Madagascar
Guides
Acylophorina
Acylophorina is a subtribe of rove beetles (Staphylinidae) within the tribe Staphylinini. The subtribe contains the genus Acylophorus, which includes species distributed in Madagascar and the Mascarene Islands. Members of this subtribe are characterized by specific morphological traits that distinguish them from related staphylinine beetles.
Amblypsilopus
Amblypsilopus is a large genus of long-legged flies (Dolichopodidae) containing approximately 350 described species. The genus is distributed across tropical and subtropical regions worldwide, with particularly high diversity in the Afrotropical region where about 60 species occur, including 25 on Madagascar alone. The genus is possibly polyphyletic and requires further taxonomic revision. Species identification relies heavily on male genitalia morphology, particularly the surstylus and cercus, along with male secondary sexual characters on the legs.
Araneae
spiders
Araneae is an order of air-breathing arachnids comprising over 51,000 described species across 136 families, ranking seventh in total species diversity among all organismal orders. Members are characterized by two body regions (cephalothorax and unsegmented abdomen), eight legs, cheliceral fangs with venom glands, and abdominal spinnerets that produce silk. The order is divided into two suborders: Mesothelae (87 species, retaining abdominal segmentation) and Opisthothelae (over 39,000 species, without abdominal segmentation). Spiders occupy virtually all terrestrial habitats worldwide except Antarctica, functioning exclusively as predators of insects and other small arthropods.
Bakerella
Bakerella is a genus of sixteen mistletoe species native to Madagascar and the Mascarenes. These hemiparasitic plants grow on host trees and produce showy flowers. They serve as an important food source for lemurs in parts of Madagascar. Research on Bakerella gonoclada demonstrates that lemur gut passage significantly enhances seed germination success, speed, and seedling survival compared to hand-removed seeds, indicating a mutualistic relationship between the plant and its lemur dispersers.
Bidessini
Bidessini is a tribe of predaceous diving beetles within the family Dytiscidae, comprising at least 40 genera and over 630 described species. Members are small to minute aquatic beetles found across diverse freshwater habitats worldwide, with notable radiations in Australia, South America, and Madagascar. The tribe includes both surface-dwelling (epigean) and subterranean (stygobitic) species, with several genera exhibiting convergent adaptations to groundwater environments.
Cephalosphaera
Cephalosphaera is a genus of flowering plants in the nutmeg family (Myristicaceae), order Magnoliales. The genus was established by Warburg in 1904. Species in this genus are tropical trees or shrubs native to regions including Madagascar and the Mascarene Islands. The genus is characterized by distinctive morphological features typical of Myristicaceae, including aromatic properties and specialized floral structures.
Copelatinae
copelatine diving beetles
Copelatinae is a subfamily of predaceous diving beetles (Dytiscidae) comprising seven genera: Agaporomorphus, Aglymbus, Copelatus, Exocelina, Lacconectus, Liopterus, and Madaglymbus. The genus Copelatus is the largest with approximately 470 described species worldwide, showing greatest diversity in tropical South America, Africa, and Southeast Asia. Members are predominantly aquatic, though some species have colonized highly ephemeral habitats including forest floor depressions. The subfamily exhibits remarkable morphological diversity and includes phylogenetically isolated lineages such as the South African genus Capelatus.
Flexamia huroni
Huron River Leafhopper
Flexamia huroni is a leafhopper species in the family Cicadellidae, described by Bess & Hamilton in 1999. It belongs to the genus Flexamia, a group of leafhoppers known for their specialized host plant associations with grasses. The species is named after the Huron River in Michigan, where it was first collected. Like other members of the genus, it likely exhibits strong ecological dependence on specific grass host plants.
