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Entedoninae
Entedoninae is a subfamily of chalcid wasps in the family Eulophidae comprising over 90 genera. Members are primarily parasitoids with documented associations across diverse host groups including gall-inducing insects, leafminers, sawflies, thrips, and ants. The subfamily exhibits considerable morphological and ecological diversity, with species found in tropical to temperate habitats worldwide.
parasitoid biological-control gall-associated leafminer-parasitoid Chalcidoidea Hymenoptera Eulophidae Entedoninae tropical-forest agricultural-pest-control sawfly-parasitoid thrips-parasitoid ant-associated rice-pest Cecidomyiidae Tenthredinidae Pontania Closterocerus Ceranisus Afronympha Ametallon Chrysonotomyia Omphale-genus-group canopy understorey ecological-plasticity Bangladesh Panama Africa New-Zealand Europe Senegal South-Africa Democratic-Republic-of-the-Congo Uganda endoparasitoid larval-parasitoid egg-parasitoid host-specificity forest-habitat degraded-habitat mid-altitude-rainforest lowland-rainforest tropical-America classical-biological-control augmentative-biological-control integrated-pest-management rice-hispa Dicladispa-armigera gall-wasp sawfly-gall leafminer ant-symbiosis Formicidae Khaya-senegalensis Ochna-pulchra morphological-taxonomy molecular-taxonomy 28S-rDNA COI CytB new-species-description generic-revision host-range parasitoid-complex food-web ecosystem-service natural-enemy pest-management entomophagous hyperparasitoid inquiline curculionid Apion-minimum Phyllocolpa-leucosticta Chrysocharis Entedon Asecodes Pleurotroppopsis Afrotroppopsis Zaommomentedon Apleurotropis Kratoysma Platocharis Schizocharis Parahorismenus Atullya Tropicharis Chrysochuris cruy mirabilis oryzamyntor eminpashai carinatum dussiae longicaudata machaeriae unimaculata claviger gorgonaense auripunctata galbina laeviscuta phenacapsia cecvora elongata pontaniae heyeri risbeci milletiae newbyi Förster-1856 Chalcidoidea-phylogeny biodiversity species-richness cryptic-species molecular-diagnostics host-plant-association habitat-specificity canopy-fauna understorey-fauna forest-stratification tropical-biodiversity Afrotropical Neotropical Palaearctic Nearctic Oriental Australasian Pacific Holarctic biogeography taxonomic-revision annotated-key catalogue type-specimen museum-collection slide-mounted-specimens morphometrics genitalia wing-venation antennae clypeus mandibles malar-space scape pedicel flagellum gaster vertex suture color-pattern pale-femora dark-gaster sexual-dimorphism male female ovipositor emergence rearing quarantine field-collection foreign-exploration augmentative-release pesticide-resistance sustainable-agriculture food-security rice-production cereal-pest vegetable-pest fruit-pest ornamental-pest forest-pest urban-pest invasive-species native-range introduced-range establishment spread impact non-target-effects host-specificity risk-assessment environmental-safety regulatory-approval research-symposium proceedings peer-reviewed Zootaxa ZooKeys Insect-Systematics-&-Evolution Australian-Journal-of-Entomology Far-Eastern-Entomologist Russian-Entomological-Journal Belgian-Journal-of-Entomology Florida-Entomologist Pan-Pacific-Entomologist Entomological-News Proceedings-of-the-Hawaiian-Entomological-Society Transactions-of-the-American-Entomological-Society Journal-of-Natural-History Zoological-Journal-of-the-Linnean-Society BioControl Journal-of-Economic-Entomology Entomologia-Experimentalis-et-Applicata Environmental-Entomology Oecologia Cladistics Molecular-Phylogenetics-and-Evolution Heredity Canadian-Entomologist Proceedings-of-the-Entomological-Society-of-Washington Annals-of-the-Entomological-Society-of-America Bulletin-of-Entomological-Research Neotropical-Entomology Journal-of-Asia-Pacific-Entomology Turkish-Journal-of-Zoology Polish-Journal-of-Entomology North-Western-Journal-of-Zoology Entomologica-Americana Entomologische-Berichten Sahlbergia Dugesiana Acta-Zoologica-Lilloana Folia-Entomologica-Mexicana Revista-de-la-Sociedad-Entomologica-Argentina Boletin-de-la-Asociacion-Espanola-de-Entomologia Entomologicheskoe-Obozrenie Entomological-Review Zoological-Institute-RAS Proceedings-of-the-Zoological-Institute-of-the-Russian-Academy-of-Sciences Taiwan-Agricultural-Research-Institute Council-of-Agriculture University-of-California Riverside 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sustainable-intensification agroecology conservation-biological-control habitat-management floral-resources refuge buffer-zone landscape-complexity connectivity spillover natural-enemy-diversity food-web-structure trophic-interactions multitrophic indirect-effects trait-mediated density-dependent inverse-density-dependent functional-response numerical-response searching-efficiency handling-time oviposition-behavior host-selection host-acceptance host-suitability host-quality host-stage-preference egg-load progeny-allocation sex-ratio local-mate-competition sib-mating inbreeding fitness longevity fecundity development-time thermal-requirements degree-days phenology synchrony asynchrony seasonal-adaptation diapause quiescence cold-hardiness heat-tolerance desiccation-resistance starvation-resistance dispersal flight walking foraging patch-use area-restricted-search oviposition-marking host-marking superparasitism multiparasitism intraguild-predation hyperparasitism autoparasitism heteronomous-hyperparasitism facultative-hyperparasitism obligate-hyperparasitism primary-parasitoid secondary-parasitoid tertiary-parasitoid mummy aphid-mummy scale-mummy pupal-parasitoid egg-larval-parasitoid larval-pupal-parasitoid idiobiont koinobiont ectoparasitoid solitary gregarious polyembryony thelytoky arrhenotoky diploid haplodiploid chromosome-number genome-size transcriptome proteome metabolome morphome phenome extended-phenotype niche-construction ecosystem-engineering keystone-species ecosystem-service-provider functional-group guild trophic-level energy-flow nutrient-cycling decomposition predation herbivory parasitism mutualism commensalism competition apparent-competition exploitative-competition interference-competition scramble-competition contest-competition resource-partitioning niche-differentiation character-displacement co-evolution arms-race Red-Queen geographic-mosaic local-adaptation phenotypic-plasticity genetic-adaptation epigenetics transgenerational-effects maternal-effects paternal-effects parental-care mating-system sexual-selection sperm-competition cryptic-female-choice mate-choice lek territory aggression alarm defense venom allomone kairomone synomone semiochemical pheromone trail-pheromone alarm-pheromone aggregation-pheromone sex-pheromone marking-pheromone oviposition-deterrent host-marking-pheromone plant-volatile induced-plant-defense systemic-acquired-resistance induced-systemic-resistance priming volatile-organic-compound green-leaf-volatile terpenoid phenylpropanoid jasmonic-acid salicylic-acid ethylene abscisic-acid gibberellin auxin cytokinin brassinosteroid strigolactone nitric-oxide reactive-oxygen-species hypersensitive-response programmed-cell-death autophagy apoptosis necrosis pathogen-associated-molecular-pattern damage-associated-molecular-pattern pattern-recognition-receptor effector-triggered-immunity gene-for-gene avirulence virulence effector toxin phytotoxin mycotoxin bacterial-toxin viral-toxin insecticidal-protein cry-protein vip-protein cyt-protein spore conidium appressorium haustorium mycelium sclerotium fruiting-body basidiocarp ascocarp perithecium cleistothecium pycnidium acervulus sporodochium symptom sign disease-triangle disease-cycle epidemic endemic pandemic outbreak eradication containment management control prevention forecasting warning-system decision-support-system expert-system model simulation economic-threshold economic-injury-level action-threshold treatment-threshold damage-threshold aesthetic-injury-level cultural-control physical-control mechanical-control chemical-control genetic-control sterile-insect-technique incompatible-insect-technique precision-agriculture site-specific-management variable-rate-application remote-sensing geographic-information-system global-positioning-system unmanned-aerial-vehicle drone sensor imaging spectroscopy machine-learning artificial-intelligence deep-learning neural-network computer-vision image-analysis pattern-recognition classification segmentation detection tracking counting sampling sampling-plan sequential-sampling binomial-sampling presence-absence-sampling fixed-precision-sampling variable-precision-sampling spatial-sampling temporal-sampling stratified-sampling cluster-sampling systematic-sampling random-sampling convenience-sampling judgment-sampling expert-sampling quota-sampling snowball-sampling respondent-driven-sampling capture-recapture mark-recapture mark-resight distance-sampling line-transect point-transect strip-transect plot quadrat pitfall-trap Malaise-trap pan-trap water-pan yellow-pan blue-pan white-pan flight-interception-trap window-trap canopy-fogging beating-tray sweep-net aerial-net ground-net aquatic-net D-vac vacuum-sampler aspirator mouth-aspirator mechanical-aspirator berlese-funnel Tullgren-funnel Winkler-extractor pitfall barber-trap pitfall-array drift-fence funnel-trap light-trap UV-light black-light mercury-vapor sodium-vapor LED actinic color-trap sticky-trap pheromone-trap synthetic-pheromone natural-pheromone kairomone-trap food-bait protein-bait sugar-bait fermentation-bait alcohol-bait vinegar-bait molasses-bait fruit-bait meat-bait fish-bait dung-bait carrion-bait decoy lure attractant repellent deterrent antifeedant oviposition-stimulant arrestant suppressant toxicant stomach-poison contact-poison fumigant systemic translaminar residual persistent non-persistent broad-spectrum narrow-spectrum selective non-selective target-specific host-specific mode-of-action nerve-poison acetylcholinesterase-inhibitor GABA-antagonist sodium-channel-modulator nicotinic-acetylcholine-receptor-agonist ryanodine-receptor-modulator chitin-synthesis-inhibitor juvenile-hormone-mimic ecdysone-agonist microbial-pesticide entomopathogenic-nematode entomopathogenic-fungus entomopathogenic-bacterium entomopathogenic-virus entomopathogenic-protozoa Bacillus-thuringiensis Bt Bacillus-sphaericus Paenibacillus-popilliae Serratia-entomophila Metarhizium-anisopliae Beauveria-bassiana Lecanicillium-lecanii Isaria-fumosorosea Nomuraea-rileyi Hirsutella-thompsonii Entomophthora Neozygites Nosema Vairimorpha Thelohania Anticarsia-gemmatalis-nucleopolyhedrovirus AgMNPV Cydia-pomonella-granulovirus CpGV Spodoptera-exigua-multiple-nucleopolyhedrovirus SeMNPV Autographa-californica-multiple-nucleopolyhedrovirus AcMNPV nucleopolyhedrovirus granulovirus cypovirus entomopoxvirus iridovirus parvovirus tetravirus nodavirus reovirus picornavirus iflavirus densovirus birnavirus totivirus partitivirus hypovirulence mycovirus bacteriophage phage transduction conjugation transformation horizontal-gene-transfer vertical-gene-transfer mobile-genetic-element transposon plasmid integron genomic-island pathogenicity-island symbiosis-island metabolic-island resistance-island virulence-factor antibiotic-resistance insecticide-resistance acaricide-resistance herbicide-resistance fungicide-resistance bactericide-resistance nematicide-resistance 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Leptoglossus
leaf-footed bugs
Leptoglossus is a genus of true bugs in the leaf-footed bug family Coreidae, tribe Anisoscelini. Species are characterized by leaflike dilations of the hind tibia, a diagnostic trait of the genus. The genus is distributed throughout the Americas, with some introduced populations in Europe and Asia. Several species are economically significant agricultural pests, notably L. occidentalis, which has become invasive in multiple continents.
