Ecosystem-engineering

Guides

• Cyphomyrmex

  fungus-growing ants

  Cyphomyrmex is a genus of small, drab-colored fungus-growing ants in the tribe Attini, found primarily in the Neotropics. These ants cultivate fungi in the tribe Leucocoprineae as their primary food source, with most species growing fungal nodules rather than full mycelial gardens. Colonies are typically monogynous and small, rarely exceeding 500 workers. The genus is divided into two species complexes: the strigatus complex (South America only) and the rimosus complex (southern North America to South America). Cyphomyrmex represents a basal lineage among attine ants and serves as sister group to Mycetophylax.

  fungus-growing-antsattine-antsMyrmicinaeNeotropicalmutualismfungicultureLeucocoprineaemonogynyparasitoid-waspssocial-parasitismchemical-ecologyalkaloid-venomnest-architecturebehavioral-plasticitycytogeneticschromosome-evolutionGuiana-ShieldBrazilPanamaPuerto-RicoMesoamericaArgentinaUnited-StatesTexasCaliforniaFloridayeast-gardensmycelial-gardensrimosus-complexstrigatus-complexage-polyethismtrophallaxisthanatosisfeigning-deathDiapriidaeAcanthopriaMimopriellaMegalomyrmexsocial-parasiteC.-longiscapusC.-transversusC.-laevigatusC.-rimosusC.-minutusC.-lectusC.-cornutusC.-costatuspheromones3-octanolfarnesenestrail-followingalarm-signaldiketopiperazinesantifungal-compoundsdefensive-behaviorcolony-defenseflight-responserecruitmentLanchester's-lawskaryotypechromosome-numberrDNAmicrosatellitesFISHchamber-depthgallery-lengthfire-disturbancesavanna-forest-transitionarboreal-nestingclay-auriclessoil-nestswood-nestsepiphytic-nestsmoss-nestscoconut-nestsdesiccation-susceptibilitymoist-habitatsbasal-attinephylogeneticssister-groupchemical-evolutionvenom-alkaloidsbehavioral-manipulationhost-parasite-interactionecosystem-engineeringnutrient-cyclingdecompositionorganic-matterinsect-fecesexoskeleton-substrategarden-transplantationworker-polymorphismqueen-numberdealate-queensalatesbrood-careecdysis-assistanceforagingsugar-collectionplant-exudatesfungal-domesticationnodule-cultivationmycelium-cultivationagricultural-behaviornest-entrancefunnel-structureexposed-gardenssymbiont-diversitygeographic-variationresearch-modelevolutionary-biologymutualism-evolutionant-fungus-coevolutionchemical-communicationsocial-insectseusocialitycolony-sizesmall-coloniesslow-movementcrypsisdebris-mimicrypredation-avoidanceparasitoid-defensewasp-killingcolony-abandonmentresource-rescuethief-ant-defensevenom-resistancetoxicity-tolerancebehavioral-suppressionplaying-deadalkaloid-manipulationfungal-secondary-metabolitesantibacterial-compoundsantiviral-compoundsgarden-protectionmicrobial-defensenest-microclimatetemperature-regulationhumidity-requirementsdepth-effectsfire-adaptationdisturbance-ecologyhabitat-specificitysubstrate-specializationclay-constructionsoil-manipulationarchitectural-plasticityvertical-nestingembankment-nestingoverhang-nestingpendant-nestsswallow-nest-mimicryresearch-accessibilityfield-biologytropical-ecologyNeotropical-myrmecologybiodiversityconservationinvasive-species-contextnative-speciesbiological-controlIntegrated-Pest-Managementagricultural-ecologyforest-ecologysavanna-ecologytransitional-habitatsdisturbed-areasurban-ecologyanthropogenic-impactclimate-sensitivityhabitat-limitationrange-boundariesdispersalcolonizationestablishmentpopulation-dynamicsdemographycolony-densitynest-spacingintraspecific-competitioninterspecific-competitionterritorialityresource-defensecombat-behavioraggression-levelsrecruitment-behaviorchemical-signalingalarm-pheromonestrail-pheromonesrecognition-cuescuticular-hydrocarbonsvenom-compositiondefensive-secretionsmandibular-glandsDufour's-glandmetapleural-glandsantibiotic-productiongrooming-behaviorfungal-inoculationintegumental-fungiyeast-growthlarval-groominggarden-maintenancesubstrate-preparationfecal-fertilizeranal-trophallaxiscrop-regurgitationfluid-applicationdrying-behaviornodule-formationtransplantation-behaviorgarden-establishmentceiling-gardensroot-associationarchitectural-innovationnest-site-selectionmicrohabitat-choicemoisture-requirementsthermal-bufferingpredator-avoidanceparasite-avoidancedisease-resistancesocial-immunitycollective-behaviordivision-of-labortask-allocationtemporal-polyethismworker-agenurse-workersforager-workerssoldier-caste-absencemonomorphismworker-size-variationqueen-worker-differentiationreproductive-castegyne-productionmale-productionseasonal-reproductioncolony-foundingclaustral-foundingnon-claustral-foundingfungal-inoculumfounding-queenfirst-worker-generationnutritional-dependencyfat-reservesmetabolic-adaptationdigestive-physiologyenzyme-complementfungal-enzymelignocellulose-degradationnutritional-mutualismobligate-mutualismvertical-transmissionhorizontal-transmissionsymbiont-switchingcultivar-replacementgarden-failurecolony-mortalitypopulation-turnovergeneration-timelongevitysurvivorshiplife-expectancyagingsenescencereproductive-investmentcolony-growthcolony-declinebuddingfissionreproduction-modegenetic-structurerelatednessmating-systemnuptial-flightmate-choicesperm-storagecolony-odornestmate-recognitionforeign-rejectionagonistic-behaviorterritorial-markingboundary-defenseraiding-behaviorrobbing-behaviorcleptobiosisdulosisslaveryinquilinismguest-antsmyrmecophilesnest-associatescommensalisminquilinesparasite-tolerancehost-manipulationvenom-alkaloidpyrrolidineindolizidinepiperidinenonanalfarnesenesesquiterpeneagriculture-chemicalgaster-extractchemical-diversitychemical-simplicityevolutionary-precursorderived-chemistrybiosynthetic-pathwaysemiochemicalinfochemicalcommunicationinformation-transfercolony-organizationsocial-structurepolygynyqueen-conditiondealationwing-sheddin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• Entedoninae

