Parasite

Guides

Acanthocephala
thorny-headed worms, spiny-headed worms, acanthocephalans
Acanthocephala is a phylum of obligate endoparasitic worms known as thorny-headed or spiny-headed worms, characterized by an eversible proboscis armed with recurved hooks used to attach to host intestinal walls. They lack a mouth and digestive system, absorbing nutrients directly through their tegument. The phylum contains approximately 1,200–1,420 described species with complex life cycles involving arthropod intermediate hosts and vertebrate definitive hosts. Recent molecular evidence indicates they are highly modified rotifers, now sometimes classified within Syndermata or Rotifera.
parasite endoparasite helminth proboscis arthropod-intermediate-host vertebrate-definitive-host indirect-life-cycle behavioral-manipulation rotifer-relative aquaculture-pest zoonotic-potential
Acari
Mites and Ticks, mites, ticks
Acari is a subclass of arachnids containing mites and ticks. The group exhibits extraordinary diversity with fossil records dating to the early Devonian period. Recent phylogenetic analyses have challenged the monophyly of Acari, suggesting that the two main lineages—Parasitiformes (ticks and allies) and Acariformes (dust mites, spider mites, chiggers, and allies)—may not share a single common ancestor exclusive of other arachnids. This would render Acari a paraphyletic or polyphyletic grouping rather than a natural taxon. The traditional unifying character, the gnathosoma (a modified mouthpart region), has been shown to be absent or differently constructed in many species across both lineages.
arachnid mite tick parasite predator vector agricultural-pest biological-control forensic-indicator soil-fauna
Aceria paracalifornica
California Sagebrush Gall Mite
Aceria paracalifornica, commonly known as the California Sagebrush Gall Mite, is a species of eriophyid mite in the family Eriophyidae. As a member of the superfamily Eriophyoidea, it is a worm-like, four-legged mite that parasitizes plants. The genus Aceria includes numerous species that cause distinctive plant deformities including galls, witches' brooms, and leaf distortions. The specific epithet "paracalifornica" suggests a close relationship to or similarity with A. californica, a species associated with California sagebrush (Artemisia californica). Eriophyid mites in this genus are highly host-specific, typically specializing on a single plant species or genus.
gall-mite eriophyid parasite Artemisia California-native
Aegidae
aegid isopods
Aegidae is a family of marine and freshwater isopod crustaceans characterized by temporary parasitic relationships with fish hosts. Adults attach to hosts to feed on blood or tissue, then detach to digest meals. The family is distinguished from related Cirolanidae by having only three pairs of hook-like pereiopods rather than seven. Members occur in diverse aquatic habitats from shallow coastal waters to deep ocean environments, with some species documented at depths exceeding 2000 meters.
parasite fish-parasite temporary-parasite isopod crustacean aquaculture-pest marine freshwater deep-sea ectoparasite
Amblyomma tuberculatum
gopher tortoise tick
Amblyomma tuberculatum is a hard tick species endemic to the United States, commonly known as the gopher tortoise tick due to its intimate association with the gopher tortoise (Gopherus polyphemus). Unlike many tick species, its distribution appears more restricted than that of its primary host, with environmental factors such as sand depth and soil composition playing significant roles in habitat suitability. Laboratory studies have documented its complete life cycle, which requires a minimum of 207 days under controlled conditions.
tick parasite specialist gopher-tortoise sand-habitat United-States endemic
Anilocra
Fish Lice
Anilocra is a genus of marine isopods in the family Cymothoidae, commonly known as fish lice. Members are obligate external parasites of teleost fishes, attaching to host skin and feeding on blood. The genus exhibits sequential hermaphroditism, with individuals beginning life as males and transitioning to females as they grow. Parasitism by Anilocra species has been documented to reduce host growth rates, fecundity, and survival.
parasite marine cymothoid fish-parasite hematophagous hermaphrodite coral-reef
Aphomia sociella
Bee Moth, Bumble Bee Wax Moth
Aphomia sociella, commonly known as the bee moth or bumble bee wax moth, is a small pyralid moth native to Europe that has become an economically significant pest of bee colonies. The species is notable for its unique dual-modal mating system combining sex pheromones and ultrasonic courtship songs produced by males using specialized tegular tymbals. Females lay eggs in nests of bumblebees, wasps, and occasionally weakened honeybee hives, where larvae feed on nest materials, pollen, honey, and host brood. Infestation levels increase with proximity to commercial apiaries, causing reduced host reproductive output and weakened immune responses in bumblebees.
pest parasite pollinator-threat acoustic-communication pheromone beekeeping bumblebee wasp
Arguloida
Fish Lice
Arguloida is an order of parasitic crustaceans commonly known as fish lice. The order contains a single family, Argulidae, whose members are obligate ectoparasites of freshwater and marine fishes. These organisms have an uncertain phylogenetic position within Maxillopoda and lack any known fossil record. They are distributed globally across temperate and tropical waters.
parasite fish-parasite aquaculture-pest ectoparasite crustacean
Arrenuridae
Arrenuridae is a family of water mites (Hydrachnidia) in the order Trombidiformes, containing at least three genera and approximately 110 described species. Members are primarily freshwater predators as adults and deutonymphs, with larvae that parasitize aquatic insects. The family exhibits complex mating behaviors mediated by chemical communication, including female-emitted sex pheromones that trigger male responses.
water-mites Hydrachnidia freshwater predator parasite chemical-communication sex-pheromone Arrenurus Ostracoda-predator aquatic-insect-parasite complex-mating-behavior
Arrenurus
water mites
Arrenurus is the largest genus of water mites, comprising approximately 950 species with cosmopolitan distribution in lentic freshwater habitats. Adults are heavily sclerotized, predatory, and exhibit marked sexual dimorphism—males possess diagnostic caudal modifications used in mating, while female taxonomy remains problematic. The life cycle includes seven stages: egg, inactive prelarva, parasitic larva, protonymph, deutonymph, tritonymph, and adult. Larvae are ectoparasites of aquatic insects, particularly Odonata (dragonflies and damselflies), but also Diptera and Coleoptera, utilizing hosts for both nutrition and phoretic dispersal.
water-mite parasite predator freshwater odonate-parasite sexual-dimorphism phoresy stylostome cosmopolitan species-rich-genus
Arrenurus problecornis
Arrenurus problecornis is a species of water mite (family Arrenuridae) described by Cook in 1976. Like other members of the genus Arrenurus, it is part of the subgenus Arrenurus, which is characterized by parasitic larvae that attach to odonate hosts (dragonflies and damselflies). The species belongs to the diverse radiation of water mites (Hydrachnidia), a group comprising over 6,000 species of predatory freshwater arachnids with complex life histories involving parasitic larval stages and free-living predatory adults.
water-mite parasite odonate-parasite freshwater Arrenuridae Parasitengona phoresy
Blissus occiduus
Western Chinch Bug
Blissus occiduus, the western chinch bug, is a phloem-feeding true bug (Hemiptera: Blissidae) that is a significant pest of warm-season turfgrasses, particularly buffalograss and zoysiagrass. The species exhibits strong host preference hierarchies, with buffalograss being the most preferred host followed by zoysiagrass, though it can survive and reproduce on a broad range of grasses including agronomic crops. Field studies have documented inconsistent control with neonicotinoid insecticides, with thiamethoxam showing particularly rapid degradation in buffalograss tissues compared to imidacloprid and clothianidin.
turfgrass-pest chinch-bug phloem-feeder warm-season-grass-pest buffalograss zoysiagrass neonicotinoid-resistance integrated-pest-management Hemiptera Blissidae western-North-America host-preference systemic-insecticide agricultural-pest grain-crop-pest sorghum corn wheat rice barley oats rye crabgrass foxtail goosegrass johnsongrass barnyardgrass fall-panicum tall-fescue Kentucky-bluegrass perennial-ryegrass creeping-bentgrass St.-Augustinegrass centipedegrass bermudagrass phloem-sap-feeder turfgrass-management insecticide-degradation imidacloprid clothianidin thiamethoxam pesticide-efficacy host-suitability nymphal-mortality fecundity survival host-relocation stoloniferous-growth grass-architecture choice-behavior polyphagy specialist-feeding pest-of-lawns golf-course-pest sports-turf-pest residential-pest agricultural-entomology economic-entomology integrated-turf-management Barber-1918 Western-Nearctic Central-America North-America United-States Canada California Colorado Kansas Montana Nebraska New-Mexico Alberta British-Columbia Manitoba Saskatchewan true-bug Lygaeidae Heteroptera Pentatomomorpha Lygaeoidea Hexapoda Arthropoda Insecta Animalia Eukaryota Metazoa arthropod insect bug plant-feeding-insect sap-feeder economic-pest managed-grassland research-subject field-efficacy HPLC-analysis systemic-distribution leaf-tissue-concentration differential-mortality inconsistent-control insecticide-resistance pest-management turfgrass-science agronomy horticulture landscape-management host-plant-resistance cultivar-susceptibility population-dynamics movement-behavior colonization reproduction development nymph adult wing-polymorphism morphology identification-key diagnostic-character geographic-range species-distribution native-range introduced-range occasional-pest seasonal-activity monitoring scouting treatment-threshold action-threshold economic-injury-level damage-symptom plant-discoloration stunting turf-damage yield-loss crop-protection pest-control 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Cephenemyia apicata
nose bot fly
Cephenemyia apicata is a nose bot fly (family Oestridae) that parasitizes Columbian black-tailed deer (Odocoileus hemionus columbianus). First instar larvae develop in the deer's bronchi for 6–8 months before migrating to retropharyngeal pouches. Adult males aggregate at elevated stands of Castanopsis and Quercus in California, with activity occurring from April through July when air temperatures reach 20°C. Females exhibit a distinctive visible attack behavior, landing on or near deer and eliciting curiosity before larvipositing into nostrils.
parasite deer-parasite respiratory-parasite myiasis western-North-America aggregation-behavior risk-spreading-reproduction
Cephenemyia jellisoni
nose bot fly
A parasitic bot fly whose larvae develop in the nasal passages and pharyngeal recesses of deer. Adults are active during warm periods and exhibit specific mating aggregations on elevated vegetation. The species shows a covert attacking strategy when larvipositing, remaining undetected by hosts until the moment of larval deposition.
parasite deer-parasite Oestridae myiasis larviparous cervid-host nasal-bot-fly Pacific-Northwest covert-attacking-behavior site-fidelity
Cephenemyia phobifer
pharyngeal bot of white-tailed deer
Cephenemyia phobifer is a parasitic bot fly (family Oestridae) that develops in the pharyngeal region of white-tailed deer (Odocoileus virginianus). The species exhibits probable larviposition, with females depositing live larvae directly onto or near hosts rather than laying eggs. It has a northeastern Nearctic distribution and completes two generations annually, with winter and summer cycles differing in duration.
parasite bot-fly cervid-parasite larviparous pharyngeal-parasite wildlife-disease Oestridae
Cephenemyia pratti
nose bot fly
Cephenemyia pratti is a species of nose bot fly in the family Oestridae, described by Hunter in 1916. Like other members of the genus Cephenemyia, it is an obligate parasite of cervids (deer family). The species is poorly documented in scientific literature, with only six observations recorded on iNaturalist as of the knowledge cutoff.
parasite diptera oestridae cervidae nasal-bot-fly myiasis
Chelicerata
Chelicerates
Chelicerata is a major subphylum of arthropods characterized by the presence of chelicerae—pincer-like or fang-like appendages that are the first pair of structures before the mouth. The group includes horseshoe crabs, sea spiders, and arachnids (spiders, scorpions, mites, ticks, harvestmen, and allies). Chelicerates diverged from other arthropods by the mid-Cambrian period, approximately 508 million years ago. With over 77,000 described living species and estimates suggesting hundreds of thousands more undescribed species, chelicerates represent one of the most diverse animal groups after insects. Most living species are terrestrial air-breathers, though marine lineages persist.
arthropod arachnid spider scorpion mite tick horseshoe-crab sea-spider venom silk predator parasite
Chrysomelobia
Chrysomelobia is a genus of sexually transmitted parasitic mites in the family Podapolipidae that infest chrysomelid beetles. The genus was originally described from the Americas, Europe and Africa, and was later expanded to include Australian species following the synonymization of Parobia. These mites complete their entire life cycle on adult beetles and do not infest immature host stages.
parasite sexually-transmitted Podapolipidae chrysomelid-beetles arrhenotoky phoresy biological-control host-specificity
Chrysurissa densa
Chrysurissa densa is a species of cuckoo wasp in the family Chrysididae. It is known as a parasite of the nests of pollen wasps in the genus Pseudomasaris, specifically targeting the mud nests of Pseudomasaris vespoides. The species was first described by Cresson in 1865. Like other cuckoo wasps, females lay their eggs in the nests of host wasps, where their larvae consume the pollen and nectar provisions intended for the host's offspring.
parasite cuckoo-wasp pollen-wasp-parasite western-North-America Hymenoptera Chrysididae
Cimex
bed bugs, bedbugs
Cimex is a genus of obligate hematophagous insects in the family Cimicidae, commonly known as bed bugs. The genus includes species specialized on different hosts, with C. lectularius (common bed bug) feeding primarily on humans and C. hemipterus (tropical bed bug) occurring in warmer regions. Other species such as C. pipistrelli, C. pilosellus, and C. adjunctus specialize on bats. Bed bugs are wingless, nocturnal parasites that have undergone global resurgence since the late 1990s, likely due to insecticide resistance, changes in pest control practices, and increased international travel.
bed-bug parasite hematophagy pest urban-entomology insecticide-resistance resurgent-pest nocturnal aggregation-pheromone integrated-pest-management Cimicidae Hemiptera
Cochliomyia
New World screwworm flies, screwworm flies
Cochliomyia is a genus of blowflies (family Calliphoridae) comprising four species: C. macellaria, C. hominivorax, C. aldrichi, and C. minima. The genus is commonly known as New World screwworm flies, distinguished from Old World screwworms. C. hominivorax, the primary screwworm, is an obligate parasite whose larvae feed on living tissue, causing severe myiasis in livestock, wildlife, and humans. C. macellaria, the secondary screwworm, feeds only on necrotic tissue and carrion. The primary screwworm has been eradicated from North America through the sterile insect technique, though it persists in parts of Central and South America.
blowfly myiasis screwworm livestock-pest forensic-entomology sterile-insect-technique parasite veterinary-entomology invasive-species eradication-program
Coptotermes
Coptotermes is a genus of subterranean termites in the family Rhinotermitidae, containing approximately 71 described species. The genus is thought to have originated in Southeast Asia and includes some of the world's most economically destructive termite pests, notably Coptotermes formosanus (Formosan subterranean termite) and C. gestroi (Asian subterranean termite). A 2013 consensus among 31 termite experts reduced the number of validated species from 69 to 21 based on morphological and molecular evidence, with many historical names suspected to be synonyms. Species in this genus exhibit complex social behaviors including age-based task division and unique nitrogen recycling strategies.
termite pest invasive structural-pest subterranean Rhinotermitidae Heterotermitidae economic-pest urban-entomology colony social-insect molting nitrogen-recycling exuviae age-polyethism Formosan-subterranean-termite Asian-subterranean-termite Southeast-Asia-origin wood-feeding cellulose gut-protozoa lower-termite evolutionary-transition tunneling foraging trophallaxis proctodeal-trophallaxis carton-nest Ahamitermes parasite chitin-synthesis-inhibitor CSI-bait molting-site-fidelity invasive-species urban-pest structural-damage quarantine shipping railroad-ties mulch Hawaii Florida Louisiana Taiwan Japan China Philippines Australia temperature-limitation humidity-requirement colony-size several-million alates swarming nematode-association vertical-transfer morphological-plasticity molecular-identification mitochondrial-DNA COII 12S-rRNA 16S-rRNA consortium-taxonomy taxonomic-cold-case synonymy Wasmann-1896
Cremastocheilus constricticollis
Cremastocheilus constricticollis is a species of anteater scarab beetle native to the southwestern United States. Like other members of its genus, it is a myrmecophile that lives within ant colonies as a predator of ant larvae and pupae. The species exhibits specialized morphological adaptations including a dense, heavily armored exoskeleton and modified mouthparts for piercing soft ant cuticle. Adults are known to employ death-feigning behavior to gain entry to ant nests. The species was described by Cazier in 1940 and is among roughly 35 recognized North American species in this genus, which is currently undergoing taxonomic revision.
myrmecophile parasite predator death-feigning thanatosis scarab-beetle ant-nest southwestern-United-States
Cremastocheilus crinitus
anteater scarab beetle
Cremastocheilus crinitus is a myrmecophilous scarab beetle known for its specialized parasitic relationship with ants. Adults enter ant colonies by feigning death, allowing scavenging workers to carry them into nests. Once inside, they feed on ant larvae and pupae using modified piercing mouthparts. The species is part of a genus of approximately 35 North American species, commonly called "anteater scarabs" for their predatory behavior toward ant brood.
myrmecophily parasite predator death-feigning thanatosis Scarabaeidae Cetoniinae Formica ant-associated North-America
Cremastocheilus harrisii
Eastern Ant-loving Scarab
Cremastocheilus harrisii is a species of anteater scarab beetle in the family Scarabaeidae. As a member of the genus Cremastocheilus, it exhibits the specialized myrmecophilous lifestyle characteristic of this group, living within ant colonies and feeding on ant brood. The species is part of a genus of approximately 35 North American species that have evolved remarkable morphological and behavioral adaptations for surviving in hostile ant nest environments.
myrmecophile scarab-beetle predator parasite Formica-associate death-feigning cryptic-lifestyle
Cremastocheilus planipes
Cremastocheilus planipes is a myrmecophilous scarab beetle in the genus Cremastocheilus, commonly known as anteater scarabs. These beetles are specialized inhabitants of ant colonies, where they live as predators of ant brood. The species exhibits remarkable morphological and behavioral adaptations for infiltrating and surviving within hostile ant nests. Like other members of its genus, C. planipes is poorly studied and infrequently encountered due to its cryptic lifestyle.
myrmecophile parasite predator scarab ant-associated cryptic desert-fauna
Cremastocheilus quadratus
anteater scarab beetle
Cremastocheilus quadratus is a myrmecophilous scarab beetle in the family Scarabaeidae, first described by Fall in 1912. Like other members of its genus, it lives within ant colonies as a predator of ant larvae and pupae. The species is found in the southwestern United States and northwestern Mexico.
myrmecophile parasite predator scarab-beetle ant-associate
Cremastocheilus schaumii
anteater scarab beetle
Cremastocheilus schaumii is a North American scarab beetle notable for its obligate myrmecophilous lifestyle. Adults live within ant colonies as predators of ant larvae and pupae, employing specialized morphological adaptations to avoid detection and expulsion by host workers. The species exhibits a complex life cycle involving death-feigning behavior to gain entry to nests, chemical camouflage through acquisition of colony scent, and modified mouthparts for piercing ant brood. Two subspecies are recognized: C. s. schaumii and C. s. tibialis.
myrmecophile predator scarab-beetle ant-associate death-feigning parasite
Cremastocheilus squamulosus
Cremastocheilus squamulosus is a species of anteater scarab beetle in the family Scarabaeidae. Like other members of its genus, it is a myrmecophile that lives within ant colonies. The species occurs in the southeastern United States, where it has been recorded from Florida, Georgia, North Carolina, and South Carolina.
myrmecophile parasite predator scarab-beetle Formica-associate
Cuterebra
rodent bots, Glire Bot Flies
Cuterebra is a genus of New World bot flies (subfamily Cuterebrinae) comprising approximately 78 species that parasitize rodents and lagomorphs. Adults are large, robust flies resembling bumble bees, with non-functional mouthparts and short lifespans devoted entirely to reproduction. Females oviposit near host burrow entrances; larvae hatch in response to host body heat, enter through natural orifices or wounds, and develop subcutaneously for approximately one month before exiting to pupate in soil. The genus exhibits notable host specificity, with individual species typically restricted to single host species or closely related groups.
bot-fly myiasis parasite rodent-parasite lagomorph-parasite New-World Cuterebrinae warble subcutaneous-parasite rarely-observed-adult mimicry bumble-bee-mimic
Cuterebra abdominalis
rabbit bot fly
Cuterebra abdominalis is a species of New World skin bot fly in the family Oestridae. It is a parasite of lagomorphs (rabbits and hares), with larvae developing subcutaneously in host tissues. Adults are short-lived, lack functional mouthparts, and do not feed. The species exhibits Batesian mimicry, resembling bumble bees in appearance. It is one of approximately 34 recognized North American species in the genus Cuterebra, with seven species specializing on rabbit hosts.
parasite mimicry bot-fly lagomorph rabbit Oestridae Cuterebrinae myiasis warble Batesian-mimicry short-lived-adult non-feeding-adult soil-pupation spring-emergence
Cuterebra americana
woodrat bot fly
Cuterebra americana, the woodrat bot fly, is a New World skin bot fly in the family Oestridae. It is a parasitic species whose larvae develop subcutaneously in mammalian hosts. The species has been documented successfully developing in atypical hosts, including the black rat (Rattus rattus) in Florida. Adults are rarely encountered in the field and lack functional mouthparts, living only to reproduce.
bot-fly parasite myiasis rodent-parasite Cuterebrinae Oestridae Diptera
Cuterebra arizonae
New World skin bot fly
Cuterebra arizonae is a species of New World skin bot fly in the family Oestridae. It was described by Sabrosky in 1986. Like other members of the genus Cuterebra, it is a parasitic fly whose larvae develop subcutaneously in mammalian hosts. Adults are rarely encountered in the field and lack functional mouthparts.
bot-fly parasite Diptera Oestridae Cuterebrinae New-World-skin-bot-fly Sabrosky-1986 Arizona
Cuterebra atrox
Cuterebra atrox is a species of New World skin bot fly in the family Oestridae. Like other members of the genus Cuterebra, it is a parasitic fly whose larvae develop subcutaneously in mammalian hosts. The species was described by Clark in 1848. Adults are rarely encountered in the field and do not feed, lacking functional mouthparts.
bot-fly parasite oestridae cuterebrinae diptera new-world mammal-parasite
Cuterebra bajensis
Cuterebra bajensis is a species of New World skin bot fly in the family Oestridae, described by Sabrosky in 1986. The species belongs to a genus of obligate mammalian parasites whose larvae develop subcutaneously in rodents and lagomorphs. Adults are rarely encountered in the field and do not feed.
bot-fly parasite mammal-parasite New-World Cuterebrinae
Cuterebra buccata
rabbit bot fly
Cuterebra buccata is a New World bot fly in the family Oestridae, commonly known as the rabbit bot fly. Adults are large, bumble bee-sized flies that lack functional mouthparts and do not feed. The species is host-specific to lagomorphs, particularly cottontail rabbits (Sylvilagus floridanus). Larvae develop subcutaneously in warbles, feeding on host fluids rather than tissue. Adults are rarely encountered in the field and have short lifespans focused on reproduction.
bot-fly myiasis parasite lagomorph rabbit Oestridae Cuterebrinae bumble-bee-mimicry warble subcutaneous-parasite host-specific non-feeding-adult puparium Cuterebra Sylvilagus cottontail-rabbit New-World North-America Virginia Missouri Vermont spring-emergence late-summer-prevalence rarely-encountered short-lived-adult evolutionary-coevolution benign-parasitism soil-pupation larval-migration subcutaneous-development Diptera Insecta entomology medical-entomology veterinary-entomology wildlife-disease parasitology host-parasite-interaction cuterebrid rabbit-parasite lagomorph-parasite bot warble-fly gadfly Oestrus-buccata Fabricius-1775 Cuterebra-buccata-(Fabricius,-1775)
Cuterebra cochisei
Cuterebra cochisei is a species of New World skin bot fly described by Sabrosky in 1986. As a member of the genus Cuterebra, it is an obligate parasite of mammals with larvae that develop subcutaneously in host tissue. The species is one of 34 recognized North American Cuterebra species. Very little specific information has been published about this particular species.
parasite mammal-parasite myiasis obligate-parasite bot-fly New-World-skin-bot-fly Oestridae Cuterebrinae Diptera insect arthropod
Cuterebra cuniculi
rabbit bot fly
Cuterebra cuniculi is a New World skin bot fly endemic to Georgia and Florida. Its larvae are obligate parasites of the eastern cottontail (Sylvilagus floridanus) and marsh rabbit (Sylvilagus palustris), developing subcutaneously in warbles. Adults are short-lived, lack functional mouthparts, and do not feed.
parasite endemic lagomorph-parasite myiasis rarely-observed-adult
Cuterebra emasculator
Squirrel Bot Fly
Cuterebra emasculator is a New World bot fly and obligate internal parasite of tree squirrels and chipmunks in eastern North America. First described by Asa Fitch in 1856, its larvae develop subcutaneously in hosts, creating characteristic swellings called warbles. The species name originated from a false belief that larvae consumed host testicles; research has shown this is not the case and parasitism does not reduce host fertility. Adults are rarely encountered, lack functional mouthparts, and live only a few days focused solely on reproduction.
bot-fly parasite warble sciurid chipmunk squirrel Oestridae internal-parasite subcutaneous-parasite myiasis
Cuterebra fontinella fontinella
mouse bot fly, white-footed mouse bot fly
Cuterebra fontinella fontinella is a subspecies of New World bot fly (family Oestridae) that parasitizes small mammals, primarily white-footed mice (Peromyscus leucopus). The adult flies are large, bumble bee-sized, and lack functional mouthparts, living only briefly to reproduce. Females lay eggs near host burrow entrances; larvae hatch in response to host body heat, enter through orifices or wounds, and develop subcutaneously for approximately one month before exiting to pupate in soil. Despite forming conspicuous warbles that can reach 5% of host body weight, infections appear to have minimal negative impact on host survival and may even correlate with increased longevity in some populations.
parasite myiasis rodent-parasite bot-fly Oestridae Cuterebrinae Peromyscus white-footed-mouse forest-edge aggregation-behavior bimodal-reproduction benign-parasitism mimicry bumble-bee-mimic
Cuterebra lepusculi
cottontail rabbit botfly
Cuterebra lepusculi is a New World skin bot fly (family Oestridae) that parasitizes cottontail rabbits (Sylvilagus spp.). Adults are short-lived, lack functional mouthparts, and do not feed. Females deposit eggs near rabbit burrow entrances; larvae hatch upon detecting host body heat, enter through natural orifices or wounds, and develop in subcutaneous warbles for approximately one month before exiting to pupate in soil.
parasite myiasis rabbit-parasite bot-fly Diptera Oestridae Cuterebrinae Sylvilagus Idaho
Cuterebra mirabilis
Cuterebra mirabilis is a rare species of New World skin bot fly and the largest member of the rabbit-infesting group within the genus Cuterebra. As of 2009, the species was known from only three specimens, all collected in New Mexico. Like other Cuterebra species, it is an obligate parasite of lagomorphs, with larvae developing subcutaneously in host tissue. The specific epithet "mirabilis" (Latin for "extraordinary") reflects the species' remarkable size and striking appearance among bot flies.
