Expanding-range

Guides

• Ixodes affinis

  Ixodes affinis is a hard-bodied tick species distributed across the southeastern United States, Mexico, Central America, and South America. Unlike its congener Ixodes scapularis, this species does not bite humans but serves as a competent vector for Borrelia burgdorferi sensu stricto, the causative agent of Lyme disease. Immature stages feed on birds and small mammals, while adults parasitize larger mammals including deer and dogs. Its expanding range into southeastern Virginia and association with migratory birds has raised concerns about its role in pathogen dispersal.

  tickvectorLyme-diseaseBorrelia-burgdorferiavian-hostrange-expansionsoutheastern-United-StatesMexicoCentral-Americanon-human-bitingenzootic-maintenanceco-feedingmigratory-birdsIxodidaeAcariparasitethree-host-tickquesting-behaviorwildlife-disease-ecologyforest-habitatdog-parasitewhite-tailed-deerCarolina-wrenwhite-throated-sparrowOld-Dominion-University-Tick-TeamHolly-GaffErin-Hellermolecular-identificationDNA-sequencingengorged-specimen-identificationpine-plantationrural-Yucatánhunting-dogsTixméhuacOpichénseasonal-activitylaboratory-rearingimmature-stageslarvanymphadultpathogen-dispersalclimate-changehabitat-fragmentationsuburbanizationdeer-populationacorn-mastingsmall-mammal-reservoirBorrelia-miyamotoico-infectiontick-borne-diseasepublic-healthveterinary-parasitologymedical-entomologyvector-ecologyhost-parasite-interactionbird-migrationlong-distance-dispersalboreal-forestCanadaManitobaSwainson's-thrushCatharus-ustulatusmigratory-birdground-foraging-birdpasserinesongbirdbrown-thrashereastern-towheeAmerican-robinnorthern-cardinalmarsh-rice-ratcotton-mouseeastern-gray-squirrelraccoonsouthern-short-tailed-shreweastern-molewhite-footed-mousePeromyscus-leucopuschipmunkTamiasoak-mastingQuercusforest-floorquesting-heighthumidity-preferencewinter-activitytick-removaltick-testingtick-borne-illness-preventionDEETpermethrintick-inspectiontick-avoidanceAIR-protocolArachnidaParasitiformesChelicerataEukaryotaMetazoaAnimaliaArthropodaIxodidahard-tickixodidNeumann-1899taxonomysystematicsdorsal-scutumcapitulumhypostomepalpsspiracular-platesfestoonsornateinornateidiosomagnathosomapodonscutumalloscutumbasis-capitulicornuaporose-areassclerotizedcuticlesetaesensillaHaller's-organtarsusclawspulvilliscutum-shapepunctationsgroovescoxaetrochantersfemoragenuatibiaetarsipretarsiadanal-platesaccessory-platesspiraclestracheaesalivary-glandsmidgutMalpighian-tubulesreproductive-systemovarytestisgenital-apertureanuspreanal-groovepostanal-groovesubanal-platesparanal-platesmarginal-groovespseudoscutumscapularscervical-groovespunctatereticulateshagreenedwrinkledsmoothglabroussetiferouspilosehispidgranularmammillatedtuberculatecarinatedsulcatestriatecostatelineatepunctate-striatereticulate-punctateIXO_affNCBI:txidGBIF:speciesCatalogue-of-Life:speciesiNaturalist:taxonWikipedia:speciesIxodesaffinisbird-tickdeer-tickdog-tickrabbit-tickshrew-tickmole-tickmouse-tickrat-ticksquirrel-tickraccoon-tickwren-tickthrasher-tickrobin-ticktowhee-tickcardinal-ticksparrow-tickthrush-tickmigratory-tickinvasive-tickexpanding-rangeemerging-vectorcryptic-speciesspecies-complexmorphological-identificationDNA-barcodingCOI16SITS2tick-microbiomepathogen-detectionBorreliaspirochetezoonosisvector-competencereservoir-competenceenzootic-cycleepizooticspilloverbridge-vectoramplifying-hostreservoir-hostmaintenance