Maternal-care
Guides
Acanthosomatidae
Shield Bugs
Acanthosomatidae is a family of shield bugs within Pentatomoidea comprising approximately 200 species in 55 genera across three subfamilies (Acanthosomatinae, Blaudusinae, Ditomotarsinae). The family is notable for its distinctive reproductive biology, with females exhibiting either maternal care (egg-nymph guarding) or egg smearing using secretions from Pendergrast's organs. Maternal care has evolved independently at least three times within the subfamily Acanthosomatinae and is correlated with reduction or loss of Pendergrast's organs, suggesting an evolutionary trade-off between these strategies.
Aconophora
Aconophora is a genus of treehoppers in the family Membracidae, tribe Aconophorini, characterized by a long, forward-projecting pronotal horn. The genus contains 13 described species distributed in the Neotropics. The most studied species, Aconophora compressa, was introduced to Australia in 1995 as a biological control agent for invasive lantana but was subsequently found to attack non-target plants, leading to its discontinued use. Members of this genus feed on plant stem sap and are not known to form mutualistic associations with ants, unlike the related genus Guayaquila.
Aconophora compressa
lantana bug, lantana treehopper, lantana stemsucking treehopper, lantana sap-sucking bug
Aconophora compressa is a treehopper (family Membracidae) native to Mexico, Central America, and Colombia. It was introduced to Australia in 1995 as a biological control agent for the invasive weed Lantana camara. The species is brown, up to 8 mm long, with a thorn-shaped body and produces copious honeydew. It has been documented to attack non-target plants, particularly fiddlewood (Citharexylum spinosum) in Australia, leading to its discontinuation as a biocontrol agent after 2001. It was also rejected for use in South Africa due to unacceptable risks to indigenous Lippia species.
treehopperbiological-controlinvasive-species-managementMembracidaephloem-feedermaternal-caregregarioushoneydew-producernon-target-effectsLantana-camaraMexicoAustraliaSouth-AfricaVerbenaceaeCitharexylum-spinosumLippiaAloysia-citrodorathermal-tolerancestem-suckingsap-feedingegg-guardinginstarsnymphal-developmentweed-biocontrolhost-range-expansionecological-risk-assessmentbiosecurityWalker-1851AconophoriniMembracinaeHemipteraAuchenorrhynchaCicadomorphaMembracoideaAetalionidae
Aetalionid Treehoppers
Aetalionidae is a family of treehoppers in the superfamily Membracoidea, distinguished from the more diverse Membracidae by morphological features including a front femur fused to the trochanter and a completely exposed scutellum. The family is predominantly Neotropical in distribution, with two recognized subfamilies: Biturritiinae (exclusively Neotropical) and Aetalioninae (containing the Neotropical genus Aetalion and the Paleotropical genus Darthula). Females exhibit maternal care by guarding egg masses, and many species form aggregations on tree branches where they produce honeydew and associate with ants and stingless bees.
Alopecosa
Fox Spiders
Alopecosa is a genus of wolf spiders (Lycosidae) comprising approximately 160 species. These ground-dwelling spiders are predominantly Eurasian in distribution, with some species extending into North Africa and North America. Most species reach up to 2 cm in body length. Females construct burrows for egg deposition and guard their egg sacs until hatching. The genus has been subject to taxonomic revision, with species historically grouped into sibling species complexes based on morphology, though courtship behavior and molecular techniques have proven more reliable for species delimitation.
Alopecosa kochi
Koch's Wolf Spider
Alopecosa kochi is a medium-sized wolf spider (Lycosidae) distributed across North America from southern Canada to northern Mexico. It is an active cursorial hunter that does not build capture webs, instead pursuing small invertebrates on open ground. The species exhibits clear sexual dimorphism, with males typically smaller, darker, and possessing more defined leg banding. Females carry egg sacs attached to their spinnerets and transport spiderlings on their backs after hatching.
Amaurobius ferox
black lace-weaver, black lace weaver
Amaurobius ferox is a nocturnal spider known for its distinctive cribellate web with woolly, lace-like silk. The species exhibits a unique subsocial organization among spiderlings, including synchronized molting, cooperative prey capture, and matriphagy—where offspring consume their mother after hatching. Native to Europe, it has been introduced to North America and is commonly found near human structures in dark, moist environments.