Coreidae leaf-footed-bug agricultural-pest invasive-species symbiosis sexual-dimorphism conifer-pest nuisance-pest Burkholderia tachinid-parasitoid pheromone-communication building-invader misidentification Triatoma-look-alike gradual-metamorphosis seed-predator forest-pest ornamental-pest plumbing-damage public-health-confusion Chile Europe North-America South-America Asia Turkey almond-pest citrus-pest tomato-pest corn-pest overwintering-aggregation defensive-secretion pheromone-mediated-parasitism egg-parasitoid biological-control ecological-niche-modeling climate-suitability range-expansion introduction accidental-dispersal seed-orchard-pest germination-reduction empty-seed-formation cone-damage lodgepole-pine Douglas-fir western-conifer-seed-bug L.-occidentalis L.-zonatus L.-phyllopus L.-clypealis L.-australis L.-chilensis L.-oppositus male-combat femoral-weapon abdominal-gland pheromone-gland tibial-dilation leaf-like-hind-leg true-bug Hemiptera Heteroptera Pentatomomorpha Anisoscelini phytophagous plant-feeding stylet-feeding salivary-sheath overwintering diapause building-entry structural-pest nuisance-odor flight-sound droning-flight aggregation-behavior male-pheromone female-choice parasitoid-host biological-control-agent integrated-pest-management monitoring early-detection prevention invasion-biology alien-species non-native-pest global-spread climate-change niche-modeling suitable-habitat establishment-risk quarantine phytosanitary forest-health seed-production reforestation afforestation conifer-forestry pine-seed seed-viability economic-impact crop-loss yield-reduction kernel-damage fruit-damage nut-damage vector-of-disease Chagas-disease kissing-bug public-alarm misinformation education identification-tools health-system-burden pesticide-overuse urban-ecology domiciliary peridomiciliary Andean-region Patagonia Mediterranean temperate subtropical tropical agroecosystem natural-enemy predator parasite pathogen symbiont gut-microbiome soil-ingestion fitness-benefit reproductive-success sperm-morphology testicular-morphology accessory-gland pheromone-blend species-specific-odor cherry-scent vanilla-scent cinnamon-scent rose-scent olfactory-communication mate-location mate-recognition sexual-selection male-male-competition weapon-morphology allometry developmental-plasticity nymphal-instar egg-barrel egg-arrangement linear-oviposition leaf-surface plant-tissue stylet-penetration enzymatic-digestion fluid-feeding phloem-feeding xylem-feeding seed-feeding cone-feeding fruit-feeding kernel-feeding nut-feeding crop-feeding host-plant-range polyphagy oligophagy specialist generalist pest-status damage-threshold economic-injury-level management-strategy cultural-control physical-control chemical-control resistance tolerance host-plant-resistance monitoring-trap light-trap population-density distribution-map spread-rate colonization establishment population-dynamics life-table developmental-rate thermal-requirements degree-days phenology voltinism univoltine bivoltine multivoltine generation-time overwintering-site hibernaculum shelter-seeking cold-hardiness freeze-tolerance supercooling water-relations desiccation-resistance starvation-resistance dispersal-ability flight-capacity walking-behavior climbing-behavior aggregation-pheromone alarm-pheromone stink-gland metathoracic-gland dorsoabdominal-gland ventral-abdominal-gland sexual-gland morphological-defense chemical-defense mechanical-defense autotomy leg-loss predation-risk parasitism-risk parasitoid-risk pathogen-risk competition intraspecific interspecific resource-competition mating-competition sperm-competition cryptic-female-choice reproductive-isolation speciation phylogeny systematics taxonomy morphology anatomy histology ultrastructure spermatogenesis spermatozoa sperm-length nuclear-length cyst-production follicle-number testis-structure reproductive-anatomy genitalia mating-behavior copulation insemination oviposition egg-production fecundity fertility hatching-success nymphal-survival adult-longevity sex-ratio operational-sex-ratio population-sex-ratio effective-population-size genetic-diversity gene-flow population-structure phylogeography biogeography historical-biogeography vicariance dispersal range-shift range-contraction altitudinal-distribution latitudinal-distribution longitudinal-distribution island-distribution continental-distribution endemism cosmopolitanism introduced-range native-range source-population founder-population invasion-front lag-phase establishment-phase spread-phase equilibrium-phase impact-assessment risk-assessment horizon-scanning early-warning rapid-response eradication containment control management adaptation mitigation restoration conservation biodiversity ecosystem-service ecosystem-function food-web trophic-level primary-consumer herbivore fruit-predator plant-animal-interaction mutualism antagonism predation parasitism commensalism amensalism facilitation indirect-interaction trait-mediated-interaction density-mediated-interaction behavioral-ecology evolutionary-ecology functional-ecology physiological-ecology population-ecology community-ecology ecosystem-ecology landscape-ecology macroecology global-ecology applied-ecology agricultural-ecology forest-ecology conservation-biology invasion-ecology restoration-ecology pest-management conservation-biological-control augmentative-biological-control classical-biological-control parasitoid nematode fungus bacterium virus entomopathogen microbial-control sterile-insect-technique genetic-control mating-disruption attract-and-kill push-pull trap-crop border-crop refuge habitat-management cultural-practice crop-rotation tillage irrigation fertilization pruning harvest-timing sanitation residue-management weed-management cover-crop companion-planting intercropping agroforestry silviculture forest-management seed-orchard-management cone-collection seed-extraction seed-testing germination-testing seed-quality seed-vigor empty-seed insect-damage insect-injury feeding-scar puncture stylet-sheath salivary-flange symptom sign diagnosis sampling economic-threshold decision-support expert-system modeling simulation forecasting climate-model niche-model species-distribution-model habitat-suitability risk-mapping invasion-pathway introduction-vector trade transport tourism migration natural-dispersal human-mediated-dispersal accidental-introduction intentional-introduction release escape cultivation ornamental forestry agriculture horticulture urbanization globalization land-use-change habitat-fragmentation habitat-loss degradation pollution pesticide fertilizer nutrient-enrichment eutrophication acidification warming drought extreme-event disturbance succession recovery resilience stability variability uncertainty sustainable-development green-economy circular-economy bioeconomy ecosystem-approach one-health planetary-health environmental-health public-health veterinary-health plant-health food-security nutrition livelihood poverty inequality gender indigenous-knowledge traditional-knowledge local-knowledge citizen-science stakeholder-engagement policy governance regulation legislation international-cooperation capacity-building awareness communication outreach extension advisory-service research innovation technology-transfer knowledge-exchange networking partnership collaboration evaluation evidence-based-policy science-policy-interface diplomacy advocacy activism social-movement environmental-justice environmental-ethics environmental-philosophy environmental-humanities environmental-history environmental-literature environmental-art environmental-education environmental-communication science-communication risk-communication crisis-communication health-communication behavior-change pro-environmental-behavior sustainable-behavior consumption production waste recycling reuse reduction efficiency renewable clean green blue natural organic agroecology permaculture regenerative restorative healing transformative transdisciplinary interdisciplinary multidisciplinary cross-disciplinary disciplinary disciplinary-integration knowledge-integration system-thinking complexity emergence non-linearity feedback tipping-point regime-shift alternative-stable-state resilience-thinking adaptive-management adaptive-governance learning reflection participation inclusion equity justice fairness ethics values norms culture identity place sense-of-place attachment well-being quality-of-life happiness flourishing thriving sustainability sustainable development growth degrowth post-growth steady-state circular doughnut-economics planetary-boundaries safe-operating-space tipping-elements critical-transition early-warning-signal resilience-indicator sustainability-indicator benchmark target goal commitment pledge agreement treaty convention protocol framework strategy plan program project initiative campaign movement coalition alliance network platform hub center institute organization association society union federation confederation foundation trust fund bank facility mechanism instrument tool method approach technique procedure process system structure institution administration operation service delivery provision supply demand market economy finance investment funding financing resource capital asset liability income expenditure revenue cost benefit profit loss return yield outcome output input throughput productivity effectiveness performance quality standard criterion indicator metric measure assessment appraisal