-hostincidental-hostdead-end-hosthost-specificityhost-rangehost-switchinghost-raceecological-fittingniche-expansionrange-shiftphenologydiapauseoverwinteringsurvivalmortalityfecundityreproductive-outputegg-massovipositioneclosionmoltingecdysisinstarstadiumdevelopmental-timegeneration-timelife-spanlongevitypopulation-dynamicsabundanceprevalenceintensityaggregationnegative-binomialzero-inflatedspatial-distributiontemporal-distributionhabitat-associationmicrohabitatvegetation-typeforest-typesuccessional-stageedge-effectfragmentationconnectivitymatrixcorridorpatchlandscape-ecologygeographic-information-systemremote-sensingmodelingdistribution-modelingecological-niche-modelingMaxEntBIOMODclimate-envelopebioclimatic-variablestemperatureprecipitationhumiditysaturation-deficitvapor-pressure-deficitevapotranspirationseasonalityphenology-mismatchglobal-warmingextreme-weatherdroughtfloodphenological-shiftrange-contractionpoleward-shiftelevation-shiftaltitudinal-gradientlatitudinal-gradientcontinentalinsularendemiccosmopolitanpantropicalholarcticnearcticneotropicalpalearcticafrotropicalorientalaustralasianoceanianantarcticbiogeographyphylogeographypopulation-geneticsgene-flowgenetic-structureisolation-by-distanceisolation-by-environmentadaptive-evolutionlocal-adaptationphenotypic-plasticitycryptic-diversityspecies-delimitationintegrative-taxonomymorphometricsgeometric-morphometricstraditional-morphometricslandmarkoutlinesemilandmarkcharacter-matrixphylogenyclassificationnomenclaturetype-specimenholotypeparatypesyntypelectotypeneotypeoriginal-descriptionredescriptionemendationsynonymyhomonymypriorityvalidityavailabilitypublished-workZooBankLSIDURNDOIORCIDauthoryearjournalvolumeissuepagesfigureplatemapdrawingphotographSEMTEMlight-microscopyconfocal-microscopymicroCTscanning-electron-microscopytransmission-electron-microscopyhistologyimmunohistochemistryin-situ-hybridizationPCRqPCRRT-PCRddPCRsequencingSangernext-generationIlluminaPacBioOxford-Nanoporegenometranscriptomeproteomemetabolomemicrobiomeviromebacteriomefungomeparasitomeholobiontsymbiosismutualismcommensalismparasitismpathogenesisvirulencetransmissionvectorial-capacitybasic-reproduction-numberR0force-of-infectionincidence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I.-malayensisI.-marxiI.-masloviI.-matopiI.-minorI.-monospinosusI.-moreliI.-moschiferiI.-murisI.-murreletiI.-myospalacisI.-nchisiensisI.-necromysI.-neitziI.-nipponensisI.-nuttalliI.-ochotonaeI.-okapiaeI.-oldiI.-ornithorhynchiI.-ovatusI.-pacificusI.-pararicinusI.-pavlovskyiI.-percavatusI.-peromysciI.-persulcatusI.-pomerantziI.-pterodromaeI.-quadratusI.-raccoonisI.-rageauiI.-randrianasoloiI.-rasusI.-redikorzeviI.-ricinusI.-rothschildiI.-rotundatusI.-rubicundusI.-rugicollisI.-rugosusI.-sapeloensisI.-sarsiI.-scapularisI.-schillingsiI.-sculptusI.-semenoviI.-soricisI.-spinaeI.-steiniI.-stilesiI.-stromiI.-subterranusI.-tancitariusI.-tanukiI.-tapirusI.-tasmaniI.-tayloriI.-tertiariusI.-texanusI.-theileraeI.-thomasaeI.-tovariI.-transvaalensisI.-triangulicepsI.-trichosuriI.-turdusI.-uriaeI.-vandenbergiI.-vanidicusI.-ventalloiI.-vespertilionisI.-victoriensisI.-walkeraeI.-werneriI.-woodiI.-zaglossiI.