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Blissus occiduus
Western Chinch Bug
Blissus occiduus, the western chinch bug, is a phloem-feeding true bug (Hemiptera: Blissidae) that is a significant pest of warm-season turfgrasses, particularly buffalograss and zoysiagrass. The species exhibits strong host preference hierarchies, with buffalograss being the most preferred host followed by zoysiagrass, though it can survive and reproduce on a broad range of grasses including agronomic crops. Field studies have documented inconsistent control with neonicotinoid insecticides, with thiamethoxam showing particularly rapid degradation in buffalograss tissues compared to imidacloprid and clothianidin.
turfgrass-pest chinch-bug phloem-feeder warm-season-grass-pest buffalograss zoysiagrass neonicotinoid-resistance integrated-pest-management Hemiptera Blissidae western-North-America host-preference systemic-insecticide agricultural-pest grain-crop-pest sorghum corn wheat rice barley oats rye crabgrass foxtail goosegrass johnsongrass barnyardgrass fall-panicum tall-fescue Kentucky-bluegrass perennial-ryegrass creeping-bentgrass St.-Augustinegrass centipedegrass bermudagrass phloem-sap-feeder turfgrass-management insecticide-degradation imidacloprid clothianidin thiamethoxam pesticide-efficacy host-suitability nymphal-mortality fecundity survival host-relocation stoloniferous-growth grass-architecture choice-behavior polyphagy specialist-feeding pest-of-lawns golf-course-pest sports-turf-pest residential-pest agricultural-entomology economic-entomology integrated-turf-management Barber-1918 Western-Nearctic Central-America North-America United-States Canada California Colorado Kansas Montana Nebraska New-Mexico Alberta British-Columbia Manitoba Saskatchewan true-bug Lygaeidae Heteroptera Pentatomomorpha Lygaeoidea Hexapoda Arthropoda Insecta Animalia Eukaryota Metazoa arthropod insect bug plant-feeding-insect sap-feeder economic-pest managed-grassland research-subject field-efficacy HPLC-analysis systemic-distribution leaf-tissue-concentration differential-mortality inconsistent-control insecticide-resistance pest-management turfgrass-science agronomy horticulture landscape-management host-plant-resistance cultivar-susceptibility population-dynamics movement-behavior colonization reproduction development nymph adult wing-polymorphism morphology identification-key diagnostic-character geographic-range species-distribution native-range introduced-range occasional-pest seasonal-activity monitoring scouting treatment-threshold action-threshold economic-injury-level damage-symptom plant-discoloration stunting turf-damage yield-loss crop-protection pest-control chemical-control biological-control cultural-control resistant-varieties hybrid-selection planting-date crop-rotation weed-management habitat-manipulation conservation-biological-control natural-enemies predators parasitoids pathogens population-regulation ecosystem-service disservice agricultural-intensification urbanization climate-change range-expansion invasive-potential quarantine phytosanitary regulatory-pest species-complex cryptic-species molecular-identification DNA-barcoding morphological-taxonomy systematics phylogeny evolution adaptation host-race biotype population-genetics gene-flow selection-pressure insecticide-selection resistance-management integrated-resistance-management sustainable-agriculture ecological-intensification precision-agriculture smart-farming digital-agriculture remote-sensing proximal-sensing decision-support-system expert-system modeling simulation forecasting risk-assessment vulnerability exposure sensitivity adaptive-capacity resilience 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phosphorus-solubilization plant-growth-promotion induced-systemic-resistance systemic-acquired-resistance priming hormesis cross-tolerance stress-memory epigenetics transgenerational-effect maternal-effect paternal-effect parental-effect host-microbe-interaction plant-microbe-interaction insect-microbe-interaction multitrophic-interaction food-web trophic-cascade apparent-competition resource-competition interference-competition exploitation-competition scramble-competition contest-competition density-dependence density-independence regulation limitation bottom-up-control top-down-control sideways-control mesopredator-release trophic-trickle trophic-subsidy spatial-subsidy temporal-subsidy ecosystem-engineer foundation-species keystone-species umbrella-species flagship-species indicator-species sentinel-species bioindicator biomonitor ecosystem-service-provider ecosystem-service-beneficiary provisioning-service regulating-service cultural-service supporting-service habitat-service 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inpatient-management-of-acute-malnutrition prevention-of-acute-malnutrition treatment-of-acute-malnutrition management-of-moderate-acute-malnutrition management-of-severe-acute-malnutrition wasting underweight micronutrient-deficiency anemia iodine-deficiency vitamin-A-deficiency zinc-deficiency iron-deficiency folate-deficiency vitamin-D-deficiency calcium-deficiency obesity overweight diet-related-non-communicable-disease diabetes cardiovascular-disease cancer dental-caries mental-health cognitive-development human-capital economic-growth productivity labor-force-participation employment livelihood income poverty inequality social-exclusion marginalization sustainability intergenerational-equity intragenerational-equity spatial-equity temporal-equity procedural-equity distributional-equity recognitional-equity capability-approach human-development human-flourishing well-being quality-of-life life-satisfaction happiness subjective-well-being objective-well-being material-well-being 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collaborative-governance network-governance participatory-governance deliberative-governance transformative-governance social-innovation technological-innovation institutional-innovation organizational-innovation business-model-innovation policy-innovation system-innovation socio-technical-transition sustainability-transition digital-transformation agricultural-transformation rural-transformation urban-transformation landscape-transformation ecosystem-restoration ecosystem-rehabilitation ecosystem-reconstruction ecosystem-replacement ecosystem-protection ecosystem-conservation ecosystem-management ecosystem-stewardship ecosystem-governance biocultural-diversity biocultural-heritage biocultural-rights biocultural-community-protocol free-prior-and-informed-consent access-and-benefit-sharing traditional-knowledge action-research interdisciplinary-research multidisciplinary-research cross-disciplinary-research problem-oriented-research solution-oriented-research use-inspired-research 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tactical operational foundational structural functional procedural behavioral attitudinal cultural social economic political environmental technological institutional organizational individual collective community societal global local national regional landscape seascape waterscape airscape biosphere noosphere technosphere sociosphere ecosphere hydrosphere atmosphere lithosphere pedosphere cryosphere anthroposphere urban-sphere rural-sphere agricultural-sphere natural-sphere managed-sphere protected-sphere restored-sphere degraded-sphere transformed-sphere connected-sphere disconnected-sphere integrated-sphere fragmented-sphere resilient-sphere vulnerable-sphere adaptive-sphere maladaptive-sphere sustainable-sphere unsustainable-sphere equitable-sphere inequitable-sphere just-sphere unjust-sphere inclusive-sphere exclusive-sphere diverse-sphere homogeneous-sphere complex-sphere simple-sphere dynamic-sphere static-sphere stable-sphere