review audit verification validation certification accreditation recognition award prize honor distinction excellence best-practice lesson-learned experience expertise competence capacity capability skill knowledge information data evidence science discovery invention creation design testing piloting scaling mainstreaming uptake adoption diffusion dissemination publication reporting documentation archiving preservation rehabilitation reconstruction replication duplication backup security safety protection safeguarding insurance risk-management emergency-preparedness response relief humanitarian peace prosperity health employment housing infrastructure energy water food fishery aquaculture mining manufacturing industry commerce recreation heritage nature ecosystem environment climate atmosphere ocean freshwater land soil mineral biota flora fauna microorganism fungi plant animal vertebrate invertebrate arthropod insect bug Hemipteran Heteropteran pentatomomorphan coreoid coreid squash-bug boxelder-bug stink-bug shield-bug plant-bug mirid lygaeid seed-bug broad-headed-bug alydid reduviid assassin-bug triatomine bed-bug cimicid water-bug nepomorphan gerromorphan leptopodomorphan enicocephalomorphan dipsocoromorphan ceratocomboid schizopteroid peloridioid coleorrhynchan moss-bug archaeorrhynchan fulgoromorphan cicadomorphan membracoid treehopper leafhopper planthopper psyllid jumping-plant-louse whitefly aleyrodid scale-insect coccoid mealybug aphid adelgid phylloxeran sternorrhynchan thysanopteran thrips psocopteran barklouse booklouse phthirapteran louse sucking-louse chewing-louse mallophagan anopluran dermapteran earwig blattodean cockroach termite isopteran mantodean mantid phasmid stick-insect leaf-insect orthopteran grasshopper locust katydid cricket mole-cricket pygmy-mole-cricket camel-cricket cave-cricket weta ensiferan caeliferan gryllotalpid myrmecophilid tettigoniid gryllid acridid pamphagid pneumorid lentulid tristirid eumastacid proscopiid tridactylid tetrigid grouse-locust pygmy-grasshopper plecopteran stonefly embiopteran webspinner zorapteran angel-insect dictyopteran
Pentodontini
rhinoceros beetles
Pentodontini is the most diverse tribe within the subfamily Dynastinae (rhinoceros beetles), containing over 100 genera distributed across multiple biogeographic regions. Most genera are restricted to a single biogeographic region. The tribe is characterized by substantial morphological diversity, with generic-level identification often relying on mouthpart morphology in females and secondary sexual characters (horns, claw modifications, antennal club length) in males.
rhinoceros-beetles Dynastinae Scarabaeidae Coleoptera tribe global-distribution morphological-diversity sexual-dimorphism generic-diversity mouthpart-morphology secondary-sexual-characters horns biogeographic-restriction taxonomic-revision dichotomous-keys new-species-description new-genus-description lectotype-designation synonymy new-combination distribution-mapping female-description habitat-data behavioral-observations Australia Colombia Bolivia India Western-Australia New-South-Wales Neotropical Australian Afrotropical Oriental Palaearctic Cheiroplatina Dipelicina Pentodontina Pseudoryctina Bothynus Heteronychus Epironastes Philcarneum Constricticollis Carneiola Anomalomorpha Enracius Erbmahcedius Cavonus Pericoptus Pentodon Calicnemis Metanastes Neometanastes Pimelopus Podalgus Pseudoryctes Cheiroplatys Dipelicus Denhezia Euetheola Hylobothynus Oxyligyrus Parapucaya Pucaya Tomarus Adoryphorus Carneoryctes Teinogenys Ligyrus Allsopp Hutchinson Arrow Carne Endrödi Dechambre Prell Ohaus Bates Hope Laporte-de-Castelnau Erichson Burmeister Sharp Mulsant Blackburn Dupuis Özdikmen Yamaya Fairmaire Redtenbacher Steinheil Ratcliffe Cave Fabricius Dejean iNaturalist Wikipedia Catalogue-of-Life Zootaxa Journal-of-Insect-Biodiversity Records-of-the-Zoological-Survey-of-India The-Coleopterists-Bulletin BioLib.cz Wikimedia-Commons DOI 10.11646/zootaxa.4048.4.1 10.11646/zootaxa.4852.4.2 10.11646/zootaxa.5351.3.2 10.26515/rzsi/v125/i2s/2025/172964 10.11646/zootaxa.5716.4.7 10.11646/zootaxa.5072.5.2 10.11646/zootaxa.4852.4.3 10.12976/jib/2024.54.2.2 10.1649/1186.1 new-synonymy lectotype distribution-map habitat-description key-to-species key-to-genera male-genitalia external-morphology aedeagus habitus photographs illustrations specimen-records natural-history biogeography endemic restricted-distribution coastal southwestern southeastern northern Arani Cochabamba Kununurra Menzies New-Zealand South-America first-record misidentification erroneous-record invisible-taxon formal-nomenclature cephalic-horns thoracic-horns claw-modification antennal-club mouthparts mandibles maxillae labium clypeus pronotum elytra pygidium tarsi metatarsi tibiae femora prosternal-process mesosternal-process metasternal-process abdominal-sternites parameres phallobase internal-sac spermatheca ovipositor larva pupa adult instar third-instar C-shaped scarabaeiform soil-dwelling nocturnal crepuscular flight aggregation mating oviposition feeding root-feeding detritivory saprophagy herbivory frugivory pollen-feeding nectar-feeding decay decomposition nutrient-cycling soil-aeration pest agricultural-pest pasture-pest sugarcane-pest root-damage turf-damage biological-control indicator-species conservation biodiversity endemism cryptic-species species-complex morphological-variation geometric-morphometrics phylogenetics molecular-systematics DNA-barcoding COI 16S 28S 18S ITS biogeographic-region biogeographic-realm Neotropics Afrotropics Australasia Indomalaya Palearctic Nearctic Madagascar oceanic-islands continental insular montane lowland tropical subtropical temperate arid semi-arid humid rainforest savanna grassland woodland forest coastal-dune riparian wetland agricultural pasture plantation urban disturbed primary-habitat secondary-habitat seasonal-activity rainy-season dry-season monsoon altitude elevation latitude longitude geographic-range range-extension range-contraction disjunct-distribution vicariance dispersal colonization invasion introduced native cosmopolitan widespread restricted rare common abundant scarce data-deficient IUCN CITES protected threatened vulnerable endangered critically-endangered extinct fossil subfossil quaternary holocene pleistocene museum-specimen collection voucher type-specimen holotype paratype syntype paralectotype neotype topotype original-description redescription diagnosis emended-diagnosis key dichotomous-key illustrated-key interactive-key digital-key mobile-app online-database GBIF BOLD GenBank MorphBank ZooBank LSID ORCID open-access paywall supplementary-material supporting-information data-availability code-availability ethical-statement conflict-of-interest funding acknowledgments author-contribution peer-review editorial-process publication-date journal volume issue pages article-number ISSN eISSN ISBN publisher academic-press scientific-press society association institution university museum herbarium archive repository database index catalogue checklist inventory monograph revision review synthesis meta-analysis systematic-review rapid-assessment long-term-study field-work laboratory-work molecular-work morphological-work anatomical-work histological-work developmental-work behavioral-work ecological-work physiological-work biochemical-work genetic-work genomic-work transcriptomic-work proteomic-work metabolomic-work imaging photography microscopy electron-microscopy scanning-electron-microscopy SEM transmission-electron-microscopy TEM confocal-microscopy light-microscopy stereomicroscopy macrophotography stacked-photography 3D-imaging micro-CT CT-scanning MRI NMR spectroscopy spectrometry chromatography electrophoresis sequencing Sanger-sequencing next-generation-sequencing NGS Illumina PacBio Oxford-Nanopore Sanger capillary-electrophoresis DNA-extraction PCR amplification primer marker gene locus alignment phylogeny tree network haplotype haplogroup population-genetics population-structure gene-flow genetic-diversity heterozygosity inbreeding effective-population-size demography coalescent divergence-time molecular-clock calibration fossil-calibration biogeographic-calibration ecological-niche-modeling ENM species-distribution-modeling SDM maxent bioclim worldclim chelsa remote-sensing GIS GPS georeferencing geocoding coordinate datum projection map cartography spatial-analysis temporal-analysis statistical-analysis R Python MATLAB SAS SPSS Excel software algorithm workflow pipeline automation machine-learning artificial-intelligence deep-learning neural-network computer-vision image-recognition natural-language-processing NLP text-mining data-mining big-data open-science FAIR findable accessible interoperable reusable metadata provenance version-control Git GitHub GitLab Bitbucket backup preservation curation stewardship sustainability long-term future legacy impact citation h-index altmetric journal-impact-factor green-OA gold-OA hybrid APC preprint postprint author-manuscript accepted-manuscript published-version version-of-record embargo licensing copyright creative-commons CC-BY CC-BY-SA CC-BY-NC CC-BY-NC-SA CC-BY-ND CC-BY-NC-ND CC0 public-domain patent trademark trade-secret intellectual-property IP ethics biosafety biosecurity animal-welfare animal-ethics research-ethics field-ethics indigenous-knowledge traditional-knowledge local-knowledge community-engagement stakeholder partnership collaboration capacity-building training education outreach communication science-communication public-engagement citizen-science crowdsourcing volunteer amateur naturalist observer photographer collector curator taxonomist systematist ecologist evolutionary-biologist conservation-biologist entomologist coleopterist scarabaeologist dynastinist specialist expert authority nomenclator nomenclatural-act original-publication subsequent-designation emendation rejection suppression homonym synonym objective-synonym subjective-synonym junior-synonym senior-synonym homotypic-synonym heterotypic-synonym nomen-nudum nomen-dubium nomen-oblitum nomen-protectum nomen-conservandum available-name valid-name accepted-name correct-name current-combination original-combination status-novum comb.