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Acentria
watermilfoil moth, water veneer
Acentria is a monotypic moth genus in the family Crambidae containing a single species, Acentria ephemerella. The genus is notable for extreme sexual dimorphism in wing development and for having the only known fully aquatic adult female moths. Most females are flightless with rudimentary wings and spend their entire lives underwater, while males are fully winged and terrestrial. The species is native to Europe and has been introduced to North America.
freshwater sexual-dimorphism brachypterous biocontrol invasive-species-management aquatic-insect case-building-larva monotypic-genus
Agroecotettix
Aridland Scrub Jumpers
Agroecotettix is a genus of medium-sized brachypterous grasshoppers in the subfamily Melanoplinae, containing 19 described species. These short-winged, flight-limited grasshoppers inhabit aridland scrub and desert regions of the southwestern United States and Mexico. The genus was revised in 2024 with the description of 16 new species, revealing previously unrecognized diversity in this cryptic group. Species identification relies primarily on male genitalia morphology.
grasshopper orthoptera acrididae melanoplinae brachypterous desert xeric cryptic-species new-species southwestern-US Mexico Texas
Agrothereutes abbreviatus
Agrothereutes abbreviatus is a parasitic wasp in the family Ichneumonidae, characterized by sexual dimorphism in wing development. Females are brachypterous with wings reduced to the postscutellum or first abdominal tergite, while males have fully developed wings. The species is active in spring and searches for hosts on the ground and in leaf litter.
brachypterous sexual-dimorphism ground-dwelling parasitoid Ichneumonidae
Amaurorhinus bewickianus
Amaurorhinus bewickianus is a weevil species in the family Curculionidae, endemic to the Madeira archipelago. It was first described by Wollaston in 1860 and has been recorded from Madeira Island and the nearby Desertas Islands. The species belongs to a small genus of flightless weevils with restricted distributions in the Macaronesian region. Its island endemism and apparent absence from mainland Europe suggest limited dispersal capability.
endemic Madeira Macaronesia flightless island-weevil brachypterous
Anafroptilum
Anafroptilum is a genus of small mayflies in the family Baetidae. The genus was first described from the East Palaearctic and Nearctic regions, with a single species A. kazlauskasi known from the former and eight species from the latter. In 2016, two new species were described from Thailand, marking the first report of the genus from the Oriental Region. Species in this genus are characterized by the absence of hind wings in adults and distinctive larval morphological features.
Ephemeroptera Baetidae mayfly Oriental-Region Thailand wingless brachypterous
Areniscythris
Areniscythris is a genus of small moths in the family Scythrididae, established by Powell in 1976. The genus contains two described species: A. brachypteris and A. whitesands, both associated with sandy habitats. The genus name references its association with sand (Latin 'arena' = sand).
moth Scythrididae sand-dune brachypterous North-America
Areniscythris brachypteris
Oso Flaco Flightless Moth, sand-dune grasshopper moth
Areniscythris brachypteris is a flightless moth in the family Scythrididae, described by Jerry A. Powell in 1976. It is endemic to coastal sand dunes of California and is notable among Lepidoptera for its reduced wings and inability to fly. The species has been observed in 90 iNaturalist records, suggesting it is documented but not abundant.
flightless endemic brachypterous coastal-dune-specialist California-endemic
Belocephalus
short-winged coneheads
Belocephalus is a genus of short-winged conehead katydids comprising approximately eight described species. Members of this genus are characterized by abbreviated wing development and are endemic to the southeastern United States. The genus was established by Scudder in 1875 and belongs to the tribe Copiphorini within the subfamily Conocephalinae.
Orthoptera Tettigoniidae Conocephalinae Copiphorini conehead-katydids brachypterous southeastern-United-States endemic
Blatta orientalis
Oriental Cockroach, Waterbug
Blatta orientalis is a large, dark-colored cockroach and a significant urban pest. Adult males measure 18–29 mm and females 20–27 mm. Females appear wingless but are brachypterous with non-functional wing pads; males have reduced wings that do not cover the abdomen. The species thrives in dark, damp environments such as basements, crawl spaces, and sewers. It is a major vector for pathogens including E. coli, Staphylococcus, and Salmonella, and proteins from this species can trigger allergic reactions and asthma.
urban-pest vector allergen cosmopolitan brachypterous
Booneacris
Wingless Mountain Grasshoppers
Booneacris is a genus of spur-throated grasshoppers in the family Acrididae, established by Rehn & Randell in 1962. The genus contains four described species, all characterized by reduced or absent wings. These grasshoppers are restricted to mountainous regions of western North America, with species found at high elevations. The common name "Wingless Mountain Grasshoppers" reflects their most distinctive trait and habitat preference.
wingless brachypterous alpine mountain endemic western-North-America Melanoplinae Podismini
Bostaera balli
Bostaera balli is a species of delphacid planthopper in the family Delphacidae, first described by Penner in 1952. It belongs to the genus Bostaera, a group of small planthoppers characterized by their reduced wings and association with grasses and sedges. The species is known from limited records in the southeastern United States.
Delphacidae planthopper North-America southeastern-United-States brachypterous grass-associated
Calosoma discors
different beautiful black searcher
Calosoma discors is a species of ground beetle in the subfamily Carabinae, described by John Lawrence LeConte in 1857. This species is known from California and Washington, where it inhabits coastal grasslands and oak savannahs in the Sierra Nevada foothills. Adults are brachypterous, meaning they have reduced wings that limit flight capability.
ground-beetle Carabidae brachypterous California Washington Sierra-Nevada coastal-grassland oak-savannah
Calosoma haydeni
Hayden's caterpillar hunter
Calosoma haydeni is a ground beetle species in the family Carabidae, described by Horn in 1870. It belongs to the genus Calosoma, commonly known as caterpillar hunters. Adults are brachypterous (short-winged), which limits their flight capability. The species occurs in Mexico and the southern United States, with two recognized subspecies showing distinct geographic distributions.
brachypterous short-winged caterpillar-hunter subspecies Horn-1870
Calosoma latipenne
broad beautiful black searcher
Calosoma latipenne is a large ground beetle in the family Carabidae, commonly known as the broad beautiful black searcher. It inhabits sandy soils in arid and semi-arid regions of the southwestern United States. Adults are brachypterous (short-winged) and have been observed preying on grasshopper species. The species belongs to a genus renowned for its caterpillar-hunting behavior, though specific ecological details for this species remain limited.
brachypterous predator arid-habitat grasshopper-predator
Calosoma luxatum
dislocated beautiful black searcher
Calosoma luxatum, commonly known as the dislocated beautiful black searcher, is a large ground beetle in the family Carabidae. Adults are brachypterous, meaning they have reduced wings and are flightless. The species is distributed across western and central North America, from the Canadian prairies through the western United States. It inhabits open short grass prairies and dry forest clearings, where it hunts as an active predator.
