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Arhopalus
burnt pine longhorn beetle (A. ferus), black pine sawyer (A. rusticus)
Arhopalus is a genus of longhorn beetles (Cerambycidae: Spondylidinae: Asemini) distributed across the Northern Hemisphere, with several species introduced to the Southern Hemisphere as invasive pests of conifers. The genus comprises approximately 20 described species, with A. rusticus and A. ferus being the most extensively studied due to their economic importance. Species develop in dead or fire-damaged conifer wood, with some exhibiting strong attraction to burnt pine volatiles. Several species produce male-emitted aggregation-sex pheromones containing fuscumol and geranylacetone. Invasive populations in New Zealand, Australia, South America, and South Africa require monitoring at ports and sawmills for timber export compliance.
Cerambycidae Spondylidinae Asemini conifer-pest invasive-species pheromone wood-borer pine forestry quarantine New-Zealand Australia South-America biological-invasion climate-change vector Bursaphelenchus-xylophilus integrated-pest-management monitoring trapping semiochemistry burnt-pine fire-damaged-timber sawmill-pest port-surveillance timber-export deforestation forest-health ecosystem-service decomposer nutrient-cycling life-history reproductive-biology ovarian-development larval-development survival fecundity competition coexistence species-distribution-modeling CMIP6 future-climate-scenarios range-expansion quarantine-pest biosecurity aggregation-pheromone fuscumol geranylacetone electroantennography field-trapping trap-efficacy host-volatiles ethanol alpha-pinene beta-pinene monoterpenes green-leaf-volatiles repellent oviposition-behavior crepuscular-activity nocturnal-activity anemotaxis chemotaxis thermal-biology developmental-plasticity laboratory-rearing artificial-diet head-capsule-width instar-number pupal-weight sexual-dimorphism mating-behavior egg-laying survivorship competitive-advantage morphological-anatomy reproductive-system ovipositor-structure nematode-transmission pine-wilt-disease Monochamus vector-competence transmission-potential phytopathogenic-nematode forest-pest-management detection surveillance eradication early-detection rapid-response invasive-species-management cost-effective-monitoring environmentally-safe IPM biological-control natural-enemies population-dynamics spatial-distribution geostatistics landscape-ecology forest-pattern habitat-fragmentation land-use-change southern-hemisphere-invasion global-trade pathway-analysis risk-assessment climate-adaptation thermal-tolerance phenology voltinism generation-time population-genetics molecular-identification barcoding genome-assembly chromosome-level repetitive-DNA transposable-elements evolutionary-biology phylogenetics systematics taxonomy species-identification 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Lygus hesperus
Western Tarnished Plant Bug
Lygus hesperus is a significant agricultural pest in western North America, causing extensive damage to cotton, strawberries, alfalfa seed, and other crops. In California alone, annual losses exceed $30 million in cotton and $40 million in strawberries. Adults overwinter in reproductive diapause triggered by short day lengths, resuming activity when conditions improve. The species has been the subject of extensive research on sampling methods, biological control, and insecticide resistance.
agricultural-pest plant-bug Miridae cotton-pest strawberry-pest alfalfa-pest western-North-America diapause biological-control integrated-pest-management insecticide-resistance mark-release-recapture salivary-enzymes extraoral-digestion Peristenus-relictus Hippodamia-convergens drone-sampling fluorescent-marking SmartWater vector Pantoea-ananatis Serratia potassium-chloride deterrent artificial-diet lentil seed-crops California Texas Arizona Nevada Washington Mexico polyphagous phytophagous stylet-probing scutellum nymph-spots economic-entomology organic-agriculture ecosystem-services bird-predation molecular-gut-analysis ELISA immunomarking parasitoid population-suppression crop-protection sampling-protocols sweep-net drop-cloth Y-tube-olfactometer gustatory-deterrent visual-cues volatile-cues reproductive-diapause postdiapause oogenesis oviposition longevity body-mass life-history mass-spectrometry salivary-proteins polygalacturonase α-amylase protease laccase glucose-dehydrogenase xanthine-dehydrogenase plant-defense host-pathogen-interaction iDrone-Bee UAV unmanned-aerial-vehicle open-source field-scouting IPM economic-threshold Bt-cotton natural-enemy conservation-biological-control landscape-ecology habitat-management weed-management border-sprays spot-treatment selective-insecticide neonicotinoid pyrethroid resistance-management sustainable-agriculture food-security economic-impact simulation-model population-dynamics immigration parasitism averted-loss value-loss cost-benefit research-methodology insect-marking dispersal fluorophore autofluorescence survivorship mortality mark-transfer crowding olfactometer feeding-behavior no-choice-assay choice-assay gustation host-selection plant-quality succulent-growth fruiting-structures square boll bloom terminal defoliation stand-loss dirty-bloom puckered-petals lesion carpel-wall lint-damage seed-damage yield-loss fiber-quality management-threshold scouting-interval action-threshold treatment-timing 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plant-microbe-insect endosymbiont gut-microbiome salivary-microbiome vector-competence transmission-efficiency acquisition inoculation retention multiplication circulative non-circulative persistent non-persistent propagative stylet-borne foregut-borne hemolymph-borne salivary-gland-borne replication barrier midgut salivary-gland escape infection incubation latent-period inoculation-threshold transmission-threshold epidemic epiphytotic disease-triangle susceptible-host virulent-pathogen favorable-environment disease-cycle primary-inoculum secondary-inoculum overwintering-pathogen survival-structure disease-reservoir alternative-host volunteer-crop weed-reservoir crop-residue soilborne seedborne insect-borne management-tactic preventive curative eradicative suppressive protectant systemic contact translaminar fumigant biological-pesticide microbial-pesticide entomopathogenic-fungus entomopathogenic-nematode entomopathogenic-virus baculovirus granulovirus nucleopolyhedrovirus bacteria Bacillus-thuringiensis Bt-toxin crystal-protein cytolytic-protein vegetative-insecticidal-protein mode-of-action gut-paralysis septicemia toxemia antibiotic symbiont antibiotic-resistance fitness-cost resistance-allele resistance-mechanism metabolic-resistance target-site-resistance penetration-resistance behavioral-resistance cross-resistance multiple-resistance resistance-evolution selection-pressure dose-response lethal-concentration lethal-dose median-lethal-concentration median-lethal-dose confidence-interval probit-analysis logit-analysis dose-mortality time-mortality resistance-ratio resistance-factor diagnostic-dose discriminating-dose baseline-susceptibility susceptible-strain reference-strain field-population resistant-population homozygous-resistant heterozygous dominance recessive incomplete-dominance gene-frequency allele-frequency selection-coefficient fitness relative-fitness population-genetics quantitative-genetics heritability genetic-correlation breeding-value 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consumer-waste plate-waste kitchen-waste food-rescue food-recovery food-redistribution food-banking composting anaerobic-digestion bioenergy biofuel bioproduct biomaterial biorefinery circular-economy circular-bioeconomy green-economy blue-economy bioeconomy nature-based-solution ecosystem-based-adaptation ecosystem-based-mitigation climate-mitigation disaster-risk-reduction extreme-event flood heat-wave cold-snap frost hail windstorm tornado hurricane typhoon cyclone storm-surge sea-level-rise salinization desertification land-degradation soil-degradation soil-erosion soil-compaction soil-sealing soil-contamination