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Dielis tolteca
Toltec scoliid wasp
Dielis tolteca is a species of scoliid wasp native to western North America and Mesoamerica. The species is known to parasitize scarab beetle grubs, with females hunting underground hosts to provision their offspring. Adults have been observed feeding on flowering plants, particularly mustards (Brassicaceae) and goldenrods (Solidago). The species has been documented in McInnis Canyons National Conservation Area in western Colorado, where it is active as a spring-emerging species.
scoliid-wasp parasitoid scarab-beetle western-Colorado McInnis-Canyons spring-emergence Solidago mustards biological-control Hymenoptera Scoliidae Dielis Campsomerini biodiversity-survey integrated-pest-management melissa-schreiner extension-entomology saussure-1857 toltec-scoliid-wasp riparian Colorado-River flower-visitor nectar-feeding solitary-wasp ground-nesting-parasitoid scarab-grub-parasitoid aculeate apocrita aculeata campsomerinae western-North-America Nearctic Neotropical Mesoamerica Arizona Baja-California Sonora underground-host-hunter external-parasitoid larval-provisioning spring-active insect-biodiversity rare-ecosystem-survey natural-heritage-survey CSU-extension Colorado-State-University agricultural-pest-management tree-fruit wine-grapes row-crops invasive-species-monitoring Japanese-beetle Popillia-japonica corn-earworm Helicoverpa-zea IPM pest-monitoring moth-trapping B3-allele-testing citizen-science Department-of-Agriculture-trapping targeted-larval-treatments 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undergraduate-research bee-pollinator-identification American-University-Beirut genetic-diversity-assessment Spartium-junceum-populations Spanish-broom rush-broom weaver's-broom insect-collection-establishment NDU-specimen-collection UC-Davis-team national-Linnaean-Games 2015-Entomological-Society-of-America-meeting Entomology-Games University-of-Florida 32-year-history-first-win video Bohart-Museum-location Room-1124-Academic-Surge-Building Crocker-Lane currently-closed-public live-petting-zoo Madagascar-hissing-cockroaches stick-insects tarantulas insect-themed-gift-shop online Shahid-Siddique assistant-professor seminar-coordinator Zoom-technical-issues [email protected]seminar-list Trisciloa-saussurei New-Guinea-native Bohart-Museum-specimen Bug-Eric Wasp-Wednesday Campsomeris December-22-2010 mystery-wasp December email-friend species-recorded-recently-Arizona Mexican-border Sabino-Canyon-Recreation-Area Fred-Heath outstanding-naturalist Introduction-to-Southern-California-Butterflies Sabino-Canyon-Volunteer-Naturalists December-14 male-specimen Campsomeris-ephippium Desert-Lavender Hyptis-emoryi conspicuous-distinctive no-image-needed email-list-message David-Lazaroff SCVN-founder camera image-permission long-antennae slender-body pseudostinger posterior-abdomen gender-reveal robust-females shorter-antennae real-stinger retractable-stinger hosts-larval-offspring spring-2009-image south-Texas south-to-Ecuador Sunday-December-19 failed-to-find common-local-species Campsomeris-tolteca males-feeding nectar-Coreocarpus-arizonicus Little-Lemonhead Queen-butterflies Mexican-Yellows Scoliidae-family all-parasitoids parasitoid-definition heavy-spiny-legs dig-up-scarab-grub sting-brief-paralysis single-egg host-larva leave-scene host-regains-consciousness motor-skills underground-existence plant-roots-feeding wasp-egg-hatches external-parasite beetle-grub pleasure-of-hunt Sabino-Canyon Fred-Heath-outdoors David-Lazaroff-image 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clavatus-Aeropedellus collare-Spharagemon coloradus-Amphitornus complanatipes-Melanoplus confusus-Melanoplus conspersa-Arphia conspersa-Chloealtis corallipes-Xanthippus coronata-Trachyrhachys crenulata-Cordillacris curtipennis-Chorthippus cyaneipennis-Trimerotropis cyaneus-Leprus dawsonii-Melanoplus delicatula-Psoloessa deorum-Ageneotettix differentialis-Melanoplus discolor-Melanoplus dodgei-Melanoplus elliotti-Aulocara enigma-Oedaloenotus equale-Spharagemon fasciatus-Melanoplus femoratum-Aulocara femurrubrum-Melanoplus flavidus-Melanoplus foedus-Melanoplus