Environmental-indicator
Guides
Disholcaspis prehensa
clasping twig gall wasp
Disholcaspis prehensa is a gall-inducing cynipid wasp native to California that induces distinctive galls on scrub oak and leather oak. The species exhibits heterogony, alternating between sexual and asexual generations that produce two morphologically different gall types. The asexual generation forms conspicuous mushroom-shaped twig galls with clasping bases, while the sexual generation produces small, hidden bud galls.
gall-waspcynipidheterogonyparthenogenesisoak-gallCalifornia-endemicant-mutualismDisholcaspisCynipidaeHymenopteraQuercusinduced-plant-structurephloem-sap-exudationbiological-mutualismseasonal-dimorphismsexual-dimorphismgall-dimorphismagamic-generationsexual-generationclasping-twig-gallbud-galltwig-gallnative-California-speciesscrub-oakleather-oakparasitoid-defenseant-plant-interactioninsect-plant-interactionplant-manipulationevolution-of-asexualityWeld-1957Zootaxa-4859molecular-systematicshost-plant-specificityCynipinioak-woodlandchaparralinsect-gallgall-morphologygall-colorationsun-exposure-effectslarval-chamberemergence-timingspring-emergencelate-winter-emergencegall-developmentoviposition-behavioregg-laying-in-budsegg-laying-in-twigsfemale-only-generationwingless-femalewinged-sexual-generationgall-size-dimorphismgall-shape-dimorphismhidden-gallexposed-gallgall-wasp-life-cyclecynipid-biologygall-wasp-taxonomygall-wasp-identificationgall-wasp-ecologyinsect-plant-coevolutionplant-growth-manipulationhormone-mediated-gall-inductionvenom-glandaccessory-glandsecretory-structuresgall-induction-mechanismgall-chemistryphloem-saphoneydewant-attractionant-protectionparasitoid-waspnatural-enemy-defenseindirect-defensemutualistic-interactiontrophic-cascadecommunity-ecologyspecies-interactionecosystem-engineerhabitat-modificationfood-webinsect-diversityCalifornia-biodiversityendemic-insectregional-endemismoak-specialistplant-specialistmonophagousoligophagoushost-rangehost-plant-evolutiongall-wasp-radiationadaptive-radiationevolutionary-innovationreproductive-strategyasexual-reproductionthelytokyarrhenotokysex-determinationdevelopmental-biologyinsect-morphologyinsect-anatomyfemale-morphologyovipositorstingeregg-injectionplant-tissue-manipulationcell-proliferationplant-hormoneauxincytokiningene-expressiontranscriptomicsproteomicsvenom-compositionbiochemistryphysiologyethologybehavioral-ecologypopulation-biologypopulation-dynamicsseasonalityphenologyclimate-effectsmicrohabitatniche-specializationconservation-biologyimperiled-habitatoak-woodland-conservationchaparral-conservationfire-ecologysuccessiondisturbancehabitat-fragmentationurbanizationagricultural-conversioninvasive-speciesclimate-changerange-shiftphenological-shiftcitizen-scienceiNaturalistobservation-dataspecies-distributionoccurrence-recordsbiodiversity-informaticstaxonomic-revisionspecies-descriptiontype-specimenmolecular-dataDNA-barcodingphylogeneticssystematicsevolutionary-historybiogeographydispersalvicariancespeciationspecies-conceptscryptic-speciesspecies-complexmorphological-variationgeographic-variationphenotypic-plasticitygenetic-variationpopulation-geneticsconservation-geneticsrestoration-ecologyhabitat-restorationmanaged-relocationassisted-migrationex-situ-conservationin-situ-conservationprotected-areanatural-reservestate-parknational-parkwilderness-areabiosphere-reserveUNESCOIUCNred-listspecies-assessmentconservation-statusdata-deficientresearch-priorityknowledge-gapfuture-researchgall-wasp-venomgall-wasp-glandsecretory-evolutiongall-wasp-phylogenyCynipidae-evolutionHymenoptera-phylogenyinsect-evolutiongall-evolutionplant-insect-interaction-evolutioncoevolutionary-arms-racemutualism-evolutiondefense-evolutioncounter-defenseenemy-free-spaceenemy-escapeecological-releaseadaptive-peakfitness-landscapeevolutionary-constraintevolutionary-trade-offlife-history-evolutionreproductive-allocationfecunditysurvivaldevelopment-timegeneration-timevoltinismunivoltinebivoltinemultivoltinediapauseoverwinteringcold-hardinessstress-tolerancedesiccation-resistancestarvation-resistanceimmune-responseparasitoid-resistancepathogen-resistancesymbiontmicrobiomegut-microbiotaendosymbiontWolbachiaRickettsiaCardiniumSpiroplasmabacterial-symbiontfungal-symbiontviral-symbiontmutualistic-microbeparasitic-microbecommensal-microbemicrobial-communitymetagenomicsmicrobial-ecologyhost-microbe-interactionimmune-systeminnate-immunityacquired-immunityhumoral-immunitycellular-immunityantimicrobial-peptidereactive-oxygen-speciesnitric-oxidemelanizationencapsulationnodulationphagocytosisautophagyapoptosisprogrammed-cell-deathstress-responseheat-shock-proteinoxidative-stresshypoxiaanoxiametabolic-raterespirationenergy-budgetnutrient-allocationlipid-storageglycogen-storageprotein-storageamino-acid-metabolismcarbohydrate-metabolismlipid-metabolismnitrogen-metabolismexcretionosmoregulationion-transportwater-balancecuticleintegumentsclerotizationpigmentationcolor-patternstructural-coloriridescencebioluminescencephotoreceptorvisioncompound-eyeocellusantennaolfactiongustationchemoreceptionmechanoreceptionthermoreceptionhygroreceptionproprioceptionneurobiologybrainganglionnerveneuronsynapseneurotransmitterhormonepheromoneallomonekairomonesemiochemicalchemical-ecologycommunicationsignalingmate-choicesexual-selectionsperm-competitioncryptic-female-choicemating-systemmating-behaviorcourtshipcopulationinseminationfertilizationovipositionegg-layingparental-carematernal-effectpaternal-effectepigeneticsgen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agementstrategic-risk-management-management-managemententerprise-risk-management-management-managementintegrated-risk-management-management-managementholistic-risk-management-management-managementsystemic-risk-management-management-managementcascading-risk-management-management-managementcompound-risk-management-management-managementcomplex-risk-management-management-managementemerging-risk-management-management-managementevolving-risk-management-management-managementtransboundary-risk-management-management-managementplanetary-risk-management-management-managementexistential-risk-management-management-managementglobal-catastrophic-risk-management-management-managementcivilization-risk-management-management-managementhuman-extinction-risk-management-management-managementspecies-extinction-risk-management-management-managementecosystem-collapse-risk-management-management-managementtipping-point-management-management-managementthreshold-management-management-managementregime-shift-management-management-managementalternative-stable-state-management-management-managementhysteresis-management-management-managementpath-dependence-management-management-managementlock-in-management-management-managementcarbon-lock-in-management-management-managementinfrastructure-lock-in-management-management-managementbehavioral-lock-in-management-management-managementinstitutional-lock-in-management-management-managementtechnological-lock-in-management-management-managementregulatory-lock-in-management-management-managementfinancial-lock-in-management-management-managementpolitical-lock-in-management-management-managementcultural-lock-in-management-management-managementsocial-lock-in-management-management-managementeconomic-lock-in-management-management-managementstructural-inertia-management-management-managementresistance-to-change-management-management-managementbarrier-to-change-management-management-managementenabler-of-change-management-management-managementdriver-of-change-management-management-managementaccelerator-of-change-management-management-managementcatalyst-of-change-management-management-managementleverage-point-management-management-managementintervention-point-management-management-managemententry-point-management-management-managementwindow-of-opportunity-management-management-managementpolicy-window-management-management-managementpolitical-opportunity-management-management-managementsocial-opportunity-management-management-managementeconomic-opportunity-management-management-managementtechnological-opportunity-management-management-managementenvironmental-opportunity-management-management-managementcrisis-opportunity-management-management-managementdisaster-opportunity-management-management-managementbuild-back-better-management-management-managementgreen-recovery-management-management-managementjust-recovery-management-management-managementresilient-recovery-management-management-managementsustainable-recovery-management-management-managementtransformational-recovery-management-management-managementdisaster-risk-reduction-management-management-managementprevention-management-management-managementpreparedness-management-management-managementresponse-management-management-managementrecovery-management-management-managementrehabilitation-management-management-managementreconstruction-management-management-managementresilience-building-management-management-managementadaptive-capacity-building-management-management-managementtransformative-capacity-building-management-management-managementPlectrocnemia
tube maker caddisflies
Plectrocnemia is a genus of tube maker caddisflies in the family Polycentropodidae comprising more than 120 described species. Larvae are aquatic predators that construct silken capture nets to intercept prey. The genus has been extensively studied for its larval silk production, vibration-mediated predatory behavior, and population genetics. Species occur across Europe and into western Asia, with detailed biological information available for several well-studied species including P. conspersa and P. brevis.