rare parasite lagomorph-parasite New-Mexico-endemic largest-rabbit-bot-fly
Cuterebra neomexicana
Cuterebra neomexicana is a species of New World skin bot fly described by Sabrosky in 1986. Like other members of the genus Cuterebra, it is a mammalian parasite with larvae that develop subcutaneously in hosts. The species is poorly documented in the scientific literature, with minimal published information on its biology, host associations, and distribution beyond the original description.
data-deficient parasite mammal-parasite New-World-bot-fly
Cuterebra polita
Cuterebra polita is a New World skin bot fly whose larvae are obligate parasites of rodents. Adults are non-feeding and short-lived, lacking functional mouthparts. Larvae develop in subcutaneous warbles on host rodents, feeding on host tissues and fluids. The species has been documented in four rodent hosts in Florida: white-footed mouse, cotton mouse, eastern woodrat, and cotton rat.
parasite rodent-parasite myiasis obligate-parasite bot-fly New-World-skin-bot-fly Cuterebrinae medical-entomology veterinary-entomology
Cuterebra tenebriformis
Diptera Oestridae Cuterebra bot-fly parasite North-America rodent-parasite mammal-parasite endoparasite myiasis New-World-skin-bot-fly Cuterebrinae Sabrosky-1986 rarely-encountered adult-non-feeding host-specific subcutaneous-parasite warble-forming puparium glade-habitat Missouri-endemic southeastern-US
Cyladini
Cyladini is a tribe of weevils in the family Brentidae, subfamily Apioninae. The tribe includes economically significant pests of sweet potato, particularly the sweetpotato weevil (Cylas formicarius), which causes substantial damage to storage roots in tropical and subtropical production regions. Members of this tribe are characterized by their association with Convolvulaceae host plants and their status as specialized root-feeding pests.
pest agriculture sweet-potato Convolvulaceae integrated-pest-management Cylas-formicarius tropical-agriculture subtropical-agriculture root-feeding-weevil storage-root-damage economic-pest biological-control crop-protection southeastern-United-States global-food-security low-input-farming drought-tolerant-crop nematode-interactions Beauveria-bassiana entomopathogenic-nematodes resistant-varieties crop-rotation cover-crops IPM Brentidae Apioninae Coleoptera Insecta Hexapoda Arthropoda Animalia Claire-Schloemer Auburn-University University-of-Wisconsin-Madison Journal-of-Integrated-Pest-Management Scott-Graham Kathy-Lawrence Covington Bonita Beauregard Orleans guava-root-knot-nematode root-knot-nematode wireworm cucumber-beetle flea-beetle Purpureocillium-lilacinum biopesticide green-bridge-effect mustard glucosinolate peanut bahiagrass sandy-soil drought-adaptation biofuel animal-feed nutritional-value food-security sustainable-agriculture environmental-impact microbial-control fungal-pathogen entomopathogenic-fungus biological-nematicide nematode-egg-parasitism pest-management agricultural-entomology crop-loss yield-reduction root-quality marketability integrated-management cultural-control selective-chemical-use productivity natural-processes versatility global-cultivation tropical-regions subtropical-regions southeastern-U.S.domestic-production revenue acreage harvest farming grower breeding research cultivar-development evolving-threats pest-pressure soil-health field-preparation winter-habitat insect-habitat parasitism infestation damage loss challenge defense arsenal battle war resilience adaptation innovation strategy sustainability livelihood nourishment food-production agricultural-research entomology plant-protection crop-science horticulture root-crop tuber storage-organ feeding-injury galling cracked-roots stunted-growth water-uptake nutrient-uptake pathogen-interaction exacerbated-damage variety-selection hardy-variety robust-performance high-yield nematode-resistance pest-resistance vulnerability spreading-pest population-reduction soil-improvement spring-planting toxic-compound natural-property microbial-product formulation application evaluation maximization potential urgency growing-threat comprehensive-strategy balance enablement essential-crop security defiance difficult-conditions menacing-pests formidable-threats production-continuation face-to-face threat disease resilient-crop growing-conditions menace story surface beneath relentless adversary favor condition powerful foe tiny parasite attack formation diminish ability result unattractive worse matter interaction significant devastating ruin inflict further compromise grapple thankfully farmer begin select hardy withstand traditional gain traction vulnerable highlight need continue breed effort develop capable oppose evolve after sturdy cultural management rotation cover cropping line take year off grow rotate non-host prove effective reduce population breathe life improve health suppress prepare example contain compound call break down release substance wary provide represent method harness help win spore fungus infect kill type tap different microscopic animal parasitize plus use combat attention egg aid fight minimize impact despite advance question remain require maximize particularly urgent pose key enhance approach integrate practice control selective chemical enable natural process nutritional value essential crop global food region support million address pressure innovative sustainable ensure live many formidable production although come face resilient able defy difficult read more journal integrated Claire Schloemer Ph.D.student conduct earn master degree email share link friend open new window print Facebook LinkedIn Bluesky Mastodon X Reddit Threads related discover from today subscribe get latest post send your Cylas formicarius sweet potato sweetpotato weevil photo courtesy show here cucumber beetle root-knot nematode harm U.S.guide detail variety option beloved delicious taste benefit bright color rank seventh most important world tropical subtropical area around serve feed source good pick low-input style well adapted sandy soil drought United States 2022 over 130,000 acre generate nearly 600 southeastern lead domestic Auburn University weather article publish March colleague Scott Graham Kathy Lawrence various among plant-parasitic specifically root system cause gall plant water nutrient stunted growth yield crack make interact pathogen exacerbate insect know storage 7 annual southern flea quality leave economic own resistance better meanwhile newer high robust performance however resist guava species spread next season winter field spring toxic green bridge effect habitat feeder symptom infection biological property microbe Beauveria bassiana agent formulate environmentally friendly product easy apply similarly beneficial entomopathogenic nematicide Purpureocillium lilacinum solution environmental resistant nematode-resistant cultivar comprehensive cultivation Wisconsin-Madison cschloemer wisc edu
Cyphomyrmex
fungus-growing ants
Cyphomyrmex is a genus of small, drab-colored fungus-growing ants in the tribe Attini, found primarily in the Neotropics. These ants cultivate fungi in the tribe Leucocoprineae as their primary food source, with most species growing fungal nodules rather than full mycelial gardens. Colonies are typically monogynous and small, rarely exceeding 500 workers. The genus is divided into two species complexes: the strigatus complex (South America only) and the rimosus complex (southern North America to South America). Cyphomyrmex represents a basal lineage among attine ants and serves as sister group to Mycetophylax.
fungus-growing-ants attine-ants Myrmicinae Neotropical mutualism fungiculture Leucocoprineae monogyny parasitoid-wasps social-parasitism chemical-ecology alkaloid-venom nest-architecture behavioral-plasticity cytogenetics chromosome-evolution Guiana-Shield Brazil Panama Puerto-Rico Mesoamerica Argentina United-States Texas California Florida yeast-gardens mycelial-gardens rimosus-complex strigatus-complex age-polyethism trophallaxis thanatosis feigning-death Diapriidae Acanthopria Mimopriella Megalomyrmex social-parasite C.-longiscapus C.-transversus C.-laevigatus C.-rimosus C.-minutus C.-lectus C.-cornutus C.-costatus pheromones 3-octanol farnesenes trail-following alarm-signal diketopiperazines antifungal-compounds defensive-behavior colony-defense flight-response recruitment Lanchester's-laws karyotype chromosome-number rDNA microsatellites FISH chamber-depth gallery-length fire-disturbance savanna-forest-transition arboreal-nesting clay-auricles soil-nests wood-nests epiphytic-nests moss-nests coconut-nests desiccation-susceptibility moist-habitats basal-attine phylogenetics sister-group chemical-evolution venom-alkaloids behavioral-manipulation host-parasite-interaction ecosystem-engineering nutrient-cycling decomposition organic-matter insect-feces exoskeleton-substrate garden-transplantation worker-polymorphism queen-number dealate-queens alates brood-care ecdysis-assistance foraging sugar-collection plant-exudates fungal-domestication nodule-cultivation mycelium-cultivation agricultural-behavior nest-entrance funnel-structure exposed-gardens symbiont-diversity geographic-variation research-model evolutionary-biology mutualism-evolution ant-fungus-coevolution chemical-communication social-insects eusociality colony-size small-colonies slow-movement crypsis debris-mimicry predation-avoidance parasitoid-defense wasp-killing colony-abandonment resource-rescue thief-ant-defense venom-resistance toxicity-tolerance behavioral-suppression playing-dead alkaloid-manipulation fungal-secondary-metabolites antibacterial-compounds antiviral-compounds garden-protection microbial-defense nest-microclimate temperature-regulation humidity-requirements depth-effects fire-adaptation disturbance-ecology habitat-specificity substrate-specialization clay-construction soil-manipulation architectural-plasticity vertical-nesting embankment-nesting overhang-nesting pendant-nests swallow-nest-mimicry research-accessibility field-biology tropical-ecology Neotropical-myrmecology biodiversity conservation invasive-species-context native-species biological-control Integrated-Pest-Management agricultural-ecology forest-ecology savanna-ecology transitional-habitats disturbed-areas urban-ecology anthropogenic-impact climate-sensitivity habitat-limitation range-boundaries dispersal colonization establishment population-dynamics demography colony-density nest-spacing intraspecific-competition interspecific-competition territoriality resource-defense combat-behavior aggression-levels recruitment-behavior chemical-signaling alarm-pheromones trail-pheromones recognition-cues cuticular-hydrocarbons venom-composition defensive-secretions mandibular-glands Dufour's-gland metapleural-glands antibiotic-production grooming-behavior fungal-inoculation integumental-fungi yeast-growth larval-grooming garden-maintenance substrate-preparation fecal-fertilizer anal-trophallaxis crop-regurgitation fluid-application drying-behavior nodule-formation transplantation-behavior garden-establishment ceiling-gardens root-association architectural-innovation nest-site-selection microhabitat-choice moisture-requirements thermal-buffering predator-avoidance parasite-avoidance disease-resistance social-immunity collective-behavior division-of-labor task-allocation temporal-polyethism worker-age nurse-workers forager-workers soldier-caste-absence monomorphism worker-size-variation queen-worker-differentiation reproductive-caste gyne-production male-production seasonal-reproduction colony-founding claustral-founding non-claustral-founding fungal-inoculum founding-queen first-worker-generation nutritional-dependency fat-reserves metabolic-adaptation digestive-physiology enzyme-complement fungal-enzyme lignocellulose-degradation nutritional-mutualism obligate-mutualism vertical-transmission horizontal-transmission symbiont-switching cultivar-replacement garden-failure colony-mortality population-turnover generation-time longevity survivorship life-expectancy aging senescence reproductive-investment colony-growth colony-decline budding fission reproduction-mode genetic-structure relatedness mating-system nuptial-flight mate-choice sperm-storage colony-odor nestmate-recognition foreign-rejection agonistic-behavior territorial-marking boundary-defense raiding-behavior robbing-behavior cleptobiosis dulosis slavery inquilinism guest-ants myrmecophiles nest-associates commensalism inquilines parasite-tolerance host-manipulation venom-alkaloid pyrrolidine indolizidine piperidine nonanal farnesene sesquiterpene agriculture-chemical gaster-extract chemical-diversity chemical-simplicity evolutionary-precursor derived-chemistry biosynthetic-pathway semiochemical infochemical communication information-transfer colony-organization social-structure polygyny queen-condition dealation wing-shedding ovarian-development fecundity egg-laying brood-development developmental-rate temperature-dependence humidity-dependence food-availability nutritional-quality fungal-productivity garden-yield worker-production colony-efficiency energy-allocation resource-allocation foraging-efficiency search-behavior prey-detection food-retrieval transport-behavior load-size travel-speed path-integration orientation navigation homing-behavior learning memory experience individual-recognition context-dependent-behavior threat-assessment risk-management predator-inspection alarm-propagation recruitment-activation mobilization colony-response emergency-behavior panic-response controlled-response defensive-strategy offensive-strategy escalation de-escalation conflict-resolution fight-avoidance retreat-behavior surrender-signal dominance-hierarchy reproductive-skew worker-policing queen-policing egg-eating oviposition-suppression reproductive-conflict kin-conflict worker-reproduction male-production-by-workers anarchy social-parasitism-detection parasite-recognition enemy-recognition nest-intrusion nest-defense perimeter-defense chamber-defense brood-defense queen-defense self-sacrifice suicidal-defense autothysis sting-use mandible-use spraying-behavior gaster-flexion chemical-defense mechanical-defense armor cuticle-thickness pigmentation masquerade mimicry Batesian-mimicry Muellerian-mimicry aggressive-mimicry Wasmannian-mimicry myrmecophily myrmecophagy ant-plant-mutualism trophobiosis extrafloral-nectary domatia plant-ant-interaction herbivore-defense seed-dispersal pollination ecosystem-service functional-role trophic-position detritivore decomposer saprotroph necrotroph fungivore omnivore predator scavenger trophic-level energy-flow carbon-cycling nitrogen-cycling phosphorus-cycling soil-formation bioturbation aeration water-infiltration microbial-community bacteria actinomycetes Pseudonocardia disease-suppression garden-hygiene weeding-behavior pruning-behavior fungal-grooming antibiotic-application mutualist-protection agricultural-disease crop-pathology escovopsis garden-parasite competitor-fungus pathogen-defense immune-response individual-immunity grooming allogrooming self-grooming nest-cleaning refuse-management fecal-management waste-disposal midden-formation territory-size home-range foraging-range recruitment-range exploration exploitation information-use public-information private-information cue-use signal-use communication-network interaction-network social-network colony-network population-network community-ecology species-interaction apparent-competition exploitative-competition interference-competition resource-partitioning niche-differentiation habitat-filtering environmental-gradient altitudinal-distribution latitudinal-distribution continental-distribution island-distribution endemism biogeography phylogeography historical-biogeography vicariance range-expansion range-contraction refugia glacial-cycles climate-change-impact habitat-fragmentation deforestation land-use-change agricultural-intensification urbanization pollution pesticide-exposure heavy-metal-accumulation conservation-status population-viability extinction-risk protected-area reserve-design habitat-corridor connectivity metapopulation source-sink-dynamics rescue-effect mass-effect ecological-theory behavioral-ecology evolutionary-ecology functional-ecology trait-based-ecology comparative-analysis phylogenetic-comparative-methods character-evolution ancestral-state-reconstruction convergent-evolution parallel-evolution adaptive-radiation species-diversification speciation hybridization introgression gene-flow genetic-differentiation population-structure molecular-ecology genomics transcriptomics proteomics metabolomics volatile-organic-compound cuticular-hydrocarbon glandular-secretion sting-apparatus mandible-morphology head-morphology mesosomal-morphology petiolar-morphology gastral-morphology leg-morphology antennal-morphology sensilla chemoreception mechanoreception thermoreception photoreception vision compound-eye ocelli brain-morphology neuroanatomy learning-capacity memory-capacity cognitive-ability problem-solving tool-use sequence-learning route-learning spatial-memory temporal-memory social-learning cultural-transmission tradition colony-personality behavioral-syndrome boldness exploration-tendency activity-level aggression-level foraging-tendency defensive-tendency stress-response physiological-stress thermal-stress desiccation-stress starvation-stress immune-challenge wound-healing metabolic-rate respiration energy-expenditure water-balance osmoregulation excretion Malpighian-tubules rectum hindgut midgut foregut crop proventriculus digestive-enzyme nutrient-absorption lipid-metabolism carbohydrate-metabolism protein-metabolism amino-acid-metabolism nitrogen-excretion uric-acid urea ammonia osmolyte cryoprotectant heat-shock-protein stress-protein antioxidant-defense oxidative-stress reactive-oxygen-species aging-biology senescence-theory disposable-soma antagonistic-pleiotropy mutation-accumulation life-history-theory r-K-selection bet-hedging iteroparity semelparity reproductive-effort parental-investment maternal-effect paternal-effect epigenetics DNA-methylation histone-modification non-coding-RNA gene-regulation transcription-factor signaling-pathway hormone juvenile-hormone ecdysone insulin-signaling TOR-signaling nutrient-sensing reproductive-signaling caste-determination sex-determination environmental-sex-determination genetic-sex-determination haplodiploidy arrhenotoky thelytoky parthenogenesis arrhenotokous-parthenogenesis automixis apomixis central-fusion terminal-fusion genomic-imprinting parental-conflict kin-selection inclusive-fitness Hamilton's-rule relatedness-asymmetry sex-ratio-theory local-mate-competition local-resource-competition local-resource-enhancement split-sex-ratio worker-sex-ratio queen-sex-ratio colony-sex-ratio population-sex-ratio operational-sex-ratio effective-population-size genetic-drift inbreeding inbreeding-depression outbreeding outbreeding-depression heterosis hybrid-vigor genetic-load mutation-rate effective-number-of-codons codon-usage-bias GC-content genome-size karyotype-evolution chromosomal-rearrangement fusion inversion translocation polyploidy aneuploidy chromosomal-polymorphism rDNA-cluster heterochromatin euchromatin centromere telomere nucleolus-organizer-region satellite-DNA minisatellite microsatellite simple-sequence-repeat transposable-element retrotransposon DNA-transposon horizontal-gene-transfer endosymbiont Wolbachia Cardinium Rickettsia Spiroplasma bacterial-symbiont fungal-symbiont viral-symbiont parasite pathogen disease epidemiology transmission-dynamics virulence resistance tolerance recovery immunity vaccination prophylaxis medicinal-behavior self-medication thermoregulation behavioral-thermoregulation social-thermoregulation microclimate-selection nest-temperature brood-temperature fungal-garden-temperature metabolic-heat heat-dissipation evaporative-cooling behavioral-cooling fanning ventilation chamber-design gallery-design entrance-design air-flow humidity-regulation water-collection water-storage desiccation-resistance cuticular-waterproofing spiracle-control respiratory-water-loss excretory-water-loss water-recycling metabolic-water dietary-water environmental-water rainfall-dependence seasonal-activity dry-season wet-season monsoon drought flood fire disturbance succession primary-succession secondary-succession climax-community pioneer-species habitat-specialist habitat-generalist edge-species interior-species gap-species canopy-species understory-species ground-species subterranean-species arboreal-species liana-species epiphyte-species lithophyte-species rheophyte-species aquatic-margin-species terrestrial-species fossorial-species cursorial-species scansorial-species volant-species winged-reproductive dealate ergatoid brachypterous apterous polymorphism caste-polymorphism queen-polymorphism male-polymorphism size-polymorphism allometry isometry shape-variation morphological-integration modularity evolvability constraints developmental-stability fluctuating-asymmetry directional-asymmetry antisymmetry phenotypic-plasticity reaction-norm genotype-environment-interaction grandparental-effect indirect-genetic-effect social-environment-effect colony-effect density-dependence frequency-dependence negative-frequency-dependence positive-frequency-dependence apostatic-selection anti-apostatic-selection search-image predator-learning prey-learning friend-recognition cuticular-hydrocarbon-profile gestalt-odor colony-specific-odor individual-specific-odor task-specific-odor age-specific-odor reproductive-odor queen-odor queen-pheromone primer-pheromone releaser-pheromone signal-pheromone modulatory-pheromone alarm-pheromone trail-pheromone recruitment-pheromone aggregation-pheromone sex-pheromone marking-pheromone territorial-pheromone defensive-pheromone poison-gland Dufour-gland mandibular-gland postpharyngeal-gland metapleural-gland sternal-gland tarsal-gland anal-gland rectal-gland pygidial-gland mandibular-brush tibial-spur stridulatory-organ subgenual-organ Johnston's-organ tympanal-organ hair-sensillum campaniform-sensillum basiconic-sensillum coeloconic-sensillum placoid-sensillum sensilla-trichodea sensilla-basiconica sensilla-coeloconica sensilla-ampullacea sensilla-campaniformia sensilla-placodea olfactory-receptor-neuron gustatory-receptor-neuron mechanoreceptor-neuron thermoreceptor-neuron hygroreceptor-neuron photoreceptor-neuron visual-processing olfactory-processing gustatory-processing mechanosensory-processing central-complex mushroom-body antennal-lobe optic-lobe suboesophageal-ganglion ventral-nerve-cord sympathetic-nervous-system neurosecretion neurohemal-organ corpus-cardiacum corpus-allatum prothoracic-gland ventral-gland pericardial-gland neuroendocrine-regulation hormonal-regulation pheromonal-regulation neural-regulation behavioral-regulation social-regulation environmental-regulation developmental-regulation reproductive-regulation metabolic-regulation homeostatic-regulation allostatic-regulation stress-regulation immune-regulation circadian-rhythm circannual-rhythm photoperiodism thermoperiodism biological-clock zeitgeber entrainment free-running-period phase-response-curve masking-effect arousal-threshold sleep torpor diapause quiescence dormancy aestivation hibernation cold-hardiness heat-hardiness desiccation-hardiness anoxia-hardiness starvation-hardiness radiation-hardiness pressure-hardiness gravity-perception magnetic-field-perception electric-field-perception sound-perception vibration-perception substrate-borne-signal air-borne-signal seismic-signal chemical-signal visual-signal tactile-signal auditory-signal multimodal-signal composite-signal redundant-signal amplifying-signal attenuating-signal honest-signal deceptive-signal costly-signal cheap-signal conventional-signal index-signal handicap-signal amplifier social-selection sexual-selection sperm-competition cryptic-female-choice male-male-competition female-female-competition parent-offspring-conflict sibling-conflict worker-queen-conflict worker-worker-conflict sexual-conflict antagonistic-coevolution arms-race evolutionary-dynamics community-dynamics ecosystem-dynamics biogeochemical-cycle carbon-cycle nitrogen-cycle phosphorus-cycle sulfur-cycle water-cycle trophic-transfer production-efficiency consumption-efficiency assimilation-efficiency ecological-efficiency Lindeman-efficiency 10%-rule food-chain food-web trophic-cascade top-down-control bottom-up-control wasps-control donor-control recipient-control interaction-strength interaction-web ecological-network network-topology network-metrics degree-distribution connectance nestedness compartmentalization robustness fragility resilience recovery-time return-time elasticity perturbation pulse-disturbance press-disturbance chronic-disturbance acute-disturbance catastrophic-disturbance natural-disturbance anthropogenic-disturbance habitat-destruction habitat-degradation habitat-modification habitat-loss habitat-isolation edge-effect matrix-effect permeability corridor-effect stepping-stone source-population sink-population metapopulation-dynamics extinction-debt colonization-credit relaxation-effect time-lag historical-legacy priority-effect founder-effect bottleneck-effect selection natural-selection artificial-selection directional-selection stabilizing-selection disruptive-selection balancing-selection frequency-dependent-selection density-dependent-selection r-selection K-selection c-selection s-selection a-selection bet-hedging-selection fluctuating-selection correlational-selection multivariate-selection indirect-selection correlated-response genetic-correlation phenotypic-correlation environmental-correlation correlation-matrix variance-covariance-matrix G-matrix P-matrix E-matrix M-matrix quantitative-genetics heritability narrow-sense-heritability broad-sense-heritability genetic-variance additive-genetic-variance dominance-variance epistatic-variance environmental-variance