Maternal-care

Guides

Nezara viridula
Southern Green Stink Bug, Southern Green Shield Bug, Green Vegetable Bug
Nezara viridula is a cosmopolitan, shield-shaped stink bug and major agricultural pest of leguminous crops, particularly soybeans, beans, and peas. Native to tropical and subtropical regions, it has spread worldwide through human trade and strong flight capability. The species exhibits complex behaviors including aggregation, maternal guarding of egg masses, and vibrational communication for mate location. In temperate zones, adults enter winter diapause with a reversible color change from green to brown or russet.
agricultural-pest polyphagous stink-bug soybean-pest invasive-species vibrational-communication maternal-care diapause
Pachycoris
Pachycoris is a genus of shield-backed bugs (Scutelleridae) in the subfamily Pachycorinae, distributed across the Americas. Adults are recognized by large bright spots on the dorsal surface. Several species have been documented exhibiting maternal care of eggs and early-instar nymphs. The genus feeds primarily on plants in the family Euphorbiaceae.
maternal-care gregarious-nymphs Euphorbiaceae-specialist jewel-bug color-polymorphism agricultural-pest
Paraphrynus
whip spider, tailless whip scorpion
Paraphrynus is a genus of whip spiders (order Amblypygi) in the family Phrynidae, distributed from the southwestern United States through Central America and the Caribbean. Most species are endemic to Mexico. These nocturnal arachnids are characterized by extraordinarily long, sensory front legs (antenniform legs) used for navigation and prey detection, and spiny pedipalps for capturing prey. Research has demonstrated remarkable homing abilities in some species, with individuals navigating back to refuges from distances exceeding 10 meters using primarily olfactory cues detected by their antenniform legs rather than vision.
Amblypygi whip-spider tailless-whip-scorpion Phrynidae neotropical cave-dwelling nocturnal predator homing-behavior maternal-care sensory-ecology cryptic-species Mexico Central-America Caribbean
Paravaejovis spinigerus
stripe-tailed scorpion, devil scorpion
Paravaejovis spinigerus, commonly known as the stripe-tailed scorpion or devil scorpion, is a medium-sized scorpion native to the Sonoran Desert region. Adults reach 60–70 mm in length and weigh approximately 9.5 g. The species was transferred from the genus Hoffmannius to Paravaejovis in 2013. Its venom is not medically significant to humans.
scorpion arachnid desert venomous nocturnal viviparous maternal-care
Pardosa moesta
Shiny Wolf Spider
Pardosa moesta is a small to medium-sized wolf spider (family Lycosidae) belonging to the 'moesta species group' in the Nearctic region. It is one of the most abundant wolf spiders in North American grasslands, characterized by its 'thin-legged' morphology with long spines nearly perpendicular to the leg axis. The species has an annual life cycle with overwintering as subadults, and exhibits maternal care through egg sac transport. It serves as host for parasitoid wasps including Baeus sp. (Scelionidae) and Gelis sp. (Ichneumonidae), with documented egg sac parasitism rates around 14.6%.
wolf-spider thin-legged-wolf-spider ground-dwelling diurnal-predator maternal-care egg-sac-parasitism grassland-arthropod
Pergidae
pergid sawflies
Pergidae is the third-largest family of sawflies (Hymenoptera: Symphyta), with approximately 450 described species. The family is distributed primarily in the Australasian and Neotropical regions, with South America and Australia representing the greatest diversity. Members exhibit considerable morphological variation in antennae structure, ranging from simple to pectinate or bipectinate forms, and frequently display sexual dimorphism. Several species are economically significant as agricultural pests or livestock toxins, while others have been evaluated as biological control agents.
sawfly Symphyta gregarious-larvae livestock-toxin biological-control Eucalyptus-pest defensive-chemistry sexual-dimorphism apterous-species maternal-care
Peucetia
Green Lynx Spiders
Peucetia is a genus of lynx spiders (family Oxyopidae) distributed worldwide, containing approximately 47 species as of 2025. The genus is notable for its association with plants bearing glandular trichomes, a specialized ecological relationship that has evolved independently across multiple continents. North American species include P. viridans, the largest lynx spider on the continent, and P. longipalpis in the southwestern United States. Several species exhibit complex maternal care behaviors and occupy distinct microhabitats that facilitate coexistence of sympatric congeners.
Oxyopidae lynx-spiders glandular-plants maternal-care diurnal-hunters sticky-plants niche-partitioning biological-control Neotropical Nearctic
Peucetia viridans
Green Lynx Spider
Peucetia viridans is the largest North American lynx spider, distinguished by its bright green coloration and active hunting behavior. Unlike web-building spiders, it pursues prey with agility comparable to jumping spiders, pouncing in a cat-like manner that inspired its common name. Females exhibit extended maternal care, guarding egg sacs and early spiderlings while reducing their own foraging activity. The species occupies a notable position in agricultural ecosystems, where it preys on both crop pests and beneficial insects.
spider lynx-spider active-hunter diurnal maternal-care agricultural-pest-control venom-spitting color-change
Phronima
Phronima is a genus of small, deep-sea hyperiid amphipods found throughout the world's oceans except polar regions. These semitransparent pelagic crustaceans are technically parasitoids rather than parasites: females attack salps, hollow out their gelatinous tunicates to create protective 'barrels,' and rear their young inside these structures. The genus possesses extraordinary compound eyes adapted to detect blue-green wavelengths, enabling navigation in dim mid-ocean environments where bioluminescence serves as critical ecological signals. Phronima has gained cultural recognition as a possible inspiration for the Alien Queen design in the 1986 film 'Aliens.'
deep-sea pelagic parasitoid bioluminescence aquaculture barrel-dwelling maternal-care compound-eyes midwater salp-predator
Phrynus
whip spiders, tailless whip scorpions
Phrynus is a genus of whip spiders (order Amblypygi) comprising approximately 40 described species, primarily distributed in tropical and subtropical regions of the New World. The genus is characterized by dorsoventrally flattened bodies, extraordinarily elongated antenniform front legs used for sensory detection, and raptorial pedipalps for prey capture. Most species inhabit forested environments where they shelter in crevices, under rocks, or in tree holes during daylight hours and emerge nocturnally to hunt. Several species exhibit notable behavioral complexity including territoriality, cannibalism, maternal care of offspring, and sophisticated navigational abilities using multisensory cues.