fontana-Trimerotropis formosus-Tropidolophus fratercula-Trimerotropis fusiformis-Ceuthophilus gladstoni-Melanoplus glaucipes-Melanoplus gracile-Stethophyma gracilis-Trimerotropis haldemanii-Pardalophora haydeni-Derotmema hirtipes-Acrolophitus humile-Spharagemon inconspicua-Trimerotropis infantilis-Melanoplus keeleri-Melanoplus kennicotti-Melanoplus kiowa-Trachyrhachys lakinus-Melanoplus latifasciata-Trimerotropis longipennis-Dissosteira magna-Brachystola magnifica-Trimerotropis melanoptera-Trimerotropis montanus-Xanthippus montezuma-Syrbula nebrascensis-Phoetaliotes neglectus-Cratypedes nevadensis-Acrolophitus niveus-Oecanthus nubilum-Boopedon obscura-Opeia occidentalis-Melanoplus occipitalis-Cordillacris ocelote-Hippiscus olivacea-Campylacantha oregonensis-Melanoplus packardii-Melanoplus pallidipennis-Trimerotropis pardalinus-Metator parviceps-Cibolacris pelidna-Orphulella pellucida-Camnula picta-Mermiria pistrinaria-Trimerotropis plattei-Mestobregma ponderosus-Melanoplus pseudonietana-Arphia quadrimaculatum-Phlibostroma quadripunctatus-Oecanthus rabula-Circotettix regalis-Melanoplus robusta-Udeopsylla robustus-Leprus rufa-Heliaula sanguinipes-Melanoplus simplex-Anabrus simplex-Arphia simplex-Eritettix sordidus-Encoptolophus sparsa-Trimerotropis speciosa-Orphulella speciosus-Hesperotettix splendidus-Melanoplus spretus-Melanoplus spurcata-Dissosteira subgracilis-Encoptolophus sulcifrons-Conozoa tenuipennis-Aeoloplides texana-Conozoa texana-Mermiria texana-Psoloessa tolteca-Trimerotropis trifasciatus-Hadrotettix tristis-Melanoplus turnbulli-Aeoloplides utahensis-Ceuthophilus verruculatus-Trimerotropis virgata-Paropomala viridifasciata-Chortophaga viridis-Hesperotettix wyomingensis-Paropomala xanthoptera-Arphia yarrowii-Melanoplus not-true-grasshoppers Acrididae discussed-manual confused-with-grasshoppers top-of-page back-to-contents navigation-main-page learn-more biology distribution-maps fact-sheets key-to-stages grasshopper-development grasshopper-names-common-scientific grasshopper-names-species-genus grasshopper-names-genus-species Acrolophitus-hirtipes Acrolophitus-nevadensis Aeoloplides-chenopodii Aeoloplides-tenuipennis Aeoloplides-tumbulli Aeropedellus-clavatus Ageneotettix-deorum Amphitomus-coloradus Anabrus-simplex Arphia-conspersa Arphia-pseudonietana Arphia-simplex Arphia-xanthoptera Aulocara-elliotti Aulocara-femoratum Boopedon-nubilum Brachystola-magna Camnula-pellucida Campylacantha-olivacea Ceuthophilus-alpinus Ceuthophilus-fusiformis Ceuthophilus-utahensis Chloealtis-abdominalis Chloealtis-conspersa Chorthippus-curtipennis Chortophaga-viridifasciata Cibolacris-parviceps Circotettix-carlinianus Circotettix-rabula Conozoa-suicifrons Conozoa-texana Cordillacris-crenulata Cordillacris-occipitalis Cratypedes-neglectus Dactylotum-bicolor Daihnia-brevipes Derotmema-haydeni Dissosteira-carolina Dissosteira-longipennis Dissosteira-spurcata Encoptolophus-sordidus Encoptolophus-subgracilis Eritettix-simplex Gryllus-spp Hadrotettix-trifasciatus Heliaula-rufa Hesperotettix-speciosus Hesperotettix-viridis Hippiscus-ocelote Hippopedon-capito Hypochlora-alba Leprus-cyaneus Leprus-robustus Melanoplus-alpinus Melanoplus-angustipennis Melanoplus-arizonae Melanoplus-bispinosus Melanoplus-bivittatus Melanoplus-borealis Melanoplus-bowditchi Melanoplus-bruneri Melanoplus-complanatipes Melanoplus-confusus Melanoplus-dawsonii Melanoplus-differentialis Melanoplus-discolor Melanoplus-dodgei Melanoplus-fasciatus Melanoplus-femurrubrum Melanoplus-flavidus Melanoplus-foedus Melanoplus-gladstoni Melanoplus-glaucipes Melanoplus-infantilis Melanoplus-keeleri Melanoplus-kennicotti Melanoplus-lakinus Melanoplus-occidentalis Melanoplus-oregonensis Melanoplus-packardii Melanoplus-ponderosus Melanoplus-regalis Melanoplus-sanguinipes Melanoplus-splendidus Melanoplus-spretus Melanoplus-tristis Melanoplus-yarrowii Mermiria-bivittata Mermiria-picta Mermiria-texana Mestobregma-plattei Metator-pardalinus Oecanthus-argentinus Oecanthus-californicus Oecanthus-niveus Oecanthus-quadripunctatus Oedaloenotus-enigma Opeia-obscura Orphulella-pelidna Orphulella-speciosa Pardalophora-apiculata Pardalophora-haidemanii Paropomala-virgata Paropomala-wyomingensis