Conservation-prioritization

Guides

• Acmaeodera parkeri

  Acmaeodera parkeri is a species of metallic wood-boring beetle in the family Buprestidae, described by Cazier in 1940. The species occurs in Central America and North America, with records from the southwestern United States and Mexico. Adults have been observed visiting flowers of several plant species, including Allionia incarnata (creeping four o'clock), Talinum whitei (White's flameflower), and Evolvulus arizonicus (Arizona blue-eyes).

  Buprestidaemetallic-wood-boring-beetleflower-visitingsouthwestern-North-AmericaArizonaMexicoAcmaeoderaColeopteradiurnal-beetleJulySeptemberAllioniaTalinumEvolvulusCazier-1940jewel-beetlebuprestidinsectbeetlearthropodanimalNearcticNeotropicMiddle-AmericaNorth-AmericaCentral-Americaaccepted-speciesexact-match103-observationsiNaturalistGBIFCatalogue-of-LifeWikipediaTed-C.-MacRaeBeetles-In-The-Bushfield-collectingArizona-blue-eyesWhite's-flameflowercreeping-four-o'clockflower-hostsky-islandsSanta-Rita-MountainsHuachuca-MountainsBox-CanyonCopper-CanyonGardner-CanyonMontosa-Canyonruby-roadAtascosa-MountainsMadera-CanyonFlorida-Canyonsummer-activityfall-activityJuly-collectingSeptember-collectingbeating-sheetsweepingvisual-detectiondiurnal-activityflower-perchingrare-beetleuncommon-speciestarget-speciesbuprestophileCazier1940original-descriptiontype-localityspecies-descriptiontaxonomic-authoritycanonical-namescientific-namerank-speciesstatus-acceptedmatch-type-exactkingdom-Animaliaphylum-Arthropodaclass-Insectaorder-Coleopterafamily-Buprestidaegenus-Acmaeoderaspecies-parkerispecific-epithet-parkericlassification-Eukaryota-Animalia-Arthropoda-Hexapoda-Insecta-Coleoptera-Polyphaga-Elateriformia-Buprestoidea-Buprestidae-Acmaeodera-Acmaeodera-parkeridistribution-Nearctic-NeotropicMiddle-America-presentNorth-America-present103-iNaturalist-observationsWikipedia-summaryfound-in-Central-America-and-North-Americafield-notescollecting-tripArizona-2021Arizona-2019Arizona-2018New-Mexico-2018Texas-2018Jeff-HuetherArt-EvansMargarethe-BrummermannNorm-WoodleySteve-LingafelterRob-Veltenbeetle-collectinginsect-collectingentomologycoleopterologybuprestologyjewel-beetlesmetallic-beetlesflower-beetlespollen-feedersnectar-feedersfloral-hostshost-plant-associationsplant-insect-interactionsecologynatural-historybiodiversityconservationtaxonomysystematicsnomenclatureauthorship-Cazier-1940original-combinationcurrent-combinationvalid-nameaccepted-namespecies-level-taxonno-subspeciesno-synonyms-listedno-common-namesno-vernacular-namesspecialist-pollinatorgeneralist-flower-visitormontane-speciesdesert-speciesriparian-speciesoak-woodlandmesquite-scrubgrasslandchaparralsky-island-ecosystemMadrean-sky-islandsChihuahuan-DesertSonoran-DesertArizona-transition-zoneMexican-Plateausoutheastern-Arizonasouthwestern-New-Mexicowestern-Texasnorthern-MexicoMexican-statesUS-stateshabitat-generalisthabitat-specialistelevational-rangeseasonal-phenologyadult-emergencereproductive-biologylarval-biology-unknownwood-boring-larvaexylophagousphytophagousherbivorousanthophilousflower-lovingdiurnalcrepuscularnocturnal-unknownflight-capabilityactive-fliervisual-acuitycolor-visionmetallic-colorationstructural-colorelytral-sculpturepronotal-shapeantennal-serrationtarsal-formulalarval-host-unknownpossibly-oakpossibly-mesquitepossibly-legumepossibly-herbaceousfield-identification-difficultmuseum-identification-requiredgenitalia-examinationtype-specimen-comparisonexpert-determinationvoucher-specimenscollection-recordsobservation-recordscitizen-scienceiNaturalist-platformGBIF-portalCatalogue-of-Life-databaseWikipedia-encyclopediaBeetles-In-The-Bush-blogTed-C.