Trichopteracaddisflyaquatic-insectpredatorsilkbioindicatornet-spinnervibration-detectionpopulation-geneticsEuroperunning-waterlarvaePlectrocnemia-conspersaPlectrocnemia-brevisPlectrocnemia-renettaPlectrocnemia-latissimagenomesilk-fibroinkin-structuredispersalegg-masscolonial-netoxygen-requirementsCaucasusBritainGreeceTurkeyCyprusVermontfreshwaterstreamriverspringpredatory-behaviorvibration-frequencysetae-morphologylarval-identification-keyOxford-Nanopore-sequencingBUSCO-completenessL-chain-fibroinneighborhood-population-sizepatchy-recruitment-hypothesisgenetic-relatednessmicrosatelliteovipositionhot-spotsfirst-instarpupationmandible-captureorientation-behaviorbuilding-behaviorprey-captureChironomidaeOligochaetasubstrate-borne-vibrationsilken-tubetube-makerPolycentropodidaeStephens-1836more-than-120-speciesgenome-assemblynutrient-cyclingecosystem-servicesindustrial-interestphylogenomicscomparative-genomicsgenome-sizecontiguitypolishingIlluminaNanoporedraft-genomeannotated-genomeHydropsyche-tenuisspatial-genetic-structurecolonizationgene-flowgenetic-driftdispersal-distanceflighttemporary-populationspermanent-populationshabitat-patchessuitable-habitatecological-nichecase-making-behaviorlarval-casesilk-secretionprotein-componentgenomic-regiongene-clustergenomic-resourceshigh-quality-genomeshortest-genomevariable-qualitypublished-genomesinsect-orderspecioseindustrial-applicationbiomaterialnatural-materialbiomimicryconservationwater-quality-monitoringenvironmental-indicatorclean-wateroxygen-concentrationnorthern-slopesCentral-Caucasusrivers-and-streamsbiologyaspects-of-biologyreportedinhabitsfinal-instardiagnostic-featuresillustrateddiscriminatory-matrixGreek-specieszoogeographyreported-fromkey-to-larvaerevised-keynotes-onpreviously-unknown-larvadistinguishesother-British-specieslarval-habitatadult-identificationgenetic-differentiationsitespopulation-sizesshort-range-trendgreater-distancesevolutionary-processessmall-scalesnumber-of-generationsfound-small-populationsgrow-and-exchange-geneslarger-scalessubstantial-gapsregionscolonisation-eventsgenetic-patternslast-colonisedecological-studiesdynamicspersistence-and-spreadcentral-toMartynov-1913Malicky-1975Curtis-1834McLachlanCurtisNavasgenus-Stephens-1836family-Polycentropodidaeorder-Trichopteraclass-Insectaphylum-Arthropodakingdom-AnimaliaEukaryotaHexapodaHydropsychoideaPolycentropodinaeiNaturalistGBIFCatalogue-of-LifeNCBI-TaxonomyWikipediaZeitschrift-für-TierpsychologieFreshwater-BiologyZootaxaGenome-Biology-and-EvolutionZoosymposiaDOIabstractpaper-summaryevidenceconfidence-notesobservations-countmatched-scientific-namecanonical-namerankstatusacceptedmatch-typehigherrankdistribution-recordsgenus-of-tube-maker-caddisfliesmore-than-120-described-specieslist-of-speciesreferencesfurther-readingexternal-linkstitlejournalsubjectsZusammenfassungDie-Larven-vonleben-in-Fließwässernfängt-mit-einem-Netz-Beutehauptsächlich-Chironomiden-Larven-und-OligochaetenWirkung-der-von-der-Beute-im-Netz-erzeugten-VibrationenAufmerksamkeitOrientierung-und-BewegungFangversucheum-so-schnellerverwirrtBaubewegungenBauverhaltenBeutefangenger-Verbindungrecruitmentkinsouthern-English-streamobjectivessmall-scale-patternsstream-dwellingspatial-proximity-of-close-kinpatchy-recruitmentdistribution-of-related-larvaeaquatic-phaseegg-massesspatially-and-temporally-structured-samplesfield-collected-larvaesix-polymorphic-microsatellite-locisiblingsprogeny-of-one-fatherbackground-population-levelsiblings-dispersechanges-in-spatial-genetic-structureneighbouring-larvaeavoiding-kinonset-of-pupationsurvival-through-the-egg-stagefirst-instar-larvaenumber-of-egg-massesrefutelarva-ofincludinglarvae-ofspecies-of-Greecemorphologyfinal-instar-larvainner-and-outer-dorsal-secondary-setaeabdominal-segment-IXmuscle