Endangered

Guides

Agonum belleri
Beller's Ground Beetle
Agonum belleri is a flightless ground beetle endemic to Pacific Northwest wetlands. It is metallic-black with copper, blue, or green reflections. The species is restricted to sphagnum bogs and associated wetlands in Oregon, Washington, and British Columbia. It is listed as endangered by the Xerces Society and a species of greatest conservation need in Washington due to habitat loss from peat mining and development.
endangered flightless wetland-specialist sphagnum-bog Pacific-Northwest-endemic conservation-concern
Anaea troglodyta floridalis
Florida Leafwing
Anaea troglodyta floridalis is a federally endangered subspecies of butterfly endemic to southern Florida. It is one of the most imperiled butterflies in North America, with populations restricted to pine rockland and tropical hardwood hammock habitats in Miami-Dade County and Everglades National Park. The subspecies is highly dependent on its host plant Croton linearis for larval development. Habitat loss, fragmentation, and fire suppression have contributed to its decline, with remaining populations concentrated in protected areas.
endangered endemic pine-rockland hardwood-hammock host-specialist conservation Florida Miami-Dade-County Everglades-National-Park leafwing-butterfly Croton-linearis dead-leaf-mimicry
Analetris
Analetris is a genus of sand-dwelling mayflies in the family Acanthametropodidae, established by Edmunds in 1972. The genus contains at least two species: the extant Analetris eximia and the extinct †Analetris secundus. Members of this genus are specialized inhabitants of riverine environments with unstable silty substrates. The best-studied species, A. eximia, is considered endangered due to habitat destruction from reservoir construction.
mayflies sand-dwelling riverine endangered North-America fossil carnivorous-nymphs chironomid-predator reservoir-impact unstable-substrate-specialist
Apodrepanulatrix liberaria
New Jersey tea inchworm, Barrens Carpet
A small geometrid moth with a wingspan of 25–35 mm, found in eastern North America. The species is notable for its specialized larval diet on New Jersey tea (Ceanothus americanus) and its endangered status in Massachusetts and Connecticut. Adults are active during the growing season, with larvae feeding on a single host plant genus.
Geometridae specialist-herbivore endangered North-America barrens-habitat
Atrytone arogos
Arogos Skipper, Beard-grass Skipper
Atrytone arogos, commonly known as the Arogos Skipper or Beard-grass Skipper, is an endangered butterfly in the family Hesperiidae. The species consists of two subspecies: the nominate A. a. arogos and A. a. iowa. It is native to grassland ecosystems of the eastern and midwestern United States, where it has experienced significant population declines due to habitat destruction and poor adaptability to environmental change.
endangered prairie-specialist grassland-dependent
Bagoinae
Bagoinae is a subfamily of weevils (Curculionidae) containing the genus Bagous and related genera. Members are primarily associated with aquatic and semi-aquatic habitats, though some occupy terrestrial environments. The group has been variously classified, with some treatments placing it within Molytinae. The subfamily includes approximately 130 species in the Palaearctic region, with the genus Bagous distributed globally except Central and South America.
weevil aquatic wetland biological-control endangered
Batrisodes venyivi
Helotes Mold Beetle
Batrisodes venyivi, commonly known as the Helotes Mold Beetle, is a small, eyeless troglobitic beetle in the family Staphylinidae. It is endemic to eight caves in Bexar County, Texas, and was listed as endangered under the U.S. Endangered Species Act in 2000. The species exhibits classic cave-adapted traits including elongated antennae, legs, and sensory setae. Very little is known about its behavior, population trends, or life history due to the logistical challenges of accessing its subterranean habitat.
troglobite cave-endemic endangered karst-invertebrate Bexar-County eyeless Staphylinidae Pselaphinae Batrisodes Texas-endemic ESA-listed
Blattellinae
wood cockroaches
Blattellinae is a subfamily of Ectobiidae containing approximately 70 genera. It includes the globally distributed German cockroach (Blattella germanica), a major household pest, as well as several endangered species such as those in Hololeptoblatta and Miriamrothschildia. The subfamily exhibits diverse ecological adaptations, with members occupying habitats ranging from wetlands and grasslands to caves and human dwellings.
cave-dwelling synanthropic wetland endangered pest
Blissus occiduus
Western Chinch Bug
Blissus occiduus, the western chinch bug, is a phloem-feeding true bug (Hemiptera: Blissidae) that is a significant pest of warm-season turfgrasses, particularly buffalograss and zoysiagrass. The species exhibits strong host preference hierarchies, with buffalograss being the most preferred host followed by zoysiagrass, though it can survive and reproduce on a broad range of grasses including agronomic crops. Field studies have documented inconsistent control with neonicotinoid insecticides, with thiamethoxam showing particularly rapid degradation in buffalograss tissues compared to imidacloprid and clothianidin.
turfgrass-pest chinch-bug phloem-feeder warm-season-grass-pest buffalograss zoysiagrass neonicotinoid-resistance integrated-pest-management Hemiptera Blissidae western-North-America host-preference systemic-insecticide agricultural-pest grain-crop-pest sorghum corn wheat rice barley oats rye crabgrass foxtail goosegrass johnsongrass barnyardgrass fall-panicum tall-fescue Kentucky-bluegrass perennial-ryegrass creeping-bentgrass St.-Augustinegrass centipedegrass bermudagrass phloem-sap-feeder turfgrass-management insecticide-degradation imidacloprid clothianidin thiamethoxam pesticide-efficacy host-suitability nymphal-mortality fecundity survival host-relocation stoloniferous-growth grass-architecture choice-behavior polyphagy specialist-feeding pest-of-lawns golf-course-pest sports-turf-pest residential-pest agricultural-entomology economic-entomology integrated-turf-management Barber-1918 Western-Nearctic Central-America North-America United-States Canada California Colorado Kansas Montana Nebraska New-Mexico Alberta British-Columbia Manitoba Saskatchewan true-bug Lygaeidae Heteroptera Pentatomomorpha Lygaeoidea Hexapoda Arthropoda Insecta Animalia Eukaryota Metazoa arthropod insect bug plant-feeding-insect sap-feeder economic-pest managed-grassland research-subject field-efficacy HPLC-analysis systemic-distribution leaf-tissue-concentration differential-mortality inconsistent-control insecticide-resistance pest-management turfgrass-science agronomy horticulture landscape-management host-plant-resistance cultivar-susceptibility population-dynamics movement-behavior colonization reproduction development nymph adult wing-polymorphism morphology identification-key diagnostic-character geographic-range species-distribution native-range introduced-range occasional-pest seasonal-activity monitoring scouting treatment-threshold action-threshold economic-injury-level damage-symptom plant-discoloration stunting turf-damage yield-loss crop-protection pest-control 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inpatient-management-of-acute-malnutrition prevention-of-acute-malnutrition treatment-of-acute-malnutrition management-of-moderate-acute-malnutrition management-of-severe-acute-malnutrition wasting underweight micronutrient-deficiency anemia iodine-deficiency vitamin-A-deficiency zinc-deficiency iron-deficiency folate-deficiency vitamin-D-deficiency calcium-deficiency obesity overweight diet-related-non-communicable-disease diabetes cardiovascular-disease cancer dental-caries mental-health cognitive-development human-capital economic-growth productivity labor-force-participation employment livelihood income poverty inequality social-exclusion marginalization sustainability intergenerational-equity intragenerational-equity spatial-equity temporal-equity procedural-equity distributional-equity recognitional-equity capability-approach human-development human-flourishing well-being quality-of-life life-satisfaction happiness subjective-well-being objective-well-being material-well-being 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tactical operational foundational structural functional procedural behavioral attitudinal cultural social economic political environmental technological institutional organizational individual collective community societal global local national regional landscape seascape waterscape airscape biosphere noosphere technosphere sociosphere ecosphere hydrosphere atmosphere lithosphere pedosphere cryosphere anthroposphere urban-sphere rural-sphere agricultural-sphere natural-sphere managed-sphere protected-sphere restored-sphere degraded-sphere transformed-sphere connected-sphere disconnected-sphere integrated-sphere fragmented-sphere resilient-sphere vulnerable-sphere adaptive-sphere maladaptive-sphere sustainable-sphere unsustainable-sphere equitable-sphere inequitable-sphere just-sphere unjust-sphere inclusive-sphere exclusive-sphere diverse-sphere homogeneous-sphere complex-sphere simple-sphere dynamic-sphere static-sphere stable-sphere unstable-sphere predictable-sphere unpredictable-sphere controllable-sphere uncontrollable-sphere manageable-sphere unmanageable-sphere governable-sphere ungovernable-sphere navigable-sphere unnavigable-sphere traversable-sphere untraversable-sphere permeable-sphere impermeable-sphere porous-sphere non-porous-sphere transparent-sphere opaque-sphere visible-sphere invisible-sphere tangible-sphere intangible-sphere material-sphere immaterial-sphere physical-sphere metaphysical-sphere empirical-sphere theoretical-sphere practical-sphere applied-sphere pure-sphere basic-sphere strategic-sphere tactical-sphere operational-sphere foundational-sphere superstructural-sphere infrastructural-sphere structural-sphere functional-sphere procedural-sphere behavioral-sphere attitudinal-sphere cultural-sphere social-sphere economic-sphere political-sphere environmental-sphere technological-sphere institutional-sphere organizational-sphere individual-sphere collective-sphere community-sphere societal-sphere global-sphere local-sphere 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airscape-sphere-sphere-sphere biosphere-sphere-sphere-sphere noosphere-sphere-sphere-sphere technosphere-sphere-sphere-sphere sociosphere-sphere-sphere-sphere ecosphere-sphere-sphere-sphere hydrosphere-sphere-sphere-sphere atmosphere-sphere-sphere-sphere lithosphere-sphere-sphere-sphere pedosphere-sphere-sphere-sphere cryosphere-sphere-sphere-sphere anthroposphere-sphere-sphere-sphere
Bombus crotchii
Crotch's bumble bee, Golden State bumble bee, Crotch's bumblebee
Bombus crotchii, commonly called Crotch's bumble bee or the Golden State bumble bee, is a short- to medium-tongue bumble bee endemic to California and adjacent regions. The species is classified as endangered under the California Endangered Species Act and listed as imperiled globally by the IUCN. It has experienced severe population declines attributed to habitat loss, pesticide exposure, climate change, and inbreeding depression. The species is a dietary generalist and important pollinator of native plants and agricultural crops.
bumble-bee endangered pollinator California-endemic conservation-priority buzz-pollination eusocial
Bombus distinguendus
Great Yellow Bumble Bee, great yellow bumblebee
Bombus distinguendus, the great yellow bumblebee, is a bumblebee species with a Palearctic distribution spanning northern and central Europe, northern Russia, and Alaska. The species has experienced significant population declines across much of its European range, with critically low numbers or local extinction in several regions. In Ireland, it is classified as Endangered and restricted to a single remaining population in the northwest. The species exhibits late-season phenology compared to other bumblebees and shows strong foraging preferences for specific plant species, particularly Common Knapweed (Centaurea nigra) in Irish populations.
bumblebee pollinator conservation endangered grassland late-emerging
Bombus franklini
Franklin's bumble bee, Franklin's bumblebee
Bombus franklini is a critically endangered bumble bee species with the most restricted range of any bumble bee in the world, confined to a 13,300-square-mile area spanning five counties in southern Oregon and northern California. The species has not been observed since 2006, despite intensive monitoring by entomologist Robbin Thorp from 1998 until his death in 2019. Listed as endangered under the U.S. Endangered Species Act in 2021, it is feared extinct due to severe population decline from an estimated 94 individuals in 1998 to zero confirmed sightings since 2006. Genomic research indicates long-term population decline driven by bottlenecks and environmental stressors rather than pathogens alone.