Amblypygi
whip spiders, tailless whip scorpions, amblypygids
Amblypygi is an ancient order of arachnids comprising approximately 280 species across five families. These nocturnal predators are characterized by extremely elongated first pair of legs modified as sensory "whips" and raptorial pedipalps for capturing prey. They lack venom glands, silk production, and the tail-like flagellum present in their relatives the whip scorpions (Uropygi). Several species exhibit unusual social behaviors including maternal care and kin recognition.
Ammophila
Thread-waisted Sand Wasps
Ammophila is a large, cosmopolitan genus of solitary hunting wasps in the family Sphecidae, comprising over 200 species distributed across warmer regions of all continents except Antarctica. These thread-waisted wasps are characterized by their elongated, slender petiole connecting the thorax and abdomen, and their distinctive nesting behavior involving burrow excavation and caterpillar provisioning. Females construct underground nests in sandy or compact soils, paralyze caterpillars with their sting, and provide this food source for their developing larvae. The genus serves as a host for Strepsiptera endoparasites and is subject to nest parasitism by satellite flies and other organisms.
Anisolabididae
Anisolabididae is a family of earwigs (order Dermaptera, suborder Neodermaptera) comprising 38 genera across 13 subfamilies. Members include both cosmopolitan and geographically restricted species, with some showing specialized adaptations for subcortical or coastal habitats. Several genera, particularly Euborellia, are recognized for their predatory behavior and potential as biological control agents of agricultural pests. The family exhibits diverse morphologies ranging from typical earwig body plans to dorsoventrally flattened forms adapted for life under bark.
Anisolabis
Anisolabis is a genus of earwigs in the subfamily Anisolabidinae, with most species occurring in Asia. The genus name derives from the asymmetrical male cerci, with the right cercus more acute than the left. The best-studied species is Anisolabis maritima, the maritime earwig, which has been extensively researched for its social behavior, sexual selection, and maternal care. Other species such as A. annulipes are known agricultural pests.
Anisolabis maritima
maritime earwig, seaside earwig
A cosmopolitan, wingless earwig species found on coastal beaches worldwide. Males possess asymmetrical forceps used for mating, prey capture, and defense. Exhibits notable maternal care behavior, with females guarding eggs and nymphs for up to 28 days and progressively provisioning offspring. Displays complex social dynamics including sex-specific aggression patterns and size-based dominance in shelter competition.
Apiomerus
Bee Assassins, Bee Assassin Bugs
Apiomerus is a genus of brightly colored, conspicuous assassin bugs in the family Reduviidae, commonly known as bee assassins. The genus contains approximately 110 described species and represents the largest clade within the monophyletic tribe Apiomerini. Members are characterized by their use of plant resin as 'sticky traps' to capture prey, a trait unique among assassin bugs. Females possess specialized tibial combs for resin collection and application onto egg masses, providing maternal care. The genus exhibits significant sexual dimorphism and extreme diversity in genital morphology, hypothesized to result from sexual selection driving speciation.
Argidae
Argid Sawflies
Argidae is a large family of sawflies (Hymenoptera: Symphyta) comprising approximately 800 species worldwide, with greatest diversity in tropical regions, particularly the Neotropics. The family is distinguished from other Symphyta by the reduction of the antenna to three flagellomeres, with the terminal segment often elongated and forked in males. Larvae are phytophagous and frequently exhibit gregarious feeding behavior; some species display maternal care of eggs and young larvae.
Attelabidae
Leaf-rolling Weevils, Giraffe Weevils
Attelabidae is a widespread family of primitive weevils characterized by straight antennae inserted near the base of the rostrum. The family includes approximately 2,500 species globally, with subfamilies exhibiting distinct reproductive behaviors. Attelabinae are true leaf rollers, where females cut and roll leaves to create cradles for egg deposition and larval development. Rhynchitinae (sometimes treated as a separate family) have larvae that develop in flower buds, fruits, terminal shoots, or as leaf miners. Some members, particularly in the genus Trachelophorus, possess elongated necks and are known as giraffe weevils.
Bethylidae
Flat wasps
Bethylidae is a family of aculeate wasps in the superfamily Chrysidoidea, commonly known as flat wasps due to their dorsoventrally compressed body form. The family exhibits a biology intermediate between parasitoid and predatory wasps: females sting and paralyze prey (primarily beetle and lepidopteran larvae), then oviposit on the immobilized hosts. Most species are small (2–10 mm), with females often wingless or apterous while males are always winged. The family contains eight recognized subfamilies and approximately 96 genera, with highest diversity in tropical and subtropical regions.
