Transition-management

Guides

• Arhopalus

  burnt pine longhorn beetle (A. ferus), black pine sawyer (A. rusticus)

  Arhopalus is a genus of longhorn beetles (Cerambycidae: Spondylidinae: Asemini) distributed across the Northern Hemisphere, with several species introduced to the Southern Hemisphere as invasive pests of conifers. The genus comprises approximately 20 described species, with A. rusticus and A. ferus being the most extensively studied due to their economic importance. Species develop in dead or fire-damaged conifer wood, with some exhibiting strong attraction to burnt pine volatiles. Several species produce male-emitted aggregation-sex pheromones containing fuscumol and geranylacetone. Invasive populations in New Zealand, Australia, South America, and South Africa require monitoring at ports and sawmills for timber export compliance.

  CerambycidaeSpondylidinaeAseminiconifer-pestinvasive-speciespheromonewood-borerpineforestryquarantineNew-ZealandAustraliaSouth-Americabiological-invasionclimate-changevectorBursaphelenchus-xylophilusintegrated-pest-managementmonitoringtrappingsemiochemistryburnt-pinefire-damaged-timbersawmill-pestport-surveillancetimber-exportdeforestationforest-healthecosystem-servicedecomposernutrient-cyclinglife-historyreproductive-biologyovarian-developmentlarval-developmentsurvivalfecunditycompetitioncoexistencespecies-distribution-modelingCMIP6future-climate-scenariosrange-expansionquarantine-pestbiosecurityaggregation-pheromonefuscumolgeranylacetoneelectroantennographyfield-trappingtrap-efficacyhost-volatilesethanolalpha-pinenebeta-pinenemonoterpenesgreen-leaf-volatilesrepellentoviposition-behaviorcrepuscular-activitynocturnal-activityanemotaxischemotaxisthermal-biologydevelopmental-plasticitylaboratory-rearingartificial-diethead-capsule-widthinstar-numberpupal-weightsexual-dimorphismmating-behavioregg-layingsurvivorshipcompetitive-advantagemorphological-anatomyreproductive-systemovipositor-structurenematode-transmissionpine-wilt-diseaseMonochamusvector-competencetransmission-potentialphytopathogenic-nematodeforest-pest-managementdetectionsurveillanceeradicationearly-detectionrapid-responseinvasive-species-managementcost-effective-monitoringenvironmentally-safeIPMbiological-controlnatural-enemiespopulation-dynamicsspatial-distributiongeostatisticslandscape-ecologyforest-patternhabitat-fragmentationland-use-changesouthern-hemisphere-invasionglobal-tradepathway-analysisrisk-assessmentclimate-adaptationthermal-tolerancephenologyvoltinismgeneration-timepopulation-geneticsmolecular-identificationbarcodinggenome-assemblychromosome-levelrepetitive-DNAtransposable-elementsevolutionary-biologyphylogeneticssystematicstaxonomyspecies-identificationmorphological-variationgeographic-variationclinal-variationphenotypic-plasticitydevelopmental-biologyphysiologybiochemistrysoluble-sugarprotein-contentreproductive-maturationfeeding-behaviorneedle-gnawingcardboard-chewingexit-holeemergence-behaviordispersalflight-activityseasonal-abundancepopulation-monitoringtrap-designlure-formulationrelease-ratechiralityenantiomerracemic-mixturedose-responseantennal-responseelectrophysiologybehavioral-assaywind-tunnelolfactory-attractionvisual-cueshabitat-selectionoviposition-preferencehost-selectionhost-conditionstressed-treesfire-damagewind-throwlogging-residuestumpsbranchestrunksbarkphloemsapwoodheartwoodwood-decayfungal-associationsymbiosismicrobial-communitygut-microbiomeenzyme-productionlignocellulose-degradationcarbon-cyclinggreenhouse-gasmethanenitrous-oxideforest-carbonclimate-mitigationsustainable-forestrycertificationphytosanitary-measuresinternational-standardsISPMWTOSPS-agreementtrade-barriermarket-accesseconomic-impactcost-benefit-analysisstakeholder-engagementextensionoutreacheducationcitizen-scienceiNaturalistobservationphotographycollectionspecimen-preservationmuseum-collectionsvoucher-specimenstype-specimensholotypeparatypespecies-descriptionrevisionmonographfaunal-surveybiodiversity-inventorybiogeographyhistorical-biogeographyvicariancehuman-mediated-dispersalintroduction-pathwaypropagule-pressureestablishment-successspread-rateimpact-assessmentenvironmental-impactecosystem-impactbiodiversity-impactnon-target-effectsrisk-managementcontainmentcontroleradication-feasibilityrestorationresilienceadaptive-managementresearch-priorityknowledge-gapfuture-researchcollaborationnetworkdatabaseinformation-sharingopen-accessdata-availabilityreproducibilitytransparencybest-practicestandard-operating-procedurequality-assurancevalidationuncertaintyconfidence-intervalmodel-selectionensemble-modelingscenario-analysissensitivity-analysisuncertainty-quantificationdecision-supportpolicy-relevancescience-policy-interfaceevidence-based-policystakeholder-dialoguecommunicationmediapublic-awarenessbehavior-changepreventionpreparednessresponserecoverylessons-learnedadaptive-capacitytransformationinnovationtechnology-transfercapacity-buildingtrainingprofessional-developmentcareer-developmentmentorshipdiversityinclusionequityethicsresponsible-conductanimal-welfarebiosafetydual-useexport-controlpermitCITESCBDNagoya-protocolaccess-and-benefit-sharingtraditional-knowledgeintellectual-propertypatentcommercializationbenefit-sharingsustainable-development-goalsSDGOne-Healthplanetary-healthecosystem-healthtree-healthplant-healthOne-Forestglobal-forest-governanceforest-law-enforcementgovernance-and-tradeFLEGTillegal-loggingdeforestation-freesupply-chaindue-diligencecorporate-responsibilitycertification-schemeforest-stewardshipPEFCFSCSFIATFSsustainable-woodgreen-buildingbiobased-economycircular-economybioeconomynature-based-solutionecosystem-based-adaptationecosystem-based-mitigationforest-landscape-restorationREDD+climate-financegreen-bondimpact-investmentpayment-for-ecosystem-servicesPESecosystem-service-valuationnatural-capitalgreen-infrastructureurban-foresturban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• Blissus occiduus