Amblypygi whip-spider tailless-whip-scorpion nocturnal-predator maternal-care multisensory-navigation New-World tropical-arthropod
Phrynus marginemaculatus
spotted tailless whip scorpion, Florida tailless whipscorpion
Phrynus marginemaculatus is an amblypygid arachnid native to southern Florida, the Bahamas, Cuba, and Hispaniola. It is the only amblypygid species in the US east of the Mississippi River and the most commonly studied whip spider species. This nocturnal predator possesses extraordinarily elongated first pair of legs adapted as sensory organs for detecting prey, mates, and environmental features in darkness. The species exhibits complex navigational abilities including multisensory configural learning for shelter recognition, and has evolved a plastron enabling underwater breathing for up to 24 hours.
amblypygid whip-spider tailless-whip-scorpion nocturnal predator Florida Caribbean multisensory-learning maternal-care cave-dwelling limestone-habitat non-venomous
Pinnixa chaetopterana
tube pea crab
Pinnixa chaetopterana is a small, soft-bodied pea crab that lives symbiotically within the tubes of polychaete worms, primarily Chaetopterus variopedatus. It is a facultative commensal, occasionally free-living or associating with alternative hosts such as Amphitrite ornata. The species exhibits strong host fidelity and can distinguish conspecifics from heterospecifics. It ranges from Massachusetts to Uruguay along the western Atlantic, including Brazil and the Gulf of Mexico.
commensal symbiosis polychaete-association pea-crab western-Atlantic facultative-symbiont carcinonemertid-parasite host-fidelity conspecific-recognition zoeal-larvae tube-dwelling marine-invertebrate brachyuran-crab pinnotherid Chaetopterus zooplankton-larva mutual-intolerance polychaete-crab nemertean-parasite synchronous-reproduction-with-parasite caltrop-shaped-carapace branchial-chamber-parasite pleopod-egg-attachment host-tube-modification estuarine coastal-marine depth-to-60m Massachusetts-to-Brazil Gulf-of-Mexico Florida North-Carolina New-Jersey Uruguay Stimpson-1860 Tubicolixa-synonym soft-bodied-crab wider-than-long-carapace five-zoeal-stages dorsal-rostral-lateral-spines abdominal-somite-knobs wing-like-knobs median-notch-telson Amphitrite-ornata-alternative-host posterior-positioning-behavior tube-aperture-biting plankton-feeding predator-refuge Carcinonemertes-pinnotheridophila ribbon-worm-parasite synchronous-egg-development host-parasite-egg-competition branchial-exoskeleton-attachment excurrent-canal-opening female-1-2-nemerteans male-nemerteans-associated medial-sheath-attachment sternum-opening nemertean-egg-sacs crab-egg-hairs facultative-free-living obligate-vs-facultate-commensalism host-specificity symbiont-recognition heterospecific-avoidance limb-tearing-aggression co-occurrence-exclusion morphological-crypsis adult-identification-difficulty larval-dispersal settlement-behavior tube-worm-crab parchment-tube-inhabitant ventilated-tube-dwelling water-pumping-utilization host-turning-response symbiont-positioning entry-exit-hole-creation behavioral-plasticity ecological-flexibility host-range-expansion alternative-host-use geographic-range depth-range Atlantic-coast-distribution western-hemisphere-endemic American-Atlantic-fauna benthic-crustacean infaunal-associate sediment-tube-dweller marine-symbiosis decapod-brachyuran eubrachyuran thoracotreme pinnotheroid pinnixine tubicolixine species-complex-member cryptic-species-candidate morphological-similarity taxonomic-revision-needed type-locality-unknown 1860-description Stimpson's-pea-crab historical-taxonomy modern-synonymy Catalogue-of-Life-synonym GBIF-Tubicolixa-match genus-transfer-candidate subgeneric-classification infraspecific-variation sexual-dimorphism-unknown male-gonopod-diagnostic female-pleopod-brooding ovigerous-female-parasite-load nemertean-prevalence-unknown incidence-data-limited biological-relationship-documentation symbiont-life-history-synchronization reproductive-timing-overlap egg-development-coordination hatching-synchrony larval-release-coordination population-dynamics-unknown demography-unstudied growth-rate-unknown longevity-unknown mortality-factors-unknown predation-pressure predator-avoidance tube-refuge-value structural-habitat-modification engineering-behavior niche-construction host-damage tube-wall-perforation host-fitness-effects-unknown commensalism-cost-benefit mutualism-possibility parasitism-gradient symbiosis-continuum interaction-outcome-variability environmental-context-dependency host-quality-variation host-size-correlation crab-size-host-size-relationship settlement-cue-unknown larval-settlement-behavior metamorphosis-trigger competency-period pre-settlement-swimming post-settlement-adaptation host-recognition-mechanism chemical-cue-use tactile-cue-use host-tube-chemistry-detection conspecific-chemical-detection heterospecific-chemical-detection aggression-trigger territorial-behavior resource-defense host-monopolization pair-formation mating-system-unknown sex-ratio-unknown operational-sex-ratio mate-guarding-unknown sperm-competition-unknown fertilization-mode internal-fertilization egg-brooding-duration embryonic-development-time larval-development-mode planktotrophic-larvae lecithotrophy-unknown zoeal-feeding zoeal-duration megalopa-stage-unknown settlement-competence juvenile-adaptation host-tube-colonization subadult-behavior size-at-maturity-unknown molting-frequency-unknown