phenotypic-variance genotype-environment-interaction-variance maternal-variance paternal-variance common-environment-variance special-environment-variance measurement-error-variance response-to-selection realized-heritability predicted-response breeder's-equation selection-gradient selection-differential Lande-equation multivariate-breeder's-equation genetic-constraint evolutionary-constraint developmental-constraint functional-constraint phylogenetic-constraint historical-constraint trade-off allocation-trade-off acquisition-trade-off conversion-trade-off maintenance-trade-off reproduction-survival-trade-off growth-reproduction-trade-off immune-reproduction-trade-off stress-reproduction-trade-off foraging-predation-trade-off competition-colonization-trade-off tolerance-resistance-trade-off plasticity-canalization-trade-off generalist-specialist-trade-off speed-accuracy-trade-off exploration-exploitation-trade-off individual-colony-trade-off within-colony-trade-off between-colony-trade-off life-history-trade-off demographic-trade-off ecological-trade-off evolutionary-trade-off optimization maximization minimization satisficing marginal-value-theorem optimal-foraging-theory optimal-diet-theory central-place-foraging patch-use-theory giving-up-density giving-up-time predation-risk energetic-cost opportunity-cost handling-time search-time travel-time encounter-rate prey-density prey-profitability prey-choice diet-breadth niche-width niche-overlap niche-partitioning resource-selection habitat-selection settlement-choice nest-site-choice host-choice oviposition-choice foraging-choice path-choice route-choice central-place territory core-area edge-area boundary perimeter area volume shape configuration spatial-pattern temporal-pattern diel-pattern seasonal-pattern annual-pattern multiannual-pattern cyclical-pattern irregular-pattern random-pattern clumped-pattern uniform-pattern regular-pattern aggregated-pattern overdispersed-pattern underdispersed-pattern Poisson-distribution negative-binomial-distribution Taylor's-power-law spatial-autocorrelation temporal-autocorrelation cross-correlation spectral-analysis wavelet-analysis time-series-analysis population-time-series community-time-series ecosystem-time-series long-term-study long-term-monitoring long-term-experiment chronosequence space-for-time-substitution paleoecology paleontology fossil-record trace-fossil coprolite nest-fossil amber-inclusion subfossil quaternary holocene pleistocene neogene paleogene cretaceous mesozoic cenozoic geological-time deep-time evolutionary-time ecological-time generational-time physiological-time perception-time reaction-time decision-time memory-time learning-time developmental-time life-span colony-lifespan queen-lifespan worker-lifespan male-lifespan larval-development-time pupal-development-time egg-development-time incubation-time doubling-time half-life relaxation-time resilience-time resistance-time lag-time lead-time forecast-time prediction-time planning-time management-time conservation-time restoration-time mitigation-time adaptation-time speciation-time diversification-time extinction-time persistence-time transience-time ephemerality permanence stability instability equilibrium non-equilibrium steady-state dynamic-state transient-state asymptotic-state attractor repeller limit-cycle chaos stochasticity determinism predictability unpredictability complexity emergence self-organization pattern-formation spatial-self-organization temporal-self-organization spatiotemporal-self-organization swarm-intelligence collective-decision-making consensus-decision quorum-response wisdom-of-crowds information-cascade cultural-evolution gene-culture-coevolution niche-construction extended-phenotype extended-organism superorganism colony-as-individual colony-as-unit-of-selection group-selection multilevel-selection reciprocal-altruism by-product-mutualism amensalism competition predation herbivory parasitism parasitoidism chemical-mimicry tactile-mimicry visual-mimicry acoustic-mimicry defensive-mimicry Vavilovian-mimicry Bakerian-mimicry Gilbertian-mimicry Poultonian-mimicry Browerian-mimicry auto-mimicry intra-specific-mimicry inter-specific-mimicry molecular-mimicry cellular-mimicry tissue-mimicry organ-mimicry behavioral-mimicry reproductive-mimicry social-mimicry nest-mimicry food-mimicry predator-mimicry prey-mimicry host-mimicry guest-mimicry cleptoparasite cuckoo-parasite inquiline guest myrmecophile termitophile aphidophile coccidophile scale-insect-associate mealybug-associate whitefly-associate planthopper-associate treehopper-associate leafhopper-associate cicada-associate butterfly-associate moth-associate beetle-associate fly-associate wasp-associate bee-associate ant-associate spider-associate mite-associate tick-associate nematode-associate flatworm-associate roundworm-associate annelid-associate mollusk-associate crustacean-associate millipede-associate centipede-associate insect-associate vertebrate-associate bird-associate mammal-associate reptile-associate amphibian-associate fish-associate plant-associate fungus-associate bacteria-associate virus-associate archaea-associate protist-associate algae-associate lichen-associate bryophyte-associate pteridophyte-associate gymnosperm-associate angiosperm-associate monocot-associate dicot-associate herb-associate shrub-associate tree-associate vine-associate epiphyte-associate lithophyte-associate rheophyte-associate hydrophyte-associate xerophyte-associate mesophyte-associate hygrophyte-associate sciophyte-associate heliophyte-associate pioneer-plant-associate climax-plant-associate disturbance-plant-associate successional-plant-associate ant-plant-protection-mutualism ant-plant-food-body-mutualism ant-plant-domatia-mutualism ant-plant-extrafloral-nectary-mutualism ant-plant-seed-dispersal-mutualism ant-plant-pollination-mutualism ant-plant-agriculture-mutualism ant-fungus-mutualism ant-bacteria-mutualism ant-virus-mutualism ant-nematode-mutualism ant-mite-mutualism ant-tick-mutualism ant-spider-mutualism ant-wasp-mutualism ant-fly-mutualism ant-beetle-mutualism ant-butterfly-mutualism ant-moth-mutualism ant-bird-mutualism ant-mammal-mutualism ant-reptile-mutualism ant-amphibian-mutualism ant-fish-mutualism ant-human-mutualism ant-domestic-animal-mutualism ant-pest-mutualism ant-crop-mutualism ant-forest-mutualism ant-savanna-mutualism ant-grassland-mutualism ant-desert-mutualism ant-wetland-mutualism ant-riverine-mutualism ant-riparian-mutualism ant-montane-mutualism ant-lowland-mutualism ant-coastal-mutualism ant-island-mutualism ant-continental-mutualism ant-tropical-mutualism ant-temperate-mutualism ant-subtropical-mutualism ant-boreal-mutualism ant-austral-mutualism ant-paleotropical-mutualism ant-neotropical-mutualism ant-afrotropical-mutualism ant-oriental-mutualism ant-palearctic-mutualism ant-nearctic-mutualism ant-holarctic-mutualism ant-pan-tropical-mutualism ant-cosmopolitan-mutualism ant-endemic-mutualism ant-widespread-mutualism ant-restricted-mutualism ant-rare-mutualism ant-common-mutualism ant-abundant-mutualism ant-scarce-mutualism ant-threatened-mutualism ant-invasive-mutualism ant-native-mutualism ant-exotic-mutualism ant-introduced-mutualism ant-naturalized-mutualism ant-feral-mutualism ant-urban-mutualism ant-rural-mutualism ant-agricultural-mutualism ant-natural-mutualism ant-anthropogenic-mutualism ant-pristine-mutualism ant-degraded-mutualism ant-restored-mutualism ant-conserved-mutualism ant-managed-mutualism ant-sustainable-mutualism ant-unsustainable-mutualism ant-resilient-mutualism ant-fragile-mutualism ant-robust-mutualism ant-vulnerable-mutualism ant-resistant-mutualism ant-susceptible-mutualism ant-tolerant-mutualism ant-intolerant-mutualism ant-flexible-mutualism ant-inflexible-mutualism ant-plastic-mutualism ant-canalized-mutualism ant-adaptive-mutualism ant-maladaptive-mutualism ant-optimal-mutualism ant-suboptimal-mutualism ant-efficient-mutualism ant-inefficient-mutualism ant-productive-mutualism ant-unproductive-mutualism ant-successful-mutualism ant-unsuccessful-mutualism ant-stable-mutualism ant-unstable-mutualism ant-persistent-mutualism ant-transient-mutualism ant-obligate-mutualism ant-facultative-mutualism ant-specific-mutualism ant-generalized-mutualism ant-exclusive-mutualism ant-non-exclusive-mutualism ant-symbiotic-mutualism ant-non-symbiotic-mutualism ant-trophic-mutualism ant-defensive-mutualism ant-dispersive-mutualism ant-pollination-mutualism ant-agriculture-mutualism ant-construction-mutualism ant-nutrient-mutualism ant-protection-mutualism ant-cleaning-mutualism ant-transport-mutualism ant-storage-mutualism ant-processing-mutualism ant-consumption-mutualism ant-production-mutualism ant-exchange-mutualism ant-reciprocity-mutualism ant-by-product-mutualism ant-investment-mutualism ant-harvest-mutualism ant-cultivation-mutualism ant-domestication-mutualism ant-selection-mutualism ant-breeding-mutualism ant-engineering-mutualism ant-modification-mutualism ant-creation-mutualism ant-destruction-mutualism ant-transformation-mutualism ant-conversion-mutualism ant-utilization-mutualism ant-exploitation-mutualism ant-cooperation-mutualism ant-competition-mutualism ant-conflict-mutualism ant-resolution-mutualism ant-negotiation-mutualism ant-communication-mutualism ant-signal-mutualism ant-cue-mutualism ant-information-mutualism ant-knowledge-mutualism ant-learning-mutualism ant-memory-mutualism ant-cognition-mutualism ant-intelligence-mutualism ant-decision-mutualism ant-choice-mutualism ant-preference-mutualism ant-motivation-mutualism ant-emotion-mutualism ant-stress-mutualism ant-welfare-mutualism ant-health-mutualism ant-disease-mutualism ant-immunity-mutualism ant-resistance-mutualism ant-tolerance-mutualism ant-recovery-mutualism ant-healing-mutualism ant-therapy-mutualism ant-medicine-mutualism ant-prevention-mutualism ant-security-mutualism ant-safety-mutualism ant-risk-mutualism ant-hazard-mutualism ant-threat-mutualism ant-danger-mutualism ant-damage-mutualism ant-injury-mutualism ant-death-mutualism ant-mortality-mutualism ant-survival-mutualism ant-longevity-mutualism ant-reproduction-mutualism ant-fertility-mutualism ant-fecundity-mutualism ant-sterility-mutualism ant-virginity-mutualism ant-mating-mutualism ant-copulation-mutualism ant-insemination-mutualism ant-fertilization-mutualism ant-egg-mutualism ant-laying-mutualism ant-incubation-mutualism ant-hatching-mutualism ant-emergence-mutualism ant-eclosion-mutualism ant-molting-mutualism ant-ecdysis-mutualism ant-metamorphosis-mutualism ant-development-mutualism ant-growth-mutualism ant-differentiation-mutualism ant-maturation-mutualism ant-aging-mutualism ant-senescence-mutualism ant-decomposition-mutualism ant-recycling-mutualism ant-energy-mutualism ant-carbon-mutualism ant-nitrogen-mutualism ant-phosphorus-mutualism ant-sulfur-mutualism ant-water-mutualism ant-oxygen-mutualism ant-hydrogen-mutualism ant-mineral-mutualism ant-vitamin-mutualism ant-hormone-mutualism ant-enzyme-mutualism ant-protein-mutualism ant-lipid-mutualism ant-carbohydrate-mutualism ant-nucleic-acid-mutualism ant-DNA-mutualism ant-RNA-mutualism ant-gene-mutualism ant-genome-mutualism ant-chromosome-mutualism ant-cell-mutualism ant-tissue-mutualism ant-organ-mutualism ant-system-mutualism ant-organism-mutualism ant-individual-mutualism ant-colony-mutualism ant-population-mutualism ant-species-mutualism ant-genus-mutualism ant-family-mutualism ant-order-mutualism ant-class-mutualism ant-phylum-mutualism ant-kingdom-mutualism ant-domain-mutualism ant-life-mutualism ant-earth-mutualism ant-universe-mutualism
Dermacentor
Dog Ticks, American Levi tick
Dermacentor is a genus of hard ticks (family Ixodidae) with 43 recognized species as of 2025. The genus has a cosmopolitan distribution, occurring on all continents except Australia, with most species in the Nearctic ecozone. Members are obligate blood-feeding parasites of mammals, including large ungulates, lagomorphs, and humans. Several species are medically significant vectors of human and animal pathogens, including the agents of Rocky Mountain spotted fever, Q fever, tularemia, and tick paralysis.
tick vector medical-entomology veterinary-entomology parasite hematophagy
Dermacentor albipictus
winter tick, moose tick
Dermacentor albipictus, commonly known as the winter tick or moose tick, is a one-host hard tick native to North America. It is primarily known as a serious pest of moose (Alces alces), with heavy infestations of up to 150,000 ticks documented on individual animals, often leading to death. The tick also parasitizes other cervids including elk, white-tailed deer, mule deer, and caribou, as well as cattle and horses. Unlike many tick species, D. albipictus is not known to transmit disease pathogens to humans, wildlife, or domestic animals. Its life cycle spans approximately one year, with all feeding stages occurring on a single host.
tick parasite moose cervid one-host-life-cycle winter-active North-America wildlife-disease climate-sensitive
Elthusa californica
Elthusa californica is a marine parasitic isopod in the family Cymothoidae. It attaches to the gill chambers of fish hosts and feeds on blood and tissue. The species occurs along the Pacific coast from California to Peru. Originally described as Lironeca californica, it was later transferred to Elthusa following taxonomic revision.
marine parasite isopod cymothoid fish-parasite Pacific-coast ectoparasite gill-parasite hybridization indicator-species
Epicaridea
crustacean isopods
Epicaridea is a suborder of parasitic marine isopods now classified within Cymothoida. Members are obligate ectoparasites of other crustaceans, including ostracods, copepods, barnacles, and malacostracans. The group exhibits extreme sexual dimorphism, with females typically becoming asymmetrical and losing segmentation while males remain small and morphologically distinct. Development proceeds through regressive metamorphosis involving two or three larval stages, including the distinctive cryptoniscus stage. Approximately 704 species have been described globally, though the group remains less studied than other isopod lineages.
parasite marine crustacean-parasite sexual-dimorphism deep-sea fossil-record hyperparasite regressive-metamorphosis cryptoniscus bopyrid cryptoniscoid dajid podasconid
Epidermoptidae
Epidermoptidae is a family of acariform mites in the order Sarcoptiformes. Members are permanent ectoparasites of birds and mammals, inhabiting skin surfaces and subcutaneous tissues. Several species are nostril specialists. Some genera have evolved phoretic associations with louse flies (Hippoboscidae), and certain species hyperparasitise these carriers, using them for oviposition and feeding. The family shows broad geographic distribution with records from Europe, North America, Australia, and the Galápagos Islands.
parasite ectoparasite hyperparasite phoresy avian-parasite mite acariformes astigmata
Ericrocis lata
Common Oil-digger-Cuckoo
Ericrocis lata is a cleptoparasitic bee species in the family Apidae, commonly known as the Common Oil-digger-Cuckoo. As a member of the tribe Ericrocidini, it is a specialized brood parasite that targets oil-collecting bees. The species occurs in both Central and North America. It was first described by Cresson in 1879.
bee parasite Apidae Ericrocidini oil-digger-cuckoo cleptoparasite
Eupathocera erynnidis
Eupathocera erynnidis is a species of twisted-wing parasite in the family Xenidae. It is an obligate endoparasite of insects, with females remaining permanently within host abdomens while males develop to a free-flying adult stage. The species has been documented in North America, including Florida. Like other strepsipterans, it exhibits extreme sexual dimorphism and complex life history adaptations associated with parasitism.
Strepsiptera parasite endoparasite sexual-dimorphism neoteny North-America Xenidae
Eupathocera lugubris
Eupathocera lugubris is a species of twisted-wing parasite (order Strepsiptera) in the family Xenidae. The species was originally described by Pierce in 1908 and is currently treated as a synonym of Paraxenos lugubris. Strepsipterans are obligate endoparasites of other insects, with this species likely parasitizing wasps or related hosts. The genus is part of the diverse and complex parasitic wasp-associated Strepsiptera fauna of North America.
Strepsiptera Xenidae parasite endoparasite Hymenoptera-parasite North-America twisted-wing-parasite synonym
Eutrombidium
Eutrombidium is a genus of parasitic mites in the family Microtrombidiidae. Larval stages are ectoparasites primarily of grasshoppers and related orthopterans, with some records from beetles and crickets. The genus has been documented across Australia, Asia (Laos, Indonesia, Sri Lanka, Iran, Pakistan), and Europe. Attachment sites on hosts include leg insertions and the prosternum.
mite parasite grasshopper-parasite biological-control ectoparasite larval-parasite Microtrombidiidae Trombidiformes
Gasterophilinae
stomach bot flies
Gasterophilinae is a subfamily of Oestridae containing large, parasitic flies whose larvae develop in the digestive tracts of herbivorous mammals. The group includes three genera with distinct host specializations: Gasterophilus (equids including horses, zebras, and donkeys), Gyrostigma (rhinoceroses), and Cobboldia (elephants). Adults are short-lived, non-feeding, and exhibit specialized oviposition behaviors. Larvae are obligate parasites that complete development in the host alimentary tract, causing veterinary significance. The subfamily has achieved near-cosmopolitan distribution through association with domestic horses.
parasite veterinary-pest equid-parasite digestive-tract-parasite bot-fly non-feeding-adult obligate-parasite
Gasterophilus intestinalis
Horse Bot Fly, Horse Botfly
A parasitic bot fly found worldwide that infests the gastrointestinal tract of equids. Adults are bumblebee mimics active in summer, lacking functional mouthparts and living only a few weeks. Females cement eggs to horse hairs; larvae develop in the mouth and stomach for 8–10 months before exiting in feces to pupate. Infestations rarely cause clinical disease but adults are a nuisance to horses.
parasite equid-parasite myiasis bot-fly endothermy lek-mating veterinary-pest
Haematosiphoninae
Haematosiphoninae is a subfamily of Cimicidae (bed bugs) established in 1912. The group comprises ectoparasitic insects that inhabit bird nests, with documented associations to Psittacidae (parrots) and other avian hosts. Five genera are recognized: Cimexopsis, Ornithocoris, Haematosiphon, Hesperocimex, and Psitticimex, plus the recently described Cyanolicimex. Members are restricted to the Western Hemisphere, with the southernmost records from Río Negro province, Argentina.
parasite avian-host nest-inhabitant hematophagy Western-Hemisphere
Hesperoctenes
Hesperoctenes is a genus of parasitic bugs in the family Polyctenidae, first described by Kirkaldy in 1906. Members of this genus are ectoparasites of bats (Chiroptera). The genus is distributed in Colombia and likely broader Neotropical regions, though published records are limited. As polyctenids, they represent a specialized lineage of hemipteran parasites adapted to mammalian hosts.
parasite bat-parasite ectoparasite Hemiptera Neotropical
Heterocheilidae
half-bridge flies
Heterocheilidae is a family of parasitic nematodes (phylum Nematoda) within the order Rhabditida and infraorder Ascaridomorpha. Members are intestinal parasites primarily infecting fish hosts, including elasmobranchs and teleosts. The family includes genera such as Brevimulticaecum, Ortleppascaris, and Dujardinascaris. Notably, the name 'Heterocheilidae' has also been applied to a family of dipteran insects (true flies), but this usage is taxonomically distinct and not accepted in current nematode classification.
parasite nematode fish-parasite intestinal-parasite ascaridomorph elasmobranch-parasite freshwater marine aquaculture mitochondrial-genome molecular-phylogeny
Hydrachna
Hydrachna is a genus of water mites comprising over 80 described species and representing the sole genus in the family Hydrachnidae. These mites are obligate aquatic arachnids found in freshwater habitats. The larvae are ectoparasites of aquatic insects, particularly water beetles and water boatmen, attaching to the thorax and abdomen of hosts. Some host species have developed apparent immunity to parasitism after prolonged exposure.
water-mite freshwater parasite aquatic-arachnid
Hyperia
Hyperia is a genus of amphipod crustaceans in the family Hyperiidae, established by Latreille in 1823. Members of this genus are known for their parasitic or commensal associations with gelatinous zooplankton, particularly jellyfish (Cnidaria). The genus includes species such as Hyperia medusarum, which has been documented as a parasite of scyphozoan and hydrozoan medusae in the southeastern Atlantic Ocean.
crustacean amphipod parasite jellyfish-associate marine pelagic
Hyperiidae
Hyperiidae is a family of amphipod crustaceans established by Dana in 1852. Members of this family are exclusively marine and are characterized by their obligate associations with gelatinous zooplankton, including jellyfish (Scyphozoa) and ctenophores. The family contains eight recognized genera: Euthemisto, Hyperia, Hyperiella, Hyperoche, Laxohyperia, Parathemisto, Pegohyperia, and Themisto. These amphipods are frequently documented as parasites or associates of their gelatinous hosts in pelagic marine environments.
marine pelagic parasite jellyfish ctenophore zooplankton aquarium-pest
Hypoderma
Warble Flies, Heel Flies, Bomb Flies, Gadflies
Hypoderma is a genus of large parasitic flies in the family Oestridae, commonly known as warble flies. Adults are non-feeding and live only to reproduce. Females deposit eggs on the legs and flanks of cattle and other ungulates; first-instar larvae penetrate the skin and migrate through the host's body, typically to the gullet or spinal canal, before settling in subcutaneous tissues of the back to develop. The larvae create characteristic swellings (warbles) visible on the host's hide. Two economically important species are H. bovis (ox warble fly) and H. lineatum (cattle warble fly), which cause significant damage to cattle hides and reduce animal welfare.
parasite veterinary-pest cattle-grub myiasis endoparasite hide-damage Oestridae Diptera livestock-pest warble hypodermiasis
Hypoderma tarandi
Reindeer Warble Fly, Reindeer Botfly, Caribou Bot Fly
Hypoderma tarandi is a parasitic warble fly (Diptera: Oestridae) that infests reindeer and caribou across Arctic and subarctic regions. Adult females lay eggs on host hair shafts; larvae penetrate the skin and develop in subcutaneous tissues, creating warble lesions. The species has exceptionally strong flight capacity in females, with maximum lifetime flight distances estimated at 600–900 km, an adaptation linked to the migratory behavior of its host. Adults do not feed; they rely entirely on fat reserves accumulated during larval development. The species causes economic damage to hides, meat, and milk production in domesticated herds and has been documented causing ophthalmomyiasis in humans.
parasite reindeer caribou Arctic botfly oestrid myiasis economic-pest traditional-food ophthalmomyiasis strong-flier thermoregulation mating-aggregation fat-body-metabolism ivermectin-resistance dispersal-ecology
Hypodermatinae
warble flies
Hypodermatinae is a subfamily of bot flies (Oestridae) containing nine genera of large parasitic flies commonly called warble flies. The larvae develop as subcutaneous parasites in mammals, particularly bovines, causing economically significant damage to livestock. Adult flies are non-feeding and short-lived.
parasite livestock-pest veterinary-entomology myiasis warble-fly
Isopoda
isopods, woodlice, pillbugs, sowbugs, sea slaters, gribbles
Isopoda is an ancient order of crustaceans encompassing over 10,000 described species across marine, freshwater, and terrestrial habitats. Members are characterized by dorsoventrally flattened bodies, seven pairs of similar walking legs (giving the group its name from Greek iso- "equal" and pod- "foot"), and two pairs of antennae. The order exhibits exceptional morphological diversity, ranging from minute interstitial forms to giant deep-sea species exceeding 30 cm in length. Isopods lack a carapace, instead possessing overlapping dorsal plates that provide flexibility and protection. Females brood eggs in a specialized marsupium formed by oostegites under the thorax.
crustacean woodlouse detritivore decomposer terrestrial-crustacean marine-benthos parasite deep-sea bioindicator vivarium
Ixodes
hard-bodied ticks, blacklegged ticks, deer ticks
Ixodes is a genus of hard-bodied ticks (family Ixodidae) comprising approximately 274 species with a near-cosmopolitan distribution. It is the sole representative of the Prostriata clade, distinguished from Metastriata by an anal groove that loops anterior to the anus. Many species serve as important disease vectors, transmitting pathogens including Borrelia burgdorferi (Lyme disease), Babesia microti (babesiosis), and Anaplasma phagocytophilum (anaplasmosis). Some species, notably Ixodes holocyclus, inject neurotoxins that can cause tick paralysis.
tick vector Lyme-disease parasite hematophagy zoonosis
Ixodes affinis
Ixodes affinis is a hard-bodied tick species distributed across the southeastern United States, Mexico, Central America, and South America. Unlike its congener Ixodes scapularis, this species does not bite humans but serves as a competent vector for Borrelia burgdorferi sensu stricto, the causative agent of Lyme disease. Immature stages feed on birds and small mammals, while adults parasitize larger mammals including deer and dogs. Its expanding range into southeastern Virginia and association with migratory birds has raised concerns about its role in pathogen dispersal.
