Parasite
Guides
Acanthocephala
thorny-headed worms, spiny-headed worms, acanthocephalans
Acanthocephala is a phylum of obligate endoparasitic worms known as thorny-headed or spiny-headed worms, characterized by an eversible proboscis armed with recurved hooks used to attach to host intestinal walls. They lack a mouth and digestive system, absorbing nutrients directly through their tegument. The phylum contains approximately 1,200–1,420 described species with complex life cycles involving arthropod intermediate hosts and vertebrate definitive hosts. Recent molecular evidence indicates they are highly modified rotifers, now sometimes classified within Syndermata or Rotifera.
Acari
Mites and Ticks, mites, ticks
Acari is a subclass of arachnids containing mites and ticks. The group exhibits extraordinary diversity with fossil records dating to the early Devonian period. Recent phylogenetic analyses have challenged the monophyly of Acari, suggesting that the two main lineages—Parasitiformes (ticks and allies) and Acariformes (dust mites, spider mites, chiggers, and allies)—may not share a single common ancestor exclusive of other arachnids. This would render Acari a paraphyletic or polyphyletic grouping rather than a natural taxon. The traditional unifying character, the gnathosoma (a modified mouthpart region), has been shown to be absent or differently constructed in many species across both lineages.
Aceria paracalifornica
California Sagebrush Gall Mite
Aceria paracalifornica, commonly known as the California Sagebrush Gall Mite, is a species of eriophyid mite in the family Eriophyidae. As a member of the superfamily Eriophyoidea, it is a worm-like, four-legged mite that parasitizes plants. The genus Aceria includes numerous species that cause distinctive plant deformities including galls, witches' brooms, and leaf distortions. The specific epithet "paracalifornica" suggests a close relationship to or similarity with A. californica, a species associated with California sagebrush (Artemisia californica). Eriophyid mites in this genus are highly host-specific, typically specializing on a single plant species or genus.
Aegidae
aegid isopods
Aegidae is a family of marine and freshwater isopod crustaceans characterized by temporary parasitic relationships with fish hosts. Adults attach to hosts to feed on blood or tissue, then detach to digest meals. The family is distinguished from related Cirolanidae by having only three pairs of hook-like pereiopods rather than seven. Members occur in diverse aquatic habitats from shallow coastal waters to deep ocean environments, with some species documented at depths exceeding 2000 meters.
Amblyomma tuberculatum
gopher tortoise tick
Amblyomma tuberculatum is a hard tick species endemic to the United States, commonly known as the gopher tortoise tick due to its intimate association with the gopher tortoise (Gopherus polyphemus). Unlike many tick species, its distribution appears more restricted than that of its primary host, with environmental factors such as sand depth and soil composition playing significant roles in habitat suitability. Laboratory studies have documented its complete life cycle, which requires a minimum of 207 days under controlled conditions.
Anilocra
Fish Lice
Anilocra is a genus of marine isopods in the family Cymothoidae, commonly known as fish lice. Members are obligate external parasites of teleost fishes, attaching to host skin and feeding on blood. The genus exhibits sequential hermaphroditism, with individuals beginning life as males and transitioning to females as they grow. Parasitism by Anilocra species has been documented to reduce host growth rates, fecundity, and survival.
Aphomia sociella
Bee Moth, Bumble Bee Wax Moth
Aphomia sociella, commonly known as the bee moth or bumble bee wax moth, is a small pyralid moth native to Europe that has become an economically significant pest of bee colonies. The species is notable for its unique dual-modal mating system combining sex pheromones and ultrasonic courtship songs produced by males using specialized tegular tymbals. Females lay eggs in nests of bumblebees, wasps, and occasionally weakened honeybee hives, where larvae feed on nest materials, pollen, honey, and host brood. Infestation levels increase with proximity to commercial apiaries, causing reduced host reproductive output and weakened immune responses in bumblebees.
Arguloida
Fish Lice
Arguloida is an order of parasitic crustaceans commonly known as fish lice. The order contains a single family, Argulidae, whose members are obligate ectoparasites of freshwater and marine fishes. These organisms have an uncertain phylogenetic position within Maxillopoda and lack any known fossil record. They are distributed globally across temperate and tropical waters.
Arrenuridae
Arrenuridae is a family of water mites (Hydrachnidia) in the order Trombidiformes, containing at least three genera and approximately 110 described species. Members are primarily freshwater predators as adults and deutonymphs, with larvae that parasitize aquatic insects. The family exhibits complex mating behaviors mediated by chemical communication, including female-emitted sex pheromones that trigger male responses.
Arrenurus
water mites
Arrenurus is the largest genus of water mites, comprising approximately 950 species with cosmopolitan distribution in lentic freshwater habitats. Adults are heavily sclerotized, predatory, and exhibit marked sexual dimorphism—males possess diagnostic caudal modifications used in mating, while female taxonomy remains problematic. The life cycle includes seven stages: egg, inactive prelarva, parasitic larva, protonymph, deutonymph, tritonymph, and adult. Larvae are ectoparasites of aquatic insects, particularly Odonata (dragonflies and damselflies), but also Diptera and Coleoptera, utilizing hosts for both nutrition and phoretic dispersal.
Arrenurus problecornis
Arrenurus problecornis is a species of water mite (family Arrenuridae) described by Cook in 1976. Like other members of the genus Arrenurus, it is part of the subgenus Arrenurus, which is characterized by parasitic larvae that attach to odonate hosts (dragonflies and damselflies). The species belongs to the diverse radiation of water mites (Hydrachnidia), a group comprising over 6,000 species of predatory freshwater arachnids with complex life histories involving parasitic larval stages and free-living predatory adults.
Blissus occiduus
Western Chinch Bug
Blissus occiduus, the western chinch bug, is a phloem-feeding true bug (Hemiptera: Blissidae) that is a significant pest of warm-season turfgrasses, particularly buffalograss and zoysiagrass. The species exhibits strong host preference hierarchies, with buffalograss being the most preferred host followed by zoysiagrass, though it can survive and reproduce on a broad range of grasses including agronomic crops. Field studies have documented inconsistent control with neonicotinoid insecticides, with thiamethoxam showing particularly rapid degradation in buffalograss tissues compared to imidacloprid and clothianidin.
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ere-spherecollective-sphere-sphere-spherecommunity-sphere-sphere-spheresocietal-sphere-sphere-sphereglobal-sphere-sphere-spherelocal-sphere-sphere-spherenational-sphere-sphere-sphereregional-sphere-sphere-spherelandscape-sphere-sphere-sphereseascape-sphere-sphere-spherewaterscape-sphere-sphere-sphereairscape-sphere-sphere-spherebiosphere-sphere-sphere-spherenoosphere-sphere-sphere-spheretechnosphere-sphere-sphere-spheresociosphere-sphere-sphere-sphereecosphere-sphere-sphere-spherehydrosphere-sphere-sphere-sphereatmosphere-sphere-sphere-spherelithosphere-sphere-sphere-spherepedosphere-sphere-sphere-spherecryosphere-sphere-sphere-sphereanthroposphere-sphere-sphere-sphereCephenemyia apicata
nose bot fly
Cephenemyia apicata is a nose bot fly (family Oestridae) that parasitizes Columbian black-tailed deer (Odocoileus hemionus columbianus). First instar larvae develop in the deer's bronchi for 6–8 months before migrating to retropharyngeal pouches. Adult males aggregate at elevated stands of Castanopsis and Quercus in California, with activity occurring from April through July when air temperatures reach 20°C. Females exhibit a distinctive visible attack behavior, landing on or near deer and eliciting curiosity before larvipositing into nostrils.
Cephenemyia jellisoni
nose bot fly
A parasitic bot fly whose larvae develop in the nasal passages and pharyngeal recesses of deer. Adults are active during warm periods and exhibit specific mating aggregations on elevated vegetation. The species shows a covert attacking strategy when larvipositing, remaining undetected by hosts until the moment of larval deposition.
Cephenemyia phobifer
pharyngeal bot of white-tailed deer
Cephenemyia phobifer is a parasitic bot fly (family Oestridae) that develops in the pharyngeal region of white-tailed deer (Odocoileus virginianus). The species exhibits probable larviposition, with females depositing live larvae directly onto or near hosts rather than laying eggs. It has a northeastern Nearctic distribution and completes two generations annually, with winter and summer cycles differing in duration.
Cephenemyia pratti
nose bot fly
Cephenemyia pratti is a species of nose bot fly in the family Oestridae, described by Hunter in 1916. Like other members of the genus Cephenemyia, it is an obligate parasite of cervids (deer family). The species is poorly documented in scientific literature, with only six observations recorded on iNaturalist as of the knowledge cutoff.
Chelicerata
Chelicerates
Chelicerata is a major subphylum of arthropods characterized by the presence of chelicerae—pincer-like or fang-like appendages that are the first pair of structures before the mouth. The group includes horseshoe crabs, sea spiders, and arachnids (spiders, scorpions, mites, ticks, harvestmen, and allies). Chelicerates diverged from other arthropods by the mid-Cambrian period, approximately 508 million years ago. With over 77,000 described living species and estimates suggesting hundreds of thousands more undescribed species, chelicerates represent one of the most diverse animal groups after insects. Most living species are terrestrial air-breathers, though marine lineages persist.
Chrysomelobia
Chrysomelobia is a genus of sexually transmitted parasitic mites in the family Podapolipidae that infest chrysomelid beetles. The genus was originally described from the Americas, Europe and Africa, and was later expanded to include Australian species following the synonymization of Parobia. These mites complete their entire life cycle on adult beetles and do not infest immature host stages.
Chrysurissa densa
Chrysurissa densa is a species of cuckoo wasp in the family Chrysididae. It is known as a parasite of the nests of pollen wasps in the genus Pseudomasaris, specifically targeting the mud nests of Pseudomasaris vespoides. The species was first described by Cresson in 1865. Like other cuckoo wasps, females lay their eggs in the nests of host wasps, where their larvae consume the pollen and nectar provisions intended for the host's offspring.
Cimex
bed bugs, bedbugs
Cimex is a genus of obligate hematophagous insects in the family Cimicidae, commonly known as bed bugs. The genus includes species specialized on different hosts, with C. lectularius (common bed bug) feeding primarily on humans and C. hemipterus (tropical bed bug) occurring in warmer regions. Other species such as C. pipistrelli, C. pilosellus, and C. adjunctus specialize on bats. Bed bugs are wingless, nocturnal parasites that have undergone global resurgence since the late 1990s, likely due to insecticide resistance, changes in pest control practices, and increased international travel.
Cochliomyia
New World screwworm flies, screwworm flies
Cochliomyia is a genus of blowflies (family Calliphoridae) comprising four species: C. macellaria, C. hominivorax, C. aldrichi, and C. minima. The genus is commonly known as New World screwworm flies, distinguished from Old World screwworms. C. hominivorax, the primary screwworm, is an obligate parasite whose larvae feed on living tissue, causing severe myiasis in livestock, wildlife, and humans. C. macellaria, the secondary screwworm, feeds only on necrotic tissue and carrion. The primary screwworm has been eradicated from North America through the sterile insect technique, though it persists in parts of Central and South America.