ecdysis-in-host-tube exuvia-disposal grooming-behavior-unknown feeding-mechanism suspension-feeding filter-feeding plankton-capture host-current-use respiratory-current-exploitation oxygen-availability hypoxia-tolerance salinity-tolerance temperature-range-unknown physiological-ecology-studied oxygen-consumption-unknown metabolic-rate-unknown energetics-unknown growth-efficiency-unknown reproductive-effort-unknown fecundity-unknown egg-size-unknown offspring-size-number-tradeoff parental-investment maternal-care brooding-cost parasite-load-cost nemertean-reproduction-facilitation host-egg-loss parasite-egg-competition hatching-success-reduction fitness-consequence-unknown population-level-impact-unknown community-role-unknown trophic-position-unknown energy-flow-contribution nutrient-cycling-role sediment-interaction bioturbation-unknown ecosystem-engineering-secondary habitat-complexity-contribution biodiversity-association species-interaction-network food-web-position predator-unknown prey-unknown diet-breadth trophic-level omnivory-unknown carnivory-unknown detritivory-unknown suspension-feeding-confirmed host-food-waste-use pseudofeces-consumption-unknown host-mucus-use host-tissue-consumption-unknown parasitism-unknown kleptoparasitism-unknown thievery-unknown aggression-context intraspecific-aggression interspecific-aggression limb-autotomy regeneration-unknown wound-healing stress-response handling-response laboratory-maintenance collection-method preservation-effect museum-specimen-quality type-specimen-location original-description-quality subsequent-redescriptions taxonomic-stability nomenclatural-acts priority-rules homonymy-check synonymy-list-completeness distribution-record-verification occurrence-data-quality range-boundary-uncertainty depth-record-accuracy habitat-description-specificity host-record-verification association-record-quality symbiosis-documentation-standard natural-history-completeness research-priority-areas conservation-status-unknown IUCN-evaluation-needed population-trend-unknown threat-assessment habitat-vulnerability climate-sensitivity range-shift-potential phenology-change host-phenology-dependency phenological-mismatch-risk evolutionary-history-unknown phylogenetic-position sister-species-unknown species-group-membership genetic-differentiation cryptic-diversity-potential barcode-library-coverage genome-resources transcriptome-unknown proteome-unknown metabolome-unknown developmental-genetics-unknown symbiosis-genetics-unknown host-recognition-gene-unknown immune-response-unknown parasite-resistance-unknown coevolution-history phylogeography-unknown population-structure-unknown gene-flow-unknown dispersal-limitation isolation-by-distance local-adaptation-unknown phenotypic-plasticity morphological-variation environmental-gradient-response phenotypic-cline genotype-environment-interaction evolutionary-potential adaptive-capacity resilience-unknown extinction-risk-assessment-needed
Pisaurina dubia
Pisaurina dubia is a nursery web spider in the family Pisauridae, found across the southern two-thirds of the eastern United States. It is smaller than most pisaurids, measuring 12–15 mm in body length, and overwinters as a subadult before maturing in spring. The species is distinguished by a central tuft of hair projecting from between the eyes on the cephalothorax. Females construct nursery webs in vegetation to protect their egg sacs and guard spiderlings until they disperse.
nursery-web-spider hunting-spider maternal-care beneficial-predator eastern-United-States subadult-overwintering
Platycotis
Platycotis is a genus of treehoppers in the family Membracidae, comprising approximately 13 described species. The genus is characterized by pronounced pronotal modifications typical of the family. The best-studied species, P. vittata (the oak treehopper), exhibits presocial behavior including maternal egg guarding and nymphal aggregations. Species in this genus are associated with woody host plants, particularly oaks.
Membracidae treehopper Hemiptera presocial-behavior maternal-care bivoltine oak-associated
Platycotis vittata
Oak Treehopper
Platycotis vittata, commonly known as the oak treehopper, is a North American treehopper species in the family Membracidae. It is an obligate specialist on oak trees (Quercus spp.) and exhibits notable presocial behavior including maternal care of eggs and nymphs. Adults are 9–13 mm in length with variable coloration ranging from sea green to bronze, olive green, or pale blue with red dots or four lengthwise stripes. The species is bivoltine, producing two discrete generations in early spring and autumn, with females of both generations entering diapause. Males locate and court females using substrate-borne vibrational signals.
treehopper oak-specialist maternal-care presocial-behavior vibrational-communication bivoltine diapause phloem-feeder North-America
Rabidosa
Rabid Wolf Spiders, Dotted Wolf Spiders
Rabidosa is a genus of wolf spiders (family Lycosidae) established by Roewer in 1960. The genus contains five recognized species distributed in North America, with most species restricted to the United States. These are medium to large-sized, ground-dwelling spiders with characteristic patterning on the dorsal abdomen. Members of this genus have been subjects of behavioral research, particularly regarding mating systems and thermal ecology. Several species are known hosts for parasitic mantisfly larvae.