Carabidae Calosoma ground-beetle brachypterous prairie predator North-America
Calosoma moniliatum
collared beautiful black searcher
Calosoma moniliatum, commonly known as the collared beautiful black searcher, is a large predatory ground beetle in the family Carabidae. Adults are brachypterous (short-winged and flightless). The species occurs in western North America, inhabiting short grass prairies and dry forests.
Carabidae Calosoma caterpillar-hunter brachypterous flightless prairie western-North-America
Calosoma monticola
mountain beautiful black searcher
Calosoma monticola is a ground beetle in the family Carabidae, described by Casey in 1913. It is found in the western United States, where it inhabits sagebrush areas. Adults are brachypterous (short-winged) and have been documented preying on various soil-dwelling insect larvae and other beetles. The species belongs to a genus known for active predation on caterpillars and other soft-bodied insects.
Carabidae ground-beetle predator brachypterous sagebrush western-North-America biological-control
Calosoma subaeneum
coppery beautiful black searcher
Calosoma subaeneum is a species of ground beetle in the subfamily Carabinae, described by Maximilien Chaudoir in 1869. It belongs to the genus Calosoma, commonly known as caterpillar hunters or searchers, which are large predatory beetles. Adults are brachypterous (short-winged), a trait that limits dispersal ability. The species occurs in lowland areas of western North America.
ground-beetle caterpillar-hunter brachypterous western-North-America lowland
Calosoma wilkesii
Wilkes's beautiful black searcher
Calosoma wilkesii is a flightless ground beetle in the family Carabidae, commonly known as Wilkes's beautiful black searcher. It is one of approximately 120 species in the genus Calosoma, a group known as "caterpillar hunters" for their predatory habits on lepidopteran larvae. The species has been the subject of genomic research due to its brachypterous (short-winged) condition, which represents an evolutionary transition to flightlessness. It occurs in western North America from British Columbia to California.
brachypterous flightless caterpillar-hunter genomic-model nocturnal western-North-America
Carabus chamissonis
Fisher's worm and slug hunter
Carabus chamissonis is a ground beetle species restricted to tundra environments across northern North America. It occupies open, dry tundra habitats with better drainage than those used by the related Carabus truncaticollis. The species exhibits brachyptery (reduced wings) and nocturnal activity patterns. Adults overwinter within their own pupal cavities, an unusual behavior among ground beetles. Disjunct populations occur on isolated mountaintops in New England, far south of the main Arctic range.
ground-beetle tundra Arctic brachypterous nocturnal alpine disjunct-distribution
Carabus finitimus
Haldeman's worm and slug hunter
Carabus finitimus is a bluish-colored ground beetle in the family Carabidae. It inhabits bottomland forests along rivers in Kansas, Oklahoma, and Texas. Adults are brachypterous (short-winged) and nocturnal. The common name "Haldeman's worm and slug hunter" reflects its predatory habits.
ground-beetle nocturnal brachypterous riparian predator
Carabus forreri
Forrer's worm and slug hunter
Carabus forreri is a ground beetle species in the family Carabidae, known from dry oak-pine forests in the southwestern United States and northern Mexico. Adults are brachypterous (short-winged) and nocturnal. The species is part of the large genus Carabus, which includes many predatory ground beetles.
brachypterous nocturnal ground-beetle oak-pine-forest southwestern-North-America
Carabus goryi
Gory's worm and slug hunter
Carabus goryi is a North American ground beetle in the family Carabidae, commonly known as Gory's worm and slug hunter. The species inhabits cool, moist forests as well as agricultural settings including orchards and cultivated fields. Adults are brachypterous (short-winged), nocturnal, and exhibit gregarious behavior. They are predatory, feeding on soft-bodied insect larvae including those of the spongy moth (Lymantria dispar).
Carabidae ground-beetle predator brachypterous nocturnal gregarious North-America forest agricultural biological-control
Carabus sylvosus
woodland ground beetle, sylvan worm and slug hunter, silvan worm and slug hunter
Carabus sylvosus, commonly known as the woodland ground beetle or sylvan worm and slug hunter, is a species of ground beetle in the family Carabidae. It is native to North America, where it inhabits wooded environments including deciduous mixed forests, coniferous forests, and orchards. Adults are brachypterous (short-winged) and nocturnal, preying on Lepidoptera pupae and scarabaeids. The species has been documented in Canada and the United States.
ground-beetle Carabidae brachypterous nocturnal predator North-America woodland
Cosumnoperla hypocrena
Cosumnes Spring Stonefly
Cosumnoperla hypocrena is a species of spring stonefly in the family Perlodidae, described from California in 1987. The genus Cosumnoperla contains species associated with spring-fed habitats. As a member of the subfamily Isoperlinae, it shares characteristics with other western North American perlodids. The specific epithet refers to the type locality near the Cosumnes River.
Plecoptera Perlodidae spring-stonefly California-endemic groundwater-dependent brachypterous
Cychrus tuberculatus
tuberculate rare snail-eating beetle
Cychrus tuberculatus is a species of ground beetle in the subfamily Carabinae, commonly known as the tuberculate rare snail-eating beetle. First described by T.W. Harris in 1839, this species is notable for its specialized diet of snails and its brachypterous (short-winged), nocturnal adults. It inhabits coniferous forests, forest clearings, and moraines in western North America, with documented populations in British Columbia, Quebec, California, Oregon, and Washington. The species is considered rare and has specific habitat requirements tied to forested environments.
ground-beetle snail-predator brachypterous nocturnal coniferous-forest rare-species western-North-America Cychrini specialized-predator
Cyphomyrmex
fungus-growing ants
Cyphomyrmex is a genus of small, drab-colored fungus-growing ants in the tribe Attini, found primarily in the Neotropics. These ants cultivate fungi in the tribe Leucocoprineae as their primary food source, with most species growing fungal nodules rather than full mycelial gardens. Colonies are typically monogynous and small, rarely exceeding 500 workers. The genus is divided into two species complexes: the strigatus complex (South America only) and the rimosus complex (southern North America to South America). Cyphomyrmex represents a basal lineage among attine ants and serves as sister group to Mycetophylax.