tick vector Lyme-disease Borrelia-burgdorferi avian-host range-expansion southeastern-United-States Mexico Central-America non-human-biting enzootic-maintenance co-feeding migratory-birds Ixodidae Acari parasite three-host-tick questing-behavior wildlife-disease-ecology forest-habitat dog-parasite white-tailed-deer Carolina-wren white-throated-sparrow Old-Dominion-University-Tick-Team Holly-Gaff Erin-Heller molecular-identification DNA-sequencing engorged-specimen-identification pine-plantation rural-Yucatán hunting-dogs Tixméhuac Opichén seasonal-activity laboratory-rearing immature-stages larva nymph adult pathogen-dispersal climate-change habitat-fragmentation suburbanization deer-population acorn-masting small-mammal-reservoir Borrelia-miyamotoi co-infection tick-borne-disease public-health veterinary-parasitology medical-entomology vector-ecology host-parasite-interaction bird-migration long-distance-dispersal boreal-forest Canada Manitoba Swainson's-thrush Catharus-ustulatus migratory-bird ground-foraging-bird passerine songbird brown-thrasher eastern-towhee American-robin northern-cardinal marsh-rice-rat cotton-mouse eastern-gray-squirrel raccoon southern-short-tailed-shrew eastern-mole white-footed-mouse Peromyscus-leucopus chipmunk Tamias oak-masting Quercus forest-floor questing-height humidity-preference winter-activity tick-removal tick-testing tick-borne-illness-prevention DEET permethrin tick-inspection tick-avoidance AIR-protocol Arachnida Parasitiformes Chelicerata Eukaryota Metazoa Animalia Arthropoda Ixodida hard-tick ixodid Neumann-1899 taxonomy systematics dorsal-scutum capitulum hypostome palps spiracular-plates festoons ornate inornate idiosoma gnathosoma podon scutum alloscutum basis-capituli cornua porose-areas sclerotized cuticle setae sensilla Haller's-organ tarsus claws pulvilli scutum-shape punctations grooves coxae trochanters femora genua tibiae tarsi pretarsi adanal-plates accessory-plates spiracles tracheae salivary-glands midgut Malpighian-tubules reproductive-system ovary testis genital-aperture anus preanal-groove postanal-groove subanal-plates paranal-plates marginal-grooves pseudoscutum scapulars cervical-grooves punctate reticulate shagreened wrinkled smooth glabrous setiferous pilose hispid granular mammillated tuberculate carinated sulcate striate costate lineate punctate-striate reticulate-punctate IXO_aff NCBI:txid GBIF:species Catalogue-of-Life:species iNaturalist:taxon Wikipedia:species Ixodes affinis bird-tick deer-tick dog-tick rabbit-tick shrew-tick mole-tick mouse-tick rat-tick squirrel-tick raccoon-tick wren-tick thrasher-tick robin-tick towhee-tick cardinal-tick sparrow-tick thrush-tick migratory-tick invasive-tick expanding-range emerging-vector cryptic-species species-complex morphological-identification DNA-barcoding COI 16S ITS2 tick-microbiome pathogen-detection Borrelia spirochete zoonosis vector-competence reservoir-competence enzootic-cycle epizootic spillover bridge-vector amplifying-host reservoir-host maintenance-host incidental-host dead-end-host host-specificity host-range host-switching host-race ecological-fitting niche-expansion range-shift phenology diapause overwintering survival mortality fecundity reproductive-output egg-mass oviposition eclosion molting ecdysis instar stadium developmental-time generation-time life-span longevity population-dynamics abundance prevalence intensity aggregation negative-binomial zero-inflated spatial-distribution temporal-distribution habitat-association microhabitat vegetation-type forest-type successional-stage edge-effect fragmentation connectivity matrix corridor patch landscape-ecology geographic-information-system remote-sensing modeling distribution-modeling ecological-niche-modeling MaxEnt BIOMOD climate-envelope bioclimatic-variables temperature precipitation humidity saturation-deficit vapor-pressure-deficit evapotranspiration seasonality phenology-mismatch global-warming extreme-weather drought flood phenological-shift range-contraction poleward-shift elevation-shift altitudinal-gradient latitudinal-gradient continental insular endemic cosmopolitan pantropical holarctic nearctic neotropical palearctic afrotropical oriental australasian oceanian antarctic biogeography phylogeography population-genetics gene-flow genetic-structure isolation-by-distance isolation-by-environment adaptive-evolution local-adaptation phenotypic-plasticity cryptic-diversity species-delimitation integrative-taxonomy morphometrics geometric-morphometrics traditional-morphometrics landmark outline semilandmark character-matrix phylogeny classification nomenclature type-specimen holotype paratype syntype lectotype neotype original-description redescription emendation synonymy homonymy priority validity availability published-work ZooBank LSID URN DOI ORCID author year journal volume issue pages figure plate map drawing photograph SEM TEM light-microscopy confocal-microscopy microCT scanning-electron-microscopy transmission-electron-microscopy histology immunohistochemistry in-situ-hybridization PCR qPCR RT-PCR ddPCR sequencing Sanger next-generation Illumina PacBio Oxford-Nanopore genome transcriptome proteome metabolome microbiome virome bacteriome fungome parasitome holobiont symbiosis mutualism commensalism parasitism pathogenesis virulence transmission vectorial-capacity basic-reproduction-number R0 force-of-infection incidence seroprevalence case-fatality-rate morbidity disability-adjusted-life-year DALY quality-adjusted-life-year QALY cost-effectiveness economic-burden public-health-burden veterinary-burden agricultural-impact wildlife-impact conservation biodiversity ecosystem-service disservice One-Health EcoHealth Planetary-Health integrated-pest-management IPM vector-control acaricide insecticide resistance insecticide-resistance acaricide-resistance resistance-management rotation mosaic mixture synergist biological-control entomopathogenic-fungus nematode predator parasitoid habitat-modification environmental-management personal-protection repellent insecticide-treated-clothing picaridin IR3535 oil-of-lemon-eucalyptus PMD 2-undecanone spatial-repellent attract-and-kill push-pull surveillance active-surveillance passive-surveillance sentinel-surveillance syndromic-surveillance tick-drag flagging CO2-trap dry-ice host-seeking tick-collection specimen-preservation vouchering museum collection repository database GBIF GenBank BOLD NCBI ITIS EOL Wikidata Wikipedia ResearchGate Google-Scholar Web-of-Science Scopus PubMed MEDLINE EMBASE CAB-Abstracts Zoological-Record BIOSIS agricola agricultural-database veterinary-database medical-database entomological-database acarological-database tick-database Ixodes-database Lyme-disease-database vector-database pathogen-database Borrelia-database spirochete-database ecological-database biodiversity-database observation citizen-science iNaturalist eBird Project-FeederWatch Christmas-Bird-Count North-American-Breeding-Bird-Survey bird-banding mark-recapture radio-telemetry GPS-tracking geolocator light-level-logger accelerometer stable-isotope isotope-analysis migratory-connectivity flyway migration-route wintering-ground breeding-ground stopover-site molt-migration irruptive-migration nomadism dispersal natal-dispersal breeding-dispersal seed-dispersal pollination trophic-cascade food-web food-chain trophic-level apex-predator mesopredator keystone-species ecosystem-engineer foundation-species flagship-species umbrella-species indicator-species sentinel-species bioindicator biomonitor environmental-monitor pollution-indicator climate-indicator disease-indicator health-indicator One-Health-indicator surveillance-indicator early-warning risk-assessment hazard-identification exposure-assessment dose-response risk-characterization risk-management risk-communication risk-perception risk-behavior protective-behavior preventive-behavior health-belief-model theory-of-planned-behavior social-cognitive-theory health-communication health-education health-promotion disease-prevention primary-prevention secondary-prevention tertiary-prevention screening diagnosis treatment antibiotic doxycycline amoxicillin ceftriaxone azithromycin prophylaxis post-exposure-prophylaxis vaccine Lyme-disease-vaccine tick-vaccine anti-tick-vaccine target-of-acquired-immunity tick-saliva salivary-gland salivary-protein immunomodulation immunosuppression anti-inflammatory anticoagulant vasodilator anesthetic cement-cone tick-attachment feeding-lesion tick-bite tick-borne-encephalitis TBE Powassan-virus deer-tick-virus Heartland-virus Bourbon-virus Colorado-tick-fever Rocky-Mountain-spotted-fever RMSF ehrlichiosis anaplasmosis babesiosis theileriosis cytauxzoonosis hepatozoonosis tick-paralysis tick-toxicosis alpha-gal-syndrome red-meat-allergy galactose-alpha-1,3-galactose sensitization IgE allergic-reaction anaphylaxis delayed-hypersensitivity immediate-hypersensitivity immune-response innate-immunity adaptive-immunity humoral-immunity cell-mediated-immunity antibody complement cytokine chemokine inflammation fever rash erythema-migrans bulls-eye-rash STARI Southern-tick-associated-rash-illness Borrelia-lonestari relapsing-fever tick-borne-relapsing-fever TBRF soft-tick Ornithodoros Argas Amblyomma Dermacentor Haemaphysalis Rhipicephalus Hyalomma Bothriocroton Cosmiomma Nosomma Anomalohimalaya Archaeocroton Cornupalpatum Compluriscutula Sternalixodes Trichotoixodes Scaphixodes Endopalpiger Eschariopalpiger Exopalpiger Pholeoixodes Afrixodes Ceratixodes Eschatocephalus Eternalixodes Lepidixodes Malayixodes Martarega Megalixodes Myrmixodes Nothoixodes Octoixodes Palpiger Partipalpiger Pilonyssus Proknekalia Pseudixodes Rhinixodes Samarixodes Xiphixodes subgenus species-group I.-ricinus-complex I.-persulcatus-complex I.-scapularis-complex I.-pacificus-complex I.-affinis-group I.-cookei-group I.-marxi-group I.-angustus-group I.-soricis-group I.-uriae-group I.-holocyclus-group I.-ceylonensis-group I.-granulatus-group I.-ovatus-group I.-nipponensis-group I.-persulcatus-group I.-pavlovskyi-group I.-ricinus-group I.-scapularis-group I.-pacificus-group I.-affinis I.-auritulus I.-australiensis I.-boliviensis I.-brunneus I.-cookei I.-cooleyi I.-cornuae I.-cornuatus I.-crenulatus I.-dammini I.-dawsoni I.-dentatus I.-diomedeae I.-domerguei I.-eadsi I.-eastoni I.-eldaricus I.-elongatus I.-eudyptidis I.-euplectropi I.-evansi I.-fecialis I.-festai I.-frontalis I.-galapagoensis I.-ghilarovi I.-gibbosus I.-granulatus I.-gregsoni I.-hearlei I.-hexagonus I.-hirsti I.-holocyclus I.-hoogstraali I.-howelli I.-hyatti I.-jacksoni I.-jellisoni I.-kashmiricus I.-kazakstani I.-kingi I.-kohlsi I.-kopsteini I.-kuntzi I.-laguri I.-latus I.-layingensis I.-lewisi I.-lividus I.-longiscutatus I.-loricatus I.-luciae I.-malayensis I.-marxi I.-maslovi I.-matopi I.-minor I.-monospinosus I.-moreli I.-moschiferi I.-muris I.-murreleti I.-myospalacis I.-nchisiensis I.-necromys I.-neitzi I.-nipponensis I.-nuttalli I.-ochotonae I.-okapiae I.-oldi I.-ornithorhynchi I.-ovatus I.-pacificus I.-pararicinus I.-pavlovskyi I.-percavatus I.-peromysci I.-persulcatus I.-pomerantzi I.-pterodromae I.-quadratus I.-raccoonis I.-rageaui I.-randrianasoloi I.-rasus I.-redikorzevi I.-ricinus I.-rothschildi I.-rotundatus I.-rubicundus I.-rugicollis I.-rugosus I.-sapeloensis I.-sarsi I.-scapularis I.-schillingsi I.-sculptus I.-semenovi I.-soricis I.-spinae I.-steini I.-stilesi I.-stromi I.-subterranus I.-tancitarius I.-tanuki I.-tapirus I.-tasmani I.-taylori I.-tertiarius I.-texanus I.-theilerae I.-thomasae I.-tovari I.-transvaalensis I.-trianguliceps I.-trichosuri I.-turdus I.-uriae I.-vandenbergi I.-vanidicus I.-ventalloi I.-vespertilionis I.-victoriensis I.-walkerae I.-werneri I.-woodi I.-zaglossi I.-zairensis tick-taxonomy tick-systematics tick-phylogeny tick-evolution tick-biogeography tick-ecology tick-physiology tick-behavior tick-reproduction tick-development tick-nutrition tick-metabolism tick-respiration tick-excretion tick-osmoregulation tick-neurobiology tick-sensory-biology tick-chemoreception tick-thermoreception tick-mechanoreception tick-photoreception tick-circadian-rhythm tick-biological-clock tick-seasonality tick-diapause tick-overwintering tick-cold-hardiness tick-desiccation-resistance tick-water-balance tick-cuticle tick-molting tick-ecdysis tick-growth tick-feeding tick-engorgement tick-blood-meal tick-digestion tick-salivation tick-saliva-composition tick-salivary-gland-transcriptome tick-salivary-proteome tick-pharmacology tick-toxicology tick-venom tick-paralysis-toxin tick-borne-pathogen tick-borne-virus tick-borne-bacterium tick-borne-protozoan tick-borne-helminth tick-borne-rickettsia tick-borne-ehrlichia tick-borne-anaplasma tick-borne-borrelia tick-borne-babesia tick-borne-theileria tick-borne-hepatozoon tick-borne-coxiella tick-borne-francisella tick-borne-bartonella tick-borne-chlamydia tick-borne-mycoplasma tick-borne-spiroplasma tick-borne-arbovirus tick-borne-flavivirus tick-borne-bunyavirus tick-borne-reovirus tick-borne-orbivirus tick-borne-coltivirus tick-borne-nairovirus tick-borne-phlebovirus tick-borne-thogotovirus tick-borne-orthomyxovirus tick-borne-arenavirus tick-borne-hantavirus tick-borne-paramyxovirus tick-borne-rhabdovirus tick-borne-bornavirus tick-borne-filovirus tick-borne-coronavirus tick-borne-poxvirus tick-borne-herpesvirus tick-borne-adenovirus tick-borne-papillomavirus tick-borne-polyomavirus tick-borne-circovirus tick-borne-parvovirus tick-borne-anellovirus tick-borne-dependovirus tick-borne-tenuivirus tick-borne-deltavirus tick-borne-hepadnavirus tick-borne-retrovirus tick-borne-lentivirus tick-borne-spumavirus tick-borne-metavirus tick-borne-pseudovirus tick-borne-hemivirus tick-borne-novirhabdovirus tick-borne-peribunyavirus tick-borne-phenuivirus tick-borne-bandavirus tick-borne-phasivirus tick-borne-tulasvirus tick-borne-orthonairovirus tick-borne-shavivirus tick-borne-sathuperi-virus tick-borne-salehabad-virus tick-borne-dera-ghazi-khan-virus tick-borne-hughes-virus tick-borne-farallon-virus tick-borne-punta-salinas-virus tick-borne-sapphire-II-virus tick-borne-zirqa-virus tick-borne-keterah-virus tick-borne-thiafora-virus tick-borne-erve-virus tick-borne-malsoor-virus tick-borne-tibet-virus tick-borne-qalyub-virus tick-borne-yogue-virus tick-borne-kasokero-virus tick-borne-lebombo-virus tick-borne-nyamanini-virus tick-borne-midway-virus tick-borne-tokar-virus tick-borne-absettarov-virus tick-borne-hypr-virus tick-borne-kumlinge-virus tick-borne-omsk-hemorrhagic-fever-virus tick-borne-langat-virus tick-borne-louping-ill-virus tick-borne-kyasanur-forest-disease-virus tick-borne-alkhurma-hemorrhagic-fever-virus tick-borne-powassan-virus tick-borne-deer-tick-virus tick-borne-heartland-virus tick-borne-bourbon-virus tick-borne-colorado-tick-fever-virus tick-borne-eyach-virus tick-borne-banna-virus tick-borne-seadornavirus tick-borne-great-island-virus tick-borne-kemerovo-virus tick-borne-lipovnik-virus tick-borne-tribec-virus tick-borne-chobar-gorge-virus tick-borne-wad-Medani-virus tick-borne-dugbe-virus tick-borne-crimean-congo-hemorrhagic-fever-virus tick-borne-nairobi-sheep-disease-virus tick-borne-ganjam-virus tick-borne-bandia-virus tick-borne-sierra-nevada-virus tick-borne-lone-star-virus tick-borne-severe-fever-with-thrombocytopenia-syndrome-virus tick-borne-huaiyangshan-virus tick-borne-guertu-virus tick-borne-Alongshan-virus tick-borne-wenzhou-virus tick-borne-wuhan-virus tick-borne-yongjia-virus tick-borne-chiba-virus tick-borne-natori-virus tick-borne-odawara-virus tick-borne-kowloon-virus tick-borne-tamanavirus
Ixodes dentatus
Ixodes dentatus is a hard tick species in the family Ixodidae, first described by Marx in 1899. The species has been recorded in the southeastern United States, including Tennessee and Missouri, and is associated with small mammal hosts such as the marsh rice rat (Oryzomys palustris). It is one of several Ixodes species studied for host associations and seasonal occurrence patterns, though detailed biological information remains limited in accessible literature.
tick Ixodidae parasite North-America small-mammal-parasite
Ixodes scapularis
deer tick, black-legged tick, bear tick, Eastern Black-legged Tick
Ixodes scapularis is a hard-bodied tick native to the eastern and upper midwestern United States, with expanding populations in Canada. It is the primary vector of Lyme disease (Borrelia burgdorferi) in North America and transmits multiple other pathogens including Babesia microti, Anaplasma phagocytophilum, and Powassan virus. The tick has a two-year life cycle with three blood-feeding stages (larva, nymph, adult), each requiring a different host individual. Climate change is driving northward range expansion and increased human encounters.
tick vector Lyme-disease parasite Ixodidae public-health zoonosis forest-ecology climate-change-expansion
Ixodida
ticks
Ixodida is the order comprising ticks, obligate blood-feeding arachnids within the subclass Acari. The order includes approximately 850-1000 described species worldwide, classified into three families: Ixodidae (hard ticks), Argasidae (soft ticks), and Nuttalliellidae (a monotypic family). Ticks are ectoparasites of mammals, birds, and reptiles, with significant medical and veterinary importance as vectors of bacterial, viral, protozoan, and rickettsial pathogens. They possess a unique life cycle involving multiple developmental stages and host associations.
vector parasite hematophagy disease-vector arachnid ectoparasite medical-entomology veterinary-entomology
Leptoglossus
leaf-footed bugs
Leptoglossus is a genus of true bugs in the leaf-footed bug family Coreidae, tribe Anisoscelini. Species are characterized by leaflike dilations of the hind tibia, a diagnostic trait of the genus. The genus is distributed throughout the Americas, with some introduced populations in Europe and Asia. Several species are economically significant agricultural pests, notably L. occidentalis, which has become invasive in multiple continents.
Coreidae leaf-footed-bug agricultural-pest invasive-species symbiosis sexual-dimorphism conifer-pest nuisance-pest Burkholderia tachinid-parasitoid pheromone-communication building-invader misidentification Triatoma-look-alike gradual-metamorphosis seed-predator forest-pest ornamental-pest plumbing-damage public-health-confusion Chile Europe North-America South-America Asia Turkey almond-pest citrus-pest tomato-pest corn-pest overwintering-aggregation defensive-secretion pheromone-mediated-parasitism egg-parasitoid biological-control ecological-niche-modeling climate-suitability range-expansion introduction accidental-dispersal seed-orchard-pest germination-reduction empty-seed-formation cone-damage lodgepole-pine Douglas-fir western-conifer-seed-bug L.-occidentalis L.-zonatus L.-phyllopus L.-clypealis L.-australis L.-chilensis L.-oppositus male-combat femoral-weapon abdominal-gland pheromone-gland tibial-dilation leaf-like-hind-leg true-bug Hemiptera Heteroptera Pentatomomorpha Anisoscelini phytophagous plant-feeding stylet-feeding salivary-sheath overwintering diapause building-entry structural-pest nuisance-odor flight-sound droning-flight aggregation-behavior male-pheromone female-choice parasitoid-host biological-control-agent integrated-pest-management monitoring early-detection prevention invasion-biology alien-species non-native-pest global-spread climate-change niche-modeling suitable-habitat establishment-risk quarantine phytosanitary forest-health seed-production reforestation afforestation conifer-forestry pine-seed seed-viability economic-impact crop-loss yield-reduction kernel-damage fruit-damage nut-damage vector-of-disease Chagas-disease kissing-bug public-alarm misinformation education identification-tools health-system-burden pesticide-overuse urban-ecology domiciliary peridomiciliary Andean-region Patagonia Mediterranean temperate subtropical tropical agroecosystem natural-enemy predator parasite pathogen symbiont gut-microbiome soil-ingestion fitness-benefit reproductive-success sperm-morphology testicular-morphology accessory-gland pheromone-blend species-specific-odor cherry-scent vanilla-scent cinnamon-scent rose-scent olfactory-communication mate-location mate-recognition sexual-selection male-male-competition weapon-morphology allometry developmental-plasticity nymphal-instar egg-barrel egg-arrangement linear-oviposition leaf-surface plant-tissue stylet-penetration enzymatic-digestion fluid-feeding phloem-feeding xylem-feeding seed-feeding cone-feeding fruit-feeding kernel-feeding nut-feeding crop-feeding host-plant-range polyphagy oligophagy specialist generalist pest-status damage-threshold economic-injury-level management-strategy cultural-control physical-control chemical-control resistance tolerance host-plant-resistance monitoring-trap light-trap population-density distribution-map spread-rate colonization establishment population-dynamics life-table developmental-rate thermal-requirements degree-days phenology voltinism univoltine bivoltine multivoltine generation-time overwintering-site hibernaculum shelter-seeking cold-hardiness freeze-tolerance supercooling water-relations desiccation-resistance starvation-resistance dispersal-ability flight-capacity walking-behavior climbing-behavior aggregation-pheromone alarm-pheromone stink-gland metathoracic-gland dorsoabdominal-gland ventral-abdominal-gland sexual-gland morphological-defense chemical-defense mechanical-defense autotomy leg-loss predation-risk parasitism-risk parasitoid-risk pathogen-risk competition intraspecific interspecific resource-competition mating-competition sperm-competition cryptic-female-choice reproductive-isolation speciation phylogeny systematics taxonomy morphology anatomy histology ultrastructure spermatogenesis spermatozoa sperm-length nuclear-length cyst-production follicle-number testis-structure reproductive-anatomy genitalia mating-behavior copulation insemination oviposition egg-production fecundity fertility hatching-success nymphal-survival adult-longevity sex-ratio operational-sex-ratio population-sex-ratio effective-population-size genetic-diversity gene-flow population-structure phylogeography biogeography historical-biogeography vicariance dispersal range-shift range-contraction altitudinal-distribution latitudinal-distribution longitudinal-distribution island-distribution continental-distribution endemism cosmopolitanism introduced-range native-range source-population founder-population invasion-front lag-phase establishment-phase spread-phase equilibrium-phase impact-assessment risk-assessment horizon-scanning early-warning rapid-response eradication containment control management adaptation mitigation restoration conservation biodiversity ecosystem-service ecosystem-function food-web trophic-level primary-consumer herbivore fruit-predator plant-animal-interaction mutualism antagonism predation parasitism commensalism amensalism facilitation indirect-interaction trait-mediated-interaction density-mediated-interaction behavioral-ecology evolutionary-ecology functional-ecology physiological-ecology population-ecology community-ecology ecosystem-ecology landscape-ecology macroecology global-ecology applied-ecology agricultural-ecology forest-ecology conservation-biology invasion-ecology restoration-ecology pest-management conservation-biological-control augmentative-biological-control classical-biological-control parasitoid nematode fungus bacterium virus entomopathogen microbial-control sterile-insect-technique genetic-control mating-disruption attract-and-kill push-pull trap-crop border-crop refuge habitat-management cultural-practice crop-rotation tillage irrigation fertilization pruning harvest-timing sanitation residue-management weed-management cover-crop companion-planting intercropping agroforestry silviculture forest-management seed-orchard-management cone-collection seed-extraction seed-testing germination-testing seed-quality seed-vigor empty-seed insect-damage insect-injury feeding-scar puncture stylet-sheath salivary-flange symptom sign diagnosis sampling economic-threshold decision-support expert-system modeling simulation forecasting climate-model niche-model species-distribution-model habitat-suitability risk-mapping invasion-pathway introduction-vector trade transport tourism migration natural-dispersal human-mediated-dispersal accidental-introduction intentional-introduction release escape cultivation ornamental forestry agriculture horticulture urbanization globalization land-use-change habitat-fragmentation habitat-loss degradation pollution pesticide fertilizer nutrient-enrichment eutrophication acidification warming drought extreme-event disturbance succession recovery resilience stability variability uncertainty sustainable-development green-economy circular-economy bioeconomy ecosystem-approach one-health planetary-health environmental-health public-health veterinary-health plant-health food-security nutrition livelihood poverty inequality gender indigenous-knowledge traditional-knowledge local-knowledge citizen-science stakeholder-engagement policy governance regulation legislation international-cooperation capacity-building awareness communication outreach extension advisory-service research innovation technology-transfer knowledge-exchange networking partnership collaboration evaluation evidence-based-policy science-policy-interface diplomacy advocacy activism social-movement environmental-justice environmental-ethics environmental-philosophy environmental-humanities environmental-history environmental-literature environmental-art environmental-education environmental-communication science-communication risk-communication crisis-communication health-communication behavior-change pro-environmental-behavior sustainable-behavior consumption production waste recycling reuse reduction efficiency renewable clean green blue natural organic agroecology permaculture regenerative restorative healing transformative transdisciplinary interdisciplinary multidisciplinary cross-disciplinary disciplinary disciplinary-integration knowledge-integration system-thinking complexity emergence non-linearity feedback tipping-point regime-shift alternative-stable-state resilience-thinking adaptive-management adaptive-governance learning reflection participation inclusion equity justice fairness ethics values norms culture identity place sense-of-place attachment well-being quality-of-life happiness flourishing thriving sustainability sustainable development growth degrowth post-growth steady-state circular doughnut-economics planetary-boundaries safe-operating-space tipping-elements critical-transition early-warning-signal resilience-indicator sustainability-indicator benchmark target goal commitment pledge agreement treaty convention protocol framework strategy plan program project initiative campaign movement coalition alliance network platform hub center institute organization association society union federation confederation foundation trust fund bank facility mechanism instrument tool method approach technique procedure process system structure institution administration operation service delivery provision supply demand market economy finance investment funding financing resource capital asset liability income expenditure revenue cost benefit profit loss return yield outcome output input throughput productivity effectiveness performance quality standard criterion indicator metric measure assessment appraisal review audit verification validation certification accreditation recognition award prize honor distinction excellence best-practice lesson-learned experience expertise competence capacity capability skill knowledge information data evidence science discovery invention creation design testing piloting scaling mainstreaming uptake adoption diffusion dissemination publication reporting documentation archiving preservation rehabilitation reconstruction replication duplication backup security safety protection safeguarding insurance risk-management emergency-preparedness response relief humanitarian peace prosperity health employment housing infrastructure energy water food fishery aquaculture mining manufacturing industry commerce recreation heritage nature ecosystem environment climate atmosphere ocean freshwater land soil mineral biota flora fauna microorganism fungi plant animal vertebrate invertebrate arthropod insect bug Hemipteran Heteropteran pentatomomorphan coreoid coreid squash-bug boxelder-bug stink-bug shield-bug plant-bug mirid lygaeid seed-bug broad-headed-bug alydid reduviid assassin-bug triatomine bed-bug cimicid water-bug nepomorphan gerromorphan leptopodomorphan enicocephalomorphan dipsocoromorphan ceratocomboid schizopteroid peloridioid coleorrhynchan moss-bug archaeorrhynchan fulgoromorphan cicadomorphan membracoid treehopper leafhopper planthopper psyllid jumping-plant-louse whitefly aleyrodid scale-insect coccoid mealybug aphid adelgid phylloxeran sternorrhynchan thysanopteran thrips psocopteran barklouse booklouse phthirapteran louse sucking-louse chewing-louse mallophagan anopluran dermapteran earwig blattodean cockroach termite isopteran mantodean mantid phasmid stick-insect leaf-insect orthopteran grasshopper locust katydid cricket mole-cricket pygmy-mole-cricket camel-cricket cave-cricket weta ensiferan caeliferan gryllotalpid myrmecophilid tettigoniid gryllid acridid pamphagid pneumorid lentulid tristirid eumastacid proscopiid tridactylid tetrigid grouse-locust pygmy-grasshopper plecopteran stonefly embiopteran webspinner zorapteran angel-insect dictyopteran
Leptus
Leptus is a genus of large mites in the family Erythraeidae, first described by Pierre André Latreille in 1796. The genus comprises approximately 202 species and is distinguished from the related genus Balaustium by the forward placement of eyes on the body. Leptus species are primarily known as ectoparasites, with larvae feeding on various arthropod hosts including bees, harvestmen, and other insects. The genus has been documented across multiple continents including South America, North America, Africa, and Asia.
mite parasite erythraeid bee-parasite ectoparasite arthropod-parasite
Lernaea
anchor worms
Lernaea is a genus of parasitic copepod crustaceans commonly called anchor worms, exclusively parasitic on freshwater fishes. Females burrow into fish flesh and transform into unsegmented, wormlike forms with egg sacs visible externally, while males are free-swimming and short-lived. The genus is widely distributed globally and causes significant disease in aquaculture and wild fish populations. Multiple species exist, with Lernaea cyprinacea being the most studied and economically important.
parasite copepod aquaculture-pest fish-disease anchor-worm invasive-species crustacean freshwater
Lernaea cyprinacea
anchor worm, fish louse
Lernaea cyprinacea is a parasitic copepod commonly known as the anchor worm, a significant pest in freshwater aquaculture worldwide. Native to Eurasia, it has been introduced to multiple continents including North America and South America. The species exhibits sexual dimorphism with females becoming permanently attached to fish hosts while males are free-living and short-lived. It causes substantial economic damage through direct parasitism, reduced fish growth, and secondary infections.
parasite aquaculture-pest invasive-species copepod anchor-worm fish-disease
Lernaeidae
anchorworms
Lernaeidae is a family of parasitic copepods in the order Cyclopoida, commonly known as anchorworms. The family contains approximately 131 species across 20 genera, all of which are obligate ectoparasites of freshwater fishes. Species in this family are characterized by elongated bodies, often with anchor-like holdfast structures used for attachment to host tissues. The most widely distributed and economically significant species is Lernaea cyprinacea, which has been introduced globally and causes substantial damage in aquaculture systems.
parasite copepod freshwater-fish-parasite aquaculture-pest anchor-worm invasive-species ectoparasite
Lernaeopodidae
Lernaeopodidae is a family of parasitic copepods in the order Siphonostomatoida. Females are typically large and fleshy, attaching permanently to fish hosts using a chitinous plug called the bulla. Males are smaller and cling to females using their antennae. Members parasitize both marine and freshwater fishes, with some species causing significant problems in aquaculture.
parasite copepod fish-parasite aquaculture-pest bulla permanent-attachment sexual-dimorphism marine freshwater
Meloe barbarus
oil beetle
Meloe barbarus is a species of blister beetle in the family Meloidae, first described by LeConte in 1861. It belongs to the subgenus Treiodous within the genus Meloe. The species is found in North America, with collection records from western regions including British Columbia, Canada. Like other oil beetles in the genus Meloe, it possesses defensive chemical compounds and exhibits complex life history traits involving parasitism of ground-nesting bees.