soil-pollution heavy-metal persistent-organic-pollutant pesticide-residue antibiotic-residue hormone-residue plastic-pollution microplastic nanoplastic particulate-matter air-pollution water-pollution groundwater-pollution surface-water-pollution eutrophication hypoxia dead-zone biodiversity-loss habitat-loss habitat-fragmentation habitat-degradation habitat-destruction invasive-species introduced-species alien-species non-native-species exotic-species naturalized-species casual-species escaped-species cultivated-species ornamental-species pet-species livestock poultry aquaculture fishery capture-fishery marine-fishery freshwater-fishery inland-fishery recreational-fishery subsistence-fishery artisanal-fishery industrial-fishery overfishing bycatch discards ghost-fishing fishing-gear fishing-method fishing-effort fishing-capacity fishing-pressure stock-assessment fishery-management marine-protected-area no-take-zone marine-reserve terrestrial-protected-area national-park nature-reserve wildlife-sanctuary biosphere-reserve world-heritage-site ramsar-site important-bird-area key-biodiversity-area ecological-network protected-area-network conservation-corridor wildlife-corridor habitat-corridor stepping-stone buffer-zone transition-zone core-zone management-zone sustainable-use-zone restoration-zone rehabilitation-zone reclamation-zone remediation-zone containment-zone eradication-zone surveillance-zone monitoring-zone quarantine-zone biosecurity phytosanitary veterinary animal-health plant-health one-health ecohealth planetary-health global-health public-health environmental-health occupational-health food-safety feed-safety water-safety air-quality environmental-quality environmental-monitoring environmental-assessment environmental-impact-assessment strategic-environmental-assessment cumulative-impact-assessment social-impact-assessment health-impact-assessment hazard-assessment exposure-assessment dose-response-assessment risk-characterization risk-management risk-communication risk-perception risk-governance precautionary-principle polluter-pays-principle user-pays-principle extended-producer-responsibility corporate-social-responsibility environmental-stewardship sustainable-livelihood sustainable-consumption sustainable-production green-procurement green-supply-chain green-logistics green-manufacturing green-chemistry green-technology clean-technology environmental-technology pollution-prevention waste-minimization resource-efficiency energy-efficiency water-efficiency material-efficiency land-use-efficiency nutrient-use-efficiency nitrogen-use-efficiency phosphorus-use-efficiency potassium-use-efficiency micronutrient-use-efficiency fertilizer-use-efficiency pesticide-use-efficiency water-productivity land-productivity labor-productivity capital-productivity total-factor-productivity partial-factor-productivity agricultural-productivity crop-productivity livestock-productivity fishery-productivity forestry-productivity economic-productivity social-productivity environmental-productivity ecological-productivity biological-productivity primary-productivity secondary-productivity net-primary-productivity gross-primary-productivity ecosystem-productivity landscape-productivity regional-productivity global-productivity carbon-productivity nitrogen-productivity water-footprint carbon-footprint nitrogen-footprint phosphorus-footprint biodiversity-footprint material-footprint land-footprint energy-footprint emission-factor intensity-factor decoupling absolute-decoupling relative-decoupling resource-decoupling impact-decoupling dematerialization rematerialization transmaterialization substitution complementarity trade-off synergy co-benefit win-win lose-lose win-lose zero-sum positive-sum negative-sum game-theory prisoner's-dilemma tragedy-of-the-commons collective-action common-pool-resource public-good private-good club-good common-good global-common global-public-good tipping-point critical-transition regime-shift hysteresis irreversibility path-dependence lock-in technological-lock-in institutional-lock-in behavioral-lock-in social-ecological-trap poverty-trap degradation-trap development-trap complex-adaptive-system social-ecological-system coupled-human-natural-system human-environment-system adaptive-capacity transformative-capacity learning-capacity innovation-capacity governance-capacity institutional-capacity organizational-capacity individual-capacity community-capacity social-capital human-capital natural-capital physical-capital financial-capital built-capital cultural-capital political-capital intellectual-capital knowledge-capital information-capital data-capital digital-capital technological-capital infrastructural-capital institutional-capital organizational-capital network-capital relationship-capital reputational-capital brand-capital symbolic-capital existential-capital spiritual-capital religious-capital moral-capital ethical-capital trust-capital reciprocity-capital solidarity-capital cooperation-capital coordination-capital collaboration-capital communication-capital information-sharing knowledge-sharing learning innovation experimentation transformation transition change evolution development growth progress improvement enhancement optimization maximization minimization satisfaction sufficiency abundance scarcity security insecurity vulnerability robustness stability instability equilibrium disequilibrium homeostasis allostasis stress distress eustress coping coping-capacity coping-strategy coping-mechanism coping-resource coping-support social-support emotional-support instrumental-support informational-support appraisal-support network-support community-support institutional-support policy-support financial-support technical-support technological-support infrastructural-support capacity-building capacity-development capacity-strengthening empowerment enablement mobilization activation engagement participation involvement inclusion exclusion marginalization discrimination inequity inequality disparity gap divide digital-divide knowledge-divide information-divide science-divide research-divide innovation-divide technology-divide development-divide north-south-divide east-west-divide urban-rural-divide gender-divide age-divide class-divide income-divide wealth-divide asset-divide opportunity-divide outcome-divide wellbeing-divide happiness-divide life-satisfaction-divide health-divide education-divide skill-divide competency-divide capability-divide functioning-divide agency-divide freedom-divide choice-divide autonomy-divide self-determination-divide dignity-divide respect-divide recognition-divide status-divide power-divide influence-divide voice-divide representation-divide participation-divide engagement-divide citizenship-divide belonging-divide identity-divide culture-divide value-divide belief-divide norm-divide practice-divide behavior-divide habit-divide custom-divide tradition-divide modernity-divide globalization-divide localization-divide universalism-divide particularism-divide cosmopolitanism-divide nationalism-divide patriotism-divide populism-divide elitism-divide democracy-divide authoritarianism-divide control-divide security-divide liberty-divide privacy-divide surveillance-divide transparency-divide accountability-divide responsibility-divide liability-divide guilt-divide blame-divide punishment-divide reward-divide incentive-divide motivation-divide intention-divide attention-divide perception-divide cognition-divide emotion-divide affect-divide attitude-divide preference-divide interest-divide need-divide want-divide desire-divide demand-divide supply-divide market-divide exchange-divide transaction-divide contract-divide agreement-divide cooperation-divide competition-divide conflict-divide coercion-divide force-divide violence-divide peace-divide harmony-divide coexistence-divide tolerance-divide acceptance-divide rejection-divide opposition-divide resistance-divide conformity-divide deviance-divide normality-divide abnormality-divide illness-divide disease-divide disorder-divide disability-divide impairment-divide handicap-divide capacity-divide