Coptotermes
Coptotermes is a genus of subterranean termites in the family Rhinotermitidae, containing approximately 71 described species. The genus is thought to have originated in Southeast Asia and includes some of the world's most economically destructive termite pests, notably Coptotermes formosanus (Formosan subterranean termite) and C. gestroi (Asian subterranean termite). A 2013 consensus among 31 termite experts reduced the number of validated species from 69 to 21 based on morphological and molecular evidence, with many historical names suspected to be synonyms. Species in this genus exhibit complex social behaviors including age-based task division and unique nitrogen recycling strategies.
termitepestinvasivestructural-pestsubterraneanRhinotermitidaeHeterotermitidaeeconomic-pesturban-entomologycolonysocial-insectmoltingnitrogen-recyclingexuviaeage-polyethismFormosan-subterranean-termiteAsian-subterranean-termiteSoutheast-Asia-originwood-feedingcellulosegut-protozoalower-termiteevolutionary-transitiontunnelingforagingtrophallaxisproctodeal-trophallaxiscarton-nestAhamitermesparasitechitin-synthesis-inhibitorCSI-baitmolting-site-fidelityinvasive-speciesurban-peststructural-damagequarantineshippingrailroad-tiesmulchHawaiiFloridaLouisianaTaiwanJapanChinaPhilippinesAustraliatemperature-limitationhumidity-requirementcolony-sizeseveral-millionalatesswarmingnematode-associationvertical-transfermorphological-plasticitymolecular-identificationmitochondrial-DNACOII12S-rRNA16S-rRNAconsortium-taxonomytaxonomic-cold-casesynonymyWasmann-1896Cremastocheilus constricticollis
Cremastocheilus constricticollis is a species of anteater scarab beetle native to the southwestern United States. Like other members of its genus, it is a myrmecophile that lives within ant colonies as a predator of ant larvae and pupae. The species exhibits specialized morphological adaptations including a dense, heavily armored exoskeleton and modified mouthparts for piercing soft ant cuticle. Adults are known to employ death-feigning behavior to gain entry to ant nests. The species was described by Cazier in 1940 and is among roughly 35 recognized North American species in this genus, which is currently undergoing taxonomic revision.
Cremastocheilus crinitus
anteater scarab beetle
Cremastocheilus crinitus is a myrmecophilous scarab beetle known for its specialized parasitic relationship with ants. Adults enter ant colonies by feigning death, allowing scavenging workers to carry them into nests. Once inside, they feed on ant larvae and pupae using modified piercing mouthparts. The species is part of a genus of approximately 35 North American species, commonly called "anteater scarabs" for their predatory behavior toward ant brood.
Cremastocheilus harrisii
Eastern Ant-loving Scarab
Cremastocheilus harrisii is a species of anteater scarab beetle in the family Scarabaeidae. As a member of the genus Cremastocheilus, it exhibits the specialized myrmecophilous lifestyle characteristic of this group, living within ant colonies and feeding on ant brood. The species is part of a genus of approximately 35 North American species that have evolved remarkable morphological and behavioral adaptations for surviving in hostile ant nest environments.
Cremastocheilus planipes
Cremastocheilus planipes is a myrmecophilous scarab beetle in the genus Cremastocheilus, commonly known as anteater scarabs. These beetles are specialized inhabitants of ant colonies, where they live as predators of ant brood. The species exhibits remarkable morphological and behavioral adaptations for infiltrating and surviving within hostile ant nests. Like other members of its genus, C. planipes is poorly studied and infrequently encountered due to its cryptic lifestyle.
Cremastocheilus quadratus
anteater scarab beetle
Cremastocheilus quadratus is a myrmecophilous scarab beetle in the family Scarabaeidae, first described by Fall in 1912. Like other members of its genus, it lives within ant colonies as a predator of ant larvae and pupae. The species is found in the southwestern United States and northwestern Mexico.
Cremastocheilus schaumii
anteater scarab beetle
Cremastocheilus schaumii is a North American scarab beetle notable for its obligate myrmecophilous lifestyle. Adults live within ant colonies as predators of ant larvae and pupae, employing specialized morphological adaptations to avoid detection and expulsion by host workers. The species exhibits a complex life cycle involving death-feigning behavior to gain entry to nests, chemical camouflage through acquisition of colony scent, and modified mouthparts for piercing ant brood. Two subspecies are recognized: C. s. schaumii and C. s. tibialis.
Cremastocheilus squamulosus
Cremastocheilus squamulosus is a species of anteater scarab beetle in the family Scarabaeidae. Like other members of its genus, it is a myrmecophile that lives within ant colonies. The species occurs in the southeastern United States, where it has been recorded from Florida, Georgia, North Carolina, and South Carolina.
Cuterebra
rodent bots, Glire Bot Flies
Cuterebra is a genus of New World bot flies (subfamily Cuterebrinae) comprising approximately 78 species that parasitize rodents and lagomorphs. Adults are large, robust flies resembling bumble bees, with non-functional mouthparts and short lifespans devoted entirely to reproduction. Females oviposit near host burrow entrances; larvae hatch in response to host body heat, enter through natural orifices or wounds, and develop subcutaneously for approximately one month before exiting to pupate in soil. The genus exhibits notable host specificity, with individual species typically restricted to single host species or closely related groups.
Cuterebra abdominalis
rabbit bot fly
Cuterebra abdominalis is a species of New World skin bot fly in the family Oestridae. It is a parasite of lagomorphs (rabbits and hares), with larvae developing subcutaneously in host tissues. Adults are short-lived, lack functional mouthparts, and do not feed. The species exhibits Batesian mimicry, resembling bumble bees in appearance. It is one of approximately 34 recognized North American species in the genus Cuterebra, with seven species specializing on rabbit hosts.
Cuterebra americana
woodrat bot fly
Cuterebra americana, the woodrat bot fly, is a New World skin bot fly in the family Oestridae. It is a parasitic species whose larvae develop subcutaneously in mammalian hosts. The species has been documented successfully developing in atypical hosts, including the black rat (Rattus rattus) in Florida. Adults are rarely encountered in the field and lack functional mouthparts, living only to reproduce.
Cuterebra arizonae
New World skin bot fly
Cuterebra arizonae is a species of New World skin bot fly in the family Oestridae. It was described by Sabrosky in 1986. Like other members of the genus Cuterebra, it is a parasitic fly whose larvae develop subcutaneously in mammalian hosts. Adults are rarely encountered in the field and lack functional mouthparts.
Cuterebra atrox
Cuterebra atrox is a species of New World skin bot fly in the family Oestridae. Like other members of the genus Cuterebra, it is a parasitic fly whose larvae develop subcutaneously in mammalian hosts. The species was described by Clark in 1848. Adults are rarely encountered in the field and do not feed, lacking functional mouthparts.
Cuterebra bajensis
Cuterebra bajensis is a species of New World skin bot fly in the family Oestridae, described by Sabrosky in 1986. The species belongs to a genus of obligate mammalian parasites whose larvae develop subcutaneously in rodents and lagomorphs. Adults are rarely encountered in the field and do not feed.
Cuterebra buccata
rabbit bot fly
Cuterebra buccata is a New World bot fly in the family Oestridae, commonly known as the rabbit bot fly. Adults are large, bumble bee-sized flies that lack functional mouthparts and do not feed. The species is host-specific to lagomorphs, particularly cottontail rabbits (Sylvilagus floridanus). Larvae develop subcutaneously in warbles, feeding on host fluids rather than tissue. Adults are rarely encountered in the field and have short lifespans focused on reproduction.
bot-flymyiasisparasitelagomorphrabbitOestridaeCuterebrinaebumble-bee-mimicrywarblesubcutaneous-parasitehost-specificnon-feeding-adultpupariumCuterebraSylvilaguscottontail-rabbitNew-WorldNorth-AmericaVirginiaMissouriVermontspring-emergencelate-summer-prevalencerarely-encounteredshort-lived-adultevolutionary-coevolutionbenign-parasitismsoil-pupationlarval-migrationsubcutaneous-developmentDipteraInsectaentomologymedical-entomologyveterinary-entomologywildlife-diseaseparasitologyhost-parasite-interactioncuterebridrabbit-parasitelagomorph-parasitebotwarble-flygadflyOestrus-buccataFabricius-1775Cuterebra-buccata-(Fabricius,-1775)Cuterebra cochisei
Cuterebra cochisei is a species of New World skin bot fly described by Sabrosky in 1986. As a member of the genus Cuterebra, it is an obligate parasite of mammals with larvae that develop subcutaneously in host tissue. The species is one of 34 recognized North American Cuterebra species. Very little specific information has been published about this particular species.
Cuterebra cuniculi
rabbit bot fly
Cuterebra cuniculi is a New World skin bot fly endemic to Georgia and Florida. Its larvae are obligate parasites of the eastern cottontail (Sylvilagus floridanus) and marsh rabbit (Sylvilagus palustris), developing subcutaneously in warbles. Adults are short-lived, lack functional mouthparts, and do not feed.
Cuterebra emasculator
Squirrel Bot Fly
Cuterebra emasculator is a New World bot fly and obligate internal parasite of tree squirrels and chipmunks in eastern North America. First described by Asa Fitch in 1856, its larvae develop subcutaneously in hosts, creating characteristic swellings called warbles. The species name originated from a false belief that larvae consumed host testicles; research has shown this is not the case and parasitism does not reduce host fertility. Adults are rarely encountered, lack functional mouthparts, and live only a few days focused solely on reproduction.
Cuterebra fontinella fontinella
mouse bot fly, white-footed mouse bot fly
Cuterebra fontinella fontinella is a subspecies of New World bot fly (family Oestridae) that parasitizes small mammals, primarily white-footed mice (Peromyscus leucopus). The adult flies are large, bumble bee-sized, and lack functional mouthparts, living only briefly to reproduce. Females lay eggs near host burrow entrances; larvae hatch in response to host body heat, enter through orifices or wounds, and develop subcutaneously for approximately one month before exiting to pupate in soil. Despite forming conspicuous warbles that can reach 5% of host body weight, infections appear to have minimal negative impact on host survival and may even correlate with increased longevity in some populations.
Cuterebra lepusculi
cottontail rabbit botfly
Cuterebra lepusculi is a New World skin bot fly (family Oestridae) that parasitizes cottontail rabbits (Sylvilagus spp.). Adults are short-lived, lack functional mouthparts, and do not feed. Females deposit eggs near rabbit burrow entrances; larvae hatch upon detecting host body heat, enter through natural orifices or wounds, and develop in subcutaneous warbles for approximately one month before exiting to pupate in soil.
Cuterebra mirabilis
Cuterebra mirabilis is a rare species of New World skin bot fly and the largest member of the rabbit-infesting group within the genus Cuterebra. As of 2009, the species was known from only three specimens, all collected in New Mexico. Like other Cuterebra species, it is an obligate parasite of lagomorphs, with larvae developing subcutaneously in host tissue. The specific epithet "mirabilis" (Latin for "extraordinary") reflects the species' remarkable size and striking appearance among bot flies.
