Semivoltine

Guides

Brachystola magna
Plains Lubber Grasshopper, Western Lubber Grasshopper, Homesteader
Brachystola magna is one of the largest grasshoppers in North America, a flightless species inhabiting the Great Plains from the Canadian border to central Mexico. Adults reach 4–6 cm in length, with females substantially larger and heavier than males. The species exhibits a two-year life cycle, with eggs overwintering twice before hatching. Though primarily herbivorous, adults display notable omnivory including scavenging and predation on other insects. The species has historical scientific significance as the subject of Walter Sutton's pioneering work on reduction division and chromosome theory.
grasshopper lubber flightless Great-Plains Romaleidae polyphagous omnivorous two-year-life-cycle semivoltine cytogenetics agricultural-pest intermediate-host thermoregulation
Ellipsoptera puritana
Puritan tiger beetle
Ellipsoptera puritana, the Puritan tiger beetle, is a federally threatened species endemic to a highly restricted range in the northeastern United States. Adults are medium-sized, fast-running predators with distinctive cream-white marginal markings on their elytra. The species occupies a narrow ecological niche: sandy beaches and riverbanks backed by eroding cliffs or bluffs, where larvae burrow into exposed sediment faces. Its two-year life cycle involves larval development in burrows followed by brief adult emergence in mid-summer. Habitat loss from shoreline development, dam construction, and bank stabilization has severely reduced populations, prompting intensive conservation efforts including captive rearing programs.
Coleoptera Cicindelidae tiger-beetle endangered-species coastal-ecology riparian-habitat North-America conservation sand-beach cliff-habitat semivoltine predator insect beetle
Flexamia huroni
Huron River Leafhopper
Flexamia huroni is a leafhopper species in the family Cicadellidae, described by Bess & Hamilton in 1999. It belongs to the genus Flexamia, a group of leafhoppers known for their specialized host plant associations with grasses. The species is named after the Huron River in Michigan, where it was first collected. Like other members of the genus, it likely exhibits strong ecological dependence on specific grass host plants.
leafhopper cicadellidae deltocephalinae paralimnini flexamia grass-specialist michigan-endemic auchenorrhyncha hemiptera insecta arthropoda animalia true-bug planthopper-relative 1999-description bess hamilton huroni huron-river usa north-america grassland-insect host-specific poorly-known rare uncommon data-deficient gbif catalogue-of-life ncbi inaturalist taxon species accepted hexapoda cicadomorpha clypeata membracoidea paralimnina flexamia-huroni bess-&-hamilton 1999 exact-match accepted-name canonical-name scientific-name authorship rank status matched taxonomy classification eukaryota metazoa distribution michigan observations 0 wikipedia none preferred-common-name huron-river-leafhopper true-bugs group kingdom phylum class order family genus authority ipt integrated-publishing-toolkit biodiversity-data-journal zookeys nature-conservation comparative-cytogenetics open-access open-access journal publication dataset specimen type nomenclatural-type herbarium university-of-granada spain fungi lichens agaricales cortinarius antonio-ortega mediterranean france italy image-collection colección-de-imágenes-de-los-tipos-nomenclaturales-de-hongos líquenes musgos-y-algas gda gdac vizoso quesada 2015 doi 10.3897 bdj 3 e5204 new-species new-jersey-pine-barrens muhlenbergia-torreyana pinebarren-smokegrass threatened-species andrew-hicks museum-of-natural-history university-of-colorado gerry-moore natural-resources-conservation-service greensboro nc uli-lorimer brooklyn-botanic-garden f.