Phlibostroma-quadrimaculatum Phoetaliotes-nebrascensis Pseudopomala-brachyptera Psoloessa-delicatula Psoloessa-texana Schistocerca-alutacea Spharagemon-bolli Spharagemon-collare Spharagemon-equale Spharagemon-humile Stenobothrus-brunneus Stethophyma-gracile Syrbula-admirabilis Syrbula-montezuma Trachyrhachys-aspera Trachyrhachys-coronata Trachyrhachys-kiowa Trimerotropis-agrestis Trimerotropis-californica Trimerotropis-campestris Trimerotropis-cincta Trimerotropis-citrina Trimerotropis-cyaneipennis Trimerotropis-fontana Trimerotropis-fratercula Trimerotropis-gracilis Trimerotropis-inconspicua Trimerotropis-latifasciata Trimerotropis-magnifica Trimerotropis-melanoptera Trimerotropis-pallidipennis Trimerotropis-pistrinaria Trimerotropis-sparsa Trimerotropis-tolteca Trimerotropis-verruculatus Tropidolophus-formosus Udeopsylla-robusta Xanthippus-corallipes Xanthippus-montanus true-grasshoppers Acrididae-discussed manual-confused grasshoppers-top page-back contents-navigation main-page learn-more-biology distribution-maps-fact sheets-key stages-grasshopper development-grasshoppers Colorado-grasshoppers Wyoming-West Entomology-GRASSHOPPERS COLORADO-GRASSHOPPER NAMES-GENUS SPECIES-GRASSHOPPER NAMES-COMMON SCIENTIFIC-GRASSHOPPER NAMES-SPECIES GENUS-abdominalis Chloealtis-Thomas Uhler-agrestis Trimerotropis-McNeill Dodge-alpinus Ceuthophilus-Scudder Scudder-alutacea Schistocerca-Scudder Dodge-apiculata Pardalophora-Harris Saussure-arizonae Melanoplus-Scudder Scudder-bicolor Dactylotum-Thomas Scudder-bivittata Mermiria-Serville Say-bolli Spharagemon-Morse Fieber-bowditchi Scudder-brevipes Daihnia-Haldeman Scudder-brunneus Stenobothrus-Thomas Bruner-californicus Oecanthus-Saussure McNeill-capito Hippopedon-Stal Thomas-carolina Dissosteira-L Bruner-cincta Trimerotropis-Thomas Scudder-clavatus Aeropedellus-Thomas Scudder-coloradus Amphitornus-Thomas Scudder-confusus Scudder-conspersa Chloealtis-Harris Haldeman-coronata Trachyrhachys-Scudder Bruner-curtipennis Chorthippus-Harris Bruner-cyaneus Leprus-Cockerell Scudder-delicatula Psoloessa-Scudder Scudder-differentialis Melanoplus-Thomas Scudder-dodgei Thomas-enigma Oedaloenotus-Scudder Say-fasciatus Melanoplus-F-Walker Scudder-femurrubrum Melanoplus-DeGeer Scudder-foedus Thomas-formosus Tropidolophus-Say McNeill-fusiformis Scudder-glaucipes Scudder-gracilis Scudder-haydeni Derotmema-Thomas Say-humile Bruner-infantilis Thomas-kennicotti Thomas-lakinus Scudder-longipennis Dissosteira-Thomas Girard-magnifica Trimerotropis-Rehn McNeill-montanus Xanthippus-Thomas Saussure-nebrascensis Phoetaliotes-Thomas Thomas-nevadensis Acrolophitus-Thomas DeGeer-nubilum Boopedon-Say Thomas-occidentalis Thomas-ocelote Hippiscus-Saussure Scudder-oregonensis Scudder-pallidipennis Trimerotropis-Burmeister Saussure-parviceps Cibolacris-Bruner Burmeister-pellucida Camnula-Scudder Walker-pistrinaria Trimerotropis-Saussure Thomas-ponderosus Thomas-quadrimaculatum Phlibostroma-Thomas Beutenmuller-rabula Circotettix-Rehn Hebard-regales Melanoplus-Dodge Haldeman-robustus Leprus-Hebard Scudder-sanguinipes Melanoplus-Fabricius Haldeman-simplex Arphia-Scudder Scudder-sordidus Encoptolophus-Burmeister Thomas-speciosa Orphulella-Scudder Scudder-splendidus Melanoplus-Hebard Walsh-spurcata Dissosteira-Saussure Caudell-sulcifrons Conozoa-Scudder Scudder-texana Conozoa-Bruner Bruner-texana Brunner-trifasciatus Hadrotettix-Say Bruner-turnbulli Aeoloplides-Caudell Thomas-verruculatus Trimerotropis-Scudder Scudder-viridifasciata Chortophaga-DeGeer Scudder-wyomingensis Paropomala-Thomas Burmeister-yarrowii species-not-true grasshoppers-Acrididae discussed-manual-because top-of-the page-back-to Contents-Navigation-Main Page-Grasshoppers-of Colorado-LEARN-MORE Biology-Distribution-Maps Fact-Sheets-Key to-Stages-of Grasshopper-Development-Biology Distribution-Maps-LEARN MORE-Distribution-Maps Fact-Sheets-LEARN MORE-Grasshoppers-of Colorado-Fact-Sheets of-Grasshopper-Development LEARN-MORE-Grasshoppers
Oreoneta
A genus of dwarf spiders (family Linyphiidae) established in 1894, comprising approximately 30 species distributed across cold-temperate and Arctic regions of the Northern Hemisphere. Most species inhabit high-latitude or high-altitude environments. The genus was revised extensively by Saaristo & Marusik in 2004, resulting in numerous new species descriptions.
