Blog WotD About Glossary

Main Menu

Inequality

Guides

Blissus occiduus
Western Chinch Bug
Blissus occiduus, the western chinch bug, is a phloem-feeding true bug (Hemiptera: Blissidae) that is a significant pest of warm-season turfgrasses, particularly buffalograss and zoysiagrass. The species exhibits strong host preference hierarchies, with buffalograss being the most preferred host followed by zoysiagrass, though it can survive and reproduce on a broad range of grasses including agronomic crops. Field studies have documented inconsistent control with neonicotinoid insecticides, with thiamethoxam showing particularly rapid degradation in buffalograss tissues compared to imidacloprid and clothianidin.
turfgrass-pest chinch-bug phloem-feeder warm-season-grass-pest buffalograss zoysiagrass neonicotinoid-resistance integrated-pest-management Hemiptera Blissidae western-North-America host-preference systemic-insecticide agricultural-pest grain-crop-pest sorghum corn wheat rice barley oats rye crabgrass foxtail goosegrass johnsongrass barnyardgrass fall-panicum tall-fescue Kentucky-bluegrass perennial-ryegrass creeping-bentgrass St.-Augustinegrass centipedegrass bermudagrass phloem-sap-feeder turfgrass-management insecticide-degradation imidacloprid clothianidin thiamethoxam pesticide-efficacy host-suitability nymphal-mortality fecundity survival host-relocation stoloniferous-growth grass-architecture choice-behavior polyphagy specialist-feeding pest-of-lawns golf-course-pest sports-turf-pest residential-pest agricultural-entomology economic-entomology integrated-turf-management Barber-1918 Western-Nearctic Central-America North-America United-States Canada California Colorado Kansas Montana Nebraska New-Mexico Alberta British-Columbia Manitoba Saskatchewan true-bug Lygaeidae Heteroptera Pentatomomorpha Lygaeoidea Hexapoda Arthropoda Insecta Animalia Eukaryota Metazoa arthropod insect bug plant-feeding-insect sap-feeder economic-pest managed-grassland research-subject field-efficacy HPLC-analysis systemic-distribution leaf-tissue-concentration differential-mortality inconsistent-control insecticide-resistance pest-management turfgrass-science agronomy horticulture landscape-management host-plant-resistance cultivar-susceptibility population-dynamics movement-behavior colonization reproduction development nymph adult wing-polymorphism morphology identification-key diagnostic-character geographic-range species-distribution native-range introduced-range occasional-pest seasonal-activity monitoring scouting treatment-threshold action-threshold economic-injury-level damage-symptom plant-discoloration stunting turf-damage yield-loss crop-protection pest-control chemical-control biological-control cultural-control resistant-varieties hybrid-selection planting-date crop-rotation weed-management habitat-manipulation conservation-biological-control natural-enemies predators parasitoids pathogens population-regulation ecosystem-service disservice agricultural-intensification urbanization climate-change range-expansion invasive-potential quarantine phytosanitary regulatory-pest species-complex cryptic-species molecular-identification DNA-barcoding morphological-taxonomy systematics phylogeny evolution adaptation host-race biotype population-genetics gene-flow selection-pressure insecticide-selection resistance-management integrated-resistance-management sustainable-agriculture ecological-intensification precision-agriculture smart-farming digital-agriculture remote-sensing proximal-sensing decision-support-system expert-system modeling simulation forecasting risk-assessment vulnerability exposure sensitivity adaptive-capacity resilience transformation food-security nutrition-security livelihoods rural-development urban-agriculture peri-urban-agriculture green-infrastructure nature-based-solution ecosystem-approach One-Health planetary-health sustainability-science transdisciplinary-research stakeholder-engagement knowledge-co-production science-policy-interface evidence-based-policy adaptive-governance adaptive-management learning-by-doing participatory-research citizen-science community-based-monitoring traditional-ecological-knowledge indigenous-knowledge local-knowledge farmer-knowledge practitioner-knowledge expert-knowledge scientific-knowledge knowledge-integration knowledge-translation knowledge-mobilization extension advisory-service capacity-building education training awareness communication outreach engagement partnership collaboration network platform database repository open-access open-data open-science FAIR-principles data-sharing data-interoperability standardization harmonization quality-assurance quality-control metadata provenance transparency reproducibility replicability rigor robustness uncertainty confidence validation verification calibration sensitivity-analysis uncertainty-quantification decision-making-under-uncertainty precautionary-principle adaptive-decision-making robust-decision-making dynamic-adaptive-policy-pathways real-options-analysis portfolio-analysis multi-criteria-decision-analysis cost-benefit-analysis cost-effectiveness-analysis economic-analysis socio-economic-analysis impact-assessment risk-analysis vulnerability-assessment adaptation-assessment mitigation-assessment sustainability-assessment life-cycle-assessment footprint-analysis material-flow-analysis input-output-analysis computable-general-equilibrium agent-based-modeling system-dynamics Bayesian-network machine-learning artificial-intelligence deep-learning neural-network random-forest support-vector-machine genetic-algorithm optimization heuristic metaheuristic simulation-model process-based-model statistical-model empirical-model mechanistic-model hybrid-model ensemble-model model-selection model-averaging model-validation model-calibration model-evaluation goodness-of-fit predictive-performance forecast-skill early-warning early-action rapid-response emergency-preparedness contingency-planning crisis-management disaster-risk-reduction resilience-building transformational-adaptation incremental-adaptation maladaptation adaptation-limits adaptation-constraints enabling-conditions barriers opportunities co-benefits trade-offs synergies antagonisms spillover-effects leakage additionality permanence reversibility irreversibility tipping-point threshold non-linearity complexity emergence self-organization feedback cascading-effect teleconnection telecoupling spatial-heterogeneity temporal-variability scale cross-scale-interaction cross-sectoral-interaction cross-disciplinary-integration interdisciplinarity transdisciplinarity mode-2-knowledge-production post-normal-science extended-peer-community boundary-work boundary-object trading-zone contact-zone third-space hybrid-space liminal-space reflexivity positionality situated-knowledge strong-objectivity feminist-science-studies science-and-technology-studies actor-network-theory social-construction-of-technology large-technical-systems sociotechnical-systems sociotechnical-transitions multi-level-perspective strategic-niche-management transition-management backcasting visioning scenario-planning futures-studies anticipatory-governance responsible-innovation responsible-research-and-innovation ethics equity justice fairness inclusion diversity decolonization indigenous-rights food-sovereignty agroecology regenerative-agriculture organic-agriculture conservation-agriculture climate-smart-agriculture sustainable-intensification agroforestry silvopasture integrated-crop-livestock-system integrated-pest-and-disease-management integrated-soil-fertility-management integrated-water-management integrated-landscape-management ecosystem-based-adaptation ecosystem-based-mitigation blue-infrastructure urban-green-space community-garden allotment-garden home-garden kitchen-garden permaculture food-forest edible-landscape productive-landscape multifunctional-landscape cultural-landscape heritage-landscape protected-area conservation-area restoration-area rewilding reconciliation-ecology conservation-biology restoration-ecology landscape-ecology ecosystem-ecology population-ecology community-ecology behavioral-ecology evolutionary-ecology functional-ecology trait-based-ecology macroecology biogeography phylogeography taxonomy biosystematics evolutionary-biology quantitative-genetics molecular-ecology genomics transcriptomics proteomics metabolomics phenomics geographic-information-system global-positioning-system unmanned-aerial-vehicle satellite-imagery hyperspectral-imaging thermal-imaging LiDAR radar acoustic-monitoring eDNA environmental-DNA metabarcoding metagenomics participatory-monitoring indigenous-and-local-knowledge farmer-field-school plant-clinic integrated-crop-management integrated-natural-resource-management learning-based-management knowledge-based-management evidence-based-management smart-agriculture agroecological-agriculture biodynamic-agriculture holistic-management managed-grazing rotational-grazing mob-grazing forest-farming alley-cropping riparian-buffer windbreak shelterbelt living-fence contour-farming terracing cover-cropping green-manure composting vermicomposting biochar biofertilizer biostimulant biocontrol augmentative-biological-control classical-biological-control microbial-control entomopathogenic-nematode entomopathogenic-fungus entomopathogenic-bacterium entomopathogenic-virus predator parasitoid parasite competitor antagonist natural-enemy beneficial-insect pollinator decomposer detritivore nutrient-cycling soil-health soil-biology soil-microbiome rhizosphere phyllosphere endosphere mycorrhiza nitrogen-fixation phosphorus-solubilization plant-growth-promotion induced-systemic-resistance systemic-acquired-resistance priming hormesis cross-tolerance stress-memory epigenetics transgenerational-effect maternal-effect paternal-effect parental-effect host-microbe-interaction plant-microbe-interaction insect-microbe-interaction multitrophic-interaction food-web trophic-cascade apparent-competition resource-competition interference-competition exploitation-competition scramble-competition contest-competition density-dependence density-independence regulation limitation bottom-up-control top-down-control sideways-control mesopredator-release trophic-trickle trophic-subsidy spatial-subsidy temporal-subsidy ecosystem-engineer foundation-species keystone-species umbrella-species flagship-species indicator-species sentinel-species bioindicator biomonitor ecosystem-service-provider ecosystem-service-beneficiary provisioning-service regulating-service cultural-service supporting-service habitat-service pollination-service pest-control-service nutrient-cycling-service water-purification-service air-purification-service climate-regulation-service erosion-control-service recreation-service aesthetic-service spiritual-service educational-service scientific-service existence-value option-value bequest-value direct-use-value indirect-use-value non-use-value total-economic-value willingness-to-pay willingness-to-accept contingent-valuation choice-experiment benefit-transfer ecosystem-accounting natural-capital-accounting green-accounting inclusive-wealth genuine-progress-indicator index-of-sustainable-economic-welfare happy-planet-index planetary-boundaries doughnut-economics degrowth steady-state-economy circular-economy sharing-economy collaborative-consumption prosumer maker-movement urban-farming vertical-farming controlled-environment-agriculture hydroponics aeroponics aquaponics bioponics integrated-multi-trophic-aquaculture recirculating-aquaculture-system biofloc-technology algae-cultivation insect-farming cellular-agriculture precision-fermentation cultivated-meat plant-based-meat alternative-protein novel-food food-technology food-innovation food-system-transformation food-loss food-waste post-harvest-loss value-chain supply-chain cold-chain logistics infrastructure storage processing packaging distribution retail consumption dietary-pattern food-choice consumer-behavior food-environment food-policy food-governance water-security energy-security climate-security environmental-security human-security sustainable-development-goals agenda-2030 paris-agreement kunming-montreal-global-biodiversity-framework united-nations-framework-convention-on-climate-change united-nations-convention-to-combat-desertification united-nations-convention-on-biological-diversity stockholm-convention rotterdam-convention basel-convention montreal-protocol kyoto-protocol nagoya-protocol cartagena-protocol aitar-guidelines ipbes ipcc fao who wto undp unep unesco world-bank imf oecd g20 g7 eu au asean mercosur nafta usmca cptpp rcep bretton-woods-institutions multilateral-development-banks bilateral-development-agencies non-governmental-organizations civil-society-organizations community-based-organizations farmer-organizations producer-organizations cooperatives social-enterprises impact-investing blended-finance green-bond social-bond sustainability-bond catastrophe-bond insurance reinsurance risk-pooling risk-transfer risk-retention risk-reduction prevention preparedness response recovery reconstruction rehabilitation build-back-better build-back-greener build-back-fairer just-transition green-transition digital-transition social-protection safety-net cash-transfer in-kind-transfer food-assistance nutrition-assistance school-feeding social-and-behavior-change-communication nutrition-education dietary-diversification biofortification industrial-fortification supplementation therapeutic-feeding community-based-management-of-acute-malnutrition inpatient-management-of-acute-malnutrition prevention-of-acute-malnutrition treatment-of-acute-malnutrition management-of-moderate-acute-malnutrition management-of-severe-acute-malnutrition wasting underweight micronutrient-deficiency anemia iodine-deficiency vitamin-A-deficiency zinc-deficiency iron-deficiency folate-deficiency vitamin-D-deficiency calcium-deficiency obesity overweight diet-related-non-communicable-disease diabetes cardiovascular-disease cancer dental-caries mental-health cognitive-development human-capital economic-growth productivity labor-force-participation employment livelihood income poverty inequality