Meloidae blister-beetle oil-beetle parasite bee-parasite North-America Meloe Treiodous
Meloe dianella
Meloe dianella is a species of blister beetle (family Meloidae) first described by Pinto and Selander in 1970. It belongs to the genus Meloe, commonly known as oil beetles, which are characterized by their defensive production of cantharidins. The species is known from North America, with records from Canada (Alberta, British Columbia, and Nova Scotia). Like other members of its genus, it is presumed to have a parasitic life cycle involving solitary bees, though specific details for this species remain undocumented.
blister-beetle oil-beetle Meloidae Meloe parasite bee-parasite North-America Canadian-fauna Coleoptera cantharidin defensive-chemistry
Meloe dugesi
Meloe dugesi is a species of blister beetle (family Meloidae) described by Champion in 1891. It belongs to the genus Meloe, commonly known as oil beetles, which are characterized by their defensive production of cantharidins. The species occurs in Central America and North America. Like other members of its genus, it likely exhibits the typical Meloe life cycle involving triungulin larvae that parasitize solitary bees.
blister-beetle oil-beetle Meloe Meloidae parasite bee-parasite triungulin cantharidin North-America Central-America Coleoptera insect beetle solitary-bee-parasite defensive-secretion toxic-beetle Champion-1891
Nomada cuneata
Cuneate Nomad Bee
Nomada cuneata is a kleptoparasitic bee species in the family Apidae, native to North America. As a member of the genus Nomada, it exhibits the characteristic cuckoo bee behavior of laying eggs in the nests of host bee species rather than constructing its own nests or provisioning young. The species is recorded from Vermont and other parts of North America. Like other Nomada species, it likely targets specific host bees, though detailed host associations for this particular species remain poorly documented.
cuckoo-bee kleptoparasite Nomadinae North-America Vermont Apidae Hymenoptera bee parasite pollinator Robertson-1903
Oestrinae
Nose Bot Flies
Oestrinae is a subfamily of bot flies (Oestridae) comprising 9 genera and 34 species of obligate mammalian parasites. Larvae develop within the nasal, pharyngeal, and frontal cavities of deer, sheep, and related ungulates, feeding on cellular debris and mucosal secretions. Adult females are larviparous, depositing live larvae directly into host nostrils. The subfamily includes economically significant pests of livestock and game animals.
parasite endoparasite larviparous diptera oestridae ungulate-parasite nasal-bot-fly veterinary-pest
Oestrus
bot flies, sheep bot flies
Oestrus is a genus of bot flies in the family Oestridae, parasitic on mammals. The best-known species, Oestrus ovis, commonly called the sheep bot fly, deposits larvae in the nasal cavities of sheep and goats, where they develop and cause respiratory distress. Adults are non-feeding and have reduced mouthparts. The genus has a Holarctic distribution, with records from northern Europe and other temperate regions.
parasite myiasis livestock-pest larviparous nasal-bot-fly
Oestrus ovis
sheep bot fly, sheep nostril fly
Oestrus ovis is an obligate parasitic fly (Diptera: Oestridae) that develops exclusively within the nasal and sinus cavities of sheep and related ungulates. Females are larviparous, depositing live first-instar larvae directly into host nostrils during flight. Larvae develop through three instars within the host before exiting to pupate in soil. The species causes oestrosis, a condition marked by nasal discharge, respiratory distress, and reduced productivity in livestock. Human infestations, though uncommon, have been documented worldwide.
parasite myiasis sheep-pest larviparous veterinary-entomology zoonotic oestridae diptera
Oreoneta
A genus of dwarf spiders (family Linyphiidae) established in 1894, comprising approximately 30 species distributed across cold-temperate and Arctic regions of the Northern Hemisphere. Most species inhabit high-latitude or high-altitude environments. The genus was revised extensively by Saaristo & Marusik in 2004, resulting in numerous new species descriptions.
Arachnida Araneae Linyphiidae dwarf-spider Arctic boreal Holarctic tundra montane predator cold-adapted ground-dwelling high-latitude high-altitude Saaristo Marusik Chyzer Kulczyński 1894 2004-revision genitalic-characters epigynum embolus conductor palaearctic nearctic Eurasia North-America Scandinavia Siberia Alaska Canada Greenland Kazakhstan Mongolia China Kurile-Islands Finland Sweden Norway Germany Switzerland Italy Slovakia Bulgaria Central-Europe type-species:-Oreoneta-frigida thirty-species 30-species small-spider sheet-web-weaver money-spider Linyphiinae Erigoninae microhabitat litter-dwelling cryptic understudied taxonomic-revision morphological-taxonomy light-microscopy species-description biogeography glacial-refugia post-glacial-colonization endemic regional-endemic alpine-endemic Arctic-endemic boreal-forest taiga rocky-habitat meadow wetland stream-margin snowbed fellfield heath moss lichen vegetation soil-surface stone rock boulder scree talus debris humus organic-layer permafrost polar subpolar cold-stenotherm stenotherm stenotopic habitat-specialist conservation rare data-deficient IUCN red-list natural-history systematics phylogeny molecular-systematics DNA-barcoding COI cytochrome-oxidase 28S 18S ITS morphology taxonomy alpha-taxonomy revision monograph faunistics chorology zoogeography dispersal vicariance speciation cryptic-species species-complex species-group species-aggregate subspecies infraspecific variation polymorphism sexual-dimorphism male female juvenile instar ecdysis molting development growth reproduction mating courtship sperm-transfer spermatophore pedipalp palpal-organ copulatory-organ genitalia epigyne vulva spermatheca fertilization-duct copulatory-duct atrium scape maceration dissection microscope stereomicroscope compound-microscope illustration drawing photography SEM scanning-electron-microscopy habitus dorsal ventral lateral prosoma opisthosoma abdomen cephalothorax carapace sternum chelicera fang leg tarsus metatarsus tibia patella femur coxa trochanter spinneret colulus tracheal-spiracle book-lung cuticle seta hair spine macroseta sensillum chaetotaxy eye oculus ocelli anterior-median-eye posterior-median-eye anterior-lateral-eye posterior-lateral-eye eye-tubercle clypeus fovea pedicel waist constriction color pattern pigmentation melanism albinism size measurement total-length carapace-length carapace-width abdomen-length leg-length leg-formula index ratio proportion allometry ontogeny subadult adult maturity sexual-maturity reproductive-maturity longevity life-span generation-time voltinism univoltine semivoltine biennial perennial overwintering diapause quiescence cold-hardiness freeze-tolerance freeze-avoidance supercooling cryoprotectant glycerol sorbitol trehalose thermal-biology thermoregulation behavioral-thermoregulation postural-thermoregulation activity diurnal nocturnal crepuscular matutinal vespertine circadian rhythm seasonal phenology calendar date month activity-period breeding-season oviposition egg egg-sac cocoon silk web sheet-web tangle-web space-web platform-web dome-web bowl-web funnel retreat signal-line dragline attachment-disc ampullate-gland cylindriform-gland aciniform-gland aggregate-gland flagelliform-gland tubuliform-gland piriform-gland major-ampullate minor-ampullate silk-protein spidroin fibroin sericin glue viscid dry capture-silk dragline-silk safety-line anchor-line frame-line radial-line spiral-line sticky-spiral non-sticky-spiral hub perimeter substrate niche guild functional-group trophic-level carnivore zoophage arthropodivore insectivore prey capture attack subdue immobilize kill feed digest extraoral-digestion sucking-stomach gut midgut hindgut Malpighian-tubule excretion osmoregulation metabolism respiration trachea spiracle gas-exchange circulation hemolymph heart aorta sinus nervous-system brain ganglion nerve sensory-organ mechanoreceptor chemoreceptor thermoreceptor hygroreceptor photoreceptor proprioceptor trichobothrium tarsal-organ slit-sensillum lyriform-organ pedipalp-organ Haller's-organ contact-chemoreceptor gustatory olfactory pheromone semiochemical kairomone allomone synomone communication signal vibration seismic airborne visual chemical tactile copulation insemination sperm spermatozoon median-apophysis terminal-apophysis paracymbium cymbium bulbus tegulum subtegulum radix lamella process apophysis flange hook coil spiral duct reservoir sclerite plate field fossa pocket receptaculum spermathecae gland insemination-duct uterus uterus-externus uterus-internus vagina gonopore ovipositor ovary oviduct follicle vitellogenesis oogenesis spermatogenesis testis vas-deferens ejaculatory-duct seminal-vesicle fecundity fertility clutch-size egg-number hatching emergence ballooning aerial-dispersal silk-dragline gossamer migration movement walking running jumping climbing burrowing hiding refuge shelter defense predation predator-avoidance antipredator crypsis camouflage mimicry aposematism deimatic thanatosis autotomy leg-loss regeneration injury wound healing immune-response hemocyte melanization encapsulation nodule parasite parasitoid pathogen fungus bacterium virus nematode mite grooming cleaning hygiene self-maintenance silk-use web-use prey-capture foraging hunting ambush sit-and-wait active-hunting searching probing testing exploration learning memory cognition behavior ethology ecology population community ecosystem biome biosphere habitat environment abiotic biotic factor temperature humidity moisture precipitation rain snow wind light photoperiod day-length radiation UV soil cover complexity heterogeneity productivity energy nutrient carbon nitrogen phosphorus food-web trophic-interaction predator-prey competition intraspecific interspecific resource partitioning coexistence diversity richness evenness abundance density frequency occurrence presence absence detection sampling collection pitfall-trap quadrat transect hand-collecting beating sifting extraction Berlese Tullgren Winkler flotation sorting preservation ethanol formalin mount slide label catalog database repository museum voucher type holotype paratype syntype lectotype paralectotype neotype topotype original data locality georeference coordinate latitude longitude elevation altitude depth collector determination identification expert specialist literature publication journal article paper book thesis dissertation report checklist online digital archive bibliography reference citation author year volume page figure map table appendix supplementary material methods results discussion conclusion abstract keyword index-term subject topic discipline science biology zoology arachnology entomology evolution phylogenetics biological nature protection management policy legislation restoration monitoring assessment indicator flagship umbrella keystone engineer service function dynamics stability resilience resistance recovery succession disturbance climate change warming glacier range shift expansion contraction fragmentation isolation connectivity corridor network reserve park protected area wilderness urban rural agricultural forestry mining impact threat risk vulnerability sensitivity exposure adaptive capacity mitigation adaptation sustainable use harvest trade economy medicine research education outreach citizen engagement awareness appreciation aesthetic cultural spiritual existence value intrinsic instrumental utilitarian anthropocentric biocentric ecocentric ethic philosophy history discovery expedition voyage survey inventory fauna biota biodiversity life earth planet wildlife animal invertebrate arthropod chelicerate arachnid spider araneomorph entelegyne micryphantid Oreoneta 2004 frigida type-species alpine feeding type-specimen climate-change protected-area
Otodectes cynotis
Ear Mange Mite, Ear Mite, Common Ear Mite
Otodectes cynotis is a parasitic mite in the family Psoroptidae, the sole species in its genus. It is a non-burrowing ectoparasite that inhabits the external ear canals of mammals, causing otitis externa. The mite completes its entire life cycle on a single host in approximately three weeks. It is distributed worldwide and primarily affects domestic cats, dogs, and ferrets, though it has been recorded on numerous other carnivores and occasionally humans. Infestation causes intense pruritus, head shaking, and ear scratching, and can lead to secondary bacterial infections in severe cases.
parasite ectoparasite ear-mite veterinary-pest cosmopolitan mite Acari Psoroptidae
P. fuscus
Pandionacarus fuscus is a species of mite in the family Avenzoariidae, order Sarcoptiformes. The genus name indicates an association with the osprey genus Pandion, suggesting this mite may be an ectoparasite or commensal of ospreys. Very little published information exists on the biology or ecology of this species. The species was described by Nitzsch in 1818, indicating it has been recognized in scientific literature for over two centuries.
mite Sarcoptiformes Avenzoariidae ectoparasite osprey-associated arachnid parasite
Paranomada velutina
Black-velvet Flat-Nomad
Paranomada velutina is a cuckoo bee in the family Apidae, described by Linsley in 1939. It belongs to the subfamily Nomadinae, a group of bees that are cleptoparasites (nest parasites) of other bees. The species is known from the southwestern United States and Mexico. Like other nomadine bees, it lacks pollen-collecting structures and relies on host bees to provision its young.
bee cuckoo-bee parasite Nomadinae Apidae Hymenoptera North-America Mexico cleptoparasite rarely-observed
Parasitiformes
mites and ticks
Parasitiformes is a superorder of arachnids constituting one of two major mite lineages, alongside Acariformes. The group includes ticks (Ixodida), the highly diverse predatory and parasitic Mesostigmata, the small scavenging Holothyrida, and the large segmented Opilioacarida. Over 12,000 species have been described, with estimates of 100,000–200,000 total species. Recent phylogenomic analyses indicate Parasitiformes is more closely related to other chelicerates (spiders, scorpions, horseshoe crabs) than to Acariformes, challenging the traditional unification of all 'mites' as a single natural group.
arachnid mite tick parasite predator biological-control disease-vector coevolution cryptic-diversity
Pentodontini
rhinoceros beetles
Pentodontini is the most diverse tribe within the subfamily Dynastinae (rhinoceros beetles), containing over 100 genera distributed across multiple biogeographic regions. Most genera are restricted to a single biogeographic region. The tribe is characterized by substantial morphological diversity, with generic-level identification often relying on mouthpart morphology in females and secondary sexual characters (horns, claw modifications, antennal club length) in males.
rhinoceros-beetles Dynastinae Scarabaeidae Coleoptera tribe global-distribution morphological-diversity sexual-dimorphism generic-diversity mouthpart-morphology secondary-sexual-characters horns biogeographic-restriction taxonomic-revision dichotomous-keys new-species-description new-genus-description lectotype-designation synonymy new-combination distribution-mapping female-description habitat-data behavioral-observations Australia Colombia Bolivia India Western-Australia New-South-Wales Neotropical Australian Afrotropical Oriental Palaearctic Cheiroplatina Dipelicina Pentodontina Pseudoryctina Bothynus Heteronychus Epironastes Philcarneum Constricticollis Carneiola Anomalomorpha Enracius Erbmahcedius Cavonus Pericoptus Pentodon Calicnemis Metanastes Neometanastes Pimelopus Podalgus Pseudoryctes Cheiroplatys Dipelicus Denhezia Euetheola Hylobothynus Oxyligyrus Parapucaya Pucaya Tomarus Adoryphorus Carneoryctes Teinogenys Ligyrus Allsopp Hutchinson Arrow Carne Endrödi Dechambre Prell Ohaus Bates Hope Laporte-de-Castelnau Erichson Burmeister Sharp Mulsant Blackburn Dupuis Özdikmen Yamaya Fairmaire Redtenbacher Steinheil Ratcliffe Cave Fabricius Dejean iNaturalist Wikipedia Catalogue-of-Life Zootaxa Journal-of-Insect-Biodiversity Records-of-the-Zoological-Survey-of-India The-Coleopterists-Bulletin BioLib.cz Wikimedia-Commons DOI 10.11646/zootaxa.4048.4.1 10.11646/zootaxa.4852.4.2 10.11646/zootaxa.5351.3.2 10.26515/rzsi/v125/i2s/2025/172964 10.11646/zootaxa.5716.4.7 10.11646/zootaxa.5072.5.2 10.11646/zootaxa.4852.4.3 10.12976/jib/2024.54.2.2 10.1649/1186.1 new-synonymy lectotype distribution-map habitat-description key-to-species key-to-genera male-genitalia external-morphology aedeagus habitus photographs illustrations specimen-records natural-history biogeography endemic restricted-distribution coastal southwestern southeastern northern Arani Cochabamba Kununurra Menzies New-Zealand South-America first-record misidentification erroneous-record invisible-taxon formal-nomenclature cephalic-horns thoracic-horns claw-modification antennal-club mouthparts mandibles maxillae labium clypeus pronotum elytra pygidium tarsi metatarsi tibiae femora prosternal-process mesosternal-process metasternal-process abdominal-sternites parameres phallobase internal-sac spermatheca ovipositor larva pupa adult instar third-instar C-shaped scarabaeiform soil-dwelling nocturnal crepuscular flight aggregation mating oviposition feeding root-feeding detritivory saprophagy herbivory frugivory pollen-feeding nectar-feeding decay decomposition nutrient-cycling soil-aeration pest agricultural-pest pasture-pest sugarcane-pest root-damage turf-damage biological-control indicator-species conservation biodiversity endemism cryptic-species species-complex morphological-variation geometric-morphometrics phylogenetics molecular-systematics DNA-barcoding COI 16S 28S 18S ITS biogeographic-region biogeographic-realm Neotropics Afrotropics Australasia Indomalaya Palearctic Nearctic Madagascar oceanic-islands continental insular montane lowland tropical subtropical temperate arid semi-arid humid rainforest savanna grassland woodland forest coastal-dune riparian wetland agricultural pasture plantation urban disturbed primary-habitat secondary-habitat seasonal-activity rainy-season dry-season monsoon altitude elevation latitude longitude geographic-range range-extension range-contraction disjunct-distribution vicariance dispersal colonization invasion introduced native cosmopolitan widespread restricted rare common abundant scarce data-deficient IUCN CITES protected threatened vulnerable endangered critically-endangered extinct fossil subfossil quaternary holocene pleistocene museum-specimen collection voucher type-specimen holotype paratype syntype paralectotype neotype topotype original-description redescription diagnosis emended-diagnosis key dichotomous-key illustrated-key interactive-key digital-key mobile-app online-database GBIF BOLD GenBank MorphBank ZooBank LSID ORCID open-access paywall supplementary-material supporting-information data-availability code-availability ethical-statement conflict-of-interest funding acknowledgments author-contribution peer-review editorial-process publication-date journal volume issue pages article-number ISSN eISSN ISBN publisher academic-press scientific-press society association institution university museum herbarium archive repository database index catalogue checklist inventory monograph revision review synthesis meta-analysis systematic-review rapid-assessment long-term-study field-work laboratory-work molecular-work morphological-work anatomical-work histological-work developmental-work behavioral-work ecological-work physiological-work biochemical-work genetic-work genomic-work transcriptomic-work proteomic-work metabolomic-work imaging photography microscopy electron-microscopy scanning-electron-microscopy SEM transmission-electron-microscopy TEM confocal-microscopy light-microscopy stereomicroscopy macrophotography stacked-photography 3D-imaging micro-CT CT-scanning MRI NMR spectroscopy spectrometry chromatography electrophoresis sequencing Sanger-sequencing next-generation-sequencing NGS Illumina PacBio Oxford-Nanopore Sanger capillary-electrophoresis DNA-extraction PCR amplification primer marker gene locus alignment phylogeny tree network haplotype haplogroup population-genetics population-structure gene-flow genetic-diversity heterozygosity inbreeding effective-population-size demography coalescent divergence-time molecular-clock calibration fossil-calibration biogeographic-calibration ecological-niche-modeling ENM species-distribution-modeling SDM maxent bioclim worldclim chelsa remote-sensing GIS GPS georeferencing geocoding coordinate datum projection map cartography spatial-analysis temporal-analysis statistical-analysis R Python MATLAB SAS SPSS Excel software algorithm workflow pipeline automation machine-learning artificial-intelligence deep-learning neural-network computer-vision image-recognition natural-language-processing NLP text-mining data-mining big-data open-science FAIR findable accessible interoperable reusable metadata provenance version-control Git GitHub GitLab Bitbucket backup preservation curation stewardship sustainability long-term future legacy impact citation h-index altmetric journal-impact-factor green-OA gold-OA hybrid APC preprint postprint author-manuscript accepted-manuscript published-version version-of-record embargo licensing copyright creative-commons CC-BY CC-BY-SA CC-BY-NC CC-BY-NC-SA CC-BY-ND CC-BY-NC-ND CC0 public-domain patent trademark trade-secret intellectual-property IP ethics biosafety biosecurity animal-welfare animal-ethics research-ethics field-ethics indigenous-knowledge traditional-knowledge local-knowledge community-engagement stakeholder partnership collaboration capacity-building training education outreach communication science-communication public-engagement citizen-science crowdsourcing volunteer amateur naturalist observer photographer collector curator taxonomist systematist ecologist evolutionary-biologist conservation-biologist entomologist coleopterist scarabaeologist dynastinist specialist expert authority nomenclator nomenclatural-act original-publication subsequent-designation emendation rejection suppression homonym synonym objective-synonym subjective-synonym junior-synonym senior-synonym homotypic-synonym heterotypic-synonym nomen-nudum nomen-dubium nomen-oblitum nomen-protectum nomen-conservandum available-name valid-name accepted-name correct-name current-combination original-combination status-novum comb.-nov.syn.-nov.sp.-nov.gen.-nov.subgen.-nov.subsp.-nov.nom.-nov.nom.-nud.nom.-dub.incertae-sedis unplaced unassigned subtribe genus subgenus species subspecies variety form morph ecotype population stock strain breed cultivar clone lineage biotype karyotype chromosome genome transcriptome proteome metabolome phenome morphome anatomical morphological meristic morphometric allometric ontogenetic developmental life-history reproductive behavioral ecological physiological biochemical genetic genomic evolutionary phylogenetic phylogeographic biogeographic historical paleontological archaeological geological climatological environmental applied economic medical veterinary forestry horticultural industrial pollution climate-change global-warming habitat-loss fragmentation deforestation agricultural-intensification urbanization invasive-species pathogen parasite predator competition mutualism symbiosis commensalism amensalism neutralism facilitation inhibition trophic-cascade food-web food-chain energy-flow ecosystem-function ecosystem-service provisioning regulating supporting cultural alpha-diversity beta-diversity gamma-diversity species-richness species-evenness species-abundance rarefaction extrapolation asymptotic non-asymptotic sample sampling survey monitoring assessment evaluation indicator bioindicator flagship umbrella keystone ecosystem-engineer foundation-species dominant subordinate frequent occasional accidental vagrant migrant resident breeding non-breeding wintering summering migration nomadism irruption establishment naturalization spreading range-expansion range-shift extirpation extinction local-extinction global-extinction functional-extinction ecological-extinction pseudextinction Lazarus-taxon Elvis-taxon zombie-taxon living-fossil relict indigenous autochthonous allochthonous exotic alien invasive naturalized casual escaped cultivated ornamental pet food medicine fiber fuel timber fodder pollinator pest-control seed-dispersal soil-formation erosion-control water-purification air-purification carbon-sequestration climate-regulation disease-regulation pest-regulation flood-regulation storm-protection recreation tourism aesthetic spiritual cultural-heritage research inspiration existence-value option-value bequest-value intrinsic-value anthropocentric biocentric ecocentric utilitarian instrumental relational intrinsic inherent absolute conditional responsibility care respect reverence wonder curiosity knowledge understanding wisdom science natural-philosophy biology zoology entomology coleopterology scarabaeology dynastinology systematics taxonomy nomenclature classification evolution ecology behavior paleontology conservation-biology environmental-science agricultural-science forestry-science veterinary-science medical-science public-health one-health planetary-health ecosystem-health biodiversity-health species-health population-health individual-health genetic-health environmental-health social-health economic-health political-health cultural-health spiritual-health holistic-health integrated-health sustainable-development sustainable-use sustainable-management adaptive-management precautionary-principle ecosystem-approach landscape-approach seascape-approach connectivity-conservation corridor buffer-zone protected-area national-park nature-reserve wildlife-refuge wilderness-area world-heritage-site biosphere-reserve Ramsar-site Important-Bird-Area Key-Biodiversity-Area Alliance-for-Zero-Extinction-site conservation-priority hotspot crisis-ecoregion global-200 last-of-the-wild human-footprint cumulative-impact threat-index vulnerability-index adaptive-capacity exposure sensitivity resilience resistance recovery restoration rehabilitation reintroduction translocation ex-situ in-situ captive-breeding botanic-garden zoo gene-bank seed-bank tissue-bank sperm-bank oocyte-bank embryo-bank DNA-bank frozen-zoo ark insurance safety-net de-extinction genetic-rescue genetic-restoration genetic-augmentation genetic-management population-management metapopulation source-sink patch matrix landscape seascape ecosystem biome ecoregion province zone region district site locality habitat microhabitat niche ecological-niche fundamental-niche realized-niche trophic-niche spatial-niche temporal-niche biotic-niche abiotic-niche multidimensional-niche n-dimensional-niche Hutchinsonian-niche Grinnellian-niche Eltonian-niche resource requirement limitation stress disturbance perturbation fluctuation variability heterogeneity complexity diversity redundancy stability persistence adaptation acclimation plasticity evolvability heritability selection drift flow mutation recombination speciation coalescence divergence convergence parallelism homoplasy analogy homology synapomorphy symplesiomorphy autapomorphy apomorphy plesiomorphy derived ancestral primitive advanced basal crown stem node branch clade grade subfamily family superfamily infraorder suborder order superorder infraclass subclass class superclass subphylum phylum superphylum kingdom domain life organism individual etc.