potential-divide performance-divide achievement-divide attainment-divide accomplishment-divide success-divide failure-divide excellence-divide mediocrity-divide average-divide standard-divide benchmark-divide threshold-divide ceiling-divide floor-divide limit-divide boundary-divide border-divide frontier-divide horizon-divide perspective-divide viewpoint-divide standpoint-divide position-divide stance-divide approach-divide method-divide technique-divide tool-divide instrument-divide device-divide apparatus-divide equipment-divide machine-divide machinery-divide mechanism-divide system-divide structure-divide organization-divide institution-divide regime-divide order-divide arrangement-divide configuration-divide pattern-divide design-divide plan-divide scheme-divide program-divide project-divide initiative-divide intervention-divide action-divide activity-divide operation-divide process-divide procedure-divide protocol-divide guideline-divide recommendation-divide requirement-divide regulation-divide rule-divide law-divide policy-divide strategy-divide tactic-divide objective-divide goal-divide target-divide aim-divide purpose-divide mission-divide vision-divide principle-divide ethic-divide moral-divide criterion-divide indicator-divide measure-divide metric-divide index-divide scale-divide score-divide rating-divide ranking-divide grade-divide level-divide stage-divide phase-divide step-divide progression-divide sequence-divide series-divide succession-divide chain-divide cycle-divide loop-divide spiral-divide wave-divide pulse-divide rhythm-divide tempo-divide pace-divide speed-divide velocity-divide acceleration-divide deceleration-divide momentum-divide inertia-divide energy-divide work-divide effort-divide strain-divide stress-divide pressure-divide tension-divide compression-divide expansion-divide contraction-divide deformation-divide distortion-divide transformation-divide conversion-divide transduction-divide transmission-divide transfer-divide transport-divide movement-divide motion-divide flow-divide flux-divide current-divide stream-divide river-divide tide-divide wind-divide storm-divide weather-divide climate-divide season-divide year-divide decade-divide century-divide millennium-divide epoch-divide era-divide period-divide time-divide duration-divide interval-divide span-divide stretch-divide spell-divide bout-divide episode-divide event-divide occurrence-divide incident-divide accident-divide coincidence-divide chance-divide luck-divide fortune-divide destiny-divide fate-divide karma-divide dharma-divide tao-divide way-divide path-divide road-divide route-divide course-divide direction-divide orientation-divide alignment-divide positioning-divide placement-divide location-divide site-divide place-divide spot-divide point-divide situation-divide context-divide environment-divide setting-divide scene-divide platform-divide base-divide foundation-divide ground-divide soil-divide earth-divide land-divide terrain-divide territory-divide domain-divide realm-divide sphere-divide world-divide globe-divide planet-divide universe-divide cosmos-divide space-divide void-divide emptiness-divide fullness-divide plenum-divide vacuum-divide atmosphere-divide air-divide gas-divide vapor-divide mist-divide fog-divide cloud-divide rain-divide snow-divide ice-divide frost-divide dew-divide humidity-divide moisture-divide wetness-divide dryness-divide aridity-divide drought-divide flood-divide deluge-divide inundation-divide submersion-divide immersion-divide saturation-divide supersaturation-divide condensation-divide evaporation-divide transpiration-divide sublimation-divide deposition-divide precipitation-divide infiltration-divide percolation-divide seepage-divide leakage-divide discharge-divide runoff-divide erosion-divide sedimentation-divide accretion-divide aggregation-divide coagulation-divide flocculation-divide crystallization-divide solidification-divide liquefaction-divide vaporization-divide boiling-divide melting-divide freezing-divide thawing-divide cooling-divide heating-divide warming-divide thermal-divide temperature-divide heat-divide cold-divide cool-divide warm-divide hot-divide thermodynamic-divide enthalpy-divide entropy-divide free-energy-divide Gibbs-free-energy-divide Helmholtz-free-energy-divide chemical-potential-divide fugacity-divide partial-pressure-divide concentration-divide dilution-divide solution-divide mixture-divide compound-divide element-divide atom-divide molecule-divide ion-divide electron-divide proton-divide neutron-divide nucleus-divide particle-divide quark-divide lepton-divide boson-divide fermion-divide hadron-divide baryon-divide meson-divide photon-divide gluon-divide weak-boson-divide Higgs-boson-divide string-divide brane-divide dimension-divide space-time-divide continuum-divide discrete-divide quantum-divide classical-divide relativistic-divide Newtonian-divide Einsteinian-divide Planckian-divide cosmological-divide astronomical-divide microscopic-divide macroscopic-divide mesoscopic-divide nanoscopic-divide atomic-divide molecular-divide cellular-divide tissue-divide organ-divide organism-divide population-divide community-divide ecosystem-divide biome-divide biosphere-divide ecosphere-divide noosphere-divide technosphere-divide anthroposphere-divide sociosphere-divide politosphere-divide econosphere-divide culturosphere-divide religiosphere-divide ideosphere-divide memesphere-divide infosphere-divide cybersphere-divide digisphere-divide virtual-divide analog-divide real-divide actual-divide possible-divide impossible-divide necessary-divide contingent-divide probable-divide improbable-divide certain-divide uncertain-divide determinate-divide indeterminate-divide definite-divide indefinite-divide specific-divide general-divide particular-divide universal-divide singular-divide plural-divide individual-divide collective-divide social-divide cultural-divide historical-divide political-divide economic-divide religious-divide philosophical-divide scientific-divide technological-divide artistic-divide literary-divide musical-divide visual-divide performing-divide cinematic-divide photographic-divide architectural-divide fashion-divide culinary-divide gastronomic-divide olfactory-divide gustatory-divide tactile-divide kinesthetic-divide proprioceptive-divide vestibular-divide auditory-divide somatosensory-divide interoceptive-divide exteroceptive-divide multisensory-divide crossmodal-divide synesthetic-divide anesthetic-divide esthetic-divide aesthetic-divide poetic-divide rhetorical-divide logical-divide mathematical-divide statistical-divide computational-divide algorithmic-divide heuristic-divide systematic-divide methodical-divide empirical-divide theoretical-divide conceptual-divide abstract-divide concrete-divide practical-divide applied-divide pure-divide fundamental-divide basic-divide strategic-divide tactical-divide operational-divide administrative-divide managerial-divide executive-divide leadership-divide followership-divide membership-divide partnership-divide ownership-divide stewardship-divide guardianship-divide trusteeship-divide custodianship-divide wardship-divide tutelage-divide mentorship-divide sponsorship-divide championship-divide friendship-divide relationship-divide kinship-divide companionship-divide fellowship-divide scholarship-divide apprenticeship-divide internship-divide residency-divide assistantship-divide associateship-divide authorship-divide editorship-divide curatorship-divide directorship-divide governorship-divide presidency-divide chancellorship-divide deanship-divide headship-divide chairmanship-divide
Pentodontini
rhinoceros beetles
Pentodontini is the most diverse tribe within the subfamily Dynastinae (rhinoceros beetles), containing over 100 genera distributed across multiple biogeographic regions. Most genera are restricted to a single biogeographic region. The tribe is characterized by substantial morphological diversity, with generic-level identification often relying on mouthpart morphology in females and secondary sexual characters (horns, claw modifications, antennal club length) in males.
