-isolation mechanical-isolation temporal-isolation habitat-isolation behavioral-isolation gametic-isolation zygotic-mortality hybrid-inviability hybrid-sterility hybrid-breakdown reinforcement character-displacement ecological-speciation allopatric-speciation sympatric-speciation parapatric-speciation peripatric-speciation centrifugal-speciation divergence-with-gene-flow speciation-continuum anagenesis cladogenesis budding-speciation branching-speciation phyletic-gradualism punctuated-equilibrium stasis trend progression orthogenesis aristogenesis mosaic-evolution co-evolution arms-race Red-Queen-hypothesis evolutionary-stable-strategy frequency-dependent-selection density-dependent-selection r-selection K-selection bet-hedging iteroparity semelparity ploidy-level chromosome-number karyotype genome-size transposable-element gene-duplication neofunctionalization subfunctionalization pseudogene regulatory-evolution cis-regulatory trans-regulatory enhancer promoter silencer insulator transcription-factor signaling-pathway developmental-pathway gene-network epigenetic-modification DNA-methylation histone-modification chromatin-remodeling genomic-imprinting parental-effect maternal-effect paternal-effect cytoplasmic-inheritance mitochondrial-DNA chloroplast-DNA endosymbiont Wolbachia Rickettsia Spiroplasma Cardinium Arsenophonus Hamiltonella Regiella Serratia Buchnera Carsonella Portiera primary-endosymbiont secondary-endosymbiont facultative-symbiont obligate-symbiont mutualistic-symbiont parasitic-symbiont commensal-symbiont pathogen vector disease-transmission zoonosis emerging-infectious-disease biosecurity quarantine eradication control management conservation-biology restoration-ecology habitat-restoration river-restoration stream-restoration floodplain-restoration riparian-restoration wetland-restoration ecosystem-restoration passive-restoration active-restoration assisted-recovery managed-relocation assisted-colonization ex-situ-conservation in-situ-conservation protected-area national-park nature-reserve wildlife-refuge biosphere-reserve world-heritage-site ramsar-site important-bird-area key-biodiversity-area Alliance-for-Zero-Extinction EDGE-species evolutionarily-distinct globally-endangered conservation-prioritization systematic-conservation-planning Marxan Zonation conservation-opportunity conservation-cost conservation-benefit trade-off socioeconomic-factor stakeholder-engagement community-based-conservation adaptive-management monitoring-program long-term-study citizen-science public-participation environmental-education outreach science-communication knowledge-transfer capacity-building institutional-strengthening policy-development legislation regulation enforcement compliance incentive payment-for-ecosystem-services carbon-credit biodiversity-offset mitigation-banking conservation-easement land-trust private-land-conservation conservation-covenant stewardship sustainable-use sustainable-management ecosystem-based-management integrated-water-resources-management basin-management catchment-management landscape-management seascape-management connectivity-conservation corridor stepping-stone buffer-zone matrix-management countryside-biogeography reconciliation-ecology novel-ecosystem designer-ecosystem ecosystem-service provisioning-service regulating-service supporting-service cultural-service food-production water-supply raw-materials genetic-resources medicinal-resources ornamental-resources air-quality-regulation climate-regulation water-regulation erosion-regulation water-purification waste-treatment disease-regulation pest-regulation pollination seed-dispersal habitat-provision nutrient-cycling soil-formation primary-production recreation tourism aesthetic-value spiritual-value educational-value scientific-value inspiration sense-of-place cultural-heritage traditional-knowledge indigenous-knowledge local-knowledge ecosystem-service-valuation economic-valuation contingent-valuation choice-experiment benefit-transfer total-economic-value use-value non-use-value direct-use indirect-use option-value bequest-value existence-value willingness-to-pay willingness-to-accept cost-benefit-analysis cost-effectiveness-analysis multi-criteria-analysis decision-support uncertainty risk-assessment scenario-analysis sensitivity-analysis robustness-analysis adaptive-capacity resilience vulnerability exposure sensitivity coping-capacity transformation social-ecological-system complex-adaptive-system tipping-point regime-shift alternative-stable-state hysteresis path-dependence legacy-effect slow-variable fast-variable panarchy adaptive-cycle exploitation release reorganization foresight anticipatory-governance transformative-change sustainability-science future-earth interdisciplinary transdisciplinary problem-oriented solution-oriented action-oriented knowledge-co-production boundary-work boundary-object boundary-organization science-policy-interface science-society-interface