endangered flagship-species pollinator-decline Siskiyou-Cascade possibly-extinct ESA-listed
Bombus fraternus
Southern Plains bumblebee
Bombus fraternus, commonly called the Southern Plains bumblebee, is a large bumblebee species native to the United States east of the Rocky Mountains. It is classified in the subgenus Cullumanobombus and is currently listed as endangered, with estimated abundance below 15% of historical numbers. The species is distinguished by its mostly black coloration with yellow thoracic and abdominal bands and unusually flattened abdominal hairs.
bumblebee pollinator endangered prairie grassland underground-nester
Bombus occidentalis
Western bumble bee, white-bottomed bee
Bombus occidentalis, the western bumble bee, is a medium to large bumble bee species native to western North America. Once among the most common bumble bees in its range, it has experienced severe population declines since the late 1990s, with losses exceeding 40% in some areas and near-disappearance from parts of California, Oregon, and Washington. The species was listed as endangered under the California Endangered Species Act in 2019. It is a generalist forager and effective pollinator of crops including tomatoes, peppers, and berries through buzz pollination.
bumble-bee pollinator endangered western-North-America buzz-pollination eusocial commercial-pollination population-decline
Bombus suckleyi
Suckley's Cuckoo Bumble Bee, Suckley's Bumble Bee
Bombus suckleyi is a cuckoo bumble bee (subgenus Psithyrus) and obligate social parasite of other Bombus species. It lacks corbiculae (pollen baskets) on its hind legs, having lost the ability to collect pollen and nectar for its own brood. The species invades host colonies, kills or subdues the resident queen, and manipulates host workers to provision its offspring. Native to northwestern North America, it has experienced severe population declines of approximately 77% and is classified as critically endangered by the IUCN and NatureServe.
bumble-bee cuckoo-bee social-parasite pollinator endangered conservation Psithyrus North-America meadow obligate-parasite
Brychius
crawling water beetles
Brychius is a genus of small aquatic beetles in the family Haliplidae, commonly known as crawling water beetles. The genus comprises five recognized species distributed across parts of Europe and North America. One species, Brychius hungerfordi, is federally endangered in the United States. Members of this genus inhabit freshwater environments and are associated with aquatic vegetation.
aquatic freshwater Haliplidae crawling-water-beetle endangered conservation Coleoptera Adephaga
Calephelis borealis
Northern Metalmark
Calephelis borealis, the northern metalmark, is a small butterfly in the family Riodinidae endemic to the eastern United States. It is one of few tropical-associated riodinids found in eastern North America. The species has declined across its range due to habitat loss from afforestation and invasive plant encroachment. Active management including canopy thinning and invasive plant removal has been shown to increase local populations.
endangered conservation-dependent limestone-barren-specialist riodinidae eastern-north-america single-brooded metapopulation habitat-management-responsive
Calephelis muticum
Swamp Metalmark
Calephelis muticum, known as the swamp metalmark, is a small riodinid butterfly with a restricted distribution across wetlands of the upper Midwest, Ohio Valley, Ozark Mountains, and parts of Arizona. The species is globally vulnerable (G3) and endangered in Illinois, with small, isolated colonies scattered across ten states. It is distinguished from the northern metalmark (Calephelis borealis) by habitat preference, wing morphology, and host plant associations. Conservation concerns center on habitat loss from wetland conversion, invasive species, pesticides, and fire suppression.
butterfly riodinidae metalmark wetland endangered thistle north-america
Callophrys hesseli
Hessel's Hairstreak
Hessel's Hairstreak is a small lycaenid butterfly endemic to Atlantic coastal wetlands of the eastern United States. The species exhibits complete ecological dependence on Atlantic white-cedar (Chamaecyparis thyoides), which serves as the exclusive larval host plant and provides adult camouflage habitat. Populations are patchily distributed and declining due to habitat loss, with state-level endangered status in Connecticut and G3 vulnerable ranking globally.
butterfly lycaenid hairstreak wetland-specialist endangered Atlantic-white-cedar host-specific diapause lek-mating
Callophrys mossii bayensis
San Bruno Elfin
The San Bruno Elfin is a federally endangered butterfly subspecies endemic to the San Francisco Peninsula in California. It inhabits rocky outcrops and cliffs in coastal scrub habitat, with its distribution directly tied to that of its host plant, broadleaf stonecrop (Sedum spathulifolium). The subspecies was described by R. Brown in 1969 and is currently listed under the U.S. Endangered Species Act.
endangered endemic California hairstreak specialist
Canthyloscelidae
Canthyloscelidae is a small family of nematoceran flies closely related to Scatopsidae. The family contains approximately 15 extant species distributed across New Zealand, North America, South America, Japan, and Russia, plus one species known from Jurassic fossils. Most species are considered endangered due to their specialized habitat requirements.
Diptera Nematocera ancient-woodland xylosaprophagous endangered decaying-wood
Catocala amestris
Three-staff Underwing
Catocala amestris, the three-staff underwing, is a North American moth in the family Erebidae. It is listed as endangered and receives legal protection in Michigan. The species has a wingspan of 4-4.5 cm and exhibits the characteristic underwing pattern of cryptic forewings concealing brightly colored hindwings. It has been documented as a new record for southern Colorado when attracted to bait stations, indicating potential range expansion or underreporting.
underwing-moth endangered specialist-herbivore North-America bait-attracted
Chytonix sensilis
barrens chytonix
Chytonix sensilis, commonly known as the barrens chytonix, is a small noctuid moth native to eastern North America. First described in 1881, it is listed as endangered in Connecticut. The species is notable for its specialized larval diet of fungi growing on leaf litter and dead wood.
moth Noctuidae fungivore endangered North-America leaf-litter dead-wood
Cicindela albissima
Coral Pink Sand Dunes tiger beetle
Cicindela albissima is a highly endemic tiger beetle restricted to the Coral Pink Sand Dunes in southern Utah. Adults are distinguished by nearly pure white elytra with reduced pigmentation, contrasting with a bronze to brown thorax. The species exhibits a bimodal adult activity pattern with peak abundance in mid-April to early May, followed by a smaller secondary emergence in late August to October. Originally described as a subspecies of C. limbata, molecular and morphological evidence elevated it to full species status. The entire global population occupies approximately 400 hectares, making it one of North America's most geographically restricted tiger beetles.
endemic endangered sand-dune-specialist spring-fall-activity white-coloration Utah conservation-priority
Cicindela ohlone
Ohlone Tiger Beetle
Cicindela ohlone is an endangered tiger beetle endemic to Santa Cruz County, California, discovered in 1987 and described in 1993. It is most closely related to C. purpurea but differs in larger body size, distinct genital morphology, and a unique late winter-spring activity period. The species occupies a restricted 24 km² range with five remaining subpopulations, having declined from 10-15 historical patches. Listed under the U.S. Endangered Species Act in 2001, it faces ongoing threats from habitat destruction, invasive vegetation, and recreational disturbance.
endangered endemic tiger-beetle California coastal-prairie conservation Cicindelidae
Cicindela patruela
Northern Barrens Tiger Beetle
Cicindela patruela, the Northern Barrens Tiger Beetle, is a medium-sized tiger beetle endemic to eastern North America with highly specific habitat requirements. Adults are metallic green with ivory elytral bands and measure 12–15 mm in length. The species has a two-year life cycle with adults active primarily in spring and fall. Populations have declined significantly due to habitat loss and fire suppression, leading to endangered status in Canada and vulnerable ranking by NatureServe.
tiger-beetle endangered fire-dependent barrens-specialist ground-beetle visual-predator
Cicindela patruela patruela
Northern Barrens Tiger Beetle
Cicindela patruela patruela is a globally rare and endangered subspecies of tiger beetle found in barrens habitats of eastern North America. The subspecies has been documented in limited populations in Canada and the United States, with particular study focus on Île-aux-Allumettes in Quebec. Research indicates small population sizes are typical for this subspecies, with individuals exhibiting moderate daily movement distances. The subspecies exhibits thermoregulatory behaviors and seasonal activity patterns that have been subjects of dedicated study.
Cicindelidae Carabidae tiger-beetle endangered barrens thermoregulation mark-recapture
Cicindelidia floridana
Miami Tiger Beetle
Cicindelidia floridana, the Miami Tiger Beetle, is a critically endangered species endemic to pine rockland habitat in Miami-Dade County, Florida. Rediscovered in 2007 after being presumed extinct since the 1940s, it is one of the rarest tiger beetles in North America. Adults are diurnal predators with brilliant coppery-green coloration, reduced elytral maculation, and distinctly punctured elytra with dense lateral pronotal setae. The species shows extreme habitat specificity to open sandy patches within fire-maintained pine rocklands, making it exceptionally vulnerable to urban development.
endangered endemic Florida pine-rockland rediscovered tiger-beetle conservation
Cicindelidia highlandensis
Highlands Tiger Beetle
Cicindelidia highlandensis is one of Florida's rarest endemic tiger beetles, restricted to remnant sand scrub and pine woodland habitats on the Lake Wales Ridge in central Florida. The species has experienced severe population declines due to habitat loss, with an estimated 90% reduction and only a few sites capable of sustaining viable populations. Adults are active during summer months, displaying characteristic thermoregulatory "stilting" behavior and extreme wariness when approached. Despite its critically imperiled global status (G1), it remains unlisted by the State of Florida.
endemic endangered Florida tiger-beetle Lake-Wales-Ridge sand-scrub fire-dependent-ecosystem
Cicindelidia marginipennis
Cobblestone Tiger Beetle
Cicindelidia marginipennis is a North American riparian specialist tiger beetle restricted to sandy cobblestone banks and bars along rivers. The species has a highly disjunct distribution spanning from southern Alabama and Mississippi through a northern corridor to New Brunswick, Canada, with a significant gap across the southeastern United States. It is state-listed as threatened or endangered throughout its U.S. range and has been periodically considered for federal listing since 1984 due to habitat decline from dam construction, channelization, and development. The species exhibits potential geographic population structure based on mitochondrial DNA, with Alabama populations possessing unique haplotypes that may represent a cryptic species.
riparian cobblestone tiger-beetle endangered indicator-species riverine metapopulation conservation laboratory-rearing cryptic-species Alabama-haplotype throw-pile mate-guarding visual-predator ice-scour seasonal-flooding
Cicurina
cave meshweaver
Cicurina is a genus of small spiders commonly known as cave meshweavers, first described by Anton Menge in 1871. The genus has undergone multiple family reassignments, most recently placed in the family Cicurinidae in 2023. Species exhibit considerable size variation, with some measuring under 2 mm and others exceeding 13 mm. Many species are specialized cave dwellers (troglobites), including several federally endangered species in the United States.
spider cave-dwelling troglobite subnivium endangered North-America Texas China conservation taxonomy
Cicurina madla
Madla Cave meshweaver, Madla's Cave Meshweaver
Cicurina madla is a federally endangered troglobitic spider endemic to karst caves in Bexar County, Texas. First described by Gertsch in 1992, it was synonymized with Cicurina venii in 2018. The species is completely eyeless and exhibits morphological adaptations to cave life, including elongated legs relative to body size. It is known from only eight to nine caves and depends on cave crickets as a food source.
troglobite endangered karst cave-spider Bexar-County Edwards-Plateau eyeless Cicurinidae
Coranarta luteola
Small Dark Yellow Underwing
Coranarta luteola is a small noctuid moth native to boreal North America. First described in 1865, it occurs from Alaska and Labrador south through the Rocky Mountains to Colorado, and eastward to Maine. The species has a forewing length of 11–12 mm. Its larvae feed on laurels (Kalmia species), while adults visit flowers, particularly Andromeda polifolia. The species is listed as endangered in Connecticut.