-whitcombi robert-whitcomb microbiology ornithology ecology host-plant warming-climate human-activities zookeys-511 69-79 zookeys.511.9572 roundworm nematode antarctic amblydorylaimus-isokaryoni pararhyssocolpus-paradoxus bulgaria scanning-electron-microscopy sem maritime-antarctic antarctic-islands lip-region spear vulva postembryonic-development molecular-analyses dorylaimida elshishka lazarova radoslavov hristov peneva 25-68 zookeys.511.9793 anidiv2 bulgarian-academy-of-sciences national-scientific-fund octocoral okinawa japan nanipora-kamurai living-fossil blue-coral heliopora aragonite-calcium-carbonate skeleton scleractinians soft-coral heliporacea lithotelestidae epiphaxum deep-sea shallow-coral-reef zamami-island national-park miyazaki reimer 1-23 zookeys.511.9432 non-biting-midge chironomus ch.-bernensis north-caucasus russia caucasian-populations europe siberia karyotype morphology mouthparts larvae chromosome genotypic-combinations mineralization eutrophication karmokov polukonova sinichkina tembotov-institute-of-ecology-of-mountain-territories saratov-state-medical-university comparative-cytogenetics-9 281-297 compcytogen.v9i3.4519 sea-turtle rescue-centre first-aid-station loggerhead green-turtle caretta-caretta chelonia-mydas bycatch mortality greece migration sexual-maturity ullmann stachowitsch uit-the-arctic-university-of-norway nature-conservation-10 45-69 natureconservation.10.4890 regional-activity-centre-for-specially-protected-areas porcupine coendou-ichillus lower-urubamba peru canopy-bridge pipeline natural-gas arboreal camera-trap dwarf-porcupine iquitos 770g gregory lunde zamora-meza carrasco-rueda repsol-exploración-perú zookeys-509 109-121 zookeys.509.9821 ant prionopelta madagascar seychelles subterranean leaf-litter dracula-ant hemolymph larval-hemolymph-feeding oophagy madagascar-biodiversity-center overson fisher zookeys-507 115-150 zookeys.507.9303 ito billen masuko spider anelosimus subsocial cobweb-spider theridiidae deforestation biodiversity-hotspot agnarsson university-of-vermont smithsonian-national-museum-of-natural-history wallace huxley buffon hooker lamarck darwin moramora eriophyoid-mite acari xinjiang china rosaceae paracolomerus gall ji-wei-li wang xue zhang zookeys-508 97-111 zookeys.508.8940 shihezi-university grasshopper wyoming melanoplus melanoplinae acrididae tetrigidae gomphocerniae oedipodinae cyrtacanthacridinae distribution-atlas field-guide wgis wyoming-grasshopper-information-system key capinera sechrist hebard helfer scudder blatchley thomas say harris degeer bruner saussure girard dodge walker fieber fabricius serville mcneill tinkham burmeister haldeman big-horn-mountains black-hills gladston indigens infantilis dodgei oregonensis marshalli yellowstone-national-park sagebrush pine elevation shortgrass-prairie mixedgrass-prairie forb grass economic-damage rangeland benefit overwinter egg hatch adult late-summer august october june life-cycle food-habit size collection survey underreported common endemic restricted-range forest-opening grassy moderate-elevation larger smaller east west united-states america north south central range extent limited restricted abundant population density occurrence presence absence habitat environment condition altitude topography terrain vegetation plant shrub tree forest opening meadow prairie steppe savanna woodland draw slope aspect soil substrate moisture temperature climate weather season phenology timing activity nymph emerge molt develop grow reproduce mate oviposit die generation voltinism univoltine bivoltine multivoltine semivoltine diapause aestivation hibernation dispersal movement behavior habit action feeding diet food host association relationship interaction specialist generalist monophagy oligophagy polyphagy herbivore detritivore predator parasitoid scavenger economic-importance pest beneficial neutral damage control management conservation threatened endangered vulnerable secure unknown global-biodiversity-information-facility es biodiversity-image-portal spanish-collections type-specimen lichen antarctica bernensis li yellowstone
Hylogomphus
clubtails
Hylogomphus is a genus of clubtail dragonflies (family Gomphidae) comprising six described species native to eastern North America. The genus was elevated from subgenus status within Gomphus in 2000, reflecting distinct morphological and phylogenetic characteristics. Species within this genus are associated with lotic freshwater habitats and exhibit conservative body plans typical of gomphid dragonflies.