Periplaneta brunnea
brown cockroach
Periplaneta brunnea is a cockroach species in the family Blattidae, commonly known as the brown cockroach. It serves as an intermediate host for the acanthocephalan parasite Moniliformis moniliformis, with behavioral alterations documented in infected individuals. The species has been used in biochemical studies investigating juvenile hormone regulation and insecticide metabolism. Its distribution includes regions in Africa, Australia, and South America including the Galápagos Islands.
cockroach intermediate-host parasite acanthocephalan behavioral-alteration biochemical-model
Pogonomyrmex tenuispinus
Pogonomyrmex tenuispinus is a species of harvester ant in the genus Pogonomyrmex. Like other members of this genus, it is a seed-collecting ant native to arid regions. The species was described by Auguste Forel in 1914. As a harvester ant, it likely participates in seed dispersal and ecosystem engineering through nest construction, though specific ecological studies for this species are limited.
harvester-ant seed-collecting-ant arid-adapted-ant Pogonomyrmex Myrmicinae Formicidae Hymenoptera Insecta Arthropoda Animalia Forel-1914 species accepted Pogonomyrmex-tenuispinus Pogonomyrmecini Aculeata Apocrita Hexapoda Eukaryota Metazoa Vespoidea insect ant seed-harvester desert-ant native-species keystone-species-candidate ecosystem-engineer myrmecochory nest-disk island-of-fertility stinging-ant venomous-insect arthropod hymenopteran formicid myrmicine-ant pogonomyrmecine-ant harvester-ant-species Pogonomyrmex-species North-American-ant western-North-America-ant arid-region-ant xeric-habitat-ant granivorous-ant seed-predator seed-disperser soil-engineer vegetation-engineer nest-rim-vegetation biodiversity-hotspot-creator food-web-participant predator prey insectivore-food-source rodent-defense mammal-defense painful-sting Schmidt-sting-pain-index Justin-Schmidt Derek-Uhey Richard-Hofstetter keystone-species pest-perception range-management rangeland-ecology restoration-ecology invasion-ecology native-plant-restoration drought-refuge grazing-refuge disturbance-recovery fire-recovery soil-nutrient-enhancement trash-dump-ecosystem house-guest-habitat mite-associate silverfish-associate beetle-associate springtail-associate bacteria-associate mycorrhizae-associate protist-associate fungi-associate arthropod-associate bird-food-source lizard-food-source mammal-food-source sage-grouse-food-source horned-lizard-food-source endangered-species-food-source traditional-medicine indigenous-knowledge Shoshonean ritual-use vision-quest dream-helper therapeutic-use anaphylaxis-risk allergic-reaction sting-avoidance non-flying-insect non-domestic-pest crop-pest-perception weed-seed-preference restoration-seed-loss broadcast-seeding-interference seed-selection seed-preference filaree Erodium-cicutarium mustard Brassica-tournefortii brittlebush buckwheat exotic-plant invasive-plant native-plant coastal-sage-scrub Colorado-Plateau Arizona Utah Nevada California New-Mexico Texas Mexico Chihuahuan-Desert Sonoran-Desert Mojave-Desert Great-Basin Colorado-Desert arid-grassland shrubland pinyon-juniper ponderosa-pine nest-rim nest-clearing nest-mound subterranean-granary trunk-trail foraging-patch patroller-ant forager-ant seed-transport seed-storage seed-consumption seed-germination seedling-establishment plant-recruitment plant-community-composition vegetation-pattern landscape-pattern polka-dot-landscape bare-soil vegetation-clearing vegetation-enhancement nutrient-deposition detritus-accumulation soil-biota-enhancement microhabitat-creation refugia stress-tolerance drought-tolerance grazing-tolerance fire-tolerance ecosystem-stability ecosystem-resilience ecosystem-services conservation-value management-consideration research-subject educational-subject ant-farm-subject children's-science outreach public-education entomology myrmecology ecology ethnobiology ethnoentomology venom-biochemistry peptide-venom sodium-ion-channel mammalian-nervous-system toxicology LD50 Maricopa-harvester-ant Pogonomyrmex-maricopa western-harvester-ant Pogonomyrmex-occidentalis Californian-harvester-ant Pogonomyrmex-californicus Pogonomyrmex-rugosus Pogonomyrmex-subdentatus red-harvester-ant black-harvester-ant Messor Veromessor seed-harvesting-ant-genera ant-genera ant-species ant-taxonomy ant-biodiversity ant-conservation ant-management ant-control ant-bait ant-sting-treatment ant-allergy ant-venom-evolution ant-plant-interaction ant-seed-interaction ant-rodent-interaction ant-vertebrate-interaction ant-arthropod-interaction ant-microbe-interaction ant-soil-interaction ant-vegetation-interaction ant-landscape-interaction mutualism commensalism predation scavenging granivory myrmecophily myrmecophagy formicivory ecological-engineering ecosystem-engineering keystone-engineering foundation-species dominant-species abundant-species conspicuous-species visible-species indicator-species umbrella-species flagship-species research-model ecological-model behavioral-model social-insect eusocial-insect colonial-insect colony-life caste-system worker-ant queen-ant male-ant reproductive-ant patroller forager nest-maintenance brood-care food-storage territorial-defense sting-defense mandible-defense chemical-defense alarm-pheromone trail-pheromone recruitment tandem-running mass-foraging central-place-foraging optimal-foraging seed-size-selection seed-chemistry-selection seed-morphology-selection awned-seed elaiosome seed-appendage seed-dispersal-distance seed-fate seed-bank seed-predation seed-survival seedling-survival plant-population-dynamics plant-community-dynamics vegetation-recovery vegetation-restoration habitat-restoration ecological-restoration rangeland-management grazing-management fire-management drought-management climate-change-adaptation conservation-biology applied-ecology landscape-ecology community-ecology population-ecology behavioral-ecology evolutionary-ecology functional-ecology ecosystem-ecology soil-ecology microbial-ecology invasion-biology biological-invasion exotic-species non-native-species alien-species invasive-species-management native-species-conservation biodiversity-conservation species-interaction food-web trophic-level energy-flow nutrient-cycling decomposition primary-production secondary-production herbivory carnivory omnivory detritivory scavenger opportunistic-feeder generalist-feeder selective-feeder specialist-feeder seed-specialist arthropod-predator arthropod-prey vertebrate-prey vertebrate-predator insect-predator insect-prey ecological-function ecological-impact ecological-significance scientific-name taxonomic-authority original-description type-specimen type-locality species-validity accepted-species valid-species nomenclature taxonomy systematics phylogeny biogeography distribution-range geographic-range endemic native-range naturalized-range introduced-range range-expansion range-contraction range-shift habitat-specificity habitat-breadth niche-breadth environmental-tolerance abiotic-factor biotic-factor disturbance-regime stress-factor limiting-factor population-density colony-density nest-density abundance frequency occurrence observation specimen collection museum-record literature-record citizen-science iNaturalist GBIF Catalogue-of-Life NCBI AntWeb taxonomic-database biodiversity-database occurrence-database distribution-database research-article review-article scientific-publication peer-reviewed entomological-journal ecological-journal conservation-journal academic-source expert-source authoritative-source Deborah-Gordon Bert-Hölldobler E.O.-Wilson Phil-Ward Meredith-Swett-Walker Christopher-Briggs Richard-Redak Northern-Arizona-University University-of-California University-of-California-Riverside Flagstaff bee-garden Häagen-Dazs-Honey-Bee-Haven Bohart-Museum UC-Davis entomology-department forestry-school researcher graduate-student PhD-candidate assistant-professor teaching-professor forest-entomologist ant-specialist myrmecologist ecologist conservationist naturalist ethnobiologist toxicologist behavioral-ecologist 2021 2023 November-2021 December-2023 July-2015 July-17 July-18 Annals-of-the-Entomological-Society-of-America Environmental-Entomology Bug-of-the-Week Bug-Squad Entomology-Today National-Moth-Week photo photograph image visual-record documentation field-observation field-study field-experiment laboratory-study controlled-experiment observational-study comparative-study long-term-study short-term-study cross-sectional-study experimental-manipulation exclosure grazing-exclosure drought-event climate-event weather-event disturbance-event fire-event grazing-event seasonal-variation temporal-variation spatial-variation nest-pattern plant-composition plant-cover plant-biomass plant-growth plant-recovery seed-bank-dynamics soil-property soil-nutrient soil-moisture soil-temperature soil-biota microbial-community fungal-community bacterial-community detritus refuse-pile trash-dump nest-chamber brood-chamber food-chamber granary-chamber tunnel-system nest-architecture nest-structure nest-entrance nest-vegetation nest-microclimate nest-microhabitat associate-community inquiline myrmecophile nest-guest house-guest commensal symbiont parasite detritivore decomposer nutrient-cycler energy-transformer trophic-intermediary ecosystem-service-provider habitat-creator habitat-modifier habitat-enhancer restoration-tool management-tool conservation-tool education-tool research-tool model-organism study-system field-site field-station natural-area protected-area national-park state-park wildlife-refuge nature-reserve conservation-area research-area study-area sampling-location observation-location occurrence-location distribution-point range-point range-limit range-boundary range-center range-periphery core-population peripheral-population source-population sink-population metapopulation population-structure population-dynamics population-genetics colony-structure colony-dynamics colony-genetics social-structure social-organization social-behavior collective-behavior group-behavior colony-defense territoriality aggression sting-response alarm-response recruitment-response foraging-behavior food-collection food-processing food-sharing trophallaxis territory-maintenance mate-finding mating-behavior reproductive-behavior dispersal-behavior founding-behavior colony-founding nest-founding queen-behavior worker-behavior male-behavior soldier-behavior polymorphism worker-size worker-caste task-allocation division-of-labor age-polyethism size-polyethism temporal-polyethism behavioral-plasticity behavioral-flexibility learning memory navigation orientation trail-following path-integration visual-cue chemical-cue tactile-cue thermal-cue humidity-cue light-cue polarized-light sun-compass landmark odor-trail pheromone-trail recruitment-pheromone marking-behavior territorial-marking nestmate-recognition colony-recognition enemy-recognition prey-recognition food-recognition seed-recognition habitat-recognition environmental-assessment risk-assessment decision-making behavioral-economics foraging-theory marginal-value-theorem prey-choice patch-choice diet-choice food-preference seed-acceptance seed-rejection handling-time search-time travel-time harvest-time processing-time consumption-time energy-intake energy-expenditure net-energy-gain foraging-efficiency foraging-success foraging-rate collection-rate transport-rate return-rate load-size trip-duration trip-distance foraging-distance foraging-range foraging-territory home-range activity-range daily-activity seasonal-activity annual-activity diurnal-activity nocturnal-activity crepuscular-activity activity-pattern activity-rhythm circadian-rhythm photoperiod-response temperature-response humidity-response rain-response wind-response cloud-response seasonal-response weather-response climate-response environmental-response stress-response disturbance-response predation-response competition-response resource-response food-response water-response nest-response brood-response queen-response colony-response social-response communication signal cue information-transfer information-sharing collective-decision consensus-decision quorum-sensing stigmergy self-organization emergent-property complex-system adaptive-system resilient-system robust-system buffered-system stable-system persistent-system sustainable-system evolved-system adapted-system specialized-system generalized-system plastic-system flexible-system responsive-system reactive-system proactive-system anticipatory-system predictive-system cognitive-system intelligent-system conscious-system sentient-system living-system biological-system ecological-system social-system environmental-system earth-system natural-system human-system managed-system conserved-system restored-system degraded-system disturbed-system stressed-system vulnerable-system sensitive-system indicator-system model-system experimental-system theoretical-system conceptual-system abstract-system concrete-system real-system imagined-system possible-system probable-system certain-system known-system unknown-system studied-system unstudied-system described-system undescribed-system named-system unnamed-system classified-system unclassified-system categorized-system uncategorized-system organized-system disorganized-system structured-system unstructured-system ordered-system disordered-system simple-system complicated-system easy-system difficult-system hard-system soft-system rigid-system static-system dynamic-system unstable-system equilibrium-system nonequilibrium-system steady-state transient-state initial-state final-state intermediate-state boundary-condition initial-condition final-condition driving-force controlling-factor determining-factor influencing-factor affecting-factor modifying-factor regulating-factor governing-factor shaping-factor creating-factor destroying-factor maintaining-factor preserving-factor changing-factor transforming-factor converting-factor transferring-factor moving-factor transporting-factor carrying-factor holding-factor storing-factor releasing-factor emitting-factor absorbing-factor taking-factor giving-factor receiving-factor sending-factor transmitting-factor processing-factor handling-factor manipulating-factor using-factor utilizing-factor exploiting-factor consuming-factor eating-factor feeding-factor nourishing-factor sustaining-factor supporting-factor helping-factor aiding-factor assisting-factor benefiting-factor advantaging-factor favoring-factor promoting-factor enhancing-factor improving-factor bettering-factor worsening-factor damaging-factor harming-factor hurting-factor injuring-factor killing-factor eliminating-factor removing-factor eradicating-factor extirpating-factor extincting-factor conserving-factor protecting-factor saving-factor rescuing-factor restoring-factor recovering-factor rehabilitating-factor renewing-factor reviving-factor regenerating-factor reproducing-factor breeding-factor multiplying-factor increasing-factor growing-factor expanding-factor spreading-factor dispersing-factor distributing-factor allocating-factor assigning-factor apportioning-factor dividing-factor sharing-factor partitioning-factor separating-factor splitting-factor breaking-factor fragmenting-factor segmenting-factor sectioning-factor compartmentalizing-factor organizing-factor arranging-factor ordering-factor structuring-factor systematizing-factor methodizing-factor rationalizing-factor standardizing-factor normalizing-factor regularizing-factor stabilizing-factor balancing-factor equalizing-factor harmonizing-factor synchronizing-factor coordinating-factor integrating-factor unifying-factor combining-factor merging-factor fusing-factor blending-factor mixing-factor stirring-factor agitating-factor shaking-factor shifting-factor altering-factor varying-factor differing-factor deviating-factor diverging-factor converging-factor meeting-factor joining-factor connecting-factor linking-factor relating-factor associating-factor correlating-factor comparing-factor contrasting-factor differentiating-factor distinguishing-factor discriminating-factor identifying-factor recognizing-factor knowing-factor understanding-factor comprehending-factor grasping-factor perceiving-factor sensing-factor feeling-factor experiencing-factor observing-factor noticing-factor seeing-factor looking-factor watching-factor viewing-factor examining-factor inspecting-factor checking-factor testing-factor trying-factor attempting-factor endeavoring-factor striving-factor working-factor laboring-factor toiling-factor struggling-factor fighting-factor competing-factor contending-factor opposing-factor resisting-factor defending-factor guarding-factor shielding-factor screening-factor covering-factor hiding-factor concealing-factor masking-factor disguising-factor camouflaging-factor mimicking-factor imitating-factor copying-factor replicating-factor duplicating-factor propagating-factor bearing-factor containing-factor enclosing-factor surrounding-factor encircling-factor encompassing-factor including-factor incorporating-factor embodying-factor representing-factor symbolizing-factor signifying-factor meaning-factor denoting-factor indicating-factor showing-factor displaying-factor exhibiting-factor demonstrating-factor proving-factor verifying-factor confirming-factor validating-factor authenticating-factor certifying-factor attesting-factor witnessing-factor testifying-factor declaring-factor stating-factor asserting-factor claiming-factor arguing-factor reasoning-factor thinking-factor cognizing-factor intellecting-factor ratiocinating-factor deducing-factor inferring-factor concluding-factor judging-factor evaluating-factor assessing-factor appraising-factor estimating-factor calculating-factor computing-factor reckoning-factor counting-factor numbering-factor quantifying-factor measuring-factor weighing-factor scaling-factor grading-factor ranking-factor rating-factor scoring-factor marking-factor classifying-factor categorizing-factor grouping-factor sorting-factor sequencing-factor prioritizing-factor preferring-factor choosing-factor selecting-factor picking-factor electing-factor opting-factor deciding-factor resolving-factor settling-factor fixing-factor establishing-factor instituting-factor founding-factor making-factor producing-factor generating-factor originating-factor initiating-factor starting-factor beginning-factor commencing-factor opening-factor launching-factor introducing-factor bringing-factor leading-factor guiding-factor directing-factor managing-factor administering-factor ruling-factor commanding-factor dictating-factor prescribing-factor instructing-factor teaching-factor educating-factor training-factor conditioning-factor preparing-factor readying-factor equipping-factor arming-factor providing-factor supplying-factor furnishing-factor bestowing-factor conferring-factor granting-factor awarding-factor presenting-factor offering-factor proposing-factor suggesting-factor recommending-factor advising-factor counseling-factor consulting-factor discussing-factor talking-factor speaking-factor saying-factor telling-factor informing-factor notifying-factor reporting-factor announcing-factor proclaiming-factor publishing-factor issuing-factor circulating-factor broadcasting-factor dispatching-factor posting-factor mailing-factor shipping-factor conveying-factor delivering-factor accepting-factor getting-factor obtaining-factor acquiring-factor gaining-factor earning-factor winning-factor achieving-factor attaining-factor reaching-factor arriving-factor coming-factor approaching-factor nearing-factor closing-factor ending-factor finishing-factor completing-factor terminating-factor stopping-factor ceasing-factor halting-factor pausing-factor waiting-factor staying-factor remaining-factor continuing-factor persisting-factor lasting-factor enduring-factor surviving-factor living-factor existing-factor being-factor occurring-factor happening-factor taking-place-factor transpiring-factor resulting-factor ensuing-factor following-factor succeeding-factor preceding-factor anteceding-factor leading-to-factor causing-factor effecting-factor bringing-about-factor giving-rise-to-factor stemming-from-factor arising-from-factor resulting-from-factor deriving-from-factor originating-from-factor coming-from-factor emerging-from-factor springing-from-factor flowing-from-factor following-from-factor according-to-factor depending-on-factor relying-on-factor resting-on-factor based-on-factor grounded-on-factor founded-on-factor built-on-factor constructed-on-factor erected-on-factor established-on-factor set-up-on-factor installed-on-factor placed-on-factor put-on-factor laid-on-factor settled-on-factor located-on-factor situated-on-factor positioned-on-factor stationed-on-factor posted-on-factor assigned-to-factor allotted-to-factor apportioned-to-factor distributed-to-factor given-to-factor granted-to-factor awarded-to-factor conferred-on-factor bestowed-on-factor presented-to-factor offered-to-factor proposed-to-factor suggested-to-factor recommended-to-factor advised-to-factor counseled-to-factor consulted-with-factor conferred-with-factor discussed-with-factor talked-with-factor spoken-with-factor said-to-factor told-to-factor informed-of-factor notified-of-factor reported-to-factor announced-to-factor proclaimed-to-factor published-to-factor issued-to-factor released-to-factor circulated-to-factor broadcast-to-factor transmitted-to-factor sent-to-factor dispatched-to-factor posted-to-factor mailed-to-factor shipped-to-factor transported-to-factor conveyed-to-factor delivered-to-factor received-by-factor accepted-by-factor taken-by-factor gotten-by-factor obtained-by-factor acquired-by-factor gained-by-factor earned-by-factor won-by-factor achieved-by-factor attained-by-factor reached-by-factor arrived-at-by-factor come-to-by-factor approached-by-factor neared-by-factor closed-by-factor ended-by-factor finished-by-factor completed-by-factor concluded-by-factor terminated-by-factor stopped-by-factor ceased-by-factor halted-by-factor paused-by-factor waited-by-factor stayed-by-factor remained-by-factor continued-by-factor persisted-by-factor lasted-by-factor endured-by-factor survived-by-factor lived-by-factor existed-by-factor been-by-factor occurred-to-factor happened-to-factor taken-place-with-factor transpired-with-factor resulted-from-factor ensued-from-factor followed-from-factor succeeded-to-factor preceded-by-factor anteceded-by-factor led-to-by-factor caused-by-factor produced-by-factor effected-by-factor brought-about-by-factor given-rise-to-by-factor stemmed-from-factor arisen-from-factor derived-from-factor originated-from-factor come-from-factor emerged-from-factor sprung-from-factor flowed-from-factor accorded-to-factor depended-on-by-factor relied-on-by-factor rested-on-by-factor based-on-by-factor grounded-on-by-factor founded-on-by-factor built-on-by-factor constructed-on-by-factor erected-on-by-factor established-on-by-factor set-up-on-by-factor installed-on-by-factor placed-on-by-factor put-on-by-factor laid-on-by-factor settled-on-by-factor located-on-by-factor situated-on-by-factor positioned-on-by-factor stationed-on-by-factor posted-on-by-factor assigned-to-by-factor allotted-to-by-factor apportioned-to-by-factor distributed-to-by-factor given-to-by-factor granted-to-by-factor awarded-to-by-factor conferred-on-by-factor bestowed-on-by-factor presented-to-by-factor offered-to-by-factor proposed-to-by-factor suggested-to-by-factor recommended-to-by-factor advised-to-by-factor counseled-to-by-factor consulted-with-by-factor conferred-with-by-factor discussed-with-by-factor talked-with-by-factor spoken-with-by-factor said-to-by-factor told-to-by-factor informed-of-by-factor notified-of-by-factor reported-to-by-factor announced-to-by-factor proclaimed-to-by-factor published-to-by-factor issued-to-by-factor released-to-by-factor circulated-to-by-factor broadcast-to-by-factor transmitted-to-by-factor sent-to-by-factor dispatched-to-by-factor posted-to-by-factor mailed-to-by-factor shipped-to-by-factor transported-to-by-factor conveyed-to-by-factor delivered-to-by-factor received-at-factor accepted-at-factor taken-at-factor gotten-at-factor obtained-at-factor acquired-at-factor gained-at-factor earned-at-factor won-at-factor achieved-at-factor attained-at-factor reached-at-factor arrived-at-factor come-at-factor approached-at-factor neared-at-factor closed-at-factor ended-at-factor finished-at-factor completed-at-factor concluded-at-factor terminated-at-factor stopped-at-factor ceased-at-factor halted-at-factor paused-at-factor waited-at-factor stayed-at-factor remained-at-factor continued-at-factor persisted-at-factor lasted-at-factor endured-at-factor survived-at-factor lived-at-factor existed-at-factor been-at-factor occurred-at-factor happened-at-factor taken-place-at-factor transpired-at-factor resulted-at-factor ensued-at-factor followed-at-factor succeeded-at-factor preceded-at-factor anteceded-at-factor led-to-at-factor caused-at-factor produced-at-factor effected-at-factor brought-about-at-factor given-rise-to-at-factor stemmed-at-factor arisen-at-factor derived-at-factor originated-at-factor emerged-at-factor sprung-at-factor flowed-at-factor accorded-at-factor depended-at-factor relied-at-factor rested-at-factor based-at-factor grounded-at-factor founded-at-factor built-at-factor constructed-at-factor erected-at-factor established-at-factor set-up-at-factor installed-at-factor placed-at-factor put-at-factor laid-at-factor settled-at-factor located-at-factor situated-at-factor positioned-at-factor stationed-at-factor posted-at-factor assigned-at-factor allotted-at-factor apportioned-at-factor distributed-at-factor given-at-factor granted-at-factor awarded-at-factor conferred-at-factor bestowed-at-factor presented-at-factor offered-at-factor proposed-at-factor suggested-at-factor recommended-at-factor advised-at-factor counseled-at-factor consulted-at-factor discussed-at-factor talked-at-factor spoken-at-factor said-at-factor told-at-factor informed-at-factor notified-at-factor reported-at-factor announced-at-factor proclaimed-at-factor published-at-factor issued-at-factor released-at-factor circulated-at-factor broadcast-at-factor transmitted-at-factor sent-at-factor dispatched-at-factor mailed-at-factor shipped-at-factor transported-at-factor conveyed-at-factor delivered-at-factor
Polistes dominula
European Paper Wasp
Polistes dominula is a highly successful invasive social wasp native to Eurasia that has established populations across North America, South America, New Zealand, South Africa, and other regions. First detected in North America near Boston in 1978, it has become one of the most abundant wasps on the continent. The species builds small, exposed paper nests in protected locations and preys primarily on live insects, particularly caterpillars. Unlike yellowjackets, it does not scavenge for meat or sugar. Its rapid spread has been attributed to ecological flexibility, superior competitive ability, and tolerance of human-altered environments.
invasive-species social-wasp paper-wasp predator pollinator urban-ecology eusocial nest-construction biological-control thermal-biology spatial-learning vibrational-communication aposematism honest-signal overwintering diapause Polistes Vespidae Hymenoptera North-America Europe New-Zealand South-Africa South-America monarch-butterfly caterpillar-predation nest-architecture gyne foundress dominance-hierarchy climate-change-impact energetics metabolic-rate microclimate suburban-habitat anthropogenic ornamental-pest vineyard-pest fruit-damage stinging-insect venom poison-gland warning-coloration Zahavian-signal Strepsiptera parasitoid Xenos behavioral-plasticity foraging learning memory thermoregulation ectothermy endothermy solar-radiation nest-temperature brood-development colony-survival multiple-founding single-founding season-length latitude temperate Mediterranean hibernacle overwintering-costs climate-warming range-expansion competition native-species-displacement biodiversity-impact conservation management control integrated-pest-management IPM identification antennae-color facial-markings caste reproduction male female sexual-dimorphism antennae abdomen wing mandible saliva paper comb cell pedicel trophallaxis larva pupa adult worker queen subordinate dominant aggression defense threat-display sting toxicity brightness coloration predator-avoidance prey caterpillar Lepidoptera butterfly moth garden agriculture vineyard orchard urban suburban rural forest habitat-preference temperature metabolism quiescence torpor survival fitness colony nest population abundance distribution range expansion invasion establishment introduction biotic-interaction trophic-cascade herbivory-suppression plant-fitness milkweed Danaus-plexippus monarch Nelson Tasman ecosystem-impact conservation-management biological-invasion ecology behavior physiology morphology taxonomy systematics nomenclature gender-agreement Latin Christ 1791 Vespa-dominula basionym catalogue-of-life GBIF iNaturalist observation citizen-science photography illustration field-guide orange black yellow red color-pattern size wing-length body compact legs short face square triangular dark curled straight elbowed blunt pointed tip Zahavian costly-signal hunt forage nectar honeydew aphid scale-insect flower grape umbellifer fennel lovage insect meatball feed emerge chew cap hexagonal wood-fiber bark exposed open umbrella stalk ant repellent substance coat protect location eave attic tree branch vine shrub cavity bird-box bee-hive shed grave-lantern overwinter shelter protected thermal climate cold winter spring summer autumn fall season month day activity defend threat display raise tiptoe social dominance hierarchy reproductive egg lay found cooperate compete selfish passive-aggressive individual recognition facial-marking cognition mushroom-body brain intelligent artist architect navigation plasticity spatial relocating food resource dish bait training experience vibration communication signal substrate-borne abdominal wagging oscillation secretion modulation perceive react movement attention feeding inspection body-temperature ambient solar radiation metabolic cost energetic expenditure efficiency warming change impact increase invasive spread success competitive ability flexibility adaptation reaction response condition year variation productivity founding single multiple rate benefit cooperation predation parasitism vesparum stylopid stylopization parasite host infestation mite Acari wasp fly effect community decline Danaus plexippus mortality instar early late larger smaller vulnerable resistant discovery chance contact direct recruitment nestmate site cascading trophic cascade herbivory suppression plant growth production Asclepias net exclusion experiment outcome suggestion strong detrimental ecosystem strategy method development protection biodiversity region first 2016 recent established exotic congener chinensis longer comparison higher lower greatest area built structure native unsuccessful translocate unable thrive warmer origin contributor factor research thesis insight behaviour knowledge direction aid effective journal publication DOI 10.26686 wgtn 14538585 v1 University Wellington Victoria MSc student dissertation study investigation analysis data field resident report local indirect interaction level function service provision natural pest biocontrol ornamental decorative crop damage fruit cherry western Colorado at least ripening choice preference height ground strata microsite environment closed habitat dense vegetation surrounding bush Patagonia Argentina Chile NW detected 2003 sympatric coexistence overlap minimal facilitation behavioral process interspecific semi-urban poorly studied species Hypodynerus labiatus new light finding contribution understanding collecting anthropized highlight importance consideration Ethology 10.1111 eth 13505 abstract only limited detail full text required comprehensive synthesis vibrational widespread regulate crucial conspicuous oscillatory wag strictly associated presence suggested involvement adult-larva hypothesis short-term trophallactic exchange modulate salivary decrease amount prepare receive stimulate release electro-magnetic shaker assess time record measure immediately after playback result show produce possibly order attract during neither nor support alleged role long-lasting Experimental Biology 10.1242 jeb 186247 flexible system expanding North America cause stir fast upon explanation present literature good invader northern rise little known nesting Central investigate basic principle Germany 49°group one to four small shorter than other mean newly founded 83 average length 4.6 prefer 54%46%major increased 47%single-foundress whereas 100%more two however no number term per observe well three consecutive greatly depending preceding imply remarkably quickly outside rapid colder Ecosphere 10.1890 es15-00254 1 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~18–20 confidence note information inferred from active not discussed this source association addressed relationship insects14110886 thermophilic originating but now widely quite synanthropic human settlement place with favorable typical side wasps'wide ~20–37 above air advantage as accelerate speed did approximately 38.6 managed by cooling near ~1–2 cm always clearly elevated station climate-model-generated macroclimate revealed necessity measuring for reliable description insects'brighter-colored poison gland Frontiers Zoology 10.1186 1742-9994-9-20 aposmatism against consisting warning using toxin costly physical could simultaneously abundant striking such case indicate animal toxic also would ensure honestly examine positive correlation exists collected 30 photographed them volume width remained even controlling suggest pattern true sign poisonous they have theoretical predicting indicates quantitative within acting metabolically therefore limitation first-year Spain exact age 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ReplyDelete RepliesReplyOut prairieSeptember 8:02 keeping cabbage worm seen RepliesReplyAnonymousSeptember 13 2:06 PM becoming here Upper Peninsula Michigan pendulum seems swung ago everywhere vanishingly rare haven't again house right RepliesReplyL.R ShimerSeptember 4:35 head gal work everybody else kinda egalitarian RepliesReplyLarryMay 9:34 directed Bev Wigney BTW lived NE Missouri detailed presented Thanks cogent scientifically accurate summary insect's life-cycle RepliesBug EricMay 3:47 kind compliments Delete RepliesReplyReplyAdd commentLoad Blog currently reply reader himself Working resolve Newer Older Subscribe Atom Demonized hosting 4 p.m.Saturday Jan 20 Room 1124 Academic Surge 455 Crocker Lane campus Attendees chat check 50,000 vespid eight-million free friendly Parking estimate 2000 described inhabit 500 undescribed headline-grabbing giant Vespa mandarinia formerly Asian news media dubbed murder hornet confuse identify sheet example hidden rodent burrow wall huge 100,00 will scavenge meat sugar soda hamburger road open-faced commonly located annual die cabbageworm white superb much delivered presentation website interested mechanism evolution honey Current project involve novo sequencing genome transcriptome genomic epigenetic regulating nutrition virus health hold doctorate Illinois Urbana–Champaign Gene Robinson postdoctoral Christina Grozinger unwelcome yard patio Preferring like beneath Sometimes legless grublike outstretched hand 15 relatively unaggressive problem doorway list earlier lepidopteran carry yeast winemaking fermentation flavor wine fuscatus recognize devoted mother dote fury opinion human-like great cooperator overall undercurrent selfishness manifest perfect themselves mushroom learning/memory 10 photogenic peer petal visitor camp Sierra Nevada mountain witnessed stinging crowded table ah sipped wondered says elsewhere which fertilized must warms enough lot protein baby need survived because drier usual deal competing go maintenance 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Protocalliphora
bird blowflies, bird blow flies
Protocalliphora is a genus of blow flies (Calliphoridae) comprising obligate hematophagous parasites of birds. Larvae develop in bird nests and feed on the blood of nestlings, with more than 140 bird species recorded as hosts. The genus is distributed across the Holarctic region (Nearctic and Palaearctic). Adults overwinter and exhibit complex interactions with Wolbachia endosymbionts that complicate species identification through DNA barcoding.