fungus-growing-ants attine-ants Myrmicinae Neotropical mutualism fungiculture Leucocoprineae monogyny parasitoid-wasps social-parasitism chemical-ecology alkaloid-venom nest-architecture behavioral-plasticity cytogenetics chromosome-evolution Guiana-Shield Brazil Panama Puerto-Rico Mesoamerica Argentina United-States Texas California Florida yeast-gardens mycelial-gardens rimosus-complex strigatus-complex age-polyethism trophallaxis thanatosis feigning-death Diapriidae Acanthopria Mimopriella Megalomyrmex social-parasite C.-longiscapus C.-transversus C.-laevigatus C.-rimosus C.-minutus C.-lectus C.-cornutus C.-costatus pheromones 3-octanol farnesenes trail-following alarm-signal diketopiperazines antifungal-compounds defensive-behavior colony-defense flight-response recruitment Lanchester's-laws karyotype chromosome-number rDNA microsatellites FISH chamber-depth gallery-length fire-disturbance savanna-forest-transition arboreal-nesting clay-auricles soil-nests wood-nests epiphytic-nests moss-nests coconut-nests desiccation-susceptibility moist-habitats basal-attine phylogenetics sister-group chemical-evolution venom-alkaloids behavioral-manipulation host-parasite-interaction ecosystem-engineering nutrient-cycling decomposition organic-matter insect-feces exoskeleton-substrate garden-transplantation worker-polymorphism queen-number dealate-queens alates brood-care ecdysis-assistance foraging sugar-collection plant-exudates fungal-domestication nodule-cultivation mycelium-cultivation agricultural-behavior nest-entrance funnel-structure exposed-gardens symbiont-diversity geographic-variation research-model evolutionary-biology mutualism-evolution ant-fungus-coevolution chemical-communication social-insects eusociality colony-size small-colonies slow-movement crypsis debris-mimicry predation-avoidance parasitoid-defense wasp-killing colony-abandonment resource-rescue thief-ant-defense venom-resistance toxicity-tolerance behavioral-suppression playing-dead alkaloid-manipulation fungal-secondary-metabolites antibacterial-compounds antiviral-compounds garden-protection microbial-defense nest-microclimate temperature-regulation humidity-requirements depth-effects fire-adaptation disturbance-ecology habitat-specificity substrate-specialization clay-construction soil-manipulation architectural-plasticity vertical-nesting embankment-nesting overhang-nesting pendant-nests swallow-nest-mimicry research-accessibility field-biology tropical-ecology Neotropical-myrmecology biodiversity conservation invasive-species-context native-species biological-control Integrated-Pest-Management agricultural-ecology forest-ecology savanna-ecology transitional-habitats disturbed-areas urban-ecology anthropogenic-impact climate-sensitivity habitat-limitation range-boundaries dispersal colonization establishment population-dynamics demography colony-density nest-spacing intraspecific-competition interspecific-competition territoriality resource-defense combat-behavior aggression-levels recruitment-behavior chemical-signaling alarm-pheromones trail-pheromones recognition-cues cuticular-hydrocarbons venom-composition defensive-secretions mandibular-glands Dufour's-gland metapleural-glands antibiotic-production grooming-behavior fungal-inoculation integumental-fungi yeast-growth larval-grooming garden-maintenance substrate-preparation fecal-fertilizer anal-trophallaxis crop-regurgitation fluid-application drying-behavior nodule-formation transplantation-behavior garden-establishment ceiling-gardens root-association architectural-innovation nest-site-selection microhabitat-choice moisture-requirements thermal-buffering predator-avoidance parasite-avoidance disease-resistance social-immunity collective-behavior division-of-labor task-allocation temporal-polyethism worker-age nurse-workers forager-workers soldier-caste-absence monomorphism worker-size-variation queen-worker-differentiation reproductive-caste gyne-production male-production seasonal-reproduction colony-founding claustral-founding non-claustral-founding fungal-inoculum founding-queen first-worker-generation nutritional-dependency fat-reserves metabolic-adaptation digestive-physiology enzyme-complement fungal-enzyme lignocellulose-degradation nutritional-mutualism obligate-mutualism vertical-transmission horizontal-transmission symbiont-switching cultivar-replacement garden-failure colony-mortality population-turnover generation-time longevity survivorship life-expectancy aging senescence reproductive-investment colony-growth colony-decline budding fission reproduction-mode genetic-structure relatedness mating-system nuptial-flight mate-choice sperm-storage colony-odor nestmate-recognition foreign-rejection agonistic-behavior territorial-marking boundary-defense raiding-behavior robbing-behavior cleptobiosis dulosis slavery inquilinism guest-ants myrmecophiles nest-associates commensalism inquilines parasite-tolerance host-manipulation venom-alkaloid pyrrolidine indolizidine piperidine nonanal farnesene sesquiterpene agriculture-chemical gaster-extract chemical-diversity chemical-simplicity evolutionary-precursor derived-chemistry biosynthetic-pathway semiochemical infochemical communication information-transfer colony-organization social-structure polygyny queen-condition dealation wing-shedding ovarian-development fecundity egg-laying brood-development developmental-rate temperature-dependence humidity-dependence food-availability nutritional-quality fungal-productivity garden-yield worker-production colony-efficiency energy-allocation resource-allocation foraging-efficiency search-behavior prey-detection food-retrieval transport-behavior load-size travel-speed path-integration orientation navigation homing-behavior learning memory experience individual-recognition context-dependent-behavior threat-assessment risk-management predator-inspection alarm-propagation recruitment-activation mobilization colony-response emergency-behavior panic-response controlled-response defensive-strategy offensive-strategy escalation de-escalation conflict-resolution fight-avoidance retreat-behavior surrender-signal dominance-hierarchy reproductive-skew worker-policing queen-policing egg-eating oviposition-suppression reproductive-conflict kin-conflict worker-reproduction male-production-by-workers anarchy social-parasitism-detection parasite-recognition enemy-recognition nest-intrusion nest-defense perimeter-defense chamber-defense brood-defense queen-defense self-sacrifice suicidal-defense autothysis sting-use mandible-use spraying-behavior gaster-flexion chemical-defense mechanical-defense armor cuticle-thickness pigmentation masquerade mimicry Batesian-mimicry Muellerian-mimicry aggressive-mimicry Wasmannian-mimicry myrmecophily myrmecophagy ant-plant-mutualism trophobiosis extrafloral-nectary domatia plant-ant-interaction herbivore-defense seed-dispersal pollination ecosystem-service