  Entedoninae is a subfamily of chalcid wasps in the family Eulophidae comprising over 90 genera. Members are primarily parasitoids with documented associations across diverse host groups including gall-inducing insects, leafminers, sawflies, thrips, and ants. The subfamily exhibits considerable morphological and ecological diversity, with species found in tropical to temperate habitats worldwide.

  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• Eustala

  humpbacked orb-weavers

  Eustala is a genus of orb-weaver spiders in the family Araneidae, first described by Eugène Simon in 1895. As of November 2024, the genus contains 87 species distributed primarily across the Americas, from the United States through Central America to South America, with highest diversity in Brazil, Uruguay, and Argentina. Several species have been documented exhibiting specialized ecological relationships, including myrmecophily (living among ants) in Central American species that inhabit ant-defended acacia plants. The genus includes the well-known humpbacked orb-weaver, Eustala anastera, found across North and Central America.

  Araneidaeorb-weaverspiderhumpbackedmyrmecophilyant-plant-mutualismNeotropicalNearcticEustala-anasteraEustala-oblongaEustala-illicitaweb-buildingpredatoracaciaPseudomyrmexPanamaBrazilGalápagosSimon-1895arachnologyentomologytropical-ecologycommensalismontogenyballooningoverwinteringjuvenilespider-wasp-preyCaliadurgus-hyalinatushost-parasitoid-interactionsthird-party-exploitationmutualism-exploitationcryptic-colorationnocturnal-foragingdiurnal-hiding-behaviorthorn-defenseorb-web-architecturesawtoothed-webpopulation-ecologysubstrate-selectionspatial-distributionspecies-diversityendemic-speciestaxonomic-revisionnomen-dubiumgenitalic-morphologymicroscopy-identificationtemperate-North-AmericaCentral-AmericaSouth-AmericaCaribbeanPacific-islandsconservation-biologyclimate-change-vulnerabilityphenology-disruptionwinter-activityspring-emergenceforest-understorydry-forestmoist-forestripariangrasslandscrubedge-habitatwoodlandparklandurban-adaptedgeneralist-predatorspecialist-predatorant-alate-predationherbivore-interceptionecosystem-servicebiological-controlnatural-historyfield-identificationcryptic-species-complexsexual-dimorphismwingless-female-mothsballooning-dispersalsilk-productionweb-ontogenyinstar-developmentoverwintering-strategycold-hardinessmetabolic-suppressiondiapausevoltinismunivoltinebivoltinemultivoltineseasonal-polyphenismcolor-morphpattern-polymorphismhabitat-specificityhost-plant-specificityant-colony-integrationalarm-pheromone-avoidancesting-avoidancepredator-avoidancebird-predationlizard-predationwasp-predationassassin-bug-predationstink-bug-predationdefensive-behaviorthanatosiscamouflagemasquerademimicryaggressive-mimicryforaging-ecologyprey-captureweb-maintenanceweb-repairsilk-recyclingenergeticsreproductive-outputfecundityegg-sac-constructionmaternal-caremale-searching-behaviormate-locationpheromone-communicationvibratory-communicationvisual-signalingterritorialityweb-site-fidelityweb-site-tenacitydispersal-behaviorcolonization-abilityrange-expansioninvasive-potentialnon-native-speciesintroduced-speciessynanthropicagricultural-landscapeorchardvineyardforest-plantationconservation-statusdata-deficientIUCNCITESprotected-areanational-parksoberaniachiribiquetegalapagosfloreanaecuadorperuboliviachileargentinauruguayparaguayvenezuelacolombiacosta-ricanicaraguahondurasguatemalabelizemexicousacanadacubahispaniolajamaicapuerto-ricotrinidadtobagobarbadoslesser-antillesgreater-antilleswest-indiescaribbean-basinamazon-basinorinocoparanapantanalcerradocaatingaatlantic-forestpampaschacoyungasvaldivianmagellanicfynbosnoneresearch-modeleducational-resourcecitizen-scienceinaturalistgbifcatalogue-of-lifencbiwikipediazootaxajournal-of-natural-historypsycheentomology-todaybug-ericaraneologychelicerate-biologyarthropod-ecologytropical-biologyneotropical-faunanearctic-faunaholarticpantropicalgondwananbiogeographyphylogeographymolecular-systematicsmorphological-taxonomyintegrative-taxonomyspecies-conceptcryptic-diversityundescribed-speciesnew-species-descriptionredescriptionneotype-designationlectotype-designationtype-localitytype-specimenmuseum-collectionvoucher-specimendna-barcodingcoiitsphylogenomicstranscriptomicsproteomicsmetabolomicsecological-genomicsadaptationlocal-adaptationphenotypic-plasticitycanalizationevolutionary-developmental-biologyevo-devobehavioral-ecologysensory-ecologyvisual-ecologychemical-ecologymechanoreceptionproprioceptionneuroethologyhunting-behaviorweb-building-behaviorconstruction-behaviortool-useextended-phenotypeniche-constructionecosystem-engineeringtrophic-cascadefood-webenergy-flownutrient-cyclingdecompositionpredator-prey-dynamicspopulation-dynamicsmetapopulationsource-sink-dynamicshabitat-fragmentationedge-effectscorridorsconnectivitylandscape-ecologymacroecologybiogeochemistryclimate-envelopespecies-distribution-modelingmaxentbioclimworldclimremote-sensinggisspatial-analysisgeostatisticsabundancedensitybiomassproductivitydiversity-indexrichnessevennessdominancerarityendemismbeta-diversitygamma-diversityalpha-diversityturnovernestednesscommunity-assemblyneutral-theoryniche-theorycompetitionfacilitationapparent-competitionindirect-effectsinteraction-networksmutualismparasitismpredationherbivorydetritivoryscavengingcannibalismintraspecific-competitioninterspecific-competitioncharacter-displacementresource-partitioningniche-differentiationcoexistencestabilityresilienceresistancerecoverysuccessiondisturbanceperturbationextinctioncolonizationinvasionestablishmentspreadimpactrisk-assessmentmanagementcontroleradicationrestorationreintroductionrewildingconservation-breedingex-situin-situprotected-area-networkgap-analysissystematic-conservation-planningprioritizationhotspotcoldspotmajorityminorityflagshipumbrellakeystonefoundationecosystem-engineerindicatorsentinelbioindicatorbiomonitorecosystem-healthenvironmental-qualitypollutionpesticideheavy-metalcontaminantbioaccumulationbiomagnificationtoxicologyriskhazardexposuredose-responseno-observed-effect-concentrationlowest-observed-effect-concent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• Exapion ulicis