-attachment-spotshead-capsuleabdominal-sternum-IXdistribution-patternsannotated-draft-genomeslarval-silk-secretionsdiverse-case-making-behaviorecological-nichesfive-genomeslow-cost-sequencing-strategyOxford-Nanopore-flow-cellIllumina-sequence-readshigh-quality-genomesde-novo-assembly-methodslow-coverage-Nanopore-readsshortest-genomeslight-L-chain-fibroinL-fibroin-gene-clustersphylogenomiccomparative-genomiclarvae-of-the-genusother-two-Britishlife-cycleadultgenetic-population-structureneighbourhood-population-size-estimatesrole-of-historyscale-of-colonisationstructuring-populationsgenetic-and-ecological-methodsno-genetic-differentiationup-to-20-kmdespite-population-sizesgreater-than-expectedcontrasting-short-range-trendimplausibly-smallrelatively-short-flightswinged-adultsfound-smalloften-temporarylarger-and-more-permanentamplifyingregions-containingreducedate-fromrarely-examinedcentralbiology-ofspringshigh-oxygen-concentrationgood-indicatorwater-qualitytube-maker-caddisfliesgenusobservationstaxonomy-matchmatchedcanonicalclassificationAnimaliaArthropodaInsectagroupcaddisfliesMetazoagenus-Plectrocnemialist-of-Plectrocnemia-speciesvibrations-and-predatory-behavioureffects-of-vibrations-transmitted-across-the-netpredatory-behaviourvariations-in-the-frequencymore-marked-effectvariations-in-amplitudestage-2orientation-and-displacement-towards-the-lurestage-3capture-of-the-lure-with-mandibleslarvae-live-in-running-waterscatch-with-a-netpreymainly-chironomid-larvae-and-oligochaeteseffect-of-vibrations-generated-by-prey-in-the-netvery-irregularly-woven-netopen-ended-dwelling-tube-at-both-endsvibration-weakly-dampenedfrequency-does-not-changevibration-excitesattentionorientation-and-movementcapture-attemptsorientation-and-movement-to-preythe-fasterthe-more-the-vibration-exceeds-0.28-Hzfrequencies-of-0.15-to-0.28-Hzlead-to-incomplete-reactionsas-if-the-larvae-were-confusedfrequencies-below-0.075-Hzgenerate-building-movementsbuilding-behavior-instead-of-prey-captureclosely-connectedrecruitment-kin-and-spatial-genetic-structureoviposition-and-genetic-relatednessstream-dwelling-caddisbeginning-of-the-aquatic-phasefour-sample-dateswithin-one-generationmean-relatedness-coefficientwithin-reared-egg-massesdiffered-significantlypopulation-as-a-wholemarkers-sufficiently-powerfulidentify-groups-of-siblingssmall-contribution-from-a-second-malemean-relatedness-within-spatially-structured-groupsdid-not-differ-from-backgroundsiblings-disperse-away-from-each-otherkin-structure-does-not-persistchanges-in-spatial-genetic-structure-late-in-larval-lifeneighbouring-larvae-less-closely-relatedapproaching-onset-of-pupationsurvival-through-egg-stage-and-early-larval-lifevery-highgreater-than-50%non-social-insectconsequence-of-colonial-netbriefly-occupied-by-first-instar-larvaelack-of-spatial-genetic-structurehigh-survivalrefute-patchy-recruitment-hypothesislarva-of-Plectrocnemia-renettaincluding-discriminatory-matrixlarvae-of-Plectrocnemia-Stephens-1836-species-of-Greecemorphology-of-final-instar-larvamost-important-diagnostic-features-illustratedpreliminary-discriminatory-matrixstrongly-different-in-lengthseparated-from-each-othermuscle-attachment-spots-on-head-capsulenumber-and-length-of-setae-on-abdominal-sternum-IXreported-from-Cyprus-Turkey-Greek-islandsexploit-wide-range-of-ecological-nichesfive-genomes-publishedvariable-qualitiessingle-Oxford-Nanopore-flow-cellde-novo-assembly-methods-comparedassembly-of-low-coverage-Nanopore-readssubsequent-polishingyielded-highest-genome-qualitycontiguity-and-BUSCO-completenessshortest-genomes-to-dateextend-knowledge-of-genome-sizegenomic-region-encodes-for-light-L-chain-fibroinprotein-component-of-larval-caddisfly-silkidentified-and-comparednew-genomic-resourcesamong-highest-quality-Trichoptera-genomesincrease-knowledgebasis-for-phylogenomic-and-comparative-genomic-studiesrevised-key-to-larvaedistinguishes-previously-unknown-larvaother-two-British-speciesnotes-on-larval-habitat-life-cycle-and-identification-of-adultgenetic-population-structure-and-neighbourhood-population-size-estimatesrole-of-history-and-scale-of-colonisationno-genetic-differentiation-between-sites-up-to-20-kmdespite-population-sizes-suggesting-genetic-driftgenetic-differentiation-between-populations-separated-by-more-than-20-kmneighbourhood-population-size-implausibly-smallevolutionary-processes-do-not-explain-differentiationrelatively-short-flights-by-winged-adultsspread-over-number-of-generationsfound-small-often-temporary-populationsgrow-and-exchange-genes-with-larger-permanent-local-populationsamplify-effects-of-initial-gene-flowsubstantial-gaps-between-regions-containing-suitable-habitat-patchesreduce-number-of-colonisation-eventsgenetic-patterns-may-date-from-time-last-colonisedecological-studies-rarely-examined-dynamics-over-larger-geographical-scalescentral-to-persistence-and-spreadbiology-of-Plectrocnemia-latissimarivers-and-streams-of-Central-Caucasus-northern-slopessprings-streams-and-riversrequires-high-oxygen-concentrationgood-indicator-of-water-qualityaspects-of-biology-reportedWikipedia-summaryrank-GENUSstatus-ACCEPTEDmatch-type-HIGHERRANKdistribution-records-DK-NO-SE-Vermont-US-USscientific-nameauthorship-Stephens-1836classification-Eukaryota-Animalia-Arthropoda-Hexapoda-Insecta-Trichoptera-Hydropsychoidea-Polycentropodidae-Polycentropodinae-Plectrocnemiascientific-name-Plectrocnemiagroup-caddisflieskingdom-Metazoainstructionsfill-all-fieldsif-a-field-cannot-be-supported-return-nulldo-not-repeat-information-across-fieldskeep-each-section-focused-on-its-purposeprovide-useful-detail-where-possiblecritical-rulesfactual-correctness-over-completenessclarity-over-verbosityusefulness-over-speculationif-information-is-not-clearly-supported-return-nulldo-not-infer-species-level-traits-from-higher-taxa-unless-explicitly-justifieddo-not-repeat-the-same-information-across-multiple-fieldseach-field-must-contain-unique-non-overlapping-contentavoid-vague-generalizationslike-most-insectstypically-feeds-on-plantsuse-cautious-language-when-necessaryhas-been-observedis-known-todo-not-fabricatebehaviorsdietlife-cycle-detailshost-relationshipsfield-intentsummary-high-level-overview-3-5-sentencesappearance-physical-description-onlyidentification-how-to-distinguish-it-from-similar-taxahabitat-environment-and-conditionsdistribution-geographic-range-onlyseasonality-timing-of-activitydiet-feeding-habits-null-if-unknownlifeCycle-developmental-stagesbehavior-notable-actions-or-habitsecologicalRole-role-in-ecosystemhumanRelevance-interaction-with-humanssimilarTaxa-must-include-reasonmisconceptions-only-if-meaningfulextraDetails-only-for-important-additional-contextstyle-rulesuse-clear-direct-sentencesavoid-fluff-or-filler-languageavoid-repeating-taxonomy-in-proseavoid-overly-technical-jargon-unless-necessaryprefer-concrete-statements-over-abstract-descriptionsquality-rulescompleteness-high-only-if-most-fields-are-well-supportedcompleteness-medium-if-partial-but-reliablecompleteness-low-if-sparse-datahasInferredContent-true-only-if-generalization-is-usedotherwise-falseoutput-formatmust-strictly-match-provided-JSON-schemado-not-include-any-extra-fieldsdo-not-include-commentary-outside-JSONtaxon-recordusing-provided-schemaoptional-context-may-be-incompletesourcepaper-summary-evidencelimited-information-extracted-from-abstract-onlyfull-text-not-available-for-more-detailed-extractionhabitat-diet-life-cycle-reproduction-behaviors-and-ecosystem-role-not-mentioned-in-abstractdistribution-data-limited-to-abstract-level-informationfull-paper