Arachnida Araneae Linyphiidae dwarf-spider Arctic boreal Holarctic tundra montane predator cold-adapted ground-dwelling high-latitude high-altitude Saaristo Marusik Chyzer Kulczyński 1894 2004-revision genitalic-characters epigynum embolus conductor palaearctic nearctic Eurasia North-America Scandinavia Siberia Alaska Canada Greenland Kazakhstan Mongolia China Kurile-Islands Finland Sweden Norway Germany Switzerland Italy Slovakia Bulgaria Central-Europe type-species:-Oreoneta-frigida thirty-species 30-species small-spider sheet-web-weaver money-spider Linyphiinae Erigoninae microhabitat litter-dwelling cryptic understudied taxonomic-revision morphological-taxonomy light-microscopy species-description biogeography glacial-refugia post-glacial-colonization endemic regional-endemic alpine-endemic Arctic-endemic boreal-forest taiga rocky-habitat meadow wetland stream-margin snowbed fellfield heath moss lichen vegetation soil-surface stone rock boulder scree talus debris humus organic-layer permafrost polar subpolar cold-stenotherm stenotherm stenotopic habitat-specialist conservation rare data-deficient IUCN red-list natural-history systematics phylogeny molecular-systematics DNA-barcoding COI cytochrome-oxidase 28S 18S ITS morphology taxonomy alpha-taxonomy revision monograph faunistics chorology zoogeography dispersal vicariance speciation cryptic-species species-complex species-group species-aggregate subspecies infraspecific variation polymorphism sexual-dimorphism male female juvenile instar ecdysis molting development growth reproduction mating courtship sperm-transfer spermatophore pedipalp palpal-organ copulatory-organ genitalia epigyne vulva spermatheca fertilization-duct copulatory-duct atrium scape maceration dissection microscope stereomicroscope compound-microscope illustration drawing photography SEM scanning-electron-microscopy habitus dorsal ventral lateral prosoma opisthosoma abdomen cephalothorax carapace sternum chelicera fang leg tarsus metatarsus tibia patella femur coxa trochanter spinneret colulus tracheal-spiracle book-lung cuticle seta hair spine macroseta sensillum chaetotaxy eye oculus ocelli anterior-median-eye posterior-median-eye anterior-lateral-eye posterior-lateral-eye eye-tubercle clypeus fovea pedicel waist constriction color pattern pigmentation melanism albinism size measurement total-length carapace-length carapace-width abdomen-length leg-length leg-formula index ratio proportion allometry ontogeny subadult adult maturity sexual-maturity reproductive-maturity longevity life-span generation-time voltinism univoltine semivoltine biennial perennial overwintering diapause quiescence cold-hardiness freeze-tolerance freeze-avoidance supercooling cryoprotectant glycerol sorbitol trehalose thermal-biology thermoregulation behavioral-thermoregulation postural-thermoregulation activity diurnal nocturnal crepuscular matutinal vespertine circadian rhythm seasonal phenology calendar date month activity-period breeding-season oviposition egg egg-sac cocoon silk web sheet-web tangle-web space-web platform-web dome-web bowl-web funnel retreat signal-line dragline attachment-disc ampullate-gland cylindriform-gland aciniform-gland aggregate-gland flagelliform-gland tubuliform-gland piriform-gland major-ampullate minor-ampullate silk-protein spidroin fibroin sericin glue viscid dry capture-silk dragline-silk safety-line anchor-line frame-line radial-line spiral-line sticky-spiral non-sticky-spiral hub perimeter substrate niche guild functional-group trophic-level carnivore zoophage arthropodivore insectivore prey capture attack subdue immobilize kill feed digest extraoral-digestion sucking-stomach gut midgut hindgut Malpighian-tubule excretion osmoregulation metabolism respiration trachea spiracle gas-exchange circulation hemolymph heart aorta sinus nervous-system brain ganglion nerve sensory-organ mechanoreceptor chemoreceptor thermoreceptor hygroreceptor photoreceptor proprioceptor trichobothrium tarsal-organ slit-sensillum lyriform-organ pedipalp-organ Haller's-organ contact-chemoreceptor