dragonfly clubtail Gomphidae Odonata North-America freshwater bioindicator semivoltine lotic-habitat ecomorphological-variation
Hylogomphus geminatus
Twin-striped Clubtail
Hylogomphus geminatus, commonly known as the Twin-striped Clubtail, is a dragonfly species in the family Gomphidae endemic to the southeastern United States. The species was transferred from genus Gomphus to Hylogomphus in 2017 when Hylogomphus was elevated from subgenus to full genus rank. It exhibits pronounced ecomorphological variation in body length across its limited geographic range, with significant differences observed between populations on either side of the Apalachicola River in Florida. The species is listed as Least Concern by IUCN, though it was previously assessed as Near Threatened in 2007.
dragonfly clubtail bioindicator endemic southeastern-US Apalachicola-River ecomorphology semivoltine
Oberea tripunctata
dogwood twig borer
Oberea tripunctata, commonly known as the dogwood twig borer, is a species of longhorn beetle in the family Cerambycidae. Adults emerge in early June and feed on dogwood twigs, causing girdling damage. Females lay eggs singly on healthy twigs; larvae bore into twigs and overwinter in stems, with some individuals taking up to two years to complete development. The species is considered a minor pest of flowering dogwood and has been recorded from additional hosts including elm, azalea, viburnum, and various fruit trees.
Cerambycidae Lamiinae Obereini longhorn-beetle pest borer dogwood Cornus North-America twig-borer minor-pest woody-plant-feeder overwintering-larvae univoltine semivoltine
Oreoneta
A genus of dwarf spiders (family Linyphiidae) established in 1894, comprising approximately 30 species distributed across cold-temperate and Arctic regions of the Northern Hemisphere. Most species inhabit high-latitude or high-altitude environments. The genus was revised extensively by Saaristo & Marusik in 2004, resulting in numerous new species descriptions.
Arachnida Araneae Linyphiidae dwarf-spider Arctic boreal Holarctic tundra montane predator cold-adapted ground-dwelling high-latitude high-altitude Saaristo Marusik Chyzer Kulczyński 1894 2004-revision genitalic-characters epigynum embolus conductor palaearctic nearctic Eurasia North-America Scandinavia Siberia Alaska Canada Greenland Kazakhstan Mongolia China Kurile-Islands Finland Sweden Norway Germany Switzerland Italy Slovakia Bulgaria Central-Europe type-species:-Oreoneta-frigida thirty-species 30-species small-spider sheet-web-weaver money-spider Linyphiinae Erigoninae microhabitat litter-dwelling cryptic understudied taxonomic-revision morphological-taxonomy light-microscopy species-description biogeography glacial-refugia post-glacial-colonization endemic regional-endemic alpine-endemic Arctic-endemic boreal-forest taiga rocky-habitat meadow wetland stream-margin snowbed fellfield heath moss lichen vegetation soil-surface stone rock boulder scree talus debris humus organic-layer permafrost polar subpolar cold-stenotherm stenotherm stenotopic habitat-specialist conservation rare data-deficient IUCN red-list natural-history systematics phylogeny molecular-systematics DNA-barcoding COI cytochrome-oxidase 28S 18S ITS morphology taxonomy alpha-taxonomy revision monograph faunistics chorology zoogeography dispersal vicariance speciation cryptic-species species-complex species-group species-aggregate subspecies infraspecific variation polymorphism sexual-dimorphism male female juvenile instar ecdysis molting development growth reproduction mating courtship sperm-transfer spermatophore pedipalp palpal-organ copulatory-organ genitalia epigyne vulva spermatheca fertilization-duct copulatory-duct atrium scape maceration dissection microscope stereomicroscope compound-microscope illustration drawing photography SEM scanning-electron-microscopy habitus dorsal ventral lateral prosoma opisthosoma abdomen cephalothorax carapace sternum chelicera fang leg tarsus metatarsus tibia patella femur coxa trochanter spinneret colulus tracheal-spiracle book-lung cuticle seta hair spine macroseta sensillum chaetotaxy eye oculus ocelli anterior-median-eye posterior-median-eye anterior-lateral-eye