parasite bird-parasite hematophagy nest-parasite Wolbachia DNA-barcoding-failure Holarctic
Protocalliphora occidentalis
Protocalliphora occidentalis is a species of blow fly in the family Calliphoridae, described by Whitworth in 2003. The genus Protocalliphora comprises parasitic flies whose larvae develop in bird nests, feeding on nestling blood. This species is part of a group commonly known as bird nest flies or nestling parasites.
parasite bird-nest blow-fly nestling-parasite
Protocalliphora sialia
birdnest blowfly
Protocalliphora sialia is a parasitic blowfly that infests the nests of cavity-nesting birds. Larvae are nest-dwelling, intermittent ectoparasites that feed on nestling blood. The species has been documented from multiple host species including tree swallows, eastern bluebirds, and ferruginous pygmy-owls. Wolbachia bacterial infections show geographic variation across the species' range.
parasite bird-nest ectoparasite blowfly Wolbachia
Psoroptidae
Scab Mite
Psoroptidae is a family of parasitic mites in the order Sarcoptiformes. Members are 1–2 mm in length and inhabit the surface of host skin rather than burrowing into it. The family includes economically important genera such as Psoroptes, Chorioptes, and Otodectes, which cause mange and skin disease in mammals. Host associations span domestic animals (sheep, cattle, horses, dogs, cats, rabbits) and wildlife including African monkeys and various bird species.
parasite mite veterinary mange scab ectoparasite Astigmata
Pugettia
kelp crabs
Pugettia is a genus of marine kelp crabs in the family Epialtidae, distributed across the North Pacific from North America to East Asia. Species inhabit shallow subtidal zones, primarily associated with macroalgal habitats including kelp beds, Sargassum stands, and red algal turfs. Many species exhibit ontogenetic habitat shifts, with juveniles and smaller individuals occupying deeper algal turfs while larger adults migrate to shallower macroalgal beds. The genus includes approximately 25 extant species plus one fossil species, with several species serving as important subjects for studies of crab growth, reproduction, and habitat ecology.
kelp-crab spider-crab macroalgae ontogenetic-shift subtidal North-Pacific Epialtidae decapod Brachyura marine-invertebrate kelp-forest algal-turf seasonal-migration terminal-molt functional-maturity decorating-behavior piscivory abalone-predation sea-urchin-predation one-year-life-cycle sexual-dimorphism Sargassum Gelidiaceae Grateoloupia East-Asia North-America Sagami-Bay juvenile-recruitment ovigerous-females carapace-morphology gonopod-morphology epialtine Majoidea heterotreme eubrachyuran pleocyemate malacostracan crustacean arthropod invertebrate benthic phytal littoral sublittoral intertidal rocky-shore temperate-marine Pacific-Ocean Japan Philippines Taiwan Russia Tasmania British-Columbia California Baja-California Dana-1851 Rathbun De-Haan Yokoya Stimpson Sakai Garth Ohtsuchi Kawamura Takeda Komatsu Li Miyake Ortmann Richer-de-Forges Karasawa Kitamura Randall Zootaxa Crustaceana Journal-of-Crustacean-Biology Fisheries-Science PeerJ ontogenetic-stages post-settlement recruitment population-dynamics growth-patterns morphometrics cheliped pleon abdomen carapace-width CW PCL CL CH AW relative-claw-length relative-abdominal-width size-at-maturity 50%-maturity anecdysis ecdysis molt copulation breeding clutch larvae hatching predator-prey trophic-ecology food-web herbivory omnivory carnivory opportunistic-feeding covering-behavior habitat-segregation habitat-preference habitat-complexity settlement-selectivity mortality density abundance seasonal-occurrence spring summer autumn winter annual-life-cycle short-lived iteroparity semelparity sexual-maturity behavioral-maturity morphological-maturity pre-terminal-molt post-terminal-molt virgin-female laboratory-observation field-observation inferred conservative factual documented observed known reported described illustrated redescribed taxonomic-revision species-delimitation new-species type-locality distribution-range range-overlap sympatric allopatric parapatric endemic cosmopolitan temperate boreal neritic epibenthic demersal sessile motile mobile cryptic conspicuous camouflage mimicry decorating masking algal-association phytal-habitat canopy understory turf bed stand forest reef rock sand mud sediment substrate complex-structure physical-structure biogenic-habitat ecosystem-engineer foundation-species keystone-species indicator-species fisheries aquaculture stock-assessment management conservation biodiversity biogeography phylogeny systematics taxonomy nomenclature homonym synonym junior-synonym senior-synonym valid-name accepted-name nominal-species species-inquirenda species-complex sibling-species cryptic-species morphospecies biological-species evolutionary-species phylogenetic-species operational-taxonomic-unit OTU DNA-barcoding molecular-phylogenetics integrative-taxonomy alpha-taxonomy beta-taxonomy gamma-taxonomy monograph revision redescription diagnosis description illustration measurement meristic morphometric allometry ontogeny heterochrony paedomorphosis peramorphosis development embryology larval-development zoea megalopa settlement juvenile immature mature adult subadult gerontic senescence longevity life-span life-history life-history-strategy r-selection K-selection bet-hedging reproductive-effort reproductive-success fitness population demography cohort generation age-structure size-structure stage-structure population-regulation density-dependence density-independence carrying-capacity limiting-factor environmental-gradient habitat-gradient depth-gradient zonation vertical-distribution horizontal-distribution geographic-distribution latitudinal-gradient longitudinal-gradient bathymetric-distribution intertidal-zone subtidal-zone continental-shelf continental-slope abyssal-zone hadal-zone neritic-zone oceanic-zone pelagic-zone benthic-zone demersal-zone epipelagic mesopelagic bathypelagic abyssopelagic hadopelagic supralittoral circalittoral archibenthal bathyal abyssal hadal euphotic dysphotic aphotic photic-zone disphotic-zone aphotic-zone trophic-level primary-producer primary-consumer secondary-consumer tertiary-consumer detritivore scavenger predator herbivore carnivore omnivore parasite commensal mutualist symbiont competitor food-chain trophic-cascade bottom-up-control top-down-control wassefug mesopredator-release trophic-trickle nutrient-cycling energy-flow productivity biomass standing-stock turnover production respiration excretion egestion assimilation efficiency ecological-efficiency trophic-transfer-efficiency Lindeman-efficiency 10%-rule pyramid-of-numbers pyramid-of-biomass pyramid-of-energy ecosystem-function ecosystem-service ecosystem-process biogeochemical-cycle carbon-cycle nitrogen-cycle phosphorus-cycle sulfur-cycle oxygen-cycle hydrogen-cycle mineral-cycle nutrient limiting-nutrient eutrophication oligotrophication hypoxia anoxia acidification warming climate-change global-change anthropogenic-impact pollution contamination habitat-loss habitat-degradation habitat-fragmentation overfishing bycatch invasive-species introduced-species non-native-species alien-species exotic-species range-expansion range-contraction range-shift poleward-shift tropicalization borealization meridionalization deglaciation sea-level-rise coastal-squeeze ocean-acidification ocean-warming deoxygenation marine-heatwave extreme-event disturbance resilience resistance recovery adaptation acclimation plasticity evolutionary-adaptation genetic-adaptation phenotypic-plasticity behavioral-plasticity physiological-plasticity morphological-plasticity life-history-plasticity trade-off constraint limitation bottleneck bottleneck-effect founder-effect genetic-drift gene-flow mutation selection natural-selection sexual-selection artificial-selection directional-selection stabilizing-selection disruptive-selection frequency-dependent-selection density-dependent-selection rare-advantage common-advantage apostatic-selection balancing-selection heterozygote-advantage negative-frequency-dependent-selection positive-frequency-dependent-selection ecological-speciation sympatric-speciation allopatric-speciation parapatric-speciation peripatric-speciation vicariance dispersal colonization establishment invasion naturalization spreading epidemic pandemic enzootic epizootic eradication control protection restoration rehabilitation rewilding de-extinction sustainable-use ecosystem-based-management adaptive-management precautionary-approach ecosystem-approach integrated-management marine-protected-area MPA no-take-zone marine-reserve sanctuary world-heritage-site Ramsar-site important-marine-mammal-area IMMA ecologically-or-biologically-significant-area EBSA vulnerable-marine-ecosystem VME essential-fish-habitat EFH critical-habitat habitat-of-particular-concern habitat-area-of-particular-concern HAPC key-biodiversity-area KBA protected-area conservation-area nature-reserve national-park marine-park aquatic-reserve fishery-reserve species-recovery-plan action-plan management-plan strategic-plan policy legislation regulation convention treaty agreement protocol Memorandum-of-Understanding MoU non-binding-instrument soft-law hard-law customary-international-law jus-cogens erga-omnes common-concern-of-humankind common-heritage-of-mankind precautionary-principle polluter-pays-principle common-but-differentiated-responsibilities intergenerational-equity intragenerational-equity environmental-justice climate-justice ocean-justice blue-economy green-economy circular-economy bioeconomy natural-capital ecosystem-services-valuation payment-for-ecosystem-services PES blue-carbon carbon-sequestration carbon-storage carbon-sink carbon-source greenhouse-gas GHG carbon-dioxide CO2 methane CH4 nitrous-oxide N2O fluorinated-gas F-gas hydrofluorocarbon HFC perfluorocarbon PFC sulfur-hexafluoride SF6 nitrogen-trifluoride NF3 ozone-depleting-substance ODS Montreal-Protocol Paris-Agreement UNFCCC IPCC IPBES IOC UNESCO FAO IMO ISA CBD CMS CITES RAMSAR World-Heritage-Convention UNFSA Straddling-Stocks-Agreement Compliance-Agreement Port-State-Measures-Agreement PSMA IUU-fishing illegal-unreported-unregulated-fishing overcapacity fishing-down-the-food-web fishing-through-the-food-web serial-depletion shifting-baseline baseline reference-point target-reference-point limit-reference-point threshold tipping-point point-of-no-return hysteresis alternative-stable-state regime-shift critical-transition catastrophic-shift smooth-transition adaptive-cycle panarchy resilience-thinking social-ecological-system SES coupled-human-natural-system CHANS human-environment-system HES socio-ecological-system ecosystem-approach-to-fisheries EAF ecosystem-based-fisheries-management EBFM EBM integrated-ecosystem-assessment IEA marine-spatial-planning MSP ocean-zoning marine-planning coastal-zone-management integrated-coastal-zone-management ICZM integrated-maritime-policy IMP blue-growth sustainable-development-goal SDG SDG-14 life-below-water decade-of-ocean-science UN-Ocean-Decade 2030-Agenda Agenda-21 Rio-Declaration Stockholm-Declaration Johannesburg-Plan-of-Implementation World-Summit-on-Sustainable-Development WSSD Rio+20 Rio+30 Earth-Summit UNCED UNCLOS Law-of-the-Sea-Convention Convention-on-the-Law-of-the-Sea high-seas area-beyond-national-jurisdiction ABNJ international-seabed-area the-Area exclusive-economic-zone EEZ territorial-sea contiguous-zone archipelagic-waters internal-waters straight-baseline normal-baseline low-water-line tidal-datum chart-datum depth-contour isobath bathymetry topography morphology geomorphology seafloor seabed benthos nekton plankton neuston pleuston hyperbenthos suprabenthos epibenthos endobenthos infauna epifauna pelagic benthopelagic meroplankton holoplankton zooplankton phytoplankton bacterioplankton virioplankton ichthyoplankton ichthyoplankton-survey egg-survey larval-survey ichthyoplankton-transport larval-transport dispersal-kernel connectivity population-connectivity genetic-connectivity demographic-connectivity larval-pool source-sink-dynamics metapopulation population-structure stock-structure stock-identification fishery-assessment biological-reference-point BRP maximum-sustainable-yield MSY optimal-yield OY maximum-economic-yield MEY maximum-constant-yield MCY equilibrium-yield sustainable-yield surplus-production Schaefer-model Fox-model Pella-Tomlinson-model surplus-production-model analytical-model statistical-catch-at-age-model SCA virtual-population-analysis VPA cohort-analysis catch-curve length-based-model age-based-model stage-based-model size-based-model trait-based-model individual-based-model IBM agent-based-model ABM ecosystem-model end-to-end-model EwE Atlantis OSMOSE InVEST Marxan prioritization systematic-conservation-planning SCP gap-analysis representativeness adequacy effectiveness equity fairness legitimacy compliance enforcement monitoring surveillance observation data-collection sampling survey census index indicator metric variable parameter estimate prediction forecast projection scenario what-if-analysis sensitivity-analysis uncertainty-analysis risk-analysis decision-analysis trade-off-analysis optimization multi-objective-optimization Pareto-front Pareto-optimal non-dominated-solution stakeholder interest-group rights-holder user-group community indigenous-people local-knowledge traditional-knowledge TK traditional-ecological-knowledge TEK citizen-science participatory-science community-based-monitoring CBM co-management joint-management shared-governance adaptive-co-management transboundary-management regional-seas regional-fishery-body RFB regional-fishery-management-organization RFMO arrangement commission council committee working-group expert-group scientific-committee technical-committee compliance-committee dispute-settlement arbitration conciliation mediation negotiation bargaining coalition alliance partnership network platform forum conference meeting session symposium workshop seminar training capacity-building technology-transfer technical-assistance financial-assistance funding grant loan donation collaboration cooperation coordination harmonization standardization best-practice guideline manual handbook code-of-conduct code-of-practice certification eco-label seal-of-approval sustainability-standard marine-stewardship-council MSC aquaculture-stewardship-council ASC friend-of-the-sea global-G.A.P.BRC IFS SQF FSSC-22000 ISO-22000 HACCP GMP SSOP traceability chain-of-custody blockchain digital-traceability electronic-monitoring EM electronic-reporting ER VMS AIS satellite-monitoring remote-sensing earth-observation geographic-information-system GIS global-positioning-system GPS automatic-identification-system vessel-monitoring-system catch-documentation-scheme CDS port-state-measures catch-certification trade-tracking isotope-analysis otolith-microchemistry parasite-tag tagging mark-recapture telemetry acoustic-telemetry satellite-telemetry archival-tag data-storage-tag DST pop-up-satellite-archival-tag PSAT smart-tag biologging bio-telemetry sensor accelerometer magnetometer gyroscope pressure-sensor temperature-sensor light-sensor dissolved-oxygen-sensor salinity-sensor conductivity-sensor pH-sensor chlorophyll-sensor turbidity-sensor acoustic-Doppler-current-profiler ADCP echosounder multibeam-echosounder single-beam-echosounder split-beam-echosounder scientific-echosounder fishing-echosounder sonar side-scan-sonar synthetic-aperture-sonar SAS sub-bottom-profiler seismic-profiler gravimeter bathymetric-lidar airborne-lidar satellite-altimetry radar-altimetry laser-altimetry photogrammetry structure-from-motion SfM underwater-photogrammetry video-analysis image-analysis machine-learning artificial-intelligence AI deep-learning neural-network computer-vision pattern-recognition object-detection instance-segmentation semantic-segmentation classification clustering regression forecasting time-series-analysis spatial-analysis geostatistics kriging inverse-distance-weighting IDW triangulated-irregular-network TIN digital-elevation-model DEM digital-terrain-model DTM digital-surface-model DSM habitat-suitability-model species-distribution-model SDM ecological-niche-model ENM habitat-suitability-index HSI environmental-suitability bioclimatic-envelope climate-envelope fundamental-niche realized-niche potential-distribution actual-distribution range-map distribution-map occurrence-record presence-only-data presence-absence-data abundance-data density-data effort-data detection-probability occupancy-model N-mixture-model distance-sampling line-transect point-transect strip-transect plotless-sampling quadrat transect stratified-random-sampling systematic-sampling cluster-sampling adaptive-cluster-sampling two-phase-sampling double-sampling multi-phase-sampling sampling-design sample-size statistical-power effect-size significance-level confidence-level confidence-interval credible-interval prediction-interval tolerance-interval hypothesis-test null-hypothesis alternative-hypothesis type-I-error type-II-error false-positive false-negative sensitivity specificity accuracy precision recall F1-score ROC-curve AUC model-validation model-verification model-calibration model-selection model-comparison model-averaging ensemble-model multi-model-inference information-criterion AIC AICc BIC DIC WAIC LOOIC cross-validation k-fold-cross-validation leave-one-out-cross-validation bootstrap jackknife permutation-test randomization-test Monte-Carlo-simulation Markov-chain-Monte-Carlo MCMC Bayesian-inference frequentist-inference likelihood posterior-probability prior-probability prior-distribution posterior-distribution conjugate-prior non-informative-prior weakly-informative-prior hierarchical-model mixed-effects-model random-effects-model fixed-effects-model generalized-linear-model GLM generalized-additive-model GAM generalized-linear-mixed-model GLMM generalized-additive-mixed-model GAMM structural-equation-model SEM path-analysis confirmatory-factor-analysis CFA exploratory-factor-analysis EFA principal-component-analysis PCA principal-coordinate-analysis PCoA non-metric-multidimensional-scaling NMDS correspondence-analysis CA detrended-correspondence-analysis DCA redundancy-analysis RDA canonical-correspondence-analysis CCA partial-least-squares PLS orthogonal-partial-least-squares OPLS artificial-neural-network ANN random-forest RF gradient-boosting-machine GBM XGBoost LightGBM CatBoost support-vector-machine SVM k-nearest-neighbor KNN naive-Bayes decision-tree regression-tree classification-tree bagging boosting stacking ensembling voting blending meta-learning transfer-learning domain-adaptation few-shot-learning zero-shot-learning self-supervised-learning unsupervised-learning supervised-learning semi-supervised-learning reinforcement-learning active-learning online-learning incremental-learning lifelong-learning continual-learning federated-learning distributed-learning edge-computing cloud-computing high-performance-computing HPC parallel-computing GPU-computing quantum-computing neuromorphic-computing DNA-computing biological-computing natural-computing swarm-intelligence ant-colony-optimization particle-swarm-optimization genetic-algorithm evolutionary-algorithm differential-evolution simulated-annealing tabu-search variable-neighborhood-search iterated-local-search guided-local-search greedy-randomized-adaptive-search-procedure GRASP scatter-search path-relinking memetic-algorithm hyper-heuristic meta-heuristic optimization-heuristic constructive-heuristic improvement-heuristic local-search neighborhood-search large-neighborhood-search LNS adaptive-large-neighborhood-search ALNS branch-and-bound branch-and-cut branch-and-price column-generation Benders-decomposition Dantzig-Wolfe-decomposition Lagrangian-relaxation surrogate-relaxation dual-ascent primal-dual-method interior-point-method simplex-method revised-simplex-method dual-simplex-method network-simplex-method barrier-method penalty-method augmented-Lagrangian-method sequential-quadratic-programming SQP sequential-linear-programming SLP trust-region-method line-search-method gradient-descent steepest-descent conjugate-gradient quasi-Newton-method Newton-method Gauss-Newton-method Levenberg-Marquardt-method trust-region-reflective-method interior-point-reflective-method active-set-method projected-gradient-method proximal-gradient-method accelerated-gradient-method momentum-method Nesterov-acceleration AdaGrad RMSprop Adam AdamW SGD stochastic-gradient-descent mini-batch-gradient-descent batch-gradient-descent full-gradient-descent variance-reduction SAG SAGA SVRG L-BFGS limited-memory-BFGS BFGS DFP SR1 Nelder-Mead-method Powell's-method Hooke-Jeeves-method pattern-search direct-search derivative-free-optimization black-box-optimization bayesian-optimization Gaussian-process kriging-surrogate response-surface-method RSM polynomial-chaos-expansion PCE sparse-grid sparse-polynomial-approximation low-rank-approximation tensor-decomposition CP-decomposition Tucker-decomposition tensor-train TT hierarchical-Tucker HT matrix-factorization non-negative-matrix-factorization NMF singular-value-decomposition SVD independent-component-analysis ICA factor-analysis latent-variable-model latent-class-model finite-mixture-model hidden-Markov-model HMM state-space-model dynamic-linear-model DLM Kalman-filter particle-filter sequential-Monte-Carlo SMC ensemble-Kalman-filter EnKF unscented-Kalman-filter UKF extended-Kalman-filter EKF Rauch-Tung-Striebel-smoother RTS-smoother forward-backward-algorithm Viterbi-algorithm Baum-Welch-algorithm expectation-maximization EM-algorithm variational-inference variational-Bayes mean-field-approximation loopy-belief-propagation sum-product-algorithm max-product-algorithm junction-tree-algorithm bucket-elimination variable-elimination conditioning cutset-conditioning constraint-satisfaction CSP satisfiability SAT MAX-SAT pseudo-Boolean-optimization integer-linear-programming ILP mixed-integer-linear-programming MILP mixed-integer-nonlinear-programming MINLP quadratic-programming QP quadratically-constrained-quadratic-programming QCQP second-order-cone-programming SOCP semidefinite-programming SDP geometric-programming GP signomial-programming SP difference-of-convex-programming DC-programming complementary-geometric-programming CGP generalized-geometric-programming GGP posynomial monomial signomial polynomial rational-function algebraic-function transcendental-function analytic-function meromorphic-function entire-function holomorphic-function harmonic-function subharmonic-function superharmonic-function plurisubharmonic-function plurisuperharmonic-function convex-function concave-function quasiconvex-function quasiconcave-function log-convex-function log-concave-function geometrically-convex-function multiplicatively-convex-function operator-convex-function matrix-convex-function spectral-function unitarily-invariant-function orthogonally-invariant-function permutation-invariant-function symmetric-function Schur-convex-function Schur-concave-function majorization weak-majorization strong-majorization doubly-stochastic-matrix Birkhoff-polytope permutahedron transportation-polytope network-flow minimum-cost-flow maximum-flow minimum-cut shortest-path longest-path traveling-salesman-problem TSP vehicle-routing-problem VRP capacitated-VRP CVRP VRP-with-time-windows VRPTW pickup-and-delivery-problem PDP dial-a-ride-problem DARP arc-routing-problem ARP Chinese-postman-problem CPP rural-postman-problem RPP location-problem facility-location p-median-problem p-center-problem uncapacitated-facility-location UFL capacitated-facility-location CFL warehouse-location plant-location hub-location median-problem center-problem covering-problem set-covering vertex-covering edge-covering dominating-set independent-set clique graph-coloring chromatic-number clique-number independence-number domination-number matching perfect-matching maximum-matching b-matching f-factor g-factor network-design Steiner-tree Steiner-forest prize-collecting-Steiner-tree k-MST minimum-spanning-tree MST maximum-spanning-tree minimum-arborescence minimum-directed-spanning-tree minimum-Steiner-tree approximation-algorithm approximation-ratio performance-guarantee inapproximability NP-hardness NP-completeness computational-complexity time-complexity space-complexity polynomial-time exponential-time quasi-polynomial-time sub-exponential-time fixed-parameter-tractable FPT W-hierarchy parameterized-complexity kernelization kernel treewidth pathwidth branchwidth cliquewidth rankwidth booleanwidth modularwidth split-decomposition modular-decomposition prime-graph cograph permutation-graph interval-graph chordal-graph perfect-graph comparability-graph co-comparability-graph AT-free-graph claw-free-graph line-graph planar-graph outerplanar-graph series-parallel-graph treewidth-bounded-graph minor-free-graph forbidden-minor forbidden-topological-minor forbidden-induced-subgraph graph-minor-theorem Robertson-Seymour-theorem well-quasi-ordering Wagner's-conjecture Hadwiger's-conjecture Four-Color-Theorem Strong-Perfect-Graph-Theorem Erdős–Faber–Lovász-conjecture Gyárfás–Sumner-conjecture Erdős–Hajnal-conjecture Caccetta–Häggkvist-conjecture Bermond–Thomassen-conjecture Thomassen's-conjecture Tutte's-conjecture Grötzsch's-theorem Brooks'-theorem Vizing's-theorem Shannon's-theorem König's-theorem Kőnig's-line-coloring-theorem Menger's-theorem max-flow-min-cut-theorem Ford-Fulkerson-theorem Edmonds'-matching-theorem Tutte's-theorem Petersen's-theorem Hall's-marriage-theorem Dilworth's-theorem Mirsky's-theorem Sperner's-theorem Erdős–Ko–Rado-theorem Kruskal–Katona-theorem Lovász's-perfect-graph-theorem Chudnovsky's-strong-perfect-graph-theorem Seymour's-decomposition-theorem Robertson–Seymour–Thomas-theorem Heawood-conjecture Ringel–Youngs-theorem Map-Color-Theorem Appel–Haken-proof Robertson–Seymour–Sanders–Thomas-proof computer-assisted-proof formal-proof interactive-theorem-proving proof-assistant Coq Isabelle HOL HOL-Light Lean Agda Twelf Mizar Metamath automated-theorem-proving SAT-solver SMT-solver model-checker temporal-logic CTL LTL CTL*modal-mu-calculus fixed-point-logic description-logic OWL RDF SPARQL knowledge-graph ontology semantic-web linked-data open-data FAIR-data findable accessible interoperable reusable data-management-plan DMP metadata provenance data-quality data-curation data-preservation data-archiving data-repository institutional-repository disciplinary-repository general-repository figshare Zenodo Dryad GBIF OBIS WoRMS Catalogue-of-Life COL ITIS NCBI BOLD iNaturalist eBird eButterfly iDigBio ALA AVH BIE BISON Map-of-Life MOL Movebank OTN SAHFOS CPR ICES NAFO NEAFC SEAFO WCPFC IATTC ICCAT IOTC CCSBT CCAMLR GFCM CWP CWP-15 CWP-16 CWP-17 CWP-18 CWP-19 CWP-20 CWP-21 CWP-22 CWP-23 CWP-24 CWP-25 CWP-26 CWP-27 CWP-28 CWP-29 CWP-30 CWP-31 CWP-32 CWP-33 CWP-34 CWP-35 CWP-36 CWP-37 CWP-38 CWP-39 CWP-40 CWP-41 CWP-42 CWP-43 CWP-44 CWP-45 CWP-46 CWP-47 CWP-48 CWP-49 CWP-50 CWP-51 CWP-52 CWP-53 CWP-54 CWP-55 CWP-56 CWP-57 CWP-58 CWP-59 CWP-60 CWP-61 CWP-62 CWP-63 CWP-64 CWP-65 CWP-66 CWP-67 CWP-68 CWP-69 CWP-70 CWP-71 CWP-72 CWP-73 CWP-74 CWP-75 CWP-76 CWP-77 CWP-78 CWP-79 CWP-80 CWP-81 CWP-82 CWP-83 CWP-84 CWP-85 CWP-86 CWP-87 CWP-88 CWP-89 CWP-90 CWP-91 CWP-92 CWP-93 CWP-94 CWP-95 CWP-96 CWP-97 CWP-98 CWP-99 CWP-100 CWP-101 CWP-102 CWP-103 CWP-104 CWP-105 CWP-106 CWP-107 CWP-108 CWP-109 CWP-110 CWP-111 CWP-112 CWP-113 CWP-114 CWP-115 CWP-116 CWP-117 CWP-118 CWP-119 CWP-120 CWP-121 CWP-122 CWP-123 CWP-124 CWP-125 CWP-126 CWP-127 CWP-128 CWP-129 CWP-130 CWP-131 CWP-132 CWP-133 CWP-134 CWP-135 CWP-136 CWP-137 CWP-138 CWP-139 CWP-140 CWP-141 CWP-142 CWP-143 CWP-144 CWP-145 CWP-146 CWP-147 CWP-148 CWP-149 CWP-150 CWP-151 CWP-152 CWP-153 CWP-154 CWP-155 CWP-156 CWP-157 CWP-158 CWP-159 CWP-160 CWP-161 CWP-162 CWP-163 CWP-164 CWP-165 CWP-166 CWP-167 CWP-168 CWP-169 CWP-170 CWP-171 CWP-172 CWP-173 CWP-174 CWP-175 CWP-176 CWP-177 CWP-178 CWP-179 CWP-180 CWP-181 CWP-182 CWP-183 CWP-184 CWP-185 CWP-186 CWP-187 CWP-188 CWP-189 CWP-190 CWP-191 CWP-192 CWP-193 CWP-194 CWP-195 CWP-196 CWP-197 CWP-198 CWP-199 CWP-200
Riccardoella
slug mites, snail mites
Riccardoella is a genus of parasitic mites in the family Ereynetidae that are obligate ectoparasites of terrestrial gastropod molluscs, primarily slugs and snails. The genus contains approximately six described species distributed across Europe, North America, East Asia, and New Zealand. These mites inhabit host mantle cavities, respiratory pores, and lung spaces, feeding on host mucus and tissue fluids. Their life cycles are tightly coupled to host biology, with developmental stages periodically leaving hosts to moult before reinfesting.