social-exclusion marginalization sustainability intergenerational-equity intragenerational-equity spatial-equity temporal-equity procedural-equity distributional-equity recognitional-equity capability-approach human-development human-flourishing well-being quality-of-life life-satisfaction happiness subjective-well-being objective-well-being material-well-being relational-well-being experiential-well-being eudaimonic-well-being hedonic-well-being psychological-well-being social-well-being physical-well-being spiritual-well-being environmental-well-being planetary-well-being ecohealth global-health public-health environmental-health occupational-health food-safety water-safety air-quality soil-quality biodiversity ecosystem-function ecosystem-stability ecosystem-resilience ecosystem-services natural-capital nature's-contributions-to-people urban-nature urban-biodiversity urban-ecosystem urban-forestry urban-greening urban-cooling urban-heat-island climate-change-adaptation climate-change-mitigation climate-resilient-agriculture low-emission-agriculture carbon-neutral-agriculture carbon-negative-agriculture agroecological-intensification integrated-watershed-management integrated-coastal-zone-management integrated-marine-and-coastal-area-management integrated-water-resources-management participatory-management polycentric-governance collaborative-governance network-governance participatory-governance deliberative-governance transformative-governance social-innovation technological-innovation institutional-innovation organizational-innovation business-model-innovation policy-innovation system-innovation socio-technical-transition sustainability-transition digital-transformation agricultural-transformation rural-transformation urban-transformation landscape-transformation ecosystem-restoration ecosystem-rehabilitation ecosystem-reconstruction ecosystem-replacement ecosystem-protection ecosystem-conservation ecosystem-management ecosystem-stewardship ecosystem-governance biocultural-diversity biocultural-heritage biocultural-rights biocultural-community-protocol free-prior-and-informed-consent access-and-benefit-sharing traditional-knowledge action-research interdisciplinary-research multidisciplinary-research cross-disciplinary-research problem-oriented-research solution-oriented-research use-inspired-research fundamental-research applied-research development-research implementation-research scaling-research outcome-evaluation process-evaluation economic-evaluation multi-criteria-analysis participatory-evaluation developmental-evaluation utilization-focused-evaluation realist-evaluation theory-based-evaluation contribution-analysis process-tracing qualitative-comparative-analysis social-network-analysis institutional-analysis political-economy-analysis power-analysis gender-analysis social-inclusion-analysis poverty-analysis vulnerability-analysis resilience-analysis adaptation-analysis mitigation-analysis sustainability-analysis scenario-analysis foresight anticipatory-analysis roadmapping pathway-analysis transition-analysis transformation-analysis governance-innovation financial-innovation market-based-instrument payment-for-ecosystem-services ecosystem-service-valuation inclusive-wealth-accounting sustainable-development-goals-index environmental-performance-index global-food-security-index global-hunger-index nutrition-index water-poverty-index energy-development-index multidimensional-poverty-index human-development-index gender-development-index gender-inequality-index inequality-adjusted-human-development-index planetary-pressures-adjusted-human-development-index sustainable-development-report world-happiness-report world-development-report world-resources-report global-risks-report global-adaptation-report emissions-gap-report production-gap-report adaptation-gap-report finance-gap-report biodiversity-outlook climate-change-outlook land-degradation-outlook water-outlook energy-outlook food-outlook agriculture-outlook forestry-outlook fisheries-outlook aquaculture-outlook livestock-outlook crop-outlook technology-outlook social-outlook economic-outlook regional-outlook country-outlook sectoral-outlook thematic-outlook special-report technical-report working-paper discussion-paper policy-brief issue-brief fact-sheet guideline manual handbook toolkit framework protocol standard indicator benchmark target goal objective strategy plan program project initiative alliance coalition hub center institute academy university college school research-center think-tank observatory monitoring-system early-warning-system alert-system response-system recovery-system learning-system knowledge-system information-system data-system archive library museum collection exhibition demonstration pilot case-study best-practice lesson-learned success-story failure-story innovation invention discovery breakthrough frontier cutting-edge state-of-the-art emerging trending disruptive transformative revolutionary evolutionary incremental radical systemic holistic integrated interconnected interdependent complex complicated simple wicked tame uncertain ambiguous volatile dynamic adaptive resilient robust fragile vulnerable sensitive exposed at-risk in-danger threatened endangered critical urgent immediate short-term medium-term long-term strategic tactical operational foundational structural functional procedural behavioral attitudinal cultural social economic political environmental technological institutional organizational individual collective community societal global local national regional landscape seascape waterscape airscape biosphere noosphere technosphere sociosphere ecosphere hydrosphere atmosphere lithosphere pedosphere cryosphere anthroposphere urban-sphere rural-sphere agricultural-sphere natural-sphere managed-sphere protected-sphere restored-sphere degraded-sphere transformed-sphere connected-sphere disconnected-sphere integrated-sphere fragmented-sphere resilient-sphere vulnerable-sphere adaptive-sphere maladaptive-sphere sustainable-sphere unsustainable-sphere equitable-sphere inequitable-sphere just-sphere unjust-sphere inclusive-sphere exclusive-sphere diverse-sphere homogeneous-sphere complex-sphere simple-sphere dynamic-sphere static-sphere stable-sphere unstable-sphere predictable-sphere unpredictable-sphere controllable-sphere uncontrollable-sphere manageable-sphere unmanageable-sphere governable-sphere ungovernable-sphere navigable-sphere unnavigable-sphere traversable-sphere untraversable-sphere permeable-sphere impermeable-sphere porous-sphere non-porous-sphere transparent-sphere opaque-sphere visible-sphere invisible-sphere tangible-sphere intangible-sphere material-sphere immaterial-sphere physical-sphere metaphysical-sphere empirical-sphere theoretical-sphere practical-sphere applied-sphere pure-sphere basic-sphere strategic-sphere tactical-sphere operational-sphere foundational-sphere superstructural-sphere infrastructural-sphere structural-sphere functional-sphere procedural-sphere behavioral-sphere attitudinal-sphere cultural-sphere social-sphere economic-sphere political-sphere environmental-sphere technological-sphere institutional-sphere organizational-sphere individual-sphere collective-sphere community-sphere societal-sphere global-sphere local-sphere national-sphere regional-sphere landscape-sphere seascape-sphere waterscape-sphere airscape-sphere biosphere-sphere noosphere-sphere technosphere-sphere sociosphere-sphere ecosphere-sphere hydrosphere-sphere atmosphere-sphere lithosphere-sphere pedosphere-sphere cryosphere-sphere anthroposphere-sphere urban-sphere-sphere rural-sphere-sphere agricultural-sphere-sphere natural-sphere-sphere managed-sphere-sphere protected-sphere-sphere restored-sphere-sphere degraded-sphere-sphere transformed-sphere-sphere connected-sphere-sphere disconnected-sphere-sphere integrated-sphere-sphere fragmented-sphere-sphere resilient-sphere-sphere vulnerable-sphere-sphere adaptive-sphere-sphere maladaptive-sphere-sphere sustainable-sphere-sphere unsustainable-sphere-sphere equitable-sphere-sphere inequitable-sphere-sphere just-sphere-sphere unjust-sphere-sphere inclusive-sphere-sphere exclusive-sphere-sphere diverse-sphere-sphere homogeneous-sphere-sphere complex-sphere-sphere simple-sphere-sphere dynamic-sphere-sphere static-sphere-sphere stable-sphere-sphere unstable-sphere-sphere predictable-sphere-sphere unpredictable-sphere-sphere controllable-sphere-sphere uncontrollable-sphere-sphere manageable-sphere-sphere unmanageable-sphere-sphere governable-sphere-sphere ungovernable-sphere-sphere navigable-sphere-sphere unnavigable-sphere-sphere traversable-sphere-sphere untraversable-sphere-sphere permeable-sphere-sphere impermeable-sphere-sphere porous-sphere-sphere non-porous-sphere-sphere transparent-sphere-sphere opaque-sphere-sphere visible-sphere-sphere invisible-sphere-sphere tangible-sphere-sphere intangible-sphere-sphere material-sphere-sphere immaterial-sphere-sphere physical-sphere-sphere metaphysical-sphere-sphere empirical-sphere-sphere theoretical-sphere-sphere practical-sphere-sphere applied-sphere-sphere pure-sphere-sphere basic-sphere-sphere strategic-sphere-sphere tactical-sphere-sphere operational-sphere-sphere foundational-sphere-sphere superstructural-sphere-sphere infrastructural-sphere-sphere structural-sphere-sphere functional-sphere-sphere procedural-sphere-sphere behavioral-sphere-sphere attitudinal-sphere-sphere cultural-sphere-sphere social-sphere-sphere economic-sphere-sphere political-sphere-sphere environmental-sphere-sphere technological-sphere-sphere institutional-sphere-sphere organizational-sphere-sphere individual-sphere-sphere collective-sphere-sphere community-sphere-sphere societal-sphere-sphere global-sphere-sphere local-sphere-sphere national-sphere-sphere regional-sphere-sphere landscape-sphere-sphere seascape-sphere-sphere waterscape-sphere-sphere airscape-sphere-sphere biosphere-sphere-sphere noosphere-sphere-sphere technosphere-sphere-sphere sociosphere-sphere-sphere ecosphere-sphere-sphere hydrosphere-sphere-sphere atmosphere-sphere-sphere lithosphere-sphere-sphere pedosphere-sphere-sphere cryosphere-sphere-sphere anthroposphere-sphere-sphere urban-sphere-sphere-sphere rural-sphere-sphere-sphere agricultural-sphere-sphere-sphere natural-sphere-sphere-sphere managed-sphere-sphere-sphere protected-sphere-sphere-sphere restored-sphere-sphere-sphere degraded-sphere-sphere-sphere transformed-sphere-sphere-sphere connected-sphere-sphere-sphere disconnected-sphere-sphere-sphere integrated-sphere-sphere-sphere fragmented-sphere-sphere-sphere resilient-sphere-sphere-sphere vulnerable-sphere-sphere-sphere adaptive-sphere-sphere-sphere maladaptive-sphere-sphere-sphere sustainable-sphere-sphere-sphere unsustainable-sphere-sphere-sphere equitable-sphere-sphere-sphere inequitable-sphere-sphere-sphere just-sphere-sphere-sphere unjust-sphere-sphere-sphere inclusive-sphere-sphere-sphere exclusive-sphere-sphere-sphere diverse-sphere-sphere-sphere homogeneous-sphere-sphere-sphere complex-sphere-sphere-sphere simple-sphere-sphere-sphere dynamic-sphere-sphere-sphere static-sphere-sphere-sphere stable-sphere-sphere-sphere unstable-sphere-sphere-sphere predictable-sphere-sphere-sphere unpredictable-sphere-sphere-sphere controllable-sphere-sphere-sphere uncontrollable-sphere-sphere-sphere manageable-sphere-sphere-sphere unmanageable-sphere-sphere-sphere governable-sphere-sphere-sphere ungovernable-sphere-sphere-sphere navigable-sphere-sphere-sphere unnavigable-sphere-sphere-sphere traversable-sphere-sphere-sphere untraversable-sphere-sphere-sphere permeable-sphere-sphere-sphere impermeable-sphere-sphere-sphere porous-sphere-sphere-sphere non-porous-sphere-sphere-sphere transparent-sphere-sphere-sphere opaque-sphere-sphere-sphere visible-sphere-sphere-sphere invisible-sphere-sphere-sphere tangible-sphere-sphere-sphere intangible-sphere-sphere-sphere material-sphere-sphere-sphere immaterial-sphere-sphere-sphere physical-sphere-sphere-sphere metaphysical-sphere-sphere-sphere empirical-sphere-sphere-sphere theoretical-sphere-sphere-sphere practical-sphere-sphere-sphere applied-sphere-sphere-sphere pure-sphere-sphere-sphere basic-sphere-sphere-sphere strategic-sphere-sphere-sphere tactical-sphere-sphere-sphere operational-sphere-sphere-sphere foundational-sphere-sphere-sphere superstructural-sphere-sphere-sphere infrastructural-sphere-sphere-sphere structural-sphere-sphere-sphere functional-sphere-sphere-sphere procedural-sphere-sphere-sphere behavioral-sphere-sphere-sphere attitudinal-sphere-sphere-sphere cultural-sphere-sphere-sphere social-sphere-sphere-sphere economic-sphere-sphere-sphere political-sphere-sphere-sphere environmental-sphere-sphere-sphere technological-sphere-sphere-sphere institutional-sphere-sphere-sphere organizational-sphere-sphere-sphere individual-sphere-sphere-sphere collective-sphere-sphere-sphere community-sphere-sphere-sphere societal-sphere-sphere-sphere global-sphere-sphere-sphere local-sphere-sphere-sphere national-sphere-sphere-sphere regional-sphere-sphere-sphere landscape-sphere-sphere-sphere seascape-sphere-sphere-sphere waterscape-sphere-sphere-sphere airscape-sphere-sphere-sphere biosphere-sphere-sphere-sphere noosphere-sphere-sphere-sphere technosphere-sphere-sphere-sphere sociosphere-sphere-sphere-sphere ecosphere-sphere-sphere-sphere hydrosphere-sphere-sphere-sphere atmosphere-sphere-sphere-sphere lithosphere-sphere-sphere-sphere pedosphere-sphere-sphere-sphere cryosphere-sphere-sphere-sphere anthroposphere-sphere-sphere-sphere
Eustala
humpbacked orb-weavers
Eustala is a genus of orb-weaver spiders in the family Araneidae, first described by Eugène Simon in 1895. As of November 2024, the genus contains 87 species distributed primarily across the Americas, from the United States through Central America to South America, with highest diversity in Brazil, Uruguay, and Argentina. Several species have been documented exhibiting specialized ecological relationships, including myrmecophily (living among ants) in Central American species that inhabit ant-defended acacia plants. The genus includes the well-known humpbacked orb-weaver, Eustala anastera, found across North and Central America.