-zairensistick-taxonomytick-systematicstick-phylogenytick-evolutiontick-biogeographytick-ecologytick-physiologytick-behaviortick-reproductiontick-developmenttick-nutritiontick-metabolismtick-respirationtick-excretiontick-osmoregulationtick-neurobiologytick-sensory-biologytick-chemoreceptiontick-thermoreceptiontick-mechanoreceptiontick-photoreceptiontick-circadian-rhythmtick-biological-clocktick-seasonalitytick-diapausetick-overwinteringtick-cold-hardinesstick-desiccation-resistancetick-water-balancetick-cuticletick-moltingtick-ecdysistick-growthtick-feedingtick-engorgementtick-blood-mealtick-digestiontick-salivationtick-saliva-compositiontick-salivary-gland-transcriptometick-salivary-proteometick-pharmacologytick-toxicologytick-venomtick-paralysis-toxintick-borne-pathogentick-borne-virustick-borne-bacteriumtick-borne-protozoantick-borne-helminthtick-borne-rickettsiatick-borne-ehrlichiatick-borne-anaplasmatick-borne-borreliatick-borne-babesiatick-borne-theileriatick-borne-hepatozoontick-borne-coxiellatick-borne-francisellatick-borne-bartonellatick-borne-chlamydiatick-borne-mycoplasmatick-borne-spiroplasmatick-borne-arbovirustick-borne-flavivirustick-borne-bunyavirustick-borne-reovirustick-borne-orbivirustick-borne-coltivirustick-borne-nairovirustick-borne-phlebovirustick-borne-thogotovirustick-borne-orthomyxovirustick-borne-arenavirustick-borne-hantavirustick-borne-paramyxovirustick-borne-rhabdovirustick-borne-bornavirustick-borne-filovirustick-borne-coronavirustick-borne-poxvirustick-borne-herpesvirustick-borne-adenovirustick-borne-papillomavirustick-borne-polyomavirustick-borne-circovirustick-borne-parvovirustick-borne-anellovirustick-borne-dependovirustick-borne-tenuivirustick-borne-deltavirustick-borne-hepadnavirustick-borne-retrovirustick-borne-lentivirustick-borne-spumavirustick-borne-metavirustick-borne-pseudovirustick-borne-hemivirustick-borne-novirhabdovirustick-borne-peribunyavirustick-borne-phenuivirustick-borne-bandavirustick-borne-phasivirustick-borne-tulasvirustick-borne-orthonairovirustick-borne-shavivirustick-borne-sathuperi-virustick-borne-salehabad-virustick-borne-dera-ghazi-khan-virustick-borne-hughes-virustick-borne-farallon-virustick-borne-punta-salinas-virustick-borne-sapphire-II-virustick-borne-zirqa-virustick-borne-keterah-virustick-borne-thiafora-virustick-borne-erve-virustick-borne-malsoor-virustick-borne-tibet-virustick-borne-qalyub-virustick-borne-yogue-virustick-borne-kasokero-virustick-borne-lebombo-virustick-borne-nyamanini-virustick-borne-midway-virustick-borne-tokar-virustick-borne-absettarov-virustick-borne-hypr-virustick-borne-kumlinge-virustick-borne-omsk-hemorrhagic-fever-virustick-borne-langat-virustick-borne-louping-ill-virustick-borne-kyasanur-forest-disease-virustick-borne-alkhurma-hemorrhagic-fever-virustick-borne-powassan-virustick-borne-deer-tick-virustick-borne-heartland-virustick-borne-bourbon-virustick-borne-colorado-tick-fever-virustick-borne-eyach-virustick-borne-banna-virustick-borne-seadornavirustick-borne-great-island-virustick-borne-kemerovo-virustick-borne-lipovnik-virustick-borne-tribec-virustick-borne-chobar-gorge-virustick-borne-wad-Medani-virustick-borne-dugbe-virustick-borne-crimean-congo-hemorrhagic-fever-virustick-borne-nairobi-sheep-disease-virustick-borne-ganjam-virustick-borne-bandia-virustick-borne-sierra-nevada-virustick-borne-lone-star-virustick-borne-severe-fever-with-thrombocytopenia-syndrome-virustick-borne-huaiyangshan-virustick-borne-guertu-virustick-borne-Alongshan-virustick-borne-wenzhou-virustick-borne-wuhan-virustick-borne-yongjia-virustick-borne-chiba-virustick-borne-natori-virustick-borne-odawara-virustick-borne-kowloon-virustick-borne-tamanavirus