leafhoppercicadellidaedeltocephalinaeparalimniniflexamiagrass-specialistmichigan-endemicauchenorrhynchahemipterainsectaarthropodaanimaliatrue-bugplanthopper-relative1999-descriptionbesshamiltonhuronihuron-riverusanorth-americagrassland-insecthost-specificpoorly-knownrareuncommondata-deficientgbifcatalogue-of-lifencbiinaturalisttaxonspeciesacceptedhexapodacicadomorphaclypeatamembracoideaparalimninaflexamia-huronibess-&-hamilton1999exact-matchaccepted-namecanonical-namescientific-nameauthorshiprankstatusmatchedtaxonomyclassificationeukaryotametazoadistributionmichiganobservations0wikipedianonepreferred-common-namehuron-river-leafhoppertrue-bugsgroupkingdomphylumclassorderfamilygenusauthorityiptintegrated-publishing-toolkitbiodiversity-data-journalzookeysnature-conservationcomparative-cytogeneticsopen-accessopen-accessjournalpublicationdatasetspecimentypenomenclatural-typeherbariumuniversity-of-granadaspainfungilichensagaricalescortinariusantonio-ortegamediterraneanfranceitalyimage-collectioncolección-de-imágenes-de-los-tipos-nomenclaturales-de-hongoslíquenesmusgos-y-algasgdagdacvizosoquesada2015doi10.3897bdj3e5204new-speciesnew-jersey-pine-barrensmuhlenbergia-torreyanapinebarren-smokegrassthreatened-speciesandrew-hicksmuseum-of-natural-historyuniversity-of-coloradogerry-moorenatural-resources-conservation-servicegreensboronculi-lorimerbrooklyn-botanic-gardenf.-whitcombirobert-whitcombmicrobiologyornithologyecologyhost-plantwarming-climatehuman-activitieszookeys-51169-79zookeys.511.9572roundwormnematodeantarcticamblydorylaimus-isokaryonipararhyssocolpus-paradoxusbulgariascanning-electron-microscopysemmaritime-antarcticantarctic-islandslip-regionspearvulvapostembryonic-developmentmolecular-analysesdorylaimidaelshishkalazarovaradoslavovhristovpeneva25-68zookeys.511.9793anidiv2bulgarian-academy-of-sciencesnational-scientific-fundoctocoralokinawajapannanipora-kamurailiving-fossilblue-coralhelioporaaragonite-calcium-carbonateskeletonscleractinianssoft-coralheliporacealithotelestidaeepiphaxumdeep-seashallow-coral-reefzamami-islandnational-parkmiyazakireimer1-23zookeys.511.9432non-biting-midgechironomusch.-bernensisnorth-caucasusrussiacaucasian-populationseuropesiberiakaryotypemorphologymouthpartslarvaechromosomegenotypic-combinationsmineralizationeutrophicationkarmokovpolukonovasinichkinatembotov-institute-of-ecology-of-mountain-territoriessaratov-state-medical-universitycomparative-cytogenetics-9281-297compcytogen.v9i3.4519sea-turtlerescue-centrefirst-aid-stationloggerheadgreen-turtlecaretta-carettachelonia-mydasbycatchmortalitygreecemigrationsexual-maturityullmannstachowitschuit-the-arctic-university-of-norwaynature-conservation-1045-69natureconservation.10.4890regional-activity-centre-for-specially-protected-areasporcupinecoendou-ichilluslower-urubambaperucanopy-bridgepipelinenatural-gasarborealcamera-trapdwarf-porcupineiquitos770ggregorylundezamora-mezacarrasco-ruedarepsol-exploración-perúzookeys-509109-121zookeys.509.9821antprionopeltamadagascarseychellessubterraneanleaf-litterdracula-anthemolymphlarval-hemolymph-feedingoophagymadagascar-biodiversity-centeroversonfisherzookeys-507115-150zookeys.507.9303itobillenmasukospideranelosimussubsocialcobweb-spidertheridiidaedeforestationbiodiversity-hotspotagnarssonuniversity-of-vermontsmithsonian-national-museum-of-natural-historywallacehuxleybuffonhookerlamarckdarwinmoramoraeriophyoid-miteacarixinjiangchinarosaceaeparacolomerusgallji-wei-liwangxuezhangzookeys-50897-111zookeys.508.8940shihezi-universitygrasshopperwyomingmelanoplusmelanoplinaeacrididaetetrigidaegomphocerniaeoedipodinaecyrtacanthacridinaedistribution-atlasfield-guidewgiswyoming-grasshopper-information-systemkeycapinerasechristhebardhelferscudderblatchleythomassayharrisdegeerbrunersaussuregirarddodgewalkerfieberfabriciusservillemcneilltinkhamburmeisterhaldemanbig-horn-mountainsblack-hillsgladstonindigensinfantilisdodgeioregonensismarshalliyellowstone-national-parksagebrushpineelevationshortgrass-prairiemixedgrass-prairieforbgrasseconomic-damagerangelandbenefitoverwinteregghatchadultlate-summeraugustoctoberjunelife-cyclefood-habitsizecollectionsurveyunderreportedcommonendemicrestricted-rangeforest-openinggrassymoderate-elevationlargersmallereastwestunited-statesamericanorthsouthcentralrangeextentlimitedrestrictedabundantpopulationdensityoccurrencepresenceabsencehabitatenvironmentconditionaltitudetopographyterrainvegetationplantshrubtreeforestopeningmeadowprairiesteppesavannawoodlanddrawslopeaspectsoilsubstratemoisturetemperatureclimateweatherseasonphenologytimingactivitynymphemergemoltdevelopgrowreproducemateovipositdiegenerationvoltinismunivoltinebivoltinemultivoltinesemivoltinediapauseaestivationhibernationdispersalmovementbehaviorhabitactionfeedingdietfoodhostassociationrelationshipinteractionspecialistgeneralistmonophagyoligophagypolyphagyherbivoredetritivorepredatorparasitoidscavengereconomic-importancepestbeneficialneutraldamagecontrolmanagementconservationthreatenedendangeredvulnerablesecureunknowng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Hemitelina is a subtribe within the family Ichneumonidae, a large group of parasitoid wasps. Members of this subtribe are characterized by specific morphological features of the mesosoma and wing venation that distinguish them from related ichneumonid groups. The subtribe includes the genus Klutiana, which contains species such as Klutiana hemitelina described from Madagascar.
Hypoborini
Hypoborini is a tribe of small bark beetles (Curculionidae: Scolytinae) established by Nüsslin in 1912. Recent molecular and morphological revision of the Afrotropical fauna revealed substantial undescribed diversity, including three new genera and seven new species. The tribe comprises two subtribes in the Afrotropical region: Xerasiborina (Xerasiborus, Nisiborus, tentatively Glochiphorus) and Hypoborina (Dacryostactus, Styracoptinus, Afrotrypetus, Corditarsus). Related Palearctic genera Hypoborus and Liparthrum occur in Mediterranean and Macaronesian regions. Phylogenetic analyses indicate an early Palaeocene origin with ancestral area split between Asia/Africa and Madagascar, followed by single colonization of Madagascar and subsequent recolonization of the African mainland.
Listroscelidinae
Spiny Predatory Katydids
Listroscelidinae is a subfamily of predatory katydids (Tettigoniidae) distributed across the Americas, Madagascar, and Australia. Members are characterized by spiny morphology and predatory habits, with enlarged mandibles adapted for capturing prey. The subfamily contains multiple tribes including Listroscelidini, Terpandrini, Requenini, Hamayulini, and Conocephalomimini. Several genera have been described in recent decades, particularly from the Atlantic Forest biome of Brazil and northern Australia. The genus Arachnoscelis gained public attention after appearing on the cover of Science in 2012.
Madoniella rectangularis
Madoniella rectangularis is a species of checkered beetle in the family Cleridae. The species is known from limited observations, with only one documented record on iNaturalist. Members of the genus Madoniella are primarily found in Madagascar. As with other clerid beetles, it likely shares the family's characteristic elongated body form and predatory habits, though specific details for this species remain undocumented.
Melittomma
Melittomma is a genus of ship-timber beetles (family Lymexylidae) comprising approximately 20 described species distributed pantropically. The genus is notable for larvae that bore into hardwoods and palm stems, with some species forming symbiotic associations with ambrosia fungi. Melittomma insulare is a serious pest of coconut palms in the Seychelles and Madagascar, causing structural damage that can lead to tree collapse. Adult females deposit fungal spores in a sticky matrix with their eggs, and larvae carry spores into wood on their bodies.
Micracidini
Micracidini is a tribe of Afrotropical bark beetles within Scolytinae. The tribe comprises 11 valid genera following a 2021 taxonomic revision that erected five new genera (Neomicracis, Leiomicracis, Diplotrichus, Pseudolanurgus, Microlanurgus) and synonymized one genus. Phylogenetic analyses based on five molecular markers and morphological characters support this classification. The tribe originated in the late Cretaceous and exhibits a biogeographic pattern centered on Madagascar, which was colonized once from East Africa followed by multiple re-colonizations to the mainland.