functional-role trophic-position detritivore decomposer saprotroph necrotroph fungivore omnivore predator scavenger trophic-level energy-flow carbon-cycling nitrogen-cycling phosphorus-cycling soil-formation bioturbation aeration water-infiltration microbial-community bacteria actinomycetes Pseudonocardia disease-suppression garden-hygiene weeding-behavior pruning-behavior fungal-grooming antibiotic-application mutualist-protection agricultural-disease crop-pathology escovopsis garden-parasite competitor-fungus pathogen-defense immune-response individual-immunity grooming allogrooming self-grooming nest-cleaning refuse-management fecal-management waste-disposal midden-formation territory-size home-range foraging-range recruitment-range exploration exploitation information-use public-information private-information cue-use signal-use communication-network interaction-network social-network colony-network population-network community-ecology species-interaction apparent-competition exploitative-competition interference-competition resource-partitioning niche-differentiation habitat-filtering environmental-gradient altitudinal-distribution latitudinal-distribution continental-distribution island-distribution endemism biogeography phylogeography historical-biogeography vicariance range-expansion range-contraction refugia glacial-cycles climate-change-impact habitat-fragmentation deforestation land-use-change agricultural-intensification urbanization pollution pesticide-exposure heavy-metal-accumulation conservation-status population-viability extinction-risk protected-area reserve-design habitat-corridor connectivity metapopulation source-sink-dynamics rescue-effect mass-effect ecological-theory behavioral-ecology evolutionary-ecology functional-ecology trait-based-ecology comparative-analysis phylogenetic-comparative-methods character-evolution ancestral-state-reconstruction convergent-evolution parallel-evolution adaptive-radiation species-diversification speciation hybridization introgression gene-flow genetic-differentiation population-structure molecular-ecology genomics transcriptomics proteomics metabolomics volatile-organic-compound cuticular-hydrocarbon glandular-secretion sting-apparatus mandible-morphology head-morphology mesosomal-morphology petiolar-morphology gastral-morphology leg-morphology antennal-morphology sensilla chemoreception mechanoreception thermoreception photoreception vision compound-eye ocelli brain-morphology neuroanatomy learning-capacity memory-capacity cognitive-ability problem-solving tool-use sequence-learning route-learning spatial-memory temporal-memory social-learning cultural-transmission tradition colony-personality behavioral-syndrome boldness exploration-tendency activity-level aggression-level foraging-tendency defensive-tendency stress-response physiological-stress thermal-stress desiccation-stress starvation-stress immune-challenge wound-healing metabolic-rate respiration energy-expenditure water-balance osmoregulation excretion Malpighian-tubules rectum hindgut midgut foregut crop proventriculus digestive-enzyme nutrient-absorption lipid-metabolism carbohydrate-metabolism protein-metabolism amino-acid-metabolism nitrogen-excretion uric-acid urea ammonia osmolyte cryoprotectant heat-shock-protein stress-protein antioxidant-defense oxidative-stress reactive-oxygen-species aging-biology senescence-theory disposable-soma antagonistic-pleiotropy mutation-accumulation life-history-theory r-K-selection bet-hedging iteroparity semelparity reproductive-effort parental-investment maternal-effect paternal-effect epigenetics DNA-methylation histone-modification non-coding-RNA gene-regulation transcription-factor signaling-pathway hormone juvenile-hormone ecdysone insulin-signaling TOR-signaling nutrient-sensing reproductive-signaling caste-determination sex-determination environmental-sex-determination genetic-sex-determination haplodiploidy arrhenotoky thelytoky parthenogenesis arrhenotokous-parthenogenesis automixis apomixis central-fusion terminal-fusion genomic-imprinting parental-conflict kin-selection inclusive-fitness Hamilton's-rule relatedness-asymmetry sex-ratio-theory local-mate-competition local-resource-competition local-resource-enhancement split-sex-ratio worker-sex-ratio queen-sex-ratio colony-sex-ratio population-sex-ratio operational-sex-ratio effective-population-size genetic-drift inbreeding inbreeding-depression outbreeding outbreeding-depression heterosis hybrid-vigor genetic-load mutation-rate effective-number-of-codons codon-usage-bias GC-content genome-size karyotype-evolution chromosomal-rearrangement fusion inversion translocation polyploidy aneuploidy chromosomal-polymorphism rDNA-cluster heterochromatin euchromatin centromere telomere nucleolus-organizer-region satellite-DNA minisatellite microsatellite simple-sequence-repeat transposable-element retrotransposon DNA-transposon horizontal-gene-transfer endosymbiont Wolbachia Cardinium Rickettsia Spiroplasma bacterial-symbiont fungal-symbiont viral-symbiont parasite pathogen disease epidemiology transmission-dynamics virulence resistance tolerance recovery immunity vaccination prophylaxis medicinal-behavior self-medication thermoregulation behavioral-thermoregulation social-thermoregulation microclimate-selection nest-temperature brood-temperature fungal-garden-temperature metabolic-heat heat-dissipation evaporative-cooling behavioral-cooling fanning ventilation chamber-design gallery-design entrance-design air-flow humidity-regulation water-collection water-storage desiccation-resistance cuticular-waterproofing spiracle-control respiratory-water-loss excretory-water-loss water-recycling metabolic-water dietary-water environmental-water rainfall-dependence seasonal-activity dry-season wet-season monsoon drought flood fire disturbance succession primary-succession secondary-succession climax-community pioneer-species habitat-specialist habitat-generalist edge-species interior-species gap-species canopy-species understory-species ground-species subterranean-species arboreal-species liana-species epiphyte-species lithophyte-species rheophyte-species aquatic-margin-species terrestrial-species fossorial-species cursorial-species scansorial-species volant-species winged-reproductive dealate ergatoid brachypterous apterous polymorphism caste-polymorphism queen-polymorphism male-polymorphism size-polymorphism allometry isometry shape-variation morphological-integration modularity evolvability constraints developmental-stability fluctuating-asymmetry directional-asymmetry antisymmetry phenotypic-plasticity reaction-norm genotype-environment-interaction grandparental-effect indirect-genetic-effect social-environment-effect colony-effect density-dependence frequency-dependence negative-frequency-dependence positive-frequency-dependence apostatic-selection anti-apostatic-selection search-image