knowledge-brokerage knowledge-exchange knowledge-integration mode-1-science mode-2-science triple-helix quadruple-helix responsible-innovation reflexive-governance adaptive-governance collaborative-governance polycentric-governance network-governance self-organization collective-action common-pool-resource tragedy-of-the-commons social-dilemma institutional-analysis institutional-design Ostrom-design-principles monitoring sanctioning conflict-resolution nested-enterprises proportionality collective-choice constitutional-choice operational-choice collective-action-theory game-theory prisoner's-dilemma chicken-game assurance-game coordination-game evolutionary-game-theory replicator-dynamics ESS Nash-equilibrium Pareto-optimality social-optimum market-failure government-failure voluntary-failure non-market-value public-good common-good club-good private-good rivalry excludability free-rider hold-up-problem moral-hazard adverse-selection information-asymmetry transaction-cost property-right common-property state-property private-property hybrid-property bundle-of-rights usufruct access withdrawal alienation tenure-security land-tenure water-tenure resource-tenure customary-tenure statutory-tenure legal-pluralism de-facto de-jure formal informal legality legitimacy recognition deviance corruption elite-capture social-capital bonding-capital bridging-capital linking-capital trust reciprocity reputation social-network social-embeddedness structural-hole brokerage closure cohesion density centrality betweenness degree eigenvector PageRank small-world scale-free random-network regular-network hierarchical-network modularity community-structure clustering assortativity homophily heterophily preferential-attachment fitness-model network-evolution network-dynamics temporal-network multilayer-network multiplex-network bipartite-network affiliation-network one-mode-network two-mode-network adjacency-matrix edge-list graph-theory vertex edge node link tie connection path cycle tree forest component giant-component percolation phase-transition criticality self-organized-criticality power-law heavy-tail Pareto-distribution lognormal-distribution exponential-distribution Poisson-distribution binomial-distribution normal-distribution Gaussian-distribution statistical-distribution probability-distribution cumulative-distribution probability-density-function cumulative-distribution-function survival-function hazard-function moment mean median mode variance standard-deviation coefficient-of-variation skewness kurtosis quantile percentile decile quartile interquartile-range range minimum maximum outlier robust-statistic parametric-test non-parametric-test hypothesis-testing null-hypothesis alternative-hypothesis significance-level p-value confidence-interval type-I-error type-II-error statistical-power effect-size sample-size sampling-design experimental-design observational-study quasi-experiment natural-experiment randomized-controlled-trial before-after-control-impact BACI impact-assessment environmental-impact-assessment strategic-environmental-assessment cumulative-impact-assessment social-impact-assessment health-impact-assessment life-cycle-assessment carbon-footprint water-footprint ecological-footprint material-flow-analysis substance-flow-analysis energy-analysis exergy-analysis emergy-analysis input-output-analysis environmentally-extended-input-output social-accounting-matrix computable-general-equilibrium agent-based-model system-dynamics-model Bayesian-network structural-equation-model path-analysis factor-analysis principal-component-analysis cluster-analysis discriminant-analysis canonical-correspondence-analysis redundancy-analysis non-metric-multidimensional-scaling ordination classification regression-analysis linear-regression nonlinear-regression generalized-linear-model generalized-additive-model mixed-effects-model multilevel-model hierarchical-model Bayesian-hierarchical-model state-space-model hidden-Markov-model time-series-analysis spatial-analysis geostatistics kriging variogram spatial-autocorrelation Moran's-I Geary's-c Getis-Ord-Gi local-indicator-of-spatial-association LISA hot-spot cold-spot spatial-cluster spatial-outlier edge-effect boundary-effect modifiable-areal-unit-problem MAUP ecological-fallacy atomistic-fallacy cross-scale-inference upscaling downscaling scaling-law allometric-scaling fractal fractal-dimension self-similarity multi-fractal lacunarity roughness texture pattern process pattern-process-relationship neutral-model null-model randomization-test permutation-test bootstrap jackknife cross-validation model-averaging model-comparison information-criterion Bayesian-model-averaging ensemble-modeling consensus-model prediction forecast projection scenario narrative-scenario quantitative-scenario storyline shared-socioeconomic-pathway representative-concentration-pathway emission-scenario climate-scenario impact-scenario adaptation-scenario mitigation-scenario