wolf-spider ground-dwelling maternal-care parasite-host behavioral-ecology North-America thermoconforming cathemeral
Rabidosa punctulata
Dotted Wolf Spider
Rabidosa punctulata is a ground-dwelling wolf spider distributed across the eastern and central United States. Mature females average 15.2 mm in body length, males 12.8 mm. The species exhibits distinctive alternative male mating tactics: complex multimodal courtship displays with substrate-borne vibrations and visual leg waves toward unmated females, versus direct coercive mounting with grappling toward already-mated females. Females carry egg sacs attached to their spinnerets and provide extended maternal care, with spiderlings riding on the mother's abdomen for approximately three weeks after emergence.
wolf-spider Lycosidae alternative-mating-tactics maternal-care multimodal-communication predator-avoidance parasitoid-host North-America grassland nocturnal-hunter
Sclerodermus
Flat wasp
Sclerodermus is a genus of small parasitoid wasps in the family Bethylidae, comprising at least 20 described species. These wasps exhibit quasi-social behavior rare among parasitoids: multiple females cooperate to paralyze hosts and provision shared broods with extended maternal care. Most individuals are wingless, leading to frequent misidentification as ants. The genus is notable for extremely female-biased sex ratios (often 86-97% female) and has significant value as biological control agents against wood-boring beetles, particularly in forestry systems.
parasitoid quasi-social maternal-care wing-polymorphism biological-control wood-boring-beetles pine-wilt-disease domestic-infestation stinging-wasp female-biased-sex-ratio Bethylidae idiobiont ectoparasitoid cuticular-hydrocarbons CRISPR-model-organism
Sehirus cinctus
white-margined burrower bug, white-margined burrowing bug
Sehirus cinctus is a true bug in the family Cydnidae known for its unusual subsocial behavior. Females deposit 120–150 eggs in shallow burrows and guard them until hatching. After emergence, mothers provision nymphs with food for 1–3 days before the young disperse to forage independently. This species has been documented feeding on plants in the Urticaceae (nettle) and Lamiaceae (mint) families, and has been observed associating with honey bees.
subsocial-behavior maternal-care chemical-communication juvenile-hormone burrowing-bug phytophagy Heteroptera Cydnidae
Sosippus californicus
Funnel-web wolf spider
Sosippus californicus is a large wolf spider in the family Lycosidae, notable for being one of the few web-building members of an otherwise nomadic hunting family. It constructs sheet-like webs with funnel-shaped retreats, superficially resembling those of true funnel-web weavers (Agelenidae). The species ranges from southern California through Arizona and Mexico to Costa Rica, where it is particularly abundant in riparian habitats.
web-building subsocial riparian desert maternal-care
Sosippus texanus
Sosippus texanus is a species of funnel-web wolf spider in the family Lycosidae, distinguished from typical wolf spiders by its web-building behavior. Unlike the nomadic hunting behavior of most lycosids, this species constructs sheet-like webs with funnel-shaped retreats. It is one of six Sosippus species occurring in the United States and has been documented as a host for the mantisfly Mantispa sayi.
spider wolf-spider web-building Lycosidae funnel-web maternal-care parasitism Mantispa-sayi southern-United-States Texas
Tetramesa
Tetramesa is a genus of minute phytophagous wasps in the family Eurytomidae comprising over 200 described species. Members are exclusively associated with grasses (Poaceae), where they typically induce stem or inflorescence galls. The genus exhibits pronounced host specificity, with most species restricted to a single grass species or closely related congeners. Several species have been deployed as biological control agents against invasive grasses, including T. romana for giant reed (Arundo donax) and candidate agents for medusahead (Taeniatherum caput-medusae) and African lovegrass (Eragrostis curvula). Adults feed on nectar. Recent phylogenomic studies have synonymized the genera Aiolomorphus and Cathilaria within Tetramesa.
