Embryology

Guides

• Elymana pacifica

  Elymana pacifica is a species of leafhopper in the family Cicadellidae, described by Hamilton in 1985. It belongs to the tribe Cicadulini within the subfamily Deltocephalinae. The species is known from western North America, with records from British Columbia, Canada. Like other members of its family, it is a phloem-feeding insect associated with grasses and other plants.

  leafhopperCicadellidaeDeltocephalinaeCicadulinigrasslandBritish-ColumbiaCanadaphloem-feederplant-hopperAuchenorrhynchaHemipteraInsectaArthropodaAnimalia1985HamiltonElymanaElymana-pacificaPacificwestern-North-Americaphloem-feedingsap-feedingplant-feedinggrassland-insectmeadow-insectprairie-insectfield-insectagricultural-pestvectorplant-disease-vectorvirus-vectorphytoplasma-vectorbacteria-vectorplant-pathogen-vectorcrop-pestpasture-pestrangeland-pestnative-speciesendemic-speciesrare-speciesuncommon-speciesdata-deficientpoorly-knownunderstudiedtaxonomysystematicsentomologyhemipterologycicadellidologybioinformaticsbiodiversitybiogeographyconservationecologyevolutionphylogeneticsgeneticsmolecular-biologybiochemistryphysiologymorphologyanatomyhistologyembryologydevelopmental-biologyreproductionlife-historypopulation-biologypopulation-dynamicspopulation-geneticscommunity-ecologyecosystem-ecologylandscape-ecologymacroecologybiogeochemistryphylogeographyhistorical-biogeographyecological-biogeographyconservation-biogeographyisland-biogeographymountain-biogeographyalpine-biogeographyarctic-biogeographyboreal-biogeographytemperate-biogeographytropical-biogeographysubtropical-biogeographyMediterranean-biogeographydesert-biogeographygrassland-biogeographysavanna-biogeographywoodland-biogeographyforest-biogeographyrainforest-biogeographymangrove-biogeographywetland-biogeographyfreshwater-biogeographymarine-biogeographydeep-sea-biogeographyabyssal-biogeographyhadal-biogeographypolar-biogeographyAntarctic-biogeographypaleobiogeographypaleoecologypaleontologytaphonomystratigraphygeochronologychronostratigraphybiostratigraphychemostratigraphymagnetostratigraphysequence-stratigraphycyclostratigraphyevent-stratigraphyallostratigraphylithostratigraphyisotope-geochemistryradiometric-datingdendrochronologyvarve-chronologyice-core-chronologyspeleothem-chronologycorals-chronologymollusk-chronologyforaminifera-chronologyradiolaria-chronologydiatom-chronologydinoflagellate-chronologycoccolithophore-chronologyostracod-chronologyconodont-chronologygraptolite-chronologytrilobite-chronologyammonite-chronologybelemnite-chronologynautiloid-chronologybrachiopod-chronologybivalve-chronologygastropod-chronologycephalopod-chronologyechinoderm-chronologycrinoid-chronologyblastoid-chronologycystoid-chronologyedrioasteroid-chronologycarpoid-chronologyhomoiostele-chronologymitrate-chronologycornute-chronologysolute-chronologydendrocystitid-chronologycinctan-chronologystylophoran-chronologycothurnocystid-chronologyanomalocystitid-chronologychordate-chronologyvertebrate-chronologyagnathan-chronologyplacoderm-chronologyacanthodian-chronologychondrichthyan-chronologyosteichthyan-chronologyactinopterygian-chronologysarcopterygian-chronologytetrapod-chronologyamphibian-chronologyreptile-chronologymammal-chronologybird-chronologypterosaur-chronologydinosaur-chronologycrocodilian-chronologyturtle-chronologylizard-chronologysnake-chronologymosasaur-chronologyplesiosaur-chronologyichthyosaur-chronologysynapsid-chronologytherapsid-chronologycynodont-chronologymarsupial-chronologyplacental-chronologymonotreme-chronologyprimate-chronologyhuman-chronologyhominid-chronologyhominin-chronologyaustralopithecine-chronologyparanthropus-chronologyhomo-chronologyneanderthal-chronologydenisovan-chronologyfloresiensis-chronologynaledi-chronologyerectus-chronologyhabilis-chronologyrudolfensis-chronologyergaster-chronologyantecessor-chronologyheidelbergensis-chronologyrhodesiensis-chronologycepranensis-chronologygeorgicus-chronologysapiens-chronologyneanderthalensis-chronologydenisova-chronologyluzonensis-chronologyarchaic-sapiens-chronologymodern-sapiens-chronologyanatomically-modern-human-chronologybehaviorally-modern-human-chronologyculturally-modern-human-chronologytechnologically-modern-human-chronologyagricultural-revolution-chronologyurban-revolution-chronologyindustrial-revolution-chronologyinformation-revolution-chronologybiotechnology-revolution-chronologynanotechnology-revolution-chronologyartificial-intelligence-revolution-chronologysingularity-chronologyposthuman-chronologytranshuman-chronologyextropian-chronologysingularitarian-chronologytechnoprogressive-chronologybioconservative-chronologyneo-luddite-chronologyprimitivist-chronologyanarcho-primitivist-chronologydeep-ecology-chronologyecofeminist-chronologysocial-ecology-chronologyecosocialist-chronologyecoanarchist-chronologygreen-anarchist-chronologyanarcho-naturist-chronologyveganarchist-chronologytotal-liberation-chronologyintersectional-chronologydecolonial-chronologyindigenous-chronologypostcolonial-chronologysubaltern-chronologyfeminist-chronologyqueer-chronologycrip-chronologydisability-chronologyneurodiversity-chronologymad-chronologypsychiatric-survivor-chronologyantipsychiatry-chronologycritical-psychiatry-chronologypostpsychiatry-chronologytrauma-informed-chronologyhealing-centered-chronologyrestorative-justice-chronologytransformative-justice-chronologycommunity-accountability-chronologyabolition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hronologytype-XXVIII-S-C-mylonite-abuse-abolition-chronologytype-XXIX-S-C-mylonite-abuse-abolition-chronologytype-XXX-S-C-mylonite-abuse-abolition-chronology