Cuterebra neomexicana
Cuterebra neomexicana is a species of New World skin bot fly described by Sabrosky in 1986. Like other members of the genus Cuterebra, it is a mammalian parasite with larvae that develop subcutaneously in hosts. The species is poorly documented in the scientific literature, with minimal published information on its biology, host associations, and distribution beyond the original description.
Cuterebra polita
Cuterebra polita is a New World skin bot fly whose larvae are obligate parasites of rodents. Adults are non-feeding and short-lived, lacking functional mouthparts. Larvae develop in subcutaneous warbles on host rodents, feeding on host tissues and fluids. The species has been documented in four rodent hosts in Florida: white-footed mouse, cotton mouse, eastern woodrat, and cotton rat.
Cuterebra tenebriformis
Cyladini
Cyladini is a tribe of weevils in the family Brentidae, subfamily Apioninae. The tribe includes economically significant pests of sweet potato, particularly the sweetpotato weevil (Cylas formicarius), which causes substantial damage to storage roots in tropical and subtropical production regions. Members of this tribe are characterized by their association with Convolvulaceae host plants and their status as specialized root-feeding pests.
pestagriculturesweet-potatoConvolvulaceaeintegrated-pest-managementCylas-formicariustropical-agriculturesubtropical-agricultureroot-feeding-weevilstorage-root-damageeconomic-pestbiological-controlcrop-protectionsoutheastern-United-Statesglobal-food-securitylow-input-farmingdrought-tolerant-cropnematode-interactionsBeauveria-bassianaentomopathogenic-nematodesresistant-varietiescrop-rotationcover-cropsIPMBrentidaeApioninaeColeopteraInsectaHexapodaArthropodaAnimaliaClaire-SchloemerAuburn-UniversityUniversity-of-Wisconsin-MadisonJournal-of-Integrated-Pest-ManagementScott-GrahamKathy-LawrenceCovingtonBonitaBeauregardOrleansguava-root-knot-nematoderoot-knot-nematodewirewormcucumber-beetleflea-beetlePurpureocillium-lilacinumbiopesticidegreen-bridge-effectmustardglucosinolatepeanutbahiagrasssandy-soildrought-adaptationbiofuelanimal-feednutritional-valuefood-securitysustainable-agricultureenvironmental-impactmicrobial-controlfungal-pathogenentomopathogenic-fungusbiological-nematicidenematode-egg-parasitismpest-managementagricultural-entomologycrop-lossyield-reductionroot-qualitymarketabilityintegrated-managementcultural-controlselective-chemical-useproductivitynatural-processesversatilityglobal-cultivationtropical-regionssubtropical-regionssoutheastern-U.S.domestic-productionrevenueacreageharvestfarminggrowerbreedingresearchcultivar-developmentevolving-threatspest-pressuresoil-healthfield-preparationwinter-habitatinsect-habitatparasitisminfestationdamagelosschallengedefensearsenalbattlewarresilienceadaptationinnovationstrategysustainabilitylivelihoodnourishmentfood-productionagricultural-researchentomologyplant-protectioncrop-sciencehorticultureroot-croptuberstorage-organfeeding-injurygallingcracked-rootsstunted-growthwater-uptakenutrient-uptakepathogen-interactionexacerbated-damagevariety-selectionhardy-varietyrobust-performancehigh-yieldnematode-resistancepest-resistancevulnerabilityspreading-pestpopulation-reductionsoil-improvementspring-plantingtoxic-compoundnatural-propertymicrobial-productformulationapplicationevaluationmaximizationpotentialurgencygrowing-threatcomprehensive-strategybalanceenablementessential-cropsecuritydefiancedifficult-conditionsmenacing-pestsformidable-threatsproduction-continuationface-to-facethreatdiseaseresilient-cropgrowing-conditionsmenacestorysurfacebeneathrelentlessadversaryfavorconditionpowerfulfoetinyparasiteattackformationdiminishabilityresultunattractiveworsematterinteractionsignificantdevastatingruininflictfurthercompromisegrapplethankfullyfarmerbeginselecthardywithstandtraditionalgaintractionvulnerablehighlightneedcontinuebreedeffortdevelopcapableopposeevolveaftersturdyculturalmanagementrotationcovercroppinglinetakeyearoffgrowrotatenon-hostproveeffectivereducepopulationbreathelifeimprovehealthsuppressprepareexamplecontaincompoundcallbreakdownreleasesubstancewaryproviderepresentmethodharnesshelpwinsporefungusinfectkilltypetapdifferentmicroscopicanimalparasitizeplususecombatattentioneggaidfightminimizeimpactdespiteadvancequestionremainrequiremaximizeparticularlyurgentposekeyenhanceapproachintegratepracticecontrolselectivechemicalenablenaturalprocessnutritionalvalueessentialcropglobalfoodregionsupportmillionaddresspressureinnovativesustainableensurelivemanyformidableproductionalthoughcomefaceresilientabledefydifficultreadmorejournalintegratedClaireSchloemerPh.D.studentconductearnmasterdegreeemailsharelinkfriendopennewwindowprintFacebookLinkedInBlueskyMastodonXRedditThreadsrelateddiscoverfromtodaysubscribegetlatestpostsendyourCylasformicariussweetpotatosweetpotatoweevilphotocourtesyshowherecucumberbeetleroot-knotnematodeharmU.S.guidedetailvarietyoptionbeloveddelicioustastebenefitbrightcolorrankseventhmostimportantworldtropicalsubtropicalareaaroundservefeedsourcegoodpicklow-inputstylewelladaptedsandysoildroughtUnitedStates2022over130,000acregeneratenearly600southeasternleaddomesticAuburnUniversityweatherarticlepublishMarchcolleagueScottGrahamKathyLawrencevariousamongplant-parasiticspecificallyrootsystemcausegallplantwaternutrientstuntedgrowthyieldcrackmakeinteractpathogenexacerbateinsectknowstorage7annualsouthernfleaqualityleaveeconomicownresistancebettermeanwhilenewerhighrobustperformancehoweverresistguavaspeciesspreadnextseasonwinterfieldspringtoxicgreenbridgeeffecthabitatfeedersymptominfectionbiologicalpropertymicrobeBeauveriabassianaagentformulateenvironmentallyfriendlyproducteasyapplysimilarlybeneficialentomopathogenicnematicidePurpureocilliumlilacinumsolutionenvironmentalresistantnematode-resistantcultivarcomprehensivecultivationWisconsin-MadisoncschloemerwisceduCyphomyrmex
fungus-growing ants
Cyphomyrmex is a genus of small, drab-colored fungus-growing ants in the tribe Attini, found primarily in the Neotropics. These ants cultivate fungi in the tribe Leucocoprineae as their primary food source, with most species growing fungal nodules rather than full mycelial gardens. Colonies are typically monogynous and small, rarely exceeding 500 workers. The genus is divided into two species complexes: the strigatus complex (South America only) and the rimosus complex (southern North America to South America). Cyphomyrmex represents a basal lineage among attine ants and serves as sister group to Mycetophylax.
fungus-growing-antsattine-antsMyrmicinaeNeotropicalmutualismfungicultureLeucocoprineaemonogynyparasitoid-waspssocial-parasitismchemical-ecologyalkaloid-venomnest-architecturebehavioral-plasticitycytogeneticschromosome-evolutionGuiana-ShieldBrazilPanamaPuerto-RicoMesoamericaArgentinaUnited-StatesTexasCaliforniaFloridayeast-gardensmycelial-gardensrimosus-complexstrigatus-complexage-polyethismtrophallaxisthanatosisfeigning-deathDiapriidaeAcanthopriaMimopriellaMegalomyrmexsocial-parasiteC.-longiscapusC.-transversusC.-laevigatusC.-rimosusC.-minutusC.-lectusC.-cornutusC.-costatuspheromones3-octanolfarnesenestrail-followingalarm-signaldiketopiperazinesantifungal-compoundsdefensive-behaviorcolony-defenseflight-responserecruitmentLanchester's-lawskaryotypechromosome-numberrDNAmicrosatellitesFISHchamber-depthgallery-lengthfire-disturbancesavanna-forest-transitionarboreal-nestingclay-auriclessoil-nestswood-nestsepiphytic-nestsmoss-nestscoconut-nestsdesiccation-susceptibilitymoist-habitatsbasal-attinephylogeneticssister-groupchemical-evolutionvenom-alkaloidsbehavioral-manipulationhost-parasite-interactionecosystem-engineeringnutrient-cyclingdecompositionorganic-matterinsect-fecesexoskeleton-substrategarden-transplantationworker-polymorphismqueen-numberdealate-queensalatesbrood-careecdysis-assistanceforagingsugar-collectionplant-exudatesfungal-domesticationnodule-cultivationmycelium-cultivationagricultural-behaviornest-entrancefunnel-structureexposed-gardenssymbiont-diversitygeographic-variationresearch-modelevolutionary-biologymutualism-evolutionant-fungus-coevolutionchemical-communicationsocial-insectseusocialitycolony-sizesmall-coloniesslow-movementcrypsisdebris-mimicrypredation-avoidanceparasitoid-defensewasp-killingcolony-abandonmentresource-rescuethief-ant-defensevenom-resistancetoxicity-tolerancebehavioral-suppressionplaying-deadalkaloid-manipulationfungal-secondary-metabolitesantibacterial-compoundsantiviral-compoundsgarden-protectionmicrobial-defensenest-microclimatetemperature-regulationhumidity-requirementsdepth-effectsfire-adaptationdisturbance-ecologyhabitat-specificitysubstrate-specializationclay-constructionsoil-manipulationarchitectural-plasticityvertical-nestingembankment-nestingoverhang-nestingpendant-nestsswallow-nest-mimicryresearch-accessibilityfield-biologytropical-ecologyNeotropical-myrmecologybiodiversityconservationinvasive-species-contextnative-speciesbiological-controlIntegrated-Pest-Managementagricultural-ecologyforest-ecologysavanna-ecologytransitional-habitatsdisturbed-areasurban-ecologyanthropogenic-impactclimate-sensitivityhabitat-limitationrange-boundariesdispersalcolonizationestablishmentpopulation-dynamicsdemographycolony-densitynest-spacingintraspecific-competitioninterspecific-competitionterritorialityresource-defensecombat-behavioraggression-levelsrecruitment-behaviorchemical-signalingalarm-pheromonestrail-pheromonesrecognition-cuescuticular-hydrocarbonsvenom-compositiondefensive-secretionsmandibular-glandsDufour's-glandmetapleural-glandsantibiotic-productiongrooming-behaviorfungal-inoculationintegumental-fungiyeast-growthlarval-groominggarden-maintenancesubstrate-preparationfecal-fertilizeranal-trophallaxiscrop-regurgitationfluid-applicationdrying-behaviornodule-formationtransplantation-behaviorgarden-establishmentceiling-gardensroot-associationarchitectural-innovationnest-site-selectionmicrohabitat-choicemoisture-requirementsthermal-bufferingpredator-avoidanceparasite-avoidancedisease-resistancesocial-immunitycollective-behaviordivision-of-labortask-allocationtemporal-polyethismworker-agenurse-workersforager-workerssoldier-caste-absencemonomorphismworker-size-variationqueen-worker-differentiationreproductive-castegyne-productionmale-productionseasonal-reproductioncolony-foundingclaustral-foundingnon-claustral-foundingfungal-inoculumfounding-queenfirst-worker-generationnutritional-dependencyfat-reservesmetabolic-adaptationdigestive-physiologyenzyme-complementfungal-enzymelignocellulose-degradationnutritional-mutualismobligate-mutualismvertical-transmissionhorizontal-transmissionsymbiont-switchingcultivar-replacementgarden-failurecolony-mortalitypopulation-turnovergeneration-timelongevitysurvivorshiplife-expectancyagingsenescencereproductive-investmentcolony-growthcolony-declinebuddingfissionreproduction-modegenetic-structurerelatednessmating-systemnuptial-flightmate-choicesperm-storagecolony-odornestmate-recognitionforeign-rejectionagonistic-behaviorterritorial-markingboundary-defenseraiding-behaviorrobbing-behaviorcleptobiosisdulosisslaveryinquilinismguest-antsmyrmecophilesnest-associatescommensalisminquilinesparasite-tolerancehost-manipulationvenom-alkaloidpyrrolidineindolizidinepiperidinenonanalfarnesenesesquiterpeneagriculture-chemicalgaster-extractchemical-diversitychemical-simplicityevolutionary-precursorderived-chemistrybiosynthetic-pathwaysemiochemicalinfochemicalcommunicationinformation-transfercolony-organizationsocial-structurepolygynyqueen-conditiondealationwing-sheddin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Dog Ticks, American Levi tick
Dermacentor is a genus of hard ticks (family Ixodidae) with 43 recognized species as of 2025. The genus has a cosmopolitan distribution, occurring on all continents except Australia, with most species in the Nearctic ecozone. Members are obligate blood-feeding parasites of mammals, including large ungulates, lagomorphs, and humans. Several species are medically significant vectors of human and animal pathogens, including the agents of Rocky Mountain spotted fever, Q fever, tularemia, and tick paralysis.