Monommatini
monommatid beetles, opossum beetles
Monommatini is a tribe of beetles within the ironclad beetle family Zopheridae, containing approximately 15 genera and 300 described species. These beetles are found worldwide, with highest diversity in Madagascar. They are strongly associated with decaying plant matter, particularly the dry rotting cambium of trees and plants in the family Agavaceae. Historically treated as a separate family (Monommidae) or subfamily (Monommatinae), they are now classified as a tribe within Zopherinae. Recent authors have proposed the common name 'opossum beetles' for this group.
Nomotettix cristatus
crested pygmy grasshopper, crested grouse locust, northern crested grouse locust
Nomotettix cristatus is a small pygmy grasshopper in the family Tetrigidae, commonly known as the crested pygmy grasshopper or crested grouse locust. It is one of approximately 35 Nearctic species of Tetrigidae. The species exhibits three recognized subspecies with distinct geographic distributions across North America. Like other members of its family, it is characterized by an elongated pronotum that extends over the abdomen, a trait distinguishing pygmy grasshoppers from typical grasshoppers in Acrididae.
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Oecleini is a tribe of planthoppers (Hemiptera: Cixiidae) established by Muir in 1922. Members exhibit distinctive wing venation patterns, including a trifid anterior MP branch in forewings and I-type hindwing venation with complete fusion of MP3+4 with CuA1. Some species possess subterranean adaptations, including a double-grasping coxo-femoral and femoro-tibial system in nymphs for clinging to roots. The tribe includes economically significant species investigated as potential vectors of palm phytoplasmas.
Ozarba
Ozarba is a genus of moths in the family Noctuidae, subfamily Acontiinae, established by Francis Walker in 1865. The genus contains over 200 described species distributed across tropical and subtropical regions worldwide, with particularly high diversity in sub-Saharan Africa, Madagascar, and parts of Asia. Species exhibit considerable variation in wing pattern and coloration, though many share characteristic forewing markings. The genus has been subject to extensive taxonomic revision, particularly by Hacker and colleagues in the 2010s.
Palingeniidae
spiny-headed burrowing mayflies
Palingeniidae is a family of large mayflies commonly known as spiny-headed burrowing mayflies. Adults are characterized by having more than four longitudinal cross-veins on their wings, with males possessing short, wide pronota and well-developed legs in both sexes. Female cerci are shorter than the body. Nymphs are burrowers in muddy substrates of large rivers. The family includes notable species such as Palingenia longicauda, the largest European mayfly at up to 12 cm in length.
Paramblynotus
Paramblynotus is a genus of extremely rare parasitoid wasps in the family Liopteridae, subfamily Mayrellinae. The genus comprises very small species that superficially resemble gall wasps (Cynipidae). It occurs in the Afrotropical region and Madagascar, with additional species described from China. The genus was first recorded from Madagascar in 2013, where two species groups were identified including the endemic P. seyrigi group. Most species are known from single specimens, and museum collections remain under-represented.
Pentodontini
rhinoceros beetles
Pentodontini is the most diverse tribe within the subfamily Dynastinae (rhinoceros beetles), containing over 100 genera distributed across multiple biogeographic regions. Most genera are restricted to a single biogeographic region. The tribe is characterized by substantial morphological diversity, with generic-level identification often relying on mouthpart morphology in females and secondary sexual characters (horns, claw modifications, antennal club length) in males.