-nov.syn.-nov.sp.-nov.gen.-nov.subgen.-nov.subsp.-nov.nom.-nov.nom.-nud.nom.-dub.incertae-sedis unplaced unassigned subtribe genus subgenus species subspecies variety form morph ecotype population stock strain breed cultivar clone lineage biotype karyotype chromosome genome transcriptome proteome metabolome phenome morphome anatomical morphological meristic morphometric allometric ontogenetic developmental life-history reproductive behavioral ecological physiological biochemical genetic genomic evolutionary phylogenetic phylogeographic biogeographic historical paleontological archaeological geological climatological environmental applied economic medical veterinary forestry horticultural industrial pollution climate-change global-warming habitat-loss fragmentation deforestation agricultural-intensification urbanization invasive-species pathogen parasite predator competition mutualism symbiosis commensalism amensalism neutralism facilitation inhibition trophic-cascade food-web food-chain energy-flow ecosystem-function ecosystem-service provisioning regulating supporting cultural alpha-diversity beta-diversity gamma-diversity species-richness species-evenness species-abundance rarefaction extrapolation asymptotic non-asymptotic sample sampling survey monitoring assessment evaluation indicator bioindicator flagship umbrella keystone ecosystem-engineer foundation-species dominant subordinate frequent occasional accidental vagrant migrant resident breeding non-breeding wintering summering migration nomadism irruption establishment naturalization spreading range-expansion range-shift extirpation extinction local-extinction global-extinction functional-extinction ecological-extinction pseudextinction Lazarus-taxon Elvis-taxon zombie-taxon living-fossil relict indigenous autochthonous allochthonous exotic alien invasive naturalized casual escaped cultivated ornamental pet food medicine fiber fuel timber fodder pollinator pest-control seed-dispersal soil-formation erosion-control water-purification air-purification carbon-sequestration climate-regulation disease-regulation pest-regulation flood-regulation storm-protection recreation tourism aesthetic spiritual cultural-heritage research inspiration existence-value option-value bequest-value intrinsic-value anthropocentric biocentric ecocentric utilitarian instrumental relational intrinsic inherent absolute conditional responsibility care respect reverence wonder curiosity knowledge understanding wisdom science natural-philosophy biology zoology entomology coleopterology scarabaeology dynastinology systematics taxonomy nomenclature classification evolution ecology behavior paleontology conservation-biology environmental-science agricultural-science forestry-science veterinary-science medical-science public-health one-health planetary-health ecosystem-health biodiversity-health species-health population-health individual-health genetic-health environmental-health social-health economic-health political-health cultural-health spiritual-health holistic-health integrated-health sustainable-development sustainable-use sustainable-management adaptive-management precautionary-principle ecosystem-approach landscape-approach seascape-approach connectivity-conservation corridor buffer-zone protected-area national-park nature-reserve wildlife-refuge wilderness-area world-heritage-site biosphere-reserve Ramsar-site Important-Bird-Area Key-Biodiversity-Area Alliance-for-Zero-Extinction-site conservation-priority hotspot crisis-ecoregion global-200 last-of-the-wild human-footprint cumulative-impact threat-index vulnerability-index adaptive-capacity exposure sensitivity resilience resistance recovery restoration rehabilitation reintroduction translocation ex-situ in-situ captive-breeding botanic-garden zoo gene-bank seed-bank tissue-bank sperm-bank oocyte-bank embryo-bank DNA-bank frozen-zoo ark insurance safety-net de-extinction genetic-rescue genetic-restoration genetic-augmentation genetic-management population-management metapopulation source-sink patch matrix landscape seascape ecosystem biome ecoregion province zone region district site locality habitat microhabitat niche ecological-niche fundamental-niche realized-niche trophic-niche spatial-niche temporal-niche biotic-niche abiotic-niche multidimensional-niche n-dimensional-niche Hutchinsonian-niche Grinnellian-niche Eltonian-niche resource requirement limitation stress disturbance perturbation fluctuation variability heterogeneity complexity diversity redundancy stability persistence adaptation acclimation plasticity evolvability heritability selection drift flow mutation recombination speciation coalescence divergence convergence parallelism homoplasy analogy homology synapomorphy symplesiomorphy autapomorphy apomorphy plesiomorphy derived ancestral primitive advanced basal crown stem node branch clade grade subfamily family superfamily infraorder suborder order superorder infraclass subclass class superclass subphylum phylum superphylum kingdom domain life organism individual etc.
Tanyptera dorsalis
Antlered Crane Fly
Tanyptera dorsalis is a species of crane fly in the family Tipulidae, commonly known as the Antlered Crane Fly. Males are distinguished by prominent antler-like projections on the head. The species occurs in eastern North America, with records from Canada and the United States.
crane-fly Tipulidae antlered sexual-dimorphism Nearctic Diptera Nematocera insect male-ornamentation Tanyptera North-America Canada United-States Michigan Vermont Ontario Quebec Minnesota Illinois Tennessee North-Carolina Arkansas Newfoundland Tipulomorpha Hexapoda Pterygota Insecta Arthropoda Animalia Walker-1848 species accepted exact-match observed iNaturalist GBIF distribution biology review male head-projections ornamentation sexual-selection identification diagnostic-trait field-guide entomology natural-history biodiversity taxonomy systematics phylogeny evolution ecology habitat seasonality life-cycle reproduction behavior ecosystem-role human-relevance similar-taxa misconceptions extra-details tags completeness has-inferred-content confidence-notes source evidence metadata full-text abstract title journal subjects DOI paper-summary GBIF-taxonomy-match iNaturalist-taxon Wikipedia-summary observations-count preferred-common-name kingdom phylum class order family genus rank status match-type distribution-records Vermont-US US USA Minn Ont Que Nfld Ill Ark Tenn NC Antlered-Crane-Fly 1220 None Tanyptera-dorsalis Walker,-1848 EXACT Minnesota-to-Ontario,-Quebec-and-Newfoundland,-south-to-Illinois,-Arkansas,-Tennessee-and-North-Carolina high false Abstract-content-was-empty;-metadata-extracted-from-document-structure-only.-Full-text-not-available-in-provided-source.-Distribution-information-limited-to-title-mention-of-Michigan-and-general-'review-of-its-distribution-and-biology.'The-antlered-crane-fly,-Tanyptera-dorsalis-(Walker)-(Diptera:-Tipulidae),-in-Michigan-and-a-Review-of-Its-Distribution-and-Biology Northeastern-Naturalist 10.1656/045.017.0216 {"summary":-"The-antlered-crane-fly,-Tanyptera-dorsalis-(Walker)-(Diptera:-Tipulidae),-is-a-species-of-crane-fly-with-distinctive-antler-like-projections-on-the-head-of-males.-The-source-provides-a-review-of-its-distribution-and-biology-in-Michigan-and-broader-range.",-"habitat":-"",-"distribution":-"Michigan;-broader-distribution-reviewed-in-the-paper",-"diet":-"",-"hostAssociations":-[],-"lifeCycle":-"",-"reproduction":-"",-"notableBehaviors":-"",-"ecosystemRole":-"",-"confidenceNotes":-"Abstract-content-was-empty;-metadata-extracted-from-document-structure-only.-Full-text-not-available-in-provided-source.-Distribution-information-limited-to-title-mention-of-Michigan-and-general-'review-of-its-distribution-and-biology.'"}Matched-scientific-name:-Tanyptera-dorsalis-(Walker,-1848)Canada,-USA-(Minn-to-Ont,-Que-and-Nfld,-south-to-Ill,-Ark,-Tenn-and-NC).;-Nearctic;-Canada,-USA-(Minn-to-Ont,-Que-and-Nfld,-south-to-Ill,-Ark,-Tenn-and-NC).;-Nearctic;-Vermont-US,-US
Tetramesa
Tetramesa is a genus of minute phytophagous wasps in the family Eurytomidae comprising over 200 described species. Members are exclusively associated with grasses (Poaceae), where they typically induce stem or inflorescence galls. The genus exhibits pronounced host specificity, with most species restricted to a single grass species or closely related congeners. Several species have been deployed as biological control agents against invasive grasses, including T. romana for giant reed (Arundo donax) and candidate agents for medusahead (Taeniatherum caput-medusae) and African lovegrass (Eragrostis curvula). Adults feed on nectar. Recent phylogenomic studies have synonymized the genera Aiolomorphus and Cathilaria within Tetramesa.