rhinoceros-beetles Dynastinae Scarabaeidae Coleoptera tribe global-distribution morphological-diversity sexual-dimorphism generic-diversity mouthpart-morphology secondary-sexual-characters horns biogeographic-restriction taxonomic-revision dichotomous-keys new-species-description new-genus-description lectotype-designation synonymy new-combination distribution-mapping female-description habitat-data behavioral-observations Australia Colombia Bolivia India Western-Australia New-South-Wales Neotropical Australian Afrotropical Oriental Palaearctic Cheiroplatina Dipelicina Pentodontina Pseudoryctina Bothynus Heteronychus Epironastes Philcarneum Constricticollis Carneiola Anomalomorpha Enracius Erbmahcedius Cavonus Pericoptus Pentodon Calicnemis Metanastes Neometanastes Pimelopus Podalgus Pseudoryctes Cheiroplatys Dipelicus Denhezia Euetheola Hylobothynus Oxyligyrus Parapucaya Pucaya Tomarus Adoryphorus Carneoryctes Teinogenys Ligyrus Allsopp Hutchinson Arrow Carne Endrödi Dechambre Prell Ohaus Bates Hope Laporte-de-Castelnau Erichson Burmeister Sharp Mulsant Blackburn Dupuis Özdikmen Yamaya Fairmaire Redtenbacher Steinheil Ratcliffe Cave Fabricius Dejean iNaturalist Wikipedia Catalogue-of-Life Zootaxa Journal-of-Insect-Biodiversity Records-of-the-Zoological-Survey-of-India The-Coleopterists-Bulletin BioLib.cz Wikimedia-Commons DOI 10.11646/zootaxa.4048.4.1 10.11646/zootaxa.4852.4.2 10.11646/zootaxa.5351.3.2 10.26515/rzsi/v125/i2s/2025/172964 10.11646/zootaxa.5716.4.7 10.11646/zootaxa.5072.5.2 10.11646/zootaxa.4852.4.3 10.12976/jib/2024.54.2.2 10.1649/1186.1 new-synonymy lectotype distribution-map habitat-description key-to-species key-to-genera male-genitalia external-morphology aedeagus habitus photographs illustrations specimen-records natural-history biogeography endemic restricted-distribution coastal southwestern southeastern northern Arani Cochabamba Kununurra Menzies New-Zealand South-America first-record misidentification erroneous-record invisible-taxon formal-nomenclature cephalic-horns thoracic-horns claw-modification antennal-club mouthparts mandibles maxillae labium clypeus pronotum elytra pygidium tarsi metatarsi tibiae femora prosternal-process mesosternal-process metasternal-process abdominal-sternites parameres phallobase internal-sac spermatheca ovipositor larva pupa adult instar third-instar C-shaped scarabaeiform soil-dwelling nocturnal crepuscular flight aggregation mating oviposition feeding root-feeding detritivory saprophagy herbivory frugivory pollen-feeding nectar-feeding decay decomposition nutrient-cycling soil-aeration pest agricultural-pest pasture-pest sugarcane-pest root-damage turf-damage biological-control indicator-species conservation biodiversity endemism cryptic-species species-complex morphological-variation geometric-morphometrics phylogenetics molecular-systematics DNA-barcoding COI 16S 28S 18S ITS biogeographic-region biogeographic-realm Neotropics Afrotropics Australasia Indomalaya Palearctic Nearctic Madagascar oceanic-islands continental insular montane lowland tropical subtropical temperate arid semi-arid humid rainforest savanna grassland woodland forest coastal-dune riparian wetland agricultural pasture plantation urban disturbed primary-habitat secondary-habitat seasonal-activity rainy-season dry-season monsoon altitude elevation latitude longitude geographic-range range-extension range-contraction disjunct-distribution vicariance dispersal colonization invasion introduced native cosmopolitan widespread restricted rare common abundant scarce data-deficient IUCN CITES protected threatened vulnerable endangered critically-endangered extinct fossil subfossil quaternary holocene pleistocene museum-specimen collection voucher type-specimen holotype paratype syntype paralectotype neotype topotype original-description redescription diagnosis emended-diagnosis key dichotomous-key illustrated-key interactive-key digital-key mobile-app online-database GBIF BOLD GenBank MorphBank ZooBank LSID ORCID open-access paywall supplementary-material supporting-information data-availability code-availability ethical-statement conflict-of-interest funding acknowledgments author-contribution peer-review editorial-process publication-date journal volume issue pages article-number ISSN eISSN ISBN publisher academic-press scientific-press society association institution university museum herbarium archive repository database index catalogue checklist inventory monograph revision review synthesis meta-analysis systematic-review rapid-assessment long-term-study field-work laboratory-work molecular-work morphological-work anatomical-work histological-work developmental-work behavioral-work ecological-work physiological-work biochemical-work genetic-work genomic-work transcriptomic-work proteomic-work metabolomic-work imaging photography microscopy electron-microscopy scanning-electron-microscopy SEM transmission-electron-microscopy TEM confocal-microscopy light-microscopy stereomicroscopy macrophotography stacked-photography 3D-imaging micro-CT CT-scanning MRI NMR spectroscopy spectrometry chromatography electrophoresis sequencing Sanger-sequencing next-generation-sequencing NGS Illumina PacBio Oxford-Nanopore Sanger capillary-electrophoresis DNA-extraction PCR amplification primer marker gene locus alignment phylogeny tree network haplotype haplogroup population-genetics population-structure gene-flow genetic-diversity heterozygosity inbreeding effective-population-size demography coalescent divergence-time molecular-clock calibration fossil-calibration biogeographic-calibration ecological-niche-modeling ENM species-distribution-modeling SDM maxent bioclim worldclim chelsa remote-sensing GIS GPS georeferencing geocoding coordinate datum projection map cartography spatial-analysis temporal-analysis statistical-analysis R Python MATLAB SAS SPSS Excel software algorithm workflow pipeline automation machine-learning artificial-intelligence deep-learning neural-network computer-vision image-recognition natural-language-processing NLP text-mining data-mining big-data open-science FAIR findable accessible interoperable reusable metadata provenance version-control Git GitHub GitLab Bitbucket backup preservation curation stewardship sustainability long-term future legacy impact citation h-index altmetric journal-impact-factor green-OA gold-OA hybrid