Noctuidae underwing-moth boreal laurel-feeder endangered Connecticut Kalmia ericaceous-host small-moth North-America pollinator wetland conservation-concern 1865-description Grote Robinson
Danaus plexippus
Monarch, Monarch butterfly, Milkweed, Common tiger, Wanderer, Black veined brown
The monarch butterfly is a large, migratory milkweed butterfly with distinctive orange, black, and white wing patterns. It is best known for its spectacular multigenerational migration between overwintering sites in central Mexico and coastal California and breeding grounds across North America. The species has declined dramatically in recent decades, with the western population declining 99.9% and the eastern population declining 84% since the 1990s. It was listed as Endangered by the IUCN in 2022, though it remains a candidate for listing under the U.S. Endangered Species Act. Conservation efforts focus on protecting milkweed host plants and overwintering habitat.
migratory endangered pollinator milkweed-dependent flagship-species multigenerational-migration chemical-defense citizen-science
Elaphrus
Marsh Ground Beetles
Elaphrus is a genus of ground beetles (Carabidae: Elaphrinae) commonly known as marsh ground beetles. The genus contains at least 40 described species distributed across the Palearctic, Nearctic, Near East, and Northern Africa. Many species are strongly associated with wetland habitats, including vernal pools and marshes. Several species exhibit strong morphological resemblance to tiger beetles (Cicindelinae), frequently leading to misidentification by collectors. At least two species, E. viridis and E. sugai, are considered threatened or endangered due to habitat loss and degradation.
Carabidae Elaphrinae wetland marsh vernal-pool predator conservation endangered misidentification tiger-beetle-mimicry
Ellipsoptera nevadica lincolniana
Salt Creek Tiger Beetle
Ellipsoptera nevadica lincolniana is a critically endangered subspecies of tiger beetle endemic to saline wetlands in Nebraska, USA. Populations remain critically low despite ongoing conservation efforts. The egg stage in June is particularly vulnerable to climatic conditions, with higher temperatures and lower precipitation negatively impacting population growth through reduced oviposition and egg survival.
endangered saline-wetland Nebraska-endemic climate-sensitive conservation-dependent
Epeoloides pilosulus
Macropis Cuckoo Bee
Epeoloides pilosulus is one of the rarest bees in North America, an obligate kleptoparasite of oil-collecting Macropis bees (Melittidae). It belongs to the tribe Osirini, a group of parasitic bees entirely dependent on oil-collecting bees as hosts. The species was thought extinct from the 1950s until its rediscovery in Nova Scotia in 2004, with subsequent records from Alberta, Ontario, Connecticut, Maine, New Hampshire, and Michigan extending its known range. Its survival depends on a three-way ecological relationship: E. pilosulus requires Macropis host bees, which in turn require oil-producing Lysimachia flowers for larval provisioning.
kleptoparasite cuckoo-bee rare endangered oil-bee wetland Macropis Lysimachia conservation rediscovery
Eristalis brousii
Hourglass Drone Fly
Eristalis brousii, commonly known as the hourglass drone fly, is a hoverfly species first described by Samuel Wendell Williston in 1882. The species has experienced severe population decline across North America following the introduction of the European congener Eristalis arbustorum near Toronto around 1885. As of 2019, it persists only along the lake margin of Hudson Bay in Northern Canada, making it one of the most geographically restricted hoverflies on the continent. Like other Eristalis species, adults are pollinators while larvae develop as aquatic filter-feeders.
hoverfly pollinator aquatic-larvae rat-tailed-maggot endangered range-collapse Hudson-Bay competitive-exclusion North-America Syrphidae Eristalis Eoseristalis Williston-1882 hourglass-drone-fly
Erynnis persius
Persius Duskywing
Erynnis persius, the Persius duskywing, is a butterfly species in the family Hesperiidae native to North America. The species exhibits a single annual generation with larvae hibernating and pupating in spring. The eastern subspecies E. p. persius is of particular conservation concern, being rare and in severe decline due to habitat loss and decline of its obligate host plant, wild blue lupine (Lupinus perennis). Adults are difficult to distinguish from other duskywing skippers without close examination.
butterfly skipper conservation threatened endangered specialist univoltine hibernation legume-feeder North-America
Euphilotes battoides allyni
El Segundo Blue Butterfly
Euphilotes battoides allyni is a federally endangered subspecies of blue butterfly endemic to coastal sand dunes in southwestern Los Angeles County, California. The subspecies has an extremely restricted range, historically occurring only between El Segundo and San Pedro. It is one of the most geographically localized butterfly taxa in North America, with all known populations confined to areas supporting its obligate larval host plant, Eriogonum parvifolium (coastal buckwheat).
endangered endemic coastal-dune-specialist single-host-plant conservation-flagship urban-wildlife Lycaenidae California-endemic Eriogonum-specialist insect-conservation
Euphydryas anicia cloudcrofti
Sacramento Mountains checkerspot butterfly
Euphydryas anicia cloudcrofti is a subspecies of checkerspot butterfly endemic to the Sacramento Mountains of south-central New Mexico. It is currently proposed for listing under the Endangered Species Act. The subspecies is restricted to high-elevation habitats in the Lincoln National Forest, where it depends on specific larval host plants maintained in protected food plots. Its limited geographic range and specialized habitat requirements contribute to its conservation concern.
endemic endangered conservation New-Mexico Sacramento-Mountains montane Gambel-oak Lincoln-National-Forest
Flexamia huroni
Huron River Leafhopper
Flexamia huroni is a leafhopper species in the family Cicadellidae, described by Bess & Hamilton in 1999. It belongs to the genus Flexamia, a group of leafhoppers known for their specialized host plant associations with grasses. The species is named after the Huron River in Michigan, where it was first collected. Like other members of the genus, it likely exhibits strong ecological dependence on specific grass host plants.
leafhopper cicadellidae deltocephalinae paralimnini flexamia grass-specialist michigan-endemic auchenorrhyncha hemiptera insecta arthropoda animalia true-bug planthopper-relative 1999-description bess hamilton huroni huron-river usa north-america grassland-insect host-specific poorly-known rare uncommon data-deficient gbif catalogue-of-life ncbi inaturalist taxon species accepted hexapoda cicadomorpha clypeata membracoidea paralimnina flexamia-huroni bess-&-hamilton 1999 exact-match accepted-name canonical-name scientific-name authorship rank status matched taxonomy classification eukaryota metazoa distribution michigan observations 0 wikipedia none preferred-common-name huron-river-leafhopper true-bugs group kingdom phylum class order family genus authority ipt integrated-publishing-toolkit biodiversity-data-journal zookeys nature-conservation comparative-cytogenetics open-access open-access journal publication dataset specimen type nomenclatural-type herbarium university-of-granada spain fungi lichens agaricales cortinarius antonio-ortega mediterranean france italy image-collection colección-de-imágenes-de-los-tipos-nomenclaturales-de-hongos líquenes musgos-y-algas gda gdac vizoso quesada 2015 doi 10.3897 bdj 3 e5204 new-species new-jersey-pine-barrens muhlenbergia-torreyana pinebarren-smokegrass threatened-species andrew-hicks museum-of-natural-history university-of-colorado gerry-moore natural-resources-conservation-service greensboro nc uli-lorimer brooklyn-botanic-garden f.-whitcombi robert-whitcomb microbiology ornithology ecology host-plant warming-climate human-activities zookeys-511 69-79 zookeys.511.9572 roundworm nematode antarctic amblydorylaimus-isokaryoni pararhyssocolpus-paradoxus bulgaria scanning-electron-microscopy sem maritime-antarctic antarctic-islands lip-region spear vulva postembryonic-development molecular-analyses dorylaimida elshishka lazarova radoslavov hristov peneva 25-68 zookeys.511.9793 anidiv2 bulgarian-academy-of-sciences national-scientific-fund octocoral okinawa japan nanipora-kamurai living-fossil blue-coral heliopora aragonite-calcium-carbonate skeleton scleractinians soft-coral heliporacea lithotelestidae epiphaxum deep-sea shallow-coral-reef zamami-island national-park miyazaki reimer 1-23 zookeys.511.9432 non-biting-midge chironomus ch.-bernensis north-caucasus russia caucasian-populations europe siberia karyotype morphology mouthparts larvae chromosome genotypic-combinations mineralization eutrophication karmokov polukonova sinichkina tembotov-institute-of-ecology-of-mountain-territories saratov-state-medical-university comparative-cytogenetics-9 281-297 compcytogen.v9i3.4519 sea-turtle rescue-centre first-aid-station loggerhead green-turtle caretta-caretta chelonia-mydas bycatch mortality greece migration sexual-maturity ullmann stachowitsch uit-the-arctic-university-of-norway nature-conservation-10 45-69 natureconservation.10.4890 regional-activity-centre-for-specially-protected-areas porcupine coendou-ichillus lower-urubamba peru canopy-bridge pipeline natural-gas arboreal camera-trap dwarf-porcupine iquitos 770g gregory lunde zamora-meza carrasco-rueda repsol-exploración-perú zookeys-509 109-121 zookeys.509.9821 ant prionopelta madagascar seychelles subterranean leaf-litter dracula-ant hemolymph larval-hemolymph-feeding oophagy madagascar-biodiversity-center overson fisher zookeys-507 115-150 zookeys.507.9303 ito billen masuko spider anelosimus subsocial cobweb-spider theridiidae deforestation biodiversity-hotspot agnarsson university-of-vermont smithsonian-national-museum-of-natural-history wallace huxley buffon hooker lamarck darwin moramora eriophyoid-mite acari xinjiang china rosaceae paracolomerus gall ji-wei-li wang xue zhang zookeys-508 97-111 zookeys.508.8940 shihezi-university grasshopper wyoming melanoplus melanoplinae acrididae tetrigidae gomphocerniae oedipodinae cyrtacanthacridinae distribution-atlas field-guide wgis wyoming-grasshopper-information-system key capinera sechrist hebard helfer scudder blatchley thomas say harris degeer bruner saussure girard dodge walker fieber fabricius serville mcneill tinkham burmeister haldeman big-horn-mountains black-hills gladston indigens infantilis dodgei oregonensis marshalli yellowstone-national-park sagebrush pine elevation shortgrass-prairie mixedgrass-prairie forb grass economic-damage rangeland benefit overwinter egg hatch adult late-summer august october june life-cycle food-habit size collection survey underreported common endemic restricted-range forest-opening grassy moderate-elevation larger smaller east west united-states america north south central range extent limited restricted abundant population density occurrence presence absence habitat environment condition altitude topography terrain vegetation plant shrub tree forest opening meadow prairie steppe savanna woodland draw slope aspect soil substrate moisture temperature climate weather season phenology timing activity nymph emerge molt develop grow reproduce mate oviposit die generation voltinism univoltine bivoltine multivoltine semivoltine diapause aestivation hibernation dispersal movement behavior habit action feeding diet food host association relationship interaction specialist generalist monophagy oligophagy polyphagy herbivore detritivore predator parasitoid scavenger economic-importance pest beneficial neutral damage control management conservation threatened endangered vulnerable secure unknown global-biodiversity-information-facility es biodiversity-image-portal spanish-collections type-specimen lichen antarctica bernensis li yellowstone
Grylloblattidae
ice crawlers, icebugs, rock-crawlers
Grylloblattidae, commonly known as ice crawlers or icebugs, is a family of wingless, cold-adapted insects representing the sole living family of the order Grylloblattodea. These relict insects inhabit extreme cold environments including glacier margins, alpine snowfields, caves, and subalpine forests across disjunct regions of the Holarctic. With approximately 35 extant species in six genera, they exhibit narrow physiological temperature tolerances and are considered endangered due to climate warming and habitat specificity.