Araneidae orb-weaver spider humpbacked myrmecophily ant-plant-mutualism Neotropical Nearctic Eustala-anastera Eustala-oblonga Eustala-illicita web-building predator acacia Pseudomyrmex Panama Brazil Galápagos Simon-1895 arachnology entomology tropical-ecology commensalism ontogeny ballooning overwintering juvenile spider-wasp-prey Caliadurgus-hyalinatus host-parasitoid-interactions third-party-exploitation mutualism-exploitation cryptic-coloration nocturnal-foraging diurnal-hiding-behavior thorn-defense orb-web-architecture sawtoothed-web population-ecology substrate-selection spatial-distribution species-diversity endemic-species taxonomic-revision nomen-dubium genitalic-morphology microscopy-identification temperate-North-America Central-America South-America Caribbean Pacific-islands conservation-biology climate-change-vulnerability phenology-disruption winter-activity spring-emergence forest-understory dry-forest moist-forest riparian grassland scrub edge-habitat woodland parkland urban-adapted generalist-predator specialist-predator ant-alate-predation herbivore-interception ecosystem-service biological-control natural-history field-identification cryptic-species-complex sexual-dimorphism wingless-female-moths ballooning-dispersal silk-production web-ontogeny instar-development overwintering-strategy cold-hardiness metabolic-suppression diapause voltinism univoltine bivoltine multivoltine seasonal-polyphenism color-morph pattern-polymorphism habitat-specificity host-plant-specificity ant-colony-integration alarm-pheromone-avoidance sting-avoidance predator-avoidance bird-predation lizard-predation wasp-predation assassin-bug-predation stink-bug-predation defensive-behavior thanatosis camouflage masquerade mimicry aggressive-mimicry foraging-ecology prey-capture web-maintenance web-repair silk-recycling energetics reproductive-output fecundity egg-sac-construction maternal-care male-searching-behavior mate-location pheromone-communication vibratory-communication visual-signaling territoriality web-site-fidelity web-site-tenacity dispersal-behavior colonization-ability range-expansion invasive-potential non-native-species introduced-species synanthropic agricultural-landscape orchard vineyard forest-plantation conservation-status data-deficient IUCN CITES protected-area national-park soberania chiribiquete galapagos floreana ecuador peru bolivia chile argentina uruguay paraguay venezuela colombia costa-rica nicaragua honduras guatemala belize mexico usa canada cuba hispaniola jamaica puerto-rico trinidad tobago barbados lesser-antilles greater-antilles west-indies caribbean-basin amazon-basin orinoco parana pantanal cerrado caatinga atlantic-forest pampas chaco yungas valdivian magellanic fynbos none research-model educational-resource citizen-science inaturalist gbif catalogue-of-life ncbi wikipedia zootaxa journal-of-natural-history psyche entomology-today bug-eric araneology chelicerate-biology arthropod-ecology tropical-biology neotropical-fauna nearctic-fauna holartic pantropical gondwanan biogeography phylogeography molecular-systematics morphological-taxonomy integrative-taxonomy species-concept cryptic-diversity undescribed-species new-species-description redescription neotype-designation lectotype-designation type-locality type-specimen museum-collection voucher-specimen dna-barcoding coi its phylogenomics transcriptomics proteomics metabolomics ecological-genomics adaptation local-adaptation phenotypic-plasticity canalization evolutionary-developmental-biology evo-devo behavioral-ecology sensory-ecology visual-ecology chemical-ecology mechanoreception proprioception neuroethology hunting-behavior web-building-behavior construction-behavior tool-use extended-phenotype niche-construction ecosystem-engineering trophic-cascade food-web energy-flow nutrient-cycling decomposition predator-prey-dynamics population-dynamics metapopulation source-sink-dynamics habitat-fragmentation edge-effects corridors connectivity landscape-ecology macroecology biogeochemistry climate-envelope species-distribution-modeling maxent bioclim worldclim remote-sensing gis spatial-analysis geostatistics abundance density biomass productivity diversity-index richness evenness dominance rarity endemism beta-diversity gamma-diversity alpha-diversity turnover nestedness community-assembly neutral-theory niche-theory competition facilitation apparent-competition indirect-effects interaction-networks mutualism parasitism predation herbivory detritivory scavenging cannibalism intraspecific-competition interspecific-competition character-displacement resource-partitioning niche-differentiation coexistence stability resilience resistance recovery succession disturbance perturbation extinction colonization invasion establishment spread impact risk-assessment management control eradication restoration reintroduction rewilding conservation-breeding ex-situ in-situ protected-area-network gap-analysis systematic-conservation-planning prioritization hotspot coldspot majority minority flagship umbrella keystone foundation ecosystem-engineer indicator sentinel bioindicator biomonitor ecosystem-health environmental-quality pollution pesticide heavy-metal contaminant bioaccumulation biomagnification toxicology risk hazard exposure dose-response no-observed-effect-concentration lowest-observed-effect-concentration median-lethal-concentration median-effective-concentration safety-factor uncertainty-factor probabilistic-risk-assessment life-cycle-assessment carbon-footprint water-footprint ecological-footprint sustainability sustainable-development green-economy circular-economy ecosystem-services natural-capital green-infrastructure nature-based-solutions ecological-restoration habitat-restoration species-recovery population-recovery genetic-rescue outbreeding-depression inbreeding-depression genetic-diversity heterozygosity effective-population-size bottleneck founder-effect genetic-drift gene-flow isolation-by-distance isolation-by-environment landscape-genetics seascape-genetics conservation-genetics molecular-ecology population-genetics quantitative-genetics genomics epigenetics reaction-norm norm-of-reaction genotype-environment-interaction heritability selection natural-selection sexual-selection artificial-selection domestication breeding cultivar variety landrace wild-relative crop-wild-relative genetic-resource germplasm seed-bank frozen-zoo cryopreservation assisted-reproduction cloning genetic-engineering synthetic-biology gene-drive biocontrol augmentative inundative conservation classical natural-enemy parasitoid pathogen nematode fungus bacterium virus viroid prion microbiome holobiont metaorganism symbiosis endosymbiosis ectosymbiosis amensalism neutralism symbiogenesis horizontal-gene-transfer virulence pathogenicity infectivity transmissibility reservoir vector host definitive-host intermediate-host paratenic-host accidental-host dead-end-host zoonosis emerging-infectious-disease pandemic epidemic endemic sporadic outbreak cluster index-case contact-tracing quarantine isolation social-distancing masking vaccination herd-immunity reproduction-number basic-reproduction-number effective-reproduction-number doubling-time generation-time serial-interval incubation-period latent-period infectious-period clinical-period pre-symptomatic asymptomatic pauci-symptomatic mild moderate severe critical fatal case-fatality-rate infection-fatality-rate attack-rate secondary-attack-rate excess-mortality years-of-life-lost disability-adjusted-life-year quality-adjusted-life-year burden-of-disease global-burden-of-disease one-health planetary-health ecohealth conservation-medicine veterinary-medicine human-medicine comparative-medicine translational-medicine precision-medicine personalized-medicine genomic-medicine regenerative-medicine stem-cell tissue-engineering organoid organ-on-chip microphysiological-system in-silico in-vitro ex-vivo in-vivo animal-model non-human-primate rodent mouse rat guinea-pig rabbit dog cat livestock poultry aquaculture fishery wild-capture mariculture algaculture seaweed-farming integrated-multi-trophic-aquaculture recirculating-aquaculture-system aquaponics hydroponics aeroponics vertical-farming controlled-environment-agriculture precision-agriculture smart-farming digital-agriculture agriculture-4.0 internet-of-things artificial-intelligence machine-learning deep-learning neural-network computer-vision drone unmanned-aerial-vehicle satellite gps gnss geographic-information-system decision-support-system expert-system knowledge-based-system ontology semantic-web linked-data open-data big-data data-mining data-science data-analytics predictive-analytics prescriptive-analytics descriptive-analytics diagnostic-analytics business-intelligence artificial-intelligence-ethics algorithmic-bias fairness accountability transparency explainability interpretability privacy security safety reliability robustness adaptability flexibility scalability efficiency effectiveness efficacy performance profitability cost-benefit cost-effectiveness economic-analysis financial-analysis investment return-on-investment net-present-value internal-rate-of-return payback-period break-even sensitivity-analysis scenario-analysis monte-carlo-simulation decision-tree real-options game-theory behavioral-economics nudge choice-architecture default framing anchoring availability-heuristic representativeness-heuristic affect-heuristic confirmation-bias availability-bias anchoring-bias framing-bias loss-aversion endowment-effect status-quo-bias sunk-cost-fallacy planning-fallacy optimism-bias pessimism-bias overconfidence underconfidence dunning-kruger-effect imposter-syndrome stereotyping prejudice discrimination implicit-bias explicit-bias structural-bias systemic-bias institutional-bias interpersonal-bias intrapersonal-bias unconscious-bias conscious-bias affinity-bias similarity-bias in-group-bias out-group-bias halo-effect horns-effect contrast-effect recency-effect primacy-effect serial-position-effect mere-exposure-effect familiarity-principle propinquity-effect social-proof authority scarcity reciprocity commitment consistency liking unity influence persuasion compliance conformity obedience empowerment agency autonomy self-determination motivation intrinsic-motivation extrinsic-motivation self-efficacy self-esteem self-concept self-identity social-identity collective-identity cultural-identity national-identity gender-identity sexual-orientation religion spirituality values beliefs attitudes norms customs traditions rituals ceremonies celebrations festivals holidays commemorations memorials monuments heritage legacy history historiography archaeology paleontology anthropology sociology psychology cognitive-science neuroscience linguistics philosophy ethics morality justice equity equality inclusion diversity belonging representation participation engagement capacity-building skill-development knowledge-transfer technology-transfer innovation invention discovery creativity imagination curiosity wonder awe beauty elegance simplicity complexity chaos order pattern structure function form process dynamics mechanism causation correlation association relationship interaction connection network system emergence self-organization autopoiesis homeostasis allostasis stress acclimation acclimatization plasticity fragility vulnerability sensitivity uncertainty ignorance surprise novelty serendipity chance luck fortune destiny fate karma dharma tao zen mindfulness meditation contemplation reflection introspection self-awareness consciousness subconscious unconscious dream sleep circadian biological-clock chronobiology seasonality phenology migration dispersal nomadism sedentism home-range habitat-use resource-selection foraging hunting gathering farming herding hunting-gathering subsistence livelihood economy market trade exchange barter currency money capital wealth poverty inequality development growth progress well-being happiness life-satisfaction quality-of-life human-development social-development economic-development millennium-development-goals sustainable-development-goals paris-agreement kyoto-protocol montreal-protocol basel-convention stockholm-convention rotterdam-convention unfccc cbd cms ramsar world-heritage man-and-biosphere ipcc ipbes encyclopedia-of-life wikidata wikimedia creative-commons open-access open-science faire-data research-data-management data-stewardship data-curation digital-preservation long-term-archiving repository archive library museum herbarium collection specimen voucher type holotype paratype syntype lectotype neotype epitype isolectotype isoneotype topotype homotype allotype cotype genotype phenotype ecotype chemotype morphotype karyotype idiogram chromosome genome proteome transcriptome metabolome lipidome glycome interactome connectome epigenome virome mycobiome bacteriome archaeome eukaryome phytobiome rhizobiome endobiome ectobiome exo-biome astrobiome planetary-biome earth-system climate-system ocean-system terrestrial-system freshwater-system atmospheric-system cryospheric-system anthroposphere technosphere noosphere biosphere hydrosphere lithosphere atmosphere magnetosphere heliosphere cosmosphere universe multiverse dimension space time spacetime relativity quantum-mechanics string-theory m-theory loop-quantum-gravity cosmology astronomy astrophysics planetary-science earth-science geology geophysics geochemistry geomorphology soil-science pedology edaphology hydrology limnology oceanography meteorology climatology paleoclimatology stratigraphy sedimentology mineralogy petrology volcanology seismology tectonics plate-tectonics continental-drift orogeny epeirogeny isostasy erosion weathering deposition transport sedimentation lithification diagenesis metamorphism melting crystallization igneous sedimentary metamorphic rock-cycle water-cycle carbon-cycle nitrogen-cycle phosphorus-cycle sulfur-cycle oxygen-cycle hydrogen-cycle biogeochemical-cycle nutrient-cycle trophic-pyramid food-chain ecological-network interaction-web