gustatory olfactory pheromone semiochemical kairomone allomone synomone communication signal vibration seismic airborne visual chemical tactile copulation insemination sperm spermatozoon median-apophysis terminal-apophysis paracymbium cymbium bulbus tegulum subtegulum radix lamella process apophysis flange hook coil spiral duct reservoir sclerite plate field fossa pocket receptaculum spermathecae gland insemination-duct uterus uterus-externus uterus-internus vagina gonopore ovipositor ovary oviduct follicle vitellogenesis oogenesis spermatogenesis testis vas-deferens ejaculatory-duct seminal-vesicle fecundity fertility clutch-size egg-number hatching emergence ballooning aerial-dispersal silk-dragline gossamer migration movement walking running jumping climbing burrowing hiding refuge shelter defense predation predator-avoidance antipredator crypsis camouflage mimicry aposematism deimatic thanatosis autotomy leg-loss regeneration injury wound healing immune-response hemocyte melanization encapsulation nodule parasite parasitoid pathogen fungus bacterium virus nematode mite grooming cleaning hygiene self-maintenance silk-use web-use prey-capture foraging hunting ambush sit-and-wait active-hunting searching probing testing exploration learning memory cognition behavior ethology ecology population community ecosystem biome biosphere habitat environment abiotic biotic factor temperature humidity moisture precipitation rain snow wind light photoperiod day-length radiation UV soil cover complexity heterogeneity productivity energy nutrient carbon nitrogen phosphorus food-web trophic-interaction predator-prey competition intraspecific interspecific resource partitioning coexistence diversity richness evenness abundance density frequency occurrence presence absence detection sampling collection pitfall-trap quadrat transect hand-collecting beating sifting extraction Berlese Tullgren Winkler flotation sorting preservation ethanol formalin mount slide label catalog database repository museum voucher type holotype paratype syntype lectotype paralectotype neotype topotype original data locality georeference coordinate latitude longitude elevation altitude depth collector determination identification expert specialist literature publication journal article paper book thesis dissertation report checklist online digital archive bibliography reference citation author year volume page figure map table appendix supplementary material methods results discussion conclusion abstract keyword index-term subject topic discipline science biology zoology arachnology entomology evolution phylogenetics biological nature protection management policy legislation restoration monitoring assessment indicator flagship umbrella keystone engineer service function dynamics stability resilience resistance recovery succession disturbance climate change warming glacier range shift expansion contraction fragmentation isolation connectivity corridor network reserve park protected area wilderness urban rural agricultural forestry mining impact threat risk vulnerability sensitivity exposure adaptive capacity mitigation adaptation sustainable use harvest trade economy medicine research education outreach citizen engagement awareness appreciation aesthetic cultural spiritual existence value intrinsic instrumental utilitarian anthropocentric biocentric ecocentric ethic philosophy history discovery expedition voyage survey inventory fauna biota biodiversity life earth planet wildlife animal invertebrate arthropod chelicerate arachnid spider araneomorph entelegyne micryphantid Oreoneta 2004 frigida type-species alpine feeding type-specimen climate-change protected-area
Pentodontini
rhinoceros beetles
Pentodontini is the most diverse tribe within the subfamily Dynastinae (rhinoceros beetles), containing over 100 genera distributed across multiple biogeographic regions. Most genera are restricted to a single biogeographic region. The tribe is characterized by substantial morphological diversity, with generic-level identification often relying on mouthpart morphology in females and secondary sexual characters (horns, claw modifications, antennal club length) in males.