predator-learning prey-learning friend-recognition cuticular-hydrocarbon-profile gestalt-odor colony-specific-odor individual-specific-odor task-specific-odor age-specific-odor reproductive-odor queen-odor queen-pheromone primer-pheromone releaser-pheromone signal-pheromone modulatory-pheromone alarm-pheromone trail-pheromone recruitment-pheromone aggregation-pheromone sex-pheromone marking-pheromone territorial-pheromone defensive-pheromone poison-gland Dufour-gland mandibular-gland postpharyngeal-gland metapleural-gland sternal-gland tarsal-gland anal-gland rectal-gland pygidial-gland mandibular-brush tibial-spur stridulatory-organ subgenual-organ Johnston's-organ tympanal-organ hair-sensillum campaniform-sensillum basiconic-sensillum coeloconic-sensillum placoid-sensillum sensilla-trichodea sensilla-basiconica sensilla-coeloconica sensilla-ampullacea sensilla-campaniformia sensilla-placodea olfactory-receptor-neuron gustatory-receptor-neuron mechanoreceptor-neuron thermoreceptor-neuron hygroreceptor-neuron photoreceptor-neuron visual-processing olfactory-processing gustatory-processing mechanosensory-processing central-complex mushroom-body antennal-lobe optic-lobe suboesophageal-ganglion ventral-nerve-cord sympathetic-nervous-system neurosecretion neurohemal-organ corpus-cardiacum corpus-allatum prothoracic-gland ventral-gland pericardial-gland neuroendocrine-regulation hormonal-regulation pheromonal-regulation neural-regulation behavioral-regulation social-regulation environmental-regulation developmental-regulation reproductive-regulation metabolic-regulation homeostatic-regulation allostatic-regulation stress-regulation immune-regulation circadian-rhythm circannual-rhythm photoperiodism thermoperiodism biological-clock zeitgeber entrainment free-running-period phase-response-curve masking-effect arousal-threshold sleep torpor diapause quiescence dormancy aestivation hibernation cold-hardiness heat-hardiness desiccation-hardiness anoxia-hardiness starvation-hardiness radiation-hardiness pressure-hardiness gravity-perception magnetic-field-perception electric-field-perception sound-perception vibration-perception substrate-borne-signal air-borne-signal seismic-signal chemical-signal visual-signal tactile-signal auditory-signal multimodal-signal composite-signal redundant-signal amplifying-signal attenuating-signal honest-signal deceptive-signal costly-signal cheap-signal conventional-signal index-signal handicap-signal amplifier social-selection sexual-selection sperm-competition cryptic-female-choice male-male-competition female-female-competition parent-offspring-conflict sibling-conflict worker-queen-conflict worker-worker-conflict sexual-conflict antagonistic-coevolution arms-race evolutionary-dynamics community-dynamics ecosystem-dynamics biogeochemical-cycle carbon-cycle nitrogen-cycle phosphorus-cycle sulfur-cycle water-cycle trophic-transfer production-efficiency consumption-efficiency assimilation-efficiency ecological-efficiency Lindeman-efficiency 10%-rule food-chain food-web trophic-cascade top-down-control bottom-up-control wasps-control donor-control recipient-control interaction-strength interaction-web ecological-network network-topology network-metrics degree-distribution connectance nestedness compartmentalization robustness fragility resilience recovery-time return-time elasticity perturbation pulse-disturbance press-disturbance chronic-disturbance acute-disturbance catastrophic-disturbance natural-disturbance anthropogenic-disturbance habitat-destruction habitat-degradation habitat-modification habitat-loss habitat-isolation edge-effect matrix-effect permeability corridor-effect stepping-stone source-population sink-population metapopulation-dynamics extinction-debt colonization-credit relaxation-effect time-lag historical-legacy priority-effect founder-effect bottleneck-effect selection natural-selection artificial-selection directional-selection stabilizing-selection disruptive-selection balancing-selection frequency-dependent-selection density-dependent-selection r-selection K-selection c-selection s-selection a-selection bet-hedging-selection fluctuating-selection correlational-selection multivariate-selection indirect-selection correlated-response genetic-correlation phenotypic-correlation environmental-correlation correlation-matrix variance-covariance-matrix G-matrix P-matrix E-matrix M-matrix quantitative-genetics heritability narrow-sense-heritability broad-sense-heritability genetic-variance additive-genetic-variance dominance-variance epistatic-variance environmental-variance phenotypic-variance genotype-environment-interaction-variance maternal-variance paternal-variance common-environment-variance special-environment-variance measurement-error-variance response-to-selection realized-heritability predicted-response breeder's-equation selection-gradient selection-differential Lande-equation multivariate-breeder's-equation genetic-constraint evolutionary-constraint developmental-constraint functional-constraint phylogenetic-constraint historical-constraint trade-off allocation-trade-off acquisition-trade-off conversion-trade-off maintenance-trade-off reproduction-survival-trade-off growth-reproduction-trade-off immune-reproduction-trade-off stress-reproduction-trade-off foraging-predation-trade-off competition-colonization-trade-off tolerance-resistance-trade-off plasticity-canalization-trade-off generalist-specialist-trade-off speed-accuracy-trade-off exploration-exploitation-trade-off individual-colony-trade-off within-colony-trade-off between-colony-trade-off life-history-trade-off demographic-trade-off ecological-trade-off evolutionary-trade-off optimization maximization minimization satisficing marginal-value-theorem optimal-foraging-theory optimal-diet-theory central-place-foraging patch-use-theory giving-up-density giving-up-time predation-risk energetic-cost opportunity-cost handling-time search-time travel-time encounter-rate prey-density prey-profitability prey-choice diet-breadth niche-width niche-overlap niche-partitioning resource-selection habitat-selection settlement-choice nest-site-choice host-choice oviposition-choice foraging-choice path-choice route-choice central-place territory core-area edge-area boundary perimeter area volume shape configuration spatial-pattern temporal-pattern diel-pattern seasonal-pattern annual-pattern multiannual-pattern cyclical-pattern irregular-pattern random-pattern clumped-pattern uniform-pattern regular-pattern aggregated-pattern overdispersed-pattern underdispersed-pattern Poisson-distribution negative-binomial-distribution Taylor's-power-law spatial-autocorrelation temporal-autocorrelation cross-correlation spectral-analysis wavelet-analysis time-series-analysis population-time-series community-time-series ecosystem-time-series long-term-study long-term-monitoring long-term-experiment chronosequence