Dermacentor albipictus
winter tick, moose tick
Dermacentor albipictus, commonly known as the winter tick or moose tick, is a one-host hard tick native to North America. It is primarily known as a serious pest of moose (Alces alces), with heavy infestations of up to 150,000 ticks documented on individual animals, often leading to death. The tick also parasitizes other cervids including elk, white-tailed deer, mule deer, and caribou, as well as cattle and horses. Unlike many tick species, D. albipictus is not known to transmit disease pathogens to humans, wildlife, or domestic animals. Its life cycle spans approximately one year, with all feeding stages occurring on a single host.
Elthusa californica
Elthusa californica is a marine parasitic isopod in the family Cymothoidae. It attaches to the gill chambers of fish hosts and feeds on blood and tissue. The species occurs along the Pacific coast from California to Peru. Originally described as Lironeca californica, it was later transferred to Elthusa following taxonomic revision.
Epicaridea
crustacean isopods
Epicaridea is a suborder of parasitic marine isopods now classified within Cymothoida. Members are obligate ectoparasites of other crustaceans, including ostracods, copepods, barnacles, and malacostracans. The group exhibits extreme sexual dimorphism, with females typically becoming asymmetrical and losing segmentation while males remain small and morphologically distinct. Development proceeds through regressive metamorphosis involving two or three larval stages, including the distinctive cryptoniscus stage. Approximately 704 species have been described globally, though the group remains less studied than other isopod lineages.
Epidermoptidae
Epidermoptidae is a family of acariform mites in the order Sarcoptiformes. Members are permanent ectoparasites of birds and mammals, inhabiting skin surfaces and subcutaneous tissues. Several species are nostril specialists. Some genera have evolved phoretic associations with louse flies (Hippoboscidae), and certain species hyperparasitise these carriers, using them for oviposition and feeding. The family shows broad geographic distribution with records from Europe, North America, Australia, and the Galápagos Islands.
Ericrocis lata
Common Oil-digger-Cuckoo
Ericrocis lata is a cleptoparasitic bee species in the family Apidae, commonly known as the Common Oil-digger-Cuckoo. As a member of the tribe Ericrocidini, it is a specialized brood parasite that targets oil-collecting bees. The species occurs in both Central and North America. It was first described by Cresson in 1879.
Eupathocera erynnidis
Eupathocera erynnidis is a species of twisted-wing parasite in the family Xenidae. It is an obligate endoparasite of insects, with females remaining permanently within host abdomens while males develop to a free-flying adult stage. The species has been documented in North America, including Florida. Like other strepsipterans, it exhibits extreme sexual dimorphism and complex life history adaptations associated with parasitism.
Eupathocera lugubris
Eupathocera lugubris is a species of twisted-wing parasite (order Strepsiptera) in the family Xenidae. The species was originally described by Pierce in 1908 and is currently treated as a synonym of Paraxenos lugubris. Strepsipterans are obligate endoparasites of other insects, with this species likely parasitizing wasps or related hosts. The genus is part of the diverse and complex parasitic wasp-associated Strepsiptera fauna of North America.
Eutrombidium
Eutrombidium is a genus of parasitic mites in the family Microtrombidiidae. Larval stages are ectoparasites primarily of grasshoppers and related orthopterans, with some records from beetles and crickets. The genus has been documented across Australia, Asia (Laos, Indonesia, Sri Lanka, Iran, Pakistan), and Europe. Attachment sites on hosts include leg insertions and the prosternum.
Gasterophilinae
stomach bot flies
Gasterophilinae is a subfamily of Oestridae containing large, parasitic flies whose larvae develop in the digestive tracts of herbivorous mammals. The group includes three genera with distinct host specializations: Gasterophilus (equids including horses, zebras, and donkeys), Gyrostigma (rhinoceroses), and Cobboldia (elephants). Adults are short-lived, non-feeding, and exhibit specialized oviposition behaviors. Larvae are obligate parasites that complete development in the host alimentary tract, causing veterinary significance. The subfamily has achieved near-cosmopolitan distribution through association with domestic horses.
Gasterophilus intestinalis
Horse Bot Fly, Horse Botfly
A parasitic bot fly found worldwide that infests the gastrointestinal tract of equids. Adults are bumblebee mimics active in summer, lacking functional mouthparts and living only a few weeks. Females cement eggs to horse hairs; larvae develop in the mouth and stomach for 8–10 months before exiting in feces to pupate. Infestations rarely cause clinical disease but adults are a nuisance to horses.
Haematosiphoninae
Haematosiphoninae is a subfamily of Cimicidae (bed bugs) established in 1912. The group comprises ectoparasitic insects that inhabit bird nests, with documented associations to Psittacidae (parrots) and other avian hosts. Five genera are recognized: Cimexopsis, Ornithocoris, Haematosiphon, Hesperocimex, and Psitticimex, plus the recently described Cyanolicimex. Members are restricted to the Western Hemisphere, with the southernmost records from Río Negro province, Argentina.
Hesperoctenes
Hesperoctenes is a genus of parasitic bugs in the family Polyctenidae, first described by Kirkaldy in 1906. Members of this genus are ectoparasites of bats (Chiroptera). The genus is distributed in Colombia and likely broader Neotropical regions, though published records are limited. As polyctenids, they represent a specialized lineage of hemipteran parasites adapted to mammalian hosts.
Heterocheilidae
half-bridge flies
Heterocheilidae is a family of parasitic nematodes (phylum Nematoda) within the order Rhabditida and infraorder Ascaridomorpha. Members are intestinal parasites primarily infecting fish hosts, including elasmobranchs and teleosts. The family includes genera such as Brevimulticaecum, Ortleppascaris, and Dujardinascaris. Notably, the name 'Heterocheilidae' has also been applied to a family of dipteran insects (true flies), but this usage is taxonomically distinct and not accepted in current nematode classification.
Hydrachna
Hydrachna is a genus of water mites comprising over 80 described species and representing the sole genus in the family Hydrachnidae. These mites are obligate aquatic arachnids found in freshwater habitats. The larvae are ectoparasites of aquatic insects, particularly water beetles and water boatmen, attaching to the thorax and abdomen of hosts. Some host species have developed apparent immunity to parasitism after prolonged exposure.
Hyperia
Hyperia is a genus of amphipod crustaceans in the family Hyperiidae, established by Latreille in 1823. Members of this genus are known for their parasitic or commensal associations with gelatinous zooplankton, particularly jellyfish (Cnidaria). The genus includes species such as Hyperia medusarum, which has been documented as a parasite of scyphozoan and hydrozoan medusae in the southeastern Atlantic Ocean.
Hyperiidae
Hyperiidae is a family of amphipod crustaceans established by Dana in 1852. Members of this family are exclusively marine and are characterized by their obligate associations with gelatinous zooplankton, including jellyfish (Scyphozoa) and ctenophores. The family contains eight recognized genera: Euthemisto, Hyperia, Hyperiella, Hyperoche, Laxohyperia, Parathemisto, Pegohyperia, and Themisto. These amphipods are frequently documented as parasites or associates of their gelatinous hosts in pelagic marine environments.
Hypoderma
Warble Flies, Heel Flies, Bomb Flies, Gadflies
Hypoderma is a genus of large parasitic flies in the family Oestridae, commonly known as warble flies. Adults are non-feeding and live only to reproduce. Females deposit eggs on the legs and flanks of cattle and other ungulates; first-instar larvae penetrate the skin and migrate through the host's body, typically to the gullet or spinal canal, before settling in subcutaneous tissues of the back to develop. The larvae create characteristic swellings (warbles) visible on the host's hide. Two economically important species are H. bovis (ox warble fly) and H. lineatum (cattle warble fly), which cause significant damage to cattle hides and reduce animal welfare.
Hypoderma tarandi
Reindeer Warble Fly, Reindeer Botfly, Caribou Bot Fly
Hypoderma tarandi is a parasitic warble fly (Diptera: Oestridae) that infests reindeer and caribou across Arctic and subarctic regions. Adult females lay eggs on host hair shafts; larvae penetrate the skin and develop in subcutaneous tissues, creating warble lesions. The species has exceptionally strong flight capacity in females, with maximum lifetime flight distances estimated at 600–900 km, an adaptation linked to the migratory behavior of its host. Adults do not feed; they rely entirely on fat reserves accumulated during larval development. The species causes economic damage to hides, meat, and milk production in domesticated herds and has been documented causing ophthalmomyiasis in humans.
Hypodermatinae
warble flies
Hypodermatinae is a subfamily of bot flies (Oestridae) containing nine genera of large parasitic flies commonly called warble flies. The larvae develop as subcutaneous parasites in mammals, particularly bovines, causing economically significant damage to livestock. Adult flies are non-feeding and short-lived.