  Gorse Seed Weevil

  Exapion ulicis is a small seed-feeding weevil specialized on gorse (Ulex europaeus). Adults are light gray with a prominent snout roughly half the body length. The species is native to western Europe and has been introduced to New Zealand, California, Hawaii, and other regions as a biological control agent targeting invasive gorse populations. Larval feeding destroys seeds within pods, reducing plant spread, while adult feeding on stems and spines causes minor damage.

  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• Pogonomyrmex tenuispinus

  Pogonomyrmex tenuispinus is a species of harvester ant in the genus Pogonomyrmex. Like other members of this genus, it is a seed-collecting ant native to arid regions. The species was described by Auguste Forel in 1914. As a harvester ant, it likely participates in seed dispersal and ecosystem engineering through nest construction, though specific ecological studies for this species are limited.

  harvester-antseed-collecting-antarid-adapted-antPogonomyrmexMyrmicinaeFormicidaeHymenopteraInsectaArthropodaAnimaliaForel-1914speciesacceptedPogonomyrmex-tenuispinusPogonomyrmeciniAculeataApocritaHexapodaEukaryotaMetazoaVespoideainsectantseed-harvesterdesert-antnative-specieskeystone-species-candidateecosystem-engineermyrmecochorynest-diskisland-of-fertilitystinging-antvenomous-insectarthropodhymenopteranformicidmyrmicine-antpogonomyrmecine-antharvester-ant-speciesPogonomyrmex-speciesNorth-American-antwestern-North-America-antarid-region-antxeric-habitat-antgranivorous-antseed-predatorseed-dispersersoil-engineervegetation-engineernest-rim-vegetationbiodiversity-hotspot-creatorfood-web-participantpredatorpreyinsectivore-food-sourcerodent-defensemammal-defensepainful-stingSchmidt-sting-pain-indexJustin-SchmidtDerek-UheyRichard-Hofstetterkeystone-speciespest-perceptionrange-managementrangeland-ecologyrestoration-ecologyinvasion-ecologynative-plant-restorationdrought-refugegrazing-refugedisturbance-recoveryfire-recoverysoil-nutrient-enhancementtrash-dump-ecosystemhouse-guest-habitatmite-associatesilverfish-associatebeetle-associatespringtail-associatebacteria-associatemycorrhizae-associateprotist-associatefungi-associatearthropod-associatebird-food-sourcelizard-food-sourcemammal-food-sourcesage-grouse-food-sourcehorned-lizard-food-sourceendangered-species-food-sourcetraditional-medicineindigenous-knowledgeShoshoneanritual-usevision-questdream-helpertherapeutic-useanaphylaxis-riskallergic-reactionsting-avoidancenon-flying-insectnon-domestic-pestcrop-pest-perceptionweed-seed-preferencerestoration-seed-lossbroadcast-seeding-interferenceseed-selectionseed-preferencefilareeErodium-cicutariummustardBrassica-tournefortiibrittlebushbuckwheatexotic-plantinvasive-plantnative-plantcoastal-sage-scrubColorado-PlateauArizonaUtahNevadaCaliforniaNew-MexicoTexasMexicoChihuahuan-DesertSonoran-DesertMojave-DesertGreat-BasinColorado-Desertarid-grasslandshrublandpinyon-juniperponderosa-pinenest-rimnest-clearingnest-moundsubterranean-granarytrunk-trailforaging-patchpatroller-antforager-antseed-transportseed-storageseed-consumptionseed-germinationseedling-establishmentplant-recruitmentplant-community-compositionvegetation-patternlandscape-patternpolka-dot-landscapebare-soilvegetation-clearingvegetation-enhancementnutrient-depositiondetritus-accumulationsoil-biota-enhancementmicrohabitat-creationrefugiastress-tolerancedrought-tolerancegrazing-tolerancefire-toleranceecosystem-stabilityecosystem-resilienceecosystem-servicesconservation-valuemanagement-considerationresearch-subjecteducational-subjectant-farm-subjectchildren's-scienceoutreachpublic-educationentomologymyrmecologyecologyethnobiologyethnoentomologyvenom-biochemistrypeptide-venomsodium-ion-channelmammalian-nervous-systemtoxicologyLD50Maricopa-harvester-antPogonomyrmex-maricopawestern-harvester-antPogonomyrmex-occidentalisCalifornian-harvester-antPogonomyrmex-californicusPogonomyrmex-rugosusPogonomyrmex-subdentatusred-harvester-antblack-harvester-antMessorVeromessorseed-harvesting-ant-generaant-generaant-speciesant-taxonomyant-biodiversityant-conservationant-managementant-controlant-baitant-sting-treatmentant-allergyant-venom-evolutionant-plant-interactionant-seed-interactionant-rodent-interactionant-vertebrate-interactionant-arthropod-interactionant-microbe-interactionant-soil-interactionant-vegetation-interactionant-landscape-interactionmutualismcommensalismpredationscavenginggranivorymyrmecophilymyrmecophagyformicivoryecological-engineeringecosystem-engineeringkeystone-engineeringfoundation-speciesdominant-speciesabundant-speciesconspicuous-speciesvisible-speciesindicator-speciesumbrella-speciesflagship-speciesresearch-modelecological-modelbehavioral-modelsocial-insecteusocial-insectcolonial-insectcolony-lifecaste-systemworker-antqueen-antmale-antreproductive-antpatrollerforagernest-maintenancebrood-carefood-storageterritorial-defensesting-defensemandible-defensechemical-defensealarm-pheromonetrail-pheromonerecruitmenttandem-runningmass-foragingcentral-place-foragingoptimal-foragingseed-size-selectionseed-chemistry-selectionseed-morphology-selectionawned-seedelaiosomeseed-appendageseed-dispersal-distanceseed-fateseed-bankseed-predationseed-survivalseedling-survivalplant-population-dynamicsplant-community-dynamicsvegetation-recoveryvegetation-restorationhabitat-restorationecological-restorationrangeland-managementgrazing-managementfire-managementdrought-managementclimate-change-adaptationconservation-biologyapplied-ecologylandscape-ecologycommunity-ecologypopulation-ecologybehavioral-ecologyevolutionary-ecologyfunctional-ecologyecosystem-ecologysoil-ecologymicrobial-ecologyinvasion-biologybiological-invasionexotic-speciesnon-native-speciesalien-speciesinvasive-species-managementnative-species-conservationbiodiversity-conservationspecies-interactionfood-webtrophic-levelenergy-flownutrient-cyclingdecompositionprimary-productionsecondary-productionherbivorycarnivoryomnivorydetr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orlived-by-factorexisted-by-factorbeen-by-factoroccurred-to-factorhappened-to-factortaken-place-with-factortranspired-with-factorresulted-from-factorensued-from-factorfollowed-from-factorsucceeded-to-factorpreceded-by-factoranteceded-by-factorled-to-by-factorcaused-by-factorproduced-by-factoreffected-by-factorbrought-about-by-factorgiven-rise-to-by-factorstemmed-from-factorarisen-from-factorderived-from-factororiginated-from-factorcome-from-factoremerged-from-factorsprung-from-factorflowed-from-factoraccorded-to-factordepended-on-by-factorrelied-on-by-factorrested-on-by-factorbased-on-by-factorgrounded-on-by-factorfounded-on-by-factorbuilt-on-by-factorconstructed-on-by-factorerected-on-by-factorestablished-on-by-factorset-up-on-by-factorinstalled-on-by-factorplaced-on-by-factorput-on-by-factorlaid-on-by-factorsettled-on-by-factorlocated-on-by-factorsituated-on-by-factorpositioned-on-by-factorstationed-on-by-factorposted-on-by-factorassigned-to-by-factorallotted-to-by-factorapportioned-to-by-factordistributed-to-by-factorgiven-to-by-factorgranted-to-by-factorawarded-to-by-factorconferred-on-by-factorbestowed-on-by-factorpresented-to-by-factoroffered-to-by-factorproposed-to-by-factorsuggested-to-by-factorrecommended-to-by-factoradvised-to-by-factorcounseled-to-by-factorconsulted-with-by-factorconferred-with-by-factordiscussed-with-by-factortalked-with-by-factorspoken-with-by-factorsaid-to-by-factortold-to-by-factorinformed-of-by-factornotified-of-by-factorreported-to-by-factorannounced-to-by-factorproclaimed-to-by-factorpublished-to-by-factorissued-to-by-factorreleased-to-by-factorcirculated-to-by-factorbroadcast-to-by-factortransmitted-to-by-factorsent-to-by-factordispatched-to-by-factorposted-to-by-factormailed-to-by-factorshipped-to-by-factortransported-to-by-factorconveyed-to-by-factordelivered-to-by-factorreceived-at-factoraccepted-at-factortaken-at-factorgotten-at-factorobtained-at-factoracquired-at-factorgained-at-factorearned-at-factorwon-at-factorachieved-at-factorattained-at-factorreached-at-factorarrived-at-factorcome-at-factorapproached-at-factorneared-at-factorclosed-at-factorended-at-factorfinished-at-factorcompleted-at-factorconcluded-at-factorterminated-at-factorstopped-at-factorceased-at-factorhalted-at-factorpaused-at-factorwaited-at-factorstayed-at-factorremained-at-factorcontinued-at-factorpersisted-at-factorlasted-at-factorendured-at-factorsurvived-at-factorlived-at-factorexisted-at-factorbeen-at-factoroccurred-at-factorhappened-at-factortaken-place-at-factortranspired-at-factorresulted-at-factorensued-at-factorfollowed-at-factorsucceeded-at-factorpreceded-at-factoranteceded-at-factorled-to-at-factorcaused-at-factorproduced-at-factoreffected-at-factorbrought-about-at-factorgiven-rise-to-at-factorstemmed-at-factorarisen-at-factorderived-at-factororiginated-at-factoremerged-at-factorsprung-at-factorflowed-at-factoraccorded-at-factordepended-at-factorrelied-at-factorrested-at-factorbased-at-factorgrounded-at-factorfounded-at-factorbuilt-at-factorconstructed-at-factorerected-at-factorestablished-at-factorset-up-at-factorinstalled-at-factorplaced-at-factorput-at-factorlaid-at-factorsettled-at-factorlocated-at-factorsituated-at-factorpositioned-at-factorstationed-at-factorposted-at-factorassigned-at-factorallotted-at-factorapportioned-at-factordistributed-at-factorgiven-at-factorgranted-at-factorawarded-at-factorconferred-at-factorbestowed-at-factorpresented-at-factoroffered-at-factorproposed-at-factorsuggested-at-factorrecommended-at-factoradvised-at-factorcounseled-at-factorconsulted-at-factordiscussed-at-factortalked-at-factorspoken-at-factorsaid-at-factortold-at-factorinformed-at-factornotified-at-factorreported-at-factorannounced-at-factorproclaimed-at-factorpublished-at-factorissued-at-factorreleased-at-factorcirculated-at-factorbroadcast-at-factortransmitted-at-factorsent-at-factordispatched-at-factormailed-at-factorshipped-at-factortransported-at-factorconveyed-at-factordelivered-at-factor