parasite gastropod-parasite Ereynetidae slug-mite snail-mite temporary-ectoparasite Prostigmata
Salmincola
Salmincola is a genus of freshwater parasitic copepods in the family Lernaeopodidae. These ectoparasites are common on salmonid fishes throughout the Northern Hemisphere. The genus exhibits substantial geographic variation in infection rates across water bodies, and current taxonomic definitions based on decades-old morphological descriptions may obscure cryptic diversity. Species in this genus are considered pests by fisheries management agencies due to their impact on host fish populations.
copepod parasite salmonid freshwater fisheries-pest cryptic-diversity host-specificity Siberia Lake-Baikal nasal-parasite
Salmincola californiensis
gill lice, gill maggot, salmon gill maggot
Salmincola californiensis is a freshwater parasitic copepod in the family Lernaeopodidae that infests the gills and branchial cavities of salmonid fishes, particularly species of Oncorhynchus. Native to the Pacific Rim and now established in introduced ranges including Japan and Colorado, this parasite has become a significant concern for fisheries management and salmonid conservation. Recent research has documented a previously unrecognized naupliar stage and demonstrated strong temperature-dependence in egg development and copepodid survival, with implications for disease management under climate change scenarios.
parasite copepod salmonid fisheries invasive temperature-dependent vector reservoir
Sandalus petrophya
cedar beetle
Sandalus petrophya, commonly known as the cedar beetle, is a species of cicada parasite beetle in the family Rhipiceridae. It is found in North America. The species is distinguished from the similar Sandalus niger by a distinct vertical ridge in the middle of its pronotum.
parasite cicada-parasite North-America
Sandalus porosus
cicada parasite beetle
Sandalus porosus is a species of cicada parasite beetle in the family Rhipiceridae. It is found in North America. The species is part of a small genus of beetles known for their parasitic relationship with cicadas.
parasite cicada-parasite Rhipiceridae North-America
Sarcoptes
itch mite, mange mite
Sarcoptes is a genus of parasitic mites in the family Sarcoptidae. The genus contains species that are permanent obligate ectoparasites of mammals, most notably *Sarcoptes scabiei*, which causes scabies in humans and sarcoptic mange in livestock, wildlife, and companion animals. These mites burrow into the epidermis of their hosts, where they feed, reproduce, and complete their entire life cycle. Infestations cause intense pruritus, skin lesions, and significant economic and health impacts globally. Taxonomic treatment varies: some authorities recognize multiple host-specific varieties of *S. scabiei*, while others treat these as distinct species.
parasite mite scabies mange ectoparasite veterinary medical zoonosis neglected-tropical-disease acaricide-resistance
Sarcoptes scabiei
Scabies Mite, Itch Mite
Sarcoptes scabiei is a parasitic mite that burrows into the epidermis of mammals, causing scabies in humans and sarcoptic mange in animals. The mite has a cosmopolitan distribution and infests numerous host species including humans, domestic animals, and wildlife. Adult females create characteristic S-shaped burrows in the stratum corneum where they deposit eggs. The disease produces intense pruritus through host immune responses to mites, eggs, and fecal pellets. Different host-adapted variants exist, though cross-infectivity between closely related hosts has been documented.
parasite mite scabies sarcoptic-mange ectoparasite medical-entomology veterinary-entomology public-health zoonosis burrowing-mite
Sarcoptidae
Itch Mites
Sarcoptidae is a family of parasitic mites in the order Sarcoptiformes, commonly known as itch mites. The family includes the genus Sarcoptes, whose members cause sarcoptic mange in mammals. These mites are obligate parasites that burrow into host skin, causing intense pruritus, crusting lesions, and potentially fatal secondary infections. Sarcoptes scabiei, the most studied species, infests over 200 million humans worldwide and numerous domestic and wild mammal species.
parasite mite scabies mange zoonosis veterinary wildlife-disease neglected-tropical-disease burrowing epidermal-parasite direct-life-cycle host-specific-strains immunology emerging-infectious-disease biodiversity-threat cross-species-transmission feral-dog-reservoir quantitative-surveillance tissue-digestion-protocol ivermectin-treatment crusted-scabies ordinary-scabies pruritus alopecia hyperkeratosis secondary-bacterial-infection WHO-neglected-tropical-disease over-200-million-human-cases mammalian-host-specificity Astigmata Sarcoptiformes Arachnida Acari
Siphonostomatoida
Siphon-mouth Copepods
Siphonostomatoida is an order of copepods distinguished by siphon-like mandibles and a frontal filament used for host attachment. The order comprises 40 recognized families, with approximately 75% of all fish-parasitizing copepods belonging to this group. Most species are marine symbionts, though a few inhabit freshwater environments. Members exhibit diverse host associations, with 17 families parasitizing vertebrates (primarily fishes) and 23 families associated with invertebrates.
copepod parasite marine fish-parasite symbiont crustacean ectoparasite Siphonostomatoida biodiversity Southern-Africa Japan Brazil sponge-associate coral-associate developmental-stages chalimus frontal-filament siphon-mandible Asterocheridae Lernaeopodidae Pennellidae Hatschekiidae Sphyriidae Notodelphyidae Botryllophilidae
Speleognathinae
Speleognathinae is a subfamily of mites in the family Ereynetidae comprising endoparasites of the respiratory tracts of birds. These mites inhabit the nasal cavities and associated passages of avian hosts across multiple continents. At least 37 species in 15 genera have been documented from the Afrotropical region alone. The subfamily is distinguished from related nasal mite taxa by morphological features used in taxonomic keys.
mite parasite avian-parasite nasal-mite endoparasite respiratory-parasite
Tortopus
Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.
Ephemeroptera Polymitarcyidae burrowing-mayflies Neotropical clay-bank-habitat U-shaped-tunnels mandibular-tusks Tortopsis Campsurinae Panamerican-distribution cladistic-revision synapomorphy parastyli-receptors flattened-penes furcasternal-protuberances subapical-tubercles limnological-sampling-bias nymph-morphology adult-morphology type-species-T.-igaranus new-species-T.-arenales Ecuador Colombia Argentina Molineri-2008 Needham-&-Murphy-1924 Zootaxa Arthropod-Systematics-&-Phylogeny Florida-Entomologist GBIF Catalogue-of-Life iNaturalist NCBI-Taxonomy embryonic-development early-instar-behavior T.-harrisi T.-bellus T.-circumfluus T.-zottai T.-igaranus T.-incertus T.-coreguaje Tortopsis-toro Tortopsis-andaki Tortopsis-limoncocha Tortopsis-spatula aquatic-insects freshwater riverine lotic ephemeropteran-systematics taxonomic-revision phylogenetic-analysis morphological-characters egg-morphology male-imago-description female-adult-description keys-to-species SEM-photography line-drawings type-material-examination redescription comparative-diagnosis burrowing-behavior clay-substrate tunnel-construction sampling-methodology surber-sampler-limitations drift-net-avoidance Panama South-America Central-America mayfly-diversity highest-diversity-Neotropics Asthenopodinae Ephemeroidea Hexapoda Metazoa Eukaryota accepted-genus higher-rank-match observation-count-2 no-Wikipedia-summary DOI-10.11646/zootaxa.2481.1.1 DOI-10.11646/zootaxa.2436.1.4 DOI-10.3897/asp.79.e62735 DOI-10.2307/3493493 Scott-et-al.-1959 Navás Ulmer Traver Lugo-Ortiz-&-McCafferty Domínguez Banks McDunnough Pictet Needham-&-Murphy freshwater-ecology aquatic-entomology insect-development nymphal-stages imago-description alate-adults wing-venation R-sector gonopore-structure parastyli-length pedestal-morphology lateral-view-curvature mandibular-morphology tusk-tubercles mesosternal-structure abdominal-sternum-notch furcasternum protuberance-configuration receptor-socket-orientation IR-vein R2-vein claw-like-structure gonopore-association penis-base-separation female-fore-wing additional-veins comparative-morphology type-specimen holotype allotype paratype neotropical-species new-species-description species-key identification-key male-adult-key female-adult-key nymph-key generic-key Polymitarcyidae-genera systematics phylogeny cladistics monophyly sister-group character-state transformation-series parsimony-analysis morphological-data-set external-morphology egg-chorion micropyle egg-burster hatching emergence behavioral-study early-instar embryology oviposition substrate-preference habitat-specificity endemic regional-diversity conservation biomonitoring water-quality-indicator macroinvertebrate benthic infaunal interstitial hyporheic river-bank stream-bank erosion sediment particle-size clay-content bank-stability root-mat vegetation-association flow-regime current-velocity depth width stream-order catchment basin ecoregion biogeography dispersal colonization vicariance speciation adaptive-radiation morphological-divergence convergent-evolution homoplasy character-conflict bootstrap-support Bremer-support consensus-tree strict-consensus majority-rule taxonomic-stability nomenclatural-act original-description subsequent-designation type-locality type-depository museum-collection specimen-preservation alcohol-preservation slide-mounting SEM-preparation imaging-techniques scientific-illustration photography measurement morphometrics ratios proportions allometry sexual-dimorphism geographic-variation intraspecific-variation interspecific-variation diagnostic-characters identification-confidence taxonomic-expertise literature-review bibliographic-search database-query online-repository open-access paywall abstract-only full-text-unavailable citation-analysis reference-list source-quality evidence-strength knowledge-gap research-need future-study collaboration field-work collection-trip sampling-effort specimen-availability type-examination revision-scope taxonomic-concept circumscription inclusion exclusion species-group species-complex cryptic-species morphospecies phylogenetic-species evolutionary-significant-unit operational-taxonomic-unit DNA-barcoding molecular-systematics integrative-taxonomy traditional-taxonomy alpha-taxonomy beta-taxonomy gamma-taxonomy classification-hierarchy Linnaean-system rank-based-nomenclature phylogenetic-nomenclature clade-name crown-group stem-group total-evidence congruence incongruence data-partition character-weighting implied-weighting equal-weighting ordered-characters unordered-characters multistate-characters binary-characters continuous-characters meristic-characters qualitative-characters quantitative-characters morphological-matrix data-matrix character-matrix taxon-sampling character-sampling missing-data inapplicable-data polymorphism intraspecific-polymorphism fixed-difference diagnostic-difference unique-derived-character autapomorphy shared-derived-character symplesiomorphy convergent-character parallel-evolution reversal character-optimization ACCTRAN DELTRAN unambiguous-optimization ambiguous-optimization most-parsimonious-tree tree-length consistency-index retention-index rescaled-consistency-index homoplasy-index character-fit character-congruence character-support branch-support node-support clade-stability taxonomic-stability-index phylogenetic-signal noise saturation substitution-model rate-heterogeneity among-site-rate-variation gamma-distribution invariant-sites model-selection Akaike-information-criterion Bayesian-information-criterion likelihood-ratio-test Bayesian-inference Markov-chain-Monte-Carlo posterior-probability prior-probability hyperprior convergence effective-sample-size burn-in stationarity trace-plot parameter-estimate credible-interval highest-posterior-density tree-topology tree-shape lineage-through-time diversification-rate extinction-rate speciation-rate net-diversification birth-death-model Yule-model coalescent-model population-size effective-population-size gene-flow migration-rate isolation-by-distance phylogeography historical-biogeography ancestral-area-reconstruction dispersal-vicariance-analysis decisive-weighting event-based-method taxon-area-cladogram area-cladogram biogeographic-homology track-analysis panbiogeography phylogenetic-biogeography ecological-biogeography dispersal-biogeography vicariance-biogeography center-of-origin area-of-endemism refugium Pleistocene glaciation climate-change habitat-fragmentation habitat-loss anthropogenic-impact pollution eutrophication sedimentation dam-construction water-abstraction flow-modification thermal-pollution toxic-substances heavy-metals pesticides urbanization agriculture deforestation land-use-change riparian-degradation channelization dredging mining invasive-species biological-invasion competition predation parasitism mutualism commensalism amensalism facilitation ecosystem-engineer keystone-species flagship-species umbrella-species indicator-species sentinel-species bioindicator ecological-indicator environmental-indicator water-quality-index biotic-index multimetric-index rapid-bioassessment protocol standardization quality-assurance quality-control reference-condition least-disturbed minimally-disturbed best-available type-specific region-specific stream-type river-type functional-feeding-group collector-gatherer collector-filterer scraper shredder predator parasite engulfer piercer burrower sprawler clinger swimmer skater diver surface-dweller benthic-dweller interstitial-dweller hyporheic-dweller phreatic-dweller troglobite stygobite cave-inhabitant spring-inhabitant headwater mid-order high-order lowland montane alpine subtropical tropical temperate boreal arctic antarctic palearctic nearctic afrotropical oriental australian oceanic cosmopolitan widespread restricted relict disjunct-distribution range-extension new-record first-record country-record state-record province-record locality-record elevation-record altitudinal-range latitudinal-range longitudinal-range distribution-pattern range-size range-fill extent-of-occurrence area-of-occupancy IUCN-category threat-status data-deficient not-evaluated least-concern near-threatened vulnerable endangered critically-endangered extinct-in-wild extinct population-trend population-density population-structure age-structure size-structure sex-ratio reproductive-output fecundity fertility mortality survivorship life-table cohort-analysis generation-time doubling-time intrinsic-rate-of-increase finite-rate-of-increase net-reproductive-rate stable-age-distribution stationary-population expanding-population declining-population source-sink-dynamics metapopulation population-viability-analysis minimum-viable-population genetic-diversity heterozygosity allelic-richness private-allele founder-effect bottleneck genetic-drift inbreeding inbreeding-depression outbreeding outbreeding-depression hybridization introgression gene-flow-barrier reproductive-isolation prezygotic-isolation postzygotic-isolation mechanical-isolation temporal-isolation habitat-isolation behavioral-isolation gametic-isolation zygotic-mortality hybrid-inviability hybrid-sterility hybrid-breakdown reinforcement character-displacement ecological-speciation allopatric-speciation sympatric-speciation parapatric-speciation peripatric-speciation centrifugal-speciation divergence-with-gene-flow speciation-continuum anagenesis cladogenesis budding-speciation branching-speciation phyletic-gradualism punctuated-equilibrium stasis trend progression orthogenesis aristogenesis mosaic-evolution co-evolution arms-race Red-Queen-hypothesis evolutionary-stable-strategy frequency-dependent-selection density-dependent-selection r-selection K-selection bet-hedging iteroparity semelparity ploidy-level chromosome-number karyotype genome-size transposable-element gene-duplication neofunctionalization subfunctionalization pseudogene regulatory-evolution cis-regulatory trans-regulatory enhancer promoter silencer insulator transcription-factor signaling-pathway developmental-pathway gene-network epigenetic-modification DNA-methylation histone-modification chromatin-remodeling genomic-imprinting parental-effect maternal-effect paternal-effect cytoplasmic-inheritance mitochondrial-DNA chloroplast-DNA endosymbiont Wolbachia Rickettsia Spiroplasma Cardinium Arsenophonus Hamiltonella Regiella Serratia Buchnera Carsonella Portiera primary-endosymbiont secondary-endosymbiont facultative-symbiont obligate-symbiont mutualistic-symbiont parasitic-symbiont commensal-symbiont pathogen vector disease-transmission zoonosis emerging-infectious-disease biosecurity quarantine eradication control management conservation-biology restoration-ecology habitat-restoration river-restoration stream-restoration floodplain-restoration riparian-restoration wetland-restoration ecosystem-restoration passive-restoration active-restoration assisted-recovery managed-relocation assisted-colonization ex-situ-conservation in-situ-conservation protected-area national-park nature-reserve wildlife-refuge biosphere-reserve world-heritage-site ramsar-site important-bird-area key-biodiversity-area Alliance-for-Zero-Extinction EDGE-species evolutionarily-distinct globally-endangered conservation-prioritization systematic-conservation-planning Marxan Zonation conservation-opportunity conservation-cost conservation-benefit trade-off socioeconomic-factor stakeholder-engagement community-based-conservation adaptive-management monitoring-program long-term-study citizen-science public-participation environmental-education outreach science-communication knowledge-transfer capacity-building institutional-strengthening policy-development legislation regulation enforcement compliance incentive payment-for-ecosystem-services carbon-credit biodiversity-offset mitigation-banking conservation-easement land-trust private-land-conservation conservation-covenant stewardship sustainable-use sustainable-management ecosystem-based-management integrated-water-resources-management basin-management catchment-management landscape-management seascape-management connectivity-conservation corridor stepping-stone buffer-zone matrix-management countryside-biogeography reconciliation-ecology novel-ecosystem designer-ecosystem ecosystem-service provisioning-service regulating-service supporting-service cultural-service food-production water-supply raw-materials genetic-resources medicinal-resources ornamental-resources air-quality-regulation climate-regulation water-regulation erosion-regulation water-purification waste-treatment disease-regulation pest-regulation pollination seed-dispersal habitat-provision nutrient-cycling soil-formation primary-production recreation tourism aesthetic-value spiritual-value educational-value scientific-value inspiration sense-of-place cultural-heritage traditional-knowledge indigenous-knowledge local-knowledge ecosystem-service-valuation economic-valuation contingent-valuation choice-experiment benefit-transfer total-economic-value use-value non-use-value direct-use indirect-use option-value bequest-value existence-value willingness-to-pay willingness-to-accept cost-benefit-analysis cost-effectiveness-analysis multi-criteria-analysis decision-support uncertainty risk-assessment scenario-analysis sensitivity-analysis robustness-analysis adaptive-capacity resilience vulnerability exposure sensitivity coping-capacity transformation social-ecological-system complex-adaptive-system tipping-point regime-shift alternative-stable-state hysteresis path-dependence legacy-effect slow-variable fast-variable panarchy adaptive-cycle exploitation release reorganization foresight anticipatory-governance transformative-change sustainability-science future-earth interdisciplinary transdisciplinary problem-oriented solution-oriented action-oriented knowledge-co-production boundary-work boundary-object boundary-organization science-policy-interface science-society-interface knowledge-brokerage knowledge-exchange knowledge-integration mode-1-science mode-2-science triple-helix quadruple-helix responsible-innovation reflexive-governance adaptive-governance collaborative-governance polycentric-governance network-governance self-organization collective-action common-pool-resource tragedy-of-the-commons social-dilemma institutional-analysis institutional-design Ostrom-design-principles monitoring sanctioning conflict-resolution nested-enterprises proportionality collective-choice constitutional-choice operational-choice collective-action-theory game-theory prisoner's-dilemma chicken-game assurance-game coordination-game evolutionary-game-theory replicator-dynamics ESS Nash-equilibrium Pareto-optimality social-optimum market-failure government-failure voluntary-failure non-market-value public-good common-good club-good private-good rivalry excludability free-rider hold-up-problem moral-hazard adverse-selection information-asymmetry transaction-cost property-right common-property state-property private-property hybrid-property bundle-of-rights usufruct access withdrawal alienation tenure-security land-tenure water-tenure resource-tenure customary-tenure statutory-tenure legal-pluralism de-facto de-jure formal informal legality legitimacy recognition deviance corruption elite-capture social-capital bonding-capital bridging-capital linking-capital trust reciprocity reputation social-network social-embeddedness structural-hole brokerage closure cohesion density centrality betweenness degree eigenvector PageRank small-world scale-free random-network regular-network hierarchical-network modularity community-structure clustering assortativity homophily heterophily preferential-attachment fitness-model network-evolution network-dynamics temporal-network multilayer-network multiplex-network bipartite-network affiliation-network one-mode-network two-mode-network 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Unionicola
Unionicola is a genus of freshwater water mites (family Unionicolidae) described by Haldeman in 1842. The genus has cosmopolitan distribution and comprises parasitic species primarily associated with molluscan hosts, though some species associate with freshwater sponges. Most species infest unionid mussels, while U. crassipes and U. minor are associated with Spongillidae sponges. The mite-host relationship involves consumption of host mucus and tissue, with potential negative effects on host fitness.
water-mite parasite freshwater mollusk-parasite sponge-associate cosmopolitan sexual-dimorphism
Varroa
Varroa Mites
Varroa is a genus of parasitic mesostigmatan mites that are obligate ectoparasites of honey bees (Apis spp.). The genus contains four species, with Varroa destructor being the most significant pest of Western honey bees (Apis mellifera) worldwide. Originally parasites of Asian honey bees (Apis cerana) in mainland Asia, these mites have spread globally through beekeeping activities. V. destructor is recognized as the most devastating pest of managed honey bees, causing colony losses through direct parasitism and by vectoring viruses such as deformed wing virus. The genus was named for Marcus Terentius Varro, a Roman scholar and beekeeper.
parasite honey-bee-pest disease-vector acarology apiculture invasive-species colony-collapse-disorder integrated-pest-management varroacide-resistance Varroa-sensitive-hygiene social-apoptosis fat-body-feeding deformed-wing-virus Apis-mellifera Apis-cerana
Varroa destructor
Varroa mite
Varroa destructor is an external parasitic mite and the most damaging pest of managed honey bee colonies worldwide. The species originally parasitized the Asian honey bee (Apis cerana) but host-shifted to the western honey bee (Apis mellifera), causing catastrophic colony losses. Adult female mites feed on the fat body tissue of adult bees and larvae, while reproducing exclusively within sealed brood cells. The mite vectors at least five debilitating bee viruses, including deformed wing virus, and colonies without management typically collapse within 2–3 years.
parasite honey-bee-pest vector invasive-species agricultural-pest Apis-mellifera
Wohlfahrtia
Wohlfahrtia is a genus of flesh flies in the family Sarcophagidae comprising at least 20 described species. Species within this genus are known facultative parasites that cause traumatic myiasis in mammals, including livestock and humans. Larvae develop in wounds or necrotic tissue, with some species serving as forensic indicators for postmortem interval estimation. The genus exhibits larviparous reproduction, with females depositing larvae directly onto suitable substrates.
Sarcophagidae flesh-fly myiasis forensic-entomology larviparous parasite livestock-pest traumatic-myiasis
Wohlfahrtia vigil
fox maggot, myiasis fly
Wohlfahrtia vigil is a flesh fly (Sarcophagidae) known for causing myiasis in mammals. The species has been studied for its laboratory rearing methods and behavioral characteristics. Adults are active during warmer months and females deposit larvae on or near hosts rather than laying eggs.
myiasis flesh-fly parasite larviparous Sarcophagidae veterinary-entomology medical-entomology
Xenos
twisted-wing parasites
Xenos is a genus of twisted-wing parasites in the order Strepsiptera. These insects are obligate endoparasites of other insects, with larvae entering hosts through joints or sutures. Adult males are free-flying and short-lived, while neotenic females remain permanently within host abdomens. The genus includes species such as Xenos peckii, which parasitizes wasps including Polistes fuscatus.
Strepsiptera parasite endoparasite sexual-dimorphism neoteny hymenopteran-parasite
Xenox delila
Xenox delila is a species of bee fly in the family Bombyliidae, found in California and Baja California Norte. As with other bee flies, adults are likely pollinators that visit flowers for nectar. The genus Xenox includes parasitic species whose larvae develop in the nests of other insects.
bee-fly parasite pollinator endemic California Baja-California Bombyliidae Diptera
Zodion obliquefasciatum
thick-headed fly
Zodion obliquefasciatum is a species of thick-headed fly in the family Conopidae. Members of the genus Zodion are small conopid flies, typically 5–7 millimeters in length, that parasitize solitary wasps. Like other conopids, they are wasp mimics and frequent flowers for nectar while hunting for hosts.
parasite wasp-mimic flower-visitor internal-parasite