mutualistic-network antagonistic-network competition-network facilitation-network ecosystem-function ecosystem-process provisioning-service regulating-service cultural-service supporting-service habitat-service refuge nursery corridor stepping-stone buffer-zone core-area periphery matrix patch landscape seascape riverscape skyscape soundscape smellscape tastescape touchscape mindscape dreamscape memoryscape heritagescape futurescape possibility-space scenario-space decision-space policy-space governance-space institutional-space social-space cultural-space economic-space political-space legal-space ethical-space moral-space spiritual-space sacred-space profane-space liminal-space transitional-space threshold boundary edge frontier margin center core hinterland heartland homeland motherland fatherland native-land ancestral-land promised-land wilderness pioneer settler colonist immigrant emigrant refugee asylum-seeker displaced-person stateless-person nomad traveler tourist pilgrim explorer adventurer discoverer inventor innovator creator maker builder craftsman artisan artist designer architect engineer scientist scholar academic researcher investigator detective journalist reporter correspondent chronicler historian biographer memoirist autobiographer diarist letter-writer poet novelist playwright screenwriter essayist critic reviewer editor publisher librarian archivist curator conservator restorer preservationist conservationist environmentalist ecologist naturalist botanist zoologist entomologist arachnologist herpetologist ichthyologist ornithologist mammalogist primatologist anthropologist archaeologist paleontologist geologist geographer cartographer surveyor navigator expedition voyage journey travel tour trip visit stay residence habitation dwelling shelter home house building construction architecture design plan blueprint model prototype mock-up draft sketch drawing painting sculpture installation exhibition display presentation demonstration show spectacle event occasion ceremony ritual celebration festival fair bazaar commerce business industry manufacturing production fabrication assembly processing refining purification synthesis analysis separation extraction identification characterization quantification measurement assessment evaluation appraisal valuation estimation calculation computation simulation modeling prediction forecast projection scenario outlook prospect expectation anticipation hope fear worry concern care attention focus concentration awareness perception sensation feeling emotion affect mood temperament personality character disposition attitude opinion belief conviction faith trust confidence reliance dependence independence self-sufficiency self-reliance hardiness toughness strength power force energy vigor vitality life health wellness fitness soundness integrity wholeness completeness perfection excellence quality standard criterion benchmark yardstick measure gauge sign signal symptom mark token symbol image icon emblem logo brand trademark copyright patent intellectual-property proprietary confidential secret classified restricted limited controlled regulated managed governed administered operated maintained sustained supported funded financed resourced staffed equipped furnished supplied provided delivered distributed allocated assigned designated appointed elected selected chosen picked preferred favored liked loved adored cherished treasured valued esteemed respected honored celebrated commemorated remembered recalled recollected reminisced reflected contemplated meditated pondered considered weighed evaluated judged assessed appraised rated ranked graded scored marked pointed noted observed noticed seen viewed looked watched gazed stared glanced peeked peered scanned surveyed examined inspected scrutinized studied investigated researched analyzed synthesized integrated compiled composed written authored penned drafted created produced made built constructed fabricated manufactured assembled put-together set-up established founded instituted organized arranged ordered structured systematized methodized rationalized streamlined optimized improved enhanced upgraded refined polished perfected completed finished concluded ended terminated closed shut sealed locked secured protected guarded defended shielded screened covered hidden concealed masked cloaked veiled shrouded wrapped enveloped surrounded encircled enclosed contained held kept retained preserved conserved saved stored stockpiled hoarded accumulated amassed gathered collected aggregated clustered grouped categorized sorted leveled tiered layered stratified hierarchized pyramided tree web mesh grid array configuration layout scheme program project enterprise venture undertaking endeavor effort attempt try test trial experiment investigation inquiry query question problem issue matter subject topic theme motif leitmotif thread strand line sequence series progression continuation extension expansion enlargement evolution change transformation conversion transition shift alteration modification adjustment accommodation reconciliation harmonization synchronization coordination integration unification combination merger fusion blend mixture compound composite alloy amalgam incorporation assimilation absorption digestion metabolism catabolism anabolism equilibrium balance steadiness constancy permanence endurance persistence continuity duration longevity lifespan lifetime existence being entity object thing item article piece part portion segment section division component element constituent ingredient factor aspect facet feature characteristic property attribute trait indication evidence proof verification validation confirmation corroboration substantiation authentication certification accreditation licensing permitting authorizing empowering enabling allowing letting tolerating accepting receiving taking getting obtaining acquiring gaining earning winning achieving attaining reaching arriving coming approaching nearing closing converging meeting joining connecting linking binding tying fastening attaching affixing appending adding supplementing complementing completing finishing perfecting consummating fulfilling satisfying gratifying pleasing delighting enjoying relishing savoring appreciating valuing prizing treasuring cherishing nurturing cultivating fostering promoting advancing furthering encouraging supporting backing assisting aiding helping serving attending tending caring nursing healing curing treating remedying correcting rectifying repairing fixing mending restoring renewing reviving resuscitating reanimating reawakening rekindling reigniting relaunching restarting rebooting refreshing rejuvenating reinvigorating reenergizing recharging refueling replenishing restocking replacing substituting exchanging swapping trading bartering negotiating bargaining dealing transacting contracting agreeing consenting approving endorsing sanctioning ratifying confirming validating authenticating verifying checking testing examining inspecting reviewing auditing monitoring supervising overseeing managing directing leading guiding steering piloting navigating driving propelling pushing pulling dragging hauling towing carrying bearing transporting conveying transmitting sending dispatching shipping delivering distributing disseminating spreading broadcasting publishing issuing releasing launching introducing presenting showing exhibiting displaying demonstrating illustrating exemplifying typifying representing symbolizing signifying meaning denoting connoting implying suggesting indicating pointing referring alluding hinting intimating insinuating inferring deducing concluding reasoning thinking cogitating cerebrating pondering musing reflecting contemplating meditating concentrating focusing listening hearing heeding noting noticing observing watching looking seeing viewing regarding considering weighing evaluating assessing judging appraising estimating guessing speculating hypothesizing theorizing postulating proposing recommending advising counseling consulting conferring discussing debating arguing disputing contesting challenging opposing resisting defying confronting facing encountering experiencing undergoing suffering enduring withstanding surviving living existing subsisting persisting persevering continuing remaining staying abiding residing inhabiting occupying possessing owning holding keeping retaining maintaining preserving conserving protecting guarding defending securing sheltering harboring entertaining welcoming greeting saluting hailing acclaiming applauding praising commending lauding extolling celebrating honoring glorifying exalting magnifying amplifying enlarging expanding extending stretching diffusing dispersing scattering allocating assigning allotting apportioning dividing partitioning separating detaching disconnecting disjoining disuniting dissociating disengaging withdrawing retreating receding retrograding reversing inverting turning rotating revolving spinning twirling whirling swirling circling orbiting looping coiling winding twisting bending curving arching vaulting bridging spanning crossing traversing passing moving going traveling journeying voyaging sailing flying gliding soaring hovering floating drifting flowing streaming pouring rushing gushing spurting spraying splashing splattering spattering uniting combining merging fusing blending mixing mingling intermingling intermixing integrating incorporating assimilating absorbing digesting metabolizing transforming converting transmuting transfiguring metamorphosing evolving developing growing maturing ripening aging declining deteriorating degenerating decaying rotting decomposing disintegrating crumbling collapsing falling failing dying perishing expiring departing leaving exiting retiring resigning abdicating relinquishing surrendering yielding submitting succumbing capitulating acquiescing complying conforming adapting adjusting accommodating reconciling harmonizing synchronizing coordinating unifying standardizing normalizing regularizing systematizing methodizing rationalizing streamlining simplifying clarifying elucidating explaining interpreting translating rendering paraphrasing summarizing abstracting condensing compressing shrinking reducing decreasing diminishing lessening lowering dropping descending sinking submerging diving plunging plummeting crashing colliding impacting hitting striking knocking tapping patting touching sensing perceiving detecting discovering finding locating positioning placing putting setting laying standing sitting lying resting relaxing reposing sleeping dreaming imagining fantasizing daydreaming woolgathering praying worshiping revering venerating adoring loving nourishing feeding eating drinking consuming ingesting energizing powering fueling motivating inspiring stimulating exciting arousing awakening alerting warning cautioning informing notifying telling reporting announcing declaring stating asserting claiming contending proving revealing disclosing exposing uncovering unmasking unveiling opening unlocking unsealing unfastening untying unbinding unwrapping unpacking unloading emptying clearing cleaning washing bathing showering rinsing soaking steeping immersing drowning suffocating choking strangling throttling squeezing pressing centering aiming targeting zeroing aligning tuning calibrating balancing leveling equalizing stabilizing steady mooring docking landing touching-down alighting perching roosting nesting denning burrowing tunneling digging excavating mining quarrying extracting removing eliminating eradicating exterminating destroying demolishing ruining wrecking damaging harming injuring hurting wounding maiming crippling laming disabling incapacitating paralyzing stunning shocking surprising astonishing amazing astounding staggering stupefying dazing bewildering confusing puzzling perplexing baffling mystifying enigmatic cryptic obscure abstruse recondite esoteric arcane occult mysterious unknown unfamiliar strange odd peculiar curious queer weird bizarre fantastic extraordinary remarkable notable noteworthy significant important momentous consequential weighty grave serious crucial vital essential necessary required needed wanted desired wished hoped expected anticipated awaited looked-for searched-for sought hunted pursued chased tracked trailed followed shadowed stalked crept crawled climbed scaled ascended mounted rose lifted elevated raised hoisted heaved hauled pulled tugged dragged drawn attracted lured enticed tempted seduced charmed enchanted fascinated captivated engrossed absorbed immersed involved engaged occupied busy active energetic dynamic vigorous forceful powerful strong mighty potent effective efficient productive fruitful fertile fecund prolific abundant plentiful ample sufficient adequate enough satisfactory acceptable tolerable bearable endurable sustainable maintainable manageable controllable governable rule govern reign dominate command direct lead guide steer pilot navigate drive propel push thrust shove press squeeze compress condense concentrate aim point target zero align adjust tune calibrate level equalize standardize normalize stabilize fix secure anchor moor dock land touch-down alight perch roost nest den burrow tunnel dig excavate mine quarry extract remove eliminate eradicate exterminate destroy demolish ruin wreck damage harm injure hurt wound maim cripple lame disable incapacitate paralyze stun shock astonish amaze astound stagger stupefy daze bewilder confuse puzzle perplex baffle mystify enigma
Leptoglossus
leaf-footed bugs
Leptoglossus is a genus of true bugs in the leaf-footed bug family Coreidae, tribe Anisoscelini. Species are characterized by leaflike dilations of the hind tibia, a diagnostic trait of the genus. The genus is distributed throughout the Americas, with some introduced populations in Europe and Asia. Several species are economically significant agricultural pests, notably L. occidentalis, which has become invasive in multiple continents.