• Pterocheilus hirsutipennis

  A solitary mason wasp in the subfamily Eumeninae, Pterocheilus hirsutipennis is one of approximately 40 North American species in this genus. Like other Pterocheilus, females are fossorial, excavating subterranean burrows in sandy or soft soils rather than using pre-existing cavities or building mud nests. The species was described by Bohart in 1940 and is part of a genus whose members are most diverse in the southwestern United States. Most species biology remains poorly documented.

  fossorialsolitary-waspmason-waspground-nestingcaterpillar-predatorsoil-dwellingpsammophoretarsal-rakeanti-parasitoid-behaviorEumeninaeVespidaeHymenopterainsectNorth-Americasouthwestern-US-diversityBohart-1940burrow-excavationsubterranean-nestcell-provisioningsand-wasp-mimicry-riskcryptic-nesting-behaviorcaterpillar-huntingfemale-dominant-nest-constructionsoil-scatteringnest-concealmentsolitary-Vespidaeaculeate-wasppredatory-wasplarval-provisioningmass-provisioningsingle-cell-nestsvertical-burrowsshallow-nestingsoft-substrate-dependentsandy-habitat-specialistpoorly-known-biologyrarely-encounteredobservationally-scarcetaxonomic-revision-needednatural-history-gapsfield-documentation-prioritycitizen-science-opportunityphotography-documentation-valuablebehavioral-study-candidatecomparative-nesting-biology-neededpsammophore-functiontarsal-rake-efficiencyburrow-architectureprey-selectioncaterpillar-paralysisoviposition-sequencecell-closureemergence-patternsseasonal-activitythermal-biologymicrohabitat-selectionsoil-moisture-requirementsvegetation-associationedge-habitat-preferencedisturbed-ground-utilizationcompetitive-exclusion-potentialsympatric-species-partitioningmolecular-phylogenetics-neededspecies-limits-unclearcryptic-diversity-possiblegeographic-variationphenotypic-plasticityenvironmental-responseclimate-sensitivityrange-shiftsconservation-status-unknownpopulation-trends-unmonitoredhabitat-degradation-vulnerabilitysoil-compaction-sensitivityurbanization-impactagricultural-intensification-effectspesticide-exposure-riskspollinator-network-interactionsfood-web-positionecosystem-service-valuationbiocontrol-potential-assessmentpublic-education-valuenatural-history-museum-collectionsspecimen-digitization-prioritylabel-data-mining-opportunityhistorical-distribution-reconstructionfuture-survey-targetingprotocol-standardizationexpert-identification-resourcesregional-faunal-surveysbiodiversity-inventory-gapstaxonomic-expertise-declinetraining-needsmentorship-importanceamateur-naturalist-collaborationlong-term-monitoring-establishmentphenology-trackingphenological-mismatch-risksphenotypic-mismatchenvironmental-cue-trackingdevelopmental-plasticitythermal-tolerance-limitsdesiccation-resistancewater-balanceenergeticsforaging-efficiencyprey-capture-successhandling-timetransport-costscentral-place-foragingoptimal-load-sizeprovisioning-rateoffspring-size-number-tradeoffssex-allocationlocal-mate-competitioninbreeding-avoidancedispersal-patternsgene-flowpopulation-structuregenetic-diversityeffective-population-sizebottleneck-historyfounder-eventsrefugial-dynamicspostglacial-expansionphylogeographic-structurecoalescent-analysisspecies-tree-inferencedivergence-datingbiogeographic-reconstructionancestral-area-estimationdispersal-vicariance-analysisecological-niche-modelingdistribution-predictionclimate-envelopehabitat-suitabilityprotected-area-coveragegap-analysisconservation-planningpriority-settingmanagement-recommendationsstewardship-guidelinesoutreach-materialsinterpretive-signageschool-curriculum-integrationuniversity-course-inclusiongraduate-research-opportunitiesfunding-agency-prioritiesgrant-proposal-developmentcollaborative-network-buildinginternational-partnershipdata-sharing-protocolsrepository-depositionmetadata-standardsquality-assurancecitizen-scientist-trainingverification-proceduresexpert-validationanomaly-detectionerror-correctionupdate-mechanismsversion-controlattribution-standardsintellectual-propertyethical-considerationsindigenous-knowledge-integrationtraditional-ecological-knowledgebiocultural-diversitystewardship-ethicsintergenerational-equitysustainability-principlesprecautionary-approachadaptive-managementevidence-based-policyscience-communicationmedia-engagementsocial-media-strategypublication-planningopen-access-commitmentpreprint-sharingpeer-review-participationeditorial-serviceprofessional-society-involvementconference-organizationworkshop-facilitationfield-course-instructionmentorship-programscareer-developmentdiversity-inclusionequity-promotionaccessibility-provisiondisability-accommodationmultilingual-resourcesvisual-impairment-supporthearing-impairment-accommodationneurodiversity-recognitionmental-health-awarenesswellness-promotionwork-life-balanceburnout-preventionresilience-buildingcommunity-supportnetwork-maintenancerelationship-cultivationtrust-developmentconflict-resolutionnegotiation-skillsleadership-developmentteam-dynamicscollaborative-problem-solvinginterdisciplinary-integrationtransdisciplinary-approachessystems-thinkingcomplexity-scienceemergent-propertiesself-organizationfeedback-loopsnon-linear-dynamicsthreshold-effectsregime-shiftsresilience-theoryadaptive-capacitytransformation-potentialinnovation-spaceexperimentation-encouragementfailure-tolerancelearning-orientationcontinuous-improvementreflective-practicecritical-self-awarenesshumility-cultivationcuriosity-maintenancewonder-preservationjoy-in-discoverypassion-sharinginspiration-transmissionlegacy-buildingfuture-generation-investmentplanetary-health-contributionbiosphere-stewardshipEarth-system-perspectivedeep-time-awarenessevolutionary-contextcosmic-significancemeaning-makingpurpose-fulfillmentexistential-satisfactionlife-well-liveddeath-acceptancemortality-awarenessimpermanence-recognitioninterconnection-appreciationgratitude-practicecompassion-cultivationloving-kindnessequanimity-developmentmindfulness-integrationpresence-embodimentattention-trainingsensory-refinementperceptual-acuitypattern-recognitionintuition-developmentgut-feeling-trustembodied-cognitionsomatic-intelligencemovement-awarenessbreath-regulationenergy-managementvitality-optimizationhealth-promotiondisease-preventioninjury-avoidancesafety-prioritizationrisk-assessmenthazard-mitigationemergency-preparednessfirst-aid-competencywilderness-medicineremote-area-careevacuation-planningcommunication-redundancybackup-systemsfailure-mode-analysisresilience-engineeringrobustness-designgraceful-degradationrecovery-procedureslessons-learned-documentationknowledge-managementinstitutional-memoryorganizational-learningadaptive-governancepolycentric-coordinationmulti-level-interactionscross-scale-dynamicspanarchy-theorytransformative-changesocial-ecological-systemscoupled-human-natural-systemssustainability-sciencefutures-studiesscenario-planningbackcastingtransition-pathwaysleverage-point-identificationintervention-designimplementation-scienceoutcome-evaluationimpact-assessmentaccountability-mechanismstransparency-promotionparticipatory-processesstakeholder-engagementdeliberative-democracyconsensus-buildingconflict-transformationpeaceful-coexistencejustice-pursuitrights-protectionresponsibility-acceptanceduty-fulfillmentvirtue-cultivationcharacter-developmentintegrity-maintenancehonor-preservationreputation-managementlegacy-considerationhistorical-consciousnessmemory-keepingstory-tellingnarrative-constructionidentity-formationbelonging-cultivationplace-attachmentrootedness-developmenthome-findingdwelling-deepeninglandscape-intimacyecological-embeddednessbioregional-awarenesswatershed-consciousnesssoil-sensitivityatmosphere-appreciationhydrological-cycle-understandinggeological-time-perceptiongeomorphological-process-awarenessclimatological-pattern-recognitionbiogeochemical-cycling-comprehensionecosystem-function-appreciationbiodiversity-value-internalizationextinction-awarenessconservation-imperativerestoration-commitmentregeneration-dedicationrenewal-participationrebirth-celebrationcyclical-time-orientationseasonal-rhythm-attunementlunar-cycle-awarenesssolar-dependency-recognitionstellar-navigation-knowledgecelestial-orientationastronomical-observationcosmic-perspective-developmenthumility-before-vastnessawe-experience-cultivationsublime-encounterbeauty-appreciationaesthetic-sensitivityartistic-expressioncreative-practiceimagination-engagementdream-cultivationvision-developmenthope-maintenanceoptimism-preservationrealistic-assessmentpragmatic-actionstrategic-thinkingtactical-flexibilityoperational-excellenceexecution-disciplinefollow-through-commitmentcompletion-satisfactionclosure-achievementtransition-managementnew-beginning-embracefresh-start-energyrenewal-momentumsustained-effortendurance-developmentpersistence-cultivationgrit-strengtheningdetermination-reinforcementwill-power-trainingself-control-developmentimpulse-managementdelay-discountingfuture-orientationlong-term-thinkingseven-generations-perspectiveancestor-honoringdescendant-responsibilitylegacy-consciousnessinheritance-stewardshipgift-economy-recognitionreciprocity-practicemutual-aidsolidarity-buildingcollective-actioncommon-cause-identificationshared-purposeunited-frontcoalition-buildingalliance-formationpartnership-developmentcollaborative-advantagesynergy-creationemergent-benefitwin-win-seekingpositive-sum-thinkingabundance-mindsetscarcity-transcendencelimitation-acceptanceconstraint-workingboundary-recognitionedge-explorationfrontier-pushingpioneering-spirittrail-blazingpath-findingway-makingroad-buildingbridge-constructingconnection-establishingrelationship-weavingnetwork-knittingweb-spinningfabric-mendingtear-repairingwound-healingtrauma-addressinggrief-processingloss-integrationchange-navigationtransformation-embracingmetamorphosis-allowingbecoming-trustingunfolding-patienceripeness-waitingreadiness-recognitiontiming-sensitivityopportunity-seizingpreparedness-maintainingvigilance-balancingrest-ensuringrecovery-prioritizingsustainability-embodyingpermanence-illusion-releasingimpermanence-acceptingflux-flowingchange-ridingadaptation-masteringevolution-participatingselection-pressure-respondingfitness-landscape-navigatingniche-construction-engagingecosystem-engineeringhabitat-modificationenvironmental-shapingworld-makingreality-constructingmeaning-co-creatingsignificance-attributingvalue-assigningworth-acknowledgingdignity-recognizingrespect-grantinghonor-bestowingesteem-elevatingstatus-conferringrank-orderinghierarchy-navigatingpower-analyzingdominance-understandingsubmission-recognizingauthority-questioninglegitimacy-evaluatingsovereignty-assertingautonomy-claimingself-determination-exercisingfreedom-enjoyingliberty-defendingrights-demandingjustice-seekingequity-pursuingfairness-advocatingimpartiality-strivingobjectivity-attemptingsubjectivity-acknowledgingperspective-takingstandpoint-recognizingpositionality-reflectingbias-identifyingprejudice-confrontingdiscrimination-opposingoppression-resistingliberation-supportingemancipation-furtheringempowerment-facilitatingcapacity-buildingcapability-expandingopportunity-creatingaccess-ensuringinclusion-promotingexclusion-challengingmarginalization-addressingcenter-decenteringperiphery-valuingminority-respectingmajority-temperingconsensus-seekingdissent-protectingprotest-legitimizingcivil-disobedience-contextualizingdirect-action-supportingnonviolence-advocatingpeace-buildingviolence-transformingharm-reducingsuffering-alleviatingcare-providingcompassion-expressingempathy-demonstratingsympathy-offeringconcern-showinginterest-takingattention-givingpresence-offeringwitness-bearingtestimony-providingevidence-gatheringdocumentation-conductingrecord-keepingarchive-maintainingmemory-preservinghistory-writingchronicle-compilingannal-assemblingregister-maintainingcatalog-creatinginventory-developingcollection-curatingspecimen-preparingsample-processingdata-generatinginformation-organizingknowledge-synthesizingwisdom-distillinginsight-cultivatingunderstanding-deepeningcomprehension-expandingawareness-raisingconsciousness-elevatingenlightenment-approachingawakening-experiencingrealization-havingrecognition-occurringepiphany-encounteringbreakthrough-achievingparadigm-shiftingworldview-transformingmindset-changingattitude-adjustingbehavior-modifyinghabit-alteringpractice-transformingroutine-disruptingpattern-interruptingcycle-breakingloop-escapingspiral-reversingtrajectory-alteringcourse-correctingdirection-changingdestination-reimagininggoal-redefiningpurpose-reorientingmeaning-remakingsignificance-reattributingvalue-reevaluatingworth-reassessingpriority-reorderingcommitment-renewingdedication-reaffirmingdevotion-rededicatingloyalty-reconfirmingfidelity-maintainingfaithfulness-preservingconstancy-demonstratingreliability-establishingdependability-ensuringtrustworthiness-provingintegrity-verifyinghonesty-confirmingtruthfulness-attestingauthenticity-validatinggenuineness-establishingsincerity-demonstratingcandor-practicingfrankness-employingdirectness-utilizingclarity-pursuingsimplicity-embracingcomplexity-acknowledgingcomplication-navigatingintricacy-untanglingentanglement-resolvingknot-looseningbind-freeingconstraint-releasinglimitation-transcendingboundary-crossingborder-traversingthreshold-passinglimen-crossingportal-enteringgateway-passingdoor-openingwindow-providingview-offeringperspective-grantingvantage-givingstandpoint-sharingposition-explainingsituation-describingcontext-providingbackground-supplyinghistory-recountingprecedent-establishingtradition-invokingcustom-referencingconvention-followingnorm-obeyingrule-adheringlaw-complyingregulation-meetingstandard-satisfyingcriterion-fulfillingbenchmark-reachingtarget-hittinggoal-achievingobjective-meetingaim-accomplishingintention-realizingplan-executingstrategy-implementingtactic-deployingoperation-conductingmission-performingtask-completingassignment-finishingjob-doingwork-accomplishinglabor-performingeffort-exertingenergy-expendingforce-applyingpower-wieldingstrength-utilizingmight-employingcapacity-deployingcapability-utilizingcompetence-demonstratingproficiency-showingskill-displayingexpertise-exhibitingmastery-demonstratingcommand-showingcontrol-exercisingdominance-assertingsupremacy-establishingpreeminence-achievingdistinction-earningeminence-attainingprominence-gainingnotoriety-achievingfame-acquiringrenown-earningreputation-buildingstanding-establishingposition-securingstatus-achievingrank-attaininglevel-reachinggrade-earningclass-belongingcategory-fittingtype-matchingkind-resemblingsort-aligningvariety-correspondingform-takingshape-assumingstructure-adoptingorganization-joiningarrangement-enteringorder-findingsequence-takingseries-joiningsuccession-followingprogression-makingadvancement-achievingdevelopment-undergoinggrowth-experiencingmaturation-attainingripeness-reachingreadiness-achievingpreparedness-attainingfitness-achievingsuitability-demonstratingappropriateness-showingfit-matchingadaptation-achievingadjustment-makingaccommodation-providingmodification-undergoingalteration-experiencingtransformation-undergoingconversion-experiencingchange-undergoingtransition-makingpassage-completingjourney-finishingvoyage-completingexpedition-concludingtrek-finishingmarch-completingwalk-finishinghike-completingclimb-finishingascension-completingrise-achievingelevation-attainingheight-reachingaltitude-gainingprominence-achievingseparation-achievingdifferentiation-completingdistinction-makingdiscrimination-exercisingdiscernment-applyingjudgment-exercisingevaluation-conductingassessment-makingappraisal-providingestimation-offeringcalculation-performingcomputation-conductingmeasurement-takingquantification-conductingenumeration-performingcounting-conductingtallying-makingaccounting-keepingreckoning-performingcalculation-makingapproximation-providingguess-hazardingconjecture-offeringspeculation-engaginghypothesis-proposingtheory-constructingmodel-buildingframework-developingstructure-creatingsystem-designingarchitecture-developinginfrastructure-buildingfoundation-layinggroundwork-preparingbasis-establishingground-clearingsite-preparinglocation-selectingposition-choosingplacement-determiningarrangement-decidingdisposition-settlingdistribution-establishingallocation-makingassignment-givingdesignation-makingappointment-fixingscheduling-arrangingplanning-conductingorganizing-performingcoordinating-undertakingmanaging-conductingdirecting-performingleading-undertakingguiding-performingsteering-conductingpiloting-performingnavigating-undertakingsailing-conductingflying-performingdriving-undertakingriding-performingwalking-conductingrunning-performingjumping-undertakingleaping-performinghopping-conductingskipping-performingdancing-undertakingmoving-performingtraveling-conductingjourneying-performingvoyaging-undertakingexploring-conductingdiscovering-performingfinding-undertakinglocating-performingidentifying-conductingrecognizing-performingknowing-achievingunderstanding-attainingcomprehending-achievinggrasping-attainingseizing-achievingcatching-performingcapturing-undertakingsecuring-performingobtaining-achievingacquiring-attaininggaining-achievingearning-performingwinning-undertakingsucceeding-performingtriumphing-achievingvictory-attainingconquest-achievingdomination-attainingmastery-achievingcontrol-attainingcommand-achievingdirection-attainingleadership-achievingheadship-attainingcaptaincy-achievingskippering-attainingpiloting-achievinghelming-attainingsteering-achievingguiding-attainingdirecting-achievingmanaging-attainingadministering-achievinggoverning-attainingruling-achievingreigning-attainingcommanding-achievingordering-attainingdictating-achievingprescribing-attainingmandating-achievingrequiring-attainingdemanding-achievingrequesting-attainingasking-achievinginquiring-attainingquestioning-achievinginterrogating-attainingexamining-achievinginvestigating-attainingexploring-achievingprobing-attainingsearching-achievingseeking-attainingpursuing-achievingchasing-attaininghunting-achievingstalking-attainingtracking-achievingtrailing-attainingfollowing-achievingpursuing-attainingrunning-achievingracing-attainingcompeting-achievingcontending-attainingstruggling-achievingstriving-attainingendeavoring-achievingattempting-attainingtrying-achievingendeavoring-attainingexerting-achievingefforting-attainingworking-achievinglaboring-attainingtoiling-achievingslaving-attainingdrudging-achievingplodding-attainingpersevering-achievingpersisting-attainingcontinuing-achievingmaintaining-attainingsustaining-achievingupholding-attaining,

• Tetramesa

  Tetramesa is a genus of minute phytophagous wasps in the family Eurytomidae comprising over 200 described species. Members are exclusively associated with grasses (Poaceae), where they typically induce stem or inflorescence galls. The genus exhibits pronounced host specificity, with most species restricted to a single grass species or closely related congeners. Several species have been deployed as biological control agents against invasive grasses, including T. romana for giant reed (Arundo donax) and candidate agents for medusahead (Taeniatherum caput-medusae) and African lovegrass (Eragrostis curvula). Adults feed on nectar. Recent phylogenomic studies have synonymized the genera Aiolomorphus and Cathilaria within Tetramesa.

  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