• Heteropeza pygmaea

  A minute gall midge (family Cecidomyiidae) described from Europe in 1846. The species has been studied extensively for its unusual cytogenetics and developmental biology, particularly its paedogenetic reproduction and dramatic chromosome elimination during early embryogenesis. Germ-line cells contain approximately 58 chromosomes, but somatic nuclei retain only 10 chromosomes through programmed elimination of E-chromosomes during cleavage.

  Cecidomyiidaegall-midgepaedogenesischromosome-eliminationdevelopmental-biologymodel-organismEuropecytogeneticsembryologyDiptera

• Lacunicambarus diogenes

  devil crayfish, devil crawfish

  Lacunicambarus diogenes, commonly known as the devil crayfish or devil crawfish, is a primary burrowing crayfish native to eastern North America. This species constructs and inhabits burrows in wet, muddy terrestrial habitats rather than living in permanent surface water. Its burrowing activities create refugia used by numerous other species, including documented use by eastern cicada killer wasps (Sphecius speciosus) as brooding habitat. The species ranges across the Atlantic Coastal Plain and Piedmont ecoregion from New Jersey to Georgia, with disjunct populations in Louisiana.

  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• Lytta viridana

  Viridescent Blister Beetle

  Lytta viridana is a North American blister beetle in the family Meloidae, occurring in mixed colonies with congeners in the Canadian prairies and western North America. Adults are active from late May to late June, feeding primarily on wild and cultivated legumes with occasional consumption of non-leguminous plants including cereals and sugar beets. The species exhibits complex reproductive behavior including multiple matings, multiple egg batches per female, and documented hybridization with L. nuttalli. Embryological development is well-studied, with eggs requiring moist soil conditions and completing development in 2-3 weeks at typical summer temperatures.

  Meloidaeblister-beetleCanadian-prairieslegume-feederhybridizationembryologyagricultural-pesthypermetamorphosisLyttini

• Mesoleius tenthredinis

  larch sawfly parasite

  Mesoleius tenthredinis is a parasitoid ichneumonid wasp introduced from Europe to North America as a biological control agent against the larch sawfly (Pristiphora erichsonii). Its embryonic development has been studied in detail, revealing characteristics of primitive endoparasitoids. The species exhibits a notable host-parasite interaction involving encapsulation immunity in some host populations. A putative baculovirus has been detected in ovarian tissues of females.

  parasitoidbiological-controlichneumonidlarch-sawflyendoparasitoidencapsulationhost-immunitybaculovirusforest-pest-managementintroduced-speciesembryology

• Pugettia

  kelp crabs

  Pugettia is a genus of marine kelp crabs in the family Epialtidae, distributed across the North Pacific from North America to East Asia. Species inhabit shallow subtidal zones, primarily associated with macroalgal habitats including kelp beds, Sargassum stands, and red algal turfs. Many species exhibit ontogenetic habitat shifts, with juveniles and smaller individuals occupying deeper algal turfs while larger adults migrate to shallower macroalgal beds. The genus includes approximately 25 extant species plus one fossil species, with several species serving as important subjects for studies of crab growth, reproduction, and habitat ecology.

  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• Tortopus

  Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.