  Western Chinch Bug

  Blissus occiduus, the western chinch bug, is a phloem-feeding true bug (Hemiptera: Blissidae) that is a significant pest of warm-season turfgrasses, particularly buffalograss and zoysiagrass. The species exhibits strong host preference hierarchies, with buffalograss being the most preferred host followed by zoysiagrass, though it can survive and reproduce on a broad range of grasses including agronomic crops. Field studies have documented inconsistent control with neonicotinoid insecticides, with thiamethoxam showing particularly rapid degradation in buffalograss tissues compared to imidacloprid and clothianidin.

  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anaged-sphere-sphereprotected-sphere-sphererestored-sphere-spheredegraded-sphere-spheretransformed-sphere-sphereconnected-sphere-spheredisconnected-sphere-sphereintegrated-sphere-spherefragmented-sphere-sphereresilient-sphere-spherevulnerable-sphere-sphereadaptive-sphere-spheremaladaptive-sphere-spheresustainable-sphere-sphereunsustainable-sphere-sphereequitable-sphere-sphereinequitable-sphere-spherejust-sphere-sphereunjust-sphere-sphereinclusive-sphere-sphereexclusive-sphere-spherediverse-sphere-spherehomogeneous-sphere-spherecomplex-sphere-spheresimple-sphere-spheredynamic-sphere-spherestatic-sphere-spherestable-sphere-sphereunstable-sphere-spherepredictable-sphere-sphereunpredictable-sphere-spherecontrollable-sphere-sphereuncontrollable-sphere-spheremanageable-sphere-sphereunmanageable-sphere-spheregovernable-sphere-sphereungovernable-sphere-spherenavigable-sphere-sphereunnavigable-sphere-spheretraversable-sphere-sphereuntraversable-sphere-spherepermeable-sphere-sphereimpermeable-sphere-sphereporous-sphere-spherenon-porous-sphere-spheretransparent-sphere-sphereopaque-sphere-spherevisible-sphere-sphereinvisible-sphere-spheretangible-sphere-sphereintangible-sphere-spherematerial-sphere-sphereimmaterial-sphere-spherephysical-sphere-spheremetaphysical-sphere-sphereempirical-sphere-spheretheoretical-sphere-spherepractical-sphere-sphereapplied-sphere-spherepure-sphere-spherebasic-sphere-spherestrategic-sphere-spheretactical-sphere-sphereoperational-sphere-spherefoundational-sphere-spheresuperstructural-sphere-sphereinfrastructural-sphere-spherestructural-sphere-spherefunctional-sphere-sphereprocedural-sphere-spherebehavioral-sphere-sphereattitudinal-sphere-spherecultural-sphere-spheresocial-sphere-sphereeconomic-sphere-spherepolitical-sphere-sphereenvironmental-sphere-spheretechnological-sphere-sphereinstitutional-sphere-sphereorganizational-sphere-sphereindividual-sphere-spherecollective-sphere-spherecommunity-sphere-spheresocietal-sphere-sphereglobal-sphere-spherelocal-sphere-spherenational-sphere-sphereregional-sphere-spherelandscape-sphere-sphereseascape-sphere-spherewaterscape-sphere-sphereairscape-sphere-spherebiosphere-sphere-spherenoosphere-sphere-