Isopoda
isopods, woodlice, pillbugs, sowbugs, sea slaters, gribbles
Isopoda is an ancient order of crustaceans encompassing over 10,000 described species across marine, freshwater, and terrestrial habitats. Members are characterized by dorsoventrally flattened bodies, seven pairs of similar walking legs (giving the group its name from Greek iso- "equal" and pod- "foot"), and two pairs of antennae. The order exhibits exceptional morphological diversity, ranging from minute interstitial forms to giant deep-sea species exceeding 30 cm in length. Isopods lack a carapace, instead possessing overlapping dorsal plates that provide flexibility and protection. Females brood eggs in a specialized marsupium formed by oostegites under the thorax.
Ixodes
hard-bodied ticks, blacklegged ticks, deer ticks
Ixodes is a genus of hard-bodied ticks (family Ixodidae) comprising approximately 274 species with a near-cosmopolitan distribution. It is the sole representative of the Prostriata clade, distinguished from Metastriata by an anal groove that loops anterior to the anus. Many species serve as important disease vectors, transmitting pathogens including Borrelia burgdorferi (Lyme disease), Babesia microti (babesiosis), and Anaplasma phagocytophilum (anaplasmosis). Some species, notably Ixodes holocyclus, inject neurotoxins that can cause tick paralysis.
Ixodes affinis
Ixodes affinis is a hard-bodied tick species distributed across the southeastern United States, Mexico, Central America, and South America. Unlike its congener Ixodes scapularis, this species does not bite humans but serves as a competent vector for Borrelia burgdorferi sensu stricto, the causative agent of Lyme disease. Immature stages feed on birds and small mammals, while adults parasitize larger mammals including deer and dogs. Its expanding range into southeastern Virginia and association with migratory birds has raised concerns about its role in pathogen dispersal.
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dentatus
Ixodes dentatus is a hard tick species in the family Ixodidae, first described by Marx in 1899. The species has been recorded in the southeastern United States, including Tennessee and Missouri, and is associated with small mammal hosts such as the marsh rice rat (Oryzomys palustris). It is one of several Ixodes species studied for host associations and seasonal occurrence patterns, though detailed biological information remains limited in accessible literature.
Ixodes scapularis
deer tick, black-legged tick, bear tick, Eastern Black-legged Tick
Ixodes scapularis is a hard-bodied tick native to the eastern and upper midwestern United States, with expanding populations in Canada. It is the primary vector of Lyme disease (Borrelia burgdorferi) in North America and transmits multiple other pathogens including Babesia microti, Anaplasma phagocytophilum, and Powassan virus. The tick has a two-year life cycle with three blood-feeding stages (larva, nymph, adult), each requiring a different host individual. Climate change is driving northward range expansion and increased human encounters.
Ixodida
ticks
Ixodida is the order comprising ticks, obligate blood-feeding arachnids within the subclass Acari. The order includes approximately 850-1000 described species worldwide, classified into three families: Ixodidae (hard ticks), Argasidae (soft ticks), and Nuttalliellidae (a monotypic family). Ticks are ectoparasites of mammals, birds, and reptiles, with significant medical and veterinary importance as vectors of bacterial, viral, protozoan, and rickettsial pathogens. They possess a unique life cycle involving multiple developmental stages and host associations.
Leptoglossus
leaf-footed bugs
Leptoglossus is a genus of true bugs in the leaf-footed bug family Coreidae, tribe Anisoscelini. Species are characterized by leaflike dilations of the hind tibia, a diagnostic trait of the genus. The genus is distributed throughout the Americas, with some introduced populations in Europe and Asia. Several species are economically significant agricultural pests, notably L. occidentalis, which has become invasive in multiple continents.
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nchanmoss-bugarchaeorrhynchanfulgoromorphancicadomorphanmembracoidtreehopperleafhopperplanthopperpsyllidjumping-plant-lousewhiteflyaleyrodidscale-insectcoccoidmealybugaphidadelgidphylloxeransternorrhynchanthysanopteranthripspsocopteranbarklousebooklousephthirapteranlousesucking-lousechewing-lousemallophagananoplurandermapteranearwigblattodeancockroachtermiteisopteranmantodeanmantidphasmidstick-insectleaf-insectorthopterangrasshopperlocustkatydidcricketmole-cricketpygmy-mole-cricketcamel-cricketcave-cricketwetaensiferancaeliferangryllotalpidmyrmecophilidtettigoniidgryllidacrididpamphagidpneumoridlentulidtristirideumastacidproscopiidtridactylidtetrigidgrouse-locustpygmy-grasshopperplecopteranstoneflyembiopteranwebspinnerzorapteranangel-insectdictyopteranLeptus
Leptus is a genus of large mites in the family Erythraeidae, first described by Pierre André Latreille in 1796. The genus comprises approximately 202 species and is distinguished from the related genus Balaustium by the forward placement of eyes on the body. Leptus species are primarily known as ectoparasites, with larvae feeding on various arthropod hosts including bees, harvestmen, and other insects. The genus has been documented across multiple continents including South America, North America, Africa, and Asia.
Lernaea
anchor worms
Lernaea is a genus of parasitic copepod crustaceans commonly called anchor worms, exclusively parasitic on freshwater fishes. Females burrow into fish flesh and transform into unsegmented, wormlike forms with egg sacs visible externally, while males are free-swimming and short-lived. The genus is widely distributed globally and causes significant disease in aquaculture and wild fish populations. Multiple species exist, with Lernaea cyprinacea being the most studied and economically important.
Lernaea cyprinacea
anchor worm, fish louse
Lernaea cyprinacea is a parasitic copepod commonly known as the anchor worm, a significant pest in freshwater aquaculture worldwide. Native to Eurasia, it has been introduced to multiple continents including North America and South America. The species exhibits sexual dimorphism with females becoming permanently attached to fish hosts while males are free-living and short-lived. It causes substantial economic damage through direct parasitism, reduced fish growth, and secondary infections.
Lernaeidae
anchorworms
Lernaeidae is a family of parasitic copepods in the order Cyclopoida, commonly known as anchorworms. The family contains approximately 131 species across 20 genera, all of which are obligate ectoparasites of freshwater fishes. Species in this family are characterized by elongated bodies, often with anchor-like holdfast structures used for attachment to host tissues. The most widely distributed and economically significant species is Lernaea cyprinacea, which has been introduced globally and causes substantial damage in aquaculture systems.
Lernaeopodidae
Lernaeopodidae is a family of parasitic copepods in the order Siphonostomatoida. Females are typically large and fleshy, attaching permanently to fish hosts using a chitinous plug called the bulla. Males are smaller and cling to females using their antennae. Members parasitize both marine and freshwater fishes, with some species causing significant problems in aquaculture.
Meloe barbarus
oil beetle
Meloe barbarus is a species of blister beetle in the family Meloidae, first described by LeConte in 1861. It belongs to the subgenus Treiodous within the genus Meloe. The species is found in North America, with collection records from western regions including British Columbia, Canada. Like other oil beetles in the genus Meloe, it possesses defensive chemical compounds and exhibits complex life history traits involving parasitism of ground-nesting bees.
Meloe dianella
Meloe dianella is a species of blister beetle (family Meloidae) first described by Pinto and Selander in 1970. It belongs to the genus Meloe, commonly known as oil beetles, which are characterized by their defensive production of cantharidins. The species is known from North America, with records from Canada (Alberta, British Columbia, and Nova Scotia). Like other members of its genus, it is presumed to have a parasitic life cycle involving solitary bees, though specific details for this species remain undocumented.
Meloe dugesi
Meloe dugesi is a species of blister beetle (family Meloidae) described by Champion in 1891. It belongs to the genus Meloe, commonly known as oil beetles, which are characterized by their defensive production of cantharidins. The species occurs in Central America and North America. Like other members of its genus, it likely exhibits the typical Meloe life cycle involving triungulin larvae that parasitize solitary bees.
Nomada cuneata
Cuneate Nomad Bee
Nomada cuneata is a kleptoparasitic bee species in the family Apidae, native to North America. As a member of the genus Nomada, it exhibits the characteristic cuckoo bee behavior of laying eggs in the nests of host bee species rather than constructing its own nests or provisioning young. The species is recorded from Vermont and other parts of North America. Like other Nomada species, it likely targets specific host bees, though detailed host associations for this particular species remain poorly documented.
Oestrinae
Nose Bot Flies
Oestrinae is a subfamily of bot flies (Oestridae) comprising 9 genera and 34 species of obligate mammalian parasites. Larvae develop within the nasal, pharyngeal, and frontal cavities of deer, sheep, and related ungulates, feeding on cellular debris and mucosal secretions. Adult females are larviparous, depositing live larvae directly into host nostrils. The subfamily includes economically significant pests of livestock and game animals.
Oestrus
bot flies, sheep bot flies
Oestrus is a genus of bot flies in the family Oestridae, parasitic on mammals. The best-known species, Oestrus ovis, commonly called the sheep bot fly, deposits larvae in the nasal cavities of sheep and goats, where they develop and cause respiratory distress. Adults are non-feeding and have reduced mouthparts. The genus has a Holarctic distribution, with records from northern Europe and other temperate regions.
Oestrus ovis
sheep bot fly, sheep nostril fly
Oestrus ovis is an obligate parasitic fly (Diptera: Oestridae) that develops exclusively within the nasal and sinus cavities of sheep and related ungulates. Females are larviparous, depositing live first-instar larvae directly into host nostrils during flight. Larvae develop through three instars within the host before exiting to pupate in soil. The species causes oestrosis, a condition marked by nasal discharge, respiratory distress, and reduced productivity in livestock. Human infestations, though uncommon, have been documented worldwide.
Oreoneta
A genus of dwarf spiders (family Linyphiidae) established in 1894, comprising approximately 30 species distributed across cold-temperate and Arctic regions of the Northern Hemisphere. Most species inhabit high-latitude or high-altitude environments. The genus was revised extensively by Saaristo & Marusik in 2004, resulting in numerous new species descriptions.
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cynotis
Ear Mange Mite, Ear Mite, Common Ear Mite
Otodectes cynotis is a parasitic mite in the family Psoroptidae, the sole species in its genus. It is a non-burrowing ectoparasite that inhabits the external ear canals of mammals, causing otitis externa. The mite completes its entire life cycle on a single host in approximately three weeks. It is distributed worldwide and primarily affects domestic cats, dogs, and ferrets, though it has been recorded on numerous other carnivores and occasionally humans. Infestation causes intense pruritus, head shaking, and ear scratching, and can lead to secondary bacterial infections in severe cases.
P. fuscus
Pandionacarus fuscus is a species of mite in the family Avenzoariidae, order Sarcoptiformes. The genus name indicates an association with the osprey genus Pandion, suggesting this mite may be an ectoparasite or commensal of ospreys. Very little published information exists on the biology or ecology of this species. The species was described by Nitzsch in 1818, indicating it has been recognized in scientific literature for over two centuries.
Paranomada velutina
Black-velvet Flat-Nomad
Paranomada velutina is a cuckoo bee in the family Apidae, described by Linsley in 1939. It belongs to the subfamily Nomadinae, a group of bees that are cleptoparasites (nest parasites) of other bees. The species is known from the southwestern United States and Mexico. Like other nomadine bees, it lacks pollen-collecting structures and relies on host bees to provision its young.
Parasitiformes
mites and ticks
Parasitiformes is a superorder of arachnids constituting one of two major mite lineages, alongside Acariformes. The group includes ticks (Ixodida), the highly diverse predatory and parasitic Mesostigmata, the small scavenging Holothyrida, and the large segmented Opilioacarida. Over 12,000 species have been described, with estimates of 100,000–200,000 total species. Recent phylogenomic analyses indicate Parasitiformes is more closely related to other chelicerates (spiders, scorpions, horseshoe crabs) than to Acariformes, challenging the traditional unification of all 'mites' as a single natural group.
Pentodontini
rhinoceros beetles
Pentodontini is the most diverse tribe within the subfamily Dynastinae (rhinoceros beetles), containing over 100 genera distributed across multiple biogeographic regions. Most genera are restricted to a single biogeographic region. The tribe is characterized by substantial morphological diversity, with generic-level identification often relying on mouthpart morphology in females and secondary sexual characters (horns, claw modifications, antennal club length) in males.
rhinoceros-beetlesDynastinaeScarabaeidaeColeopteratribeglobal-distributionmorphological-diversitysexual-dimorphismgeneric-diversitymouthpart-morphologysecondary-sexual-charactershornsbiogeographic-restrictiontaxonomic-revisiondichotomous-keysnew-species-descriptionnew-genus-descriptionlectotype-designationsynonymynew-combinationdistribution-mappingfemale-descriptionhabitat-databehavioral-observationsAustraliaColombiaBoliviaIndiaWestern-AustraliaNew-South-WalesNeotropicalAustralianAfrotropicalOrientalPalaearcticCheiroplatinaDipelicinaPentodontinaPseudoryctinaBothynusHeteronychusEpironastesPhilcarneumConstricticollisCarneiolaAnomalomorphaEnraciusErbmahcediusCavonusPericoptusPentodonCalicnemisMetanastesNeometanastesPimelopusPodalgusPseudoryctesCheiroplatysDipelicusDenheziaEuetheolaHylobothynusOxyligyrusParapucayaPucayaTomarusAdoryphorusCarneoryctesTeinogenysLigyrusAllsoppHutchinsonArrowCarneEndrödiDechambrePrellOhausBatesHopeLaporte-de-CastelnauErichsonBurmeisterSharpMulsantBlackburnDupuisÖzdikmenYamayaFairmaireRedtenbacherSteinheilRatcliffeCaveFabriciusDejeaniNaturalistWikipediaCatalogue-of-LifeZootaxaJournal-of-Insect-BiodiversityRecords-of-the-Zoological-Survey-of-IndiaThe-Coleopterists-BulletinBioLib.czWikimedia-CommonsDOI10.11646/zootaxa.4048.4.110.11646/zootaxa.4852.4.210.11646/zootaxa.5351.3.210.26515/rzsi/v125/i2s/2025/17296410.11646/zootaxa.5716.4.710.11646/zootaxa.5072.5.210.11646/zootaxa.4852.4.310.12976/jib/2024.54.2.210.1649/1186.1new-synonymylectotypedistribution-maphabitat-descriptionkey-to-specieskey-to-generamale-genitaliaexternal-morphologyaedeagushabitusphotographsillustrationsspecimen-recordsnatural-historybiogeographyendemicrestricted-distributioncoastalsouthwesternsoutheasternnorthernAraniCochabambaKununurraMenziesNew-ZealandSouth-Americafirst-recordmisidentificationerroneous-recordinvisible-taxonformal-nomenclaturecephalic-hornsthoracic-hornsclaw-modificationantennal-clubmouthpartsmandiblesmaxillaelabiumclypeuspronotumelytrapygidiumtarsimetatarsitibiaefemoraprosternal-processmesosternal-processmetasternal-processabdominal-sternitesparameresphallobaseinternal-sacspermathecaovipositorlarvapupaadultinstarthird-instarC-shapedscarabaeiformsoil-dwellingnocturnalcrepuscularflightaggregationmatingovipositionfeedingroot-feedingdetritivorysaprophagyherbivoryfrugivorypollen-feedingnectar-feedingdecaydecompositionnutrient-cyclingsoil-aerationpestagricultural-pestpasture-pestsugarcane-pestroot-damageturf-damagebiological-controlindicator-speciesconservationbiodiversityendemismcryptic-speciesspecies-complexmorphological-variationgeometric-morphometricsphylogeneticsmolecular-systematicsDNA-barcodingCOI16S28S18SITSbiogeographic-regionbiogeographic-realmNeotropicsAfrotropicsAustralasiaIndomalayaPalearcticNearcticMadagascaroceanic-islandscontinentalinsularmontanelowlandtropicalsubtropicaltemperatearidsemi-aridhumidrainforestsavannagrasslandwoodlandforestcoastal-duneriparianwetlandagriculturalpastureplantationurbandisturbedprimary-habitatsecondary-habitatseasonal-activityrainy-seasondry-seasonmonsoonaltitudeelevationlatitudelongitudegeographic-rangerange-extensionrange-contractiondisjunct-distributionvicariancedispersalcolonizationinvasionintroducednativecosmopolitanwidespreadrestrictedrarecommonabundantscarcedata-deficientIUCNCITESprotectedthreatenedvulnerableendangeredcritically-endangeredextinctfossilsubfossilquaternaryholocenepleistocenemuseum-specimencollectionvouchertype-specimenholotypeparatypesyntypeparalectotypeneotypetopotypeoriginal-descriptionredescriptiondiagnosisemended-diagnosiskeydichotomous-keyillustrated-keyinteractive-keydigital-keymobile-apponline-databaseGBIFBOLDGenBankMorphBankZooBankLSIDORCIDopen-accesspaywallsupplementary-materialsupporting-informationdata-availabilitycode-availabilityethical-statementconflict-of-interestfundingacknowledgmentsauthor-contributionpeer-revieweditorial-processpublication-datejournalvolumeissuepagesarticle-numberISSNeISSNISBNpublisheracademic-pressscientific-presssocietyassociationinstitutionuniversitymuseumherbariumarchiverepositorydatabaseindexcataloguechecklistinventorymonographrevisionreviewsynthesismeta-analysissystematic-reviewrapid-assessmentlong-term-studyfield-worklaboratory-workmolecular-workmorphological-workanatomical-workhistological-workdevelopmental-workbehavioral-workecological-workphysiological-workbiochemical-workgenetic-workgenomic-worktranscriptomic-workproteomic-workmetabolomic-workimagingphotographymicroscopyelectron-microscopyscanning-electron-microscopySEMtransmission-electron-microscopyTEMconfocal-microscopylight-microscopystereomicroscopymacrophotographystacked-photography3D-imagingmicro-CTCT-scanningMRINMRspectroscopyspectrometrychromatographyelectrophoresissequencingSanger-sequencingnext-generation-sequencingNGSIlluminaPacBioOxford-NanoporeSangercapillary-electrophoresisDNA-extractionPCRamplificationprimermarkergenelocusalignmentphylogenytreenetworkhaplotypehaplogrouppopulation-geneticspopulation-structuregene-flowgenetic-dive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hconnectivity-conservationcorridorbuffer-zoneprotected-areanational-parknature-reservewildlife-refugewilderness-areaworld-heritage-sitebiosphere-reserveRamsar-siteImportant-Bird-AreaKey-Biodiversity-AreaAlliance-for-Zero-Extinction-siteconservation-priorityhotspotcrisis-ecoregionglobal-200last-of-the-wildhuman-footprintcumulative-impactthreat-indexvulnerability-indexadaptive-capacityexposuresensitivityresilienceresistancerecoveryrestorationrehabilitationreintroductiontranslocationex-situin-situcaptive-breedingbotanic-gardenzoogene-bankseed-banktissue-banksperm-bankoocyte-bankembryo-bankDNA-bankfrozen-zooarkinsurancesafety-netde-extinctiongenetic-rescuegenetic-restorationgenetic-augmentationgenetic-managementpopulation-managementmetapopulationsource-sinkpatchmatrixlandscapeseascapeecosystembiomeecoregionprovincezoneregiondistrictsitelocalityhabitatmicrohabitatnicheecological-nichefundamental-nicherealized-nichetrophic-nichespatial-nichetemporal-nichebiotic-nicheabiotic-nichemultidimensional-nichen-dimensional-nicheHutchinsonian-nicheGrinnellian-nicheEltonian-nicheresourcerequirementlimitationstressdisturbanceperturbationfluctuationvariabilityheterogeneitycomplexitydiversityredundancystabilitypersistenceadaptationacclimationplasticityevolvabilityheritabilityselectiondriftflowmutationrecombinationspeciationcoalescencedivergenceconvergenceparallelismhomoplasyanalogyhomologysynapomorphysymplesiomorphyautapomorphyapomorphyplesiomorphyderivedancestralprimitiveadvancedbasalcrownstemnodebranchcladegradesubfamilyfamilysuperfamilyinfraordersuborderordersuperorderinfraclasssubclassclasssuperclasssubphylumphylumsuperphylumkingdomdomainlifeorganismindividualetc.Periplaneta brunnea
brown cockroach
Periplaneta brunnea is a cockroach species in the family Blattidae, commonly known as the brown cockroach. It serves as an intermediate host for the acanthocephalan parasite Moniliformis moniliformis, with behavioral alterations documented in infected individuals. The species has been used in biochemical studies investigating juvenile hormone regulation and insecticide metabolism. Its distribution includes regions in Africa, Australia, and South America including the Galápagos Islands.