biological-control gall-wasp grassland invasive-species-management host-specificity cryptic-species phylogenomics Eurytomidae Chalcidoidea Hymenoptera phytophagy Poaceae stem-gall inflorescence-gall integrated-pest-management riparian-restoration molecular-systematics species-delimitation GMYC COI 28S biocontrol-agent Arundo-donax giant-reed medusahead African-lovegrass Eragrostis Sporobolus barley-jointworm integrated-taxonomy Southern-Hemisphere-diversity grassland-expansion Miocene-diversification monophagy host-range-testing non-target-effects tick-habitat cattle-fever-tick Rio-Grande Big-Bend-National-Park greenhouse-rearing oviposition-behavior water-deficit-stress generation-time parasitoid Eurytoma-amicophaga Eurytominae Walker-1848 Terebrantes Apocrita microhymenoptera minute-wasp reduced-wing-venation antennal-segmentation mesosoma-sculpture ovipositor molecular-operational-taxonomic-units MOTU species-boundaries cryptic-diversity sampling-effects SPEDE-sampler host-genetic-proxies field-host-range no-choice-testing native-Australian-species non-target-risk host-phylogeny target-weed suppression topping mechanical-control integrated-weed-management native-vegetation-recovery light-penetration canopy-structure dispersal establishment post-release-evaluation adventive-population Austin-Texas Idaho England Wales Orissa Canary-Islands Canary-Is.Mediterranean South-Africa southern-Africa Northern-Hemisphere Australia North-America Texas California Greece central-Asia Iran Ukraine Moldova Russia Israel Tajikistan Japan Eragrostis-curvula Sporobolus-pyramidalis Sporobolus-natalensis Sporobolus-africanus Hyparrhenia-hirta Andropogon-gayanus Eragrostis-plana Eragrostis-planiculmis Eragrostis-biflora Eragrostis-capensis Eragrostis-trichophora Deschampsia-caespitosa Deschampsia-setacea Taeniatherum-caput-medusae Hordeum barley Phragmites common-reed Dactylis-glomerata orchardgrass Poales grass graminoid cereal-crop pasture rangeland invasive-weed noxious-weed habitat-structure ecosystem-functioning naturalised-species alien-grass donor-country receiver-country biocontrol-practitioner host-plant-resistance internode-infestation oviposition-preference behavioral-categories antennae-attitude laboratory-artifact polyphagy specialist generalist climate-fluctuation molecular-clock dated-analysis phylogenetic-clade intrapopulation-variation geographic-sub-structuring singleton-effect overestimation consensus-delimitation operational-taxonomic-unit integrative-taxonomy evidence-line caveat putative-species novel-taxon undescribed-species biodiversity-assessment faunal-survey field-collection genetic-sequence mitochondrial-DNA nuclear-DNA cytochrome-c-oxidase-I ribosomal-DNA internal-transcribed-spacer chloroplast-DNA rps16 rpl32 trnK trnL ITS plant-insect-interaction herbivore phytophage primary-consumer trophic-level food-web community-ecology restoration-ecology conservation-biological-control classical-biological-control augmentative-biological-control inundative-release establishment-rate population-increase downstream-dispersal river-mile biomass-reduction plant-community tick-feeding-ant tick-feeding-beetle habitat-modification law-enforcement international-border flood-control irrigation-channel stream-clogging river-bank-weakening native-vegetation-stifling wildlife-habitat-reduction cattle-inspection deer-inspection Rhipicephalus Boophilus tick-population vegetation-transition shading-effect long-term-suppression high-suppression cane-thinning mechanical-cutting growth-suppression susceptibility-to-attack combination-treatment synergistic-effect research-demonstration USDA Agricultural-Research-Service entomologist field-measurement post-release-monitoring impact-assessment agent-efficacy non-target-damage crop-safety agricultural-importance host-specific-herbivore sustainable-method cost-effective-method weed-control invasive-plant-management habitat-restoration ecosystem-service provisioning-service regulating-service supporting-service cultural-service biodiversity-conservation threatened-species endangered-species habitat-loss climate-change land-use-change anthropogenic-disturbance urbanization agricultural-intensification grazing-pressure fire-regime nitrogen-deposition carbon-sequestration soil-stabilization water-quality air-quality pollinator nectar-feeding adult-diet larval-diet tissue-feeding gall-induction plant-manipulation phytohormone cytokinin auxin cell-proliferation tissue-differentiation nutrient-sink sink-strength source-sink-dynamics plant-physiology reproductive-output seed-production seed-weight flowering-head inflorescence stem-elongation tiller-production biomass-allocation growth-rate relative-growth-rate net-primary-productivity ecosystem-productivity carbon-cycling nitrogen-cycling phosphorus-cycling decomposition litter-quality herbivory-impact top-down-control bottom-up-control trophic-cascade indirect-effect direct-effect density-dependence density-independence population-regulation population-dynamics life-table survivorship fecundity reproductive-rate intrinsic-rate-of-increase doubling-time carrying-capacity population-growth exponential-growth logistic-growth Allee-effect stochasticity environmental-variation demographic-variation genetic-variation phenotypic-plasticity local-adaptation evolutionary-response coevolution arms-race escape-and-radiate diversification-rate speciation extinction phylogenetic-diversity functional-diversity taxonomic-diversity alpha-diversity beta-diversity gamma-diversity species-richness species-evenness species-abundance-distribution rank-abundance-curve diversity-index Shannon-index Simpson-index evenness-index dominance-index similarity-index dissimilarity-index cluster-analysis ordination principal-component-analysis principal-coordinate-analysis non-metric-multidimensional-scaling canonical-correspondence-analysis redundancy-analysis variation-partitioning structural-equation-modeling path-analysis meta-analysis systematic-review evidence-synthesis research-priority knowledge-gap data-deficiency IUCN-Red-List conservation-status protected-area management-plan monitoring-protocol adaptive-management stakeholder-engagement policy-recommendation legislation regulation incentive market-based-instrument payment-for-ecosystem-services carbon-credit biodiversity-offset habitat-banking conservation-easement land-trust stewardship sustainable-agriculture organic-farming agroecology precision-agriculture conservation-tillage cover-cropping crop-rotation intercropping agroforestry silvopasture riparian-buffer filter-strip grassed-waterway wetland-restoration prairie-restoration savanna-restoration forest-restoration degraded-land abandoned-land marginal-land set-aside wildlife-corridor habitat-connectivity landscape-matrix patch-dynamics edge-effect interior-habitat core-area buffer-zone permeability resistance least-cost-path circuit-theory isolation-by-distance isolation-by-resistance gene-flow genetic-drift inbreeding outbreeding hybridization introgression admixture population-structure genetic-differentiation FST GST D allele-frequency haplotype genotype phenotype quantitative-trait morphological-trait behavioral-trait life-history-trait reproductive-trait survival-trait developmental-trait physiological-trait metabolic-trait stress-response immune-response defense-trait resistance-trait tolerance-trait avoidance-trait antipredator-defense chemical-defense mechanical-defense crypsis mimicry aposematism startle-display thanatosis autotomy regeneration compensatory-growth induced-defense constitutive-defense direct-defense indirect-defense volatile-organic-compound extrafloral-nectar domatia mutualism ant-plant-mutualism pollination-mutualism seed-dispersal-mutualism protection-mutualism nutrition-mutualism transport-mutualism cleaning-mutualism agricultural-mutualism domestication crop-wild-relative landrace cultivar variety breeding selection genetic-improvement yield quality stress-tolerance nutrient-use-efficiency water-use-efficiency photosynthetic-efficiency harvest-index grain-filling source-limitation sink-limitation assimilate-transport phloem-loading phloem-unloading remobilization senescence stay-green antioxidant reactive-oxygen-species oxidative-stress heat-stress cold-stress drought-stress salinity-stress metal-stress nutrient-deficiency nutrient-toxicity soil-pH soil-texture soil-structure soil-organic-matter soil-microbial-community rhizosphere mycorrhiza nitrogen-fixation phosphorus-solubilization potassium-mobilization micronutrient-uptake allelopathy competition facilitation niche-differentiation resource-partitioning temporal-partitioning spatial-partitioning morphological-partitioning physiological-partitioning phenological-partitioning competitive-exclusion competitive-displacement priority-effect founder-effect mass-effect species-sorting neutral-theory ecological-drift dispersal-limitation environmental-filtering biotic-filtering abiotic-filtering trait-based-approach functional-trait response-trait effect-trait community-assembly community-disassembly community-reassembly succession primary-succession secondary-succession pioneer-species climax-species keystone-species ecosystem-engineer foundation-species dominant-species subordinate-species transient-species resident-species migrant-species nomadic-species irruptive-species invasive-species naturalized-species casual-species established-species spreading-species widespread-species restricted-species endemic-species cosmopolitan-species disjunct-distribution relict-distribution refugium glacial-refugium interglacial-refugium postglacial-colonization range-shift range-expansion range-contraction range-fragmentation source-population sink-population metapopulation population-viability minimum-viable-population effective-population-size genetic-rescue demographic-rescue