Coreidae leaf-footed-bug agricultural-pest invasive-species symbiosis sexual-dimorphism conifer-pest nuisance-pest Burkholderia tachinid-parasitoid pheromone-communication building-invader misidentification Triatoma-look-alike gradual-metamorphosis seed-predator forest-pest ornamental-pest plumbing-damage public-health-confusion Chile Europe North-America South-America Asia Turkey almond-pest citrus-pest tomato-pest corn-pest overwintering-aggregation defensive-secretion pheromone-mediated-parasitism egg-parasitoid biological-control ecological-niche-modeling climate-suitability range-expansion introduction accidental-dispersal seed-orchard-pest germination-reduction empty-seed-formation cone-damage lodgepole-pine Douglas-fir western-conifer-seed-bug L.-occidentalis L.-zonatus L.-phyllopus L.-clypealis L.-australis L.-chilensis L.-oppositus male-combat femoral-weapon abdominal-gland pheromone-gland tibial-dilation leaf-like-hind-leg true-bug Hemiptera Heteroptera Pentatomomorpha Anisoscelini phytophagous plant-feeding stylet-feeding salivary-sheath overwintering diapause building-entry structural-pest nuisance-odor flight-sound droning-flight aggregation-behavior male-pheromone female-choice parasitoid-host biological-control-agent integrated-pest-management monitoring early-detection prevention invasion-biology alien-species non-native-pest global-spread climate-change niche-modeling suitable-habitat establishment-risk quarantine phytosanitary forest-health seed-production reforestation afforestation conifer-forestry pine-seed seed-viability economic-impact crop-loss yield-reduction kernel-damage fruit-damage nut-damage vector-of-disease Chagas-disease kissing-bug public-alarm misinformation education identification-tools health-system-burden pesticide-overuse urban-ecology domiciliary peridomiciliary Andean-region Patagonia Mediterranean temperate subtropical tropical agroecosystem natural-enemy predator parasite pathogen symbiont gut-microbiome soil-ingestion fitness-benefit reproductive-success sperm-morphology testicular-morphology accessory-gland pheromone-blend species-specific-odor cherry-scent vanilla-scent cinnamon-scent rose-scent olfactory-communication mate-location mate-recognition sexual-selection male-male-competition weapon-morphology allometry developmental-plasticity nymphal-instar egg-barrel egg-arrangement linear-oviposition leaf-surface plant-tissue stylet-penetration enzymatic-digestion fluid-feeding phloem-feeding xylem-feeding seed-feeding cone-feeding fruit-feeding kernel-feeding nut-feeding crop-feeding host-plant-range polyphagy oligophagy specialist generalist pest-status damage-threshold economic-injury-level management-strategy cultural-control physical-control chemical-control resistance tolerance host-plant-resistance monitoring-trap light-trap population-density distribution-map spread-rate colonization establishment population-dynamics life-table developmental-rate thermal-requirements degree-days phenology voltinism univoltine bivoltine multivoltine generation-time overwintering-site hibernaculum shelter-seeking cold-hardiness freeze-tolerance supercooling water-relations desiccation-resistance starvation-resistance dispersal-ability flight-capacity walking-behavior climbing-behavior aggregation-pheromone alarm-pheromone stink-gland metathoracic-gland dorsoabdominal-gland ventral-abdominal-gland sexual-gland morphological-defense chemical-defense mechanical-defense autotomy leg-loss predation-risk parasitism-risk parasitoid-risk pathogen-risk competition intraspecific interspecific resource-competition mating-competition sperm-competition cryptic-female-choice reproductive-isolation speciation phylogeny systematics taxonomy morphology anatomy histology ultrastructure spermatogenesis spermatozoa sperm-length nuclear-length cyst-production follicle-number testis-structure reproductive-anatomy genitalia mating-behavior copulation insemination oviposition egg-production fecundity fertility hatching-success nymphal-survival adult-longevity sex-ratio operational-sex-ratio population-sex-ratio effective-population-size genetic-diversity gene-flow population-structure phylogeography biogeography historical-biogeography vicariance dispersal range-shift range-contraction altitudinal-distribution latitudinal-distribution longitudinal-distribution island-distribution continental-distribution endemism cosmopolitanism introduced-range native-range source-population founder-population invasion-front lag-phase establishment-phase spread-phase equilibrium-phase impact-assessment risk-assessment horizon-scanning early-warning rapid-response eradication containment control management adaptation mitigation restoration conservation biodiversity ecosystem-service ecosystem-function food-web trophic-level primary-consumer herbivore fruit-predator plant-animal-interaction mutualism antagonism predation parasitism commensalism amensalism facilitation indirect-interaction trait-mediated-interaction density-mediated-interaction behavioral-ecology evolutionary-ecology functional-ecology physiological-ecology population-ecology community-ecology ecosystem-ecology landscape-ecology macroecology global-ecology applied-ecology agricultural-ecology forest-ecology conservation-biology invasion-ecology restoration-ecology pest-management conservation-biological-control augmentative-biological-control classical-biological-control parasitoid nematode fungus bacterium virus entomopathogen microbial-control sterile-insect-technique genetic-control mating-disruption attract-and-kill push-pull trap-crop border-crop refuge habitat-management cultural-practice crop-rotation tillage irrigation fertilization pruning harvest-timing sanitation residue-management weed-management cover-crop companion-planting intercropping agroforestry silviculture forest-management seed-orchard-management cone-collection seed-extraction seed-testing germination-testing seed-quality seed-vigor empty-seed insect-damage insect-injury feeding-scar puncture stylet-sheath salivary-flange symptom sign diagnosis sampling economic-threshold decision-support expert-system modeling simulation forecasting climate-model niche-model species-distribution-model habitat-suitability risk-mapping invasion-pathway introduction-vector trade transport tourism migration natural-dispersal human-mediated-dispersal accidental-introduction intentional-introduction release escape cultivation ornamental forestry agriculture horticulture urbanization globalization land-use-change habitat-fragmentation habitat-loss degradation pollution pesticide fertilizer nutrient-enrichment eutrophication acidification warming drought extreme-event disturbance succession recovery resilience stability variability uncertainty sustainable-development green-economy circular-economy bioeconomy ecosystem-approach one-health planetary-health environmental-health public-health veterinary-health plant-health food-security nutrition livelihood poverty inequality gender indigenous-knowledge traditional-knowledge local-knowledge citizen-science stakeholder-engagement policy governance regulation legislation international-cooperation capacity-building awareness communication outreach extension advisory-service research innovation technology-transfer knowledge-exchange networking partnership collaboration evaluation evidence-based-policy science-policy-interface diplomacy advocacy activism social-movement environmental-justice environmental-ethics environmental-philosophy environmental-humanities environmental-history environmental-literature environmental-art environmental-education environmental-communication science-communication risk-communication crisis-communication health-communication behavior-change pro-environmental-behavior sustainable-behavior consumption production waste recycling reuse reduction efficiency renewable clean green blue natural organic agroecology permaculture regenerative restorative healing transformative transdisciplinary interdisciplinary multidisciplinary cross-disciplinary disciplinary disciplinary-integration knowledge-integration system-thinking complexity emergence non-linearity feedback tipping-point regime-shift alternative-stable-state resilience-thinking adaptive-management adaptive-governance learning reflection participation inclusion equity justice fairness ethics values norms culture identity place sense-of-place attachment well-being quality-of-life happiness flourishing thriving sustainability sustainable development growth degrowth post-growth steady-state circular doughnut-economics planetary-boundaries safe-operating-space tipping-elements critical-transition early-warning-signal resilience-indicator sustainability-indicator benchmark target goal commitment pledge agreement treaty convention protocol framework strategy plan program project initiative campaign movement coalition alliance network platform hub center institute organization association society union federation confederation foundation trust fund bank facility mechanism instrument tool method approach technique procedure process system structure institution administration operation service delivery provision supply demand market economy finance investment funding financing resource capital asset liability income expenditure revenue cost benefit profit loss return yield outcome output input throughput productivity effectiveness performance quality standard criterion indicator metric measure assessment appraisal review audit verification validation certification accreditation recognition award prize honor distinction excellence best-practice lesson-learned experience expertise competence capacity capability skill knowledge information data evidence science discovery invention creation design testing piloting scaling mainstreaming uptake adoption diffusion dissemination publication reporting documentation archiving preservation rehabilitation reconstruction replication duplication backup security safety protection safeguarding insurance risk-management emergency-preparedness response relief humanitarian peace prosperity health employment housing infrastructure energy water food fishery aquaculture mining manufacturing industry commerce recreation heritage nature ecosystem environment climate atmosphere ocean freshwater land soil mineral biota flora fauna microorganism fungi plant animal vertebrate invertebrate arthropod insect bug Hemipteran Heteropteran pentatomomorphan coreoid coreid squash-bug boxelder-bug stink-bug shield-bug plant-bug mirid lygaeid seed-bug broad-headed-bug alydid reduviid assassin-bug triatomine bed-bug cimicid water-bug nepomorphan gerromorphan leptopodomorphan enicocephalomorphan dipsocoromorphan ceratocomboid schizopteroid peloridioid coleorrhynchan moss-bug archaeorrhynchan fulgoromorphan cicadomorphan membracoid treehopper leafhopper planthopper psyllid jumping-plant-louse whitefly aleyrodid scale-insect coccoid mealybug aphid adelgid phylloxeran sternorrhynchan thysanopteran thrips psocopteran barklouse booklouse phthirapteran louse sucking-louse chewing-louse mallophagan anopluran dermapteran earwig blattodean cockroach termite isopteran mantodean mantid phasmid stick-insect leaf-insect orthopteran grasshopper locust katydid cricket mole-cricket pygmy-mole-cricket camel-cricket cave-cricket weta ensiferan caeliferan gryllotalpid myrmecophilid tettigoniid gryllid acridid pamphagid pneumorid lentulid tristirid eumastacid proscopiid tridactylid tetrigid grouse-locust pygmy-grasshopper plecopteran stonefly embiopteran webspinner zorapteran angel-insect dictyopteran
Lygus hesperus
Western Tarnished Plant Bug
Lygus hesperus is a significant agricultural pest in western North America, causing extensive damage to cotton, strawberries, alfalfa seed, and other crops. In California alone, annual losses exceed $30 million in cotton and $40 million in strawberries. Adults overwinter in reproductive diapause triggered by short day lengths, resuming activity when conditions improve. The species has been the subject of extensive research on sampling methods, biological control, and insecticide resistance.
agricultural-pest plant-bug Miridae cotton-pest strawberry-pest alfalfa-pest western-North-America diapause biological-control integrated-pest-management insecticide-resistance mark-release-recapture salivary-enzymes extraoral-digestion Peristenus-relictus Hippodamia-convergens drone-sampling fluorescent-marking SmartWater vector Pantoea-ananatis Serratia potassium-chloride deterrent artificial-diet lentil seed-crops California Texas Arizona Nevada Washington Mexico polyphagous phytophagous stylet-probing scutellum nymph-spots economic-entomology organic-agriculture ecosystem-services bird-predation molecular-gut-analysis ELISA immunomarking parasitoid population-suppression crop-protection sampling-protocols sweep-net drop-cloth Y-tube-olfactometer gustatory-deterrent visual-cues volatile-cues reproductive-diapause postdiapause oogenesis oviposition longevity body-mass life-history mass-spectrometry salivary-proteins polygalacturonase α-amylase protease laccase glucose-dehydrogenase xanthine-dehydrogenase plant-defense host-pathogen-interaction iDrone-Bee UAV unmanned-aerial-vehicle open-source field-scouting IPM economic-threshold Bt-cotton natural-enemy conservation-biological-control landscape-ecology habitat-management weed-management border-sprays spot-treatment selective-insecticide neonicotinoid pyrethroid resistance-management sustainable-agriculture food-security economic-impact simulation-model population-dynamics immigration parasitism averted-loss value-loss cost-benefit research-methodology insect-marking dispersal fluorophore autofluorescence survivorship mortality mark-transfer crowding olfactometer feeding-behavior no-choice-assay choice-assay gustation host-selection plant-quality succulent-growth fruiting-structures square boll bloom terminal defoliation stand-loss dirty-bloom puckered-petals lesion carpel-wall lint-damage seed-damage yield-loss fiber-quality management-threshold scouting-interval action-threshold treatment-timing insecticide-efficacy labeled-rate difficult-to-control species-complex identification-key morphology color-variation wing-development antennae thorax abdomen instar developmental-stage laboratory-rearing oligidic-diet semisolid-slurry chicken-egg soybean wheat-germ lima-bean cost-reduction biomass-production survival fecundity egg-production generation-time population-growth log-phase colony-maintenance genetic-diversity field-collection adaptation stress-tolerance oxidative-stress temperature-extreme food-deficit resource-allocation reproductive-senescence endogenous-reserves environmental-stressor climate-adaptation phenology migration dispersal-pattern habitat-patch source-sink population-source population-sink crop-border field-edge non-crop-habitat refuge overwintering-site spring-emergence summer-population fall-population density-dependence aggregation spatial-distribution sampling-accuracy pest-detection monitoring-technology precision-agriculture digital-agriculture automation robotics machine-learning computer-vision sensor-technology data-collection decision-support grower-education extension stakeholder-engagement multidisciplinary-research academia-industry-collaboration sustainable-intensification agroecosystem food-system plant-insect-interaction insect-plant-coevolution salivary-secretion digestive-enzyme lytic-enzyme tissue-maceration feeding-injury physiological-damage mechanical-damage wound-response plant-defense-response induced-resistance systemic-acquired-resistance jasmonic-acid ethylene salicylic-acid secondary-metabolite phenolic-compound oxidative-enzyme redox-enzyme bacterial-transmission phytopathogen opportunistic-pathogen infection-court inoculum disease-vector epidemiology crop-hygiene sanitation crop-rotation cover-crop trap-crop push-pull habitat-manipulation conservation-biocontrol augmentative-biocontrol classical-biocontrol importation establishment non-target-effect host-specificity parasitoid-biology reproductive-parasitoid idiobiont koinobiont mummy-formation emergence-rate parasitoid-impact population-regulation top-down-control bottom-up-control trophic-cascade food-web ecosystem-function ecosystem-service disservice net-effect risk-assessment management-strategy policy regulation pesticide-registration resistance-monitoring stewardship integrated-resistance-management IRM refuge-strategy pyramid-strategy multiple-toxins transgenic-crop GMO biosafety environmental-impact non-target-organism pollinator-protection beneficial-insect natural-enemy-conservation floral-resource nectar pollen alternative-prey habitat-diversification agri-environment-scheme green-infrastructure ecological-compensation landscape-complexity connectivity matrix-quality spillover source-habitat ecological-trap population-viability extinction-risk invasion-biology range-expansion climate-change warming drought irrigation water-stress nitrogen-fertilization crop-vigor plant-nutrition host-plant-quality tritrophic-interaction plant-microbe-insect endosymbiont gut-microbiome salivary-microbiome vector-competence transmission-efficiency acquisition inoculation retention multiplication circulative non-circulative persistent non-persistent propagative stylet-borne foregut-borne hemolymph-borne salivary-gland-borne replication barrier midgut salivary-gland escape infection incubation latent-period inoculation-threshold transmission-threshold epidemic epiphytotic disease-triangle susceptible-host virulent-pathogen favorable-environment disease-cycle primary-inoculum secondary-inoculum overwintering-pathogen survival-structure disease-reservoir alternative-host volunteer-crop weed-reservoir crop-residue soilborne seedborne insect-borne management-tactic preventive curative eradicative suppressive protectant systemic contact translaminar fumigant biological-pesticide microbial-pesticide entomopathogenic-fungus entomopathogenic-nematode entomopathogenic-virus baculovirus granulovirus nucleopolyhedrovirus bacteria Bacillus-thuringiensis Bt-toxin crystal-protein cytolytic-protein vegetative-insecticidal-protein mode-of-action gut-paralysis septicemia toxemia antibiotic symbiont antibiotic-resistance fitness-cost resistance-allele resistance-mechanism metabolic-resistance target-site-resistance penetration-resistance behavioral-resistance cross-resistance multiple-resistance resistance-evolution selection-pressure dose-response lethal-concentration lethal-dose median-lethal-concentration median-lethal-dose confidence-interval probit-analysis logit-analysis dose-mortality time-mortality resistance-ratio resistance-factor diagnostic-dose discriminating-dose baseline-susceptibility susceptible-strain reference-strain field-population resistant-population homozygous-resistant heterozygous dominance recessive incomplete-dominance gene-frequency allele-frequency selection-coefficient fitness relative-fitness population-genetics quantitative-genetics heritability genetic-correlation breeding-value genetic-gain marker-assisted-selection genomic-selection transcriptomics proteomics metabolomics genomics functional-genomics comparative-genomics evolutionary-genomics population-genomics phylogenomics bioinformatics computational-biology systems-biology network-analysis pathway-analysis gene-ontology protein-protein-interaction signal-transduction hormone juvenile-hormone ecdysone insulin-like-peptide neuropeptide pheromone sex-pheromone aggregation-pheromone alarm-pheromone trail-pheromone kairomone allomone synomone semiochemical olfaction mechanoreception vision phototaxis chemotaxis anemotaxis thermotaxis hygrotaxis geotaxis kinesis taxis orthokinesis klinokinesis rheotaxis thigmotaxis contact-chemoreception olfactory-receptor gustatory-receptor ionotropic-receptor odorant-binding-protein chemosensory-protein sensillum sensory-neuron antennal-lobe mushroom-body lateral-horn central-complex optical-lobe compound-eye ocellus simple-eye photoreceptor rhabdom ommatidium facet visual-acuity motion-detection polarized-light-detection color-vision ultraviolet-vision infrared-detection thermoreception hygroreception mechanoreceptor chordotonal-organ campaniform-sensillum trichoid-sensillum basiconic-sensillum coeloconic-sensillum placoid-sensillum sensory-ecology foraging-behavior host-location host-acceptance host-suitability oviposition-preference oviposition-deterrence antixenosis antibiosis tolerance resistance-category gall-midge hessian-fly gene-for-gene avirulence-gene resistance-gene R-gene effector-triggered-immunity pattern-triggered-immunity pathogen-associated-molecular-pattern damage-associated-molecular-pattern microbe-associated-molecular-pattern recognition perception defense-response hypersensitive-response cell-death programmed-cell-death autophagy necrosis oxidative-burst reactive-oxygen-species nitric-oxide calcium-signaling mitogen-activated-protein-kinase transcription-factor defense-gene pathogenesis-related-protein antimicrobial-peptide phytoalexin lignin suberin callose papilla cell-wall-reinforcement hypersensitive-cell-death induced-systemic-resistance priming defense-elicitor plant-activator biostimulant plant-strengthener organic-amendment compost manure biochar vermicompost compost-tea microbial-inoculant plant-growth-promoting-rhizobacteria mycorrhizal-fungus endophyte epiphyte phytosphere rhizosphere phyllosphere spermosphere anthroposphere ecosystem-process nutrient-cycling carbon-sequestration nitrogen-fixation phosphorus-solubilization potassium-mobilization micronutrient-availability soil-structure water-infiltration water-holding-capacity erosion-control pollination pest-suppression disease-suppression weed-suppression decomposition detritivory predation herbivory mutualism commensalism competition facilitation inhibition amensalism neutralism symbiosis antagonism trophic-interaction food-chain energy-flow material-cycling biomass-pyramid trophic-pyramid ecological-efficiency production-efficiency assimilation-efficiency ecological-footprint sustainability resilience resistance adaptability transformability socio-ecological-system adaptive-management participatory-research citizen-science knowledge-co-production transdisciplinary-research interdisciplinary-research disciplinary-research basic-research applied-research development-research action-research participatory-action-research farmer-participatory-research on-farm-research station-research controlled-experiment field-experiment observational-study correlational-study manipulative-experiment natural-experiment quasi-experiment longitudinal-study cross-sectional-study case-study comparative-study meta-analysis systematic-review scoping-review rapid-review living-review evidence-synthesis knowledge-synthesis research-synthesis systematic-map bibliometric-analysis citation-analysis co-citation-analysis bibliographic-coupling keyword-analysis topic-modeling science-mapping research-front emerging-topic hot-topic research-gap future-direction research-priority research-agenda strategic-research mission-oriented-research societal-challenge sustainable-development-goal nutrition-security climate-smart-agriculture climate-resilient-agriculture low-input-agriculture regenerative-agriculture agroecology integrated-farming mixed-farming diversified-farming specialized-farming intensive-farming extensive-farming conventional-farming alternative-farming urban-agriculture peri-urban-agriculture rural-agriculture smallholder-agriculture large-scale-agriculture family-farming corporate-farming contract-farming tenant-farming subsistence-farming commercial-farming export-oriented-farming import-substituting-farming local-food-system global-food-system food-value-chain supply-chain value-chain commodity-chain agribusiness agrifood-system food-processing food-distribution food-retail food-service food-consumption food-waste food-loss post-harvest-loss storage-loss processing-loss distribution-loss retail-loss consumer-waste plate-waste kitchen-waste food-rescue food-recovery food-redistribution food-banking composting anaerobic-digestion bioenergy biofuel bioproduct biomaterial biorefinery circular-economy circular-bioeconomy green-economy blue-economy bioeconomy nature-based-solution ecosystem-based-adaptation ecosystem-based-mitigation climate-mitigation disaster-risk-reduction extreme-event flood heat-wave cold-snap frost hail windstorm tornado hurricane typhoon cyclone storm-surge sea-level-rise salinization desertification land-degradation soil-degradation soil-erosion soil-compaction soil-sealing soil-contamination soil-pollution heavy-metal persistent-organic-pollutant pesticide-residue antibiotic-residue hormone-residue plastic-pollution microplastic nanoplastic particulate-matter air-pollution water-pollution groundwater-pollution surface-water-pollution eutrophication hypoxia dead-zone biodiversity-loss habitat-loss habitat-fragmentation habitat-degradation habitat-destruction invasive-species introduced-species alien-species non-native-species exotic-species naturalized-species casual-species escaped-species cultivated-species ornamental-species pet-species livestock poultry aquaculture fishery capture-fishery marine-fishery freshwater-fishery inland-fishery recreational-fishery subsistence-fishery artisanal-fishery industrial-fishery overfishing bycatch discards ghost-fishing fishing-gear fishing-method fishing-effort fishing-capacity fishing-pressure stock-assessment fishery-management marine-protected-area no-take-zone marine-reserve terrestrial-protected-area national-park nature-reserve wildlife-sanctuary biosphere-reserve world-heritage-site ramsar-site important-bird-area key-biodiversity-area ecological-network protected-area-network conservation-corridor wildlife-corridor habitat-corridor stepping-stone buffer-zone transition-zone core-zone management-zone sustainable-use-zone restoration-zone rehabilitation-zone reclamation-zone remediation-zone containment-zone eradication-zone surveillance-zone monitoring-zone quarantine-zone biosecurity phytosanitary veterinary animal-health plant-health one-health ecohealth planetary-health global-health public-health environmental-health occupational-health food-safety feed-safety water-safety air-quality environmental-quality environmental-monitoring environmental-assessment environmental-impact-assessment strategic-environmental-assessment cumulative-impact-assessment social-impact-assessment health-impact-assessment hazard-assessment exposure-assessment dose-response-assessment