transformation-scenario backcasting forecasting trend-extrapolation expert-judgment Delphi-method participatory-modeling stakeholder-analysis influence-diagram conceptual-model mental-model system-archetype leverage-point intervention-point policy-instrument economic-instrument regulatory-instrument voluntary-instrument information-instrument market-based-instrument command-and-control-instrument subsidy tax charge fee levy deposit-refund tradable-permit emission-trading carbon-tax biodiversity-banking conditional-cash-transfer community-based-natural-resource-management joint-forest-management co-management adaptive-co-management transboundary-management international-regime global-governance multilateral-environmental-agreement convention treaty cartagena-protocol nagoya-protocol Aichi-target sustainable-development-goal SDG post-2020-global-biodiversity-framework Kunming-Montreal-global-biodiversity-framework 30-by-30-target nature-based-solution NbS ecosystem-based-adaptation ecosystem-based-mitigation green-infrastructure blue-infrastructure natural-infrastructure hybrid-infrastructure climate-smart-agriculture climate-smart-landscape climate-smart-development low-carbon-development green-economy circular-economy bioeconomy blue-economy sustainable-consumption sustainable-production life-cycle-thinking cradle-to-cradle industrial-ecology industrial-symbiosis urban-metabolism socio-ecological-metabolism material-intensity energy-intensity carbon-intensity decoupling absolute-decoupling relative-decoupling resource-efficiency eco-efficiency eco-innovation green-technology clean-technology environmental-technology appropriate-technology participatory-technology technology-transfer technology-diffusion innovation-system national-innovation-system sectoral-innovation-system technological-innovation-system socio-technical-system socio-technical-transition multi-level-perspective niche-innovation regime landscape transition-management strategic-niche-management transition-experiment living-lab real-world-laboratory urban-living-lab rural-living-lab sustainability-transition fundamental-change system-innovation directionality reflexivity responsiveness anticipation learning-by-doing doing-by-learning social-learning organizational-learning policy-learning transformative-capacity capacity-development institutional-learning evolutionary-governance institutional-change path-dependency lock-in carbon-lock-in technological-lock-in institutional-lock-in behavioral-lock-in cultural-lock-in unlocking destabilization phase-out divestment stranded-asset just-transition transition-pathway transition-scenario deep-decarbonization net-zero carbon-neutral climate-neutral nature-positive biodiversity-positive restoration-positive regenerative circular doughnut-economics planetary-boundary safe-operating-space environmental-ceiling social-foundation intergenerational-equity intragenerational-equity distributive-justice procedural-justice recognition-justice environmental-justice climate-justice energy-justice water-justice food-justice spatial-justice territorial-justice ecological-justice species-justice intersectionality adaptation mitigation sustainability sustainable-development well-being human-development capability-approach human-security environmental-security climate-security water-security food-security energy-security livelihood-security disaster-risk-reduction climate-change-adaptation climate-change-mitigation loss-and-damage climate-finance green-climate-fund adaptation-fund global-environment-facility biodiversity-finance nature-finance blended-finance impact-investment ESG environmental-social-governance sustainable-finance green-bond social-bond sustainability-bond blue-bond nature-bond biodiversity-credit offset inset nature-based-credit ecosystem-service-credit payment-for-results results-based-payment conditional-payment performance-based-payment outcome-based-payment environmental-service-payment PSA PES REDD REDD+AFOLU LULUCF NAMA NAMC NDC NAP NBSAP national-adaptation-plan national-biodiversity-strategy-and-action-plan nationally-determined-contribution nationally-appropriate-mitigation-action intended-nationally-determined-contribution long-term-low-emission-development-strategy net-zero-target carbon-budget remaining-carbon-budget emission-gap adaptation-gap finance-gap implementation-gap ambition-loop ratchet-mechanism global-stocktake transparency-framework enhanced-transparency-framework common-but-differentiated-responsibilities respective-capabilities climate-equity climate-litigation climate-governance climate-diplomacy climate-negotiation UNFCCC COP CMP CMA SBSTA SBI IPCC IPBES UNCCD CBD CITES Ramsar World-Heritage Man-and-the-Biosphere MAB UNESCO FAO WHO ILO UNDP UNEP UN-Habitat World-Bank IMF WTO OECD GEF GCF AF multilateral-development-bank regional-development-bank national-development-bank development-finance-institution export-credit-agency