• Listronotus maculicollis

  annual bluegrass weevil

  Listronotus maculicollis, the annual bluegrass weevil, is a highly destructive pest of short-mown turfgrass in eastern North America, particularly golf course putting greens, fairways, and tee boxes. Adults are small (up to 4.5 mm), dark-colored weevils with a distinct spotted or maculate pronotum. The species has expanded its range from the northeastern United States into the Midwest and Southeast since the 2000s. Larval feeding causes severe damage, with early instars acting as stem-borers and later instars feeding on the crown, resulting in irregular patches of dead turf. The weevil shows strong preference for Poa annua (annual bluegrass) over other turfgrass species, though it can develop on several cool-season grasses. Temperature, not photoperiod, drives daily activity patterns, with peak activity occurring at 14–17°C.

  turfgrass-pestgolf-course-peststem-borercrown-feedertemperature-dependent-activityinsecticide-resistancePoa-annua-specialistexpanding-rangeweevilColeopteraCurculionidaeintegrated-pest-managementbiological-controlnematode-controlfluorescent-marking-technique

• Spoladea

  beet webworm, Hawaiian beet webworm, amaranth leaf-webber

  Spoladea is a genus of Crambidae moths containing two described species, with S. recurvalis being a significant agricultural pest of amaranth, spinach, and sugar beet across tropical and subtropical regions worldwide. The genus was established by Guenée in 1854. S. recurvalis has achieved nearly cosmopolitan distribution through human-mediated dispersal and is subject to intensive study for molecular mechanisms of pheromone production, flight behavior, and host plant interactions to support integrated pest management.

  agricultural-pestamaranth-pestbiological-controltranscriptomepheromoneflight-behaviorexpanding-rangecitizen-science

• Trichonephila clavata

  Jorō spider, Joro Spider, Parachute spider

  Trichonephila clavata, commonly known as the Jorō spider, is a large orb-weaving spider native to East Asia that has become established as an invasive species in the southeastern United States since approximately 2010. First confirmed in Georgia in 2014, it has expanded rapidly across multiple states through a combination of ballooning dispersal and human-mediated transport. The species is notable for its substantial size, striking coloration, and extensive golden webs, but poses minimal risk to humans due to small fangs and docile behavior. Its physiological adaptations—including higher metabolic rate, faster heart rate, and greater cold tolerance than its congener Trichonephila clavipes—suggest potential for continued northward range expansion.