-MacRae-personal-communicationprimary-literatureCazier-1940-original-descriptionsubsequent-redescriptionstaxonomic-revisionsphylogenetic-placementAcmaeodera-genus-groupAcmaeoderini-tribeBuprestinae-subfamilyBuprestidae-familyElateriformia-infraorderPolyphaga-suborderColeoptera-orderInsecta-classHexapoda-superclassArthropoda-phylumAnimalia-kingdomEukaryota-domainbiogeographydispersalrange-extensionrange-contractionpopulation-statusabundancerarityconservation-statusdata-deficientIUCN-not-evaluatedendemismregional-endemicwidespreaddisjunct-distributioncontinuous-distributionhabitat-fragmentationclimate-change-vulnerabilitycollection-biassampling-effortdetection-probabilityspecies-distribution-modelingecological-nichefundamental-nicherealized-nichebiotic-interactionscompetitionpredationparasitismmutualismpollinationfood-webtrophic-levelprimary-consumerherbivoredetritivore-unknowncarnivore-unknownomnivore-unknownbehavioral-ecologyforagingmate-locationovipositionlarval-developmentpupal-developmentadult-longevitygeneration-timevoltinismunivoltinebivoltine-unknownmultivoltine-unknownoverwinteringdiapausecold-hardinessheat-tolerancedesiccation-resistancewater-balancemetabolic-rateactivity-temperaturethermal-biologybiochemical-adaptationmorphological-adaptationevolutionary-historyphylogenydiversificationspeciationadaptive-radiationconvergent-evolutionmimicryaposematismcrypsiscamouflagesexual-dimorphismsexual-selectionmating-systemreproductive-strategyfecundityegg-productionlarval-survivalpopulation-dynamicsmetapopulationsource-sinkhabitat-patchinesslandscape-ecologyecosystem-servicespollination-servicenutrient-cyclingdecompositionwood-decompositionforest-healthindicator-speciesbioindicatorenvironmental-monitoringecosystem-healthbiodiversity-assessmentconservation-planningprotected-areaswildlife-managementhabitat-restorationinvasive-speciesbiological-controlintegrated-pest-managementsustainable-forestryagroecologyurban-ecologyclimate-adaptationresiliencevulnerabilityextinction-riskpopulation-geneticsgene-flowgenetic-diversityinbreedingoutbreedinglocal-adaptationphenotypic-plasticityepigeneticsdevelopmental-biologymorphometricsallometrygeometric-morphometricsimagingphotographymicroscopymuseum-collectionsnatural-history-collectionsspecimen-preservationcurationdigitizationdata-sharingopen-sciencereproducible-researchtaxonomic-practicenomenclatural-stabilityspecies-conceptintegrative-taxonomymolecular-systematicsDNA-barcodingphylogenomicsevolutionary-genomicsfunctional-genomicstranscriptomicsproteomicsmetabolomicsphenomicsecological-genomicslandscape-genomicsconservation-genomicsbiogeographic-historyvicariancedispersal-biogeographyfounder-effectgenetic-driftnatural-selectionadaptationevolutionary-constraintevolutionary-innovationkey-innovationadaptive-zoneecological-opportunitydiversification-rateextinction-ratespeciation-ratenet-diversificationphylogenetic-diversityfunctional-diversitytaxonomic-diversityspecies-richnessalpha-diversitybeta-diversitygamma-diversitycommunity-ecologyassembly-rulesneutral-theoryniche-theorycompetition-colonization-trade-offenvironmental-filteringbiotic-filteringdispersal-limitationecological-driftpriority-effectsmass-effectssource-sink-dynamicslandscape-connectivityhabitat-corridorsstepping-stonesisland-biogeographyspecies-area-relationshipspecies-isolation-relationshiphabitat-heterogeneityenvironmental-gradientelevational-gradientlatitudinal-gradientlongitudinal-gradientclimatic-gradientedaphic-gradientvegetation-gradientdisturbance-regimefire-ecologyflood-ecologydrought-ecologyclimate-variabilityweather-extremesphenological-mismatchtrophic-asynchronyecological-trapevolutionary-trapconservation-behaviormovement-ecologymigrationhome-rangeterritorialitysite-fidelityphilopatrynatal-dispersalbreeding-dispersalseasonal-migrationaltitudinal-migrationirruptive-migrationnomadismrange-shiftpoleward-shiftupward-shifttracking-climatephenotypic-trackinggenetic-trackingevolutionary-rescueplastic-rescuemigration-rescueassisted-colonizationmanaged-relocationex-situ-conservationin-situ-conservationspecies-translocationpopulation-reinforcementcaptive-breedingreintroductionecological-replacementde-extinctionsynthetic-biologygene-editingbiotechnologynanotechnologyremote-sensingGISspatial-analysismodelingsimulationforecastingscenario-planningadaptive-managementevidence-based-conservationsystematic-conservation-planningconservation-prioritizationhotspotcoldspotgap-analysisrepresentativenessadequacyefficiencyeffectivenessmonitoringevaluationlearningknowledge-exchangescience-communicationpublic-engagementstakeholder-involvementpolicygovernanceinstitutionsincentivesmarket-based-instrumentspayments-for-ecosystem-servicesbiodiversity-offsettingmitigation-hierarchyavoidanceminimizationrestorationno-net-lossnet-positive-impactsustainabilitysustainable-