Species-group

Guides

• Ixodes affinis

  Ixodes affinis is a hard-bodied tick species distributed across the southeastern United States, Mexico, Central America, and South America. Unlike its congener Ixodes scapularis, this species does not bite humans but serves as a competent vector for Borrelia burgdorferi sensu stricto, the causative agent of Lyme disease. Immature stages feed on birds and small mammals, while adults parasitize larger mammals including deer and dogs. Its expanding range into southeastern Virginia and association with migratory birds has raised concerns about its role in pathogen dispersal.

  tickvectorLyme-diseaseBorrelia-burgdorferiavian-hostrange-expansionsoutheastern-United-StatesMexicoCentral-Americanon-human-bitingenzootic-maintenanceco-feedingmigratory-birdsIxodidaeAcariparasitethree-host-tickquesting-behaviorwildlife-disease-ecologyforest-habitatdog-parasitewhite-tailed-deerCarolina-wrenwhite-throated-sparrowOld-Dominion-University-Tick-TeamHolly-GaffErin-Hellermolecular-identificationDNA-sequencingengorged-specimen-identificationpine-plantationrural-Yucatánhunting-dogsTixméhuacOpichénseasonal-activitylaboratory-rearingimmature-stageslarvanymphadultpathogen-dispersalclimate-changehabitat-fragmentationsuburbanizationdeer-populationacorn-mastingsmall-mammal-reservoirBorrelia-miyamotoico-infectiontick-borne-diseasepublic-healthveterinary-parasitologymedical-entomologyvector-ecologyhost-parasite-interactionbird-migrationlong-distance-dispersalboreal-forestCanadaManitobaSwainson's-thrushCatharus-ustulatusmigratory-birdground-foraging-birdpasserinesongbirdbrown-thrashereastern-towheeAmerican-robinnorthern-cardinalmarsh-rice-ratcotton-mouseeastern-gray-squirrelraccoonsouthern-short-tailed-shreweastern-molewhite-footed-mousePeromyscus-leucopuschipmunkTamiasoak-mastingQuercusforest-floorquesting-heighthumidity-preferencewinter-activitytick-removaltick-testingtick-borne-illness-preventionDEETpermethrintick-inspectiontick-avoidanceAIR-protocolArachnidaParasitiformesChelicerataEukaryotaMetazoaAnimaliaArthropodaIxodidahard-tickixodidNeumann-1899taxonomysystematicsdorsal-scutumcapitulumhypostomepalpsspiracular-platesfestoonsornateinornateidiosomagnathosomapodonscutumalloscutumbasis-capitulicornuaporose-areassclerotizedcuticlesetaesensillaHaller's-organtarsusclawspulvilliscutum-shapepunctationsgroovescoxaetrochantersfemoragenuatibiaetarsipretarsiadanal-platesaccessory-platesspiraclestracheaesalivary-glandsmidgutMalpighian-tubulesreproductive-systemovarytestisgenital-apertureanuspreanal-groovepostanal-groovesubanal-platesparanal-platesmarginal-groovespseudoscutumscapularscervical-groovespunctatereticulateshagreenedwrinkledsmoothglabroussetiferouspilosehispidgranularmammillatedtuberculatecarinatedsulcatestriatecostatelineatepunctate-striatereticulate-punctateIXO_affNCBI:txidGBIF:speciesCatalogue-of-Life:speciesiNaturalist:taxonWikipedia:speciesIxodesaffinisbird-tickdeer-tickdog-tickrabbit-tickshrew-tickmole-tickmouse-tickrat-ticksquirrel-tickraccoon-tickwren-tickthrasher-tickrobin-ticktowhee-tickcardinal-ticksparrow-tickthrush-tickmigratory-tickinvasive-tickexpanding-rangeemerging-vectorcryptic-speciesspecies-complexmorphological-identificationDNA-barcodingCOI16SITS2tick-microbiomepathogen-detectionBorreliaspirochetezoonosisvector-competencereservoir-competenceenzootic-cycleepizooticspilloverbridge-vectoramplifying-hostreservoir-hostmaintenance-hostincidental-hostdead-end-hosthost-specificityhost-rangehost-switchinghost-raceecological-fittingniche-expansionrange-shiftphenologydiapauseoverwinteringsurvivalmortalityfecundityreproductive-outputegg-massovipositioneclosionmoltingecdysisinstarstadiumdevelopmental-timegeneration-timelife-spanlongevitypopulation-dynamicsabundanceprevalenceintensityaggregationnegative-binomialzero