Centrotinae
Centrotinae is the largest subfamily of treehoppers (Membracidae) and the only subfamily with a worldwide distribution. It comprises approximately 1,350 species in 216 genera. The subfamily is characterized by morphological features including a posterior pronotal process that typically does not conceal the scutellum, and forewings with a truncated clavus and broad apical limbus. Centrotinae is considered the most primitive membracid subfamily based on fossil evidence and biogeographic patterns.
Chelisoches morio
black earwig
Chelisoches morio, commonly known as the black earwig, is a cosmopolitan earwig species in the family Chelisochidae. Adults are uniformly jet black and among the largest earwigs, reaching up to 36 mm in length. The species exhibits extended maternal care, with females guarding eggs and feeding first-instar nymphs before they disperse. Males possess distinctive widely separated, serrated cerci that differ markedly from female morphology. The species has been investigated as a potential biological control agent due to its predatory habits and high consumption rates of agricultural pests.
Chrysomelina
Chrysomelina is a subtribe of leaf beetles within the family Chrysomelidae, subfamily Chrysomelinae. Members of this subtribe are characterized by their ability to produce chemical defenses, including de novo synthesis of iridoids (cyclic monoterpenes) and host-plant-dependent compounds such as salicylaldehyde. Larvae possess specialized glandular secretions used to repel predators. The subtribe exhibits diverse defensive strategies that have evolved through recruitment of oxidases from the glucose-methanol-choline (GMC) oxidoreductase superfamily. Some species display subsocial behavior with maternal care of offspring.
Chrysomelinae
broad-bodied leaf beetles, broad-shouldered leaf beetles
Chrysomelinae is a subfamily of leaf beetles (Chrysomelidae) comprising approximately 3,000 species worldwide, commonly known as broad-bodied or broad-shouldered leaf beetles. The subfamily includes the economically significant Colorado potato beetle (Leptinotarsa decemlineata), a major agricultural pest. Chrysomelinae exhibits remarkable diversity in form and coloration, with adults typically displaying convex, rounded bodies often with bright coloration and variable patterns. The subfamily is distinguished by several unique morphological features including antennae inserted on or adjacent to the anterior head edge, mandibles with large membranous prosthecae, and a single anal cell in each wing. Larvae possess six pairs of stemmata, palmate mandibles, and annular spiracles. Both life stages possess defensive glands that secrete protective chemicals.
Coleothorpa dominicana
Lord Leaf Beetle
Coleothorpa dominicana is a case-bearing leaf beetle in the family Chrysomelidae. It is native to North America. The species is notable for its maternal egg protection behavior, in which females construct funnel-shaped structures to shield their eggs. Development from egg to adult takes approximately one year, with a larval period lasting 5 to 8 months.
Coriarachne brunneipes
Brown-legged Crab Spider
Coriarachne brunneipes is a species of crab spider in the family Thomisidae, commonly known as the Brown-legged Crab Spider. It occurs across the United States and Canada. The species is known for its habitat association with tree bark and documented maternal care behavior.
Ctenus
Tropical Wandering Spiders
Ctenus is a genus of wandering spiders in the family Ctenidae, first described by Walckenaer in 1805. It comprises approximately 155 species with a broad distribution spanning South America, Africa, and East Asia. Species in this genus are abundant in tropical rainforest leaf litter, where they serve as active predators and have become model organisms for studying predator ecology in ground-dwelling arthropod communities. Some species, particularly C. medius, possess venom with documented proteolytic, hyaluronidase, and phospholipase activities, with potential medical significance due to complement system interference.
Cydnidae
burrowing bugs, burrower bugs
Cydnidae is a family of pentatomoid true bugs commonly known as burrowing or burrower bugs. Many species are fossorial, burrowing into soil using their head and forelegs, emerging primarily to mate and lay eggs. Other members live above ground, often associated with plants. The family includes agricultural pests and exhibits diverse habitat preferences across subfamilies.
Diplocentrus spitzeri
Ruby Rock Scorpion
Diplocentrus spitzeri is a scorpion species in the family Diplocentridae, endemic to the Sierra Madre Occidental in Sonora, Mexico. The species exhibits distinctive maternal care behaviors, including a specialized birth posture where the female elevates her body on her legs and pedipalps while young emerge from the genital opercula and climb onto her back. First instar young remain on the mother's back until their first molt, during which time the female displays defensive behaviors to protect them. The species inhabits rocky pine-oak forest environments, where it shelters under stones and in rock crevices.