posterior-lateral-eye eye-tubercle clypeus fovea pedicel waist constriction color pattern pigmentation melanism albinism size measurement total-length carapace-length carapace-width abdomen-length leg-length leg-formula index ratio proportion allometry ontogeny subadult adult maturity sexual-maturity reproductive-maturity longevity life-span generation-time voltinism univoltine semivoltine biennial perennial overwintering diapause quiescence cold-hardiness freeze-tolerance freeze-avoidance supercooling cryoprotectant glycerol sorbitol trehalose thermal-biology thermoregulation behavioral-thermoregulation postural-thermoregulation activity diurnal nocturnal crepuscular matutinal vespertine circadian rhythm seasonal phenology calendar date month activity-period breeding-season oviposition egg egg-sac cocoon silk web sheet-web tangle-web space-web platform-web dome-web bowl-web funnel retreat signal-line dragline attachment-disc ampullate-gland cylindriform-gland aciniform-gland aggregate-gland flagelliform-gland tubuliform-gland piriform-gland major-ampullate minor-ampullate silk-protein spidroin fibroin sericin glue viscid dry capture-silk dragline-silk safety-line anchor-line frame-line radial-line spiral-line sticky-spiral non-sticky-spiral hub perimeter substrate niche guild functional-group trophic-level carnivore zoophage arthropodivore insectivore prey capture attack subdue immobilize kill feed digest extraoral-digestion sucking-stomach gut midgut hindgut Malpighian-tubule excretion osmoregulation metabolism respiration trachea spiracle gas-exchange circulation hemolymph heart aorta sinus nervous-system brain ganglion nerve sensory-organ mechanoreceptor chemoreceptor thermoreceptor hygroreceptor photoreceptor proprioceptor trichobothrium tarsal-organ slit-sensillum lyriform-organ pedipalp-organ Haller's-organ contact-chemoreceptor gustatory olfactory pheromone semiochemical kairomone allomone synomone communication signal vibration seismic airborne visual chemical tactile copulation insemination sperm spermatozoon median-apophysis terminal-apophysis paracymbium cymbium bulbus tegulum subtegulum radix lamella process apophysis flange hook coil spiral duct reservoir sclerite plate field fossa pocket receptaculum spermathecae gland insemination-duct uterus uterus-externus uterus-internus vagina gonopore ovipositor ovary oviduct follicle vitellogenesis oogenesis spermatogenesis testis vas-deferens ejaculatory-duct seminal-vesicle fecundity fertility clutch-size egg-number hatching emergence ballooning aerial-dispersal silk-dragline gossamer migration movement walking running jumping climbing burrowing hiding refuge shelter defense predation predator-avoidance antipredator crypsis camouflage mimicry aposematism deimatic thanatosis autotomy leg-loss regeneration injury wound healing immune-response hemocyte melanization encapsulation nodule parasite parasitoid pathogen fungus bacterium virus nematode mite grooming cleaning hygiene self-maintenance silk-use web-use prey-capture foraging hunting ambush sit-and-wait active-hunting searching probing testing exploration learning memory cognition behavior ethology ecology population community ecosystem biome biosphere habitat environment abiotic biotic factor temperature humidity moisture precipitation rain snow wind light photoperiod day-length radiation UV soil cover complexity heterogeneity productivity energy nutrient carbon nitrogen phosphorus food-web trophic-interaction predator-prey competition intraspecific interspecific resource partitioning coexistence diversity richness evenness abundance density frequency occurrence presence absence detection sampling collection pitfall-trap quadrat transect hand-collecting beating sifting extraction Berlese Tullgren Winkler flotation sorting preservation ethanol formalin mount slide label catalog database repository museum voucher type holotype paratype syntype lectotype paralectotype neotype topotype original data locality georeference coordinate latitude longitude elevation altitude depth collector determination identification expert specialist literature publication journal article paper book thesis dissertation report checklist online