-inflatedspatial-distributiontemporal-distributionhabitat-associationmicrohabitatvegetation-typeforest-typesuccessional-stageedge-effectfragmentationconnectivitymatrixcorridorpatchlandscape-ecologygeographic-information-systemremote-sensingmodelingdistribution-modelingecological-niche-modelingMaxEntBIOMODclimate-envelopebioclimatic-variablestemperatureprecipitationhumiditysaturation-deficitvapor-pressure-deficitevapotranspirationseasonalityphenology-mismatchglobal-warmingextreme-weatherdroughtfloodphenological-shiftrange-contractionpoleward-shiftelevation-shiftaltitudinal-gradientlatitudinal-gradientcontinentalinsularendemiccosmopolitanpantropicalholarcticnearcticneotropicalpalearcticafrotropicalorientalaustralasianoceanianantarcticbiogeographyphylogeographypopulation-geneticsgene-flowgenetic-structureisolation-by-distanceisolation-by-environmentadaptive-evolutionlocal-adaptationphenotypic-plasticitycryptic-diversityspecies-delimitationintegrative-taxonomymorphometricsgeometric-morphometricstraditional-morphometricslandmarkoutlinesemilandmarkcharacter-matrixphylogenyclassificationnomenclaturetype-specimenholotypeparatypesyntypelectotypeneotypeoriginal-descriptionredescriptionemendationsynonymyhomonymypriorityvalidityavailabilitypublished-workZooBankLSIDURNDOIORCIDauthoryearjournalvolumeissuepagesfigureplatemapdrawingphotographSEMTEMlight-microscopyconfocal-microscopymicroCTscanning-electron-microscopytransmission-electron-microscopyhistologyimmunohistochemistryin-situ-hybridizationPCRqPCRRT-PCRddPCRsequencingSangernext-generationIlluminaPacBioOxford-Nanoporegenometranscriptomeproteomemetabolomemicrobiomeviromebacteriomefungomeparasitomeholobiontsymbiosismutualismcommensalismparasitismpathogenesisvirulencetransmissionvectorial-capacitybasic-reproduction-numberR0force-of-infectionincidenceseroprevalencecase-fatality-ratemorbiditydisability-adjusted-life-yearDALYquality-adjusted-life-yearQALYcost-effectivenesseconomic-burdenpublic-health-burdenveterinary-burdenagricultural-impactwildlife-impactconservationbiodiversityecosystem-servicedisserviceOne-HealthEcoHealthPlanetary-Healthintegrated-pest-managementIPMvector-controlacaricideinsecticideresistanceinsecticide-resistanceacaricide-resistanceresistance-managementrotationmosaicmixturesynergistbiological-controlentomopathogenic-fungusnematodepredatorparasitoidhabitat-modificationenvironmental-managementpersonal-protectionrepellentinsecticide-treated-clothingpicaridinIR3535oil-of-lemon-eucalyptusPMD2-undecanonespatial-repellentattract-and-killpush-pullsurveillanceactive-surveillancepassive-surveillancesentinel-surveillancesyndromic-surveillancetick-dragflaggingCO2-trapdry-icehost-seekingtick-collectionspecimen-preservationvoucheringmuseumcollectionrepositorydatabaseGBIFGenBankBOLDNCBIITISEOLWikidataWikipediaResearchGateGoogle-ScholarWeb-of-ScienceScopusPubMedMEDLINEEMBASECAB-AbstractsZoological-RecordBIOSISagricolaagricultural-databaseveterinary-databasemedical-databaseentomological-databaseacarological-databasetick-databaseIxodes-databaseLyme-disease-databasevector-databasepathogen-databaseBorrelia-databasespirochete-databaseecological-databasebiodiversity-databaseobservationcitizen-scienceiNaturalisteBirdProject-FeederWatchChristmas-Bird-CountNorth-American-Breeding-Bird-Surveybird-bandingmark-recaptureradio-telemetryGPS-trackinggeolocatorlight-level-loggeraccelerometerstable-isotopeisotope-analysismigratory-connectivityflywaymigration-routewintering-groundbreeding-groundstopover-sitemolt-migrationirruptive-migrationnomadismdispersalnatal-dispersalbreeding-dispersalseed-dispersalpollinationtrophic-cascadefood-webfood-chaintrophic-levelapex-predatormesopredatorkeystone-speciesecosystem-engineerfoundation-speciesflagship-speciesumbrella-speciesindicator-speciessentinel-speciesbioindicatorbiomonitorenvironmental-monitorpollution-indicatorclimate-indicatordisease-indicatorhealth-indicatorOne-Health-indicatorsurveillance-indicatorearly-warningrisk-assessmenthazard-identificationexposure-assessmentdose-responserisk-characterizationrisk-managementrisk-communicationrisk-perceptionrisk-behaviorprotective-behaviorpreventive-behaviorhealth-belief-modeltheory-of-planned-behaviorsocial-cognitive-theoryhealth-communicationhealth-educationhealth-promotiondisease-preventionprimary-preventionsecondary-preventiontertiary-preventionscreeningdiagnosistreatmentantibioticdoxycyclineamoxicillinceftriaxoneazithromycinprophylaxispost-exposure-prophylaxisvaccineLyme-disease-vaccinetick-vaccineanti-tick-vaccinetarget-of-acquired-immunitytick-salivasalivary-glandsalivary-proteinimmunomodulationimmunosuppressionanti-inflammatoryanticoagulantvasodilatoranestheticcement-conetick-attachmentfeeding-lesiontick-bitetick-borne-encephalitisTBEPowassan-virusdeer-tick-virusHeartland-virusBourbon-virusColorado-tick-feverRocky-Mountain-spotted-feverRMSFehrlichiosisanaplasmosisbabesiosistheileriosiscytauxzoonosishepatozoonosistick-paralysistick-toxicosisalpha-gal-syndromered-meat-allergygalactose-alpha-1,3-galactosesensitizationIgEallergic-reactionanaphylaxisdelayed-hypersensitivityimmediate-hypersensitivityimmune-responseinnate-immunityadaptive-immunityhumoral-immunitycell-mediated-immunityantibodycomplementcytokinechemokineinflammationfeverrasherythema-migransbulls-eye-rashSTARISouthern-tick-associated-rash-illnessBorrelia-lonestarirelapsing-fevertick-borne-relapsing-feverTBRFsoft-tickOrnithodorosArgasAmblyommaDermacentorHaemaphysalisRhipicephalusHyalommaBothriocrotonCosmiommaNosommaAnomalohimalayaArchaeocrotonCornupalpatumCompluriscutulaSternalixodesTrichotoixodesScaphixodesEndopalpigerEschariopalpigerExopalpigerPholeoixodesAfrixodesCeratixodesEschatocephalusEternalixodesLepidixodesMalayixodesMartaregaMegalixodesMyrmixodesNothoixodesOctoixodesPalpigerPartipalpigerPilonyssusProknekaliaPseudixodesRhinixodesSamarixodesXiphixodessubgenusspecies-groupI.-ricinus-complexI.-persulcatus-complexI.-scapularis-complexI.-pacificus-complexI.-affinis-groupI.-cookei-groupI.-marxi-groupI.-angustus-groupI.-soricis-groupI.-uriae-groupI.-holocyclus-groupI.-ceylonensis-groupI.-granulatus-groupI.-ovatus-groupI.-nipponensis-groupI.-persulcatus-groupI.-pavlovskyi-groupI.-ricinus-groupI.-scapularis-groupI.-pacificus-groupI.-affinisI.-auritulusI.-australiensisI.-boliviensisI.-brunneusI.-cookeiI.-cooleyiI.-cornuaeI.-cornuatusI.-crenulatusI.-damminiI.-dawsoniI.-dentatusI.-diomedeaeI.-domergueiI.-eadsiI.-eastoniI.-eldaricusI.-elongatusI.-eudyptidisI.-euplectropiI.-evansiI.-fecialisI.-festaiI.-frontalisI.-galapagoensisI.-ghilaroviI.-gibbosusI.-granulatusI.-gregsoniI.-hearleiI.-hexagonusI.-hirstiI.-holocyclusI.-hoogstraaliI.-howelliI.-hyattiI.-jacksoniI.-jellisoniI.-kashmiricusI.-kazakstaniI.-kingiI.-kohlsiI.-kopsteiniI.-kuntziI.-laguriI.-latusI.-layingensisI.-lewisiI.-lividusI.-longiscutatusI.-loricatusI.-luciaeI.-malayensisI.-marxiI.-masloviI.-matopiI.-minorI.-monospinosusI.-moreliI.-moschiferiI.-murisI.-murreletiI.-myospalacisI.-nchisiensisI.-necromysI.-neitziI.-nipponensisI.-nuttalliI.-ochotonaeI.-okapiaeI.-oldiI.-ornithorhynchiI.-ovatusI.-pacificusI.-pararicinusI.-pavlovskyiI.-percavatusI.-peromysciI.-persulcatusI.-pomerantziI.-pterodromaeI.-quadratusI.-raccoonisI.-rageauiI.-randrianasoloiI.-rasusI.-redikorzeviI.-ricinusI.-rothschildiI.-rotundatusI.-rubicundusI.-rugicollisI.-rugosusI.-sapeloensisI.-sarsiI.-scapularisI.-schillingsiI.-sculptusI.-semenoviI.-soricisI.-spinaeI.-steiniI.-stilesiI.-stromiI.-subterranusI.-tancitariusI.-tanukiI.-tapirusI.-tasmaniI.-tayloriI.-tertiariusI.-texanusI.-theileraeI.-thomasaeI.-tovariI.-transvaalensisI.-triangulicepsI.-trichosuriI.-turdusI.-uriaeI.