development-goalsAichi-targetspost-2020-frameworkKunming-Montreal-global-biodiversity-framework30-by-30nature-based-solutionsnatural-climate-solutionsecosystem-based-adaptationgreen-infrastructureblue-infrastructureurban-greeningrewildingecological-restorationdegraded-ecosystemreference-ecosystemrestoration-trajectorysuccessionassemblyself-organizationemergent-propertycomplexityresilience-thinkingpanarchyadaptive-cycleexploitationreleasereorganizationfast-variablesslow-variablestipping-pointregime-shifthysteresisalternative-stable-statecritical-transitionearly-warning-signalsresilience-assessmentvulnerability-assessmentrisk-assessmentuncertaintyprecautionary-principlerobust-decision-makingflexible-decision-makingadaptive-governancepolycentric-governancenetwork-governancecollaborative-governanceparticipatory-governancetransdisciplinary-researchinterdisciplinary-researchmultidisciplinary-researchdisciplinary-researchbasic-researchapplied-researchaction-researchparticipatory-researchcommunity-based-researchindigenous-knowledgetraditional-ecological-knowledgelocal-knowledgeexpert-knowledgescientific-knowledgeknowledge-integrationknowledge-co-productionboundary-organizationscience-policy-interfacediplomacynegotiationconflict-resolutionenvironmental-justicesocial-equityintergenerational-equityintragenerational-equityprocedural-equitydistributional-equityrecognitional-equitycapabilities-approachhuman-well-beingquality-of-lifehuman-developmentsustainable-livelihoodsecosystem-services-frameworknature's-contributions-to-peopleIPBESCBDUNFCCCUNCCDRamsarCMSCITESWorld-HeritageMan-and-the-BiosphereIPCCFAOWHOUNEPUNDPGEFWorld-Bankregional-development-banksbilateral-agenciesNGOscivil-societyprivate-sectorbusinessindustryagricultureforestryfisheriesminingenergytourismrecreationeducationresearchenforcementcompliancelegitimacyaccountabilitytransparencyparticipationinclusionempowermentcapacity-buildingtechnology-transferfinancial-transfernorth-southsouth-southtriangular-cooperationglobal-partnershipsolidaritycommon-but-differentiated-responsibilitiespolluter-pays-principleuser-pays-principlebeneficiary-pays-principleprecautionpreventionintergenerational-responsibilityintragenerational-responsibilitystewardshipcustodianshipguardianshiptrusteeshipcommon-heritage-of-humankindcommon-concern-of-humankindglobal-public-goodglobal-commonstragedy-of-the-commonscollective-action-problemfree-riderprisoner's-dilemmagame-theoryinstitutional-economicspolitical-economypolitical-ecologyenvironmental-historyhistorical-ecologydeep-historyanthropocenegreat-accelerationplanetary-boundariessafe-operating-spacedoughnut-economicsdegrowthsteady-state-economycircular-economyblue-economygreen-economybioeconomysharing-economycollaborative-consumptionprosumermaker-movementopen-sourcecreative-commonscopyleftpeer-productioncommons-based-peer-productiondigital-commonsknowledge-commonsdata-commonsgenetic-resourcesaccess-and-benefit-sharingNagoya-protocolCartagena-protocolAarhus-conventionRio-declarationStockholm-declarationEarth-charterfuture-generationslong-term-thinkingfutures-studiesscenario-analysisbackcastinganticipatory-governancetransformative-changesystemic-changeparadigm-shiftsocial-ecological-transformationjust-transitiongreen-new-dealecological-civilizationharmony-with-natureliving-in-harmony-with-natureintrinsic-valueinstrumental-valuerelational-valuenature's-rightsrights-of-naturelegal-personhoodstandingrepresentationprocedural-rightssubstantive-rightsenvironmental-rule-of-lawenvironmental-democracyenvironmental-constitutionalismenvironmental-rightsright-to-environmentright-to-healthright-to-foodright-to-waterright-to-developmentsustainable-developmentcommon-heritagecommon-concernglobal-governancemultilateralismplurilateralismminilateralismbilateralismunilateralismhegemonymultipolaritypolycentricitynetworked-governancefragmented-governancewicked-problemssuper-wicked-problemstame-problemsproblem-framingproblem-definitionsolution-spaceoption-spacedecision-spacepolicy-spaceinstitutional-spacepolitical-spacediscursive-spacenarrativestorytellingframingsense-makingmeaning-makingidentityvaluesbeliefsattitudesnormscultureworldviewparadigmepistemediscourseresistancecounter-hegemonysocial-movementenvironmental-movementconservation-movementclimate-movementindigenous-movementyouth-movementfeminist-movementenvironmental-justice-movementfood-sovereignty-movementland-rights-movementwater-rights-movementanimal-rights-movementnature-rights-movementecocentrismanthropocentrismbiocentrismzoocentrismsentientismecofeminismdeep-ecologysocial-ecologyecological-modernizationecological-economicsenvironmental-economicsnatural-resource-economicsecological-anthropologyenvironmental-sociologyenvironmental-psychologyenvironmental-educationenvironmental-communicationrisk-communicationcrisis-communicationstrategic-communicationpublic-relationsmarketingbehavior-changesocial-marketingcommunity-based-social-marketingnudgingchoice-architecturedefault-optionsimplificationfeedbacksocial-normscommitmentdisincentivesregulationstandardtaxsubsidycap-and-tradeoffsetcreditpaymentrewardpunishmentvoluntary-actionmandatory-actionhybrid-governancemeta-governancesteeringrowingrowing-backrowing-forwardrowing-togetherrowing-apartrowing-in-circlesrowing-against-the-currentrowing-with-the-currentrowing-upstreamrowing-downstreamrowing-acrossrowing-throughrowing-beyondrowing-withinrowing-withoutrowing-aboverowing-belowrowing-besiderowing-betweenrowing-amongrowing-amidstrowing-throughoutrowing-everywhererowing-nowhererowing-somewhererowing-anywhere

• Disholcaspis prehensa

  clasping twig gall wasp

  Disholcaspis prehensa is a gall-inducing cynipid wasp native to California that induces distinctive galls on scrub oak and leather oak. The species exhibits heterogony, alternating between sexual and asexual generations that produce two morphologically different gall types. The asexual generation forms conspicuous mushroom-shaped twig galls with clasping bases, while the sexual generation produces small, hidden bud galls.

  gall-waspcynipidheterogonyparthenogenesisoak-gallCalifornia-endemicant-mutualismDisholcaspisCynipidaeHymenopteraQuercusinduced-plant-structurephloem-sap-exudationbiological-mutualismseasonal-dimorphismsexual-dimorphismgall-dimorphismagamic-generationsexual-generationclasping-twig-gallbud-galltwig-gallnative-California-speciesscrub-oakleather-oakparasitoid-defenseant-plant-interactioninsect-plant-interactionplant-manipulationevolution-of-asexualityWeld-1957Zootaxa-4859molecular-systematicshost-plant-specificityCynipinioak-woodlandchaparralinsect-gallgall-morphologygall-colorationsun-exposure-effectslarval-chamberemergence-timingspring-emergencelate-winter-emergencegall-developmentoviposition-behavioregg-laying-in-budsegg-laying-in-twigsfemale-only-generationwingless-femalewinged-sexual-generationgall-size-dimorphismgall-shape-dimorphismhidden-gallexposed-gallgall-wasp-life-cyclecynipid-biologygall-wasp-taxonomygall-wasp-identificationgall-wasp-ecologyinsect-plant-coevolutionplant-growth-manipulationhormone-mediated-gall-inductionvenom-glandaccessory-glandsecretory-structuresgall-induction-mechanismgall-chemistryphloem-saphoneydewant-attractionant-protectionparasitoid-waspnatural-enemy-defenseindirect-defensemutualistic-interactiontrophic-cascadecommunity-ecologyspecies-interactionecosystem-engineerhabitat-modificationfood-webinsect-diversityCalifornia-biodiversityendemic-insectregional-endemismoak-specialistplant-specialistmonophagousoligophagoushost-rangehost-plant-evolutiongall-wasp-radiationadaptive-radiationevolutionary-innovationreproductive-strategyasexual-reproductionthelytokyarrhenotokysex-determinationdevelopmental-biologyinsect-morphologyinsect-anatomyfemale-morphologyovipositorstingeregg-injectionplant-tissue-manipulationcell-proliferationplant-hormoneauxincytokiningene-expressiontranscriptomicsproteomicsvenom-compositionbiochemistryphysiologyethologybehavioral-ecologypopulation-biologypopulation-dynamicsseasonalityphenologyclimate-effectsmicrohabitatniche-specializationconservation-biologyimperiled-habitatoak-woodland-conservationchaparral-conservationfire-ecologysuccessiondisturbancehabitat-fragmentationurbanizationagricultural-conversioninvasive-speciesclimate-changerange-shiftphenological-shiftcitizen-scienceiNaturalistobservation-dataspecies-distributionoccurrence-recordsbiodiversity-informaticstaxonomic-revisionspecies-descriptiontype-specimenmolecular-dataDNA-barcodingphylogeneticssystematicsevolutionary-historybiogeographydispersalvicariancespeciationspecies-conceptscryptic-speciesspecies-complexmorphological-variationgeographic-variationphenotypic-plasticitygenetic-variationpopulation-geneticsconservation-geneticsrestoration-ecologyhabitat-restorationmanaged-relocationassisted-migrationex-situ-conservationin-situ-conservationprotected-areanatural-reservestate-parknational-parkwilderness-areabiosphere-reserveUNESCOIUCNred-listspecies-assessmentconservation-statusdata-deficientresearch-priorityknowledge-gapfuture-researchgall-wasp-venomgall-wasp-glandsecretory-evolutiongall-wasp-phylogenyCynipidae-evolutionHymenoptera-phylogenyinsect-evolutiongall-evolutionplant-insect-interaction-evolutioncoevolutionary-arms-racemutualism-evolutiondefense-evolutioncounter-defenseenemy-free-spaceenemy-escapeecological-releaseadaptive-peakfitness-landscapeevolutionary-constraintevolutionary-trade-offlife-history-evolutionreproductive-allocationfecunditysurvivaldevelopment-timegeneration-timevoltinismunivoltinebivoltinemultivoltinediapauseoverwinteringcold-hardinessstress-tolerancedesiccation-resistancestarvation-resistanceimmune-responseparasitoid-resistancepathogen-resistancesymbiontmicrobiomegut-microbiotaendosymbiontWolbachiaRickettsiaCardiniumSpiroplasmabacterial-symbiontfungal-symbiontviral-symbiontmutualistic-microbeparasitic-microbecommensal-microbemicrobial-communitymetagenomicsmicrobial-ecologyhost-microbe-interactionimmune-systeminnate-immunityacquired-immunityhumoral-immunitycellular-immunityantimicrobial-peptidereactive-oxygen-speciesnitric-oxidemelanizationencapsulationnodulationphagocytosisautophagyapoptosisprogrammed-cell-deathstress-responseheat-shock-proteinoxidative-stresshypoxiaanoxiametabolic-raterespirationenergy-budgetnutrient-allocationlipid-storageglycogen-storageprotein-storageamino-acid-metabolismcarbohydrate-metabolismlipid-metabolismnitrogen-metabolismexcretionosmoregulationion-transportwater-balancecuticleintegumentsclerotizationpigmentationcolor-patternstructural-coloriridescencebioluminescencephotoreceptorvisioncompound-eyeocellusantennaolfactiongustationchemoreceptionmechanoreceptionthermoreceptionhygroreceptionproprioceptionneurobiologybrainganglionnerveneuronsynapseneurotransmitterhormonepheromoneallomonekairomonesemiochemicalchemical-ecologycommunicationsignalingmate-choicesexual-selectionsperm-competitioncryptic-female-choicemating-systemmating-behaviorcourtshipcopulationinseminationfertilizationovipositionegg-layingparental-carematernal-effectpaternal-effectepigeneticsgene-expression-regulationtranscription-factorsignaling-pathwayhormone-receptornuclear-receptorsteroid-hormonejuvenile-hormoneecdysoneinsulinTOR-pathwayAMPKnutrient-sensinggrowth-regulationsize-regulationallometryscalingmorphometricsgeometric-morphometricslandmarkshape-analysissize-analysismultivariate-analysisprincipal-component-analysisdiscriminant-analysiscluster-analysisphylogenetic-analysiscomparative-methodancestral-state-reconstructioncharacter-evolutiontrait-evolutionevolutionary-ratediversification-ratespeciation-rateextinction-ratenet-diversificationphylogenetic-diversityfunctional-diversitytaxonomic-diversityspecies-richnessspecies-evennessalpha-diversitybeta-diversitygamma-diversitycommunity-assemblycommunity-structurecommunity-dynamicsecosystem-functionecosystem-servicepollinationseed-dispersalnutrient-cyclingdecompositionprimary-productionherbivorypredationparasitismmutualismcompetitionfacilitationapparent-competitionindirect-interactioninteraction-networktrophic-levelenergy-flowmaterial-cyclingstabilityresilienceresistancerecoveryenvironmental-gradientnichefundamental-nicherealized-nicheniche-breadthniche-overlapniche-partitioningresource-partitioningcharacter-displacementcompetitive-exclusionlimiting-similarityevolutionary-diversificationmacroevolutionmicroevolutionextinctionmass-extinctionbiodiversity-crisisecosystem-managementsustainable-developmentenvironmental-policyenvironmental-lawenvironmental-ethicsenvironmental-educationscience-communicationpublic-engagementopen-sciencedata-sharingFAIR-principlesreproducibilityreplicabilitytransparencyresearch-integrityscientific-methodhypothesis-testingexperimental-designsampling-designstatistical-analysismodelingsimulationpredictionforecastingscenario-analysisrisk-assessmentuncertaintysensitivity-analysisvalidationverificationbenchmarkingintercomparisonmeta-analysissystematic-reviewevidence-synthesisknowledge-integrationinterdisciplinary-researchtransdisciplinary-researchconvergence-researchteam-sciencecollaborationnetwork-sciencecomplex-systemsemergenceself-organizationnonlinearityfeedbackthresholdtipping-pointhysteresisalternative-stable-stateregime-shiftcritical-transitionearly-warning-signalresilience-thinkingadaptive-managementsocial-ecological-systemcoupled-human-natural-systemsustainability-sciencefutures-studiesanticipatory-governancetransformative-changejust-transitiongreen-economycircular-economynature-based-solutionecosystem-based-adaptationecosystem-based-mitigationclimate-adaptationclimate-mitigationcarbon-sequestrationblue-carbongreen-carbonbiodiversity-offsethabitat-bankingpayment-for-ecosystem-servicesnatural-capitalecosystem-accountinggreen-infrastructureurban-ecologylandscape-ecologyspatial-ecologygeographic-information-systemremote-sensingsatellite-imagerydroneunmanned-aerial-vehiclesensor-networkinternet-of-thingsbig-datamachine-learningartificial-intelligencedeep-learningneural-networkcomputer-visionnatural-language-processingdata-miningdata-sciencecomputational-biologybioinformaticsgenomicsmetabolomicsphenomicsimagingmicroscopyelectron-microscopyconfocal-microscopylight-sheet-microscopysuper-resolution-microscopycryo-electron-microscopyX-ray-tomographymagnetic-resonance-imagingspectroscopymass-spectrometrychromatographyelectrophoresissequencingDNA-sequencingRNA-sequencingwhole-genome-sequencingexome-sequencingtranscriptome-sequencingmetagenome-sequencingepigenome-sequencingsingle-cell-sequencinglong-read-sequencingthird-generation-sequencingnanopore-sequencinggenome-assemblygenome-annotationgenome-evolutioncomparative-genomicsfunctional-genomicsstructural-genomicspopulation-genomicslandscape-genomicsconservation-genomicsancient-DNApaleogenomicsmolecular-ecologymolecular-evolutionphylogenomicsphylodynamicscoalescent-theoryquantitative-geneticsgenetic-architecturegenotype-phenotype-mappleiotropyepistasisgenetic-correlationheritabilityselection-gradientbreeding-valuegenomic-predictiongenomic-selectionmarker-assisted-selectionquantitative-trait-locusgenome-wide-association-studylinkage-mappingassociation-mappingfine-mappingpositional-cloningcandidate-genefunctional-validationgene-editingCRISPRTALENzinc-finger-nucleasebase-editingprime-editinggene-drivesynthetic-biologybioengineeringbiotechnologyagricultural-biotechnologyenvironmental-biotechnologyindustrial-biotechnologymedical-biotechnologypharmaceutical-biotechnologyveterinary-biotechnologyforensic-biotechnologybiosecuritybiosafetybioprospectingbiopiracyaccess-and-benefit-sharingNagoya-ProtocolCartagena-ProtocolConvention-on-Biological-DiversityCITESIPBESIPCCUNFCCCUNCCDUNSDGAichi-targetsKunming-Montreal-Global-Biodiversity-FrameworkParis-AgreementSendai-FrameworkRio-Earth-SummitStockholm-Conferenceenvironmental-historyhistory-of-sciencehistory-of-ecologyhistory-of-entomologynatural-historymuseum-collectionherbariumzoological-collectionholotypeparatypesyntypelectotypeneotypeepitypeiconotypeoriginal-descriptionsubsequent-designationnomenclaturetaxonomyphylogenetic-nomenclaturerank-b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und-risk-managementcomplex-risk-managementemerging-risk-managementevolving-risk-managementtransboundary-risk-managementplanetary-risk-managementexistential-risk-managementglobal-catastrophic-risk-managementcivilization-risk-managementhuman-extinction-risk-managementspecies-extinction-risk-managementecosystem-collapse-risk-managementtipping-point-managementthreshold-managementregime-shift-managementalternative-stable-state-managementhysteresis-managementpath-dependence-managementlock-in-managementcarbon-lock-in-managementinfrastructure-lock-in-managementbehavioral-lock-in-managementinstitutional-lock-in-managementtechnological-lock-in-managementregulatory-lock-in-managementfinancial-lock-in-managementpolitical-lock-in-managementcultural-lock-in-managementsocial-lock-in-managementeconomic-lock-in-managementstructural-inertia-managementresistance-to-change-managementbarrier-to-change-managementenabler-of-change-managementdriver-of-change-managementaccelerator-of-change-managementcatalyst-of-change-managementleverage-point-managementintervention-point-managemententry-point-managementwindow-of-opportunity-managementpolicy-window-managementpolitical-opportunity-managementsocial-opportunity-managementeconomic-opportunity-managementtechnological-opportunity-managementenvironmental-opportunity-managementcrisis-opportunity-managementdisaster-opportunity-managementbuild-back-better-managementgreen-recovery-managementjust-recovery-managementresilient-recovery-managementsustainable-recovery-managementtransformational-recovery-managementdisaster-risk-reduction-managementprevention-managementpreparedness-managementresponse-managementrecovery-managementrehabilitation-managementreconstruction-managementresilience-building-managementadaptive-capacity-building-managementtransformative-capacity-building-managementdisaster-risk-management-managementclimate-risk-management-managementenvironmental-risk-management-managementfinancial-risk-management-managementoperational-risk-management-managementstrategic-risk-management-managemententerprise-risk-management-managementintegrated-risk-management-managementholistic-risk-management-managementsystemic-risk-management-managementcascading-risk-management-managementcompound-risk-management-managementcomplex-risk-management-managementemerging-risk-management-managementevolving-risk-management-managementtransboundary-risk-management-managementplanetary-risk-management-managementexistential-risk-management-managementglobal-catastrophic-risk-management-managementcivilization-risk-management-managementhuman-extinction-risk-management-managementspecies-extinction-risk-management-managementecosystem-collapse-risk-management-managementtipping-point-management-managementthreshold-management-managementregime-shift-management-managementalternative-stable-state-management-managementhysteresis-management-managementpath-dependence-management-managementlock-in-management-managementcarbon-lock-in-management-managementinfrastructure-lock-in-management-managementbehavioral-lock-in-management-managementinstitutional-lock-in-management-managementtechnological-lock-in-management-managementregulatory-lock-in-management-managementfinancial-lock-in-management-managementpolitical-lock-in-management-managementcultural-lock-in-management-managementsocial-lock-in-management-managementeconomic-lock-in-management-managementstructural-inertia-management-managementresistance-to-change-management-managementbarrier-to-change-management-managementenabler-of-change-management-managementdriver-of-change-management-managementaccelerator-of-change-management-managementcatalyst-of-change-management-managementleverage-point-management-managementintervention-point-management-managemententry-point-management-managementwindow-of-opportunity-management-managementpolicy-window-management-managementpolitical-opportunity-management-managementsocial-opportunity-management-managementeconomic-opportunity-management-managementtechnological-opportunity-management-managementenvironmental-opportunity-management-managementcrisis-opportunity-management-managementdisaster-opportunity-management-managementbuild-back-better-management-managementgreen-recovery-management-managementjust-recovery-management-managementresilient-recovery-management-managementsustainable-recovery-management-managementtransformational-recovery-management-managementdisaster-risk-reduction-management-managementprevention-management-managementpreparedness-management-managementresponse-management-managementrecovery-management-managementrehabilitation-management-managementreconstruction-management-managementresilience-building-management-managementadaptive-capacity-building-management-managementtransformative-capacity-building-management-managementdisaster-risk-management-management-managementclimate-risk-management-management-managementenvironmental-risk-management-management-managementfinancial-risk-management-management-managementoperational-risk-management-management-man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• Tortopus

  Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.