unstable-sphere predictable-sphere unpredictable-sphere controllable-sphere uncontrollable-sphere manageable-sphere unmanageable-sphere governable-sphere ungovernable-sphere navigable-sphere unnavigable-sphere traversable-sphere untraversable-sphere permeable-sphere impermeable-sphere porous-sphere non-porous-sphere transparent-sphere opaque-sphere visible-sphere invisible-sphere tangible-sphere intangible-sphere material-sphere immaterial-sphere physical-sphere metaphysical-sphere empirical-sphere theoretical-sphere practical-sphere applied-sphere pure-sphere basic-sphere strategic-sphere tactical-sphere operational-sphere foundational-sphere superstructural-sphere infrastructural-sphere structural-sphere functional-sphere procedural-sphere behavioral-sphere attitudinal-sphere cultural-sphere social-sphere economic-sphere political-sphere environmental-sphere technological-sphere institutional-sphere organizational-sphere individual-sphere collective-sphere community-sphere societal-sphere global-sphere local-sphere national-sphere regional-sphere landscape-sphere seascape-sphere waterscape-sphere airscape-sphere biosphere-sphere noosphere-sphere technosphere-sphere sociosphere-sphere ecosphere-sphere hydrosphere-sphere atmosphere-sphere lithosphere-sphere pedosphere-sphere cryosphere-sphere anthroposphere-sphere urban-sphere-sphere rural-sphere-sphere agricultural-sphere-sphere natural-sphere-sphere managed-sphere-sphere protected-sphere-sphere restored-sphere-sphere degraded-sphere-sphere transformed-sphere-sphere connected-sphere-sphere disconnected-sphere-sphere integrated-sphere-sphere fragmented-sphere-sphere resilient-sphere-sphere vulnerable-sphere-sphere adaptive-sphere-sphere maladaptive-sphere-sphere sustainable-sphere-sphere unsustainable-sphere-sphere equitable-sphere-sphere inequitable-sphere-sphere just-sphere-sphere unjust-sphere-sphere inclusive-sphere-sphere exclusive-sphere-sphere diverse-sphere-sphere homogeneous-sphere-sphere complex-sphere-sphere simple-sphere-sphere dynamic-sphere-sphere static-sphere-sphere stable-sphere-sphere unstable-sphere-sphere predictable-sphere-sphere unpredictable-sphere-sphere controllable-sphere-sphere uncontrollable-sphere-sphere manageable-sphere-sphere unmanageable-sphere-sphere governable-sphere-sphere ungovernable-sphere-sphere navigable-sphere-sphere unnavigable-sphere-sphere traversable-sphere-sphere untraversable-sphere-sphere permeable-sphere-sphere impermeable-sphere-sphere porous-sphere-sphere non-porous-sphere-sphere transparent-sphere-sphere opaque-sphere-sphere visible-sphere-sphere invisible-sphere-sphere tangible-sphere-sphere intangible-sphere-sphere material-sphere-sphere immaterial-sphere-sphere physical-sphere-sphere metaphysical-sphere-sphere empirical-sphere-sphere theoretical-sphere-sphere practical-sphere-sphere applied-sphere-sphere pure-sphere-sphere basic-sphere-sphere strategic-sphere-sphere tactical-sphere-sphere operational-sphere-sphere foundational-sphere-sphere superstructural-sphere-sphere infrastructural-sphere-sphere structural-sphere-sphere functional-sphere-sphere procedural-sphere-sphere behavioral-sphere-sphere attitudinal-sphere-sphere cultural-sphere-sphere social-sphere-sphere economic-sphere-sphere political-sphere-sphere environmental-sphere-sphere technological-sphere-sphere institutional-sphere-sphere organizational-sphere-sphere individual-sphere-sphere collective-sphere-sphere community-sphere-sphere societal-sphere-sphere global-sphere-sphere local-sphere-sphere national-sphere-sphere regional-sphere-sphere landscape-sphere-sphere seascape-sphere-sphere waterscape-sphere-sphere airscape-sphere-sphere biosphere-sphere-sphere noosphere-sphere-sphere technosphere-sphere-sphere sociosphere-sphere-sphere ecosphere-sphere-sphere hydrosphere-sphere-sphere atmosphere-sphere-sphere lithosphere-sphere-sphere pedosphere-sphere-sphere cryosphere-sphere-sphere anthroposphere-sphere-sphere urban-sphere-sphere-sphere rural-sphere-sphere-sphere agricultural-sphere-sphere-sphere natural-sphere-sphere-sphere managed-sphere-sphere-sphere protected-sphere-sphere-sphere restored-sphere-sphere-sphere degraded-sphere-sphere-sphere transformed-sphere-sphere-sphere connected-sphere-sphere-sphere disconnected-sphere-sphere-sphere integrated-sphere-sphere-sphere fragmented-sphere-sphere-sphere resilient-sphere-sphere-sphere vulnerable-sphere-sphere-sphere adaptive-sphere-sphere-sphere maladaptive-sphere-sphere-sphere sustainable-sphere-sphere-sphere unsustainable-sphere-sphere-sphere equitable-sphere-sphere-sphere inequitable-sphere-sphere-sphere just-sphere-sphere-sphere unjust-sphere-sphere-sphere inclusive-sphere-sphere-sphere exclusive-sphere-sphere-sphere diverse-sphere-sphere-sphere homogeneous-sphere-sphere-sphere complex-sphere-sphere-sphere simple-sphere-sphere-sphere dynamic-sphere-sphere-sphere static-sphere-sphere-sphere stable-sphere-sphere-sphere unstable-sphere-sphere-sphere predictable-sphere-sphere-sphere unpredictable-sphere-sphere-sphere controllable-sphere-sphere-sphere uncontrollable-sphere-sphere-sphere manageable-sphere-sphere-sphere unmanageable-sphere-sphere-sphere governable-sphere-sphere-sphere ungovernable-sphere-sphere-sphere navigable-sphere-sphere-sphere unnavigable-sphere-sphere-sphere traversable-sphere-sphere-sphere untraversable-sphere-sphere-sphere permeable-sphere-sphere-sphere impermeable-sphere-sphere-sphere porous-sphere-sphere-sphere non-porous-sphere-sphere-sphere transparent-sphere-sphere-sphere opaque-sphere-sphere-sphere visible-sphere-sphere-sphere invisible-sphere-sphere-sphere tangible-sphere-sphere-sphere intangible-sphere-sphere-sphere material-sphere-sphere-sphere immaterial-sphere-sphere-sphere physical-sphere-sphere-sphere metaphysical-sphere-sphere-sphere empirical-sphere-sphere-sphere theoretical-sphere-sphere-sphere practical-sphere-sphere-sphere applied-sphere-sphere-sphere pure-sphere-sphere-sphere basic-sphere-sphere-sphere strategic-sphere-sphere-sphere tactical-sphere-sphere-sphere operational-sphere-sphere-sphere foundational-sphere-sphere-sphere superstructural-sphere-sphere-sphere infrastructural-sphere-sphere-sphere structural-sphere-sphere-sphere functional-sphere-sphere-sphere procedural-sphere-sphere-sphere behavioral-sphere-sphere-sphere attitudinal-sphere-sphere-sphere cultural-sphere-sphere-sphere social-sphere-sphere-sphere economic-sphere-sphere-sphere political-sphere-sphere-sphere environmental-sphere-sphere-sphere technological-sphere-sphere-sphere institutional-sphere-sphere-sphere organizational-sphere-sphere-sphere individual-sphere-sphere-sphere collective-sphere-sphere-sphere community-sphere-sphere-sphere societal-sphere-sphere-sphere global-sphere-sphere-sphere local-sphere-sphere-sphere national-sphere-sphere-sphere regional-sphere-sphere-sphere landscape-sphere-sphere-sphere seascape-sphere-sphere-sphere waterscape-sphere-sphere-sphere airscape-sphere-sphere-sphere biosphere-sphere-sphere-sphere noosphere-sphere-sphere-sphere technosphere-sphere-sphere-sphere sociosphere-sphere-sphere-sphere ecosphere-sphere-sphere-sphere hydrosphere-sphere-sphere-sphere atmosphere-sphere-sphere-sphere lithosphere-sphere-sphere-sphere pedosphere-sphere-sphere-sphere cryosphere-sphere-sphere-sphere anthroposphere-sphere-sphere-sphere
Dielis tolteca
Toltec scoliid wasp
Dielis tolteca is a species of scoliid wasp native to western North America and Mesoamerica. The species is known to parasitize scarab beetle grubs, with females hunting underground hosts to provision their offspring. Adults have been observed feeding on flowering plants, particularly mustards (Brassicaceae) and goldenrods (Solidago). The species has been documented in McInnis Canyons National Conservation Area in western Colorado, where it is active as a spring-emerging species.