-vandenbergiI.-vanidicusI.-ventalloiI.-vespertilionisI.-victoriensisI.-walkeraeI.-werneriI.-woodiI.-zaglossiI.-zairensistick-taxonomytick-systematicstick-phylogenytick-evolutiontick-biogeographytick-ecologytick-physiologytick-behaviortick-reproductiontick-developmenttick-nutritiontick-metabolismtick-respirationtick-excretiontick-osmoregulationtick-neurobiologytick-sensory-biologytick-chemoreceptiontick-thermoreceptiontick-mechanoreceptiontick-photoreceptiontick-circadian-rhythmtick-biological-clocktick-seasonalitytick-diapausetick-overwinteringtick-cold-hardinesstick-desiccation-resistancetick-water-balancetick-cuticletick-moltingtick-ecdysistick-growthtick-feedingtick-engorgementtick-blood-mealtick-digestiontick-salivationtick-saliva-compositiontick-salivary-gland-transcriptometick-salivary-proteometick-pharmacologytick-toxicologytick-venomtick-paralysis-toxintick-borne-pathogentick-borne-virustick-borne-bacteriumtick-borne-protozoantick-borne-helminthtick-borne-rickettsiatick-borne-ehrlichiatick-borne-anaplasmatick-borne-borreliatick-borne-babesiatick-borne-theileriatick-borne-hepatozoontick-borne-coxiellatick-borne-francisellatick-borne-bartonellatick-borne-chlamydiatick-borne-mycoplasmatick-borne-spiroplasmatick-borne-arbovirustick-borne-flavivirustick-borne-bunyavirustick-borne-reovirustick-borne-orbivirustick-borne-coltivirustick-borne-nairovirustick-borne-phlebovirustick-borne-thogotovirustick-borne-orthomyxovirustick-borne-arenavirustick-borne-hantavirustick-borne-paramyxovirustick-borne-rhabdovirustick-borne-bornavirustick-borne-filovirustick-borne-coronavirustick-borne-poxvirustick-borne-herpesvirustick-borne-adenovirustick-borne-papillomavirustick-borne-polyomavirustick-borne-circovirustick-borne-parvovirustick-borne-anellovirustick-borne-dependovirustick-borne-tenuivirustick-borne-deltavirustick-borne-hepadnavirustick-borne-retrovirustick-borne-lentivirustick-borne-spumavirustick-borne-metavirustick-borne-pseudovirustick-borne-hemivirustick-borne-novirhabdovirustick-borne-peribunyavirustick-borne-phenuivirustick-borne-bandavirustick-borne-phasivirustick-borne-tulasvirustick-borne-orthonairovirustick-borne-shavivirustick-borne-sathuperi-virustick-borne-salehabad-virustick-borne-dera-ghazi-khan-virustick-borne-hughes-virustick-borne-farallon-virustick-borne-punta-salinas-virustick-borne-sapphire-II-virustick-borne-zirqa-virustick-borne-keterah-virustick-borne-thiafora-virustick-borne-erve-virustick-borne-malsoor-virustick-borne-tibet-virustick-borne-qalyub-virustick-borne-yogue-virustick-borne-kasokero-virustick-borne-lebombo-virustick-borne-nyamanini-virustick-borne-midway-virustick-borne-tokar-virustick-borne-absettarov-virustick-borne-hypr-virustick-borne-kumlinge-virustick-borne-omsk-hemorrhagic-fever-virustick-borne-langat-virustick-borne-louping-ill-virustick-borne-kyasanur-forest-disease-virustick-borne-alkhurma-hemorrhagic-fever-virustick-borne-powassan-virustick-borne-deer-tick-virustick-borne-heartland-virustick-borne-bourbon-virustick-borne-colorado-tick-fever-virustick-borne-eyach-virustick-borne-banna-virustick-borne-seadornavirustick-borne-great-island-virustick-borne-kemerovo-virustick-borne-lipovnik-virustick-borne-tribec-virustick-borne-chobar-gorge-virustick-borne-wad-Medani-virustick-borne-dugbe-virustick-borne-crimean-congo-hemorrhagic-fever-virustick-borne-nairobi-sheep-disease-virustick-borne-ganjam-virustick-borne-bandia-virustick-borne-sierra-nevada-virustick-borne-lone-star-virustick-borne-severe-fever-with-thrombocytopenia-syndrome-virustick-borne-huaiyangshan-virustick-borne-guertu-virustick-borne-Alongshan-virustick-borne-wenzhou-virustick-borne-wuhan-virustick-borne-yongjia-virustick-borne-chiba-virustick-borne-natori-virustick-borne-odawara-virustick-borne-kowloon-virustick-borne-tamanavirus