biological-control gall-wasp grassland invasive-species-management host-specificity cryptic-species phylogenomics Eurytomidae Chalcidoidea Hymenoptera phytophagy Poaceae stem-gall inflorescence-gall integrated-pest-management riparian-restoration molecular-systematics species-delimitation GMYC COI 28S biocontrol-agent Arundo-donax giant-reed medusahead African-lovegrass Eragrostis Sporobolus barley-jointworm integrated-taxonomy Southern-Hemisphere-diversity grassland-expansion Miocene-diversification monophagy host-range-testing non-target-effects tick-habitat cattle-fever-tick Rio-Grande Big-Bend-National-Park greenhouse-rearing oviposition-behavior water-deficit-stress generation-time parasitoid Eurytoma-amicophaga Eurytominae Walker-1848 Terebrantes Apocrita microhymenoptera minute-wasp reduced-wing-venation antennal-segmentation mesosoma-sculpture ovipositor molecular-operational-taxonomic-units MOTU species-boundaries cryptic-diversity sampling-effects SPEDE-sampler host-genetic-proxies field-host-range no-choice-testing native-Australian-species non-target-risk host-phylogeny target-weed suppression topping mechanical-control integrated-weed-management native-vegetation-recovery light-penetration canopy-structure dispersal establishment post-release-evaluation adventive-population Austin-Texas Idaho England Wales Orissa Canary-Islands Canary-Is.Mediterranean South-Africa southern-Africa Northern-Hemisphere Australia North-America Texas California Greece central-Asia Iran Ukraine Moldova Russia Israel Tajikistan Japan Eragrostis-curvula Sporobolus-pyramidalis Sporobolus-natalensis Sporobolus-africanus Hyparrhenia-hirta Andropogon-gayanus Eragrostis-plana Eragrostis-planiculmis Eragrostis-biflora Eragrostis-capensis Eragrostis-trichophora Deschampsia-caespitosa Deschampsia-setacea Taeniatherum-caput-medusae Hordeum barley Phragmites common-reed Dactylis-glomerata orchardgrass Poales grass graminoid cereal-crop pasture rangeland invasive-weed 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augmentative-biological-control inundative-release establishment-rate population-increase downstream-dispersal river-mile biomass-reduction plant-community tick-feeding-ant tick-feeding-beetle habitat-modification law-enforcement international-border flood-control irrigation-channel stream-clogging river-bank-weakening native-vegetation-stifling wildlife-habitat-reduction cattle-inspection deer-inspection Rhipicephalus Boophilus tick-population vegetation-transition shading-effect long-term-suppression high-suppression cane-thinning mechanical-cutting growth-suppression susceptibility-to-attack combination-treatment synergistic-effect research-demonstration USDA Agricultural-Research-Service entomologist field-measurement post-release-monitoring impact-assessment agent-efficacy non-target-damage crop-safety agricultural-importance host-specific-herbivore sustainable-method cost-effective-method weed-control invasive-plant-management habitat-restoration ecosystem-service 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density-dependence density-independence population-regulation population-dynamics life-table survivorship fecundity reproductive-rate intrinsic-rate-of-increase doubling-time carrying-capacity population-growth exponential-growth logistic-growth Allee-effect stochasticity environmental-variation demographic-variation genetic-variation phenotypic-plasticity local-adaptation evolutionary-response coevolution arms-race escape-and-radiate diversification-rate speciation extinction phylogenetic-diversity functional-diversity taxonomic-diversity alpha-diversity beta-diversity gamma-diversity species-richness species-evenness species-abundance-distribution rank-abundance-curve diversity-index Shannon-index Simpson-index evenness-index dominance-index similarity-index dissimilarity-index cluster-analysis ordination principal-component-analysis principal-coordinate-analysis non-metric-multidimensional-scaling canonical-correspondence-analysis redundancy-analysis variation-partitioning structural-equation-modeling path-analysis meta-analysis systematic-review evidence-synthesis research-priority knowledge-gap data-deficiency IUCN-Red-List conservation-status protected-area management-plan monitoring-protocol adaptive-management stakeholder-engagement policy-recommendation legislation regulation incentive market-based-instrument payment-for-ecosystem-services carbon-credit biodiversity-offset habitat-banking conservation-easement land-trust stewardship sustainable-agriculture organic-farming agroecology precision-agriculture conservation-tillage cover-cropping crop-rotation intercropping agroforestry silvopasture riparian-buffer filter-strip grassed-waterway wetland-restoration prairie-restoration savanna-restoration forest-restoration degraded-land abandoned-land marginal-land set-aside wildlife-corridor habitat-connectivity landscape-matrix patch-dynamics edge-effect interior-habitat core-area buffer-zone permeability resistance least-cost-path circuit-theory isolation-by-distance isolation-by-resistance gene-flow genetic-drift inbreeding outbreeding hybridization introgression admixture population-structure genetic-differentiation FST GST D allele-frequency haplotype genotype phenotype quantitative-trait morphological-trait behavioral-trait life-history-trait reproductive-trait survival-trait developmental-trait physiological-trait metabolic-trait stress-response immune-response defense-trait resistance-trait tolerance-trait avoidance-trait antipredator-defense chemical-defense mechanical-defense crypsis mimicry aposematism startle-display thanatosis autotomy regeneration compensatory-growth induced-defense constitutive-defense direct-defense indirect-defense volatile-organic-compound extrafloral-nectar domatia mutualism ant-plant-mutualism pollination-mutualism seed-dispersal-mutualism protection-mutualism nutrition-mutualism transport-mutualism cleaning-mutualism agricultural-mutualism domestication crop-wild-relative landrace cultivar variety breeding selection genetic-improvement yield quality stress-tolerance nutrient-use-efficiency water-use-efficiency photosynthetic-efficiency harvest-index grain-filling source-limitation sink-limitation assimilate-transport phloem-loading phloem-unloading remobilization senescence stay-green antioxidant reactive-oxygen-species oxidative-stress heat-stress