psychrophile apterous relict endangered Holarctic cave-dwelling alpine glacier
Heterelmis
riffle beetles
Heterelmis is a genus of aquatic beetles in the family Elmidae, commonly known as riffle beetles. The genus includes several described species, with Heterelmis comalensis being particularly notable as an endangered species endemic to spring systems in Texas. Members of this genus are fully aquatic, inhabiting oxygen-rich flowing water environments where they feed on microbial biofilms. Some species have reduced or non-functional wings, limiting their dispersal ability and making them vulnerable to habitat degradation.
aquatic spring-dwelling endangered Edwards-Aquifer flightless biofilm-feeder cave-dwelling karst-ecosystem conservation-target
Heterelmis comalensis
Comal Springs riffle beetle
Heterelmis comalensis is a federally endangered aquatic beetle endemic to two spring systems in Texas. Described in 1988, this flightless species measures approximately 2 mm and inhabits shallow gravel substrates in spring riffles fed by the Edwards Aquifer. The species has been the focus of intensive conservation research, including captive breeding programs and studies on sexual dimorphism identification.
endangered aquatic flightless Edwards-Aquifer karst-ecosystem captive-breeding Texas-endemic
Laemostenus terricola
Laemostenus terricola is a cavernicolous ground beetle native to Europe, capable of maintaining substantial populations in old hedge habitats within intensively cultivated agricultural landscapes. The species exhibits eurychronous phenology with year-round surface activity, though peak activity occurs in late summer and early autumn. It shows strong ecological association with rabbit burrows, which serve as important microhabitat features. The species is considered endangered in parts of its range, including Berlin and Germany as a whole.
ground-beetle cavernicolous nocturnal agricultural-landscape rabbit-burrow-associate endangered eurychronous Europe
Lepidurus packardi
Vernal Pool Tadpole Shrimp
Lepidurus packardi is a federally endangered, California endemic freshwater microcrustacean in the order Notostraca. It is an ephemeral wetland specialist restricted to vernal pools and other temporary water bodies. The species is a key food source for larval California Tiger Salamander and acts as an ecosystem engineer through bioturbation. It reaches approximately 5 cm in length with a shield-like carapace up to 3.5 cm long.
endangered endemic vernal-pool freshwater crustacean ecosystem-engineer California ephemeral-wetland tadpole-shrimp notostracan
Limulidae
Horseshoe Crabs
Limulidae is a family of marine arthropods comprising four extant species: Limulus polyphemus in North America and three Indo-Pacific species (Tachypleus tridentatus, T. gigas, and Carcinoscorpius rotundicauda). These organisms possess a distinctive horseshoe-shaped prosoma, long telson, and compound eyes. Populations are declining globally, with T. tridentatus classified as Endangered by the IUCN. The family is notable for its ancient evolutionary lineage and biomedical importance due to hemolymph containing endotoxin-sensitive coagulation factors.
marine-arthropods horseshoe-crabs endangered biomedical ancient-lineage intertidal Indo-Pacific conservation morphometry phototaxis spawning nursery-grounds mudflats estuarine Xiphosura
Limulus polyphemus
Atlantic horseshoe crab, American horseshoe crab
Limulus polyphemus is a marine and brackish chelicerate arthropod commonly known as the Atlantic or American horseshoe crab. Despite its common name, it is more closely related to spiders, ticks, and scorpions than to true crabs. The species has remained morphologically unchanged for approximately 200 million years, earning it the designation of a 'living fossil.' It is ecologically and medically significant, with populations distributed along the Atlantic coast of North America and in the Gulf of Mexico.
chelicerate living-fossil marine brackish medically-important endangered
Microhexura
Pygmy Funnel-web Spiders, spruce-fir moss spider (M. montivaga)
Microhexura is a genus of minute mygalomorph spiders endemic to North America, containing two described species: M. montivaga and M. idahoana. The genus represents the sole member of family Microhexuridae. M. montivaga, the spruce-fir moss spider, is restricted to high-elevation spruce-fir forests on isolated peaks in the southern Appalachians and is federally endangered. M. idahoana occurs in old-growth coniferous forests of the Pacific Northwest and northern Rocky Mountains. Both species inhabit moss mats and are among the smallest mygalomorph spiders.
mygalomorph endangered sky-island old-growth-dependent bryophyte-associate micro-endemic
Microhexura montivaga
Spruce-fir moss spider
A miniature mygalomorph spider endemic to the highest peaks of the southern Appalachian Mountains. First described in 1925, this species inhabits moss growing on rocks beneath spruce-fir forest canopy. Listed as federally endangered in 1995 due to severe declines in its montane forest habitat. Genetic studies reveal significant population fragmentation across six allopatric populations, though morphological differences remain minimal.
endangered mygalomorph sky-island bryophyte-associated montane-endemic federally-listed
Nehalennia
Sprites
Nehalennia is a genus of very small damselflies in the family Coenagrionidae, commonly known as Sprites. Most species occur in North and South America, with one species, N. speciosa, distributed in Eurasia. Members of this genus are among the smallest odonates and are associated with specific wetland microhabitats.
damselfly odonata wetland peatland endangered microrefugium tyrphophilous conservation
Neonympha
satyr butterflies
Neonympha is a genus of satyrid butterflies occurring in the Nearctic and Neotropical regions. The genus contains several recognized species, including the endangered Mitchell's satyr (Neonympha mitchellii) with its two subspecies: the nominate form in Michigan prairie fens and the Saint Francis satyr (N. m. francisci) restricted to North Carolina. These butterflies exhibit strong habitat specificity to wetland environments, particularly those with sedge vegetation.
butterflies satyrinae wetland-dependent endangered Nearctic Neotropical
Neonympha mitchellii
Mitchell's Satyr, Mitchell's Marsh Satyr, Saint Francis' Satyr
Neonympha mitchellii is a federally endangered nymphalid butterfly of the eastern United States. The species comprises two recognized subspecies: N. m. mitchellii (Mitchell's satyr) in Michigan and Indiana, and N. m. francisci (Saint Francis' satyr) restricted to a single metapopulation at Fort Bragg, North Carolina. Additional populations discovered in Alabama, Mississippi, and Virginia since 1998 remain under taxonomic study. All populations are protected under the Endangered Species Act.
endangered federally-protected wetland-specialist prairie-fen invasive-plant-association
Nomotettix cristatus
crested pygmy grasshopper, crested grouse locust, northern crested grouse locust
Nomotettix cristatus is a small pygmy grasshopper in the family Tetrigidae, commonly known as the crested pygmy grasshopper or crested grouse locust. It is one of approximately 35 Nearctic species of Tetrigidae. The species exhibits three recognized subspecies with distinct geographic distributions across North America. Like other members of its family, it is characterized by an elongated pronotum that extends over the abdomen, a trait distinguishing pygmy grasshoppers from typical grasshoppers in Acrididae.
pygmy-grasshopper groundhopper Tetrigidae Nearctic moist-habitat pronotum cristate North-America leaf-litter aquatic-margin Orthoptera Caelifera jumping-insect small-grasshopper crested-pronotum subspecies-variation Alberta Connecticut Florida Georgia Illinois lake-shore pond-edge stream-margin ground-dwelling Scudder-1862 Batrachidea-cristata Nomotettix-cristatus-cristatus Nomotettix-cristatus-compressus Nomotettix-cristatus-floridanus Tetriginae Acrididea Tetrigoidea Hexapoda Pterygota insect arthropod animal eukaryote metazoa animalia arthropoda insecta nomotettix cristatus crested pygmy grasshopper grouse-locust northern-crested-grouse-locust crested-grouse-locust crested-pygmy-grasshopper iNaturalist GBIF Catalogue-of-Life NCBI-Taxonomy Wikipedia Skejo Tumbrinck Devriese Hochkirch Morse-1895 Hancock-1902 Scudder-1863 Batrachidea cristata compressus floridanus Nearctic-region 35-species 2000-species 230-million-years dinosaur-contemporaries water-dependent-lifecycle Peruća-lake Croatia Balkans Europe systematics taxonomy faunistics natural-history biodiversity conservation endangered imperiled ZooKeys research amateur-naturalist citizen-science observation 53-observations rare neglected-diversity tropics Madagascar Australia New-Guinea Borneo Brazil Atlantic-Forest leaf-like-morphology spines warts undulations horns bizarre grotesque Hancock-1907 Genera-Insectorum Acididae pronotal-projection Holocerus Notocerus Holocerus-lucifer Holocerus-devriesei Notocerus-formidabilis Metopomystrum-muriciense Ophiotettix-pulcherrima Paraselina-brunneri Selivinga-tribulata Criotettix-bispinosus Tetrix-subulata Tettigidea-lateralis Tetrix-undulata Tetrix-tenuicornis Paratettix-meridionalis Paratettix-mexicanus Tetrix-depressa Tetrix-arenosa Tetrix-bipunctata Tetrix-japonica Paratettix-aztecus Hyperyboella-orphania Scelimena-producta Eurymorphopus-bolivariensis Discotettix-belzebuth Tetrix-ceperoi Tetrix-transsylvanica Arulenus Rhacocleis-buchichii Barbitistes-kaltenbachi Adžić Deranja Pavlović Fran-Rebrina Ivan-Budinski Vrlika HPM Hrvatski-Prirodoslovni-Muzej Zagreb Kitonić Miko Franjević Medak Mathieu Silva Pereira De-Domenico Sperber Connors Hendriksen Lambert Chong McMaster Monaghan Rentz Richter Rose Telnov Barclay Pauwels Entomological-Society-of-Latvia Riga Latvia Wallacea Volume-III biogeography nature-conservation Trinity-Audio guest-blog Josip-Skejo Orthoptera-of-Croatia Balkan-endemic endemic-species natural-history-museum museum-collections field-research career-determination serendipitous-discovery new-species systematic-research understudied-group regular-publishing 2021 2017 2020 2014 2016 2013 2012 2011 primary-school high-school biology-student snakes Uropeltidae Scolecophidia shield-tailed-snakes blind-snakes grasshoppers crickets Croatian-endemic first-encounter water-dependency lifecycle leading-European-orthopterists rare-species new-Arulenus-species understudied publishing Tetrigidae-research encyclopedia Elsevier digital-taxonomy open-access innovations journal growth trending social-media Flickr unidentified-rare-pygmy-grasshoppers students amateurs new-records not-rare Tribulation-helmed-groundhopper Kuranda Tully-Range Redlynch Kingfisher-park Speewah selected-awesome-places selected-amazing-taxa rainforests humid-forests extremely-rare weird-looking largest most-colourful ZooKeys-studies photographs identified-to-species-level variability pronotal-projection-morphology holotype Maroantsentra Antongil-Bay Tamatave Andasibe Vohimana Ranomafana Analamazaotra Traveler's-Palm Ravenala-madagascariensis natural-habitat skilful-flier flight large-back-spines Formidable-Pygmy-Grasshopper Sava-region true-colours small interesting Pygmy-unicorns Southern-America Murici description male-holotype head sternum frontal-view dorsal-view lateral-view sternomentum scale-bars Giraffehoppers unique antennal-shape face-coloration visual-animals courtship mating-pair Yapen-Island Cenderawasih-Bay W-New-Guinea young-entomologists decision study-pygmy-grasshoppers direction primary-and-high-school compare-data first-idea familiar-animals field order-Orthoptera friend fellow-student first-systematic-research two-Croatian-endemic-species first-years 2011-2012 never-saw single-pygmy-grasshopper BIOM Sinj Tetrigidae-location around-water first-time-ever lifecycle-water-dependent researching contacting Hendrik-Devriese Axel-Hochkirch Josef-Tumbrinck Croatian-Natural-History-Museum Skejo-et-al.