• Melanoplus dodgei group

  The Melanoplus dodgei-species-group is a taxonomic grouping within the genus Melanoplus (spurthroated grasshoppers). Species groups in Melanoplus represent clusters of morphologically similar species that share diagnostic characteristics, particularly in male genitalia and pronotal structure. The group is part of the diverse western North American Melanoplus fauna documented in regional field guides and distribution atlases. Individual species within this group are recorded from Wyoming and Colorado, with M. dodgei itself being a documented western species.

  grasshopperspurthroatedwestern-North-Americaspecies-groupMelanoplinae

• Melanoplus viridipes group

  The Melanoplus viridipes-species-group is a species group within the genus Melanoplus (subfamily Melanoplinae), a diverse genus of spur-throated grasshoppers found in North America. Species groups in Melanoplus represent informal taxonomic aggregations of morphologically similar species that share diagnostic characteristics. Members of this group are characterized by green legs (viridipes = "green-footed"), a trait that distinguishes them from related species groups within the genus. The group occurs within the western and central United States, with species typically inhabiting grassland ecosystems. Like other Melanoplus species, members are medium-sized grasshoppers with a distinct conical spur between the front legs, perpendicular facial orientation, and strong flight capabilities.

  Melanoplusspecies-groupviridipesspur-throated-grasshopperAcrididaeMelanoplinaeNorth-Americagrasslandtaxonomymorphological-group

• Oreoneta

  A genus of dwarf spiders (family Linyphiidae) established in 1894, comprising approximately 30 species distributed across cold-temperate and Arctic regions of the Northern Hemisphere. Most species inhabit high-latitude or high-altitude environments. The genus was revised extensively by Saaristo & Marusik in 2004, resulting in numerous new species descriptions.