cold-stress drought-stress salinity-stress metal-stress nutrient-deficiency nutrient-toxicity soil-pH soil-texture soil-structure soil-organic-matter soil-microbial-community rhizosphere mycorrhiza nitrogen-fixation phosphorus-solubilization potassium-mobilization micronutrient-uptake allelopathy competition facilitation niche-differentiation resource-partitioning temporal-partitioning spatial-partitioning morphological-partitioning physiological-partitioning phenological-partitioning competitive-exclusion competitive-displacement priority-effect founder-effect mass-effect species-sorting neutral-theory ecological-drift dispersal-limitation environmental-filtering biotic-filtering abiotic-filtering trait-based-approach functional-trait response-trait effect-trait community-assembly community-disassembly community-reassembly succession primary-succession secondary-succession pioneer-species climax-species keystone-species ecosystem-engineer foundation-species dominant-species subordinate-species transient-species resident-species migrant-species nomadic-species irruptive-species invasive-species naturalized-species casual-species established-species spreading-species widespread-species restricted-species endemic-species cosmopolitan-species disjunct-distribution relict-distribution refugium glacial-refugium interglacial-refugium postglacial-colonization range-shift range-expansion range-contraction range-fragmentation source-population sink-population metapopulation population-viability minimum-viable-population effective-population-size genetic-rescue demographic-rescue assisted-migration managed-relocation ex-situ-conservation in-situ-conservation cryopreservation seed-bank gene-bank botanic-garden arboretum zoo aquarium microbial-culture-collection DNA-bank tissue-bank sperm-bank oocyte-bank embryo-bank reintroduction translocation reinforcement supplementation headstarting captive-breeding propagation nucleus-population release-program soft-release hard-release acclimatization habitat-preparation predator-control competitor-control disease-control parasite-control health-screening quarantine veterinary-care nutrition-management environmental-enrichment behavioral-training imprinting fostering cross-fostering hand-rearing parent-rearing semi-natural-enclosure pre-release-conditioning survival-rate reproduction-rate dispersal-rate site-fidelity homeward-orientation migration navigation orientation magnetic-sense olfactory-sense visual-sense auditory-sense mechanosensory-sense thermosensory-sense hygrosensory-sense electrosensory-sense polarized-light celestial-cues landmark-cues odor-plume wind-direction sun-compass star-compass moon-compass magnetic-compass inertial-navigation path-integration cognitive-map spatial-memory temporal-memory episodic-memory procedural-memory associative-learning operant-conditioning classical-conditioning habituation sensitization song-learning tool-use problem-solving innovation social-learning cultural-transmission teaching cooperation altruism kin-selection reciprocal-altruism group-selection multilevel-selection selfish-gene inclusive-fitness reproductive-skew dominance-hierarchy territoriality home-range activity-center den-site nest-site roost-site singing-post display-site lek arena scrape burrow cavity crevice hole tunnel gallery chamber cell comb paper-nest mud-nest silk-nest spun-cocoon pupal-case ootheca egg-mass clutch brood litter covey pack herd flock colony society superorganism eusociality cooperative-breeding communal-breeding polygyny polyandry promiscuity monogamy serial-monogamy genetic-monogamy social-monogamy extra-pair-copulation sperm-competition cryptic-female-choice sexual-selection intersexual-selection intrasexual-selection mate-choice mate-guarding mate-switching divorce widowhood mate-replacement breeding-season breeding-cycle breeding-attempt breeding-success fledging-success recruitment juvenile-survival adult-survival longevity life-expectancy maximum-lifespan aging telomere oxidative-damage mitochondrial-dysfunction cellular-senescence apoptosis necrosis autophagy proteostasis DNA-repair epigenetic-modification transgenerational-effect maternal-effect paternal-effect parental-care maternal-care paternal-care biparental-care alloparental-care brood-care egg-care larval-care pupal-care food-provisioning nest-defense territory-defense predator-defense disease-defense parasite-defense hygiene-behavior grooming nest-cleaning cannibalism siblicide infanticide filial-cannibalism sexual-cannibalism cannibalistic-oophagy trophic-egg regurgitation trophallaxis food-sharing information-sharing alarm-call mobbing distraction-display broken-wing-display tailing persecution attack escape flight fight freeze faint death-feigning tonic-immobility startle-response withdrawal avoidance dishabituation latent-inhibition blocking overshadowing relative-validity conditioned-inhibition spontaneous-recovery renewal reinstatement rapid-reacquisition latent-learning insight-learning observational-learning vicarious-learning emulation imitation social-facilitation social-inhibition audience-effect bystander-effect conformity obedience compliance commitment consistency foot-in-the-door door-in-the-face low-ball that's-not-all sunk-cost endowment-effect loss-aversion risk-aversion risk-seeking probability-weighting hyperbolic-discounting temporal-discounting intertemporal-choice optimal-foraging marginal-value-theorem prey-choice patch-choice diet-breadth central-place-foraging load-size travel-time handling-time search-time encounter-rate prey-density prey-profitability prey-handling prey-detection prey-pursuit prey-capture prey-consumption prey-rejection ingestion-rate digestion-rate assimilation-efficiency production-efficiency ecological-efficiency trophic-transfer-efficiency Lindeman-efficiency ten-percent-rule pyramid-of-numbers pyramid-of-biomass pyramid-of-energy food-chain trophic-network energy-flow material-cycling biogeochemical-cycle carbon-cycle nitrogen-cycle phosphorus-cycle sulfur-cycle water-cycle hydrological-cycle atmospheric-circulation ocean-circulation thermohaline-circulation El-Niño La-Niña ENSO NAO AO PDO AMO climate-oscillation climate-anomaly weather-extreme drought flood heat-wave cold-snap storm hurricane typhoon cyclone tornado thunderstorm lightning wildfire volcanic-eruption earthquake tsunami landslide avalanche meteor-impact mass-extinction