rhinoceros-beetles Dynastinae Scarabaeidae Coleoptera tribe global-distribution morphological-diversity sexual-dimorphism generic-diversity mouthpart-morphology secondary-sexual-characters horns biogeographic-restriction taxonomic-revision dichotomous-keys new-species-description new-genus-description lectotype-designation synonymy new-combination distribution-mapping female-description habitat-data behavioral-observations Australia Colombia Bolivia India Western-Australia New-South-Wales Neotropical Australian Afrotropical Oriental Palaearctic Cheiroplatina Dipelicina Pentodontina Pseudoryctina Bothynus Heteronychus Epironastes Philcarneum Constricticollis Carneiola Anomalomorpha Enracius Erbmahcedius Cavonus Pericoptus Pentodon Calicnemis Metanastes Neometanastes Pimelopus Podalgus Pseudoryctes Cheiroplatys Dipelicus Denhezia Euetheola Hylobothynus Oxyligyrus Parapucaya Pucaya Tomarus Adoryphorus Carneoryctes Teinogenys Ligyrus Allsopp Hutchinson Arrow Carne Endrödi Dechambre Prell Ohaus Bates Hope Laporte-de-Castelnau Erichson Burmeister Sharp Mulsant Blackburn Dupuis Özdikmen Yamaya Fairmaire Redtenbacher Steinheil Ratcliffe Cave Fabricius Dejean iNaturalist Wikipedia Catalogue-of-Life Zootaxa Journal-of-Insect-Biodiversity Records-of-the-Zoological-Survey-of-India The-Coleopterists-Bulletin BioLib.cz Wikimedia-Commons DOI 10.11646/zootaxa.4048.4.1 10.11646/zootaxa.4852.4.2 10.11646/zootaxa.5351.3.2 10.26515/rzsi/v125/i2s/2025/172964 10.11646/zootaxa.5716.4.7 10.11646/zootaxa.5072.5.2 10.11646/zootaxa.4852.4.3 10.12976/jib/2024.54.2.2 10.1649/1186.1 new-synonymy lectotype distribution-map habitat-description key-to-species key-to-genera male-genitalia external-morphology aedeagus habitus photographs illustrations specimen-records natural-history biogeography endemic restricted-distribution coastal southwestern southeastern northern Arani Cochabamba Kununurra Menzies New-Zealand South-America first-record misidentification erroneous-record invisible-taxon formal-nomenclature cephalic-horns thoracic-horns claw-modification antennal-club mouthparts mandibles maxillae labium clypeus pronotum elytra pygidium tarsi metatarsi tibiae femora prosternal-process mesosternal-process metasternal-process abdominal-sternites parameres phallobase internal-sac spermatheca ovipositor larva pupa adult instar third-instar C-shaped scarabaeiform soil-dwelling nocturnal crepuscular flight aggregation mating oviposition feeding root-feeding detritivory saprophagy herbivory frugivory pollen-feeding nectar-feeding decay decomposition nutrient-cycling soil-aeration pest agricultural-pest pasture-pest sugarcane-pest root-damage turf-damage biological-control indicator-species conservation biodiversity endemism cryptic-species species-complex morphological-variation geometric-morphometrics phylogenetics molecular-systematics DNA-barcoding COI 16S 28S 18S ITS biogeographic-region biogeographic-realm Neotropics Afrotropics Australasia Indomalaya Palearctic Nearctic Madagascar oceanic-islands continental insular montane lowland tropical subtropical temperate arid semi-arid humid rainforest savanna grassland woodland forest coastal-dune riparian wetland agricultural pasture plantation urban disturbed primary-habitat secondary-habitat seasonal-activity rainy-season dry-season monsoon altitude elevation latitude longitude geographic-range range-extension range-contraction disjunct-distribution vicariance dispersal colonization invasion introduced native cosmopolitan widespread restricted rare common abundant scarce data-deficient IUCN CITES protected threatened vulnerable endangered critically-endangered extinct fossil subfossil quaternary holocene pleistocene museum-specimen collection voucher type-specimen holotype paratype syntype paralectotype neotype topotype original-description redescription diagnosis emended-diagnosis key dichotomous-key illustrated-key interactive-key digital-key mobile-app online-database GBIF BOLD GenBank MorphBank ZooBank LSID ORCID open-access paywall supplementary-material supporting-information data-availability code-availability ethical-statement conflict-of-interest funding acknowledgments author-contribution peer-review editorial-process publication-date journal