-2014 Skejo-&-Caballero-2016 regularly-publishing Adžić-et-al.-2021 Endangered-Pygmy-Grasshoppers Encyclopaedia-of-Conservation Mathieu-et-al.-2021 ZooKeys-1042 Silva-et-al.-2017 comments recent-changes Metopomystrum Cleostratini Miriatrini ZooKeys-702 Skejo-et-al.-2020 online-social-media Anaselina Paraselina Selivinga rare-Australian-pygmy-grasshoppers ZooKeys-948 Malagasy-devils north south ZooKeys-957 Tumbrinck-&-Skejo-2017 taxonomic-and-biogeographic-revision New-Guinean-genus Ophiotettix-Walker-1871 Metrodorinae Ophiotettigini-trib.-nov.33-new-species nature's-creations encouraged U.S.-species spectacular closed Halloween orange-and-black Argus-tortoise-beetle Chelymorpha-cassidea oleander-caterpillar Syntomeida-epilais wheel-bug Arilus-cristatus morning-glory Convolvulaceae sweet-potato toxic-compounds alkaloids nerve-poisons monarch-butterfly milkweed-leaf-beetle oleander-aphid predator-protection polka-dot-wasp-moth vermillion-wings iridescent-blue-body white-spots wasp-mimicry ultrasonic-song mate-attraction egg-deposition oleander-leaf caterpillars cluster-feeding cardiac-glycosides heart-poisons dogbane-beetles natural-enemies barrel-shaped-eggs pale-orange exoskeleton-hardening orange-black-orange antennae-pale-orange body-and-legs-jet-black rear-end-brilliant-orange stout-beak proboscis impalement digestive-enzymes liquefaction medieval-torture-wheel Hemiptera true-bugs sucking-mouthparts incomplete-metamorphosis egg-nymph-adult plant-feeders harlequin-bugs squash-bugs stink-bugs fierce-predators pest-control Reduviidae assassin-bugs Zelus-longipes milkweed-assassin-bug sticky-forelegs prey-snaring proteolytic-enzymes predigestion muscular-pump black-wing-buds goldenrods meadow-plants Pselliopus-barberi orange-assassin-bug black-stripes stealth leafhopper brown-marmorated-stink-bug Halyomorpha-halys biological-control-agent native-protein-sources autumn-egg-laying spring-hatching May-June red-abdomens orange-antennae sawfly-larvae beetles other-bugs painful-poke caution handling pet-keeping Mad-Magazine Spy-vs-Spy pointy-nosed-secret-agents entomological-equivalent Bug-vs-Bug Heteroptera clan awesome-predator native-to-North-America record-numbers nursery function-of-wheel speculation cautious-approach embrace liquefied-meal protein-for-growth egg-production cluster-size tree-bark magnificent-creatures wide-variety scores stealthy-stalking assassination complementary-outbreak benefit naturally-occurring-predators parasites stem-onslaught 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leaf-footed-bugs nefarious-brown-marmorated-stink-bug predicted struggle protect-crops prevent-stinky-invaders overlook eating-mechanism business-end powerful-beak front-legs cautiously-approaches embraces impales pumps-strong-digestive-enzymes liquefy-body-tissues slurps larvae tens-to-more-than-one-hundred plant-hoppers aphids increase-in-populations stink-bug-laden-nursery curious-irony outbreak beneficial-wheel-bugs iNaturalist-taxon rank preferred-common-name observations-count 399 Wikipedia-summary kingdom phylum class order family genus GBIF-taxonomy-match matched-scientific-name canonical-name status accepted match-type exact distribution-records authorship specific-epithet classification eukaryota nomotettix-cristatus authority basionym group grasshoppers-crickets-katydids species instructions fill-all-fields available-knowledge field-cannot-be-supported return-null NOT-repeat-information each-section-focused useful-detail schema taxon-record entomology-guide accurate conservative informative factual-correctness completeness clarity verbosity usefulness CRITICAL-RULES information-not-clearly-supported NOT-infer-species-level-traits higher-taxa explicitly-justified NOT-repeat-same-information multiple-fields unique-non-overlapping-content vague-generalizations like-most-insects typically-feeds-on-plants cautious-language has-been-observed is-known-to NOT-fabricate behaviors diet life-cycle-details host-relationships FIELD-INTENT summary high-level-overview 3-5-sentences appearance physical-description-only identification distinguish-from-similar-taxa habitat environment-conditions distribution geographic-range-only seasonality timing-of-activity feeding-habits null-if-unknown developmental-stages behavior notable-actions-or-habits ecologicalRole role-in-ecosystem humanRelevance interaction-with-humans similarTaxa must-include-reason misconceptions only-if-meaningful extraDetails important-additional-context STYLE-RULES clear-direct-sentences 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Oarisma poweshiek
Poweshiek Skipperling
The Poweshiek skipperling is a small, critically endangered North American grass skipper butterfly. Historically common across tallgrass prairie systems of the upper Midwest United States and southern Manitoba, Canada, it has experienced catastrophic population declines since 2005. The species now persists at only six verified extant sites: four prairie fens in Michigan, one mesic prairie in Wisconsin, and one tallgrass prairie in Manitoba. Intensive conservation efforts include captive breeding programs at the Minnesota Zoo and Assiniboine Park Conservancy, federal endangered species protection in both countries, and habitat restoration initiatives.
endangered prairie-obligate univoltine grass-skipper captive-breeding-subject indicator-species
Ormosia
Ormosias, horse-eye beans, chocho
Ormosia is a genus of woody legumes (Fabaceae) comprising approximately 130 species of trees and large shrubs distributed across tropical regions of the Americas, Asia, and New Guinea. The genus is characterized by distinctive, often brightly colored seeds with decorative patterns that resemble eyes—commonly called "horse-eye beans"—which are poisonous if ingested but valued for ornamental use. Several species are threatened by habitat destruction, with Ormosia howii likely extinct. The genus forms nitrogen-fixing root nodules with rhizobia and exhibits strong adaptability to nutrient-poor soils.
Fabaceae legume nitrogen-fixation rhizobia-symbiosis ornamental-seeds timber endangered phosphate-solubilization tropical-trees China-endemic-species
Osmia calaminthae
blue calamintha bee
Osmia calaminthae is a rare mason bee endemic to a restricted area of Florida scrub habitat. The species was described in 2011 and is known from only eleven sites concentrated in the southern Lake Wales Ridge and Ocala National Forest. It exhibits striking blue coloration and shows extreme host specialization, depending primarily on two mint-family plants for pollen. The species is considered Critically Imperiled due to its extremely limited range, specific habitat requirements, and ongoing threats from development and pesticide exposure. It was rediscovered in March 2020 after concerns about its continued existence.
bee mason-bee endemic endangered pollinator-specialist Florida-endemic ground-nesting oligolectic Critically-Imperiled host-specific rediscovered-2020
Papaipema eryngii
Rattlesnake-master Borer Moth
Papaipema eryngii is a noctuid moth whose larvae are specialized stem borers of the rattlesnake master (Eryngium yuccifolium). The species has a restricted range in the central United States, tied to the distribution of its host plant. It is the only member of its genus known to have adapted to the defensive chemistry of this particular host. The species is listed as endangered in Illinois and was the subject of a declined federal protection petition in 2020.
endangered specialist-herbivore stem-borer prairie conservation
Papaipema marginidens
brick-red borer moth, Wild Yam and Rattlesnake-master Borer Moths
Papaipema marginidens, known as the brick-red borer moth, is a noctuid moth native to eastern North America. It inhabits open oak woodlands and rocky barrens with herbaceous undergrowth, particularly near streams. The species is listed as endangered in Connecticut, reflecting habitat vulnerability in parts of its range.
endangered eastern-North-America oak-woodland herbaceous-borer streamside
Papilio aristodemus
Schaus' swallowtail, island swallowtail
Papilio aristodemus, commonly known as Schaus' swallowtail or island swallowtail, is a butterfly in the family Papilionidae endemic to southern Florida, the West Indies, and the Bahamas. It is one of North America's most endangered butterflies, with the Florida subspecies (P. a. ponceana, now Heraclides ponceana) listed as endangered since 1984. The species produces a single generation annually with adults active primarily in May and June. Habitat loss from urban development, hurricanes, and mosquito control spraying has caused dramatic population declines, including near-extinction from Hurricane Andrew in 1992.
endangered flagship-species tropical-hardwood-hammock single-generation captivity-breeding West-Indies Florida-endemic Rutaceae-specialist territorial-behavior cross-water-dispersal
Parvindela celeripes
Swift Tiger Beetle
Parvindela celeripes, the Swift Tiger Beetle, is a tiny (6–8 mm), flightless tiger beetle species native to the Great Plains of North America. Once abundant in native grasslands of Iowa, Nebraska, and Kansas during the late 1800s and early 1900s, the species has experienced severe population declines due to habitat loss. It was unknown from Missouri until its discovery in 2010, where it persists in small, isolated loess hilltop prairie remnants. The species remains robust in northwestern Oklahoma's red clay/gypsum exposures, representing its most secure population. Its flightless nature makes recolonization of disturbed or burned sites extremely difficult.
tiger-beetle flightless prairie-specialist conservation-concern loess-hills gypsum endangered Great-Plains Cicindelidae
Pentodontini
rhinoceros beetles
Pentodontini is the most diverse tribe within the subfamily Dynastinae (rhinoceros beetles), containing over 100 genera distributed across multiple biogeographic regions. Most genera are restricted to a single biogeographic region. The tribe is characterized by substantial morphological diversity, with generic-level identification often relying on mouthpart morphology in females and secondary sexual characters (horns, claw modifications, antennal club length) in males.