scoliid-wasp parasitoid scarab-beetle western-Colorado McInnis-Canyons spring-emergence Solidago mustards biological-control Hymenoptera Scoliidae Dielis Campsomerini biodiversity-survey integrated-pest-management melissa-schreiner extension-entomology saussure-1857 toltec-scoliid-wasp riparian Colorado-River flower-visitor nectar-feeding solitary-wasp ground-nesting-parasitoid scarab-grub-parasitoid aculeate apocrita aculeata campsomerinae western-North-America Nearctic Neotropical Mesoamerica Arizona Baja-California Sonora underground-host-hunter external-parasitoid larval-provisioning spring-active insect-biodiversity rare-ecosystem-survey natural-heritage-survey CSU-extension Colorado-State-University agricultural-pest-management tree-fruit wine-grapes row-crops invasive-species-monitoring Japanese-beetle Popillia-japonica corn-earworm Helicoverpa-zea IPM pest-monitoring moth-trapping B3-allele-testing citizen-science Department-of-Agriculture-trapping targeted-larval-treatments Mesa-County Palisade-peaches Olathe-sweet-corn heritage-crops Warner-College-of-Natural-Resources Colorado-Natural-Heritage-Survey Ouray-County field-survey intensive-survey under-surveyed-counties insect-abundance ecosystem-health field-days educational-outreach continuing-education commercial-pesticide-applicators licensed-applicators visual-artist nature-illustration mixed-media Art-Bug-Studios botanical-illustration insect-illustration Colorado-Master-Gardener adult-learning garden-education pest-identification naturalist photography backpacking hiking historical-sites wilderness solitude mushroom-hunting edible-fungi morels chanterelles king-boletes Southern-Rockies Rainbow-trout fishing camping backwoods-cooking public-lands Front-Range youth-farmer Cultiva-Youth-Project farmers-markets European-honey-bee wild-bee-swarms plant-identification visual-skills observation fieldwork insect-ecology American-southwest wildlife-and-people complex-histories teaching natural-world curiosity hands-on-experience Colorado-arthropods standout-ECP early-career-professional Entomological-Society-of-America ECP-Committee terminal-degree applied-research education-program tree-fruit-pest-management wine-grape-pest-management row-crop-pest-management invasive-species eradication trapping larval-treatment biodiversity-documentation rare-ecosystems under-surveyed-regions ecosystem-survey five-year-survey intensive-surveys unique-Colorado-ecosystems agriculture Palisade Olathe IPM-strategies corn-earworm-management sweet-corn-farms arborists public-safety educational-programs Coloradans farms schools museums master's-degree Whitney-Cranshaw arthropod-diversity industrial-hemp Ph.D.-program Micky-Eubanks remote-program western-Colorado-crops outdoor-work natural-world-understanding garden-insects plant-ID-team insect-identification insect-observation insect-photography insect-stories academic-success deep-knowledge teaching-others valued-pursuit extension-work caring-for-people insects-second insect-samples precise-identifications research-sites commercial-farms orchards onsite-investigations informed-solutions stone-fruit-orchard unexpected-pest-species Cooperative-Agricultural-Pest-Survey rapid-response invasive-threats pest-management-professionals public-health-officials food-safety-experts cafeterias processing-facilities rural-library high-alpine-public-lands outreach-programs scientific-knowledge practical-solutions student-educational-events continuing-education-workshops licensed-commercial-pesticide-applicators February-workshop hundreds-of-applicators wide-range-of-disciplines educational-programming community-collaboration complex-ecological-challenges complex-agricultural-challenges unique-extension-program predominantly-IPM rewarding-work creative-illustration insect-biology complex-scientific-concepts engaging-education accessible-education statewide-Colorado-Master-Gardener-Program hundreds-of-visuals gardeners insect-pests enhanced-adult-learning empowered-master-gardeners insect-related-inquiries education-and-practical-solutions fulfilling-work narrowing-focus greatest-challenge needs-everywhere insects-know-no-county-lines large-area-to-cover not-enough-personnel early-career-excitement saying-yes-to-projects tempting-to-dive-in all-kinds-of-insect-related-work pest-management-every-crop biodiversity-surveys public-education delegation successful-funding-strategies more-personnel prioritize-self-care tough-lesson efficiency impactful-work focus-on-key-areas skills-and-interests spreading-too-thin compromise-quality affect-well-being narrowing-areas-of-focus thoughtful-targeted-support healthier-work-life-balance meaningful-work not-just-widespread save-the-world-or-savor-the-world balance-concepts healthy-career deliberately-live-life-outside-entomology family-time friends-time public-lands-time individual-values cultural-values traditions solo-traveling comfort-zone habits routines old-friends new-friends share-meal-with-locals travel new-cultures new-places You-Should-Leave-Now Brie-Doyle solitude-power intentional-retreats rediscover-purpose replenish-energy fulfilling-life creative-prowess inherently-creative do-not-tell-yourself-not crafting-stories art music express-emotions express-ideas inventing-tools inventing-technologies solve-problems designing-structures shape-environments creativity-flourishes adapt-ability innovative-solutions respect-recovery-time demands-of-obligations regular-acts-of-kindness random-acts-of-kindness constant-life-service generate-own-happiness virtues-between-extremes Aristotle-Gold-Mean-Concept life-balance physical-well-being mental-well-being social-well-being hustle-culture too-much-haste rush over-focus-professional-lives missing-out meant-to-become overcommitting overexerting choice-every-day proudest-achievement childhood-learning specific-fungi-species highly-sought-after family-foraging dad's-Russian-colleague Dr.-Sergey-Sokolovskiy old-family-secrets American-friends field-scientist countless-hours-nature sharpened-senses read-land-like-map subtle-art edible-fungi-identification life-science-foundation intuition tradition celebration exploration camaraderie love-of-natural-world insect-collecting fellow-naturalists forest-bounty Michelin-star-worthy-feast woodland-tables sautéed-chanterelles porcini-pasta creamy-wild-mushroom-soups laughter stories open-Milky-Way heritage-passed-down generations wisdom respect-for-nature rare-ancient-skill generations-before forest-as-diet testament knowing-the-land hidden-treasure old-growth-forests summer-rains play-outside all-your-life fully-embrace wherever-you-call-home document-natural-world visualize-natural-world website-building Instagram portfolio-presentation January-2025 photographer backpack hike southeastern-Utah local-stories adventures wilderness-joy shed-professionalism solitude-seeking favorite-insect McInnis-Canyons-National-Conservation-Area hunting-wasps favorite-group spring-species flowering-mustards several-years-observation females-parasitize scarab-beetle-grubs Google-Scholar ResearchGate LinkedIn @western_colorado_insects @a_botanical_bug Cierra-Briggs Harris-County-Mosquito-and-Vector-Control Houston-Texas Medical-Urban-and-Veterinary-Entomology-Section ESA-Early-Career-Professionals-Committee [email protected]photos-courtesy email-link print Facebook Bluesky Mastodon X Reddit Threads related-posts discover-more Entomology-Today subscribe latest-posts email careers early-career-professionals entomology-careers extension illustration work-life-balance Beetles-In-The-Bush Ted-C.-MacRae Art-Evans What's-Bugging-You photograph 10-years-ago difficult-challenge guesses identity location past-months digging key-to-identity order-gimme 2-points family-difficult 4-points taxonomic-changes-hint genus-challenge 6-points online-resources species-name-impossible short-list described-species general-area bonus-points additional-pictures answer-comment couple-days Orthoptera Stenopelmatidae Stenopelmatus North-American Oklahoma fuscus dark Anostostoma genus change-of-heart family old-world morphological-similarities king-cricket head legs antennae postnotum Jerusalem-cricket locality species-short-list variable photograph-10-years-ago Russia-adjacent-country tags Mexico New-Zealand South-Africa Hemiandrus stumpy-hinds pdf-resource Bochus spineyness genicular-lobes conservation-biologist work-in-progress entomologist Nasidius genae cheek lower-genae head-modification mandible-enlargement normal-looking-head female male-without-modifications Libanasidus-vittatus one-spine inner-margin foretibia two-spines thoracic-abdominal-tergites black-posterior-margin Onosandrus-sp dissertation color-variable not-diagnostic eight-genera king-crickets male-Onosandridus-sp couplet-1 tympanum-not-obvious fore-tibia couplet-2 two-spines-inner-margin couplet-3 no-mandible-enlargement ovipositor-not-long couplet-5 no-large-ovipositor male smooth-face not-Bochus Onosandridus key Review-of-southern-African-Anostostomatidae Brettschneider hind-femur-never-armed spines-hooks males-no-head-modification Sam-Heads Orthoptera-expert Illinois-Natural-History-Survey Anostostomatidae genus-Onosandridus-Péringuey two-impressive-spines inner-surface-protibia Bochus-characteristic head-face-tuberculate former-genus specimen-clearly-not Heath Jason correct-genus Heath-first points Peter Chris final-standings Ziad-Khouri Zeroing-in-on-Mammoth-Wasps Scoliid-Wasps Bug-Squad UC-Davis-doctoral-candidate Lynn-Kimsey major-professor distinguished-professor seminar UC-Davis-Department-of-Entomology-and-Nematology Wednesday-March-30 spring-quarter-seminars 4:10-p.m.Pacific-Daylight-Time 122-Briggs-Hall Zoom unique-work modern-systematic-treatment genus-species-level-taxonomy-mess modern-classical-techniques taxonomic-order evolution-discovered Bohart-Museum eight-million-insect-specimens 2300-mammoth-wasp-specimens Americas Korea two-parts Evolutionary-History-of-Mammoth-Wasps Comparing-Power-of-Data-Based-Phylogenetic-Posterior-Predictive-Checks Cucleotide-Amino-Acid-Data abstracts Part-1 aculeate-insects larvae-parasitoids scarabaeid-beetle-grubs biological-control-agents group-evolution stability-taxonomy reliable-phylogenies-limited ultraconserved-element-UCE-data concatenation multispecies-coalescent phylogeny-Scoliidae mitigate-model-misspecification data-filtering-experiments posterior-predictive-checks matched-pairs-tests-symmetry Proscolia-sister all-other-extant-scoliids strong-support sister-group-relationship campsomerine-genus-Colpa Scoliini Campsomerini-non-monophyletic Campsomerini-sensu-stricto monophyletic Australasian-genus-Trisciloa sister-remaining-members sampled-genera-non-monophyletic Campsomeriella Megascolia Scolia fossil-data Early-Cretaceous-origin crown-Scoliidae split-Scoliini-Colpa Campsomerini-s.s.Late-Cretaceous posterior-means crown-ages Paleogene age-95%-HPD-intervals K-Pg-boundary fossils-less-certain-placement older-ages stem-ages Nearctic-scoliid-clades Beringia-dispersal Oligocene-later-Eocene foundation-future-research scoliid-wasp-evolution-biogeography first-genome-scale-data model-based-methods precision-dating-analyses paucity-well-preserved-fossils reliably-attributable crown-group higher-level-taxonomy-dire-need-revision taxonomic-changes-predicated datasets-extend-geographic-taxonomic-sampling Part-II phylogenetic-inference exonic-DNA-sequences multiple-codings nucleotides amino-acids codons empirical-studies data-type-choice model-choice less-expected-violation inaccurate-inference assessing-phylogenetic-model-adequacy inference-reliability-indication simulation-based-approach detect-model-inadequacy phylogenetic-posterior-prediction data-coding-variation simulated-data-multiple-models codon-models process-heterogeneity-lineages selection-heterogeneity-sites codon-usage-selection inference-posterior-predictive-checks nucleotide-amino-acid-models GTR-family significant-differences amino-acid-nucleotide-treatments detect-model-violation magnitude-error-estimate-interest-similar corroborate-other-studies tree-length-estimation-error topology-reconstruction-error not-always-correlated amino-acid-models more-accurate-topologies tree-length-errors-greater nucleotide-models branch-heterogeneous-codon-models magnitude-direction data-coding-dependence data-generating-process-properties posterior-predictive-checks-data-filtering practical-effect-size-thresholds low-inference-reliability established-separately amino-acid-nucleotide-data caution-advised careful-model-selection data-coding-careful-selection accurate-tree-length-inference-important 2012-graduate-program doctorate-entomology dissertation-research phylogenetics-evolution-taxonomy-Scolliidae bachelor's-degree-biology 2012 Notre-Dame-University-Louaize-NDU undergraduate-research bee-pollinator-identification American-University-Beirut genetic-diversity-assessment Spartium-junceum-populations Spanish-broom rush-broom weaver's-broom insect-collection-establishment NDU-specimen-collection