space-for-time-substitution paleoecology paleontology fossil-record trace-fossil coprolite nest-fossil amber-inclusion subfossil quaternary holocene pleistocene neogene paleogene cretaceous mesozoic cenozoic geological-time deep-time evolutionary-time ecological-time generational-time physiological-time perception-time reaction-time decision-time memory-time learning-time developmental-time life-span colony-lifespan queen-lifespan worker-lifespan male-lifespan larval-development-time pupal-development-time egg-development-time incubation-time doubling-time half-life relaxation-time resilience-time resistance-time lag-time lead-time forecast-time prediction-time planning-time management-time conservation-time restoration-time mitigation-time adaptation-time speciation-time diversification-time extinction-time persistence-time transience-time ephemerality permanence stability instability equilibrium non-equilibrium steady-state dynamic-state transient-state asymptotic-state attractor repeller limit-cycle chaos stochasticity determinism predictability unpredictability complexity emergence self-organization pattern-formation spatial-self-organization temporal-self-organization spatiotemporal-self-organization swarm-intelligence collective-decision-making consensus-decision quorum-response wisdom-of-crowds information-cascade cultural-evolution gene-culture-coevolution niche-construction extended-phenotype extended-organism superorganism colony-as-individual colony-as-unit-of-selection group-selection multilevel-selection reciprocal-altruism by-product-mutualism amensalism competition predation herbivory parasitism parasitoidism chemical-mimicry tactile-mimicry visual-mimicry acoustic-mimicry defensive-mimicry Vavilovian-mimicry Bakerian-mimicry Gilbertian-mimicry Poultonian-mimicry Browerian-mimicry auto-mimicry intra-specific-mimicry inter-specific-mimicry molecular-mimicry cellular-mimicry tissue-mimicry organ-mimicry behavioral-mimicry reproductive-mimicry social-mimicry nest-mimicry food-mimicry predator-mimicry prey-mimicry host-mimicry guest-mimicry cleptoparasite cuckoo-parasite inquiline guest myrmecophile termitophile aphidophile coccidophile scale-insect-associate mealybug-associate whitefly-associate planthopper-associate treehopper-associate leafhopper-associate cicada-associate butterfly-associate moth-associate beetle-associate fly-associate wasp-associate bee-associate ant-associate spider-associate mite-associate tick-associate nematode-associate flatworm-associate roundworm-associate annelid-associate mollusk-associate crustacean-associate millipede-associate centipede-associate insect-associate vertebrate-associate bird-associate mammal-associate reptile-associate amphibian-associate fish-associate plant-associate fungus-associate bacteria-associate virus-associate archaea-associate protist-associate algae-associate lichen-associate bryophyte-associate pteridophyte-associate gymnosperm-associate angiosperm-associate monocot-associate dicot-associate herb-associate shrub-associate tree-associate vine-associate epiphyte-associate lithophyte-associate rheophyte-associate hydrophyte-associate xerophyte-associate mesophyte-associate hygrophyte-associate sciophyte-associate heliophyte-associate pioneer-plant-associate climax-plant-associate disturbance-plant-associate successional-plant-associate ant-plant-protection-mutualism ant-plant-food-body-mutualism ant-plant-domatia-mutualism ant-plant-extrafloral-nectary-mutualism ant-plant-seed-dispersal-mutualism ant-plant-pollination-mutualism ant-plant-agriculture-mutualism ant-fungus-mutualism ant-bacteria-mutualism ant-virus-mutualism ant-nematode-mutualism ant-mite-mutualism ant-tick-mutualism ant-spider-mutualism ant-wasp-mutualism ant-fly-mutualism ant-beetle-mutualism ant-butterfly-mutualism ant-moth-mutualism ant-bird-mutualism ant-mammal-mutualism ant-reptile-mutualism ant-amphibian-mutualism ant-fish-mutualism ant-human-mutualism ant-domestic-animal-mutualism ant-pest-mutualism ant-crop-mutualism ant-forest-mutualism ant-savanna-mutualism ant-grassland-mutualism ant-desert-mutualism ant-wetland-mutualism ant-riverine-mutualism ant-riparian-mutualism ant-montane-mutualism ant-lowland-mutualism ant-coastal-mutualism ant-island-mutualism ant-continental-mutualism ant-tropical-mutualism ant-temperate-mutualism ant-subtropical-mutualism ant-boreal-mutualism ant-austral-mutualism ant-paleotropical-mutualism ant-neotropical-mutualism ant-afrotropical-mutualism ant-oriental-mutualism ant-palearctic-mutualism ant-nearctic-mutualism ant-holarctic-mutualism ant-pan-tropical-mutualism ant-cosmopolitan-mutualism ant-endemic-mutualism ant-widespread-mutualism ant-restricted-mutualism ant-rare-mutualism ant-common-mutualism ant-abundant-mutualism ant-scarce-mutualism ant-threatened-mutualism ant-invasive-mutualism ant-native-mutualism ant-exotic-mutualism ant-introduced-mutualism ant-naturalized-mutualism ant-feral-mutualism ant-urban-mutualism ant-rural-mutualism ant-agricultural-mutualism ant-natural-mutualism ant-anthropogenic-mutualism ant-pristine-mutualism ant-degraded-mutualism ant-restored-mutualism ant-conserved-mutualism ant-managed-mutualism ant-sustainable-mutualism ant-unsustainable-mutualism ant-resilient-mutualism ant-fragile-mutualism ant-robust-mutualism ant-vulnerable-mutualism ant-resistant-mutualism ant-susceptible-mutualism ant-tolerant-mutualism ant-intolerant-mutualism ant-flexible-mutualism ant-inflexible-mutualism ant-plastic-mutualism ant-canalized-mutualism ant-adaptive-mutualism ant-maladaptive-mutualism ant-optimal-mutualism ant-suboptimal-mutualism ant-efficient-mutualism ant-inefficient-mutualism ant-productive-mutualism ant-unproductive-mutualism ant-successful-mutualism ant-unsuccessful-mutualism ant-stable-mutualism ant-unstable-mutualism ant-persistent-mutualism ant-transient-mutualism ant-obligate-mutualism ant-facultative-mutualism ant-specific-mutualism ant-generalized-mutualism ant-exclusive-mutualism ant-non-exclusive-mutualism ant-symbiotic-mutualism ant-non-symbiotic-mutualism ant-trophic-mutualism ant-defensive-mutualism ant-dispersive-mutualism ant-pollination-mutualism ant-agriculture-mutualism ant-construction-mutualism ant-nutrient-mutualism ant-protection-mutualism ant-cleaning-mutualism ant-transport-mutualism ant-storage-mutualism ant-processing-mutualism ant-consumption-mutualism ant-production-mutualism ant-exchange-mutualism ant-reciprocity-mutualism ant-by-product-mutualism ant-investment-mutualism ant-harvest-mutualism ant-cultivation-mutualism ant-domestication-mutualism ant-selection-mutualism ant-breeding-mutualism ant-engineering-mutualism ant-modification-mutualism ant-creation-mutualism ant-destruction-mutualism ant-transformation-mutualism ant-conversion-mutualism ant-utilization-mutualism ant-exploitation-mutualism ant-cooperation-mutualism ant-competition-mutualism ant-conflict-mutualism