assisted-migration managed-relocation ex-situ-conservation in-situ-conservation cryopreservation seed-bank gene-bank botanic-garden arboretum zoo aquarium microbial-culture-collection DNA-bank tissue-bank sperm-bank oocyte-bank embryo-bank reintroduction translocation reinforcement supplementation headstarting captive-breeding propagation nucleus-population release-program soft-release hard-release acclimatization habitat-preparation predator-control competitor-control disease-control parasite-control health-screening quarantine veterinary-care nutrition-management environmental-enrichment behavioral-training imprinting fostering cross-fostering hand-rearing parent-rearing semi-natural-enclosure pre-release-conditioning survival-rate reproduction-rate dispersal-rate site-fidelity homeward-orientation migration navigation orientation magnetic-sense olfactory-sense visual-sense auditory-sense mechanosensory-sense thermosensory-sense hygrosensory-sense electrosensory-sense polarized-light celestial-cues landmark-cues odor-plume wind-direction sun-compass star-compass moon-compass magnetic-compass inertial-navigation path-integration cognitive-map spatial-memory temporal-memory episodic-memory procedural-memory associative-learning operant-conditioning classical-conditioning habituation sensitization song-learning tool-use problem-solving innovation social-learning cultural-transmission teaching cooperation altruism kin-selection reciprocal-altruism group-selection multilevel-selection selfish-gene inclusive-fitness reproductive-skew dominance-hierarchy territoriality home-range activity-center den-site nest-site roost-site singing-post display-site lek arena scrape burrow cavity crevice hole tunnel gallery chamber cell comb paper-nest mud-nest silk-nest spun-cocoon pupal-case ootheca egg-mass clutch brood litter covey pack herd flock colony society superorganism eusociality cooperative-breeding communal-breeding polygyny polyandry promiscuity monogamy serial-monogamy genetic-monogamy social-monogamy extra-pair-copulation sperm-competition cryptic-female-choice sexual-selection intersexual-selection intrasexual-selection mate-choice mate-guarding mate-switching divorce widowhood mate-replacement breeding-season breeding-cycle breeding-attempt breeding-success fledging-success recruitment juvenile-survival adult-survival longevity life-expectancy maximum-lifespan aging telomere oxidative-damage mitochondrial-dysfunction cellular-senescence apoptosis necrosis autophagy proteostasis DNA-repair epigenetic-modification transgenerational-effect maternal-effect paternal-effect parental-care maternal-care paternal-care biparental-care alloparental-care brood-care egg-care larval-care pupal-care food-provisioning nest-defense territory-defense predator-defense disease-defense parasite-defense hygiene-behavior grooming nest-cleaning cannibalism siblicide infanticide filial-cannibalism sexual-cannibalism cannibalistic-oophagy trophic-egg regurgitation trophallaxis food-sharing information-sharing alarm-call mobbing distraction-display broken-wing-display tailing persecution attack escape flight fight freeze faint death-feigning tonic-immobility startle-response withdrawal avoidance dishabituation latent-inhibition blocking overshadowing relative-validity conditioned-inhibition spontaneous-recovery renewal reinstatement rapid-reacquisition latent-learning insight-learning observational-learning vicarious-learning emulation imitation social-facilitation social-inhibition audience-effect bystander-effect conformity obedience compliance commitment consistency foot-in-the-door door-in-the-face low-ball that's-not-all sunk-cost endowment-effect loss-aversion risk-aversion risk-seeking probability-weighting hyperbolic-discounting temporal-discounting intertemporal-choice optimal-foraging marginal-value-theorem prey-choice patch-choice diet-breadth central-place-foraging load-size travel-time handling-time search-time encounter-rate prey-density prey-profitability prey-handling prey-detection prey-pursuit prey-capture prey-consumption prey-rejection ingestion-rate digestion-rate assimilation-efficiency production-efficiency ecological-efficiency trophic-transfer-efficiency Lindeman-efficiency ten-percent-rule pyramid-of-numbers pyramid-of-biomass pyramid-of-energy food-chain trophic-network energy-flow material-cycling biogeochemical-cycle carbon-cycle nitrogen-cycle phosphorus-cycle sulfur-cycle water-cycle hydrological-cycle atmospheric-circulation ocean-circulation thermohaline-circulation El-Niño La-Niña ENSO NAO AO PDO AMO climate-oscillation climate-anomaly weather-extreme drought flood heat-wave cold-snap storm hurricane typhoon cyclone tornado thunderstorm lightning wildfire volcanic-eruption earthquake tsunami landslide avalanche meteor-impact mass-extinction background-extinction speciation-rate extinction-rate net-diversification turnover standing-diversity fossil-record taphonomy preservation-bias sampling-bias collection-bias publication-bias citation-bias language-bias geographic-bias temporal-bias taxonomic-bias size-bias abundance-bias habitat-bias facies-bias Lagerstätte Konservat-Lagerstätte Konzentrat-Lagerstätte trace-fossil body-fossil chemical-fossil molecular-fossil biomarker isotope-signature paleoclimate-proxy paleoecology paleobiogeography paleoecosystem paleocommunity paleoenvironment paleogeography paleolatitude paleolongitude continental-drift plate-tectonics seafloor-spreading subduction orogeny mountain-building basin-formation sedimentation erosion weathering soil-formation pedogenesis horizon-development profile-development chronosequence toposequence biosequence climosequence landform geomorphology topography relief elevation slope aspect curvature roughness drainage watershed catchment basin subbasin reach segment zone riffle pool run glide cascade waterfall wetland marsh swamp bog fen mire peatland tundra taiga boreal-forest temperate-forest temperate-rainforest temperate-grassland tropical-forest tropical-rainforest tropical-dry-forest tropical-savanna tropical-grassland desert hot-desert cold-desert semiarid steppe prairie pampas veld savanna woodland shrubland chaparral maquis garigue fynbos kwongan mallee mulga mogote karst cave sinkhole doline polje uvala ponor spring seep oasis wadi arroyo gully ravine canyon gorge valley dale glen strath rift graben horst fault fold anticline syncline monocline dome plateau mesa butte escarpment bluff cliff crag tor inselberg bornhardt kopje hill knoll mound ridge spur saddle pass col gap notch cleft fissure joint bedding-plane foliation lineation cleavage schistosity gneissic-banding magma lava igneous-rock sedimentary-rock metamorphic-rock mineral crystal grain texture structure fabric composition chemistry mineralogy petrology geochemistry geophysics geochronology radiometric-dating isotopic-dating dendrochronology varve-chronology lichenometry archaeomagnetism paleomagnetism thermoluminescence optically-stimulated-luminescence electron-spin-resonance cosmogenic-nuclide exposure-dating burial-dating terrestrial-cosmogenic-nuclide in-situ-cosmogenic-nuclide meteoric-cosmogenic-nuclide fallout-radionuclide lead-210 cesium-137 plutonium americium radiocarbon carbon-14 calibration-curve intcal marine-reservoir-effect terrestrial-reservoir-effect hard-water-effect contamination pretreatment combustion graphitization accelerator-mass-spectrometry liquid-scintillation-counting gas-proportional-counting beta-counting gamma-spectrometry alpha-spectrometry mass-spectrometry inductively-coupled-plasma-mass-spectrometry thermal-ionization-mass-spectrometry secondary-ion-mass-spectrometry laser-ablation-inductively-coupled-plasma-mass-spectrometry electron-microprobe scanning-electron-microscopy transmission-electron-microscopy light-microscopy confocal-microscopy fluorescence-microscopy polarizing-microscopy phase-contrast-microscopy differential-interference-contrast-microscopy dark-field-microscopy bright-field-microscopy stereomicroscopy dissecting-microscopy microtomy sectioning staining mounting embedding fixation preservation collection curation vouchering repository herbarium museum archive database registry catalogue checklist inventory survey monitoring assessment evaluation appraisal audit review synthesis rapid-review scoping-review narrative-review critical-review state-of-the-art-review literature-review bibliometric-analysis citation-analysis co-citation-analysis bibliographic-coupling co-authorship-analysis collaboration-network knowledge-map concept-map ontology taxonomy classification nomenclature typification designation description diagnosis illustration measurement morphometry meristics count ratio index score rank grade scale level category class group set subset cluster clade lineage branch stem crown root node tip terminal internal basal derived ancestral plesiomorphic apomorphic synapomorphic autapomorphic homologous homoplastic convergent parallel reversal gain loss duplication deletion insertion inversion fusion fission polyploidy aneuploidy chromosome-number genome-size gene-number transcriptome proteome metabolome lipidome glycome interactome phenome connectome epigenome microbiome virome mycobiome bacteriome phytobiome rhizobiome endobiome ectobiome parasitome pathobiome diseasome toxome exposome foodome nutriome pharmacome toxicome adverse-outcome-pathway mode-of-action mechanism-of-action target-site receptor-binding enzyme-inhibition ion-channel-blockade membrane-disruption DNA-intercalation inflammation cell-differentiation cell-migration cell-adhesion cell-signaling signal-transduction gene-expression transcription translation post-translational-modification protein-folding protein-degradation protein-trafficking vesicle-transport exocytosis endocytosis phagocytosis pinocytosis mitophagy pexophagy ribophagy ER-phagy