APC preprint postprint author-manuscript accepted-manuscript published-version version-of-record embargo licensing copyright creative-commons CC-BY CC-BY-SA CC-BY-NC CC-BY-NC-SA CC-BY-ND CC-BY-NC-ND CC0 public-domain patent trademark trade-secret intellectual-property IP ethics biosafety biosecurity animal-welfare animal-ethics research-ethics field-ethics indigenous-knowledge traditional-knowledge local-knowledge community-engagement stakeholder partnership collaboration capacity-building training education outreach communication science-communication public-engagement citizen-science crowdsourcing volunteer amateur naturalist observer photographer collector curator taxonomist systematist ecologist evolutionary-biologist conservation-biologist entomologist coleopterist scarabaeologist dynastinist specialist expert authority nomenclator nomenclatural-act original-publication subsequent-designation emendation rejection suppression homonym synonym objective-synonym subjective-synonym junior-synonym senior-synonym homotypic-synonym heterotypic-synonym nomen-nudum nomen-dubium nomen-oblitum nomen-protectum nomen-conservandum available-name valid-name accepted-name correct-name current-combination original-combination status-novum comb.-nov.syn.-nov.sp.-nov.gen.-nov.subgen.-nov.subsp.-nov.nom.-nov.nom.-nud.nom.-dub.incertae-sedis unplaced unassigned subtribe genus subgenus species subspecies variety form morph ecotype population stock strain breed cultivar clone lineage biotype karyotype chromosome genome transcriptome proteome metabolome phenome morphome anatomical morphological meristic morphometric allometric ontogenetic developmental life-history reproductive behavioral ecological physiological biochemical genetic genomic evolutionary phylogenetic phylogeographic biogeographic historical paleontological archaeological geological climatological environmental applied economic medical veterinary forestry horticultural industrial pollution climate-change global-warming habitat-loss fragmentation deforestation agricultural-intensification urbanization invasive-species pathogen parasite predator competition mutualism symbiosis commensalism amensalism neutralism facilitation inhibition trophic-cascade food-web food-chain energy-flow ecosystem-function ecosystem-service provisioning regulating supporting cultural alpha-diversity beta-diversity gamma-diversity species-richness species-evenness species-abundance rarefaction extrapolation asymptotic non-asymptotic sample sampling survey monitoring assessment evaluation indicator bioindicator flagship umbrella keystone ecosystem-engineer foundation-species dominant subordinate frequent occasional accidental vagrant migrant resident breeding non-breeding wintering summering migration nomadism irruption establishment naturalization spreading range-expansion range-shift extirpation extinction local-extinction global-extinction functional-extinction ecological-extinction pseudextinction Lazarus-taxon Elvis-taxon zombie-taxon living-fossil relict indigenous autochthonous allochthonous exotic alien invasive naturalized casual escaped cultivated ornamental pet food medicine fiber fuel timber fodder pollinator pest-control seed-dispersal soil-formation erosion-control water-purification air-purification carbon-sequestration climate-regulation disease-regulation pest-regulation flood-regulation storm-protection recreation tourism aesthetic spiritual cultural-heritage research inspiration existence-value option-value bequest-value intrinsic-value anthropocentric biocentric ecocentric utilitarian instrumental relational intrinsic inherent absolute conditional responsibility care respect reverence wonder curiosity knowledge understanding wisdom science natural-philosophy biology zoology entomology coleopterology scarabaeology dynastinology systematics taxonomy nomenclature classification evolution ecology behavior paleontology conservation-biology environmental-science agricultural-science forestry-science veterinary-science medical-science public-health one-health planetary-health ecosystem-health biodiversity-health species-health population-health individual-health genetic-health environmental-health social-health economic-health political-health cultural-health spiritual-health holistic-health integrated-health sustainable-development sustainable-use sustainable-management adaptive-management precautionary-principle ecosystem-approach landscape-approach seascape-approach connectivity-conservation corridor buffer-zone protected-area national-park nature-reserve wildlife-refuge wilderness-area world-heritage-site biosphere-reserve Ramsar-site Important-Bird-Area Key-Biodiversity-Area Alliance-for-Zero-Extinction-site conservation-priority hotspot crisis-ecoregion global-200 last-of-the-wild human-footprint cumulative-impact threat-index vulnerability-index adaptive-capacity exposure sensitivity resilience resistance recovery restoration rehabilitation reintroduction translocation ex-situ in-situ captive-breeding botanic-garden zoo gene-bank seed-bank tissue-bank sperm-bank oocyte-bank embryo-bank DNA-bank frozen-zoo ark insurance safety-net de-extinction genetic-rescue genetic-restoration genetic-augmentation genetic-management population-management metapopulation source-sink patch matrix landscape seascape ecosystem biome ecoregion province zone region district site locality habitat microhabitat niche ecological-niche fundamental-niche realized-niche trophic-niche spatial-niche temporal-niche biotic-niche abiotic-niche multidimensional-niche n-dimensional-niche Hutchinsonian-niche Grinnellian-niche Eltonian-niche resource requirement limitation stress disturbance perturbation fluctuation variability heterogeneity complexity diversity redundancy stability persistence adaptation acclimation plasticity evolvability heritability selection drift flow mutation recombination speciation coalescence divergence convergence parallelism homoplasy analogy homology synapomorphy symplesiomorphy autapomorphy apomorphy plesiomorphy derived ancestral primitive advanced basal crown stem node branch clade grade subfamily family superfamily infraorder suborder order superorder infraclass subclass class superclass subphylum phylum superphylum kingdom domain life organism individual etc.
Tortopus
Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.