Pogonomyrmex tenuispinus
Pogonomyrmex tenuispinus is a species of harvester ant in the genus Pogonomyrmex. Like other members of this genus, it is a seed-collecting ant native to arid regions. The species was described by Auguste Forel in 1914. As a harvester ant, it likely participates in seed dispersal and ecosystem engineering through nest construction, though specific ecological studies for this species are limited.
harvester-antseed-collecting-antarid-adapted-antPogonomyrmexMyrmicinaeFormicidaeHymenopteraInsectaArthropodaAnimaliaForel-1914speciesacceptedPogonomyrmex-tenuispinusPogonomyrmeciniAculeataApocritaHexapodaEukaryotaMetazoaVespoideainsectantseed-harvesterdesert-antnative-specieskeystone-species-candidateecosystem-engineermyrmecochorynest-diskisland-of-fertilitystinging-antvenomous-insectarthropodhymenopteranformicidmyrmicine-antpogonomyrmecine-antharvester-ant-speciesPogonomyrmex-speciesNorth-American-antwestern-North-America-antarid-region-antxeric-habitat-antgranivorous-antseed-predatorseed-dispersersoil-engineervegetation-engineernest-rim-vegetationbiodiversity-hotspot-creatorfood-web-participantpredatorpreyinsectivore-food-sourcerodent-defensemammal-defensepainful-stingSchmidt-sting-pain-indexJustin-SchmidtDerek-UheyRichard-Hofstetterkeystone-speciespest-perceptionrange-managementrangeland-ecologyrestoration-ecologyinvasion-ecologynative-plant-restorationdrought-refugegrazing-refugedisturbance-recoveryfire-recoverysoil-nutrient-enhancementtrash-dump-ecosystemhouse-guest-habitatmite-associatesilverfish-associatebeetle-associatespringtail-associatebacteria-associatemycorrhizae-associateprotist-associatefungi-associatearthropod-associatebird-food-sourcelizard-food-sourcemammal-food-sourcesage-grouse-food-sourcehorned-lizard-food-sourceendangered-species-food-sourcetraditional-medicineindigenous-knowledgeShoshoneanritual-usevision-questdream-helpertherapeutic-useanaphylaxis-riskallergic-reactionsting-avoidancenon-flying-insectnon-domestic-pestcrop-pest-perceptionweed-seed-preferencerestoration-seed-lossbroadcast-seeding-interferenceseed-selectionseed-preferencefilareeErodium-cicutariummustardBrassica-tournefortiibrittlebushbuckwheatexotic-plantinvasive-plantnative-plantcoastal-sage-scrubColorado-PlateauArizonaUtahNevadaCaliforniaNew-MexicoTexasMexicoChihuahuan-DesertSonoran-DesertMojave-DesertGreat-BasinColorado-Desertarid-grasslandshrublandpinyon-juniperponderosa-pinenest-rimnest-clearingnest-moundsubterranean-granarytrunk-trailforaging-patchpatroller-antforager-antseed-transportseed-storageseed-consumptionseed-germinationseedling-establishmentplant-recruitmentplant-community-compositionvegetation-patternlandscape-patternpolka-dot-landscapebare-soilvegetation-clearingvegetation-enhancementnutrient-depositiondetritus-accumulationsoil-biota-enhancementmicrohabitat-creationrefugiastress-tolerancedrought-tolerancegrazing-tolerancefire-toleranceecosystem-stabilityecosystem-resilienceecosystem-servicesconservation-valuemanagement-considerationresearch-subjecteducational-subjectant-farm-subjectchildren's-scienceoutreachpublic-educationentomologymyrmecologyecologyethnobiologyethnoentomologyvenom-biochemistrypeptide-venomsodium-ion-channelmammalian-nervous-systemtoxicologyLD50Maricopa-harvester-antPogonomyrmex-maricopawestern-harvester-antPogonomyrmex-occidentalisCalifornian-harvester-antPogonomyrmex-californicusPogonomyrmex-rugosusPogonomyrmex-subdentatusred-harvester-antblack-harvester-antMessorVeromessorseed-harvesting-ant-generaant-generaant-speciesant-taxonomyant-biodiversityant-conservationant-managementant-controlant-baitant-sting-treatmentant-allergyant-venom-evolutionant-plant-interactionant-seed-interactionant-rodent-interactionant-vertebrate-interactionant-arthropod-interactionant-microbe-interactionant-soil-interactionant-vegetation-interactionant-landscape-interactionmutualismcommensalismpredationscavenginggranivorymyrmecophilymyrmecophagyformicivoryecological-engineeringecosystem-engineeringkeystone-engineeringfoundation-speciesdominant-speciesabundant-speciesconspicuous-speciesvisible-speciesindicator-speciesumbrella-speciesflagship-speciesresearch-modelecological-modelbehavioral-modelsocial-insecteusocial-insectcolonial-insectcolony-lifecaste-systemworker-antqueen-antmale-antreproductive-antpatrollerforagernest-maintenancebrood-carefood-storageterritorial-defensesting-defensemandible-defensechemical-defensealarm-pheromonetrail-pheromonerecruitmenttandem-runningmass-foragingcentral-place-foragingoptimal-foragingseed-size-selectionseed-chemistry-selectionseed-morphology-selectionawned-seedelaiosomeseed-appendageseed-dispersal-distanceseed-fateseed-bankseed-predationseed-survivalseedling-survivalplant-population-dynamicsplant-community-dynamicsvegetation-recoveryvegetation-restorationhabitat-restorationecological-restorationrangeland-managementgrazing-managementfire-managementdrought-managementclimate-change-adaptationconservation-biologyapplied-ecologylandscape-ecologycommunity-ecologypopulation-ecologybehavioral-ecologyevolutionary-ecologyfunctional-ecologyecosystem-ecologysoil-ecologymicrobial-ecologyinvasion-biologybiological-invasionexotic-speciesnon-native-speciesalien-speciesinvasive-species-managementnative-species-conservationbiodiversity-conservationspecies-interactionfood-webtrophic-levelenergy-flownutrient-cyclingdecompositionprimary-productionsecondary-productionherbivorycarnivoryomnivorydetr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dominula
European Paper Wasp
Polistes dominula is a highly successful invasive social wasp native to Eurasia that has established populations across North America, South America, New Zealand, South Africa, and other regions. First detected in North America near Boston in 1978, it has become one of the most abundant wasps on the continent. The species builds small, exposed paper nests in protected locations and preys primarily on live insects, particularly caterpillars. Unlike yellowjackets, it does not scavenge for meat or sugar. Its rapid spread has been attributed to ecological flexibility, superior competitive ability, and tolerance of human-altered environments.
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bird blowflies, bird blow flies
Protocalliphora is a genus of blow flies (Calliphoridae) comprising obligate hematophagous parasites of birds. Larvae develop in bird nests and feed on the blood of nestlings, with more than 140 bird species recorded as hosts. The genus is distributed across the Holarctic region (Nearctic and Palaearctic). Adults overwinter and exhibit complex interactions with Wolbachia endosymbionts that complicate species identification through DNA barcoding.
Protocalliphora occidentalis
Protocalliphora occidentalis is a species of blow fly in the family Calliphoridae, described by Whitworth in 2003. The genus Protocalliphora comprises parasitic flies whose larvae develop in bird nests, feeding on nestling blood. This species is part of a group commonly known as bird nest flies or nestling parasites.
Protocalliphora sialia
birdnest blowfly
Protocalliphora sialia is a parasitic blowfly that infests the nests of cavity-nesting birds. Larvae are nest-dwelling, intermittent ectoparasites that feed on nestling blood. The species has been documented from multiple host species including tree swallows, eastern bluebirds, and ferruginous pygmy-owls. Wolbachia bacterial infections show geographic variation across the species' range.
Psoroptidae
Scab Mite
Psoroptidae is a family of parasitic mites in the order Sarcoptiformes. Members are 1–2 mm in length and inhabit the surface of host skin rather than burrowing into it. The family includes economically important genera such as Psoroptes, Chorioptes, and Otodectes, which cause mange and skin disease in mammals. Host associations span domestic animals (sheep, cattle, horses, dogs, cats, rabbits) and wildlife including African monkeys and various bird species.
Pugettia
kelp crabs
Pugettia is a genus of marine kelp crabs in the family Epialtidae, distributed across the North Pacific from North America to East Asia. Species inhabit shallow subtidal zones, primarily associated with macroalgal habitats including kelp beds, Sargassum stands, and red algal turfs. Many species exhibit ontogenetic habitat shifts, with juveniles and smaller individuals occupying deeper algal turfs while larger adults migrate to shallower macroalgal beds. The genus includes approximately 25 extant species plus one fossil species, with several species serving as important subjects for studies of crab growth, reproduction, and habitat ecology.
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slug mites, snail mites
Riccardoella is a genus of parasitic mites in the family Ereynetidae that are obligate ectoparasites of terrestrial gastropod molluscs, primarily slugs and snails. The genus contains approximately six described species distributed across Europe, North America, East Asia, and New Zealand. These mites inhabit host mantle cavities, respiratory pores, and lung spaces, feeding on host mucus and tissue fluids. Their life cycles are tightly coupled to host biology, with developmental stages periodically leaving hosts to moult before reinfesting.
Salmincola
Salmincola is a genus of freshwater parasitic copepods in the family Lernaeopodidae. These ectoparasites are common on salmonid fishes throughout the Northern Hemisphere. The genus exhibits substantial geographic variation in infection rates across water bodies, and current taxonomic definitions based on decades-old morphological descriptions may obscure cryptic diversity. Species in this genus are considered pests by fisheries management agencies due to their impact on host fish populations.
Salmincola californiensis
gill lice, gill maggot, salmon gill maggot
Salmincola californiensis is a freshwater parasitic copepod in the family Lernaeopodidae that infests the gills and branchial cavities of salmonid fishes, particularly species of Oncorhynchus. Native to the Pacific Rim and now established in introduced ranges including Japan and Colorado, this parasite has become a significant concern for fisheries management and salmonid conservation. Recent research has documented a previously unrecognized naupliar stage and demonstrated strong temperature-dependence in egg development and copepodid survival, with implications for disease management under climate change scenarios.
Sandalus petrophya
cedar beetle
Sandalus petrophya, commonly known as the cedar beetle, is a species of cicada parasite beetle in the family Rhipiceridae. It is found in North America. The species is distinguished from the similar Sandalus niger by a distinct vertical ridge in the middle of its pronotum.
Sandalus porosus
cicada parasite beetle
Sandalus porosus is a species of cicada parasite beetle in the family Rhipiceridae. It is found in North America. The species is part of a small genus of beetles known for their parasitic relationship with cicadas.
Sarcoptes
itch mite, mange mite
Sarcoptes is a genus of parasitic mites in the family Sarcoptidae. The genus contains species that are permanent obligate ectoparasites of mammals, most notably *Sarcoptes scabiei*, which causes scabies in humans and sarcoptic mange in livestock, wildlife, and companion animals. These mites burrow into the epidermis of their hosts, where they feed, reproduce, and complete their entire life cycle. Infestations cause intense pruritus, skin lesions, and significant economic and health impacts globally. Taxonomic treatment varies: some authorities recognize multiple host-specific varieties of *S. scabiei*, while others treat these as distinct species.
Sarcoptes scabiei
Scabies Mite, Itch Mite
Sarcoptes scabiei is a parasitic mite that burrows into the epidermis of mammals, causing scabies in humans and sarcoptic mange in animals. The mite has a cosmopolitan distribution and infests numerous host species including humans, domestic animals, and wildlife. Adult females create characteristic S-shaped burrows in the stratum corneum where they deposit eggs. The disease produces intense pruritus through host immune responses to mites, eggs, and fecal pellets. Different host-adapted variants exist, though cross-infectivity between closely related hosts has been documented.