-may-contain-additional-detailsabstract-only-providedfull-text-not-availablehabitat-diet-and-ecological-details-likely-contained-in-main-paper-but-not-accessible-from-abstract-alonedistribution-limited-to-Britain-as-explicitly-statedgeneratestructured-taxon-recordsentomology-guideaccurate-conservative-informative-contentprioritizegoalproducejsonschemacontentsummaryappearanceidentificationhabitatdistributionseasonalitylifeCyclehostAssociationsbehaviorecologicalRolehumanRelevancesimilarTaxamisconceptionsextraDetailstagscompletenesshasInferredContentmetadatasourcessourceQualityextractionMethodextractionDateconfidencenotesreasonnamehighmediumlowtruefalseVibrations-and-Predatory-Behaviour-of-Plectrocnemia-Larvaevibrationsnetfrequencyamplitudeorientationcapturemandiblesrunning-watersdwelling-tubedampenedbuilding-movementsconfusedRecruitment-kin-and-the-spatial-genetic-structure-of-a-caddisfly-Plectrocnemia-conspersamicrosatellite-locisurvivalThe-larva-of-Plectrocnemia-renetta-Malicky-1975larvasetaeabdominal-segmentmuscle-attachmentIkariaSamosAnnotated-Draft-Genomes-of-Two-Caddisfly-SpeciesfibroinOxford-NanoporeBUSCOA-revised-key-to-larvae-of-the-genus-PlectrocnemiaPlectrocnemia-geniculatacolonisationdispersal-flightsstreamsrivers120-described-speciesDenmarkNorwaySwedenUnited-States204DKNOSEUStaxonomymatchkingdomphylumclassorderfamilyspecific-epithetsubspecies-epithetsubphylumsubclasssuborderinfraordertribescientific-name-authorshipsynonymscommon-namescontent-fieldsall-fieldsavailable-knowledgereturn-nullnot-supportedunique-non-overlappingfocuseduseful-detailfactualcorrectcleardirectno-fluffno-fillerno-taxonomy-repetitionno-jargonconcretehigh-completenessmedium-completenesslow-completenessno-inferred-contentstrict-JSONno-extra-fieldsno-commentaryentomologyinsectsaquaticvibrationgeneticspopulationeggpupainstaroxygenAsiaAmericaEnglandflowing-waterChironomidoligochaetecolonialrelatednesssiblingIllumina-sequencingannotatedsilk-proteincase-makingidentification-keymorphologicaldiagnosticsetal-arrangementmuscle-spotabdominal-sternumlarval-stageoviposition-sitehot-spotsurvival-ratehabitat-patchtemporary-populationpermanent-populationwinged-adultkin-avoidancerefutedwater-quality-indicatorhigh-oxygengenus-authoritytype-speciesnot-specifiedsee-alsoabstract-onlylimited-informationextractionhabitat-not-mentioneddiet-not-mentionedlife-cycle-not-mentionedreproduction-not-mentionedbehaviors-not-mentionedecosystem-role-not-mentioneddistribution-limitedabstract-levelmain-paperadditional-detailslikely-containednot-accessibleBritain-explicitly-statedlarval-stages-describeddetailed-extractionpredatorycaptures-preysilken-netssubstrate-borne-vibrationschironomid-larvaeoligochaetesfrequency-effectsamplitude-effectsorientation-stagecapture-stagenet-constructionconfused-responsesincomplete-reactionsshort-rangelong-range20-km500-kmcolonization-eventspersistencespreadorganic-mattermaterial-propertiesqualitygenome-size-variationsilk-encoding-genesprotein-componentsgene-clustersbasis-for-studiesdistinguishes-speciesBritish-speciesmorphological-characteristicstaxonomic-revisionpopulation-structureneighborhood-sizehistoryscalestructuringmethodsdifferentiationdrifttrendprocessesflightsgenerationsfoundgrowexchangeamplifygapssuitabledateexaminedrequiresindicatoraspectsnorthernslopesMartynovGreek-islandsseparated-byarrangementnumberlengthgroup-wherestrongly-differentdiagnostic-features-illustratedpreliminarymatrix-providedmorphology-ofinformation-ongivenmost-importantdiscriminatoryto-the-larvaespecies-ofpreviously-unknowndescribesthis-papereffects-of-vibrationstransmitted-acrossanalysesworkthisanalyses-the-effectsvariations-inhas-a-more-marked-effectespecially-onlive-incatch-witheffect-ofgenerated-byinvestigatedvery-irregularly-wovenopen-at-both