Elasmucha
Parent shieldbugs
Elasmucha is a genus of shield bugs in the family Acanthosomatidae, commonly known as parent shieldbugs. Species in this genus are notable for exhibiting subsocial behavior, specifically extended maternal care of eggs and early nymphal stages. Females guard egg masses and remain with offspring through early instars, actively defending against predators and parasitoids. The genus includes arboreal species associated with specific host trees, with documented seasonal host plant switching in some species.
Embiidina
webspinners, footspinners
Embiidina is a small order of cryptic, soft-bodied insects known as webspinners or footspinners, characterized by their unique ability to produce silk from specialized glands located in their swollen foretarsi. They construct extensive silk galleries or tunnels under bark, in leaf litter, or within soil crevices, which serve as protective shelters and foraging sites. The order exhibits pronounced sexual dimorphism: females are wingless and neotenic, remaining in galleries throughout their lives, while males of most species develop wings and disperse to locate mates. Webspinners are primarily tropical in distribution and display facultatively communal behavior with maternal care of eggs and young.
Enoplognatha ovata
candy-striped spider, common candy-striped spider, polymorphic spider
Enoplognatha ovata is a small theridiid spider native to Europe and introduced to North America. The species exhibits striking color polymorphism, with three main morphs varying in abdominal pigmentation from plain to striped patterns. Despite its diminutive size (4-7 mm), it is an effective predator capable of capturing prey substantially larger than itself. Females construct tangled webs on the undersides of leaves and demonstrate maternal care by guarding egg sacs and provisioning spiderlings.
Epimetopidae
Hooded Shore Beetles
Epimetopidae is a small family of semi-aquatic beetles in the superfamily Hydrophiloidea, comprising approximately 72 described species across three genera. The family exhibits a disjunct distribution: Epimetopus is restricted to the New World (Neotropical and Nearctic regions), Eumetopus occurs in Asia, and Eupotemus is found in Africa. A distinctive feature of the family is the pronotal hood—a central projection of the pronotum that forms a shelf above the head. Females of all three genera carry egg cases on the underside of the abdomen, representing one of three independent origins of maternal care in Hydrophiloidea.
Eremobates
sun spiders, camel spiders, wind scorpions
Eremobates is a genus of solifuges (sun spiders/camel spiders) in the family Eremobatidae, distributed primarily in western North America. These fast-moving arachnids are notable for having the largest jaw-to-body ratio of any animal. They are non-venomous predators that rely on powerful chelicerae to subdue prey. Most species are nocturnal and navigate primarily using their pedipalps rather than vision. The genus contains approximately 97 accepted species.
Eustala
humpbacked orb-weavers
Eustala is a genus of orb-weaver spiders in the family Araneidae, first described by Eugène Simon in 1895. As of November 2024, the genus contains 87 species distributed primarily across the Americas, from the United States through Central America to South America, with highest diversity in Brazil, Uruguay, and Argentina. Several species have been documented exhibiting specialized ecological relationships, including myrmecophily (living among ants) in Central American species that inhabit ant-defended acacia plants. The genus includes the well-known humpbacked orb-weaver, Eustala anastera, found across North and Central America.
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htperchroostnestdenburrowtunneldigexcavateminequarryextractremoveeliminateeradicateexterminatedestroydemolishruinwreckdamageharminjurehurtwoundmaimcripplelamedisableincapacitateparalyzestunshockastonishamazeastoundstaggerstupefydazebewilderconfusepuzzleperplexbafflemystifyenigmaForficula
earwigs
Forficula is the largest genus of earwigs, containing at least 68 species. The genus was established by Linnaeus in 1758 and is native to Afro-Eurasia, primarily Europe, though it now occurs on all continents except Antarctica. The best known species is Forficula auricularia, a common household and agricultural pest that has been introduced to North America, Australia, and New Zealand. Members of this genus exhibit maternal care of eggs and nymphs, a rare trait among insects.