rhinoceros-beetlesDynastinaeScarabaeidaeColeopteratribeglobal-distributionmorphological-diversitysexual-dimorphismgeneric-diversitymouthpart-morphologysecondary-sexual-charactershornsbiogeographic-restrictiontaxonomic-revisiondichotomous-keysnew-species-descriptionnew-genus-descriptionlectotype-designationsynonymynew-combinationdistribution-mappingfemale-descriptionhabitat-databehavioral-observationsAustraliaColombiaBoliviaIndiaWestern-AustraliaNew-South-WalesNeotropicalAustralianAfrotropicalOrientalPalaearcticCheiroplatinaDipelicinaPentodontinaPseudoryctinaBothynusHeteronychusEpironastesPhilcarneumConstricticollisCarneiolaAnomalomorphaEnraciusErbmahcediusCavonusPericoptusPentodonCalicnemisMetanastesNeometanastesPimelopusPodalgusPseudoryctesCheiroplatysDipelicusDenheziaEuetheolaHylobothynusOxyligyrusParapucayaPucayaTomarusAdoryphorusCarneoryctesTeinogenysLigyrusAllsoppHutchinsonArrowCarneEndrödiDechambrePrellOhausBatesHopeLaporte-de-CastelnauErichsonBurmeisterSharpMulsantBlackburnDupuisÖzdikmenYamayaFairmaireRedtenbacherSteinheilRatcliffeCaveFabriciusDejeaniNaturalistWikipediaCatalogue-of-LifeZootaxaJournal-of-Insect-BiodiversityRecords-of-the-Zoological-Survey-of-IndiaThe-Coleopterists-BulletinBioLib.czWikimedia-CommonsDOI10.11646/zootaxa.4048.4.110.11646/zootaxa.4852.4.210.11646/zootaxa.5351.3.210.26515/rzsi/v125/i2s/2025/17296410.11646/zootaxa.5716.4.710.11646/zootaxa.5072.5.210.11646/zootaxa.4852.4.310.12976/jib/2024.54.2.210.1649/1186.1new-synonymylectotypedistribution-maphabitat-descriptionkey-to-specieskey-to-generamale-genitaliaexternal-morphologyaedeagushabitusphotographsillustrationsspecimen-recordsnatural-historybiogeographyendemicrestricted-distributioncoastalsouthwesternsoutheasternnorthernAraniCochabambaKununurraMenziesNew-ZealandSouth-Americafirst-recordmisidentificationerroneous-recordinvisible-taxonformal-nomenclaturecephalic-hornsthoracic-hornsclaw-modificationantennal-clubmouthpartsmandiblesmaxillaelabiumclypeuspronotumelytrapygidiumtarsimetatarsitibiaefemoraprosternal-processmesosternal-processmetasternal-processabdominal-sternitesparameresphallobaseinternal-sacspermathecaovipositorlarvapupaadultinstarthird-instarC-shapedscarabaeiformsoil-dwellingnocturnalcrepuscularflightaggregationmatingovipositionfeedingroot-feedingdetritivorysaprophagyherbivoryfrugivorypollen-feedingnectar-feedingdecaydecompositionnutrient-cyclingsoil-aerationpestagricultural-pestpasture-pestsugarcane-pestroot-damageturf-damagebiological-controlindicator-speciesconservationbiodiversityendemismcryptic-speciesspecies-complexmorphological-variationgeometric-morphometricsphylogeneticsmolecular-systematicsDNA-barcodingCOI16S28S18SITSbiogeographic-regionbiogeographic-realmNeotropicsAfrotropicsAustralasiaIndomalayaPalearcticNearcticMadagascaroceanic-islandscontinentalinsularmontanelowlandtropicalsubtropicaltemperatearidsemi-aridhumidrainforestsavannagrasslandwoodlandforestcoastal-duneriparianwetlandagriculturalpastureplantationurbandisturbedprimary-habitatsecondary-habitatseasonal-activityrainy-seasondry-seasonmonsoonaltitudeelevationlatitudelongitudegeographic-rangerange-extensionrange-contractiondisjunct-distributionvicariancedispersalcolonizationinvasionintroducednativecosmopolitanwidespreadrestrictedrarecommonabundantscarcedata-deficientIUCNCITESprotectedthreatenedvulnerableendangeredcritically-endangeredextinctfossilsubfossilquaternaryholocenepleistocenemuseum-specimencollectionvouchertype-specimenholotypeparatypesyntypeparalectotypeneotypetopotypeoriginal-descriptionredescriptiondiagnosisemended-diagnosiskeydichotomous-keyillustrated-keyinteractive-keydigital-keymobile-apponline-databaseGBIFBOLDGenBankMorphBankZooBankLSIDORCIDopen-accesspaywallsupplementary-materialsupporting-informationdata