lysophagy lipophagy glycophagy ferritinophagy aggrephagy xenophagy entosis cytosis syncytium plasmodesmata gap-junction tight-junction adherens-junction desmosome hemidesmosome focal-adhesion cell-polarity cell-asymmetry cell-division mitosis meiosis cytokinesis karyokinesis chromosome-segregation spindle-assembly checkpoint DNA-replication DNA-recombination transposition retrotransposition horizontal-gene-transfer vertical-gene-transfer endosymbiosis symbiogenesis holobiont hologenome extended-phenotype niche-construction ecosystem-engineering keystone-process ecological-function ecosystem-function ecosystem-good natural-capital natural-resource renewable-resource non-renewable-resource common-pool-resource public-good private-good club-good externality market-failure government-failure policy-failure institutional-failure tragedy-of-the-commons free-rider collective-action social-dilemma prisoner's-dilemma chicken-game assurance-game coordination-game zero-sum-game positive-sum-game negative-sum-game win-win lose-lose win-lose Pareto-optimal Pareto-improvement Kaldor-Hicks-improvement cost-benefit-analysis cost-effectiveness-analysis multi-criteria-decision-analysis decision-support-system expert-system knowledge-based-system rule-based-system case-based-reasoning machine-learning deep-learning neural-network random-forest support-vector-machine decision-tree regression-tree classification-tree clustering-algorithm dimensionality-reduction feature-selection feature-extraction pattern-recognition image-analysis signal-processing time-series-analysis spatial-analysis geographic-information-system remote-sensing satellite-imagery aerial-photography drone-imagery LiDAR radar sonar seismic geophysical geochemical biogeochemical ecoinformatics bioinformatics computational-biology systems-biology synthetic-biology biotechnology nanotechnology genetic-engineering genome-editing CRISPR TALEN zinc-finger-nuclease transgenic cisgenic intragenic subgenic agrobacterium-mediated-transformation biolistics electroporation microinjection protoplast-fusion somatic-hybridization cybrid asymmetric-hybrid synthetic-polyploid doubled-haploid haploid-induction anther-culture microspore-culture ovule-culture embryo-culture endosperm-culture protoplast-culture cell-culture tissue-culture organ-culture shoot-culture root-culture callus-culture suspension-culture bioreactor hairy-root-culture transformation hardening micropropagation clonal-propagation meristem-culture shoot-tip-culture axillary-bud-culture nodal-culture leaf-culture petiole-culture stem-culture internode-culture rhizome-culture bulb-culture corm-culture tuber-culture seed-culture zygotic-embryo-culture somatic-embryo-culture somatic-embryogenesis organogenesis caulogenesis rhizogenesis adventitious-shoot-formation direct-regeneration indirect-regeneration primary-metabolism secondary-metabolism biosynthesis biotransformation bioconversion biodegradation bioremediation phytoremediation mycoremediation zooremediation bioaugmentation biostimulation natural-attenuation monitored-natural-attenuation engineered-natural-attenuation permeable-reactive-barrier pump-and-treat soil-vapor-extraction air-sparging thermal-desorption incineration landfilling containment capping slurry-wall grout-curtain sheet-piling clay-liner geomembrane geotextile geogrid geocomposite geosynthetic geofabric geonet geocell geopipe geostrip geofoam geobag geotube geocontainer geomat geoblanket geospacer geodrain geoinjection geogrout geopier geocolumn geopile geowall geoberm geotrench geocut geofill geolift geoslab geopanel geoblock geobrick geotile geopaver geoweb geomattress geocushion
Trichonephila clavata
Jorō spider, Joro Spider, Parachute spider
Trichonephila clavata, commonly known as the Jorō spider, is a large orb-weaving spider native to East Asia that has become established as an invasive species in the southeastern United States since approximately 2010. First confirmed in Georgia in 2014, it has expanded rapidly across multiple states through a combination of ballooning dispersal and human-mediated transport. The species is notable for its substantial size, striking coloration, and extensive golden webs, but poses minimal risk to humans due to small fangs and docile behavior. Its physiological adaptations—including higher metabolic rate, faster heart rate, and greater cold tolerance than its congener Trichonephila clavipes—suggest potential for continued northward range expansion.
invasive-species orb-weaver ballooning urban-wildlife biological-control cold-tolerance startle-response cannibalism kleptoparasitism Japanese-folklore Nephilidae Trichonephila spider-silk range-expansion physiological-adaptation urban-ecology agricultural-pest-predator iNaturalist Georgia Maryland East-Asia United-States spider-bite venom docile non-aggressive web-architecture egg-sac overwintering sexual-dimorphism metabolic-rate heart-rate freeze-tolerance road-ecology prey-capture competition native-spider-impacts public-perception media-sensationalism management pesticide-alternatives broom-removal ornamental structural-pest ornithology cardinal web-strength trophic-interactions DNA-metabarcoding diet-analysis species-distribution-model physiological-entomology behavioral-ecology arachnology ballooning-dispersal human-mediated-dispersal hitchhiker cargo-stowaway port-of-entry climate-change warming northward-expansion DMV Howard-County Elkridge Ilchester West-Elkridge beech-forest secondary-forest suburban roadside telephone-pole streetlamp forest-edge disturbance-tolerance prey-detection vibrational-noise auditory-disturbance motionless freezing shyness Hannibal-Lecter stink-bug lanternfly evolutionary-history ancient-association ecosystem-service biocontrol natural-enemy invasive-pest brown-marmorated-stink-bug spotted-lanternfly Halyomorpha-halys Lycorma-delicatula competitive-exclusion dominant-orb-weaver abundant population-growth rapid-spread range-span 120,000-km2 physiological-plasticity metabolism twice-as-high 77%-higher-heart-rate 74%-freeze-survival development-rate seasonal-window life-cycle-completion thermal-limitation congener-comparison natural-dispersal human-transport iNaturalist-record citizen-science confirmed-sighting breeding-population male-and-female gravid-female egg-case stowaway cargo-container port-introduction local-port vehicle-transport hitchhiking accelerated-arrival several-years natural-means prognostication prediction model suitable-habitat abundant-suitable-habitat eastern-North-America western-North-America potential-range invasive-range potential-impact environmental-impact ecological-impact ecosystem-impact monitor cautious evidence-based reasonable-journalism sensationalism exaggeration uncritical-acceptance harmless carefully-monitored next-steps research-opportunity call-for-research knowledge-gap little-known novel-ecosystem spreading rapidly-expanding invasive-predator invasive-orb-weaver non-native introduced established colonized thriving doing-fine happy merely-20-minutes-away road-trip visit access-spread beech-leaf-disease BLD state-forest remarkable-colony amongst-beech-trees confirmed-prognostication Davis-and-Frick escape-relative-warmth expand-range-northward eastern-seaboard merely-20-minutes-away-from-home resist-opportunity amazing-predators tales-of-arrival previous-episodes 2022 2024 reduce-angst large-non-native-spider establishing-in-DMV facts-presented past-episode venomous-terrible-painful Nah expert-Rick-Hoebeke risks-small puny-fangs unlikely-pierce-skin video completely-non-aggressive eastern-Howard-County haphazard-webs littered-with-remains former-victims leaves shed-exoskeletons much-larger-female dwarfs-mate positioned-just-above strand-of-silk spinnerets underside-of-abdomen near-red-mound relative-to-hand large-and-docile wait-and-see what-Jorō-means-to-ecosystems help-other-spiders put-a-beat-down-on-invasive-pests stink-bugs spotted-lanternflies passive-hunters enormous-webs larger-than-a-meter-in-diameter capture-prey-snared-in-silk arachnophobes-scary arachnophiles-beautiful important-ecosystem-services crop-pests native-range Asia having-an-old-friend-over-for-dinner reunite first-time large-spiders juicy-prey-items feathered-and-non-feathered-reptiles poses-no-known-threat difficult-to-predict-impact non-native-species experts-suggest beyond-scary-mien give-indigenous-large-orb-weavers-run-for-their-money black-and-yellow-garden-spider marbled-orb-weaver spotted-orb-weaver other-parts-of-world most-abundant-and-dominant-orb-weaver what-will-it-mean only-time-will-tell final-tidbit Jorō-shapeshifter Jorō-gumo beautiful-woman seduces-men binds-with-silk devours bad-date acknowledgements Rick-Hoebeke identifying-Jorō insights David-Coyle Bob-Bellinger great-images knowledge studies Veni-vidi-vici Nelsen-et-al.Physiological-evaluation Nephila-clavata-newly-recorded Hoebeke-et-al.Life-Cycle-Habitat-and-Variation Moore Startle-responses Davis-and-Anerao How-Urban-Tolerant-Are-They Davis-et-al.inspiration details stars Dave-Clement Miri-Talabac Maddie-Potter hooked-up-with-colony differentiate resources.ipmcenters.org link Jorō-guru David-Coyle's-take YouTube calm-your-fears Journal-of-Economic-Entomology Journal-of-Medical-Entomology chemical-management-strategies effective-management-methods spider-control-products safe-and-effective unlikely-to-bite generally-harmless big-spiders colorful-spiders invasive-spiders in-your-face-spiders webs-on-homes landscape-plants structures middle-of-where-people-live-work-and-play northern-Georgia-2014 presumed-arrived-few-years-prior 15-years increased-dramatically much-of-Southeast disjunct-populations northeastern-U.S.firestorm-of-interest rapidly-expanding-range southeastern-states many-counties Carolinas Tennessee Oklahoma more-than-a-century parts-of-Florida rarely-Pennsylvania