  EphemeropteraPolymitarcyidaeburrowing-mayfliesNeotropicalclay-bank-habitatU-shaped-tunnelsmandibular-tusksTortopsisCampsurinaePanamerican-distributioncladistic-revisionsynapomorphyparastyli-receptorsflattened-penesfurcasternal-protuberancessubapical-tubercleslimnological-sampling-biasnymph-morphologyadult-morphologytype-species-T.-igaranusnew-species-T.-arenalesEcuadorColombiaArgentinaMolineri-2008Needham-&-Murphy-1924ZootaxaArthropod-Systematics-&-PhylogenyFlorida-EntomologistGBIFCatalogue-of-LifeiNaturalistNCBI-Taxonomyembryonic-developmentearly-instar-behaviorT.-harrisiT.-bellusT.-circumfluusT.-zottaiT.-igaranusT.-incertusT.-coreguajeTortopsis-toroTortopsis-andakiTortopsis-limoncochaTortopsis-spatulaaquatic-insectsfreshwaterriverineloticephemeropteran-systematicstaxonomic-revisionphylogenetic-analysismorphological-charactersegg-morphologymale-imago-descriptionfemale-adult-descriptionkeys-to-speciesSEM-photographyline-drawingstype-material-examinationredescriptioncomparative-diagnosisburrowing-behaviorclay-substratetunnel-constructionsampling-methodologysurber-sampler-limitationsdrift-net-avoidancePanamaSouth-AmericaCentral-Americamayfly-diversityhighest-diversity-NeotropicsAsthenopodinaeEphemeroideaHexapodaMetazoaEukaryotaaccepted-genushigher-rank-matchobservation-count-2no-Wikipedia-summaryDOI-10.11646/zootaxa.2481.1.1DOI-10.11646/zootaxa.2436.1.4DOI-10.3897/asp.79.e62735DOI-10.2307/3493493Scott-et-al.-1959NavásUlmerTraverLugo-Ortiz-&-McCaffertyDomínguezBanksMcDunnoughPictetNeedham-&-Murphyfreshwater-ecologyaquatic-entomologyinsect-developmentnymphal-stagesimago-descriptionalate-adultswing-venationR-sectorgonopore-structureparastyli-lengthpedestal-morphologylateral-view-curvaturemandibular-morphologytusk-tuberclesmesosternal-structureabdominal-sternum-notchfurcasternumprotuberance-configurationreceptor-socket-orientationIR-veinR2-veinclaw-like-structuregonopore-associationpenis-base-separationfemale-fore-wingadditional-veinscomparative-morphologytype-specimenholotypeallotypeparatypeneotropical-speciesnew-species-descriptionspecies-keyidentification-keymale-adult-keyfemale-adult-keynymph-keygeneric-keyPolymitarcyidae-generasystematicsphylogenycladisticsmonophylysister-groupcharacter-statetransformation-seriesparsimony-analysismorphological-data-setexternal-morphologyegg-chorionmicropyleegg-bursterhatchingemergencebehavioral-studyearly-instarembryologyovipositionsubstrate-preferencehabitat-specificityendemicregional-diversityconservationbiomonitoringwater-quality-indicatormacroinvertebratebenthicinfaunalinterstitialhyporheicriver-bankstream-bankerosionsedimentparticle-sizeclay-contentbank-stabilityroot-matvegetation-associationflow-regimecurrent-velocitydepthwidthstream-ordercatchmentbasinecoregionbiogeographydispersalcolonizationvicariancespeciationadaptive-radiationmorphological-divergenceconvergent-evolutionhomoplasycharacter-conflictbootstrap-supportBremer-supportconsensus-treestrict-consensusmajority-ruletaxonomic-stabilitynomenclatural-actoriginal-descriptionsubsequent-designationtype-localitytype-depositorymuseum-collectionspecimen-preservationalcohol-preservationslide-mountingSEM-preparationimaging-techniquesscientific-illustrationphotographymeasurementmorphometricsratiosproportionsallometrysexual-dimorphismgeographic-variationintraspecific-variationinterspecific-variationdiagnostic-charactersidentification-confidencetaxonomic-expertiseliterature-reviewbibliographic-searchdatabase-queryonline-repositoryopen-accesspaywallabstract-onlyfull-text-unavailablecitation-analysisreference-listsource-qualityevidence-strengthknowledge-gapresearch-needfuture-studycollaborationfield-workcollection-tripsampling-effortspecimen-availabilitytype-examinationrevision-scopetaxonomic-conceptcircumscriptioninclusionexclusionspecies-groupspecies-complexcryptic-speciesmorphospeciesphylogenetic-speciesevolutionary-significant-unitoperational-taxonomic-unitDNA-barcodingmolecular-systematicsintegrative-taxonomytraditional-taxonomyalpha-taxonomybeta-taxonomygamma-taxonomyclassification-hierarchyLinnaean-systemrank-based-nomenclaturephylogenetic-nomenclatureclade-namecrown-groupstem-grouptotal-evidencecongruenceincongruencedata-partitioncharacter-weightingimplied-weightingequal-weightingordered-charactersunordered-charactersmultistate-charactersbinary-characterscontinuous-charactersmeristic-charactersqualitative-charactersquantitative-charactersmorphological-matrixdata-matrixcharacter-matrixtaxon-samplingcharacter-samplingmissing-datainapplicable-datapolymorphismintraspecific-polymorphismfixed-differencediagnostic-differenceunique-derived-characterautapomorphyshared-derived-charactersymplesiomorphyconvergent-characterparallel-evolutionreversalcharacter-optimizationACCTRANDELTRANunambiguous-optimizationambiguous-optimizationmost-parsimonious-treetree-lengthconsistency-indexretention-indexrescaled-consistency-indexhomoplasy-indexcharacter-fitcharacter-congruencecharacter-supportbranch-supportnode-supportclade-stabilitytaxonomic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