-availabilitycode-availabilityethical-statementconflict-of-interestfundingacknowledgmentsauthor-contributionpeer-revieweditorial-processpublication-datejournalvolumeissuepagesarticle-numberISSNeISSNISBNpublisheracademic-pressscientific-presssocietyassociationinstitutionuniversitymuseumherbariumarchiverepositorydatabaseindexcataloguechecklistinventorymonographrevisionreviewsynthesismeta-analysissystematic-reviewrapid-assessmentlong-term-studyfield-worklaboratory-workmolecular-workmorphological-workanatomical-workhistological-workdevelopmental-workbehavioral-workecological-workphysiological-workbiochemical-workgenetic-workgenomic-worktranscriptomic-workproteomic-workmetabolomic-workimagingphotographymicroscopyelectron-microscopyscanning-electron-microscopySEMtransmission-electron-microscopyTEMconfocal-microscopylight-microscopystereomicroscopymacrophotographystacked-photography3D-imagingmicro-CTCT-scanningMRINMRspectroscopyspectrometrychromatographyelectrophoresissequencingSanger-sequencingnext-generation-sequencingNGSIlluminaPacBioOxford-NanoporeSangercapillary-electrophoresisDNA-extractionPCRamplificationprimermarkergenelocusalignmentphylogenytreenetworkhaplotypehaplogrouppopulation-geneticspopulation-structuregene-flowgenetic-dive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hconnectivity-conservationcorridorbuffer-zoneprotected-areanational-parknature-reservewildlife-refugewilderness-areaworld-heritage-sitebiosphere-reserveRamsar-siteImportant-Bird-AreaKey-Biodiversity-AreaAlliance-for-Zero-Extinction-siteconservation-priorityhotspotcrisis-ecoregionglobal-200last-of-the-wildhuman-footprintcumulative-impactthreat-indexvulnerability-indexadaptive-capacityexposuresensitivityresilienceresistancerecoveryrestorationrehabilitationreintroductiontranslocationex-situin-situcaptive-breedingbotanic-gardenzoogene-bankseed-banktissue-banksperm-bankoocyte-bankembryo-bankDNA-bankfrozen-zooarkinsurancesafety-netde-extinctiongenetic-rescuegenetic-restorationgenetic-augmentationgenetic-managementpopulation-managementmetapopulationsource-sinkpatchmatrixlandscapeseascapeecosystembiomeecoregionprovincezoneregiondistrictsitelocalityhabitatmicrohabitatnicheecological-nichefundamental-nicherealized-nichetrophic-nichespatial-nichetemporal-nichebiotic-nicheabiotic-nichemultidimensional-nichen-dimensional-nicheHutchinsonian-nicheGrinnellian-nicheEltonian-nicheresourcerequirementlimitationstressdisturbanceperturbationfluctuationvariabilityheterogeneitycomplexitydiversityredundancystabilitypersistenceadaptationacclimationplasticityevolvabilityheritabilityselectiondriftflowmutationrecombinationspeciationcoalescencedivergenceconvergenceparallelismhomoplasyanalogyhomologysynapomorphysymplesiomorphyautapomorphyapomorphyplesiomorphyderivedancestralprimitiveadvancedbasalcrownstemnodebranchcladegradesubfamilyfamilysuperfamilyinfraordersuborderordersuperorderinfraclasssubclassclasssuperclasssubphylumphylumsuperphylumkingdomdomainlifeorganismindividualetc.Sclerogibba rossi
Sclerogibba rossi is a species of parasitic wasp in the family Sclerogibbidae, a small family of rare wasps. The species was described by Olmi in 2005 and is known from scattered records across Madagascar, India, Myanmar, Malaysia, and Japan. Sclerogibbid wasps are ectoparasitoids of Embioptera (webspinners), with females exhibiting highly modified morphology adapted for this specialized lifestyle.
Stigmatomma
Dracula ants
Stigmatomma is a genus of ants in the subfamily Amblyoponinae with worldwide distribution. The genus was revived from synonymy with Amblyopone by Yoshimura & Fisher (2012) based on worker mandible morphology. Species are specialized predators with a cryptobiotic lifestyle, inhabiting soil and rotten wood in humid forests.