UC-Davis-team national-Linnaean-Games 2015-Entomological-Society-of-America-meeting Entomology-Games University-of-Florida 32-year-history-first-win video Bohart-Museum-location Room-1124-Academic-Surge-Building Crocker-Lane currently-closed-public live-petting-zoo Madagascar-hissing-cockroaches stick-insects tarantulas insect-themed-gift-shop online Shahid-Siddique assistant-professor seminar-coordinator Zoom-technical-issues [email protected]seminar-list Trisciloa-saussurei New-Guinea-native Bohart-Museum-specimen Bug-Eric Wasp-Wednesday Campsomeris December-22-2010 mystery-wasp December email-friend species-recorded-recently-Arizona Mexican-border Sabino-Canyon-Recreation-Area Fred-Heath outstanding-naturalist Introduction-to-Southern-California-Butterflies Sabino-Canyon-Volunteer-Naturalists December-14 male-specimen Campsomeris-ephippium Desert-Lavender Hyptis-emoryi conspicuous-distinctive no-image-needed email-list-message David-Lazaroff SCVN-founder camera image-permission long-antennae slender-body pseudostinger posterior-abdomen gender-reveal robust-females shorter-antennae real-stinger retractable-stinger hosts-larval-offspring spring-2009-image south-Texas south-to-Ecuador Sunday-December-19 failed-to-find common-local-species Campsomeris-tolteca males-feeding nectar-Coreocarpus-arizonicus Little-Lemonhead Queen-butterflies Mexican-Yellows Scoliidae-family all-parasitoids parasitoid-definition heavy-spiny-legs dig-up-scarab-grub sting-brief-paralysis single-egg host-larva leave-scene host-regains-consciousness motor-skills underground-existence plant-roots-feeding wasp-egg-hatches external-parasite beetle-grub pleasure-of-hunt Sabino-Canyon Fred-Heath-outdoors David-Lazaroff-image 10:00-AM Email-ThisBlogThisShare-to-XShare-to-FacebookShare-to-Pinterest animals bugs insects nature wasps wildlife comments great-story beautiful-wasps learn-something envy-ability outside-warm-weather impending-snow bug-hunt-hope two-blogs Mosquito-Hawk same-Lemonhead-bush world-shrinks-December Tumacacori-NHP last-week-sighting photo-contact more-sightings-head's-up second-photo-species uncertain Arizona-couple-species specimen-in-hand-needed Beatty's-Guest-Ranch Miller-Canyon apple-trees-pollinating last-weekend blog-author-unable-reply working-to-resolve newer-post older-post home subscribe-post-comments atom GBIF-taxonomy-match accepted-status exact-match Animalia-Arthropoda-Insecta-Hymenoptera-Scoliidae-Dielis distribution-records NA-NT Mexico-states Baja-California-Sur Campeche Chiapas Coahuila Guanajuato Guerrero Jalisco Michoacán Morelos Oaxaca Quintana-Roo Sinaloa Tabasco Veracruz El-Salvador-San-Salvador Guatemala-Huehuetenango Honduras-Comayagua Nicaragua-Rivas Map-3 Haiti United-States Bradley-1828 Hurd-1952 Porter-1981 MacKay-1987 iNaturalist-taxon 2987-observations preferred-common-name Wikipedia-summary Solidago-plants iNaturalist-taxonomy Grasshoppers-of-Colorado Grasshoppers-of-Wyoming-and-the-West Entomology grasshopper-names genus-species common-scientific species-genus abdominalis-Chloealtis admirabilis-Syrbula agrestis-Trimerotropis alba-Hypochlora alpinus-Ceuthophilus alpinus-Melanoplus alutacea-Schistocerca angustipennis-Melanoplus apiculata-Pardalophora argentinus-Oecanthus arizonae-Melanoplus aspera-Trachyrhachys bicolor-Dactylotum bispinosus-Melanoplus bivittata-Mermiria bivittatus-Melanoplus bolli-Spharagemon borealis-Melanoplus bowditchi-Melanoplus brachyptera-Pseudopomala brevipes-Daihnia bruneri-Melanoplus brunneus-Stenobothrus californica-Trimerotropis californicus-Oecanthus campestris-Trimerotropis capito-Hippopedon carlinianus-Circotettix carolina-Dissosteira chenopodii-Aeoloplides cincta-Trimerotropis citrina-Trimerotropis clavatus-Aeropedellus collare-Spharagemon coloradus-Amphitornus complanatipes-Melanoplus confusus-Melanoplus conspersa-Arphia conspersa-Chloealtis corallipes-Xanthippus coronata-Trachyrhachys crenulata-Cordillacris curtipennis-Chorthippus cyaneipennis-Trimerotropis cyaneus-Leprus dawsonii-Melanoplus delicatula-Psoloessa deorum-Ageneotettix differentialis-Melanoplus discolor-Melanoplus dodgei-Melanoplus elliotti-Aulocara enigma-Oedaloenotus equale-Spharagemon fasciatus-Melanoplus femoratum-Aulocara femurrubrum-Melanoplus flavidus-Melanoplus foedus-Melanoplus fontana-Trimerotropis formosus-Tropidolophus fratercula-Trimerotropis fusiformis-Ceuthophilus gladstoni-Melanoplus glaucipes-Melanoplus gracile-Stethophyma gracilis-Trimerotropis haldemanii-Pardalophora haydeni-Derotmema hirtipes-Acrolophitus humile-Spharagemon inconspicua-Trimerotropis infantilis-Melanoplus keeleri-Melanoplus kennicotti-Melanoplus kiowa-Trachyrhachys lakinus-Melanoplus latifasciata-Trimerotropis longipennis-Dissosteira magna-Brachystola magnifica-Trimerotropis melanoptera-Trimerotropis montanus-Xanthippus montezuma-Syrbula nebrascensis-Phoetaliotes neglectus-Cratypedes nevadensis-Acrolophitus niveus-Oecanthus nubilum-Boopedon obscura-Opeia occidentalis-Melanoplus occipitalis-Cordillacris ocelote-Hippiscus olivacea-Campylacantha oregonensis-Melanoplus packardii-Melanoplus pallidipennis-Trimerotropis pardalinus-Metator parviceps-Cibolacris pelidna-Orphulella pellucida-Camnula picta-Mermiria pistrinaria-Trimerotropis plattei-Mestobregma ponderosus-Melanoplus pseudonietana-Arphia quadrimaculatum-Phlibostroma quadripunctatus-Oecanthus rabula-Circotettix regalis-Melanoplus robusta-Udeopsylla robustus-Leprus rufa-Heliaula sanguinipes-Melanoplus simplex-Anabrus simplex-Arphia simplex-Eritettix sordidus-Encoptolophus sparsa-Trimerotropis speciosa-Orphulella speciosus-Hesperotettix splendidus-Melanoplus spretus-Melanoplus spurcata-Dissosteira subgracilis-Encoptolophus sulcifrons-Conozoa tenuipennis-Aeoloplides texana-Conozoa texana-Mermiria texana-Psoloessa tolteca-Trimerotropis trifasciatus-Hadrotettix tristis-Melanoplus turnbulli-Aeoloplides utahensis-Ceuthophilus verruculatus-Trimerotropis virgata-Paropomala viridifasciata-Chortophaga viridis-Hesperotettix wyomingensis-Paropomala xanthoptera-Arphia yarrowii-Melanoplus not-true-grasshoppers Acrididae discussed-manual confused-with-grasshoppers top-of-page back-to-contents navigation-main-page learn-more biology distribution-maps fact-sheets key-to-stages grasshopper-development grasshopper-names-common-scientific grasshopper-names-species-genus grasshopper-names-genus-species Acrolophitus-hirtipes Acrolophitus-nevadensis Aeoloplides-chenopodii Aeoloplides-tenuipennis Aeoloplides-tumbulli Aeropedellus-clavatus Ageneotettix-deorum Amphitomus-coloradus