ant-resolution-mutualism ant-negotiation-mutualism ant-communication-mutualism ant-signal-mutualism ant-cue-mutualism ant-information-mutualism ant-knowledge-mutualism ant-learning-mutualism ant-memory-mutualism ant-cognition-mutualism ant-intelligence-mutualism ant-decision-mutualism ant-choice-mutualism ant-preference-mutualism ant-motivation-mutualism ant-emotion-mutualism ant-stress-mutualism ant-welfare-mutualism ant-health-mutualism ant-disease-mutualism ant-immunity-mutualism ant-resistance-mutualism ant-tolerance-mutualism ant-recovery-mutualism ant-healing-mutualism ant-therapy-mutualism ant-medicine-mutualism ant-prevention-mutualism ant-security-mutualism ant-safety-mutualism ant-risk-mutualism ant-hazard-mutualism ant-threat-mutualism ant-danger-mutualism ant-damage-mutualism ant-injury-mutualism ant-death-mutualism ant-mortality-mutualism ant-survival-mutualism ant-longevity-mutualism ant-reproduction-mutualism ant-fertility-mutualism ant-fecundity-mutualism ant-sterility-mutualism ant-virginity-mutualism ant-mating-mutualism ant-copulation-mutualism ant-insemination-mutualism ant-fertilization-mutualism ant-egg-mutualism ant-laying-mutualism ant-incubation-mutualism ant-hatching-mutualism ant-emergence-mutualism ant-eclosion-mutualism ant-molting-mutualism ant-ecdysis-mutualism ant-metamorphosis-mutualism ant-development-mutualism ant-growth-mutualism ant-differentiation-mutualism ant-maturation-mutualism ant-aging-mutualism ant-senescence-mutualism ant-decomposition-mutualism ant-recycling-mutualism ant-energy-mutualism ant-carbon-mutualism ant-nitrogen-mutualism ant-phosphorus-mutualism ant-sulfur-mutualism ant-water-mutualism ant-oxygen-mutualism ant-hydrogen-mutualism ant-mineral-mutualism ant-vitamin-mutualism ant-hormone-mutualism ant-enzyme-mutualism ant-protein-mutualism ant-lipid-mutualism ant-carbohydrate-mutualism ant-nucleic-acid-mutualism ant-DNA-mutualism ant-RNA-mutualism ant-gene-mutualism ant-genome-mutualism ant-chromosome-mutualism ant-cell-mutualism ant-tissue-mutualism ant-organ-mutualism ant-system-mutualism ant-organism-mutualism ant-individual-mutualism ant-colony-mutualism ant-population-mutualism ant-species-mutualism ant-genus-mutualism ant-family-mutualism ant-order-mutualism ant-class-mutualism ant-phylum-mutualism ant-kingdom-mutualism ant-domain-mutualism ant-life-mutualism ant-earth-mutualism ant-universe-mutualism
Diplotaxis brachyptera
A small scarab beetle in the genus Diplotaxis, family Scarabaeidae, described by Patricia Vaurie in 1960. The species name 'brachyptera' (short-winged) suggests reduced flight capability. Records indicate presence in arid and semi-arid regions of the southwestern United States and northern Mexico. Like other Diplotaxis species, adults are likely nocturnal and may aggregate on vegetation.
Scarabaeidae Melolonthinae Diplotaxis Coleoptera beetle scarab nocturnal arid sand-dunes New-Mexico Chihuahua brachypterous aggregation-behavior
Elachiptera
frit flies
Elachiptera is a genus of small frit flies in the family Chloropidae, subfamily Oscinellinae. The genus contains approximately 81 valid species distributed across North America, Europe, and Asia, with some species extending into the Oriental Region. Several species are known agricultural pests of cereal crops, with larvae feeding internally within stems of wheat and maize. The genus includes both fully winged and wingless (brachypterous) species.
Chloropidae Oscinellinae cereal-pest stem-borer wheat-pest maize-pest invasive-species China Holarctic brachypterous wingless agricultural-pest grass-fly frit-fly
Eucnecosum brachypterum
Eucnecosum brachypterum is a small rove beetle (Staphylinidae: Omaliinae) with a Holarctic distribution, occurring across boreal and temperate regions of North America and Eurasia. The species was originally described in the genus Arpedium and later transferred to Eucnecosum, a generic placement confirmed by taxonomic revision. It is one of three holarctic Eucnecosum species recognized in North America. The species name refers to its short-winged (brachypterous) condition.
rove-beetle Omaliinae Holarctic boreal brachypterous taxonomic-revision
Eupelmus messene
Eupelmus messene is a tiny parasitoid wasp in the family Eupelmidae. Females are brachypterous (short-winged) and reproduce via thelytoky, producing only daughters. The species is a specialized idiobiont ectoparasitoid of the gall wasp Aulacidea hieracii, attacking both concealed and exposed larvae and pupae of its host. Females possess a remarkably thin, flexible, and mobile ovipositor capable of drilling through hard substrates, including polystyrene Petri dish walls in laboratory observations.
parasitoid gall-wasp-associate ovipositor-drilling thelytoky brachypterous biomimetics idiobiont ectoparasitoid
Gymnoscirtetes
Gymnoscirtetes is a genus of small (11–22 mm), slender, flightless grasshoppers endemic to the southeastern North American Coastal Plain. The genus was revised from two to six species based on male genital morphology and geographic data. Species are divided into two groups: the western morsei group (G. morsei, G. rex) and the eastern pusillus group (G. pusillus, G. pageae, G. wadeorum, G. georgiaensis). The genus is characterized by vestigial wings, linear body shape, and brownish-green or bronze coloration with a black lateral stripe.
endemic flightless grasshopper southeastern-USA brachypterous
Hormiinae
Hormiinae is a subfamily of Braconidae (Hymenoptera) comprising approximately 8 species in 4 genera and 2 tribes in Iran, with a cosmopolitan genus Hormius. Members include brachypterous and micropterous species with reduced wings. Some species are parasitoids of lepidopteran pupae.
Braconidae Hymenoptera parasitoid wing-reduction brachypterous micropterous Lepidoptera-parasitoid
Hysteropterini
Hysteropterini is a tribe of planthoppers within the family Issidae, subfamily Hysteropterinae. Members of this tribe are characterized by their compact body form and reduced wings, adaptations associated with a ground-dwelling or low-vegetation lifestyle. The tribe was established by Melichar in 1906 and contains multiple genera distributed primarily in the Palearctic region. These insects are part of the diverse fulgoromorph fauna but are less frequently encountered than their more volant relatives.
planthopper Issidae brachypterous ground-dwelling Palearctic grassland-insect
Leistus ferrugineus
Palearctic rusty basket-throat
Leistus ferrugineus is a species of ground beetle in the family Carabidae, native to the Palearctic region and introduced to northeastern North America. It inhabits mixed forests, where it occurs on moist gravelly ground, grassy areas, hedges, and open woodland. Adults are brachypterous, possessing reduced wings that limit flight capability. The species is part of the subfamily Nebriinae, a group of ground beetles often associated with cool, moist habitats.