spheretechnosphere-sphere-spheresociosphere-sphere-sphereecosphere-sphere-spherehydrosphere-sphere-sphereatmosphere-sphere-spherelithosphere-sphere-spherepedosphere-sphere-spherecryosphere-sphere-sphereanthroposphere-sphere-sphereurban-sphere-sphere-sphererural-sphere-sphere-sphereagricultural-sphere-sphere-spherenatural-sphere-sphere-spheremanaged-sphere-sphere-sphereprotected-sphere-sphere-sphererestored-sphere-sphere-spheredegraded-sphere-sphere-spheretransformed-sphere-sphere-sphereconnected-sphere-sphere-spheredisconnected-sphere-sphere-sphereintegrated-sphere-sphere-spherefragmented-sphere-sphere-sphereresilient-sphere-sphere-spherevulnerable-sphere-sphere-sphereadaptive-sphere-sphere-spheremaladaptive-sphere-sphere-spheresustainable-sphere-sphere-sphereunsustainable-sphere-sphere-sphereequitable-sphere-sphere-sphereinequitable-sphere-sphere-spherejust-sphere-sphere-sphereunjust-sphere-sphere-sphereinclusive-sphere-sphere-sphereexclusive-sphere-sphere-spherediverse-sphere-sphere-spherehomogeneous-sphere-sphere-spherecomplex-sphere-sphere-spheresimple-sphere-sphere-spheredynamic-sphere-sphere-spherestatic-sphere-sphere-spherestable-sphere-sphere-sphereunstable-sphere-sphere-spherepredictable-sphere-sphere-sphereunpredictable-sphere-sphere-spherecontrollable-sphere-sphere-sphereuncontrollable-sphere-sphere-spheremanageable-sphere-sphere-sphereunmanageable-sphere-sphere-spheregovernable-sphere-sphere-sphereungovernable-sphere-sphere-spherenavigable-sphere-sphere-sphereunnavigable-sphere-sphere-spheretraversable-sphere-sphere-sphereuntraversable-sphere-sphere-spherepermeable-sphere-sphere-sphereimpermeable-sphere-sphere-sphereporous-sphere-sphere-spherenon-porous-sphere-sphere-spheretransparent-sphere-sphere-sphereopaque-sphere-sphere-spherevisible-sphere-sphere-sphereinvisible-sphere-sphere-spheretangible-sphere-sphere-sphereintangible-sphere-sphere-spherematerial-sphere-sphere-sphereimmaterial-sphere-sphere-spherephysical-sphere-sphere-spheremetaphysical-sphere-sphere-sphereempirical-sphere-sphere-spheretheoretical-sphere-sphere-spherepractical-sphere-sphere-sphereapplied-sphere-sphere-spherepure-sphere-sphere-spherebasic-sphere-sphere-spherestrategic-sphere-sphere-spheretactical-sphere-sphere-sphereoperational-sphere-sphere-spherefoundational-sphere-sphere-spheresuperstructural-sphere-sphere-sphereinfrastructural-sphere-sphere-spherestructural-sphere-sphere-spherefunctional-sphere-sphere-sphereprocedural-sphere-sphere-spherebehavioral-sphere-sphere-sphereattitudinal-sphere-sphere-spherecultural-sphere-sphere-spheresocial-sphere-sphere-sphereeconomic-sphere-sphere-spherepolitical-sphere-sphere-sphereenvironmental-sphere-sphere-spheretechnological-sphere-sphere-sphereinstitutional-sphere-sphere-sphereorganizational-sphere-sphere-sphereindividual-sphere-sph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• Pterocheilus hirsutipennis