Ephemeroptera Polymitarcyidae burrowing-mayflies Neotropical clay-bank-habitat U-shaped-tunnels mandibular-tusks Tortopsis Campsurinae Panamerican-distribution cladistic-revision synapomorphy parastyli-receptors flattened-penes furcasternal-protuberances subapical-tubercles limnological-sampling-bias nymph-morphology adult-morphology type-species-T.-igaranus new-species-T.-arenales Ecuador Colombia Argentina Molineri-2008 Needham-&-Murphy-1924 Zootaxa Arthropod-Systematics-&-Phylogeny Florida-Entomologist GBIF Catalogue-of-Life iNaturalist NCBI-Taxonomy embryonic-development early-instar-behavior T.-harrisi T.-bellus T.-circumfluus T.-zottai T.-igaranus T.-incertus T.-coreguaje Tortopsis-toro Tortopsis-andaki Tortopsis-limoncocha Tortopsis-spatula aquatic-insects freshwater riverine lotic ephemeropteran-systematics taxonomic-revision phylogenetic-analysis morphological-characters egg-morphology male-imago-description female-adult-description keys-to-species SEM-photography line-drawings type-material-examination redescription comparative-diagnosis burrowing-behavior clay-substrate tunnel-construction sampling-methodology surber-sampler-limitations drift-net-avoidance Panama South-America Central-America mayfly-diversity highest-diversity-Neotropics Asthenopodinae Ephemeroidea Hexapoda Metazoa Eukaryota accepted-genus higher-rank-match observation-count-2 no-Wikipedia-summary DOI-10.11646/zootaxa.2481.1.1 DOI-10.11646/zootaxa.2436.1.4 DOI-10.3897/asp.79.e62735 DOI-10.2307/3493493 Scott-et-al.-1959 Navás Ulmer Traver Lugo-Ortiz-&-McCafferty Domínguez Banks McDunnough Pictet Needham-&-Murphy freshwater-ecology aquatic-entomology insect-development nymphal-stages imago-description alate-adults wing-venation R-sector gonopore-structure parastyli-length pedestal-morphology lateral-view-curvature mandibular-morphology tusk-tubercles mesosternal-structure abdominal-sternum-notch furcasternum protuberance-configuration receptor-socket-orientation IR-vein R2-vein claw-like-structure gonopore-association penis-base-separation female-fore-wing additional-veins comparative-morphology type-specimen holotype allotype paratype neotropical-species new-species-description species-key identification-key male-adult-key female-adult-key nymph-key generic-key Polymitarcyidae-genera systematics phylogeny cladistics monophyly sister-group character-state transformation-series parsimony-analysis morphological-data-set external-morphology egg-chorion micropyle egg-burster hatching emergence behavioral-study early-instar embryology oviposition substrate-preference habitat-specificity endemic regional-diversity conservation biomonitoring water-quality-indicator macroinvertebrate benthic infaunal interstitial hyporheic river-bank stream-bank erosion sediment particle-size clay-content bank-stability root-mat vegetation-association flow-regime current-velocity depth width stream-order catchment basin ecoregion biogeography dispersal colonization vicariance speciation adaptive-radiation morphological-divergence convergent-evolution homoplasy character-conflict bootstrap-support Bremer-support consensus-tree strict-consensus majority-rule taxonomic-stability nomenclatural-act original-description subsequent-designation type-locality type-depository museum-collection specimen-preservation alcohol-preservation slide-mounting SEM-preparation imaging-techniques scientific-illustration photography measurement morphometrics ratios proportions allometry sexual-dimorphism geographic-variation intraspecific-variation interspecific-variation diagnostic-characters identification-confidence taxonomic-expertise literature-review bibliographic-search database-query online-repository open-access paywall abstract-only full-text-unavailable citation-analysis reference-list source-quality evidence-strength knowledge-gap research-need future-study collaboration field-work collection-trip sampling-effort specimen-availability type-examination revision-scope taxonomic-concept circumscription inclusion exclusion species-group species-complex cryptic-species morphospecies phylogenetic-species evolutionary-significant-unit operational-taxonomic-unit DNA-barcoding molecular-systematics integrative-taxonomy traditional-taxonomy alpha-taxonomy beta-taxonomy gamma-taxonomy classification-hierarchy Linnaean-system rank-based-nomenclature phylogenetic-nomenclature clade-name crown-group stem-group total-evidence congruence incongruence data-partition character-weighting implied-weighting equal-weighting ordered-characters unordered-characters multistate-characters binary-characters continuous-characters meristic-characters qualitative-characters quantitative-characters morphological-matrix data-matrix character-matrix taxon-sampling character-sampling missing-data inapplicable-data polymorphism intraspecific-polymorphism fixed-difference diagnostic-difference unique-derived-character autapomorphy shared-derived-character symplesiomorphy convergent-character parallel-evolution reversal character-optimization ACCTRAN DELTRAN unambiguous-optimization ambiguous-optimization most-parsimonious-tree tree-length consistency-index retention-index rescaled-consistency-index homoplasy-index character-fit character-congruence character-support branch-support node-support clade-stability taxonomic-stability-index phylogenetic-signal noise saturation substitution-model rate-heterogeneity among-site-rate-variation gamma-distribution invariant-sites model-selection Akaike-information-criterion Bayesian-information-criterion likelihood-ratio-test Bayesian-inference Markov-chain-Monte-Carlo posterior-probability prior-probability hyperprior convergence effective-sample-size burn-in stationarity trace-plot parameter-estimate credible-interval highest-posterior-density tree-topology tree-shape lineage-through-time diversification-rate extinction-rate speciation-rate net-diversification birth-death-model Yule-model coalescent-model population-size effective-population-size gene-flow migration-rate isolation-by-distance phylogeography historical-biogeography ancestral-area-reconstruction dispersal-vicariance-analysis decisive-weighting event-based-method taxon-area-cladogram area-cladogram biogeographic-homology track-analysis panbiogeography phylogenetic-biogeography ecological-biogeography dispersal-biogeography vicariance-biogeography center-of-origin area-of-endemism refugium Pleistocene glaciation climate-change habitat-fragmentation habitat-loss anthropogenic-impact pollution eutrophication sedimentation dam-construction water-abstraction flow-modification thermal-pollution toxic-substances heavy-metals pesticides urbanization agriculture deforestation land-use-change riparian-degradation channelization dredging mining invasive-species biological-invasion competition predation parasitism mutualism commensalism amensalism facilitation ecosystem-engineer keystone-species flagship-species umbrella-species indicator-species sentinel-species bioindicator ecological-indicator environmental-indicator water-quality-index biotic-index multimetric-index rapid-bioassessment protocol standardization quality-assurance quality-control reference-condition least-disturbed minimally-disturbed best-available type-specific region-specific stream-type river-type functional-feeding-group collector-gatherer collector-filterer scraper shredder predator parasite engulfer piercer burrower sprawler clinger swimmer skater diver surface-dweller benthic-dweller interstitial-dweller hyporheic-dweller phreatic-dweller troglobite stygobite cave-inhabitant spring-inhabitant headwater mid-order high-order lowland montane alpine subtropical tropical temperate boreal arctic antarctic palearctic nearctic afrotropical oriental australian oceanic cosmopolitan widespread restricted relict disjunct-distribution range-extension new-record first-record country-record state-record province-record locality-record elevation-record altitudinal-range latitudinal-range longitudinal-range distribution-pattern range-size range-fill extent-of-occurrence area-of-occupancy IUCN-category threat-status data-deficient not-evaluated least-concern near-threatened vulnerable endangered critically-endangered extinct-in-wild extinct population-trend population-density population-structure age-structure size-structure sex-ratio reproductive-output fecundity fertility mortality survivorship life-table cohort-analysis generation-time doubling-time intrinsic-rate-of-increase finite-rate-of-increase net-reproductive-rate stable-age-distribution stationary-population expanding-population declining-population