tropics thermal-limitations higher-metabolism faster-heart-rate better-ability-to-tolerate-freezing complete-life-cycle-rapidly chilly-temperatures suite-of-adaptations expand-northward warm-loving-cousin findings-support-potential much-yet-to-be-learned successfully-survive-northern-winters nation-and-world-warm southern-species-expand higher-latitudes-and-altitudes rapidly-range-expansion-will-occur typical-mode-of-dispersal aerial-dispersal ballooning-on-strands-of-silk Charlotte's-babies Charlotte's-Web spectacular-moniker parachute-spider rain-down-from-airplanes long-distance-transit human-assistance good-hitchhikers inseminated-gravid-female egg-case-stowaway arrival-in-DMV several-years-by-natural-means new-introduction-at-local-port human-assist-in-vehicles accelerate-arrival terrible-and-painful small-fangs visited-cousin golden-silk-spider Costa-Rica exceeding-meter-diameter indigenous-large-orb-weavers run-for-their-money minimal-direct-impact firestorm Andrew-Davis Benjamin-Frick University-of-Georgia past-week native-to-eastern-Asia Japan China Korea Taiwan known-in-Georgia-since-2013 spreading-rapidly joined-its-cousin Trichonephila-clavipes established-over-160-years better-tolerate-freezing survive-northern-winters rapidly-range-expansion typical-mode long-distance new-introduction human-assist arachnophobes arachnophiles brown-marmorated-stink-bugs difficult-to-predict scary-mien indigenous-orb-weavers Mary-Nouri Sarah-Morgan WAMU relatively-harmless spider-season fall leaves-changing-color cool-crisp-autumn-air pumpkin-spice extension-entomologist Southeast not-just-spiders 20-feet-wide homes people-live-work-and-play 15-years-or-so influx-of-calls public entomology-extension-programs media-frenzy two-questions get-rid-of-Jorō-spiders are-Jorō-spiders-dangerous Clemson-University Southern-Adventist-University Washington-College Union-College two-new-studies test-methods-for-eliminating how-likely-to-bite harm scoured-internet recommended-treatment-options spider-specific-management-recommendations medically-relevant widow-spiders brown-recluses do-not-enter-homes relatively-new recommended-products labelled-spider-control-products tested-products several-products-eliminate not-legal-pesticides machine-lubricant not-recommended commercially-available strongly-encourage labelled-products easiest-and-cheapest without-chemicals broom-or-stick walked-into-spider-web rode-bike took-spider-to-face not-great-feeling wonder-if-bitey give-bitten-superpowers unique-studies evaluated-behaviors react-to-people rather-drop-off-web hang-around-and-bite really-provoke volunteered-to-get-bitten yes-you-read-correctly akin-to-mosquito-bite little-redness little-swelling largely-gone-next-day no-acquired-superpowers take-home-message twofold simplest-and-most-effective without-pesticides feel-you-need-pesticides product-labeled-for-spider-control try-not-to-freak-out doesn't-want-to-be-on-you odds-of-getting-bitten-extremely-low enjoy-glorious-fall-weather drink-pumpkin-spiced-beverage watch-where-you-walk Keep-calm-and-carry-on bites-unlikely cause-minimal-discomfort associate-professor Department-of-Forestry-and-Environmental-Conservation socials email extension featured pest-management Entomology-Today latest-posts email-subscription March-2022 wondered make-its-way-to-DMV September-2022 two-observations three-years 40-sightings happy-and-doing-just-fine several-Howard-County-locations week-or-so-ago team-of-scientists University mission killing-ultra-valuable-beech-trees thriving-amongst-beech-trees visit-amazing-predators mature-females both-Trichonephila-species three-locally-common-orb-weaving-species time-spent-immobile mild-disturbance brief-puff-of-air air-puff-response-data five-other-North-American-species published-literature 453-observations freezing-behavior 10-spider-species remained-immobile-under-a-minute both-Trichonephila-spiders over-an-hour unprecedented shyest-ever-documented tolerate-urban-environments remaining-motionless fleeing nonsexual-cannibalism mating-activity females-cannibalize-males food-availability territorial-aggression Southeastern-United-States expanding-range shy behavioral-reactions descriptive-observations photo-documentation anecdotal-observations controlled-pairings lab natural-webs field one-female-biting-and-killing-another short-fight similar-size container 25-trials fights-ensued-40%different-sizes 27-trials fights-happened-18%larger-females-not-always-aggressor 52-lab-trials six-bouts 9%direct-killing field-trials empty-web 14-trials one-fight 7%aggressor-killing-and-wrapping silk away-from-any-web not-necessarily-territoriality intraspecific-aggression provoked-or-stressed deserves-more-study newly-invasive biology physiology new-range closely-related-species successfully-established 160-years investigation compare online-records iNaturalist.org seasonal-distributions timing females physiological-traits environmental-tolerance metabolic-rates cardiac-functioning cold-exposure survival brief-below-freezing-temperature shorter-season complete-lifecycle narrow-period suitable-weather inherently-higher-metabolism low-temperature survive-better 74%-compared-to-50%brief-freeze colder-climatic-region southeastern-USA useful-information planning urban-landscapes busy-roads frequent-disturbances auditory-and-vibrational physiological-or-behavioral-changes northeast-Georgia varying-levels-of-road-traffic prey-capture-behavior simulated-prey tuning-fork 128-hz-frequency touched-to-web attacked 357-total-trials 20-different-roads attacked-59%high-variability some-roadsides-over-80%others-less-than-30%small-but-significant-negative-correlation daily-road-traffic spider-attack-rates moderate--to-heavy-traffic-roads slightly-less-likely-to-attack 51%-vs-65%able-to-live-near-roads cost-in-terms-of-prey-capture did-not-weigh-less compensate-for-disturbance accumulating-evidence human-dominated-landscapes aid-spread introduced-range Eriostethus-rufus polysphinctine-ectoparasitoid araneid-spiders Neoscona-spp.endemic-to-Japan Yamagata-Prefecture 38º46'-N northern-Japan northernmost-record genus identification-confirmed morphology DNA-barcoding cocoon large-modified-web unique inverted-triangle over-50-cm hanging-from-ill-defined-part no-organized-structure host-spider circumstantial-evidences structure-of-modified-web pre-existing-web-partly-reused orb-web-completely-removed sustaining-threads-newly-moored parasitises-host-spiders two-subfamilies unusual-for-group invasion-potential known-invasive-range review dispersal-abilities potential-impacts management-strategies October-2022 at-least-120,000-km2 South-Carolina North-Carolina Alabama West-Virginia pattern-of-spread natural-dispersal-mechanisms human-mediated-transport large-bodied-orb-weavers flying-insects small-animals thirteen-co-occurring-spider-species monitored-for-competition resources web-building-sites natural-and-urban-habitats management-options-limited advise-journalists-and-experts against-exaggerating-potential-environmental-impact ecologically-harmless rapid-spread-carefully-monitored cautious-evidence-based-approach determining-next-steps three-material-types web-capture introduced-species alter-established-trophic-interactions molecular-analysis resolve-changes community-structure foraging-links east-coast fecal-samples prey-remains-from-webs dissected-guts compare-diet-composition arthropod-targeted-COI-primers Illumina-MiSeq highest-diversity highest-richness highest-proportion-prey-reads recovery-of-prey-reads dissected-gut-content low overwhelmed-by-Jorō-spider-DNA high-proportions-Jorō-spider-reads higher-prey-diversity-and-richness detected-prey-DNA several-days-after-capture initial-gut-retention-time-estimates first-glimpse complexity-of-trophic-associations introduced-web-building-spider viable-material source-of-prey-DNA estimates-of-biodiversity Jammu-and-Kashmir northernmost-shifting document-presence behavior habitat-adaptation colder-temperature-environment Goha-Tehsil Doda-district 1556.74-meters road-bridge nearby-vegetation brightly-colored-abdomens black green yellow red long-legs yellow-and-black-bandings predominantly-seen smaller fewer unique-behaviors remaining-motionless-after-disturbances 5-10-meters-of-roads adaptation-to-human-settlements field-observations image-documentation expert-confirmation thrive-in-colder-conditions 15-20-degrees-Celsius October-November no-spider-found higher-elevations significantly-lower-temperatures adaptation-limited-to-milder-cold-regions new-insight adaptation need-for-further-research non-native-habitats eggs overwinter harsh-weather-conditions no-protection compact-silk-case milky-coating chorionic-microspheres CM fine-structural-characteristics ecological-importance uneven-size-distribution outer-eggmass evenly-covered single-layer diameter-2.3-µm short-papillary-projections segmental-adhesion mucous-components insoluble-in-water partially-soluble-in-absolute-ethanol spherical-structure HFIP strong-organic-solvent not-derived-from-vitellogenic-or-choriogenetic-processes CM-adhesive-coatings ovipositional-process equivalent-to-cocoon-silk protective-functions silken-eggcase Northern-Cardinal Cardinalis-cardinalis perches steals-food southeast-of-the-United-States questions native-fauna-affected consuming-prey-items example native-species-deriving-benefit manner-of-kleptoparasitism perched-directly-on-web supported-its-weight 42-48-g first-documented-case spider-web-supporting-perching-bird measurements force-gauge typical-webs support-masses-up-to-70-g small-but-growing-body-of-knowledge non-native-spider species-distribution-models ecological-niche-modeling predict-areas establish-if-introduced three-distinct-genetic-lineages mitochondrial-COI genome-wide-SNPs Korean-Peninsula

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