Anabrus-simplex Arphia-conspersa Arphia-pseudonietana Arphia-simplex Arphia-xanthoptera Aulocara-elliotti Aulocara-femoratum Boopedon-nubilum Brachystola-magna Camnula-pellucida Campylacantha-olivacea Ceuthophilus-alpinus Ceuthophilus-fusiformis Ceuthophilus-utahensis Chloealtis-abdominalis Chloealtis-conspersa Chorthippus-curtipennis Chortophaga-viridifasciata Cibolacris-parviceps Circotettix-carlinianus Circotettix-rabula Conozoa-suicifrons Conozoa-texana Cordillacris-crenulata Cordillacris-occipitalis Cratypedes-neglectus Dactylotum-bicolor Daihnia-brevipes Derotmema-haydeni Dissosteira-carolina Dissosteira-longipennis Dissosteira-spurcata Encoptolophus-sordidus Encoptolophus-subgracilis Eritettix-simplex Gryllus-spp Hadrotettix-trifasciatus Heliaula-rufa Hesperotettix-speciosus Hesperotettix-viridis Hippiscus-ocelote Hippopedon-capito Hypochlora-alba Leprus-cyaneus Leprus-robustus Melanoplus-alpinus Melanoplus-angustipennis Melanoplus-arizonae Melanoplus-bispinosus Melanoplus-bivittatus Melanoplus-borealis Melanoplus-bowditchi Melanoplus-bruneri Melanoplus-complanatipes Melanoplus-confusus Melanoplus-dawsonii Melanoplus-differentialis Melanoplus-discolor Melanoplus-dodgei Melanoplus-fasciatus Melanoplus-femurrubrum Melanoplus-flavidus Melanoplus-foedus Melanoplus-gladstoni Melanoplus-glaucipes Melanoplus-infantilis Melanoplus-keeleri Melanoplus-kennicotti Melanoplus-lakinus Melanoplus-occidentalis Melanoplus-oregonensis Melanoplus-packardii Melanoplus-ponderosus Melanoplus-regalis Melanoplus-sanguinipes Melanoplus-splendidus Melanoplus-spretus Melanoplus-tristis Melanoplus-yarrowii Mermiria-bivittata Mermiria-picta Mermiria-texana Mestobregma-plattei Metator-pardalinus Oecanthus-argentinus Oecanthus-californicus Oecanthus-niveus Oecanthus-quadripunctatus Oedaloenotus-enigma Opeia-obscura Orphulella-pelidna Orphulella-speciosa Pardalophora-apiculata Pardalophora-haidemanii Paropomala-virgata Paropomala-wyomingensis Phlibostroma-quadrimaculatum Phoetaliotes-nebrascensis Pseudopomala-brachyptera Psoloessa-delicatula Psoloessa-texana Schistocerca-alutacea Spharagemon-bolli Spharagemon-collare Spharagemon-equale Spharagemon-humile Stenobothrus-brunneus Stethophyma-gracile Syrbula-admirabilis Syrbula-montezuma Trachyrhachys-aspera Trachyrhachys-coronata Trachyrhachys-kiowa Trimerotropis-agrestis Trimerotropis-californica Trimerotropis-campestris Trimerotropis-cincta Trimerotropis-citrina Trimerotropis-cyaneipennis Trimerotropis-fontana Trimerotropis-fratercula Trimerotropis-gracilis Trimerotropis-inconspicua Trimerotropis-latifasciata Trimerotropis-magnifica Trimerotropis-melanoptera Trimerotropis-pallidipennis Trimerotropis-pistrinaria Trimerotropis-sparsa Trimerotropis-tolteca Trimerotropis-verruculatus Tropidolophus-formosus Udeopsylla-robusta Xanthippus-corallipes Xanthippus-montanus true-grasshoppers Acrididae-discussed manual-confused grasshoppers-top page-back contents-navigation main-page learn-more-biology distribution-maps-fact sheets-key stages-grasshopper development-grasshoppers Colorado-grasshoppers Wyoming-West Entomology-GRASSHOPPERS COLORADO-GRASSHOPPER NAMES-GENUS SPECIES-GRASSHOPPER NAMES-COMMON SCIENTIFIC-GRASSHOPPER NAMES-SPECIES GENUS-abdominalis Chloealtis-Thomas Uhler-agrestis Trimerotropis-McNeill Dodge-alpinus Ceuthophilus-Scudder Scudder-alutacea Schistocerca-Scudder Dodge-apiculata Pardalophora-Harris Saussure-arizonae Melanoplus-Scudder Scudder-bicolor Dactylotum-Thomas Scudder-bivittata Mermiria-Serville Say-bolli Spharagemon-Morse Fieber-bowditchi Scudder-brevipes Daihnia-Haldeman Scudder-brunneus Stenobothrus-Thomas Bruner-californicus Oecanthus-Saussure McNeill-capito Hippopedon-Stal Thomas-carolina Dissosteira-L Bruner-cincta Trimerotropis-Thomas Scudder-clavatus Aeropedellus-Thomas Scudder-coloradus Amphitornus-Thomas Scudder-confusus Scudder-conspersa Chloealtis-Harris Haldeman-coronata Trachyrhachys-Scudder Bruner-curtipennis Chorthippus-Harris Bruner-cyaneus Leprus-Cockerell Scudder-delicatula Psoloessa-Scudder Scudder-differentialis Melanoplus-Thomas Scudder-dodgei Thomas-enigma Oedaloenotus-Scudder Say-fasciatus Melanoplus-F-Walker Scudder-femurrubrum Melanoplus-DeGeer Scudder-foedus Thomas-formosus Tropidolophus-Say McNeill-fusiformis Scudder-glaucipes Scudder-gracilis Scudder-haydeni Derotmema-Thomas Say-humile Bruner-infantilis Thomas-kennicotti Thomas-lakinus Scudder-longipennis Dissosteira-Thomas Girard-magnifica Trimerotropis-Rehn McNeill-montanus Xanthippus-Thomas Saussure-nebrascensis Phoetaliotes-Thomas Thomas-nevadensis Acrolophitus-Thomas DeGeer-nubilum Boopedon-Say Thomas-occidentalis Thomas-ocelote Hippiscus-Saussure Scudder-oregonensis Scudder-pallidipennis Trimerotropis-Burmeister Saussure-parviceps Cibolacris-Bruner Burmeister-pellucida Camnula-Scudder Walker-pistrinaria Trimerotropis-Saussure Thomas-ponderosus Thomas-quadrimaculatum Phlibostroma-Thomas Beutenmuller-rabula Circotettix-Rehn Hebard-regales Melanoplus-Dodge Haldeman-robustus Leprus-Hebard Scudder-sanguinipes Melanoplus-Fabricius Haldeman-simplex Arphia-Scudder Scudder-sordidus Encoptolophus-Burmeister Thomas-speciosa Orphulella-Scudder Scudder-splendidus Melanoplus-Hebard Walsh-spurcata Dissosteira-Saussure Caudell-sulcifrons Conozoa-Scudder Scudder-texana Conozoa-Bruner Bruner-texana Brunner-trifasciatus Hadrotettix-Say Bruner-turnbulli Aeoloplides-Caudell Thomas-verruculatus Trimerotropis-Scudder Scudder-viridifasciata Chortophaga-DeGeer Scudder-wyomingensis Paropomala-Thomas Burmeister-yarrowii species-not-true grasshoppers-Acrididae discussed-manual-because top-of-the page-back-to Contents-Navigation-Main Page-Grasshoppers-of Colorado-LEARN-MORE Biology-Distribution-Maps Fact-Sheets-Key to-Stages-of Grasshopper-Development-Biology Distribution-Maps-LEARN MORE-Distribution-Maps Fact-Sheets-LEARN MORE-Grasshoppers-of Colorado-Fact-Sheets of-Grasshopper-Development LEARN-MORE-Grasshoppers
Microterys
Microterys is a large genus of parasitoid wasps in the family Encyrtidae (Chalcidoidea), with its center of distribution in the northern parts of the Northern Hemisphere. Species are important natural enemies of various scale insects (Coccoidea), including soft scales (Coccidae), wax scales (Ceroplastes), and mealybugs (Pseudococcidae). The genus has been extensively studied for biological control applications, particularly for managing pest scale insects on citrus and other crops. Several species have been introduced to new regions as biocontrol agents, including Microterys flavus in California.