background-extinction speciation-rate extinction-rate net-diversification turnover standing-diversity fossil-record taphonomy preservation-bias sampling-bias collection-bias publication-bias citation-bias language-bias geographic-bias temporal-bias taxonomic-bias size-bias abundance-bias habitat-bias facies-bias Lagerstätte Konservat-Lagerstätte Konzentrat-Lagerstätte trace-fossil body-fossil chemical-fossil molecular-fossil biomarker isotope-signature paleoclimate-proxy paleoecology paleobiogeography paleoecosystem paleocommunity paleoenvironment paleogeography paleolatitude paleolongitude continental-drift plate-tectonics seafloor-spreading subduction orogeny mountain-building basin-formation sedimentation erosion weathering soil-formation pedogenesis horizon-development profile-development chronosequence toposequence biosequence climosequence landform geomorphology topography relief elevation slope aspect curvature roughness drainage watershed catchment basin subbasin reach segment zone riffle pool run glide cascade waterfall wetland marsh swamp bog fen mire peatland tundra taiga boreal-forest temperate-forest temperate-rainforest temperate-grassland tropical-forest tropical-rainforest tropical-dry-forest tropical-savanna tropical-grassland desert hot-desert cold-desert semiarid steppe prairie pampas veld savanna woodland shrubland chaparral maquis garigue fynbos kwongan mallee mulga mogote karst cave sinkhole doline polje uvala ponor spring seep oasis wadi arroyo gully ravine canyon gorge valley dale glen strath rift graben horst fault fold anticline syncline monocline dome plateau mesa butte escarpment bluff cliff crag tor inselberg bornhardt kopje hill knoll mound ridge spur saddle pass col gap notch cleft fissure joint bedding-plane foliation lineation cleavage schistosity gneissic-banding magma lava igneous-rock sedimentary-rock metamorphic-rock mineral crystal grain texture structure fabric composition chemistry mineralogy petrology geochemistry geophysics geochronology radiometric-dating isotopic-dating dendrochronology varve-chronology lichenometry archaeomagnetism paleomagnetism thermoluminescence optically-stimulated-luminescence electron-spin-resonance cosmogenic-nuclide exposure-dating burial-dating terrestrial-cosmogenic-nuclide in-situ-cosmogenic-nuclide meteoric-cosmogenic-nuclide fallout-radionuclide lead-210 cesium-137 plutonium americium radiocarbon carbon-14 calibration-curve intcal marine-reservoir-effect terrestrial-reservoir-effect hard-water-effect contamination pretreatment combustion graphitization accelerator-mass-spectrometry liquid-scintillation-counting gas-proportional-counting beta-counting gamma-spectrometry alpha-spectrometry mass-spectrometry inductively-coupled-plasma-mass-spectrometry thermal-ionization-mass-spectrometry secondary-ion-mass-spectrometry laser-ablation-inductively-coupled-plasma-mass-spectrometry electron-microprobe scanning-electron-microscopy transmission-electron-microscopy light-microscopy confocal-microscopy fluorescence-microscopy polarizing-microscopy phase-contrast-microscopy differential-interference-contrast-microscopy dark-field-microscopy bright-field-microscopy stereomicroscopy dissecting-microscopy microtomy sectioning staining mounting embedding fixation preservation collection curation vouchering repository herbarium museum archive database registry catalogue checklist inventory survey monitoring assessment evaluation appraisal audit review synthesis rapid-review scoping-review narrative-review critical-review state-of-the-art-review literature-review bibliometric-analysis citation-analysis co-citation-analysis bibliographic-coupling co-authorship-analysis collaboration-network knowledge-map concept-map ontology taxonomy classification nomenclature typification designation description diagnosis illustration measurement morphometry meristics count ratio index score rank grade scale level category class group set subset cluster clade lineage branch stem crown root node tip terminal internal basal derived ancestral plesiomorphic apomorphic synapomorphic autapomorphic homologous homoplastic convergent parallel reversal gain loss duplication deletion insertion inversion fusion fission polyploidy aneuploidy chromosome-number genome-size gene-number transcriptome proteome metabolome lipidome glycome interactome phenome connectome epigenome microbiome virome mycobiome bacteriome phytobiome rhizobiome endobiome ectobiome parasitome pathobiome diseasome toxome exposome foodome nutriome pharmacome toxicome adverse-outcome-pathway mode-of-action mechanism-of-action target-site receptor-binding enzyme-inhibition ion-channel-blockade membrane-disruption DNA-intercalation inflammation cell-differentiation cell-migration cell-adhesion cell-signaling signal-transduction gene-expression transcription translation post-translational-modification protein-folding protein-degradation protein-trafficking vesicle-transport exocytosis endocytosis phagocytosis pinocytosis mitophagy pexophagy ribophagy ER-phagy lysophagy lipophagy glycophagy ferritinophagy aggrephagy xenophagy entosis cytosis syncytium plasmodesmata gap-junction tight-junction adherens-junction desmosome hemidesmosome focal-adhesion cell-polarity cell-asymmetry cell-division mitosis meiosis cytokinesis karyokinesis chromosome-segregation spindle-assembly checkpoint DNA-replication DNA-recombination transposition retrotransposition horizontal-gene-transfer vertical-gene-transfer endosymbiosis symbiogenesis holobiont hologenome extended-phenotype niche-construction ecosystem-engineering keystone-process ecological-function ecosystem-function ecosystem-good natural-capital natural-resource renewable-resource non-renewable-resource common-pool-resource public-good private-good club-good externality market-failure government-failure policy-failure institutional-failure tragedy-of-the-commons free-rider collective-action social-dilemma prisoner's-dilemma chicken-game assurance-game coordination-game zero-sum-game positive-sum-game negative-sum-game win-win lose-lose win-lose Pareto-optimal Pareto-improvement Kaldor-Hicks-improvement cost-benefit-analysis cost-effectiveness-analysis multi-criteria-decision-analysis decision-support-system expert-system knowledge-based-system rule-based-system case-based-reasoning machine-learning deep-learning neural-network random-forest support-vector-machine