source-sink-dynamics metapopulation population-viability-analysis minimum-viable-population genetic-diversity heterozygosity allelic-richness private-allele founder-effect bottleneck genetic-drift inbreeding inbreeding-depression outbreeding outbreeding-depression hybridization introgression gene-flow-barrier reproductive-isolation prezygotic-isolation postzygotic-isolation mechanical-isolation temporal-isolation habitat-isolation behavioral-isolation gametic-isolation zygotic-mortality hybrid-inviability hybrid-sterility hybrid-breakdown reinforcement character-displacement ecological-speciation allopatric-speciation sympatric-speciation parapatric-speciation peripatric-speciation centrifugal-speciation divergence-with-gene-flow speciation-continuum anagenesis cladogenesis budding-speciation branching-speciation phyletic-gradualism punctuated-equilibrium stasis trend progression orthogenesis aristogenesis mosaic-evolution co-evolution arms-race Red-Queen-hypothesis evolutionary-stable-strategy frequency-dependent-selection density-dependent-selection r-selection K-selection bet-hedging iteroparity semelparity ploidy-level chromosome-number karyotype genome-size transposable-element gene-duplication neofunctionalization subfunctionalization pseudogene regulatory-evolution cis-regulatory trans-regulatory enhancer promoter silencer insulator transcription-factor signaling-pathway developmental-pathway gene-network epigenetic-modification DNA-methylation histone-modification chromatin-remodeling genomic-imprinting parental-effect maternal-effect paternal-effect cytoplasmic-inheritance mitochondrial-DNA chloroplast-DNA endosymbiont Wolbachia Rickettsia Spiroplasma Cardinium Arsenophonus Hamiltonella Regiella Serratia Buchnera Carsonella Portiera primary-endosymbiont secondary-endosymbiont facultative-symbiont obligate-symbiont mutualistic-symbiont parasitic-symbiont commensal-symbiont pathogen vector disease-transmission zoonosis emerging-infectious-disease biosecurity quarantine eradication control management conservation-biology restoration-ecology habitat-restoration river-restoration stream-restoration floodplain-restoration riparian-restoration wetland-restoration ecosystem-restoration passive-restoration active-restoration assisted-recovery managed-relocation assisted-colonization ex-situ-conservation in-situ-conservation protected-area national-park nature-reserve wildlife-refuge biosphere-reserve world-heritage-site ramsar-site important-bird-area key-biodiversity-area Alliance-for-Zero-Extinction EDGE-species evolutionarily-distinct globally-endangered conservation-prioritization systematic-conservation-planning Marxan Zonation conservation-opportunity conservation-cost conservation-benefit trade-off socioeconomic-factor stakeholder-engagement community-based-conservation adaptive-management monitoring-program long-term-study citizen-science public-participation environmental-education outreach science-communication knowledge-transfer capacity-building institutional-strengthening policy-development legislation regulation enforcement compliance incentive payment-for-ecosystem-services carbon-credit biodiversity-offset mitigation-banking conservation-easement land-trust private-land-conservation conservation-covenant stewardship sustainable-use sustainable-management ecosystem-based-management integrated-water-resources-management basin-management catchment-management landscape-management seascape-management connectivity-conservation corridor stepping-stone buffer-zone matrix-management countryside-biogeography reconciliation-ecology novel-ecosystem designer-ecosystem ecosystem-service provisioning-service regulating-service supporting-service cultural-service food-production water-supply raw-materials genetic-resources medicinal-resources ornamental-resources air-quality-regulation climate-regulation water-regulation erosion-regulation water-purification waste-treatment disease-regulation pest-regulation pollination seed-dispersal habitat-provision nutrient-cycling soil-formation primary-production recreation tourism aesthetic-value spiritual-value educational-value scientific-value inspiration sense-of-place cultural-heritage traditional-knowledge indigenous-knowledge local-knowledge ecosystem-service-valuation economic-valuation contingent-valuation choice-experiment benefit-transfer total-economic-value use-value non-use-value direct-use indirect-use option-value bequest-value existence-value willingness-to-pay willingness-to-accept cost-benefit-analysis cost-effectiveness-analysis multi-criteria-analysis decision-support uncertainty risk-assessment scenario-analysis sensitivity-analysis robustness-analysis adaptive-capacity resilience vulnerability exposure sensitivity coping-capacity transformation social-ecological-system complex-adaptive-system tipping-point regime-shift alternative-stable-state hysteresis path-dependence legacy-effect slow-variable fast-variable panarchy adaptive-cycle exploitation release reorganization foresight anticipatory-governance transformative-change sustainability-science future-earth interdisciplinary transdisciplinary problem-oriented solution-oriented action-oriented knowledge-co-production boundary-work boundary-object boundary-organization science-policy-interface science-society-interface knowledge-brokerage knowledge-exchange knowledge-integration mode-1-science mode-2-science triple-helix quadruple-helix responsible-innovation reflexive-governance adaptive-governance collaborative-governance polycentric-governance network-governance self-organization collective-action common-pool-resource tragedy-of-the-commons social-dilemma institutional-analysis institutional-design Ostrom-design-principles monitoring sanctioning conflict-resolution nested-enterprises proportionality collective-choice constitutional-choice operational-choice collective-action-theory game-theory prisoner's-dilemma chicken-game assurance-game coordination-game evolutionary-game-theory replicator-dynamics ESS Nash-equilibrium Pareto-optimality social-optimum market-failure government-failure voluntary-failure non-market-value public-good common-good club-good private-good rivalry excludability free-rider hold-up-problem moral-hazard adverse-selection information-asymmetry transaction-cost property-right common-property state-property private-property hybrid-property bundle-of-rights usufruct access withdrawal alienation tenure-security land-tenure water-tenure resource-tenure customary-tenure statutory-tenure legal-pluralism de-facto de-jure formal informal legality legitimacy recognition deviance corruption elite-capture social-capital bonding-capital bridging-capital linking-capital trust reciprocity reputation social-network social-embeddedness structural-hole brokerage closure cohesion density centrality betweenness degree eigenvector PageRank small-world scale-free random-network regular-network hierarchical-network modularity community-structure clustering assortativity homophily heterophily preferential-attachment fitness-model network-evolution network-dynamics temporal-network multilayer-network multiplex-network bipartite-network affiliation-network one-mode-network two-mode-network adjacency-matrix edge-list graph-theory vertex edge node link tie connection path cycle tree forest component giant-component percolation phase-transition criticality self-organized-criticality power-law heavy-tail Pareto-distribution lognormal-distribution exponential-distribution Poisson-distribution binomial-distribution normal-distribution Gaussian-distribution statistical-distribution probability-distribution cumulative-distribution probability-density-function cumulative-distribution-function survival-function hazard-function moment mean median mode variance standard-deviation coefficient-of-variation skewness kurtosis quantile percentile decile quartile interquartile-range range minimum maximum outlier robust-statistic parametric-test non-parametric-test hypothesis-testing null-hypothesis alternative-hypothesis significance-level p-value confidence-interval type-I-error type-II-error statistical-power effect-size sample-size sampling-design experimental-design observational-study quasi-experiment natural-experiment randomized-controlled-trial before-after-control-impact BACI impact-assessment environmental-impact-assessment strategic-environmental-assessment cumulative-impact-assessment social-impact-assessment health-impact-assessment life-cycle-assessment carbon-footprint water-footprint ecological-footprint material-flow-analysis