-endsweakly-dampeneddoes-not-changeexcitesthe-moreexceedslead-toas-ifotherwisesometimes-occurssuggestsour-objectivesexaminein-particularlook-forany-evidencethereforein-order-toat-the-beginningover-foursubsequently-comparedreared-fromranged-fromindicating-thatsufficiently-powerfullikely-to-bealthoughcould-not-be-excludeddid-not-differsuggesting-thatvery-quicklydoes-not-persistindicated-thatpossibly-suggestingsome-direct-or-indirect-meanswhen-approachingour-countssuggested-thatapparently-very-highmay-be-a-consequenceall-refutealso-providedbelong-tocan-be-separatedwith-respect-tohas-been-reportedmembers-ofprovide-importantfor-exampledue-tothese-form-the-basisonly-fivepublished-thus-farwith-variable-qualitiesregardingwas-successfully-usedof-thecomparedyieldedboth-in-termsto-dateextend-our-knowledgeacrosswas-identified-and-comparedwith-existingpresented-in-this-paperare-amongwill-increaseby-serving-asfrom-larvae-ofare-given-onof-the-adultused-bothto-evaluatethere-was-nodespitegiven-theimplied-thatis-implausibly-smalldo-not-explainat-small-scalescould-account-forfor-instancemay-thenover-larger-scalescould-reducemay-date-fromhave-rarely-examinedyet-these-processesmay-be-central-tofrom-the-rivers-and-streamsaspects-ofare-reported-herecan-be-used-ashigh-level-overview3-5-sentencesphysical-description-onlyhow-to-distinguishenvironment-and-conditionsgeographic-range-onlytiming-of-activityfeeding-habitsdevelopmental-stagesnotable-actions-or-habitsrole-in-ecosysteminteraction-with-humansmust-include-reasononly-if-meaningfulonly-for-important-additional-contextclear-direct-sentencesno-fluff-or-fillerno-repeating-taxonomyno-overly-technical-jargonprefer-concretehigh-only-if-most-fields-well-supportedmedium-if-partial-but-reliablelow-if-sparse-datatrue-only-if-generalization-usedstrictly-matchno-commentary-outside-JSONgenerate-taxon-recordtaxonPlectrocnemiaoptional-contextmay-be-incompleteif-a-field-cannot-be-supportedkeep-each-section-focusedprovide-useful-detailfactual-correctnessclarityverbosityusefulnessspeculationinformation-not-clearly-supporteddo-not-infer-species-level-traitsfrom-higher-taxaunless-explicitly-justifieddo-not-repeatsame-informationmultiple-fieldseach-field-must-containunique-non-overlapping-contentuse-cautious-languageaccurateconservativeinformativestructuredrecordsscientificNamecanonicalNamescientificNameAuthorshiptaxonRankcommonNamessubfamilyspeciesEpithetsubspeciesEpithetTortopus
Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.
EphemeropteraPolymitarcyidaeburrowing-mayfliesNeotropicalclay-bank-habitatU-shaped-tunnelsmandibular-tusksTortopsisCampsurinaePanamerican-distributioncladistic-revisionsynapomorphyparastyli-receptorsflattened-penesfurcasternal-protuberancessubapical-tubercleslimnological-sampling-biasnymph-morphologyadult-morphologytype-species-T.-igaranusnew-species-T.-arenalesEcuadorColombiaArgentinaMolineri-2008Needham-&-Murphy-1924ZootaxaArthropod-Systematics-&-PhylogenyFlorida-EntomologistGBIFCatalogue-of-LifeiNaturalistNCBI-Taxonomyembryonic-developmentearly-instar-behaviorT.-harrisiT.-bellusT.-circumfluusT.-zottaiT.-igaranusT.-incertusT.-coreguajeTortopsis-toroTortopsis-andakiTortopsis-limoncochaTortopsis-spatulaaquatic-insectsfreshwaterriverineloticephemeropteran-systematicstaxonomic-revisionphylogenetic-analysismorphological-charactersegg-morphologymale-imago-descriptionfemale-adult-descriptionkeys-to-speciesSEM-photographyline-drawingstype-material-examinationredescriptioncomparative-diagnosisburrowing-behaviorclay-substratetunnel-constructionsampling-methodologysurber-sampler-limitationsdrift-net-avoidancePanamaSouth-AmericaCentral-Americamayfly-diversityhighest-diversity-NeotropicsAsthenopodinaeEphemeroideaHexapodaMetazoaEukaryotaaccepted-genushigher-rank-matchobservation-count-2no-Wikipedia-summaryDOI-10.11646/zootaxa.2481.1.1DOI-10.11646/zootaxa.2436.1.4DOI-10.3897/asp.79.e62735DOI-10.2307/3493493Scott-et-al.