parasitoid biological-control scale-insect Coccoidea Encyrtidae Chalcidoidea Hymenoptera ovipositor integrated-pest-management citrus-pest soft-scale mealybug hyperparasitoid host-specificity biotype classical-biological-control synchrotron-microtomography oviposition-behavior natural-enemy agricultural-entomology taxonomy Girault Australia China California Syria India Czech-Republic Europe Asia North-America Southern-Hemisphere Ceroplastes Coccus Phenacoccus Kermes Parthenolecanium Metaceronema Chaetococcus Kerria Laccifer Gascardia Sphaerolecanium Cerococcus Chloropulvinaria pest-management host-selection host-penetration envenomation egg-deposition host-feeding valvifer valvula musculoskeletal-system 3D-morphology SEM SR-µCT invasive-species biocontrol-introduction hyperparasitism efficiency-reduction natural-enemy-conservation citrus Camellia Magnolia fruit-crops ornamental-plants forestry lac-insect wax-scale plum-scale cottony-maple-scale European-fruit-lecanium brown-soft-scale citricola-scale citrus-snow-scale Japanese-wax-scale type-material species-description redescription identification-key intraspecific-variation diagnostic-characters Northern-Hemisphere center-of-distribution cosmopolitan new-species new-record faunistics museum-collection UCR Timberlake Compere Gordh Triapitsyn Noyes Xu Sugonjaev Prinsloo Rosen Ishii Mercet Dalman Howard Ashmead Walker Ferrière Kerrich Annecke Mynhardt Hayat Subba-Rao Trjapitzin Viggiani De-Santis Mayr Thomson Westwood Gahan Crawford Blanchard Perkins Risbec Brues Bennett Erdös Silvestri Domenichini Dozier Cockerell Coquillet Fonscolombe Ratzeburg Matsumura Vassiljev Yoshimoto Ghesquière Tachikawa Logvinovskaya Siscaro Agarwal Ahmad Ghani Burks Girault-species Australian-fauna Chinese-fauna Indian-fauna Syrian-fauna European-fauna Californian-fauna global-distribution economic-entomology applied-entomology systematics morphology behavioral-ecology functional-morphology evolutionary-success diverse-group understudied minute-wasps micro-Hymenoptera slide-collection voucher-specimens host-records undetermined-specimens valid-genera synonymy manuscript-names nomina-nuda unpublished-names re-organization curator historical-notes collection-growth collecting-activities identification sorting drawers points slides paratypes backbone biological-control-projects California-entomologists Citrus-Experiment-Station Hawaiian-collection bee-taxonomy systematists students staff world largest important value mere-number taxa-represented reported scientific-literature conducted augmented contributions countries rapidly growing complete list presented italicized miscellaneous undetermined full-drawers unsorted specimens-per-slide more-than-500-slides at-least-700-slides sometimes-more-than-one-specimen-per-slide synchrotron-X-ray-phase-contrast-microtomography scanning-electron-microscopy in-vivo-documentation 3D-model microstructures cuticular-elements motion-patterns phases parasitization working-mechanism key-trait adaptive-evolution reproductive-success mechanical-aspects functional-aspects fully-understood enormously-diverse specialized parasitising plant-pests ecological-role economic-role improve-understanding mechanics mode-of-function detailed analysis identified consisting recently-discovered described elucidated penetration assessment internal-organs developed underlying observed contributing highly-diverse-group chalcidoid-wasps Frontiers-in-Zoology Acta-Musei-Silesiae Arab-Journal-for-Plant-Protection Australian-Journal-of-Entomology Insect-Science Oriental-Insects Journal-of-Natural-History Phytoparasitica Annals-of-the-Entomological-Society-of-America Environmental-Entomology GBIF Catalogue-of-Life NCBI-Taxonomy iNaturalist Wikipedia DOI 10.1186/s12983-025-00575-1 10.2478/cszma-2021-0007 10.22268/AJPP-41.3.292305 10.1111/j.1440-6055.1975.tb02058.x 10.1111/j.1744-7917.2000.tb00345.x 10.1080/00305316.2011.646827 10.1111/j.1440-6055.1973.tb01670.x 10.1080/00222933.2020.1786608 10.1007/s12600-019-00727-0 10.1007/s12600-018-0691-5 10.1093/aesa/54.2.222 10.1093/ee/7.6.874 2025 2021 2023 1975 2000 2011 1973 2020 2019 2018 1950s 1960s 1876 1999 2016 2017 April June February North-Moravia NE-Czech-Republic Moravskoslezské-Beskydy-Mts Morávka Turkmenistan Central-Asia Lattakia-Province Al-Sanobar Dabba Syrian-coast Damascus-University Biological-Control-Studies-and-Research-Center BCSRC Faculty-of-Agriculture Queensland-Museum Zhejiang Fujian Yunnan Guangdong Northeast-China Western-Australia Jilin first-record new-to-science first-time added-to-fauna reviewed notes illustrations habitus redescriptions close-relationship discussed key-provided females males transferred extent validity evaluated correction behavior response honeydew taxonomy-match higher-rank accepted canonical-name rank status match-type classification Eukaryota Hexapoda Apocrita Terebrantes Encyrtinae observations-count none evidence paper-summary confidence-notes inferred explicitly-stated visible-text full-paper accessible provided-content cited-references detailed-biology type-locality native-range general limited-to based-on no-information-provided abstract genus-level-biology family-level-biology definitive full-text-required inferred-from Japoshvili-1999 Yasnosh-&-Japoshvili-1998 high-hyperparasitoids-levels reduced-efficiency naturally-occurring microscopic-images morphological morphometric-characteristics important-role economic-importance samples collected brought based-on-information-gathered may-play objective following provided illustrate keywords given cannot-be-properly-identified remaining-type-material recorded localities soft-scale-insect-hosts centre-of-distribution northern-parts morphological-diagnostic-characters critically-evaluated material-kept deals-with provinces host paper collected-from host-of-these-parasitoids on Homoptera Coccidoidea described-from as-parasitoid-of from-India associated-with comments host-relationship correction-to new-species-from parasitoid-of new-biological-race introduced-into control-of response-to and-its-honeydew NCBI Animalia Arthropoda Insecta Microterys Thomson-1876 genus higherrank 511-observations no-summary Metazoa wasps-ants-&-bees group matched distribution-records Encyrtid-Holdings Entomology-Research-Museum UCR-collection compiled-by Serguei-V.-Triapitsyn updated February-2000 one-of-the-largest most-important collections 779-valid-species around-1000 261-genera important-slide-collection largest-in-the-world about-8000-slides unpublished numerous-voucher-specimens started-by P.H.-Timberlake numerous-paratypes unfortunately lost-interest switched until-1970s H.-Compere notable-contributions P.-Timberlake J.-Maple P.-DeBach C.-Clausen 1970s-1980s G.-Gordh J.-LaSalle J.-Woolley J.-Pinto J.-Beardsley Mexico US other-countries presently growing-rapidly J.-Heraty majority V.-Trjapitzin J.-Noyes complete-re-organization S.