substance-flow-analysis energy-analysis exergy-analysis emergy-analysis input-output-analysis environmentally-extended-input-output social-accounting-matrix computable-general-equilibrium agent-based-model system-dynamics-model Bayesian-network structural-equation-model path-analysis factor-analysis principal-component-analysis cluster-analysis discriminant-analysis canonical-correspondence-analysis redundancy-analysis non-metric-multidimensional-scaling ordination classification regression-analysis linear-regression nonlinear-regression generalized-linear-model generalized-additive-model mixed-effects-model multilevel-model hierarchical-model Bayesian-hierarchical-model state-space-model hidden-Markov-model time-series-analysis spatial-analysis geostatistics kriging variogram spatial-autocorrelation Moran's-I Geary's-c Getis-Ord-Gi local-indicator-of-spatial-association LISA hot-spot cold-spot spatial-cluster spatial-outlier edge-effect boundary-effect modifiable-areal-unit-problem MAUP ecological-fallacy atomistic-fallacy cross-scale-inference upscaling downscaling scaling-law allometric-scaling fractal fractal-dimension self-similarity multi-fractal lacunarity roughness texture pattern process pattern-process-relationship neutral-model null-model randomization-test permutation-test bootstrap jackknife cross-validation model-averaging model-comparison information-criterion Bayesian-model-averaging ensemble-modeling consensus-model prediction forecast projection scenario narrative-scenario quantitative-scenario storyline shared-socioeconomic-pathway representative-concentration-pathway emission-scenario climate-scenario impact-scenario adaptation-scenario mitigation-scenario transformation-scenario backcasting forecasting trend-extrapolation expert-judgment Delphi-method participatory-modeling stakeholder-analysis influence-diagram conceptual-model mental-model system-archetype leverage-point intervention-point policy-instrument economic-instrument regulatory-instrument voluntary-instrument information-instrument market-based-instrument command-and-control-instrument subsidy tax charge fee levy deposit-refund tradable-permit emission-trading carbon-tax biodiversity-banking conditional-cash-transfer community-based-natural-resource-management joint-forest-management co-management adaptive-co-management transboundary-management international-regime global-governance multilateral-environmental-agreement convention treaty cartagena-protocol nagoya-protocol Aichi-target sustainable-development-goal SDG post-2020-global-biodiversity-framework Kunming-Montreal-global-biodiversity-framework 30-by-30-target nature-based-solution NbS ecosystem-based-adaptation ecosystem-based-mitigation green-infrastructure blue-infrastructure natural-infrastructure hybrid-infrastructure climate-smart-agriculture climate-smart-landscape climate-smart-development low-carbon-development green-economy circular-economy bioeconomy blue-economy sustainable-consumption sustainable-production life-cycle-thinking cradle-to-cradle industrial-ecology industrial-symbiosis urban-metabolism socio-ecological-metabolism material-intensity energy-intensity carbon-intensity decoupling absolute-decoupling relative-decoupling resource-efficiency eco-efficiency eco-innovation green-technology clean-technology environmental-technology appropriate-technology participatory-technology technology-transfer technology-diffusion innovation-system national-innovation-system sectoral-innovation-system technological-innovation-system socio-technical-system socio-technical-transition multi-level-perspective niche-innovation regime landscape transition-management strategic-niche-management transition-experiment living-lab real-world-laboratory urban-living-lab rural-living-lab sustainability-transition fundamental-change system-innovation directionality reflexivity responsiveness anticipation learning-by-doing doing-by-learning social-learning organizational-learning policy-learning transformative-capacity capacity-development institutional-learning evolutionary-governance institutional-change path-dependency lock-in carbon-lock-in technological-lock-in institutional-lock-in behavioral-lock-in cultural-lock-in unlocking destabilization phase-out divestment stranded-asset just-transition transition-pathway transition-scenario deep-decarbonization net-zero carbon-neutral climate-neutral nature-positive biodiversity-positive restoration-positive regenerative circular doughnut-economics planetary-boundary safe-operating-space environmental-ceiling social-foundation intergenerational-equity intragenerational-equity distributive-justice procedural-justice recognition-justice environmental-justice climate-justice energy-justice water-justice food-justice spatial-justice territorial-justice ecological-justice species-justice intersectionality adaptation mitigation sustainability sustainable-development well-being human-development capability-approach human-security environmental-security climate-security water-security food-security energy-security livelihood-security disaster-risk-reduction climate-change-adaptation climate-change-mitigation loss-and-damage climate-finance green-climate-fund adaptation-fund global-environment-facility biodiversity-finance nature-finance blended-finance impact-investment ESG environmental-social-governance sustainable-finance green-bond social-bond sustainability-bond blue-bond nature-bond biodiversity-credit offset inset nature-based-credit ecosystem-service-credit payment-for-results results-based-payment conditional-payment performance-based-payment outcome-based-payment environmental-service-payment PSA PES REDD REDD+AFOLU LULUCF NAMA NAMC NDC NAP NBSAP national-adaptation-plan national-biodiversity-strategy-and-action-plan nationally-determined-contribution nationally-appropriate-mitigation-action intended-nationally-determined-contribution long-term-low-emission-development-strategy net-zero-target carbon-budget remaining-carbon-budget emission-gap adaptation-gap finance-gap implementation-gap ambition-loop ratchet-mechanism global-stocktake transparency-framework enhanced-transparency-framework common-but-differentiated-responsibilities respective-capabilities climate-equity climate-litigation climate-governance climate-diplomacy climate-negotiation UNFCCC COP CMP CMA SBSTA SBI IPCC IPBES UNCCD CBD CITES Ramsar World-Heritage Man-and-the-Biosphere MAB UNESCO FAO WHO ILO UNDP UNEP UN-Habitat World-Bank IMF WTO OECD GEF GCF AF multilateral-development-bank regional-development-bank national-development-bank development-finance-institution export-credit-agency sovereign-wealth-fund pension-fund insurance-company asset-manager asset-owner corporate-investor retail-investor impact-investor philanthropic-investor development-partner bilateral-donor multilateral-donor private-donor foundation NGO CSO civil-society-organization non-governmental-organization community-based-organization indigenous-peoples-organization women's-organization youth-organization faith-based-organization research-organization think-tank university academic-institution knowledge-institution capacity-building-institution training-institution extension-service advisory-service consulting-firm engineering-firm technology-provider equipment-supplier service-provider contractor operator maintainer utility municipality local-government provincial-government national-government federal-government ministry department agency authority commission 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seed-commons water-commons fisheries-commons forest-commons pasture-commons land-commons urban-commons atmospheric-commons global-commons common-heritage-of-mankind common-concern-of-humankind restorative-justice multispecies-justice more-than-human anthropocene capitalocene plantationocene chthulucene symbiocene good-anthropocene great-acceleration great-transformation great-decoupling great-convergence great-divergence great-fragmentation great-integration great-transition great-work long-now deep-time futures-thinking future-studies strategic-foresight scenario-planning trend-analysis weak-signal wild-card black-swan grey-rhino blue-sky green-swan systemic-change structural-change cultural-change behavioral-change individual-change collective-change social-change organizational-change technological-change economic-change political-change ecological-change planetary-change earth-system-change biodiversity-change ocean-change cryosphere-change atmospheric-change biogeochemical-change 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