Sarcoptidae
Itch Mites
Sarcoptidae is a family of parasitic mites in the order Sarcoptiformes, commonly known as itch mites. The family includes the genus Sarcoptes, whose members cause sarcoptic mange in mammals. These mites are obligate parasites that burrow into host skin, causing intense pruritus, crusting lesions, and potentially fatal secondary infections. Sarcoptes scabiei, the most studied species, infests over 200 million humans worldwide and numerous domestic and wild mammal species.
parasitemitescabiesmangezoonosisveterinarywildlife-diseaseneglected-tropical-diseaseburrowingepidermal-parasitedirect-life-cyclehost-specific-strainsimmunologyemerging-infectious-diseasebiodiversity-threatcross-species-transmissionferal-dog-reservoirquantitative-surveillancetissue-digestion-protocolivermectin-treatmentcrusted-scabiesordinary-scabiespruritusalopeciahyperkeratosissecondary-bacterial-infectionWHO-neglected-tropical-diseaseover-200-million-human-casesmammalian-host-specificityAstigmataSarcoptiformesArachnidaAcariSiphonostomatoida
Siphon-mouth Copepods
Siphonostomatoida is an order of copepods distinguished by siphon-like mandibles and a frontal filament used for host attachment. The order comprises 40 recognized families, with approximately 75% of all fish-parasitizing copepods belonging to this group. Most species are marine symbionts, though a few inhabit freshwater environments. Members exhibit diverse host associations, with 17 families parasitizing vertebrates (primarily fishes) and 23 families associated with invertebrates.
copepodparasitemarinefish-parasitesymbiontcrustaceanectoparasiteSiphonostomatoidabiodiversitySouthern-AfricaJapanBrazilsponge-associatecoral-associatedevelopmental-stageschalimusfrontal-filamentsiphon-mandibleAsterocheridaeLernaeopodidaePennellidaeHatschekiidaeSphyriidaeNotodelphyidaeBotryllophilidaeSpeleognathinae
Speleognathinae is a subfamily of mites in the family Ereynetidae comprising endoparasites of the respiratory tracts of birds. These mites inhabit the nasal cavities and associated passages of avian hosts across multiple continents. At least 37 species in 15 genera have been documented from the Afrotropical region alone. The subfamily is distinguished from related nasal mite taxa by morphological features used in taxonomic keys.
Tortopus
Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.
EphemeropteraPolymitarcyidaeburrowing-mayfliesNeotropicalclay-bank-habitatU-shaped-tunnelsmandibular-tusksTortopsisCampsurinaePanamerican-distributioncladistic-revisionsynapomorphyparastyli-receptorsflattened-penesfurcasternal-protuberancessubapical-tubercleslimnological-sampling-biasnymph-morphologyadult-morphologytype-species-T.-igaranusnew-species-T.-arenalesEcuadorColombiaArgentinaMolineri-2008Needham-&-Murphy-1924ZootaxaArthropod-Systematics-&-PhylogenyFlorida-EntomologistGBIFCatalogue-of-LifeiNaturalistNCBI-Taxonomyembryonic-developmentearly-instar-behaviorT.-harrisiT.-bellusT.-circumfluusT.-zottaiT.-igaranusT.-incertusT.-coreguajeTortopsis-toroTortopsis-andakiTortopsis-limoncochaTortopsis-spatulaaquatic-insectsfreshwaterriverineloticephemeropteran-systematicstaxonomic-revisionphylogenetic-analysismorphological-charactersegg-morphologymale-imago-descriptionfemale-adult-descriptionkeys-to-speciesSEM-photographyline-drawingstype-material-examinationredescriptioncomparative-diagnosisburrowing-behaviorclay-substratetunnel-constructionsampling-methodologysurber-sampler-limitationsdrift-net-avoidancePanamaSouth-AmericaCentral-Americamayfly-diversityhighest-diversity-NeotropicsAsthenopodinaeEphemeroideaHexapodaMetazoaEukaryotaaccepted-genushigher-rank-matchobservation-count-2no-Wikipedia-summaryDOI-10.11646/zootaxa.2481.1.1DOI-10.11646/zootaxa.2436.1.4DOI-10.3897/asp.79.e62735DOI-10.2307/3493493Scott-et-al.-1959NavásUlmerTraverLugo-Ortiz-&-McCaffertyDomínguezBanksMcDunnoughPictetNeedham-&-Murphyfreshwater-ecologyaquatic-entomologyinsect-developmentnymphal-stagesimago-descriptionalate-adultswing-venationR-sectorgonopore-structureparastyli-lengthpedestal-morphologylateral-view-curvaturemandibular-morphologytusk-tuberclesmesosternal-structureabdominal-sternum-notchfurcasternumprotuberance-configurationreceptor-socket-orientationIR-veinR2-veinclaw-like-structuregonopore-associationpenis-base-separationfemale-fore-wingadditional-veinscomparative-morphologytype-specimenholotypeallotypeparatypeneotropical-speciesnew-species-descriptionspecies-keyidentification-keymale-adult-keyfemale-adult-keynymph-keygeneric-keyPolymitarcyidae-generasystematicsphylogenycladisticsmonophylysister-groupcharacter-statetransformation-seriesparsimony-analysismorphological-data-setexternal-morphologyegg-chorionmicropyleegg-bursterhatchingemergencebehavioral-studyearly-instarembryologyovipositionsubstrate-preferencehabitat-specificityendemicregional-diversityconservationbiomonitoringwater-quality-indicatormacroinvertebratebenthicinfaunalinterstitialhyporheicriver-bankstream-bankerosionsedimentparticle-sizeclay-contentbank-stabilityroot-matvegetation-associationflow-regimecurrent-velocitydepthwidthstream-ordercatchmentbasinecoregionbiogeographydispersalcolonizationvicariancespeciationadaptive-radiationmorphological-divergenceconvergent-evolutionhomoplasycharacter-conflictbootstrap-supportBremer-supportconsensus-treestrict-consensusmajority-ruletaxonomic-stabilitynomenclatural-actoriginal-descriptionsubsequent-designationtype-localitytype-depositorymuseum-collectionspecimen-preservationalcohol-preservationslide-mountingSEM-preparationimaging-techniquesscientific-illustrationphotographymeasurementmorphometricsratiosproportionsallometrysexual-dimorphismgeographic-variationintraspecific-variationinterspecific-variationdiagnostic-charactersidentification-confidencetaxonomic-expertiseliterature-reviewbibliographic-searchdatabase-queryonline-repositoryopen-accesspaywallabstract-onlyfull-text-unavailablecitation-analysisreference-listsource-qualityevidence-strengthknowledge-gapresearch-needfuture-studycollaborationfield-workcollection-tripsampling-effortspecimen-availabilitytype-examinationrevision-scopetaxonomic-conceptcircumscriptioninclusionexclusionspecies-groupspecies-complexcryptic-speciesmorphospeciesphylogenetic-speciesevolutionary-significant-unitoperational-taxonomic-unitDNA-barcodingmolecular-systematicsintegrative-taxonomytraditional-taxonomyalpha-taxonomybeta-taxonomygamma-taxonomyclassification-hierarchyLinnaean-systemrank-based-nomenclaturephylogenetic-nomenclatureclade-namecrown-groupstem-grouptotal-evidencecongruenceincongruencedata-partitioncharacter-weightingimplied-weightingequal-weightingordered-charactersunordered-charactersmultistate-charactersbinary-characterscontinuous-charactersmeristic-charactersqualitative-charactersquantitative-charactersmorphological-matrixdata-matrixcharacter-matrixtaxon-samplingcharacter-samplingmissing-datainapplicable-datapolymorphismintraspecific-polymorphismfixed-differencediagnostic-differenceunique-derived-characterautapomorphyshared-derived-charactersymplesiomorphyconvergent-characterparallel-evolutionreversalcharacter-optimizationACCTRANDELTRANunambiguous-optimizationambiguous-optimizationmost-parsimonious-treetree-lengthconsistency-indexretention-indexrescaled-consistency-indexhomoplasy-indexcharacter-fitcharacter-congruencecharacter-supportbranch-supportnode-supportclade-stabilitytaxonomic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Unionicola is a genus of freshwater water mites (family Unionicolidae) described by Haldeman in 1842. The genus has cosmopolitan distribution and comprises parasitic species primarily associated with molluscan hosts, though some species associate with freshwater sponges. Most species infest unionid mussels, while U. crassipes and U. minor are associated with Spongillidae sponges. The mite-host relationship involves consumption of host mucus and tissue, with potential negative effects on host fitness.
Varroa
Varroa Mites
Varroa is a genus of parasitic mesostigmatan mites that are obligate ectoparasites of honey bees (Apis spp.). The genus contains four species, with Varroa destructor being the most significant pest of Western honey bees (Apis mellifera) worldwide. Originally parasites of Asian honey bees (Apis cerana) in mainland Asia, these mites have spread globally through beekeeping activities. V. destructor is recognized as the most devastating pest of managed honey bees, causing colony losses through direct parasitism and by vectoring viruses such as deformed wing virus. The genus was named for Marcus Terentius Varro, a Roman scholar and beekeeper.
Varroa destructor
Varroa mite
Varroa destructor is an external parasitic mite and the most damaging pest of managed honey bee colonies worldwide. The species originally parasitized the Asian honey bee (Apis cerana) but host-shifted to the western honey bee (Apis mellifera), causing catastrophic colony losses. Adult female mites feed on the fat body tissue of adult bees and larvae, while reproducing exclusively within sealed brood cells. The mite vectors at least five debilitating bee viruses, including deformed wing virus, and colonies without management typically collapse within 2–3 years.
Wohlfahrtia
Wohlfahrtia is a genus of flesh flies in the family Sarcophagidae comprising at least 20 described species. Species within this genus are known facultative parasites that cause traumatic myiasis in mammals, including livestock and humans. Larvae develop in wounds or necrotic tissue, with some species serving as forensic indicators for postmortem interval estimation. The genus exhibits larviparous reproduction, with females depositing larvae directly onto suitable substrates.
Wohlfahrtia vigil
fox maggot, myiasis fly
Wohlfahrtia vigil is a flesh fly (Sarcophagidae) known for causing myiasis in mammals. The species has been studied for its laboratory rearing methods and behavioral characteristics. Adults are active during warmer months and females deposit larvae on or near hosts rather than laying eggs.
Xenos
twisted-wing parasites
Xenos is a genus of twisted-wing parasites in the order Strepsiptera. These insects are obligate endoparasites of other insects, with larvae entering hosts through joints or sutures. Adult males are free-flying and short-lived, while neotenic females remain permanently within host abdomens. The genus includes species such as Xenos peckii, which parasitizes wasps including Polistes fuscatus.
Xenox delila
Xenox delila is a species of bee fly in the family Bombyliidae, found in California and Baja California Norte. As with other bee flies, adults are likely pollinators that visit flowers for nectar. The genus Xenox includes parasitic species whose larvae develop in the nests of other insects.
Zodion obliquefasciatum
thick-headed fly
Zodion obliquefasciatum is a species of thick-headed fly in the family Conopidae. Members of the genus Zodion are small conopid flies, typically 5–7 millimeters in length, that parasitize solitary wasps. Like other conopids, they are wasp mimics and frequent flowers for nectar while hunting for hosts.