risk-characterization risk-management risk-communication risk-perception risk-governance precautionary-principle polluter-pays-principle user-pays-principle extended-producer-responsibility corporate-social-responsibility environmental-stewardship sustainable-livelihood sustainable-consumption sustainable-production green-procurement green-supply-chain green-logistics green-manufacturing green-chemistry green-technology clean-technology environmental-technology pollution-prevention waste-minimization resource-efficiency energy-efficiency water-efficiency material-efficiency land-use-efficiency nutrient-use-efficiency nitrogen-use-efficiency phosphorus-use-efficiency potassium-use-efficiency micronutrient-use-efficiency fertilizer-use-efficiency pesticide-use-efficiency water-productivity land-productivity labor-productivity capital-productivity total-factor-productivity partial-factor-productivity agricultural-productivity crop-productivity livestock-productivity fishery-productivity forestry-productivity economic-productivity social-productivity environmental-productivity ecological-productivity biological-productivity primary-productivity secondary-productivity net-primary-productivity gross-primary-productivity ecosystem-productivity landscape-productivity regional-productivity global-productivity carbon-productivity nitrogen-productivity water-footprint carbon-footprint nitrogen-footprint phosphorus-footprint biodiversity-footprint material-footprint land-footprint energy-footprint emission-factor intensity-factor decoupling absolute-decoupling relative-decoupling resource-decoupling impact-decoupling dematerialization rematerialization transmaterialization substitution complementarity trade-off synergy co-benefit win-win lose-lose win-lose zero-sum positive-sum negative-sum game-theory prisoner's-dilemma tragedy-of-the-commons collective-action common-pool-resource public-good private-good club-good common-good global-common global-public-good tipping-point critical-transition regime-shift hysteresis irreversibility path-dependence lock-in technological-lock-in institutional-lock-in behavioral-lock-in social-ecological-trap poverty-trap degradation-trap development-trap complex-adaptive-system social-ecological-system coupled-human-natural-system human-environment-system adaptive-capacity transformative-capacity learning-capacity innovation-capacity governance-capacity institutional-capacity organizational-capacity individual-capacity community-capacity social-capital human-capital natural-capital physical-capital financial-capital built-capital cultural-capital political-capital intellectual-capital knowledge-capital information-capital data-capital digital-capital technological-capital infrastructural-capital institutional-capital organizational-capital network-capital relationship-capital reputational-capital brand-capital symbolic-capital existential-capital spiritual-capital religious-capital moral-capital ethical-capital trust-capital reciprocity-capital solidarity-capital cooperation-capital coordination-capital collaboration-capital communication-capital information-sharing knowledge-sharing learning innovation experimentation transformation transition change evolution development growth progress improvement enhancement optimization maximization minimization satisfaction sufficiency abundance scarcity security insecurity vulnerability robustness stability instability equilibrium disequilibrium homeostasis allostasis stress distress eustress coping coping-capacity coping-strategy coping-mechanism coping-resource coping-support social-support emotional-support instrumental-support informational-support appraisal-support network-support community-support institutional-support policy-support financial-support technical-support technological-support infrastructural-support capacity-building capacity-development capacity-strengthening empowerment enablement mobilization activation engagement participation involvement inclusion exclusion marginalization discrimination inequity inequality disparity gap divide digital-divide knowledge-divide information-divide science-divide research-divide innovation-divide technology-divide development-divide north-south-divide east-west-divide urban-rural-divide gender-divide age-divide class-divide income-divide wealth-divide asset-divide opportunity-divide outcome-divide wellbeing-divide happiness-divide life-satisfaction-divide health-divide education-divide skill-divide competency-divide capability-divide functioning-divide agency-divide freedom-divide choice-divide autonomy-divide self-determination-divide dignity-divide respect-divide recognition-divide status-divide power-divide influence-divide voice-divide representation-divide participation-divide engagement-divide citizenship-divide belonging-divide identity-divide culture-divide value-divide belief-divide norm-divide practice-divide behavior-divide habit-divide custom-divide tradition-divide modernity-divide globalization-divide localization-divide universalism-divide particularism-divide cosmopolitanism-divide nationalism-divide patriotism-divide populism-divide elitism-divide democracy-divide authoritarianism-divide control-divide security-divide liberty-divide privacy-divide surveillance-divide transparency-divide accountability-divide responsibility-divide liability-divide guilt-divide blame-divide punishment-divide reward-divide incentive-divide motivation-divide intention-divide attention-divide perception-divide cognition-divide emotion-divide affect-divide attitude-divide preference-divide interest-divide need-divide want-divide desire-divide demand-divide supply-divide market-divide exchange-divide transaction-divide contract-divide agreement-divide cooperation-divide competition-divide conflict-divide coercion-divide force-divide violence-divide peace-divide harmony-divide coexistence-divide tolerance-divide acceptance-divide rejection-divide opposition-divide resistance-divide conformity-divide deviance-divide normality-divide abnormality-divide illness-divide disease-divide disorder-divide disability-divide impairment-divide handicap-divide capacity-divide potential-divide performance-divide achievement-divide attainment-divide accomplishment-divide success-divide failure-divide excellence-divide mediocrity-divide average-divide standard-divide benchmark-divide threshold-divide ceiling-divide floor-divide limit-divide boundary-divide border-divide frontier-divide horizon-divide perspective-divide viewpoint-divide standpoint-divide position-divide stance-divide approach-divide method-divide technique-divide tool-divide instrument-divide device-divide apparatus-divide equipment-divide machine-divide machinery-divide mechanism-divide system-divide structure-divide organization-divide institution-divide regime-divide order-divide arrangement-divide configuration-divide pattern-divide design-divide plan-divide scheme-divide program-divide project-divide initiative-divide intervention-divide action-divide activity-divide operation-divide process-divide procedure-divide protocol-divide guideline-divide recommendation-divide requirement-divide regulation-divide rule-divide law-divide policy-divide strategy-divide tactic-divide objective-divide goal-divide target-divide aim-divide purpose-divide mission-divide vision-divide principle-divide ethic-divide moral-divide criterion-divide indicator-divide measure-divide metric-divide index-divide scale-divide score-divide rating-divide ranking-divide grade-divide level-divide stage-divide phase-divide step-divide progression-divide sequence-divide series-divide succession-divide chain-divide cycle-divide loop-divide spiral-divide wave-divide pulse-divide rhythm-divide tempo-divide pace-divide speed-divide velocity-divide acceleration-divide deceleration-divide momentum-divide inertia-divide energy-divide work-divide effort-divide strain-divide stress-divide pressure-divide tension-divide compression-divide expansion-divide contraction-divide deformation-divide distortion-divide transformation-divide conversion-divide transduction-divide transmission-divide transfer-divide transport-divide movement-divide motion-divide flow-divide flux-divide current-divide stream-divide river-divide tide-divide wind-divide storm-divide weather-divide climate-divide season-divide year-divide decade-divide century-divide millennium-divide epoch-divide era-divide period-divide time-divide duration-divide interval-divide span-divide stretch-divide spell-divide bout-divide episode-divide event-divide occurrence-divide incident-divide accident-divide coincidence-divide chance-divide luck-divide fortune-divide destiny-divide fate-divide karma-divide dharma-divide tao-divide way-divide path-divide road-divide route-divide course-divide direction-divide orientation-divide alignment-divide positioning-divide placement-divide location-divide site-divide place-divide spot-divide point-divide situation-divide context-divide environment-divide setting-divide scene-divide platform-divide base-divide foundation-divide ground-divide soil-divide earth-divide land-divide terrain-divide territory-divide domain-divide realm-divide sphere-divide world-divide globe-divide planet-divide universe-divide cosmos-divide space-divide void-divide emptiness-divide fullness-divide plenum-divide vacuum-divide atmosphere-divide air-divide gas-divide vapor-divide mist-divide fog-divide cloud-divide rain-divide snow-divide ice-divide frost-divide dew-divide humidity-divide moisture-divide wetness-divide dryness-divide aridity-divide drought-divide flood-divide deluge-divide inundation-divide submersion-divide immersion-divide saturation-divide supersaturation-divide condensation-divide evaporation-divide transpiration-divide sublimation-divide deposition-divide precipitation-divide infiltration-divide percolation-divide seepage-divide leakage-divide discharge-divide runoff-divide erosion-divide sedimentation-divide accretion-divide aggregation-divide coagulation-divide flocculation-divide crystallization-divide solidification-divide liquefaction-divide vaporization-divide boiling-divide melting-divide freezing-divide thawing-divide cooling-divide heating-divide warming-divide thermal-divide temperature-divide heat-divide cold-divide cool-divide warm-divide hot-divide thermodynamic-divide enthalpy-divide entropy-divide free-energy-divide Gibbs-free-energy-divide Helmholtz-free-energy-divide chemical-potential-divide fugacity-divide partial-pressure-divide concentration-divide dilution-divide solution-divide mixture-divide compound-divide element-divide atom-divide molecule-divide ion-divide electron-divide proton-divide neutron-divide nucleus-divide particle-divide quark-divide lepton-divide boson-divide fermion-divide hadron-divide baryon-divide meson-divide photon-divide gluon-divide weak-boson-divide Higgs-boson-divide string-divide brane-divide dimension-divide space-time-divide continuum-divide discrete-divide quantum-divide classical-divide relativistic-divide Newtonian-divide Einsteinian-divide Planckian-divide cosmological-divide astronomical-divide microscopic-divide macroscopic-divide mesoscopic-divide nanoscopic-divide atomic-divide molecular-divide cellular-divide tissue-divide organ-divide organism-divide population-divide community-divide ecosystem-divide biome-divide biosphere-divide ecosphere-divide noosphere-divide technosphere-divide anthroposphere-divide sociosphere-divide politosphere-divide econosphere-divide culturosphere-divide religiosphere-divide ideosphere-divide memesphere-divide infosphere-divide cybersphere-divide digisphere-divide virtual-divide analog-divide real-divide actual-divide possible-divide impossible-divide necessary-divide contingent-divide probable-divide improbable-divide certain-divide uncertain-divide determinate-divide indeterminate-divide definite-divide indefinite-divide specific-divide general-divide particular-divide universal-divide singular-divide plural-divide individual-divide collective-divide social-divide cultural-divide historical-divide political-divide economic-divide religious-divide philosophical-divide scientific-divide technological-divide artistic-divide literary-divide musical-divide visual-divide performing-divide cinematic-divide photographic-divide architectural-divide fashion-divide culinary-divide gastronomic-divide olfactory-divide gustatory-divide tactile-divide kinesthetic-divide proprioceptive-divide vestibular-divide auditory-divide somatosensory-divide interoceptive-divide exteroceptive-divide multisensory-divide crossmodal-divide synesthetic-divide anesthetic-divide esthetic-divide aesthetic-divide poetic-divide rhetorical-divide logical-divide mathematical-divide statistical-divide computational-divide algorithmic-divide heuristic-divide systematic-divide methodical-divide empirical-divide theoretical-divide conceptual-divide abstract-divide concrete-divide practical-divide applied-divide pure-divide fundamental-divide basic-divide strategic-divide tactical-divide operational-divide administrative-divide managerial-divide executive-divide leadership-divide followership-divide membership-divide partnership-divide ownership-divide stewardship-divide guardianship-divide trusteeship-divide custodianship-divide wardship-divide tutelage-divide mentorship-divide sponsorship-divide championship-divide friendship-divide relationship-divide kinship-divide companionship-divide fellowship-divide scholarship-divide apprenticeship-divide internship-divide residency-divide assistantship-divide associateship-divide authorship-divide editorship-divide curatorship-divide directorship-divide governorship-divide presidency-divide chancellorship-divide deanship-divide headship-divide chairmanship-divide
Polistes dominula
European Paper Wasp
Polistes dominula is a highly successful invasive social wasp native to Eurasia that has established populations across North America, South America, New Zealand, South Africa, and other regions. First detected in North America near Boston in 1978, it has become one of the most abundant wasps on the continent. The species builds small, exposed paper nests in protected locations and preys primarily on live insects, particularly caterpillars. Unlike yellowjackets, it does not scavenge for meat or sugar. Its rapid spread has been attributed to ecological flexibility, superior competitive ability, and tolerance of human-altered environments.