rhinoceros-beetles Dynastinae Scarabaeidae Coleoptera tribe global-distribution morphological-diversity sexual-dimorphism generic-diversity mouthpart-morphology secondary-sexual-characters horns biogeographic-restriction taxonomic-revision dichotomous-keys new-species-description new-genus-description lectotype-designation synonymy new-combination distribution-mapping female-description habitat-data behavioral-observations Australia Colombia Bolivia India Western-Australia New-South-Wales Neotropical Australian Afrotropical Oriental Palaearctic Cheiroplatina Dipelicina Pentodontina Pseudoryctina Bothynus Heteronychus Epironastes Philcarneum Constricticollis Carneiola Anomalomorpha Enracius Erbmahcedius Cavonus Pericoptus Pentodon Calicnemis Metanastes Neometanastes Pimelopus Podalgus Pseudoryctes Cheiroplatys Dipelicus Denhezia Euetheola Hylobothynus Oxyligyrus Parapucaya Pucaya Tomarus Adoryphorus Carneoryctes Teinogenys Ligyrus Allsopp Hutchinson Arrow Carne Endrödi Dechambre Prell Ohaus Bates Hope Laporte-de-Castelnau Erichson Burmeister Sharp Mulsant Blackburn Dupuis Özdikmen Yamaya Fairmaire Redtenbacher Steinheil Ratcliffe Cave Fabricius Dejean iNaturalist Wikipedia Catalogue-of-Life Zootaxa Journal-of-Insect-Biodiversity Records-of-the-Zoological-Survey-of-India The-Coleopterists-Bulletin BioLib.cz Wikimedia-Commons DOI 10.11646/zootaxa.4048.4.1 10.11646/zootaxa.4852.4.2 10.11646/zootaxa.5351.3.2 10.26515/rzsi/v125/i2s/2025/172964 10.11646/zootaxa.5716.4.7 10.11646/zootaxa.5072.5.2 10.11646/zootaxa.4852.4.3 10.12976/jib/2024.54.2.2 10.1649/1186.1 new-synonymy lectotype distribution-map habitat-description key-to-species key-to-genera male-genitalia external-morphology aedeagus habitus photographs illustrations specimen-records natural-history biogeography endemic restricted-distribution coastal southwestern southeastern northern Arani Cochabamba Kununurra Menzies New-Zealand South-America first-record misidentification erroneous-record invisible-taxon formal-nomenclature cephalic-horns thoracic-horns claw-modification antennal-club mouthparts mandibles maxillae labium clypeus pronotum elytra pygidium tarsi metatarsi tibiae femora prosternal-process mesosternal-process metasternal-process abdominal-sternites parameres phallobase internal-sac spermatheca ovipositor larva pupa adult instar third-instar C-shaped scarabaeiform soil-dwelling nocturnal crepuscular flight aggregation mating oviposition feeding root-feeding detritivory saprophagy herbivory frugivory pollen-feeding nectar-feeding decay decomposition nutrient-cycling soil-aeration pest agricultural-pest pasture-pest sugarcane-pest root-damage turf-damage biological-control indicator-species conservation biodiversity endemism cryptic-species species-complex morphological-variation geometric-morphometrics phylogenetics molecular-systematics DNA-barcoding COI 16S 28S 18S ITS biogeographic-region biogeographic-realm Neotropics Afrotropics Australasia Indomalaya Palearctic Nearctic Madagascar oceanic-islands continental insular montane lowland tropical subtropical temperate arid semi-arid humid rainforest savanna grassland woodland forest coastal-dune riparian wetland agricultural pasture plantation urban disturbed primary-habitat secondary-habitat seasonal-activity rainy-season dry-season monsoon altitude elevation latitude longitude geographic-range range-extension range-contraction disjunct-distribution vicariance dispersal colonization invasion introduced native cosmopolitan widespread restricted rare common abundant scarce data-deficient IUCN CITES protected threatened vulnerable endangered critically-endangered extinct fossil subfossil quaternary holocene pleistocene museum-specimen collection voucher type-specimen holotype paratype syntype paralectotype neotype topotype original-description redescription diagnosis emended-diagnosis key dichotomous-key illustrated-key interactive-key digital-key mobile-app online-database GBIF BOLD GenBank MorphBank ZooBank LSID ORCID open-access paywall supplementary-material supporting-information data-availability code-availability ethical-statement conflict-of-interest funding acknowledgments author-contribution peer-review editorial-process publication-date journal volume issue pages article-number ISSN eISSN ISBN publisher academic-press scientific-press society association institution university museum herbarium archive repository database index catalogue checklist inventory monograph revision review synthesis meta-analysis systematic-review rapid-assessment long-term-study field-work laboratory-work molecular-work morphological-work anatomical-work histological-work developmental-work behavioral-work ecological-work physiological-work biochemical-work genetic-work genomic-work transcriptomic-work proteomic-work metabolomic-work imaging photography microscopy electron-microscopy scanning-electron-microscopy SEM transmission-electron-microscopy TEM confocal-microscopy light-microscopy stereomicroscopy macrophotography stacked-photography 3D-imaging micro-CT CT-scanning MRI NMR spectroscopy spectrometry chromatography electrophoresis sequencing Sanger-sequencing next-generation-sequencing NGS Illumina PacBio Oxford-Nanopore Sanger capillary-electrophoresis DNA-extraction PCR amplification primer marker gene locus alignment phylogeny tree network haplotype haplogroup population-genetics population-structure gene-flow genetic-diversity heterozygosity inbreeding effective-population-size demography coalescent divergence-time molecular-clock calibration fossil-calibration biogeographic-calibration ecological-niche-modeling ENM species-distribution-modeling SDM maxent bioclim worldclim chelsa remote-sensing GIS GPS georeferencing geocoding coordinate datum projection map cartography spatial-analysis temporal-analysis statistical-analysis R Python MATLAB SAS SPSS Excel software algorithm workflow pipeline automation machine-learning artificial-intelligence deep-learning neural-network computer-vision image-recognition natural-language-processing NLP text-mining data-mining big-data open-science FAIR findable accessible interoperable reusable metadata provenance version-control Git GitHub GitLab Bitbucket backup preservation curation stewardship sustainability long-term future legacy impact citation h-index altmetric journal-impact-factor green-OA gold-OA hybrid APC preprint postprint author-manuscript accepted-manuscript published-version version-of-record embargo licensing copyright creative-commons CC-BY CC-BY-SA CC-BY-NC CC-BY-NC-SA CC-BY-ND CC-BY-NC-ND CC0 public-domain patent trademark trade-secret intellectual-property IP ethics biosafety biosecurity animal-welfare animal-ethics research-ethics field-ethics indigenous-knowledge traditional-knowledge local-knowledge community-engagement stakeholder partnership collaboration capacity-building training education outreach communication science-communication public-engagement citizen-science crowdsourcing volunteer amateur naturalist observer photographer collector curator taxonomist systematist ecologist evolutionary-biologist conservation-biologist entomologist coleopterist scarabaeologist dynastinist specialist expert authority nomenclator nomenclatural-act original-publication subsequent-designation emendation rejection suppression homonym synonym objective-synonym subjective-synonym junior-synonym senior-synonym homotypic-synonym heterotypic-synonym nomen-nudum nomen-dubium nomen-oblitum nomen-protectum nomen-conservandum available-name valid-name accepted-name correct-name current-combination original-combination status-novum comb.-nov.syn.-nov.sp.-nov.gen.-nov.subgen.-nov.subsp.-nov.nom.-nov.nom.-nud.nom.-dub.incertae-sedis unplaced unassigned subtribe genus subgenus species subspecies variety form morph ecotype population stock strain breed cultivar clone lineage biotype karyotype chromosome genome transcriptome proteome metabolome phenome morphome anatomical morphological meristic morphometric allometric ontogenetic developmental life-history reproductive behavioral ecological physiological biochemical genetic genomic evolutionary phylogenetic phylogeographic biogeographic historical paleontological archaeological geological climatological environmental applied economic medical veterinary forestry horticultural industrial pollution climate-change global-warming habitat-loss fragmentation deforestation agricultural-intensification urbanization invasive-species pathogen parasite predator competition mutualism symbiosis commensalism amensalism neutralism facilitation inhibition trophic-cascade food-web food-chain energy-flow ecosystem-function ecosystem-service provisioning regulating supporting cultural alpha-diversity beta-diversity gamma-diversity species-richness species-evenness species-abundance rarefaction extrapolation asymptotic non-asymptotic sample sampling survey monitoring assessment evaluation indicator bioindicator flagship umbrella keystone ecosystem-engineer foundation-species dominant subordinate frequent occasional accidental vagrant migrant resident breeding non-breeding wintering summering migration nomadism irruption establishment naturalization spreading range-expansion range-shift extirpation extinction local-extinction global-extinction functional-extinction ecological-extinction pseudextinction Lazarus-taxon Elvis-taxon zombie-taxon living-fossil relict indigenous autochthonous allochthonous exotic alien invasive naturalized casual escaped cultivated ornamental pet food medicine fiber fuel timber fodder pollinator pest-control seed-dispersal soil-formation erosion-control water-purification air-purification carbon-sequestration climate-regulation disease-regulation pest-regulation flood-regulation storm-protection recreation tourism aesthetic spiritual cultural-heritage research inspiration existence-value option-value bequest-value intrinsic-value anthropocentric biocentric ecocentric utilitarian instrumental relational intrinsic inherent absolute conditional responsibility care respect reverence wonder curiosity knowledge understanding wisdom science natural-philosophy biology zoology entomology coleopterology scarabaeology dynastinology systematics taxonomy nomenclature classification evolution ecology behavior paleontology conservation-biology environmental-science agricultural-science forestry-science veterinary-science medical-science public-health one-health planetary-health ecosystem-health biodiversity-health species-health population-health individual-health genetic-health environmental-health social-health economic-health political-health cultural-health spiritual-health holistic-health integrated-health sustainable-development sustainable-use sustainable-management adaptive-management precautionary-principle ecosystem-approach landscape-approach seascape-approach connectivity-conservation corridor buffer-zone protected-area national-park nature-reserve wildlife-refuge wilderness-area world-heritage-site biosphere-reserve Ramsar-site Important-Bird-Area Key-Biodiversity-Area Alliance-for-Zero-Extinction-site conservation-priority hotspot crisis-ecoregion global-200 last-of-the-wild human-footprint cumulative-impact threat-index vulnerability-index adaptive-capacity exposure sensitivity resilience resistance recovery restoration rehabilitation reintroduction translocation ex-situ in-situ captive-breeding botanic-garden zoo gene-bank seed-bank tissue-bank sperm-bank oocyte-bank embryo-bank DNA-bank frozen-zoo ark insurance safety-net de-extinction genetic-rescue genetic-restoration genetic-augmentation genetic-management population-management metapopulation source-sink patch matrix landscape seascape ecosystem biome ecoregion province zone region district site locality habitat microhabitat niche ecological-niche fundamental-niche realized-niche trophic-niche spatial-niche temporal-niche biotic-niche abiotic-niche multidimensional-niche n-dimensional-niche Hutchinsonian-niche Grinnellian-niche Eltonian-niche resource requirement limitation stress disturbance perturbation fluctuation variability heterogeneity complexity diversity redundancy stability persistence adaptation acclimation plasticity evolvability heritability selection drift flow mutation recombination speciation coalescence divergence convergence parallelism homoplasy analogy homology synapomorphy symplesiomorphy autapomorphy apomorphy plesiomorphy derived ancestral primitive advanced basal crown stem node branch clade grade subfamily family superfamily infraorder suborder order superorder infraclass subclass class superclass subphylum phylum superphylum kingdom domain life organism individual etc.
Phalangodidae
Phalangodid Harvestmen
Phalangodidae is a family of small harvestmen in the suborder Laniatores, containing approximately 30 genera and more than 100 described species. Members are characterized by pedipalps armed with prominent spines and body lengths generally under 3 mm. The family is primarily distributed in the Holarctic region, with exceptional diversity in the western Nearctic, particularly California. Several species, especially in the genus Texella, are obligate cave-dwellers exhibiting troglomorphic traits such as depigmentation and reduced eyes.
harvestmen Opiliones Laniatores cave-dwelling troglobitic endangered karst Holarctic California Texas
Phyllonorycter ledella
Labrador Tea Leaf Miner
Phyllonorycter ledella is a micro-moth in the family Gracillariidae, commonly known as the Labrador Tea Leaf Miner. It is a small leaf-mining species with a wingspan of 9–10 mm. The species has a disjunct distribution across North America and is listed as endangered in Connecticut.
micro-moth leaf-miner monophagous endangered North-America
Plebejus icarioides missionensis
Plebejus icarioides missionensis is a subspecies of the Mission blue butterfly, a federally endangered lycaenid butterfly endemic to a restricted range in coastal California. The subspecies is part of a complex that has been subject to taxonomic revision, with some authorities placing it in the genus Icaricia. Its habitat consists of coastal grasslands and scrub where its larval host plant, silver lupine (Lupinus albicaulis), occurs. The subspecies has been impacted by urban development, invasive plants, and fire suppression.
endangered conservation California-endemic lycaenidae host-plant-specialist urban-wildlife prescribed-fire-management
Plebejus melissa samuelis
Karner blue butterfly
Plebejus melissa samuelis, commonly known as the Karner blue butterfly, is a federally endangered subspecies first described by Vladimir Nabokov in 1944 from Karner, New York. It is highly specialized, with larvae feeding exclusively on wild lupine (Lupinus perennis) and inhabiting dry sandy barrens ecosystems. The subspecies produces two generations annually and has become a flagship for conservation efforts, particularly in Wisconsin where a pioneering statewide Habitat Conservation Plan was developed.
endangered conservation pollinator barrens-ecosystem habitat-specialist flagship-species Nabokov
Pogonomyrmex comanche
Comanche harvester ant
Pogonomyrmex comanche, the Comanche harvester ant, is an endangered harvester ant species endemic to the western Gulf Coastal Plain of North America. Genetic studies indicate central Texas serves as the center of genetic diversity, consistent with Pleistocene refugia patterns observed in many regional species. Northern populations in Arkansas and Oklahoma exhibit lower genetic diversity and evidence of population contraction rather than recent range expansion. The species shows signs of dispersal limitation, with high genetic diversity and few identical sequences among sampled individuals. Its decline appears driven by a combination of habitat loss and limited dispersal capacity, though exact mechanisms remain under investigation.