Carabidae ground-beetle brachypterous Palearctic introduced-species forest-habitat Nebriinae
Litaneutria obscura
obscure ground mantis
Litaneutria obscura is a small ground-dwelling mantid native to the desert southwest of North America. Unlike most mantids that employ ambush predation from vegetation, this species is cursorial, actively running down prey on the ground. It is one of two Litaneutria species in the United States, with a more restricted range than its congener L. minor. The species is currently treated as a synonym of L. ocularis by some taxonomic authorities.
ground-mantis cursorial desert arid synonymized native predator cryptic brachypterous small-wings
Meconematinae
quiet-calling katydids, quiet crickets
Meconematinae is a subfamily of katydids (Tettigoniidae) commonly known as quiet-calling katydids or quiet crickets. The subfamily has a worldwide distribution with notable gaps in Antarctica and North America. It comprises three tribes: Meconematini (mostly Europe and Asia, with some African representation), Phisidini (pan-tropical), and Phlugidini (pan-tropical). Members are characterized by their ability to produce ultrasonic sound, with some species being miniature and brachypterous.
katydids bush-crickets ultrasonic brachypterous humid-habitat worldwide-distribution tropical subtropical mountainous soft-cuticle
Melanoliarus forcipatus
Melanoliarus forcipatus is a species of planthopper in the family Cixiidae, first described by Caldwell in 1947. Species in this genus are small to medium-sized fulgoroid insects characterized by their brachypterous (short-winged) condition in females and macropterous (long-winged) condition in males. Cixiidae planthoppers are generally associated with woody or herbaceous host plants, with nymphs often feeding on roots. The specific epithet "forcipatus" refers to a forceps-like structure, likely describing a morphological feature of the male genitalia or other appendage.
planthopper Cixiidae Auchenorrhyncha Fulgoromorpha brachypterous macropterous dimorphism Caldwell-1947
Melanoplus huroni
Huron Short-winged Locust, Huron Short-wing Grasshopper
Melanoplus huroni is a North American grasshopper in the spur-throated grasshopper group, first described by Blatchley in 1898. The species is characterized by its short-winged morphology, which distinguishes it from many congeneric species. It belongs to the large genus Melanoplus, which contains over 200 species across North America. The specific epithet refers to Lake Huron, reflecting its original discovery location in the Great Lakes region.
brachypterous spur-throated Great-Lakes-region Blatchley-1898
Melanoplus mirus
Weldon short-wing grasshopper
Melanoplus mirus is a species of spur-throated grasshopper in the family Acrididae. It is endemic to North America, with confirmed records from North Carolina. The species is characterized by reduced wing development, as indicated by its common name. It was described by Rehn and Hebard in 1916.
Melanoplus Acrididae Orthoptera grasshopper North-America North-Carolina brachypterous short-winged
Melanoplus oreophilus
Mount Hood Short-wing Grasshopper
Melanoplus oreophilus is a spur-throated grasshopper species endemic to the Mount Hood region of Oregon. The species is characterized by reduced wing development, a trait reflected in its common name. It inhabits high-elevation alpine environments and has a restricted geographic range.
brachypterous endemic alpine Cascade-Range Oregon spur-throated-grasshopper
Minitingis
Minitingis is a genus of lace bugs (Tingidae) comprising two described species: M. minusculus (Barbados, 1954) and M. elsae (West Indies, 1968). Originally synonymized with Zetekella by Drake & Ruhoff (1960), it was revalidated by Froeschner (1968) based on distinct morphological characters and zoogeographical separation. The genus is restricted to the West Indies, contrasting with Zetekella's Neotropical mainland distribution (Panama, Brazil, Ecuador). Both species are known only from brachypterous forms.
lace-bugs Tingidae Heteroptera West-Indies brachypterous revalidated-genus zoogeography
Nabis rufusculus
Short-winged Nabis
Nabis rufusculus is a species of damsel bug in the family Nabidae, commonly known as the Short-winged Nabis. It is a small predatory true bug found in North American agroecosystems, particularly alfalfa and soybean fields. The species has been studied for its phenology and abundance patterns alongside congeners N. americoferus and N. roseipennis. Like other damsel bugs, it is an economically important predator in agricultural systems.
predator agricultural-pest-control brachypterous nocturnal North-America
Nebria appalachia
southern Appalachian gazelle beetle
Nebria appalachia is a flightless ground beetle endemic to the southern Appalachian Mountains. Adults are brachypterous, with reduced wings preventing aerial dispersal. The species occupies a restricted geographic range limited to mid- to upland mixed forests in North Carolina and Tennessee.
brachypterous flightless endemic riparian southern-Appalachians
Nebria coloradensis
Colorado gazelle beetle
Nebria coloradensis is a flightless ground beetle endemic to the mountainous regions of Colorado. Adults possess reduced wings (brachypterous) and are active nocturnally. The species inhabits wet ground in alpine and subalpine environments.
brachypterous endemic montane nocturnal Carabidae Nebria
Nebria diversa
sea beach gazelle beetle
Nebria diversa is a coastal ground beetle endemic to sandy Pacific beaches of western North America. Adults are brachypterous (short-winged), restricting dispersal to coastal habitats. The species is nocturnal and occupies a narrow ecological niche beneath driftwood and debris on open sandy shorelines.
ground-beetle brachypterous coastal-endemic nocturnal Pacific-beach Carabidae carnivorous driftwood-habitat sand-beach western-North-America
Nebria gregaria
gregarious gazelle beetle
Nebria gregaria is a ground beetle in the family Carabidae, commonly known as the gregarious gazelle beetle. Adults are brachypterous (short-winged), nocturnal, and carnivorous. The species is restricted to riparian habitats and is found in Alaska, the Aleutian Islands, eastern Russia, and China.
Carabidae ground-beetle riparian brachypterous nocturnal carnivorous Alaska Aleutian-Islands Russia China
Nebria kincaidi
Kincaid's gazelle beetle
Nebria kincaidi is a ground beetle species in the family Carabidae, first described by Schwarz in 1900. Adults are brachypterous (short-winged), nocturnal, and carnivorous. The species comprises two recognized subspecies with distinct geographic distributions in the Pacific Northwest and Alaska.
ground-beetle brachypterous nocturnal carnivorous Pacific-Northwest subspecies Nebriinae
Nebria lituyae
Lituya's gazelle beetle
Nebria lituyae, known as Lituya's gazelle beetle, is a ground beetle species in the subfamily Nebriinae. It is endemic to a restricted alpine region spanning Alaska and British Columbia. The species inhabits moist ground on hill summits at high elevations. Adults are brachypterous (short-winged), nocturnal, and carnivorous.
endemic alpine brachypterous nocturnal carnivorous
Nebria ovipennis
oval gazelle beetle
Nebria ovipennis is a ground beetle endemic to California and Nevada, where it inhabits meadows and snowfield margins in upland and mountainous areas. Adults are brachypterous (short-winged), nocturnal, and carnivorous. The species belongs to the subfamily Nebriinae, a group often associated with cool, moist habitats at higher elevations.
Carabidae Nebriinae ground-beetle endemic brachypterous montane California Nevada nocturnal predator