invasive-species social-wasp paper-wasp predator pollinator urban-ecology eusocial nest-construction biological-control thermal-biology spatial-learning vibrational-communication aposematism honest-signal overwintering diapause Polistes Vespidae Hymenoptera North-America Europe New-Zealand South-Africa South-America monarch-butterfly caterpillar-predation nest-architecture gyne foundress dominance-hierarchy climate-change-impact energetics metabolic-rate microclimate suburban-habitat anthropogenic ornamental-pest vineyard-pest fruit-damage stinging-insect venom poison-gland warning-coloration Zahavian-signal Strepsiptera parasitoid Xenos behavioral-plasticity foraging learning memory thermoregulation ectothermy endothermy solar-radiation nest-temperature brood-development colony-survival multiple-founding single-founding season-length latitude temperate Mediterranean hibernacle overwintering-costs climate-warming range-expansion competition native-species-displacement biodiversity-impact conservation management control integrated-pest-management IPM identification antennae-color facial-markings caste reproduction male female sexual-dimorphism antennae abdomen wing mandible saliva paper comb cell pedicel trophallaxis larva pupa adult worker queen subordinate dominant aggression defense threat-display sting toxicity brightness coloration predator-avoidance prey caterpillar Lepidoptera butterfly moth garden agriculture vineyard orchard urban suburban rural forest habitat-preference temperature metabolism quiescence torpor survival fitness colony nest population abundance distribution range expansion invasion establishment introduction biotic-interaction trophic-cascade herbivory-suppression plant-fitness milkweed Danaus-plexippus monarch Nelson Tasman ecosystem-impact conservation-management biological-invasion ecology behavior physiology morphology taxonomy systematics nomenclature gender-agreement Latin Christ 1791 Vespa-dominula basionym catalogue-of-life GBIF iNaturalist observation citizen-science photography illustration field-guide orange black yellow red color-pattern size wing-length body compact legs short face square triangular dark curled straight elbowed blunt pointed tip Zahavian costly-signal hunt forage nectar honeydew aphid scale-insect flower grape umbellifer fennel lovage insect meatball feed emerge chew cap hexagonal wood-fiber bark exposed open umbrella stalk ant repellent substance coat protect location eave attic tree branch vine shrub cavity bird-box bee-hive shed grave-lantern overwinter shelter protected thermal climate cold winter spring summer autumn fall season month day activity defend threat display raise tiptoe social dominance hierarchy reproductive egg lay found cooperate compete selfish passive-aggressive individual recognition facial-marking cognition mushroom-body brain intelligent artist architect navigation plasticity spatial relocating food resource dish bait training experience vibration communication signal substrate-borne abdominal wagging oscillation secretion modulation perceive react movement attention feeding inspection body-temperature ambient solar radiation metabolic cost energetic expenditure efficiency warming change impact increase invasive spread success competitive ability flexibility adaptation reaction response condition year variation productivity founding single multiple rate benefit cooperation predation parasitism vesparum stylopid stylopization parasite host infestation mite Acari wasp fly effect community decline Danaus plexippus mortality instar early late larger smaller vulnerable resistant discovery chance contact direct recruitment nestmate site cascading trophic cascade herbivory suppression plant growth production Asclepias net exclusion experiment outcome suggestion strong detrimental ecosystem strategy method development protection biodiversity region first 2016 recent established exotic congener chinensis longer comparison higher lower greatest area built structure native unsuccessful translocate unable thrive warmer origin contributor factor research thesis insight behaviour knowledge direction aid effective journal publication DOI 10.26686 wgtn 14538585 v1 University Wellington Victoria MSc student dissertation study investigation analysis data field resident report local indirect interaction level function service provision natural pest biocontrol ornamental decorative crop damage fruit cherry western Colorado at least ripening choice preference height ground strata microsite environment closed habitat dense vegetation surrounding bush Patagonia Argentina Chile NW detected 2003 sympatric coexistence overlap minimal facilitation behavioral process interspecific semi-urban poorly studied species Hypodynerus labiatus new light finding contribution understanding collecting anthropized highlight importance consideration Ethology 10.1111 eth 13505 abstract only limited detail full text required comprehensive synthesis vibrational widespread regulate crucial conspicuous oscillatory wag strictly associated presence suggested involvement adult-larva hypothesis short-term trophallactic exchange modulate salivary decrease amount prepare receive stimulate release electro-magnetic shaker assess time record measure immediately after playback result show produce possibly order attract during neither nor support alleged role long-lasting Experimental Biology 10.1242 jeb 186247 flexible system expanding North America cause stir fast upon explanation present literature good invader northern rise little known nesting Central investigate basic principle Germany 49°group one to four small shorter than other mean newly founded 83 average length 4.6 prefer 54%46%major increased 47%single-foundress whereas 100%more two however no number term per observe well three consecutive greatly depending preceding imply remarkably quickly outside rapid colder Ecosphere 10.1890 es15-00254 1 thermoregulatory polistine different collect demand brood mainly ectothermic task living differing reveal environmental influence warm Southern infrared thermography entire usually ~20–40 °C nearly linearly thorax warmest part achieve optimal preferably use model enable evaluation endothermic effort exhibit weak almost constant performance about contrast mostly minor avoid high way reduce Insects 10.3390 insects10070187 trait strongly Comparative B 10.1007 s00360-016-1041-x provided gallicus insects14110849 adverse weather offer low sheltered live energy store Italy Austria describe calculate according standard differ significantly between 3.2 ±5.71 8.5 5.29 variance correspond closely meteorological cumulative mass-specific over period lowest compared both acclimation calculation up 3 due dramatic 40%additional exponentially temperature-dependent fluctuation disproportionately Jensen's inequality wooden bird box empty bee hive under grapevine grave lantern cemetery begin when exceed ~18–20 confidence note information inferred from active not discussed this source association addressed relationship insects14110886 thermophilic originating but now widely quite synanthropic human settlement place with favorable typical side wasps'wide ~20–37 above air advantage as accelerate speed did approximately 38.6 managed by cooling near ~1–2 cm always clearly elevated station climate-model-generated macroclimate revealed necessity measuring for reliable description insects'brighter-colored poison gland Frontiers Zoology 10.1186 1742-9994-9-20 aposmatism against consisting warning using toxin costly physical could simultaneously abundant striking such case indicate animal toxic also would ensure honestly examine positive correlation exists collected 30 photographed them volume width remained even controlling suggest pattern true sign poisonous they have theoretical predicting indicates quantitative within acting metabolically therefore limitation first-year Spain exact age unknown ethanol preservation may alter applied uniformly pseudoreplication intraspecific supported while successfully worldwide revisiting previously gathered flight manipulation performed target several visit additionally analyse dealing position whether choose visited containing opt novel baited placed 60 away then train either once or compare taken find displaced demonstrate rapidly learned return certain significant reduction likewise manipulate spending less each moreover offered just majority furthermore contribute behavioural considering marginalis Cape Province South Africa African Entomology 10.4001 003 022 0104 Caucasian Entomological Bulletin 10.23885 1814-3326-2015-11-1-85-89 degree Hungary Natura Somogyiensis 10.24394 natsom 2011 19 229 many European country observed 1993 documentation photograph 5 of examined stylopized these exemplar parasitized 18 strepsipteran specimen some taxon scientific name rank preferred common Wonderful World Wasps Bug Squad Article Chris Alice Kratzer author LinkedIn Photo terrible jam-packed pain anger love us out blue minding own business eat picnic frighten child build home complicate already-stressful life Everyone knows be better without So preface book published company 2020 Owlfly LLC what write believe point essential complicated beautiful Indeed play important provide beneficial Look ma pesticide labor passionate dedicated written conversational tone long admirer supporter who grew Jersey continue remember interacting yes getting stung assorted including come territory become entomologist opted engineer received bachelor science 2019 mechanical engineering Rochester Institute Technology worked York until Today executive director managing division Publishing opportunity develop market sustainable potential mitigation fascinate fold into want why how do if tell building understand cycle organism before trying entangle ecological meaning inquiline called cuckoo exploit young call fact catch dismember tiny waist prevent eating sold fluid primary most grasshopper cricket katydid beetle earthworm lanternfly cockroach spider cicada aren't engineering-trained professional knowledgeably began her 2018 January 25 path included holing Cornell Collection networking expert teaching herself software align genetic barcode spent close thousand hour poring platform identifying mapping studying color incredible 400-page billed guide all arctic Greenland Alaska tropical Panama Grenada cover 208 22 genera feature 900 image macro photographer posting bring back Iowa State Professor Amy Toth's seminar Department Nematology told crowd attack extremely complex fascinating creature coined hashtag wasplove wrote illustrated It variety yellowjacket Eurasian red-banded American blackjacket volcano Guatemalian Eastern widow polka-dot prairie California form downy German knew there 'jackets familiar elaborate know dominulo means mistress implies Introduced 1970s since continent Woman Lynn Kimsey Bohart Museum distinguished telling difference scavenger take particularly serious mention UC Statewide Integrated Program wealth PestNote distinct type far troublesome especially ground-cavity-nesting tend vigorously disturbed Defensive increases progresses scarcer primarily start around garbage can dog cat where ripe overripe accessible large colorful tidbit sprinkle throughout intriguing instance Pennsylvania despite Someone mislabeled original And rated 2/5 painful Schmidt Index Bottom line eye-opener page-turner mind-boggler offers approach belongs library scientist citizen curious person wants incredibly diverse amazing next gathering anti-social folk ask anyone sharing rose Davis Calif thirsty sipping water fountain Agraulis vanillae Vacaville tucked inside Wednesday Cincinnati Ohio among changed recently overtaken counterpart Still United adjacent belong same family horn potter mason boreal cooler decidedly handful reaching rarely exceeding 200 unprotected suspend twig stem semi-protected hang sturdy often coated repel chief construction apparent destined likely determined stage act eventually assert leave performs securing developing offspring harvesting fiber surface dead fence post scrape ball deftly plies strip added existing worth having heavily Webworm grab hairy butcher kill spot stripping off any irritating hair spine easily consumed directly share turn distribute occupy reach maturity spin silken dome top molt shortly thereafter freedom sole fertilize Watch August September easy though genders elbow taper goldenrod Solidago throughwort Eupatorium evident extreme variability pair deep marking vary considerably DNA adding mud situation instead clarifying shown below going lump Enjoy watching unless antagonize give standing tip-toe raising make perfectly peaceful neighbor Posted 2:40 AM Email ThisBlogThis!Share XShare FacebookShare Pinterest Labels nature 6 comment ShandaSeptember 8 2010 5:26 loved get never faced can't wait kid question left end hibernate produced ReplyDelete RepliesReplyOut prairieSeptember 8:02 keeping cabbage worm seen RepliesReplyAnonymousSeptember 13 2:06 PM becoming here Upper Peninsula Michigan pendulum seems swung ago everywhere vanishingly rare haven't again house right RepliesReplyL.R ShimerSeptember 4:35 head gal work everybody else kinda egalitarian RepliesReplyLarryMay 9:34 directed Bev Wigney BTW lived NE Missouri detailed presented Thanks cogent scientifically accurate summary insect's life-cycle RepliesBug EricMay 3:47 kind compliments Delete RepliesReplyReplyAdd commentLoad Blog currently reply reader himself Working resolve Newer Older Subscribe Atom Demonized hosting 4 p.m.Saturday Jan 20 Room 1124 Academic Surge 455 Crocker Lane campus Attendees chat check 50,000 vespid eight-million free friendly Parking estimate 2000 described inhabit 500 undescribed headline-grabbing giant Vespa mandarinia formerly Asian news media dubbed murder hornet confuse identify sheet example hidden rodent burrow wall huge 100,00 will scavenge meat sugar soda hamburger road open-faced commonly located annual die cabbageworm white superb much delivered presentation website interested mechanism evolution honey Current project involve novo sequencing genome transcriptome genomic epigenetic regulating nutrition virus health hold doctorate Illinois Urbana–Champaign Gene Robinson postdoctoral Christina Grozinger unwelcome yard patio Preferring like beneath Sometimes legless grublike outstretched hand 15 relatively unaggressive problem doorway list earlier lepidopteran carry yeast winemaking fermentation flavor wine fuscatus recognize devoted mother dote fury opinion human-like great cooperator overall undercurrent selfishness manifest perfect themselves mushroom learning/memory 10 photogenic peer petal visitor camp Sierra Nevada mountain witnessed stinging crowded table ah sipped wondered says elsewhere which fertilized must warms enough lot protein baby need survived because drier usual deal competing go maintenance mode maintain fuel largely useless started mid trap spraying removed--this done night hollow available nearby hope wet barbecue shy except door window high-traffic Typically mated midsummer phase require cease sweet thing normally mild coastal San Francisco survive informative YouTube video Distinguishing see those distinctive put another Oakland Raiders'logo Giants soaking sun abandoned Pollinators Artists delight overhang neighborhood thought Wow hanging lip trash vanished Twitter associate Organismal

Subscribe to the newsletter

Email address

youtube tiktok snapchat instagram threads facebook twitter

Mike Porter

•

© 2026

•

•

Tailwind Nextjs Theme