digital archive bibliography reference citation author year volume page figure map table appendix supplementary material methods results discussion conclusion abstract keyword index-term subject topic discipline science biology zoology arachnology entomology evolution phylogenetics biological nature protection management policy legislation restoration monitoring assessment indicator flagship umbrella keystone engineer service function dynamics stability resilience resistance recovery succession disturbance climate change warming glacier range shift expansion contraction fragmentation isolation connectivity corridor network reserve park protected area wilderness urban rural agricultural forestry mining impact threat risk vulnerability sensitivity exposure adaptive capacity mitigation adaptation sustainable use harvest trade economy medicine research education outreach citizen engagement awareness appreciation aesthetic cultural spiritual existence value intrinsic instrumental utilitarian anthropocentric biocentric ecocentric ethic philosophy history discovery expedition voyage survey inventory fauna biota biodiversity life earth planet wildlife animal invertebrate arthropod chelicerate arachnid spider araneomorph entelegyne micryphantid Oreoneta 2004 frigida type-species alpine feeding type-specimen climate-change protected-area
Peltoperla
Peltoperla is a genus of stoneflies in the family Peltoperlidae, found in Appalachian headwater streams of eastern North America. Species in this genus have semivoltine life cycles, typically developing over two years with egg diapause periods of approximately six months. Nymphs are strongly associated with leaf pack habitats in small forested streams. The genus exhibits 'leaky' cohort dynamics, where some individuals complete development in one year while others take two years, resulting in overlapping generations and high gene flow among cohorts.
Plecoptera stonefly detritivore headwater-streams Appalachia leaf-packs semivoltine
Peltoperlidae
roach-like stoneflies, roachflies
Peltoperlidae is a family of stoneflies comprising 11 genera and approximately 46 known species. Larvae are dorsoventrally flattened with expanded thoracic plates, giving them a distinctive roach-like appearance. The family exhibits semivoltine life cycles lasting one to two years, with adult emergence typically occurring from April through June. Members are classified as shredder-detritivores, playing a significant role in leaf litter decomposition in lotic systems.
Plecoptera stoneflies aquatic-insects shredders bioindicators drumming-behavior lotic-systems leaf-litter-processing semivoltine Appalachian-streams Cretaceous-fossil-record
Quesada gigas
Giant cicada, chichara grande, coyoyo, coyuyo, coffee cicada
Quesada gigas is a large cicada species native to North, Central, and South America, with the widest geographic range of any cicada in the Western Hemisphere. It is a significant agricultural pest, particularly of coffee (Coffea) in Brazil and paricá (Schizolobium parahyba var. amazonicum) in the Amazon region, where nymphal feeding on roots can cause substantial yield losses or tree mortality. The species exhibits low selectivity in oviposition site choice, with females documented laying eggs in dry branches of non-host plants such as Conyza spp. weeds, which cannot support complete nymphal development due to their annual life cycle.
agricultural-pest coffee-pest semivoltine root-feeder subterranean-nymph gregarious low-oviposition-selectivity climate-change-model Neotropical wide-distribution
Recurvaria
Recurvaria is a genus of small moths in the family Gelechiidae, established by Haworth in 1828. The genus contains approximately 40 described species distributed across the Holarctic region. Several species are economically significant as pests of coniferous trees and pistachio, with larvae that mine needles or feed internally within fruits. The genus exhibits diverse life history strategies, including univoltine and bivoltine cycles, with some species requiring two years to complete development.
Gelechiidae Litini needle-miner pistachio-pest forest-pest Holarctic internal-feeder diapause multivoltine semivoltine