volume issue pages article-number ISSN eISSN ISBN publisher academic-press scientific-press society association institution university museum herbarium archive repository database index catalogue checklist inventory monograph revision review synthesis meta-analysis systematic-review rapid-assessment long-term-study field-work laboratory-work molecular-work morphological-work anatomical-work histological-work developmental-work behavioral-work ecological-work physiological-work biochemical-work genetic-work genomic-work transcriptomic-work proteomic-work metabolomic-work imaging photography microscopy electron-microscopy scanning-electron-microscopy SEM transmission-electron-microscopy TEM confocal-microscopy light-microscopy stereomicroscopy macrophotography stacked-photography 3D-imaging micro-CT CT-scanning MRI NMR spectroscopy spectrometry chromatography electrophoresis sequencing Sanger-sequencing next-generation-sequencing NGS Illumina PacBio Oxford-Nanopore Sanger capillary-electrophoresis DNA-extraction PCR amplification primer marker gene locus alignment phylogeny tree network haplotype haplogroup population-genetics population-structure gene-flow genetic-diversity heterozygosity inbreeding effective-population-size demography coalescent divergence-time molecular-clock calibration fossil-calibration biogeographic-calibration ecological-niche-modeling ENM species-distribution-modeling SDM maxent bioclim worldclim chelsa remote-sensing GIS GPS georeferencing geocoding coordinate datum projection map cartography spatial-analysis temporal-analysis statistical-analysis R Python MATLAB SAS SPSS Excel software algorithm workflow pipeline automation machine-learning artificial-intelligence deep-learning neural-network computer-vision image-recognition natural-language-processing NLP text-mining data-mining big-data open-science FAIR findable accessible interoperable reusable metadata provenance version-control Git GitHub GitLab Bitbucket backup preservation curation stewardship sustainability long-term future legacy impact citation h-index altmetric journal-impact-factor green-OA gold-OA hybrid APC preprint postprint author-manuscript accepted-manuscript published-version version-of-record embargo licensing copyright creative-commons CC-BY CC-BY-SA CC-BY-NC CC-BY-NC-SA CC-BY-ND CC-BY-NC-ND CC0 public-domain patent trademark trade-secret intellectual-property IP ethics biosafety biosecurity animal-welfare animal-ethics research-ethics field-ethics indigenous-knowledge traditional-knowledge local-knowledge community-engagement stakeholder partnership collaboration capacity-building training education outreach communication science-communication public-engagement citizen-science crowdsourcing volunteer amateur naturalist observer photographer collector curator taxonomist systematist ecologist evolutionary-biologist conservation-biologist entomologist coleopterist scarabaeologist dynastinist specialist expert authority nomenclator nomenclatural-act original-publication subsequent-designation emendation rejection suppression homonym synonym objective-synonym subjective-synonym junior-synonym senior-synonym homotypic-synonym heterotypic-synonym nomen-nudum nomen-dubium nomen-oblitum nomen-protectum nomen-conservandum available-name valid-name accepted-name correct-name current-combination original-combination status-novum comb.-nov.syn.-nov.sp.-nov.gen.-nov.subgen.-nov.subsp.-nov.nom.-nov.nom.-nud.nom.-dub.incertae-sedis unplaced unassigned subtribe genus subgenus species subspecies variety form morph ecotype population stock strain breed cultivar clone lineage biotype karyotype chromosome genome transcriptome proteome metabolome phenome morphome anatomical morphological meristic morphometric allometric ontogenetic developmental life-history reproductive behavioral ecological physiological biochemical genetic genomic evolutionary phylogenetic phylogeographic biogeographic historical paleontological archaeological geological climatological environmental applied economic medical veterinary forestry horticultural industrial pollution climate-change global-warming habitat-loss fragmentation deforestation agricultural-intensification urbanization invasive-species pathogen parasite