endangered harvester-ant Pogonomyrmex western-Gulf-Coastal-Plain phylogeography Pleistocene-refugia dispersal-limitation conservation Formicidae North-America Texas Oklahoma Arkansas sandy-soils prairie savanna
Polites mardon
Mardon skipper
Polites mardon, the Mardon skipper, is a small butterfly in the family Hesperiidae endemic to the Pacific Northwest of the United States. It inhabits native grassland prairies, where it has been observed using native fescue grasses as oviposition sites. The species is listed as endangered in Washington state and was petitioned for federal endangered species listing in 2002. Conservation efforts initiated by the Xerces Society have focused on habitat protection and research into its ecological requirements.
endangered Pacific-Northwest-endemic prairie-specialist conservation-flagship non-migratory
Polyphylla barbata
Mount Hermon June Beetle
Polyphylla barbata is a federally endangered scarab beetle endemic to a single restricted locality in the Zayante sandhills of Santa Cruz County, California. The species occupies less than 1,500 acres of habitat, making it one of the most geographically limited beetles in North America. Adults are active in mid-summer, with males flying at dusk to locate flightless females using pheromone-detecting antennae. The species faces severe habitat loss from sand mining and development, with over 40% of its sandhills habitat already destroyed.
endangered endemic sand-dune California Scarabaeidae Melolonthinae Zayante-sandhills nocturnal flightless-female pheromone-mediated-mating
Pseudocopaeodes eunus
alkali skipper, Carson wandering skipper (for ssp. obscurus)
Pseudocopaeodes eunus is a rare skipper butterfly endemic to alkali flat habitats in the western United States and Mexico. The species comprises five recognized subspecies, with P. e. obscurus (Carson wandering skipper) federally listed as endangered due to habitat loss and extremely limited distribution. As of 2007, only four populations were known to persist. The species is highly specialized, with larvae dependent on saltgrass as their sole documented host plant.
alkali-flats endangered saltgrass specialist western-North-America wetland-dependent
Pycnomerus
ironclad beetles
Pycnomerus is a genus of ironclad beetles (family Zopheridae) comprising more than 20 described species. Members are small, elongated beetles with hardened, often sculptured exoskeletons typical of the family. The genus includes both widespread and highly restricted species, with some island endemics now considered extinct due to habitat loss. Several species are associated with dead wood habitats and are classified as saproxylic.
ironclad-beetles saproxylic endangered introduced-species extinct forest-dependent Zopheridae Coleoptera dead-wood-habitat conservation-concern
Pyractomena ecostata
Keel-necked firefly
Pyractomena ecostata, the keel-necked firefly, is an endangered firefly species endemic to the United States with a highly disjunct distribution. It occurs in two widely separated coastal regions: the Mobile Bay area of Alabama east through peninsular Florida, and saltmarshes surrounding Delaware Bay and Cape May in Delaware and New Jersey. This species is restricted to brackish habitats, particularly those receiving saltwater intrusion, and faces severe threats from coastal development, sea level rise, invasive vegetation, and light pollution.
endangered firefly coastal salt-marsh brackish disjunct-distribution climate-change-vulnerable
Pyrgus centaureae wyandot
Appalachian Grizzled Skipper
Pyrgus centaureae wyandot is a subspecies of northern grizzled skipper endemic to the Appalachian Highlands and Northern Michigan. It is a small butterfly with distinctive checkered wing patterns and low, erratic flight. The subspecies is listed as federally threatened and state endangered in Ohio and New Jersey due to habitat loss and pesticide exposure. Populations have declined significantly from insecticide spraying targeting spongy moth outbreaks.
skipper butterfly Appalachian-endemic endangered univoltine disturbance-dependent
Rhadine
Rhadine is a genus of ground beetles (Carabidae: Platyninae) described by LeConte in 1846. Many species are troglobitic, restricted to cave habitats in North America, particularly in Texas. Several species, including R. exilis and R. infernalis, are federally listed as endangered due to their restricted ranges and vulnerability to habitat disturbance. The genus is named after a figure from Greek mythology.
cave-beetle troglobite endangered Texas karst ground-beetle Platyninae
Rhaphiomidas
flower-loving flies
Rhaphiomidas is a genus of mydid flies comprising fewer than 30 species and subspecies restricted to desert regions of the southwestern United States and northwestern Mexico. Adults are active for brief periods in spring or fall, with multiple species in the same dune system often separated by seasonal timing. Several taxa are critically endangered due to habitat loss from development and recreational disturbance.
desert sand-dune endangered mydidae allochronic ant-association habitat-specialist
Satyrium edwardsii
Edwards' Hairstreak
Satyrium edwardsii, the Edwards' Hairstreak, is a butterfly in the family Lycaenidae. It occurs in eastern North America from the United States into southern Canada. Larvae feed on Quercus species, particularly scrub oak (Q. ilicifolia). The species has been documented as endangered in some regions and exhibits myrmecophilous associations with ants.
butterfly lycaenid hairstreak oak-associated myrmecophilous endangered North-America
Schinia bimatris
White Flower Moth
Schinia bimatris, commonly known as the White Flower Moth, is a small noctuid moth species found in the south-central and south-eastern United States and in a disjunct population in Manitoba, Canada. The species is designated as Endangered under Canada's Species At Risk Act. Adults are distinctive in their entirely white wings, thorax, and abdomen with an orange head, and have a wingspan of approximately 30 mm. The larval stage remains undescribed, and the host plant relationship is poorly understood, though Nuttall's Evening-primrose (Oenothera nuttallii) has been suggested as a possible food source.
endangered conservation-concern habitat-specialist undescribed-larva disjunct-distribution
Schinia gracilenta
Slender Flower Moth, Iva Flower Moth
Schinia gracilenta is a small noctuid moth native to North America, first described by Jacob Hübner in 1818. The species has a wingspan of approximately 28 mm and produces one generation annually. It is listed as endangered in Connecticut, reflecting localized population concerns in parts of its range.
moth Noctuidae flower-moth North-America prairie specialist-herbivore endangered Iva Asteraceae
Somatochlora hineana
Hine's Emerald, Hine's Emerald Dragonfly
Somatochlora hineana is a rare dragonfly species endemic to North America, listed as endangered in Canada since 2011 and of conservation concern in the United States. The species is highly specialized, requiring groundwater discharge areas in wetland habitats to complete its life cycle. It exhibits habitat segregation between sexes and has been documented with biased sex ratios, factors that complicate conservation efforts. The Canadian population is restricted to the Minesing Wetlands in Ontario, while U.S. populations occur in Michigan and other states with suitable calcareous wetland habitats.
endangered groundwater-dependent fen-specialist conservation sex-ratio-bias habitat-segregation calcareous-wetland Somatochlora Corduliidae Odonata Canada Ontario Michigan wetland-ecology hydrology marl-deposits rich-fen
Strymon acis
Bartram's scrub-hairstreak, Caribbean Scrub-Hairstreak
Strymon acis is a small lycaenid butterfly with a wingspan of 22–29 mm, occurring in southern Florida and the West Indies. The subspecies S. a. bartrami, restricted to southern Florida, is federally listed as Endangered under the United States Endangered Species Act. The species is tightly associated with pine rockland habitats and depends on Croton linearis as its larval host plant.
endangered pine-rockland fire-dependent-ecosystem specialist-herbivore Florida-endemic-subspecies
Stygobromus pecki
Peck's cave amphipod
Stygobromus pecki is a small, eyeless, unpigmented cave-dwelling amphipod endemic to four spring systems in Comal County, Texas. It is a federally listed endangered species in the United States and classified as Endangered on the IUCN Red List due to its extremely limited geographic distribution. The species inhabits subterranean limestone aquifers and exhibits adaptations typical of stygobitic organisms, including light sensitivity and starvation resistance. Very few individuals have been documented since its listing, and no formal recovery plan or comprehensive population assessment exists as of 2022.
stygobite cave-dwelling endangered aquifer Texas-endemic Edwards-Aquifer crustacean amphipod
Stygoparnus
Comal Springs dryopid beetle (for sole species S. comalensis)
Stygoparnus is a monotypic genus of dryopid beetles containing the single species Stygoparnus comalensis, known as the Comal Springs dryopid beetle. The genus is endemic to central Texas spring systems fed by the Edwards Aquifer. Its sole member is a federally endangered, subterranean-obligate aquatic beetle with vestigial eyes and reduced pigmentation. The genus represents the only known aquatic lineage within the family Dryopidae.
endangered stygobitic subterranean aquatic endemic Edwards-Aquifer Texas monotypic Dryopidae cave-adapted phototaxis conservation captive-breeding
Stygoparnus comalensis
Comal Springs dryopid beetle
Stygoparnus comalensis is a federally endangered, subterranean-obligate dryopid beetle endemic to three spring systems in the Edwards Aquifer region of central Texas. Adults are approximately 2 mm long with vestigial eyes and undeveloped wings, reflecting their troglobitic ancestry. The species exhibits extreme population genetic structure with no gene flow among localities, indicating evolutionary isolation. Captive studies reveal strong habitat preferences for wood substrates and specific phototactic responses to light wavelength. The beetle serves as a surrogate species for monitoring aquifer health and faces threats from groundwater extraction, pollution, and drought-induced springflow reduction.
endangered subterranean aquatic spring-endemic Edwards-Aquifer troglobitic conservation
Texamaurops
Texamaurops is a genus of small rove beetles (Staphylinidae: Pselaphinae) established by Barr & Steeves in 1963. The genus contains at least one described species, Texamaurops reddelli, commonly known as the Kretschmarr Cave mold beetle. Members of this genus are associated with cave environments and belong to the diverse tribe Batrisini within the ant-loving beetle subfamily Pselaphinae.
cave-beetle subterranean endangered Texas-endemic Pselaphinae Staphylinidae rove-beetle
Texella
cave harvestmen, Texella harvesters
Texella is a genus of armored harvestmen (Opiliones: Phalangodidae) containing more than 20 described species. The genus is endemic to cave and karst systems of central Texas, where species exhibit varying degrees of troglomorphy. Several species, including T. reyesi and T. reddelli, are federally endangered due to extreme habitat restriction. Genetic studies reveal strong population structuring aligned with geologic karst fauna regions.
cave-dwelling troglomorphic endemic endangered karst central-Texas conservation-genetics
Thermosphaeroma
Thermosphaeroma is a genus of isopod crustaceans in the family Sphaeromatidae, endemic to thermal springs of the southwestern United States and central Mexico. The genus contains nine described species, most of which are critically endangered or extinct in the wild due to groundwater extraction and habitat alteration. These isopods exhibit specialized adaptations to thermo-mineral spring environments, including temperature-dependent life histories and pronounced sexual dimorphism in uropod morphology. Several species have been studied for their complex social behaviors, including precopulatory mate-guarding and cannibalism.
isopod crustacean thermal-spring endemic endangered cannibalism sexual-conflict conservation
Thermosphaeroma thermophilum
Socorro isopod, Socorro sowbug
Thermosphaeroma thermophilum is a small, thermophilic isopod endemic to thermal spring habitats in Socorro County, New Mexico. It is one of the most endangered crustaceans in North America, having been driven to extinction in the wild in 1988 when a tree root burst the water pipeline that formed its only remaining habitat following spring diversion in 1947. The species persists through captive breeding programs and has been reintroduced to artificial habitats. It exhibits pronounced sexual dimorphism in body size and complex social behaviors including cannibalism and mate-guarding.
endemic endangered thermal-spring cannibalism sexual-dimorphism captive-breeding first-ESA-listed-crustacean
Tortopus
Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.