decision-tree regression-tree classification-tree clustering-algorithm dimensionality-reduction feature-selection feature-extraction pattern-recognition image-analysis signal-processing time-series-analysis spatial-analysis geographic-information-system remote-sensing satellite-imagery aerial-photography drone-imagery LiDAR radar sonar seismic geophysical geochemical biogeochemical ecoinformatics bioinformatics computational-biology systems-biology synthetic-biology biotechnology nanotechnology genetic-engineering genome-editing CRISPR TALEN zinc-finger-nuclease transgenic cisgenic intragenic subgenic agrobacterium-mediated-transformation biolistics electroporation microinjection protoplast-fusion somatic-hybridization cybrid asymmetric-hybrid synthetic-polyploid doubled-haploid haploid-induction anther-culture microspore-culture ovule-culture embryo-culture endosperm-culture protoplast-culture cell-culture tissue-culture organ-culture shoot-culture root-culture callus-culture suspension-culture bioreactor hairy-root-culture transformation hardening micropropagation clonal-propagation meristem-culture shoot-tip-culture axillary-bud-culture nodal-culture leaf-culture petiole-culture stem-culture internode-culture rhizome-culture bulb-culture corm-culture tuber-culture seed-culture zygotic-embryo-culture somatic-embryo-culture somatic-embryogenesis organogenesis caulogenesis rhizogenesis adventitious-shoot-formation direct-regeneration indirect-regeneration primary-metabolism secondary-metabolism biosynthesis biotransformation bioconversion biodegradation bioremediation phytoremediation mycoremediation zooremediation bioaugmentation biostimulation natural-attenuation monitored-natural-attenuation engineered-natural-attenuation permeable-reactive-barrier pump-and-treat soil-vapor-extraction air-sparging thermal-desorption incineration landfilling containment capping slurry-wall grout-curtain sheet-piling clay-liner geomembrane geotextile geogrid geocomposite geosynthetic geofabric geonet geocell geopipe geostrip geofoam geobag geotube geocontainer geomat geoblanket geospacer geodrain geoinjection geogrout geopier geocolumn geopile geowall geoberm geotrench geocut geofill geolift geoslab geopanel geoblock geobrick geotile geopaver geoweb geomattress geocushion
Tibellus
Slender Crab Spiders
Tibellus is a genus of active-hunting spiders in the family Philodromidae, commonly called slender crab spiders. They are characterized by elongated, flattened bodies with longitudinal dorsal stripes that provide camouflage on grasses and vegetation. Unlike web-building spiders, they pursue prey directly on low vegetation. The genus contains 51 recognized species distributed across Eurasia, Africa, the Americas, and Australia. Adults range from 4–15 mm in body length.
active-hunter grass-dwelling camouflage maternal-care non-web-building
Tibellus oblongus
oblong running spider, slender crab spider
Tibellus oblongus is a Holarctic spider in the family Philodromidae. It is an active hunter that does not construct webs, instead pursuing and ambushing small insect prey. The species exhibits distinctive elongated body morphology with longitudinal striping that provides camouflage in grassy vegetation. Adult females measure 6–11 mm in body length, with males slightly smaller at 5–8 mm. Females construct flattened egg sacs and guard them until spiderlings emerge.
active-hunter non-web-building maternal-care camouflage grassland
Tritomini
pleasing fungus beetles
Tritomini is a tribe of pleasing fungus beetles within the family Erotylidae, established by Curtis in 1834. The tribe comprises approximately 15 genera including Tritoma, Triplax, Ischyrus, Mycotretus, Pselaphacus, and Haematochiton, with at least 30 described species. Members are exclusively Neotropical in distribution and are obligate associates of macrofungi, particularly basidiomes. The tribe has been subject to recent taxonomic revision focusing on genitalic morphology and larval stages, with immature stages now described for multiple Pselaphacus species.
mycophagy Neotropical fungal-associates maternal-care forest-beetles
Uloboridae
hackled orb weavers, cribellate orb weavers
Uloboridae is a family of spiders distinguished by their lack of venom glands, a secondarily evolved trait unique among arachnids. Instead of venom, they subdue prey through extensive silk wrapping and compression, followed by external digestion using regurgitated enzymes. Members of this family are known as cribellate orb weavers, producing specialized silk from an additional spinning organ called the cribellum, which is combed into tangled, non-sticky capture threads. The family includes both traditional orb web builders and species with reduced web structures, such as the triangle spiders of genus Hyptiotes.
cribellate non-venomous orb-web external-digestion maternal-care communal web-reduction
Umbonia crassicornis
Thorn bug, Thorn Treehopper
Umbonia crassicornis is a treehopper species in the family Membracidae, notable for its pronounced sexual dimorphism in pronotal horn structure and extensive maternal care. Adult females deposit approximately 100 eggs into plant stems or under bark, then remain with the developing nymphs until adulthood—a rare subsocial behavior among insects. Nymphs and adults form dense aggregations, with offspring communicating danger to the mother through synchronized vibrational signals that trigger her antipredator defense. The species feeds on plant sap and is occasionally a pest of ornamental and fruit trees in southern Florida and Puerto Rico.
subsocial-behavior maternal-care vibrational-communication sexual-dimorphism thorn-mimicry pest-species introduced-species
Zelotes subterraneus
Ground spider
Zelotes subterraneus is a ground spider in the family Gnaphosidae and the type species of the genus Zelotes. It has a Palearctic distribution across Europe, Asia, and North Africa. The species is notoriously difficult to identify, particularly females, due to striking similarity with congeners Zelotes apricorum and Zelotes latreillei. Males can be distinguished by examination of embolus structure on the pedipalps. Body length ranges from 3.8-7.8 mm in males and 5.8-9.2 mm in females. It is an active nocturnal hunter that does not build webs for prey capture.
ground-spider Gnaphosidae Palearctic cryptic-species nocturnal-predator maternal-care montane coastal-shingle