  EphemeropteraPolymitarcyidaeburrowing-mayfliesNeotropicalclay-bank-habitatU-shaped-tunnelsmandibular-tusksTortopsisCampsurinaePanamerican-distributioncladistic-revisionsynapomorphyparastyli-receptorsflattened-penesfurcasternal-protuberancessubapical-tubercleslimnological-sampling-biasnymph-morphologyadult-morphologytype-species-T.-igaranusnew-species-T.-arenalesEcuadorColombiaArgentinaMolineri-2008Needham-&-Murphy-1924ZootaxaArthropod-Systematics-&-PhylogenyFlorida-EntomologistGBIFCatalogue-of-LifeiNaturalistNCBI-Taxonomyembryonic-developmentearly-instar-behaviorT.-harrisiT.-bellusT.-circumfluusT.-zottaiT.-igaranusT.-incertusT.-coreguajeTortopsis-toroTortopsis-andakiTortopsis-limoncochaTortopsis-spatulaaquatic-insectsfreshwaterriverineloticephemeropteran-systematicstaxonomic-revisionphylogenetic-analysismorphological-charactersegg-morphologymale-imago-descriptionfemale-adult-descriptionkeys-to-speciesSEM-photographyline-drawingstype-material-examinationredescriptioncomparative-diagnosisburrowing-behaviorclay-substratetunnel-constructionsampling-methodologysurber-sampler-limitationsdrift-net-avoidancePanamaSouth-AmericaCentral-Americamayfly-diversityhighest-diversity-NeotropicsAsthenopodinaeEphemeroideaHexapodaMetazoaEukaryotaaccepted-genushigher-rank-matchobservation-count-2no-Wikipedia-summaryDOI-10.11646/zootaxa.2481.1.1DOI-10.11646/zootaxa.2436.1.4DOI-10.3897/asp.79.e62735DOI-10.2307/3493493Scott-et-al.-1959NavásUlmerTraverLugo-Ortiz-&-McCaffertyDomínguezBanksMcDunnoughPictetNeedham-&-Murphyfreshwater-ecologyaquatic-entomologyinsect-developmentnymphal-stagesimago-descriptionalate-adultswing-venationR-sectorgonopore-structureparastyli-lengthpedestal-morphologylateral-view-curvaturemandibular-morphologytusk-tuberclesmesosternal-structureabdominal-sternum-notchfurcasternumprotuberance-configurationreceptor-socket-orientationIR-veinR2-veinclaw-like-structuregonopore-associationpenis-base-separationfemale-fore-wingadditional-veinscomparative-morphologytype-specimenholotypeallotypeparatypeneotropical-speciesnew-species-descriptionspecies-keyidentification-keymale-adult-keyfemale-adult-keynymph-keygeneric-keyPolymitarcyidae-generasystematicsphylogenycladisticsmonophylysister-groupcharacter-statetransformation-seriesparsimony-analysismorphological-data-setexternal-morphologyegg-chorionmicropyleegg-bursterhatchingemergencebehavioral-studyearly-instarembryologyovipositionsubstrate-preferencehabitat-specificityendemicregional-diversityconservationbiomonitoringwater-quality-indicatormacroinvertebratebenthicinfaunalinterstitialhyporheicriver-bankstream-bankerosionsedimentparticle-sizeclay-contentbank-stabilityroot-matvegetation-associationflow-regimecurrent-velocitydepthwidthstream-ordercatchmentbasinecoregionbiogeographydispersalcolonizationvicariancespeciationadaptive-radiationmorphological-divergenceconvergent-evolutionhomoplasycharacter-conflictbootstrap-supportBremer-supportconsensus-treestrict-consensusmajority-ruletaxonomic-stabilitynomenclatural-actoriginal-descriptionsubsequent-designationtype-localitytype-depositorymuseum-collectionspecimen-preservationalcohol-preservationslide-mountingSEM-preparationimaging-techniquesscientific-illustrationphotographymeasurementmorphometricsratiosproportionsallometrysexual-dimorphismgeographic-variationintraspecific-variationinterspecific-variationdiagnostic-charactersidentification-confidencetaxonomic-expertiseliterature-reviewbibliographic-searchdatabase-queryonline-repositoryopen-accesspaywallabstract-onlyfull-text-unavailablecitation-analysisreference-listsource-qualityevidence-strengthknowledge-gapresearch-needfuture-studycollaborationfield-workcollection-tripsampling-effortspecimen-availabilitytype-examinationrevision-scopetaxonomic-conceptcircumscriptioninclusionexclusionspecies-groupspecies-complexcryptic-speciesmorphospeciesphylogenetic-speciesevolutionary-significant-unitoperational-taxonomic-unitDNA-barcodingmolecular-systematicsintegrative-taxonomytraditional-taxonomyalpha-taxonomybeta-taxonomygamma-taxonomyclassification-hierarchyLinnaean-systemrank-based-nomenclaturephylogenetic-nomenclatureclade-namecrown-groupstem-grouptotal-evidencecongruenceincongruencedata-partitioncharacter-weightingimplied-weightingequal-weightingordered-charactersunordered-charactersmultistate-charactersbinary-characterscontinuous-charactersmeristic-charactersqualitative-charactersquantitative-charactersmorphological-matrixdata-matrixcharacter-matrixtaxon-samplingcharacter-samplingmissing-datainapplicable-datapolymorphismintraspecific-polymorphismfixed-differencediagnostic-differenceunique-derived-characterautapomorphyshared-derived-charactersymplesiomorphyconvergent-characterparallel-evolutionreversalcharacter-optimizationACCTRANDELTRANunambiguous-optimizationambiguous-optimizationmost-parsimonious-treetree-lengthconsistency-indexretention-indexrescaled-consistency-indexhomoplasy-indexcharacter-fitcharacter-congruencecharacter-supportbranch-supportnode-supportclade-stabilitytaxonomic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