  A solitary mason wasp in the subfamily Eumeninae, Pterocheilus hirsutipennis is one of approximately 40 North American species in this genus. Like other Pterocheilus, females are fossorial, excavating subterranean burrows in sandy or soft soils rather than using pre-existing cavities or building mud nests. The species was described by Bohart in 1940 and is part of a genus whose members are most diverse in the southwestern United States. Most species biology remains poorly documented.

  fossorialsolitary-waspmason-waspground-nestingcaterpillar-predatorsoil-dwellingpsammophoretarsal-rakeanti-parasitoid-behaviorEumeninaeVespidaeHymenopterainsectNorth-Americasouthwestern-US-diversityBohart-1940burrow-excavationsubterranean-nestcell-provisioningsand-wasp-mimicry-riskcryptic-nesting-behaviorcaterpillar-huntingfemale-dominant-nest-constructionsoil-scatteringnest-concealmentsolitary-Vespidaeaculeate-wasppredatory-wasplarval-provisioningmass-provisioningsingle-cell-nestsvertical-burrowsshallow-nestingsoft-substrate-dependentsandy-habitat-specialistpoorly-known-biologyrarely-encounteredobservationally-scarcetaxonomic-revision-needednatural-history-gapsfield-documentation-prioritycitizen-science-opportunityphotography-documentation-valuablebehavioral-study-candidatecomparative-nesting-biology-neededpsammophore-functiontarsal-rake-efficiencyburrow-architectureprey-selectioncaterpillar-paralysisoviposition-sequencecell-closureemergence-patternsseasonal-activitythermal-biologymicrohabitat-selectionsoil-moisture-requirementsvegetation-associationedge-habitat-preferencedisturbed-ground-utilizationcompetitive-exclusion-potentialsympatric-species-partitioningmolecular-phylogenetics-neededspecies-limits-unclearcryptic-diversity-possiblegeographic-variationphenotypic-plasticityenvironmental-responseclimate-sensitivityrange-shiftsconservation-status-unknownpopulation-trends-unmonitoredhabitat-degradation-vulnerabilitysoil-compaction-sensitivityurbanization-impactagricultural-intensification-effectspesticide-exposure-riskspollinator-network-interactionsfood-web-positionecosystem-service-valuationbiocontrol-potential-assessmentpublic-education-valuenatural-history-museum-collectionsspecimen-digitization-prioritylabel-data-mining-opportunityhistorical-distribution-reconstructionfuture-survey-targetingprotocol-standardizationexpert-identification-resourcesregional-faunal-surveysbiodiversity-inventory-gapstaxonomic-expertise-declinetraining-needsmentorship-importanceamateur-naturalist-collaborationlong-term-monitoring-establishmentphenology-trackingphenological-mismatch-risksphenotypic-mismatchenvironmental-cue-trackingdevelopmental-plasticitythermal-tolerance-limitsdesiccation-resistancewater-balanceenergeticsforaging-efficiencyprey-capture-successhandling-timetransport-costscentral-place-foragingoptimal-load-sizeprovisioning-rateoffspring-size-number-tradeoffssex-allocationlocal-mate-competitioninbreeding-avoidancedispersal-patternsgene-flowpopulation-structuregenetic-diversityeffective-population-sizebottleneck-historyfounder-eventsrefugial-dynamicspostglacial-expansionphylogeographic-structurecoalescent-analysisspecies-tree-inferencedivergence-datingbiogeographic-reconstructionancestral-area-estimationdispersal-vicariance-analysisecological-niche-modelingdistribution-predictionclimate-envelopehabitat-suitabilityprotected-area-coveragegap-analysisconservation-planningpriority-settingmanagement-recommendationsstewardship-guidelinesoutreach-materialsinterpretive-signageschool-curriculum-integrationuniversity-course-inclusiongraduate-research-opportunitiesfunding-agency-prioritiesgrant-proposal-developmentcollaborative-network-buildinginternational-partnershipdata-sharing-protocolsrepository-depositionmetadata-standardsquality-assurancecitizen-scientist-trainingverification-proceduresexpert-validationanomaly-detectionerror-correctionupdate-mechanismsversion-controlattribution-standardsintellectual-propertyethical-considerationsindigenous-knowledge-integrationtraditional-ecological-knowledgebiocultural-diversitystewardship-ethicsintergenerational-equitysustainability-principlesprecautionary-approachadaptive-managementevidence-based-policyscience-communicationmedia-engagementsocial-media-strategypublication-planningopen-access-commitmentpreprint-sharingpeer-review-participationeditorial-serviceprofessional-society-involvementconference-organizationworkshop-facilitationfield-course-instructionmentorship-programscareer-developmentdiversity-inclusionequity-promotionaccessibility-provisiondisability-accommodationmultilingual-resourcesvisual-impairment-supporthearing-impairment-accommodationneurodiversity-recognitionmental-health-awarenesswellness-promotionwork-life-balanceburnout-preventionresilience-buildingcommunity-supportnetwork-maintenancerelationship-cultivationtrust-developmentconflict-resolutionnegotiation-skillsleadership-developmentteam-dynamicscollaborative-problem-solvinginterdisciplinary-integrationtransdisciplinary-approachessystems-thinkingcomplexity-scienceemergent-propertiesself-organizationfeedback-loopsnon-linear-dynamicsthreshold-effectsregime-shiftsresilience-theoryadaptive-capacitytransformation-potentialinnovation-spaceexperimentation-encouragementfailure-tolerancelearning-orientationcontinuous-improvementreflective-practicecritical-self-awarenesshumility-cultivationcuriosity-maintenancewonder-preservationjoy-in-discoverypassion-sharinginspiration-transmissionlegacy-buildingfuture-generation-inves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gcalculation-performingcomputation-conductingmeasurement-takingquantification-conductingenumeration-performingcounting-conductingtallying-makingaccounting-keepingreckoning-performingcalculation-makingapproximation-providingguess-hazardingconjecture-offeringspeculation-engaginghypothesis-proposingtheory-constructingmodel-buildingframework-developingstructure-creatingsystem-designingarchitecture-developinginfrastructure-buildingfoundation-layinggroundwork-preparingbasis-establishingground-clearingsite-preparinglocation-selectingposition-choosingplacement-determiningarrangement-decidingdisposition-settlingdistribution-establishingallocation-makingassignment-givingdesignation-makingappointment-fixingscheduling-arrangingplanning-conductingorganizing-performingcoordinating-undertakingmanaging-conductingdirecting-performingleading-undertakingguiding-performingsteering-conductingpiloting-performingnavigating-undertakingsailing-conductingflying-performingdriving-undertakingriding-performingwalking-conductingrunning-performingjumping-undertakingleaping-performinghopping-conductingskipping-performingdancing-undertakingmoving-performingtraveling-conductingjourneying-performingvoyaging-undertakingexploring-conductingdiscovering-performingfinding-undertakinglocating-performingidentifying-conductingrecognizing-performingknowing-achievingunderstanding-attainingcomprehending-achievinggrasping-attainingseizing-achievingcatching-performingcapturing-undertakingsecuring-performingobtaining-achievingacquiring-attaininggaining-achievingearning-performingwinning-undertakingsucceeding-performingtriumphing-achievingvictory-attainingconquest-achievingdomination-attainingmastery-achievingcontrol-attainingcommand-achievingdirection-attainingleadership-achievingheadship-attainingcaptaincy-achievingskippering-attainingpiloting-achievinghelming-attainingsteering-achievingguiding-attainingdirecting-achievingmanaging-attainingadministering-achievinggoverning-attainingruling-achievingreigning-attainingcommanding-achievingordering-attainingdictating-achievingprescribing-attainingmandating-achievingrequiring-attainingdemanding-achievingrequesting-attainingasking-achievinginquiring-attainingquestioning-achievinginterrogating-attainingexamining-achievinginvestigating-attainingexploring-achievingprobing-attainingsearching-achievingseeking-attainingpursuing-achievingchasing-attaininghunting-achievingstalking-attainingtracking-achievingtrailing-attainingfollowing-achievingpursuing-attainingrunning-achievingracing-attainingcompeting-achievingcontending-attainingstruggling-achievingstriving-attainingendeavoring-achievingattempting-attainingtrying-achievingendeavoring-attainingexerting-achievingefforting-attainingworking-achievinglaboring-attainingtoiling-achievingslaving-attainingdrudging-achievingplodding-attainingpersevering-achievingpersisting-attainingcontinuing-achievingmaintaining-attainingsustaining-achievingupholding-attaining,