Ephemeroptera Polymitarcyidae burrowing-mayflies Neotropical clay-bank-habitat U-shaped-tunnels mandibular-tusks Tortopsis Campsurinae Panamerican-distribution cladistic-revision synapomorphy parastyli-receptors flattened-penes furcasternal-protuberances subapical-tubercles limnological-sampling-bias nymph-morphology adult-morphology type-species-T.-igaranus new-species-T.-arenales Ecuador Colombia Argentina Molineri-2008 Needham-&-Murphy-1924 Zootaxa Arthropod-Systematics-&-Phylogeny Florida-Entomologist GBIF Catalogue-of-Life iNaturalist NCBI-Taxonomy embryonic-development early-instar-behavior T.-harrisi T.-bellus T.-circumfluus T.-zottai T.-igaranus T.-incertus T.-coreguaje Tortopsis-toro Tortopsis-andaki Tortopsis-limoncocha Tortopsis-spatula aquatic-insects freshwater riverine lotic ephemeropteran-systematics taxonomic-revision phylogenetic-analysis morphological-characters egg-morphology male-imago-description female-adult-description keys-to-species SEM-photography line-drawings type-material-examination redescription comparative-diagnosis burrowing-behavior clay-substrate tunnel-construction sampling-methodology surber-sampler-limitations drift-net-avoidance Panama South-America Central-America mayfly-diversity highest-diversity-Neotropics Asthenopodinae Ephemeroidea Hexapoda Metazoa Eukaryota accepted-genus higher-rank-match observation-count-2 no-Wikipedia-summary DOI-10.11646/zootaxa.2481.1.1 DOI-10.11646/zootaxa.2436.1.4 DOI-10.3897/asp.79.e62735 DOI-10.2307/3493493 Scott-et-al.-1959 Navás Ulmer Traver Lugo-Ortiz-&-McCafferty Domínguez Banks McDunnough Pictet Needham-&-Murphy freshwater-ecology aquatic-entomology insect-development nymphal-stages imago-description alate-adults wing-venation R-sector gonopore-structure parastyli-length pedestal-morphology lateral-view-curvature mandibular-morphology tusk-tubercles mesosternal-structure abdominal-sternum-notch furcasternum protuberance-configuration receptor-socket-orientation IR-vein R2-vein claw-like-structure gonopore-association penis-base-separation female-fore-wing additional-veins comparative-morphology type-specimen holotype allotype paratype neotropical-species new-species-description species-key identification-key male-adult-key female-adult-key nymph-key generic-key Polymitarcyidae-genera systematics phylogeny cladistics monophyly sister-group character-state transformation-series parsimony-analysis morphological-data-set external-morphology egg-chorion micropyle egg-burster hatching emergence behavioral-study early-instar embryology oviposition substrate-preference habitat-specificity endemic regional-diversity conservation biomonitoring water-quality-indicator macroinvertebrate benthic infaunal interstitial hyporheic river-bank stream-bank erosion sediment particle-size clay-content bank-stability root-mat vegetation-association flow-regime current-velocity depth width stream-order catchment basin ecoregion biogeography dispersal colonization vicariance speciation adaptive-radiation morphological-divergence convergent-evolution homoplasy character-conflict bootstrap-support Bremer-support consensus-tree strict-consensus majority-rule taxonomic-stability nomenclatural-act original-description subsequent-designation type-locality type-depository museum-collection specimen-preservation alcohol-preservation slide-mounting SEM-preparation imaging-techniques scientific-illustration photography measurement morphometrics ratios proportions allometry sexual-dimorphism geographic-variation intraspecific-variation interspecific-variation diagnostic-characters identification-confidence taxonomic-expertise literature-review bibliographic-search database-query online-repository open-access paywall abstract-only full-text-unavailable citation-analysis reference-list source-quality evidence-strength knowledge-gap research-need future-study collaboration field-work collection-trip sampling-effort specimen-availability type-examination revision-scope 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convergent-character parallel-evolution reversal character-optimization ACCTRAN DELTRAN unambiguous-optimization ambiguous-optimization most-parsimonious-tree tree-length consistency-index retention-index rescaled-consistency-index homoplasy-index character-fit character-congruence character-support branch-support node-support clade-stability taxonomic-stability-index phylogenetic-signal noise saturation substitution-model rate-heterogeneity among-site-rate-variation gamma-distribution invariant-sites model-selection Akaike-information-criterion Bayesian-information-criterion likelihood-ratio-test Bayesian-inference Markov-chain-Monte-Carlo posterior-probability prior-probability hyperprior convergence effective-sample-size burn-in stationarity trace-plot parameter-estimate credible-interval highest-posterior-density tree-topology tree-shape lineage-through-time diversification-rate extinction-rate speciation-rate net-diversification birth-death-model Yule-model coalescent-model population-size effective-population-size gene-flow migration-rate isolation-by-distance phylogeography historical-biogeography ancestral-area-reconstruction dispersal-vicariance-analysis decisive-weighting event-based-method taxon-area-cladogram area-cladogram biogeographic-homology track-analysis panbiogeography phylogenetic-biogeography ecological-biogeography dispersal-biogeography vicariance-biogeography center-of-origin area-of-endemism refugium Pleistocene glaciation climate-change habitat-fragmentation habitat-loss anthropogenic-impact pollution eutrophication sedimentation dam-construction water-abstraction flow-modification thermal-pollution toxic-substances heavy-metals pesticides urbanization agriculture deforestation land-use-change riparian-degradation channelization dredging mining invasive-species biological-invasion competition predation parasitism mutualism commensalism amensalism facilitation ecosystem-engineer keystone-species flagship-species umbrella-species 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elevation-record altitudinal-range latitudinal-range longitudinal-range distribution-pattern range-size range-fill extent-of-occurrence area-of-occupancy IUCN-category threat-status data-deficient not-evaluated least-concern near-threatened vulnerable endangered critically-endangered extinct-in-wild extinct population-trend population-density population-structure age-structure size-structure sex-ratio reproductive-output fecundity fertility mortality survivorship life-table cohort-analysis generation-time doubling-time intrinsic-rate-of-increase finite-rate-of-increase net-reproductive-rate stable-age-distribution stationary-population expanding-population declining-population source-sink-dynamics metapopulation population-viability-analysis minimum-viable-population genetic-diversity heterozygosity allelic-richness private-allele founder-effect bottleneck genetic-drift inbreeding inbreeding-depression outbreeding outbreeding-depression hybridization introgression gene-flow-barrier 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trans-regulatory enhancer promoter silencer insulator transcription-factor signaling-pathway developmental-pathway gene-network epigenetic-modification DNA-methylation histone-modification chromatin-remodeling genomic-imprinting parental-effect maternal-effect paternal-effect cytoplasmic-inheritance mitochondrial-DNA chloroplast-DNA endosymbiont Wolbachia Rickettsia Spiroplasma Cardinium Arsenophonus Hamiltonella Regiella Serratia Buchnera Carsonella Portiera primary-endosymbiont secondary-endosymbiont facultative-symbiont obligate-symbiont mutualistic-symbiont parasitic-symbiont commensal-symbiont pathogen vector disease-transmission zoonosis emerging-infectious-disease biosecurity quarantine eradication control management conservation-biology restoration-ecology habitat-restoration river-restoration stream-restoration floodplain-restoration riparian-restoration wetland-restoration ecosystem-restoration passive-restoration active-restoration assisted-recovery managed-relocation 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ecosystem-service-valuation economic-valuation contingent-valuation choice-experiment benefit-transfer total-economic-value use-value non-use-value direct-use indirect-use option-value bequest-value existence-value willingness-to-pay willingness-to-accept cost-benefit-analysis cost-effectiveness-analysis multi-criteria-analysis decision-support uncertainty risk-assessment scenario-analysis sensitivity-analysis robustness-analysis adaptive-capacity resilience vulnerability exposure sensitivity coping-capacity transformation social-ecological-system complex-adaptive-system tipping-point regime-shift alternative-stable-state hysteresis path-dependence legacy-effect slow-variable fast-variable panarchy adaptive-cycle exploitation release reorganization foresight anticipatory-governance transformative-change sustainability-science future-earth interdisciplinary transdisciplinary problem-oriented solution-oriented action-oriented knowledge-co-production boundary-work boundary-object 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multilevel-model hierarchical-model Bayesian-hierarchical-model state-space-model hidden-Markov-model time-series-analysis spatial-analysis geostatistics kriging variogram spatial-autocorrelation Moran's-I Geary's-c Getis-Ord-Gi local-indicator-of-spatial-association LISA hot-spot cold-spot spatial-cluster spatial-outlier edge-effect boundary-effect modifiable-areal-unit-problem MAUP ecological-fallacy atomistic-fallacy cross-scale-inference upscaling downscaling scaling-law allometric-scaling fractal fractal-dimension self-similarity multi-fractal lacunarity roughness texture pattern process pattern-process-relationship neutral-model null-model randomization-test permutation-test bootstrap jackknife cross-validation model-averaging model-comparison information-criterion Bayesian-model-averaging ensemble-modeling consensus-model prediction forecast projection scenario narrative-scenario quantitative-scenario storyline shared-socioeconomic-pathway representative-concentration-pathway emission-scenario climate-scenario impact-scenario adaptation-scenario mitigation-scenario transformation-scenario backcasting forecasting trend-extrapolation expert-judgment Delphi-method participatory-modeling stakeholder-analysis influence-diagram conceptual-model mental-model system-archetype leverage-point intervention-point policy-instrument economic-instrument regulatory-instrument voluntary-instrument information-instrument market-based-instrument command-and-control-instrument subsidy tax charge fee levy deposit-refund tradable-permit emission-trading carbon-tax biodiversity-banking conditional-cash-transfer community-based-natural-resource-management joint-forest-management co-management adaptive-co-management transboundary-management international-regime global-governance multilateral-environmental-agreement convention treaty cartagena-protocol nagoya-protocol Aichi-target sustainable-development-goal SDG post-2020-global-biodiversity-framework Kunming-Montreal-global-biodiversity-framework 30-by-30-target nature-based-solution NbS ecosystem-based-adaptation ecosystem-based-mitigation green-infrastructure blue-infrastructure natural-infrastructure hybrid-infrastructure climate-smart-agriculture climate-smart-landscape climate-smart-development low-carbon-development green-economy circular-economy bioeconomy blue-economy sustainable-consumption sustainable-production life-cycle-thinking cradle-to-cradle industrial-ecology industrial-symbiosis urban-metabolism socio-ecological-metabolism material-intensity energy-intensity carbon-intensity decoupling absolute-decoupling relative-decoupling resource-efficiency eco-efficiency eco-innovation green-technology clean-technology environmental-technology appropriate-technology participatory-technology technology-transfer technology-diffusion innovation-system national-innovation-system sectoral-innovation-system technological-innovation-system socio-technical-system 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Williamsonia fletcheri
Ebony Boghaunter
Williamsonia fletcheri, commonly known as the ebony boghaunter, is a small dragonfly in the emerald family Corduliidae. It is one of North America's earliest-flying dragonflies, active in May and June. The species is strongly associated with acidic bog habitats and has a restricted range in the northeastern United States and southeastern Canada. It is considered rare across most of its range and is classified as endangered in Massachusetts.
dragonfly Odonata Corduliidae bog-specialist early-season rare endangered acidic-habitat North-America
Zale curema
Black-eyed Zale Moth, Northeastern Pine Zale, Black-eyed Zale
Zale curema is a moth in the family Erebidae, first described by John Bernhard Smith in 1908. The species is strongly associated with pitch pine forests in eastern North America. It has one generation per year, with larvae feeding preferentially on young needles of pitch pine. The species is listed as endangered in Connecticut but considered globally secure.
moth Erebidae pitch-pine univoltine endangered Connecticut forest-specialist North-America herbivore conifer-feeder