Forficula auricularia
European earwig, common earwig
Forficula auricularia is an omnivorous earwig native to Europe, western Asia, and North Africa, now invasive across North America, Australia, and New Zealand. It is recognized by forceps-like cerci—curved in males, straight in females—and concealed hindwings that unfold to resemble human ears, inspiring both its common name and specific epithet. The species exhibits extended maternal care, with females guarding eggs and nymphs in underground nests. Population dynamics vary geographically: temperate populations typically produce one brood annually, while warmer regions may support two broods.
Gargaphia
Gargaphia is a genus of lace bugs (family Tingidae) containing more than 70 described species. Members are small, phytophagous true bugs characterized by intricate reticulated forewings. Several species are economically significant as agricultural pests, particularly on solanaceous crops and passion fruit. The genus is notable for exhibiting maternal care behaviors, including egg guarding and defensive responses to predators.
Gargaphia solani
eggplant lace bug
Gargaphia solani is a subsocial lace bug in the family Tingidae, described by Heidemann in 1914 after emerging as a pest of eggplant in Norfolk, Virginia. It is the first species in its family in which maternal care was discovered. Females guard circular egg masses on leaf undersides and defend nymphs against predators, a behavior essential for offspring survival given heavy predation pressure. Development from egg to adult takes approximately 20 days, with nymphs passing through five instars. The species is a significant agricultural pest of solanaceous crops and has been extensively studied for its reproductive trade-offs and the evolution of parental care.
Geotomini
Geotomini is a tribe of burrower bugs within the subfamily Cydninae. Members are ground-dwelling heteropterans with documented maternal care behaviors, including seed-carrying by females to provision nymphs. The tribe has been studied primarily through the species Melanaethus crenatus, with limited tribe-wide behavioral characterization.
Hogna
wolf spiders
Hogna is a large genus of wolf spiders (Lycosidae) comprising more than 200 described species distributed across all continents except Antarctica. The genus is taxonomically problematic, having never been properly revised even in Europe, and lacks a reliable diagnostic framework. Species are generally large, ground-dwelling hunters that construct open silk-lined burrows rather than using trapdoors. Hogna carolinensis is among the largest wolf spiders in North America, with females reaching 35 mm body length. The genus exhibits notable ecological diversity, with parallel habitat specialization documented in island radiations such as the Galápagos and Madeira.
Hogna carolinensis
Carolina wolf spider, giant wolf spider
Hogna carolinensis, the Carolina wolf spider, is the largest wolf spider in North America. Females reach 22–35 mm in body length, males 18–20 mm. This species inhabits self-made or appropriated burrows in open habitats across a broad geographic range. Females carry egg sacs attached to their abdomen during the approximately two-week incubation period, and spiderlings cling to the mother's back for several days after hatching. The species is notable for its eye shine when illuminated at night, effective thermoregulation abilities, and unique venom containing lycotoxins with antimicrobial properties.
Hogna lenta
Field Wolf Spider
Hogna lenta is a species of wolf spider in the family Lycosidae, commonly known as the Field Wolf Spider. It is native to the southeastern United States, with particular abundance in Florida. This large, ground-dwelling spider constructs vertical burrows in sandy substrates and hunts using vibration detection. The species has been used in neuroanatomical research to study catecholaminergic neuron distribution in spider central nervous systems.
Homalonychus selenopoides
Homalonychus selenopoides is a true spider in the family Homalonychidae, found in desert regions of the southwestern United States and northwestern Mexico. Males perform distinctive courtship behaviors involving vibratory drumming on the substrate and spinning silk threads embedded with sand grains. Females construct silk retreats incorporating sand for egg-laying and exhibit maternal care by guarding egg sacs.
Labia minor
lesser earwig, small earwig
Labia minor is a small earwig species, 4–7 mm in length including pincers, distinguished by its dull yellow-brown to chocolate brown coloration and less shiny appearance compared to the common earwig. It is notable as the first Dermapteran species to have its genome sequenced by the Darwin Tree of Life project. The species exhibits exceptional maternal care behavior, with females guarding eggs and feeding offspring for 1–2 weeks after hatching—a rarity among insects. Unlike many earwigs, it flies frequently and possesses visible hind wings.
Labidura
striped earwigs
Labidura is a genus of earwigs in the family Labiduridae, established by Leach in 1815. The genus includes approximately nine species, with Labidura riparia being the most widespread and cosmopolitan member. Labidura riparia is notable as a predator of agricultural pests, including the Asian corn borer and Mediterranean fruit fly pupae. The Saint Helena earwig (Labidura herculeana), now possibly extinct, was the largest known earwig species. Several Labidura species exhibit subsocial maternal care of eggs and young.
Labidura riparia
shore earwig, tawny earwig, striped earwig, riparian earwig
Labidura riparia is a cosmopolitan earwig species found primarily in tropical to subtropical regions. It exhibits complex subsocial behavior with maternal care of eggs and young. The species is a generalist predator valued for biological pest control, with documented predation on Lepidoptera larvae, insect eggs, and crop pests including the Asian corn borer and Mediterranean fruit fly pupae. Males possess two functional penises but show a 90% preference for the right penis during copulation, a behavioral asymmetry unique among earwigs.
Labiduridae
striped earwigs
Labiduridae is a family of earwigs commonly known as striped earwigs, containing approximately 72 species across seven genera in three subfamilies. Members are moderate to large in size with cylindrical bodies, well-developed wings, and notably long antennae. The family exhibits cosmopolitan distribution and includes well-known species such as Labidura riparia (tawny earwig) and Nala lividipes. Many species display subsocial maternal care and predatory habits, with some serving as biological control agents of agricultural pests.
Lycosidae
Wolf Spiders
Wolf spiders (family Lycosidae) are robust, agile hunters characterized by excellent eyesight and solitary behavior. The family is distinguished by a unique eye arrangement featuring two large, forward-facing posterior median eyes that produce distinctive reflective eyeshine. Most species are ground-dwelling and do not construct webs for prey capture, instead actively hunting or ambushing prey. Females carry egg sacs attached to their spinnerets and subsequently transport spiderlings on their backs. The family exhibits considerable ecological diversity, with species occupying habitats ranging from sandy beaches to forest floors and agricultural fields.
Mastigoproctus
whip scorpion, vinegaroon
Mastigoproctus is a genus of large whip scorpions (vinegaroons) in the family Thelyphonidae. These arachnids are characterized by their elongated antenniform front legs, robust pedipalps, and ability to spray defensive acetic acid. The genus includes approximately 25 accepted species distributed across the Americas, from the southwestern United States through Mexico, Central America, the Caribbean, and into northern South America. Despite their intimidating appearance, they are not venomous and lack venom glands entirely.
Mastigoproctus giganteus
giant whip scorpion, giant vinegaroon, grampus, Texas giant vinegaroon
Mastigoproctus giganteus is a large arachnid in the order Uropygi, commonly known as the giant whip scorpion or vinegaroon. It is the only member of family Thelyphonidae found in the United States, occurring from the southern U.S. through Mexico. The species is nocturnal and occupies humid microhabitats in arid and semi-arid environments. It is notable for its ability to spray a defensive secretion containing 85% acetic acid, which produces a strong vinegar-like odor. The species is long-lived, with females producing a single brood of approximately 40 offspring that receive extended maternal care.
Melanaethus crenatus
Melanaethus crenatus is a burrower bug (Hemiptera: Cydnidae) described by Signoret in 1883. The species exhibits distinctive maternal behavior in which females carry seeds to oviposition sites, a trait documented in a detailed study of its immature stages. The egg and five nymphal instars have been formally described.
Membracidae
treehoppers, thorn bugs, typical treehoppers
Membracidae, commonly called treehoppers or thorn bugs, is a family of approximately 3,200–3,500 species in over 400 genera within the order Hemiptera. The family is distinguished by extraordinary morphological diversity, particularly the pronounced enlargement and modification of the pronotum—the dorsal plate of the first thoracic segment—which can form thorns, horns, keels, or bizarre projections that often obscure the body and wings. This family represents the most diverse lineage within the superfamily Membracoidea, with its center of diversity in the New World tropics. Most species exhibit phytophagous habits, feeding on plant sap with piercing-sucking mouthparts. Many species engage in mutualistic relationships with ants, which protect them from predators in exchange for honeydew.
Mesomphaliini
tortoise beetles
Mesomphaliini is a tribe of tortoise beetles (Cassidinae) erected by Hope in 1840, though some authorities recognize the alternative name Stolaini (Hincks, 1952). The tribe is predominantly Neotropical, with most species occurring in the Americas. Some genera, including Chelymorpha, Stolas, and Cyrtonota, have representatives extending into Europe and Japan. Members exhibit distinctive larval behaviors including construction of exuvio-fecal shields and documented subsocial maternal care in at least some species.