• Tetropium parvulum

  Northern Spruce Borer

  Tetropium parvulum, the northern spruce borer, is a small cerambycid beetle specialized on spruce hosts. It was elevated from synonymy with T. cinnamopterum based on distinct morphological differences in larvae and adults, and exhibits strict host specificity to Picea compared to the conifer generalist T. cinnamopterum. The species has a one-year life cycle and is an economically important borer of white and Engelmann spruce logs in western Canada.

  Cerambycidaespruce-borerwood-boring-beetleforest-pestwestern-Canadaconifer-specialistTetropiumPiceaeconomic-pestone-year-life-cycleTetropium-parvulumNorthern-Spruce-BorerTetropium-cinnamopterum-complexmorphological-diagnosisstrict-host-specificityTetropium-alaskanum-synonymRhimphoctona-alaskensis-parasitoidspruce-log-pestAlberta-forestryboreal-forest-insectphloem-feedercambium-borerL-shaped-galleriesearly-summer-emergencewoodpecker-predationdisease-mortalityexcess-heat-mortalityTetropium-velutinum-comparisonlectotype-designationTetropium-parallelumTetropium-schwarzianumconiferous-forest-decomposerprimary-borerTetropium-taxonomyCasey-1891Canadian-Entomologisteconomic-entomologyforest-healthtimber-pestspruce-regenerationPicea-glauca-pestPicea-engelmannii-pestwestern-Canadian-distributionNorth-American-CerambycidaeTetropium-morphologylarval-urogomphiadult-genitaliapronotal-punctationeye-morphologyscutellum-structurehost-preference-divergenceconifer-specialist-vs-generalistsympatric-speciesspecies-elevation-from-synonymytype-specimen-examinationdiagnostic-keyforest-insect-natural-enemiesinsect-disease-mortalityparasitoid-waspwoodpecker-insectivorythermal-mortalitygallery-depthoverwintering-biologyspring-pupationSeptember-larval-maturityone-year-univoltinespruce-logging-impactforest-managementintegrated-pest-managementboreal-forest-ecologysubcortical-insectphloeophagous-beetlexylophagous-larvaewood-degradationnutrient-cyclingforest-successiondisturbance-ecologyfire-killed-sprucewindthrow-sprucesalvage-logging-insectstanding-dead-timberspruce-beetle-complexScolytinae-competitorCerambycidae-communityconiferous-wood-borer-guildNorth-American-forest-insectswestern-Canadian-entomologyAlberta-entomologyBritish-Columbia-entomologyManitoba-entomologyLabrador-entomologytranscontinental-distributionboreal-forest-biodiversityspruce-ecosystem-engineertree-killing-insectsublethal-host-stressinduced-host-defenseinsect-fungus-interactionblue-stain-fungus-vectorwood-discolorationlumber-degradeveneer-quality-impactpulpwood-qualitybiomass-energy-feedstock-qualitycarbon-sequestration-impactclimate-change-forest-healthwarming-thermal-mortalitydrought-stress-predispositionspruce-decline-agentmulti-stressor-forest-dynamicsinsect-population-regulationnatural-enemy-complextrophic-cascadeforest-food-webavian-insectivorywoodpecker-foragingsapsucker-drillingnuthatch-predationparasitoid-communityichneumonid-waspbraconid-wasptachinid-flynematode-parasitebacterial-diseasefungal-entomopathogenviral-diseaseabiotic-mortality-factorcompetitive-exclusioninterspecific-competitionresource-partitioninghost-tree-chemistrymonoterpene-defensephenolic-compoundresin-duct-responseinduced-resistanceconstitutive-defensespruce-genotype-resistanceprovenance-variationlocal-adaptationevolutionary-ecologyspeciation-mechanismhost-race-formationreproductive-isolationmorphological-divergencegenitalic-evolutionallopatric-divergencesympatric-maintenancehybridization-potentialintrogression-riskconservation-geneticspopulation-structuregene-floweffective-population-sizeinbreeding-depressiongenetic-diversityadaptation-potentialevolutionary-rescueassisted-migrationex-situ-conservationcryopreservationmuseum-specimentype-localityholotypelectotypeparalectotypesyntypeoriginal-descriptiontaxonomic-revisionspecies-conceptbiological-speciesmorphological-speciesecological-speciesphylogenetic-speciesintegrative-taxonomyDNA-barcodingCOI-sequencenuclear-markermitochondrial-genomephylogenomicssystematicsclassification-stabilitynomenclatural-actsynonymyhomonymyavailabilityvaliditypriorityPrinciple-of-PriorityInternational-Code-of-Zoological-Nomenclaturetype-fixationtype-depositioncollection-repositoryNational-Museum-of-Natural-HistoryCanadian-National-CollectionRoyal-Ontario-MuseumUniversity-of-Alberta-collectionforestry-researchentomological-surveyinsect-monitoringpheromone-trappingflight-intercept-trapLindgren-funnel-trapemergence-cagerearing-logdissection-microscopegenitalic-preparationmicroscopic-photographyscanning-electron-microscopyconfocal-microscopycomputed-tomography3D-reconstructionmorphometric-analysisgeometric-morphometricslandmark-analysisshape-variationsize-allometrysexual-dimorphismintraspecific-variationphenotypic-plasticityenvironmental-gradientclinal-variationaltitudinal-gradientlatitudinal-gradientphenologydegree-day-modelthermal-accumulationdevelopment-ratevoltinismdiapausequiescencecold-hardinessfreezing-tolerancesupercooling-pointcryoprotectantglycerolsorbitoltrehaloseantifreeze-proteinwinter-survivalspring-emergenceadult-longevityoviposition-behavioregg-gallerylarval-feedingfrass-productiongallery-architecturepupal-chambercallus-tissuewound-responsetree-defensecompartmentalizationCODIT-modeldecay-columnheartwood-rotsapwood-stainstructural-integrity-losswindthrow-riskstand-dynamicssuccessional-trajectoryforest-regenerationplantation-forestrysilvicultural-practicethinningpruningfertilizationsite-preparationgenetic-improvementresistance-breedingclonal-forestryseed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• Tortopus

  Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.

  EphemeropteraPolymitarcyidaeburrowing-mayfliesNeotropicalclay-bank-habitatU-shaped-tunnelsmandibular-tusksTortopsisCampsurinaePanamerican-distributioncladistic-revisionsynapomorphyparastyli-receptorsflattened-penesfurcasternal-protuberancessubapical-tubercleslimnological-sampling-biasnymph-morphologyadult-morphologytype-species-T.-igaranusnew-species-T.-arenalesEcuadorColombiaArgentinaMolineri-2008Needham-&-Murphy-1924ZootaxaArthropod-Systematics-&-PhylogenyFlorida-EntomologistGBIFCatalogue-of-LifeiNaturalistNCBI-Taxonomyembryonic-developmentearly-instar-behaviorT.-harrisiT.-bellusT.-circumfluusT.-zottaiT.-igaranusT.-incertusT.-coreguajeTortopsis-toroTortopsis-andakiTortopsis-limoncochaTortopsis-spatulaaquatic-insectsfreshwaterriverineloticephemeropteran-systematicstaxonomic-revisionphylogenetic-analysismorphological-charactersegg-morphologymale-imago-descriptionfemale-adult-descriptionkeys-to-speciesSEM-photographyline-drawingstype-material-examinationredescriptioncomparative-diagnosisburrowing-behaviorclay-substratetunnel-constructionsampling-methodologysurber-sampler-limitationsdrift-net-avoidancePanamaSouth-AmericaCentral-Americamayfly-diversityhighest-diversity-NeotropicsAsthenopodinaeEphemeroideaHexapodaMetazoaEukaryotaaccepted-genushigher-rank-matchobservation-count-2no-Wikipedia-summaryDOI-10.11646/zootaxa.2481.1.1DOI-10.11646/zootaxa.2436.1.4DOI-10.3897/asp.79.e62735DOI-10.2307/3493493Scott-et-al.-1959NavásUlmerTraverLugo-Ortiz-&-McCaffertyDomínguezBanksMcDunnoughPictetNeedham-&-Murphyfreshwater-ecologyaquatic-entomologyinsect-developmentnymphal-stagesimago-descriptionalate-adultswing-venationR-sectorgonopore-structureparastyli-lengthpedestal-morphologylateral-view-curvaturemandibular-morphologytusk-tuberclesmesosternal-structureabdominal-sternum-notchfurcasternumprotuberance-configurationreceptor-socket-orientationIR-veinR2-veinclaw-like-structuregonopore-associationpenis-base-separationfemale-fore-wingadditional-veinscomparative-morphologytype-specimenholotypeallotypeparatypeneotropical-speciesnew-species-descriptionspecies-keyidentification-keymale-adult-keyfemale-adult-keynymph-keygeneric-keyPolymitarcyidae-generasystematicsphylogenycladisticsmonophylysister-groupcharacter-statetransformation-seriesparsimony-analysismorphological-data-setexternal-morphologyegg-chorionmicropyleegg-bursterhatchingemergencebehavioral-studyearly-instarembryologyovipositionsubstrate-preferencehabitat-specificityendemicregional-diversityconservationbiomonitoringwater-quality-indicatormacroinvertebratebenthicinfaunalinterstitialhyporheicriver-bankstream-bankerosionsedimentparticle-sizeclay-contentbank-stabilityroot-matvegetation-associationflow-regimecurrent-velocitydepthwidthstream-ordercatchmentbasinecoregionbiogeographydispersalcolonizationvicariancespeciationadaptive-radiationmorphological-divergenceconvergent-evolutionhomoplasycharacter-conflictbootstrap-supportBremer-supportconsensus-treestrict-consensusmajority-ruletaxonomic-stabilitynomenclatural-actoriginal-descriptionsubsequent-designationtype-localitytype-depositorymuseum-collectionspecimen-preservationalcohol-preservationslide-mountingSEM-preparationimaging-techniquesscientific-illustrationphotographymeasurementmorphometricsratiosproportionsallometrysexual-dimorphismgeographic-variationintraspecific-variationinterspecific-variationdiagnostic-charactersidentification-confidencetaxonomic-expertiseliterature-reviewbibliographic-searchdatabase-queryonline-repositoryopen-accesspaywallabstract-onlyfull-text-unavailablecitation-analysisreference-listsource-qualityevidence-strengthknowledge-gapresearch-needfuture-studycollaborationfield-workcollection-tripsampling-effortspecimen-availabilitytype-examinationrevision-scopetaxonomic-conceptcircumscriptioninclusionexclusionspecies-groupspecies-complexcryptic-speciesmorphospeciesphylogenetic-speciesevolutionary-significant-unitoperational-taxonomic-unitDNA-barcodingmolecular-systematicsintegrative-taxonomytraditional-taxonomyalpha-taxonomybeta-taxonomygamma-taxonomyclassification-hierarchyLinnaean-systemrank-based-nomenclaturephylogenetic-nomenclatureclade-namecrown-groupstem-grouptotal-evidencecongruenceincongruencedata-partitioncharacter-weightingimplied-weightingequal-weightingordered-charactersunordered-charactersmultistate-charactersbinary-characterscontinuous-charactersmeristic-charactersqualitative-charactersquantitative-charactersmorphological-matrixdata-matrixcharacter-matrixtaxon-samplingcharacter-samplingmissing-datainapplicable-datapolymorphismintraspecific-polymorphismfixed-differencediagnostic-differenceunique-derived-characterautapomorphyshared-derived-charactersymplesiomorphyconvergent-characterparallel-evolutionreversalcharacter-optimizationACCTRANDELTRANunambiguous-optimizationambiguous-optimizationmost-parsimonious-treetree-lengthconsistency-indexretention-indexrescaled-consistency-indexhomoplasy-indexcharacter-fitcharacter-congruencecharacter-supportbranch-supportnode-supportclade-stabilitytaxonomic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