  ArachnidaAraneaeLinyphiidaedwarf-spiderArcticborealHolarctictundramontanepredatorcold-adaptedground-dwellinghigh-latitudehigh-altitudeSaaristoMarusikChyzerKulczyński18942004-revisiongenitalic-charactersepigynumembolusconductorpalaearcticnearcticEurasiaNorth-AmericaScandinaviaSiberiaAlaskaCanadaGreenlandKazakhstanMongoliaChinaKurile-IslandsFinlandSwedenNorwayGermanySwitzerlandItalySlovakiaBulgariaCentral-Europetype-species:-Oreoneta-frigidathirty-species30-speciessmall-spidersheet-web-weavermoney-spiderLinyphiinaeErigoninaemicrohabitatlitter-dwellingcrypticunderstudiedtaxonomic-revisionmorphological-taxonomylight-microscopyspecies-descriptionbiogeographyglacial-refugiapost-glacial-colonizationendemicregional-endemicalpine-endemicArctic-endemicboreal-foresttaigarocky-habitatmeadowwetlandstream-marginsnowbedfellfieldheathmosslichenvegetationsoil-surfacestonerockboulderscreetalusdebrishumusorganic-layerpermafrostpolarsubpolarcold-stenothermstenothermstenotopichabitat-specialistconservationraredata-deficientIUCNred-listnatural-historysystematicsphylogenymolecular-systematicsDNA-barcodingCOIcytochrome-oxidase28S18SITSmorphologytaxonomyalpha-taxonomyrevisionmonographfaunisticschorologyzoogeographydispersalvicariancespeciationcryptic-speciesspecies-complexspecies-groupspecies-aggregatesubspeciesinfraspecificvariationpolymorphismsexual-dimorphismmalefemalejuvenileinstarecdysismoltingdevelopmentgrowthreproductionmatingcourtshipsperm-transferspermatophorepedipalppalpal-organcopulatory-organgenitaliaepigynevulvaspermathecafertilization-ductcopulatory-ductatriumscapemacerationdissectionmicroscopestereomicroscopecompound-microscopeillustrationdrawingphotographySEMscanning-electron-microscopyhabitusdorsalventrallateralprosomaopisthosomaabdomencephalothoraxcarapacesternumchelicerafanglegtarsusmetatarsustibiapatellafemurcoxatrochanterspinneretcolulustracheal-spiraclebook-lungcuticlesetahairspinemacrosetasensillumchaetotaxyeyeoculusocellianterior-median-eyeposterior-median-eyeanterior-lateral-eyeposterior-lateral-eyeeye-tubercleclypeusfoveapedicelwaistconstrictioncolorpatternpigmentationmelanismalbinismsizemeasurementtotal-lengthcarapace-lengthcarapace-widthabdomen-lengthleg-lengthleg-formulaindexratioproportionallometryontogenysubadultadultmaturitysexual-maturityreproductive-maturitylongevitylife-spangeneration-timevoltinismunivoltinesemivoltinebiennialperennialoverwinteringdiapausequiescencecold-hardinessfreeze-tolerancefreeze-avoidancesupercoolingcryoprotectantglycerolsorbitoltrehalosethermal-biologythermoregulationbehavioral-thermoregulationpostural-thermoregulationactivitydiurnalnocturnalcrepuscularmatutinalvespertinecircadianrhythmseasonalphenologycalendardatemonthactivity-periodbreeding-seasonovipositioneggegg-saccocoonsilkwebsheet-webtangle-webspace-webplatform-webdome-webbowl-webfunnelretreatsignal-linedraglineattachment-discampullate-glandcylindriform-glandaciniform-glandaggregate-glandflagelliform-glandtubuliform-glandpiriform-glandmajor-ampullateminor-ampullatesilk-proteinspidroinfibroinsericingluevisciddrycapture-silkdragline-silksafety-lineanchor-lineframe-lineradial-linespiral-linesticky-spiralnon-sticky-spiralhubperimetersubstratenicheguildfunctional-grouptrophic-levelcarnivorezoophagearthropodivoreinsectivorepreycaptureattacksubdueimmobilizekillfeeddigestextraoral-digestionsucking-stomachgutmidguthindgutMalpighian-tubuleexcretionosmoregulationmetabolismrespirationtracheaspiraclegas-exchangecirculationhemolymphheartaortasinusnervous-systembrainganglionnervesensory-organmechanoreceptorchemoreceptorthermoreceptorhygroreceptorphotoreceptorproprioceptortrichobothriumtarsal-organslit-sensillumlyriform-organpedipalp-organHaller's-organcontact-chemoreceptorgustatoryolfactorypheromonesemiochemicalkairomoneallomonesynomonecommunicationsignalvibrationseismicairbornevisualchemicaltactilecopulationinseminationspermspermatozoonmedian-apophysisterminal-apophysisparacymbiumcymbiumbulbustegulumsubtegulumradixlamellaprocessapophysisflangehookcoilspiralductreservoirscleriteplatefieldfossapocketreceptaculumspermathecaeglandinsemination-ductuterusuterus-externusuterus-internusvaginagonoporeovipositorovaryoviductfolliclevitellogenesisoogenesisspermatogenesistestisvas-deferensejaculatory-ductseminal-vesiclefecundityfertilityclutch-sizeegg-numberhatchingemergenceballooningaerial-dispersalsilk-draglinegossamermigrationmovementwalkingrunningjumpingclimbingburrowinghidingrefugeshelterdefensepredationpredator-avoidanceantipredatorcrypsiscamouflagemimicryaposematismdeimaticthanatosisautotomyleg-lossregenerationinjurywoundhealingimmune-responsehemocytemelanizationencapsulationnoduleparasiteparasitoidpathogenfungusbacteriumvirusnematodemitegroomingcleaninghygieneself-maintenancesilk-useweb-useprey-captureforaginghuntingambushsit-and-waitactive-huntingsearchingprobingtestingexplorationlearningmemorycognitionbehaviorethologyecologypopulationcommunityecosystembiomebiospherehabitatenvironmentabioticbioticfactortemperaturehumiditymoistureprecipitationrainsnowwindlightphotoperiodday-lengthradiationUVsoilcovercomplexityheterogeneityproductivityenergynutrientcarbonnitrogenphosphorusfood-webtrophic-interactionpredator-preycompetitionintraspecificinterspecificresourcepartitioningcoexistencediversityrichnessevennessabundancedensityfrequencyoccurrencepresenceabsencedetectionsamplingcollectionpitfall-trapquadrattransecthand-collectingbeatingsiftingextractionBerleseTullgrenWinklerflotationsortingpreservationethanolformalinmountslidelabelcatalogdatabaserepositorymuseumvouchertypeholotypeparatypesyntypelectotypeparalectotypeneotypetopotypeoriginaldatalocalitygeoreferencecoordinatelatitudelongitudeelevationaltitudedepthcollectordeterminationidentificationexpertspecialistliteraturepublicationjournalarticlepaperbookthesisdissertationreportchecklistonlinedigitalarchivebibliographyreferencecitationauthoryearvolumepagefiguremaptableappendixsupplementarymaterialmethodsresultsdiscussionconclusionabstractkeywordindex-termsubjecttopicdisciplinesciencebiologyzoologyarachnologyentomologyevolutionphylogeneticsbiologicalnatureprotectionmanagementpolicylegislationrestorationmonitoringassessmentindicatorflagshipumbrellakeystoneengineerservicefunctiondynamicsstabilityresilienceresistancerecoverysuccessiondisturbanceclimatechangewarmingglacierrangeshiftexpansioncontractionfragmentationisolationconnectivitycorridornetworkreserveparkprotectedareawildernessurbanruralagriculturalforestryminingimpactthreatriskvulnerabilitysensitivityexposureadaptivecapacitymitigationadaptationsustainableuseharvesttradeeconomymedicineresearcheducationoutreachcitizenengagementawarenessappreciationaestheticculturalspiritualexistencevalueintrinsicinstrumentalutilitariananthropocentricbiocentricecocentricethicphilosophyhistorydiscoveryexpeditionvoyagesurveyinventoryfaunabiotabiodiversitylifeearthplanetwildlifeanimalinvertebratearthropodcheliceratearachnidspideraraneomorphentelegynemicryphantidOreoneta2004frigidatype-speciesalpinefeedingtype-specimenclimate-changeprotected-area