sovereign-wealth-fund pension-fund insurance-company asset-manager asset-owner corporate-investor retail-investor impact-investor philanthropic-investor development-partner bilateral-donor multilateral-donor private-donor foundation NGO CSO civil-society-organization non-governmental-organization community-based-organization indigenous-peoples-organization women's-organization youth-organization faith-based-organization research-organization think-tank university academic-institution knowledge-institution capacity-building-institution training-institution extension-service advisory-service consulting-firm engineering-firm technology-provider equipment-supplier service-provider contractor operator maintainer utility municipality local-government provincial-government national-government federal-government ministry department agency authority commission board council committee parliament congress legislature executive judiciary media journalist reporter editor publisher broadcaster platform social-media influencer celebrity activist advocate campaigner mobilizer organizer networker connector bridge-builder boundary-spanner knowledge-broker science-communicator extension-agent development-practitioner policy-maker decision-maker planner manager administrator leader champion entrepreneur innovator intrapreneur social-entrepreneur systems-entrepreneur institutional-entrepreneur policy-entrepreneur political-entrepreneur business-model business-model-innovation social-business inclusive-business base-of-pyramid fortune-at-bottom-of-pyramid shared-value creating-shared-value stakeholder-capitalism purpose-driven-business B-Corp benefit-corporation social-enterprise cooperative mutual association collective commons commoning peer-production open-source creative-commons knowledge-commons digital-commons data-commons genetic-commons seed-commons water-commons fisheries-commons forest-commons pasture-commons land-commons urban-commons atmospheric-commons global-commons common-heritage-of-mankind common-concern-of-humankind restorative-justice multispecies-justice more-than-human anthropocene capitalocene plantationocene chthulucene symbiocene good-anthropocene great-acceleration great-transformation great-decoupling great-convergence great-divergence great-fragmentation great-integration great-transition great-work long-now deep-time futures-thinking future-studies strategic-foresight scenario-planning trend-analysis weak-signal wild-card black-swan grey-rhino blue-sky green-swan systemic-change structural-change cultural-change behavioral-change individual-change collective-change social-change organizational-change technological-change economic-change political-change ecological-change planetary-change earth-system-change biodiversity-change ocean-change cryosphere-change atmospheric-change biogeochemical-change nitrogen-cycle-change phosphorus-cycle-change carbon-cycle-change water-cycle-change hydrological-change soil-change vegetation-change ecosystem-change landscape-change seascape-change biosphere-change noosphere-change technosphere-change anthroposphere-change sociosphere-change econosphere-change politosphere-change culturosphere-change semiosphere-change infosphere-change digisphere-change cybersphere-change urban-sphere-change agricultural-sphere-change industrial-sphere-change extractive-sphere-change manufacturing-sphere-change service-sphere-change financial-sphere-change knowledge-sphere-change innovation-sphere-change creativity-sphere-change culture-sphere-change identity-sphere-change meaning-sphere-change value-sphere-change purpose-sphere-change vision-sphere-change mission-sphere-change strategy-sphere-change tactic-sphere-change operation-sphere-change action-sphere-change practice-sphere-change habit-sphere-change routine-sphere-change ritual-sphere-change tradition-sphere-change custom-sphere-change convention-sphere-change norm-sphere-change rule-sphere-change law-sphere-change regulation-sphere-change standard-sphere-change protocol-sphere-change procedure-sphere-change process-sphere-change system-sphere-change structure-sphere-change function-sphere-change pattern-sphere-change dynamic-sphere-change state-sphere-change phase-sphere-change regime-sphere-change equilibrium-sphere-change stability-sphere-change instability-sphere-change chaos-sphere-change order-sphere-change complexity-sphere-change simplicity-sphere-change diversity-sphere-change uniformity-sphere-change heterogeneity-sphere-change homogeneity-sphere-change variation-sphere-change invariance-sphere-change change-sphere-change continuity-sphere-change discontinuity-sphere-change emergence-sphere-change submergence-sphere-change evolution-sphere-change devolution-sphere-change revolution-sphere-change involution-sphere-change convolution-sphere-change resolution-sphere-change dissolution-sphere-change absolution-sphere-change solution-sphere-change

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