predator competition mutualism symbiosis commensalism amensalism neutralism facilitation inhibition trophic-cascade food-web food-chain energy-flow ecosystem-function ecosystem-service provisioning regulating supporting cultural alpha-diversity beta-diversity gamma-diversity species-richness species-evenness species-abundance rarefaction extrapolation asymptotic non-asymptotic sample sampling survey monitoring assessment evaluation indicator bioindicator flagship umbrella keystone ecosystem-engineer foundation-species dominant subordinate frequent occasional accidental vagrant migrant resident breeding non-breeding wintering summering migration nomadism irruption establishment naturalization spreading range-expansion range-shift extirpation extinction local-extinction global-extinction functional-extinction ecological-extinction pseudextinction Lazarus-taxon Elvis-taxon zombie-taxon living-fossil relict indigenous autochthonous allochthonous exotic alien invasive naturalized casual escaped cultivated ornamental pet food medicine fiber fuel timber fodder pollinator pest-control seed-dispersal soil-formation erosion-control water-purification air-purification carbon-sequestration climate-regulation disease-regulation pest-regulation flood-regulation storm-protection recreation tourism aesthetic spiritual cultural-heritage research inspiration existence-value option-value bequest-value intrinsic-value anthropocentric biocentric ecocentric utilitarian instrumental relational intrinsic inherent absolute conditional responsibility care respect reverence wonder curiosity knowledge understanding wisdom science natural-philosophy biology zoology entomology coleopterology scarabaeology dynastinology systematics taxonomy nomenclature classification evolution ecology behavior paleontology conservation-biology environmental-science agricultural-science forestry-science veterinary-science medical-science public-health one-health planetary-health ecosystem-health biodiversity-health species-health population-health individual-health genetic-health environmental-health social-health economic-health political-health cultural-health spiritual-health holistic-health integrated-health sustainable-development sustainable-use sustainable-management adaptive-management precautionary-principle ecosystem-approach landscape-approach seascape-approach connectivity-conservation corridor buffer-zone protected-area national-park nature-reserve wildlife-refuge wilderness-area world-heritage-site biosphere-reserve Ramsar-site Important-Bird-Area Key-Biodiversity-Area Alliance-for-Zero-Extinction-site conservation-priority hotspot crisis-ecoregion global-200 last-of-the-wild human-footprint cumulative-impact threat-index vulnerability-index adaptive-capacity exposure sensitivity resilience resistance recovery restoration rehabilitation reintroduction translocation ex-situ in-situ captive-breeding botanic-garden zoo gene-bank seed-bank tissue-bank sperm-bank oocyte-bank embryo-bank DNA-bank frozen-zoo ark insurance safety-net de-extinction genetic-rescue genetic-restoration genetic-augmentation genetic-management population-management metapopulation source-sink patch matrix landscape seascape ecosystem biome ecoregion province zone region district site locality habitat microhabitat niche ecological-niche fundamental-niche realized-niche trophic-niche spatial-niche temporal-niche biotic-niche abiotic-niche multidimensional-niche n-dimensional-niche Hutchinsonian-niche Grinnellian-niche Eltonian-niche resource requirement limitation stress disturbance perturbation fluctuation variability heterogeneity complexity diversity redundancy stability persistence adaptation acclimation plasticity evolvability heritability selection drift flow mutation recombination speciation coalescence divergence convergence parallelism homoplasy analogy homology synapomorphy symplesiomorphy autapomorphy apomorphy plesiomorphy derived ancestral primitive advanced basal crown stem node branch clade grade subfamily family superfamily infraorder suborder order superorder infraclass subclass class superclass subphylum phylum superphylum kingdom domain life organism individual etc.

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