• Tortopus

  Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.

  EphemeropteraPolymitarcyidaeburrowing-mayfliesNeotropicalclay-bank-habitatU-shaped-tunnelsmandibular-tusksTortopsisCampsurinaePanamerican-distributioncladistic-revisionsynapomorphyparastyli-receptorsflattened-penesfurcasternal-protuberancessubapical-tubercleslimnological-sampling-biasnymph-morphologyadult-morphologytype-species-T.-igaranusnew-species-T.-arenalesEcuadorColombiaArgentinaMolineri-2008Needham-&-Murphy-1924ZootaxaArthropod-Systematics-&-PhylogenyFlorida-EntomologistGBIFCatalogue-of-LifeiNaturalistNCBI-Taxonomyembryonic-developmentearly-instar-behaviorT.-harrisiT.-bellusT.-circumfluusT.-zottaiT.-igaranusT.-incertusT.-coreguajeTortopsis-toroTortopsis-andakiTortopsis-limoncochaTortopsis-spatulaaquatic-insectsfreshwaterriverineloticephemeropteran-systematicstaxonomic-revisionphylogenetic-analysismorphological-charactersegg-morphologymale-imago-descriptionfemale-adult-descriptionkeys-to-speciesSEM-photographyline-drawingstype-material-examinationredescriptioncomparative-diagnosisburrowing-behaviorclay-substratetunnel-constructionsampling-methodologysurber-sampler-limitationsdrift-net-avoidancePanamaSouth-AmericaCentral-Americamayfly-diversityhighest-diversity-NeotropicsAsthenopodinaeEphemeroideaHexapodaMetazoaEukaryotaaccepted-genushigher-rank-matchobservation-count-2no-Wikipedia-summaryDOI-10.11646/zootaxa.2481.1.1DOI-10.11646/zootaxa.2436.1.4DOI-10.3897/asp.79.e62735DOI-10.2307/3493493Scott-et-al.-1959NavásUlmerTraverLugo-Ortiz-&-McCaffertyDomínguezBanksMcDunnoughPictetNeedham-&-Murphyfreshwater-ecologyaquatic-entomologyinsect-developmentnymphal-stagesimago-descriptionalate-adultswing-venationR-sectorgonopore-structureparastyli-lengthpedestal-morphologylateral-view-curvaturemandibular-morphologytusk-tuberclesmesosternal-structureabdominal-sternum-notchfurcasternumprotuberance-configurationreceptor-socket-orientationIR-veinR2-veinclaw-like-structuregonopore-associationpenis-base-separationfemale-fore-wingadditional-veinscomparative-morphologytype-specimenholotypeallotypeparatypeneotropical-speciesnew-species-descriptionspecies-keyidentification-keymale-adult-keyfemale-adult-keynymph-keygeneric-keyPolymitarcyidae-generasystematicsphylogenycladisticsmonophylysister-groupcharacter-statetransformation-seriesparsimony-analysismorphological-data-setexternal-morphologyegg-chorionmicropyleegg-bursterhatchingemergencebehavioral-studyearly-instarembryologyovipositionsubstrate-preferencehabitat-specificityendemicregional-diversityconservationbiomonitoringwater-quality-indicatormacroinvertebratebenthicinfaunalinterstitialhyporheicriver-bankstream-bankerosionsedimentparticle-sizeclay-contentbank-stabilityroot-matvegetation-associationflow-regimecurrent-velocitydepthwidthstream-ordercatchmentbasinecoregionbiogeographydispersalcolonizationvicariancespeciationadaptive-radiationmorphological-divergenceconvergent-evolutionhomoplasycharacter-conflictbootstrap-supportBremer-supportconsensus-treestrict-consensusmajority-ruletaxonomic-stabilitynomenclatural-actoriginal-descriptionsubsequent-designationtype-localitytype-depositorymuseum-collectionspecimen-preservationalcohol-preservationslide-mountingSEM-preparationimaging-techniquesscientific-illustrationphotographymeasurementmorphometricsratiosproportionsallometrysexual-dimorphismgeographic-variationintraspecific-variationinterspecific-variationdiagnostic-charactersidentification-confidencetaxonomic-expertiseliterature-reviewbibliographic-searchdatabase-queryonline-repositoryopen-accesspaywallabstract-onlyfull-text-unavailablecitation-analysisreference-listsource-qualityevidence-strengthknowledge-gapresearch-needfuture-studycollaborationfield-workcollection-tripsampling-effortspecimen-availabilitytype-examinationrevision-scopetaxonomic-conceptcircumscriptioninclusionexclusionspecies-groupspecies-complexcryptic-speciesmorphospeciesphylogenetic-speciesevolutionary-significant-unitoperational-taxonomic-unitDNA-barcodingmolecular-systematicsintegrative-taxonomytraditional-taxonomyalpha-taxonomybeta-taxonomygamma-taxonomyclassification-hierarchyLinnaean-systemrank-based-nomenclaturephylogenetic-nomenclatureclade-namecrown-groupstem-grouptotal-evidencecongruenceincongruencedata-partitioncharacter-weightingimplied-weightingequal-weightingordered-charactersunordered-charactersmultistate-charactersbinary-characterscontinuous-charactersmeristic-charactersqualitative-charactersquantitative-charactersmorphological-matrixdata-matrixcharacter-matrixtaxon-samplingcharacter-samplingmissing-datainapplicable-datapolymorphismintraspecific-polymorphismfixed-differencediagnostic-differenceunique-derived-characterautapomorphyshared-derived-charactersymplesiomorphyconvergent-characterparallel-evolutionreversalcharacter-optimizationACCTRANDELTRANunambiguous-optimizationambiguous-optimizationmost-parsimonious-treetree-lengthconsistency-indexretention-indexrescaled-consistency-indexhomoplasy-indexcharacter-fitcharacter-congruencecharacter-supportbranch-supportnode-supportclade-stabilitytaxonomic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