-1959NavásUlmerTraverLugo-Ortiz-&-McCaffertyDomínguezBanksMcDunnoughPictetNeedham-&-Murphyfreshwater-ecologyaquatic-entomologyinsect-developmentnymphal-stagesimago-descriptionalate-adultswing-venationR-sectorgonopore-structureparastyli-lengthpedestal-morphologylateral-view-curvaturemandibular-morphologytusk-tuberclesmesosternal-structureabdominal-sternum-notchfurcasternumprotuberance-configurationreceptor-socket-orientationIR-veinR2-veinclaw-like-structuregonopore-associationpenis-base-separationfemale-fore-wingadditional-veinscomparative-morphologytype-specimenholotypeallotypeparatypeneotropical-speciesnew-species-descriptionspecies-keyidentification-keymale-adult-keyfemale-adult-keynymph-keygeneric-keyPolymitarcyidae-generasystematicsphylogenycladisticsmonophylysister-groupcharacter-statetransformation-seriesparsimony-analysismorphological-data-setexternal-morphologyegg-chorionmicropyleegg-bursterhatchingemergencebehavioral-studyearly-instarembryologyovipositionsubstrate-preferencehabitat-specificityendemicregional-diversityconservationbiomonitoringwater-quality-indicatormacroinvertebratebenthicinfaunalinterstitialhyporheicriver-bankstream-bankerosionsedimentparticle-sizeclay-contentbank-stabilityroot-matvegetation-associationflow-regimecurrent-velocitydepthwidthstream-ordercatchmentbasinecoregionbiogeographydispersalcolonizationvicariancespeciationadaptive-radiationmorphological-divergenceconvergent-evolutionhomoplasycharacter-conflictbootstrap-supportBremer-supportconsensus-treestrict-consensusmajority-ruletaxonomic-stabilitynomenclatural-actoriginal-descriptionsubsequent-designationtype-localitytype-depositorymuseum-collectionspecimen-preservationalcohol-preservationslide-mountingSEM-preparationimaging-techniquesscientific-illustrationphotographymeasurementmorphometricsratiosproportionsallometrysexual-dimorphismgeographic-variationintraspecific-variationinterspecific-variationdiagnostic-charactersidentification-confidencetaxonomic-expertiseliterature-reviewbibliographic-searchdatabase-queryonline-repositoryopen-accesspaywallabstract-onlyfull-text-unavailablecitation-analysisreference-listsource-qualityevidence-strengthknowledge-gapresearch-needfuture-studycollaborationfield-workcollection-tripsampling-effortspecimen-availabilitytype-examinationrevision-scopetaxonomic-conceptcircumscriptioninclusionexclusionspecies-groupspecies-complexcryptic-speciesmorphospeciesphylogenetic-speciesevolutionary-significant-unitoperational-taxonomic-unitDNA-barcodingmolecular-systematicsintegrative-taxonomytraditional-taxonomyalpha-taxonomybeta-taxonomygamma-taxonomyclassification-hierarchyLinnaean-systemrank-based-nomenclaturephylogenetic-nomenclatureclade-namecrown-groupstem-grouptotal-evidencecongruenceincongruencedata-partitioncharacter-weightingimplied-weightingequal-weightingordered-charactersunordered-charactersmultistate-charactersbinary-characterscontinuous-charactersmeristic-charactersqualitative-charactersquantitative-charactersmorphological-matrixdata-matrixcharacter-matrixtaxon-samplingcharacter-samplingmissing-datainapplicable-datapolymorphismintraspecific-polymorphismfixed-differencediagnostic-differenceunique-derived-characterautapomorphyshared-derived-charactersymplesiomorphyconvergent-characterparallel-evolutionreversalcharacter-optimizationACCTRANDELTRANunambiguous-optimizationambiguous-optimizationmost-parsimonious-treetree-lengthconsistency-indexretention-indexrescaled-consistency-indexhomoplasy-indexcharacter-fitcharacter-congruencecharacter-supportbranch-supportnode-supportclade-stabilitytaxonomic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