-Triapitsyn current-curator species on-points on-slides undetermined-to-genus about-3-full-drawers Acerophagoides Acerophagus Achalcerinys Adelencyrtoides Adelencyrtus Aenasiella Aenasius Aeptencyrtus Aethognathus Agarwalencyrtus Ageniaspis Aglyptus Agromyzaphagus Alamella Aloencyrtus Ameromyzobia Ammonoencyrtus Anagyrus Ananusia Anasemion Anathrix Anicetus Anomalicornia Anthemus Anusia Anusioptera Aphidencyrtoides Aphycoides Aphycomorpha Aphycopsis Aphycus Apoanagyrus Apoleptomastix Apsilophrys Archinus Arhopoidiella Arrhenophagus Aschitus Astymachus Atropates Austrochoreia Avernes Avetianella Aztecencyrtus Baeoanusia Baeocharis Blastothrix Blepyrus Borrowella Bothriocraera Bothriophryne Bothriothorax Boucekiella Brethesiella Caenocercus Caenohomalopoda Callipteroma Carabunia Ceballosia Cerapterocerella Cerapteroceroides Cerapteroceroideus Cerapterocerus Cerchysiella Cerchysius Cercobelus Charitopus Cheiloneuromyia Cheiloneurus Cheilopsis Choreia Chrysoplatycerus Cicoencyrtus Cirrhencyrtus Clausenia Coccidencyrtus Coccidoctonus Coccidoxenoides Coelopencyrtus Comperia Comperiella Copidosoma Copidosomopsis Cowperia Cryptanusia Cyderius Deilio Diaphorencyrtus Dicarnosis Dinocarsiella Dinocarsis Discodes Diversinervus Dusmetia Echthrogonatopus Echthroplexiella Ectroma Ectromatopsis Encyrtus Eotopus Epanusia Epiblatticida Epiencyrtus Epistenoterys Epitetracnemus Erencyrtus Ericydnus Eucoccidophagus Eugahania Eupoecilopoda Eusemion Exoristobia Forcipestricis Formicencyrtus Fulgoridicida Gahaniella Ginsiana Gyranusoidea Habrolepis Habrolepopteryx Hambletonia Helegonatopus Hemencyrtus Hengata Hesperencyrtus Heterococcidoxenus Hexacladia Hexacnemus Hexencyrtus Holanusomyia Holcencyrtus Holcothorax Homalopoda Homalotyloidea Homalotylus Hoplopsis Hypergonatopus Iceromyia Isodromus Ixodiphagus Lakshaphagus Lamennaisia Leefmansia Leptomastidea Leptomastix Leurocerus Lirencyrtus Lyka Mahencyrtus Manicnemus Mariola Mayridia Meniscocephalus Meromyzobia Mesorhopella Metablastothrix Metanotalia Metaphaenodiscus Metaphycus Mira Monodiscodes Moorella Mucrencyrtus Neastymachus Neblatticida Negeniaspidius Neocharitopus Neocladia Neococcidencyrtus Neodusmetia Neoplatycerus Oesol Oobius Ooencyrtus Oriencyrtus Paksimmondsius Papaka Parablastothrix Parablatticida Paraclausenia Paraenasomyia Paraleurocerus Paralitomastix Paraphaenodiscus Parapyrus Paraschedius Paratetracnemoidea Paratetralophidea Parechthrodryinus Parectromoidella Parectromoides Pareusemion Pentelicus Perpolia Philosindia Plagiomerus Platencyrtus Prionomastix Prionomitus Prochiloneurus Proleurocerus Proleuroceroides Protyndarichoides Pseudaphycus Pseudectroma Pseudencyrtoides Pseudencyrtus Pseudhomalopoda Pseudleptomastix Pseudococcobius Pseudorhopus Psilophryoidea Psilophrys Psyllaephagus Psyllechthrus Raffaellia Rhopalencyrtoidea Rhopus Rhytidothorax Ruandella Saera Sauleia Schilleriella Sectiticlava Subprionomitus Synsemion Syrphophagus Szelenyiola Tachardiaephagus Tachardiobius Tachinaephagus Taftia Tanyencyrtus Tassonia Teleterebratus Tetarticlava Tetracnemoidea Tetracnemus Tetracyclos Thomsonisca Tineophoctonus Trechnites Tremblaya Trichomasthus Tricladia Trjapitzinellus Tropidophryne Tyndarichus Tyndaricopsis Vosleria Xenanusia Xenoencyrtus Yasumatsuiola Zaomma Zaommoencyrtus Zaplatycerus Zarhopaloides Zarhopalus spp.sp.sp(p).?nr.ms-name manuscript-name nomen-nudum including probably-mixed-with unsorted-miscellaneous University-of-California-Riverside collection museum specimens vouchers hosts parasitoids scale-insects mealybugs soft-scales wax-scales citrus-pests agricultural-pests natural-enemies hyperparasitoids efficiency oviposition anatomy 3D-reconstruction microtomography in-vivo-observation evolution adaptive-radiation diversity Africa South-America Middle-East Hawaii horticulture crop-protection augmentative-biological-control conservation-biological-control egg-laying population-dynamics ecosystem-services natural-pest-control sustainable-agriculture organic-farming biodiversity invasive-species-management quarantine phylogeny diagnosis description new-records biogeography museum-collections historical-collections type-specimens curatorial-practices collection-management data-quality scientific-infrastructure research-capacity collaboration knowledge-transfer education outreach public-engagement science-communication open-access data-sharing FAIR-principles digitalization imaging microscopy tomography 3D-modeling computational-biology biomechanics kinematics muscle-physiology neuroethology sensory-ecology chemical-ecology host-location host-recognition host-acceptance host-suitability foraging-behavior learning memory decision-making life-history development reproduction survival mortality population-ecology community-ecology trophic-interactions food-webs ecosystem-functioning ecosystem-stability resilience sustainability environmental-change climate-change land-use-change habitat-fragmentation pollution pesticide-use biological-invasions conservation restoration rewilding nature-based-solutions green-infrastructure urban-ecology agroecology ecological-intensification sustainable-intensification climate-smart-agriculture precision-agriculture digital-agriculture smart-farming agricultural-technology biotechnology genetic-improvement breeding selection biocontrol-agent-improvement mass-rearing release-strategies monitoring evaluation impact-assessment risk-assessment regulatory-frameworks policy governance stakeholder-engagement farmer-adoption socio-economics cost-benefit-analysis return-on-investment livelihoods food-security nutrition health wellbeing equity justice ethics responsible-innovation precautionary-principle biosafety biosecurity One-Health planetary-health sustainability-science futures-studies scenario-planning adaptive-management transdisciplinary-research participatory-research citizen-science indigenous-knowledge traditional-ecological-knowledge local-knowledge expert-knowledge scientific-knowledge knowledge-integration knowledge-co-production boundary-work science-policy-interface evidence-based-policy policy-relevant-research research-impact societal-impact transformational-change system-change paradigm-shift new-normal Anthropocene Great-Acceleration planetary-boundaries safe-operating-space Doughnut-Economics circular-economy regenerative-agriculture agroforestry permaculture biodynamic-agriculture nature-farming ecological-agriculture organic-agriculture low-input-agriculture conservation-agriculture sustainable-food-systems healthy-diets sustainable-consumption-and-production Sustainable-Development-Goals Agenda-2030 Paris-Agreement Convention-on-Biological-Diversity IPBES IPCC FAO WHO UNEP UNDP World-Bank CGIAR national-agricultural-research-systems universities museums herbaria botanical-gardens zoos aquaria arboreta seed-banks gene-banks culture-collections microbial-resource-centers biorepositories cryobanks digital-repositories data-banks knowledge-banks 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