  invasive-speciesorb-weaverballooningurban-wildlifebiological-controlcold-tolerancestartle-responsecannibalismkleptoparasitismJapanese-folkloreNephilidaeTrichonephilaspider-silkrange-expansionphysiological-adaptationurban-ecologyagricultural-pest-predatoriNaturalistGeorgiaMarylandEast-AsiaUnited-Statesspider-bitevenomdocilenon-aggressiveweb-architectureegg-sacoverwinteringsexual-dimorphismmetabolic-rateheart-ratefreeze-toleranceroad-ecologyprey-capturecompetitionnative-spider-impactspublic-perceptionmedia-sensationalismmanagementpesticide-alternativesbroom-removalornamentalstructural-pestornithologycardinalweb-strengthtrophic-interactionsDNA-metabarcodingdiet-analysisspecies-distribution-modelphysiological-entomologybehavioral-ecologyarachnologyballooning-dispersalhuman-mediated-dispersalhitchhikercargo-stowawayport-of-entryclimate-changewarmingnorthward-expansionDMVHoward-CountyElkridgeIlchesterWest-Elkridgebeech-forestsecondary-forestsuburbanroadsidetelephone-polestreetlampforest-edgedisturbance-toleranceprey-detectionvibrational-noiseauditory-disturbancemotionlessfreezingshynessHannibal-Lecterstink-buglanternflyevolutionary-historyancient-associationecosystem-servicebiocontrolnatural-enemyinvasive-pestbrown-marmorated-stink-bugspotted-lanternflyHalyomorpha-halysLycorma-delicatulacompetitive-exclusiondominant-orb-weaverabundantpopulation-growthrapid-spreadrange-span120,000-km2physiological-plasticitymetabolismtwice-as-high77%-higher-heart-rate74%-freeze-survivaldevelopment-rateseasonal-windowlife-cycle-completionthermal-limitationcongener-comparisonnatural-dispersalhuman-transportiNaturalist-recordcitizen-scienceconfirmed-sightingbreeding-populationmale-and-femalegravid-femaleegg-casestowawaycargo-containerport-introductionlocal-portvehicle-transporthitchhikingaccelerated-arrivalseveral-yearsnatural-meansprognosticationpredictionmodelsuitable-habitatabundant-suitable-habitateastern-North-Americawestern-North-Americapotential-rangeinvasive-rangepotential-impactenvironmental-impactecological-impactecosystem-impactmonitorcautiousevidence-basedreasonable-journalismsensationalismexaggerationuncritical-acceptanceharmlesscarefully-monitorednext-stepsresearch-opportunitycall-for-researchknowledge-gaplittle-knownnovel-ecosystemspreadingrapidly-expandinginvasive-predatorinvasive-orb-weavernon-nativeintroducedestablishedcolonizedthrivingdoing-finehappymerely-20-minutes-awayroad-tripvisitaccess-spreadbeech-leaf-diseaseBLDstate-forestremarkable-colonyamongst-beech-treesconfirmed-prognosticationDavis-and-Frickescape-relative-warmthexpand-range-northwardeastern-seaboardmerely-20-minutes-away-from-homeresist-opportunityamazing-predatorstales-of-arrivalprevious-episodes20222024reduce-angstlarge-non-native-spiderestablishing-in-DMVfacts-presentedpast-episodevenomous-terrible-painfulNahexpert-Rick-Hoebekerisks-smallpuny-fangsunlikely-pierce-skinvideocompletely-non-aggressiveeastern-Howard-Countyhaphazard-webslittered-with-remainsformer-victimsleavesshed-exoskeletonsmuch-larger-femaledwarfs-matepositioned-just-abovestrand-of-silkspinneretsunderside-of-abdomennear-red-moundrelative-to-handlarge-and-docilewait-and-seewhat-Jorō-means-to-ecosystemshelp-other-spidersput-a-beat-down-on-invasive-pestsstink-bugsspotted-lanternfliespassive-huntersenormous-webslarger-than-a-meter-in-diametercapture-prey-snared-in-silkarachnophobes-scaryarachnophiles-beautifulimportant-ecosystem-servicescrop-pestsnative-rangeAsiahaving-an-old-friend-over-for-dinnerreunitefirst-timelarge-spidersjuicy-prey-itemsfeathered-and-non-feathered-reptilesposes-no-known-threatdifficult-to-predict-impactnon-native-speciesexperts-suggestbeyond-scary-miengive-indigenous-large-orb-weavers-run-for-their-moneyblack-and-yellow-garden-spidermarbled-orb-weaverspotted-orb-weaverother-parts-of-worldmost-abundant-and-dominant-orb-weaverwhat-will-it-meanonly-time-will-tellfinal-tidbitJorō-shapeshifterJorō-gumobeautiful-womanseduces-menbinds-with-silkdevoursbad-dateacknowledgementsRick-Hoebekeidentifying-JorōinsightsDavid-CoyleBob-Bellingergreat-imagesknowledgestudiesVeni-vidi-viciNelsen-et-al.Physiological-evaluationNephila-clavata-newly-recordedHoebeke-et-al.Life-Cycle-Habitat-and-VariationMooreStartle-responsesDavis-and-AneraoHow-Urban-Tolerant-Are-TheyDavis-et-al.inspirationdetailsstarsDave-ClementMiri-TalabacMaddie-Potterhooked-up-with-colonydifferentiateresources.ipmcenters.orglinkJorō-guruDavid-Coyle's-takeYouTubecalm-your-fearsJournal-of-Economic-EntomologyJournal-of-Medical-Entomologychemical-management-strategieseffective-management-methodsspider-control-productssafe-and-effectiveunlikely-to-bitegenerally-harmlessbig-spiderscolorful-spidersinvasive-spidersin-your-face-spiderswebs-on-homeslandscape-plantsstructuresmiddle-of-where-people-live-work-and-playnorthern-Georgia-2014presumed-arrived-few-years-prior15-yearsincreased-dramaticallymuch-of-Southeastdisjunct-populationsnortheastern-U.S.firestorm-of-interestrapidly-expanding-rangesoutheastern-statesmany-countiesCarol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