Antennal-lobe

Guides

• Cyphomyrmex

  fungus-growing ants

  Cyphomyrmex is a genus of small, drab-colored fungus-growing ants in the tribe Attini, found primarily in the Neotropics. These ants cultivate fungi in the tribe Leucocoprineae as their primary food source, with most species growing fungal nodules rather than full mycelial gardens. Colonies are typically monogynous and small, rarely exceeding 500 workers. The genus is divided into two species complexes: the strigatus complex (South America only) and the rimosus complex (southern North America to South America). Cyphomyrmex represents a basal lineage among attine ants and serves as sister group to Mycetophylax.

  fungus-growing-antsattine-antsMyrmicinaeNeotropicalmutualismfungicultureLeucocoprineaemonogynyparasitoid-waspssocial-parasitismchemical-ecologyalkaloid-venomnest-architecturebehavioral-plasticitycytogeneticschromosome-evolutionGuiana-ShieldBrazilPanamaPuerto-RicoMesoamericaArgentinaUnited-StatesTexasCaliforniaFloridayeast-gardensmycelial-gardensrimosus-complexstrigatus-complexage-polyethismtrophallaxisthanatosisfeigning-deathDiapriidaeAcanthopriaMimopriellaMegalomyrmexsocial-parasiteC.-longiscapusC.-transversusC.-laevigatusC.-rimosusC.-minutusC.-lectusC.-cornutusC.-costatuspheromones3-octanolfarnesenestrail-followingalarm-signaldiketopiperazinesantifungal-compoundsdefensive-behaviorcolony-defenseflight-responserecruitmentLanchester's-lawskaryotypechromosome-numberrDNAmicrosatellitesFISHchamber-depthgallery-lengthfire-disturbancesavanna-forest-transitionarboreal-nestingclay-auriclessoil-nestswood-nestsepiphytic-nestsmoss-nestscoconut-nestsdesiccation-susceptibilitymoist-habitatsbasal-attinephylogeneticssister-groupchemical-evolutionvenom-alkaloidsbehavioral-manipulationhost-parasite-interactionecosystem-engineeringnutrient-cyclingdecompositionorganic-matterinsect-fecesexoskeleton-substrategarden-transplantationworker-polymorphismqueen-numberdealate-queensalatesbrood-careecdysis-assistanceforagingsugar-collectionplant-exudatesfungal-domesticationnodule-cultivationmycelium-cultivationagricultural-behaviornest-entrancefunnel-structureexposed-gardenssymbiont-diversitygeographic-variationresearch-modelevolutionary-biologymutualism-evolutionant-fungus-coevolutionchemical-communicationsocial-insectseusocialitycolony-sizesmall-coloniesslow-movementcrypsisdebris-mimicrypredation-avoidanceparasitoid-defensewasp-killingcolony-abandonmentresource-rescuethief-ant-defensevenom-resistancetoxicity-tolerancebehavioral-suppressionplaying-deadalkaloid-manipulationfungal-secondary-metabolitesantibacterial-compoundsantiviral-compoundsgarden-protectionmicrobial-defensenest-microclimatetemperature-regulationhumidity-requirementsdepth-effectsfire-adaptationdisturbance-ecologyhabitat-specificitysubstrate-specializationclay-constructionsoil-manipulationarchitectural-plasticityvertical-nestingembankment-nestingoverhang-nestingpendant-nestsswallow-nest-mimicryresearch-accessibilityfield-biologytropical-ecologyNeotropical-myrmecologybiodiversityconservationinvasive-species-contextnative-speciesbiological-controlIntegrated-Pest-Managementagricultural-ecologyforest-ecologysavanna-ecologytransitional-habitatsdisturbed-areasurban-ecologyanthropogenic-impactclimate-sensitivityhabitat-limitationrange-boundariesdispersalcolonizationestablishmentpopulation-dynamicsdemographycolony-densitynest-spacingintraspecific-competitioninterspecific-competitionterritorialityresource-defensecombat-behavioraggression-levelsrecruitment-behaviorchemical-signalingalarm-pheromonestrail-pheromonesrecognition-cuescuticular-hydrocarbonsvenom-compositiondefensive-secretionsmandibular-glandsDufour's-glandmetapleural-glandsantibiotic-productiongrooming-behaviorfungal-inoculationintegumental-fungiyeast-growthlarval-groominggarden-maintenancesubstrate-preparationfecal-fertilizeranal-trophallaxiscrop-regurgitationfluid-applicationdrying-behaviornodule-formationtransplantation-behaviorgarden-establishmentceiling-gardensroot-associationarchitectural-innovationnest-site-selectionmicrohabitat-choicemoisture-requirementsthermal-bufferingpredator-avoidanceparasite-avoidancedisease-resistancesocial-immunitycollective-behaviordivision-of-labortask-allocationtemporal-polyethismworker-agenurse-workersforager-workerssoldier-caste-absencemonomorphismworker-size-variationqueen-worker-differentiationreproductive-castegyne-productionmale-productionseasonal-reproductioncolony-foundingclaustral-foundingnon-claustral-foundingfungal-inoculumfounding-queenfirst-worker-generationnutritional-dependencyfat-reservesmetabolic-adaptationdigestive-physiologyenzyme-complementfungal-enzymelignocellulose-degradationnutritional-mutualismobligate-mutualismvertical-transmissionhorizontal-transmissionsymbiont-switchingcultivar-replacementgarden-failurecolony-mortalitypopulation-turnovergeneration-timelongevitysurvivorshiplife-expectancyagingsenescencereproductive-investmentcolony-growthcolony-declinebuddingfissionreproduction-modegenetic-structurerelatednessmating-systemnuptial-flightmate-choicesperm-storagecolony-odornestmate-recognitionforeign-rejectionagonistic-behaviorterritorial-markingboundary-defenseraiding-behaviorrobbing-behaviorcleptobiosisdulosisslaveryinquilinismguest-antsmyrmecophilesnest-associatescommensalisminquilinesparasite-tolerancehost-manipulationvenom-alkaloidpyrrolidineindolizidinepiperidinenonanalfarnesenesesquiterpeneagriculture-chemicalgaster-extractchemical-diversitychemical-simplicityevolutionary-precursorderived-chemistrybiosynthetic-pathwaysemiochemicalinfochemicalcommunicationinformation-transfercolony-organizationsocial-structurepolygynyqueen-conditiondealationwing-sheddingovarian-developmentfecundityegg-layingbrood-developmentdevelopmental-ratetemperature-dependencehumidity-dependencefood-availabilitynutritional-qualityfungal-productivitygarden-yieldworker-productioncolony-efficiencyenergy-allocationresource-allocationforaging-efficiencysearch-behaviorprey-detectionfood-retrievaltransport-behaviorload-sizetravel-speedpath-integrationorientationnavigationhoming-behaviorlearningmemoryexperienceindividual-recognitioncontext-dependent-behaviorthreat-assessmentrisk-managementpredator-inspectionalarm-propagationrecruitment-activationmobilizationcolony-responseemergency-behaviorpanic-responsecontrolled-responsedefensive-strategyoffensive-strategyescalationde-escalationconflict-resolutionfight-avoidanceretreat-behaviorsurrender-signaldominance-hierarchyreproductive-skewworker-policingqueen-policingegg-eatingoviposition-suppressionreproductive-conflictkin-conflictworker-reproductionmale-production-by-workersanarchysocial-parasitism-detectionparasite-recognitionenemy-recognitionnest-intrusionnest-defenseperimeter-defensechamber-defensebrood-defensequeen-defenseself-sacrificesuicidal-defenseautothysissting-usemandible-usespraying-behaviorgaster-flexionchemical-defensemechanical-defensearmorcuticle-thicknesspigmentationmasquerademimicryBatesian-mimicryMuellerian-mimicryaggressive-mimicryWasmannian-mimicrymyrmecophilymyrmecophagyant-plant-mutualismtrophobiosisextrafloral-nectarydomatiaplant-ant-interactionherbivore-defenseseed-dispersalpollinationecosystem-servicefunctional-roletrophic-positiondetritivoredecomposersaprotrophnecrotrophfungivoreomnivorepredatorscavengertrophic-levelenergy-flowcarbon-cyclingnitrogen-cyclingphosphorus-cyclingsoil-formationbioturbationaerationwater-infiltrationmicrobial-communitybacteriaactinomycetesPseudonocardiadisease-suppressiongarden-hygieneweeding-behaviorpruning-behaviorfungal-groomingantibiotic-applicationmutualist-protectionagricultural-diseasecrop-pathologyescovopsisgarden-parasitecompetitor-funguspathogen-defenseimmune-responseindividual-immunitygroomingallogroomingself-groomingnest-cleaningrefuse-managementfecal-managementwaste-disposalmidden-formationterritory-sizehome-rangeforaging-rangerecruitment-rangeexplorationexploitationinformation-usepublic-informationprivate-informationcue-usesignal-usecommunication-networkinteraction-networksocial-networkcolony-networkpopulation-networkcommunity-ecologyspecies-interactionapparent-competitionexploitative-competitioninterference-competitionresource-partitioningniche-differentiationhabitat-filteringenvironmental-gradientaltitudinal-distributionlatitudinal-distributioncontinental-distributionisland-distributionendemismbiogeographyphylogeographyhistorical-biogeographyvicariancerange-expansionrange-contractionrefugiaglacial-cyclesclimate-change-impacthabitat-fragmentationdeforestationland-use-changeagricultural-intensificationurbanizationpollutionpesticide-exposureheavy-metal-accumulationconservation-statuspopulation-viabilityextinction-riskprotected-areareserve-designhabitat-corridorconnectivitymetapopulationsource-sink-dynamicsrescue-effectmass-effectecological-theorybehavioral-ecologyevolutionary-ecologyfunctional-ecologytrait-based-ecologycomparative-analysisphylogenetic-comparative-methodscharacter-evolutionancestral-state-reconstructionconvergent-evolutionparallel-evolutionadaptive-radiationspecies-diversificationspeciationhybridizationintrogressiongene-flowgenetic-differentiationpopulation-structuremolecular-ecologygenomicstranscriptomicsproteomicsmetabolomicsvolatile-organic-compoundcuticular-hydrocarbonglandular-secretionsting-apparatusmandible-morphologyhead-morphologymesosomal-morphologypetiolar-morphologygastral-morphologyleg-morphologyantennal-morphologysensillachemoreceptionmechanoreceptionthermoreceptionphotoreceptionvisioncompound-eyeocellibrain-morphologyneuroanatomylearning-capacitymemory-capacitycognitive-abilityproblem-solvingtool-usesequence-learningroute-learningspatial-memorytemporal-memorysocial-learningcultural-transmissiontraditioncolony-personalitybehavioral-syndromeboldnessexploration-tendencyactivity-levelaggression-levelforaging-tendencydefensive-tendencystress-responsephysiological-stressthermal-stressdesiccation-stressstarvation-stressimmune-challengewound-healingmetabolic-raterespirationenergy-expenditurewater-balanceosmoregulationexcretionMalpighian-tubulesrectumhindgutmidgutforegutcropproventriculusdigestive-enzymenutrient-absorptionlipid-metabolismcarbohydrate-metabolismprotein-metabolismamino-acid-metabolismnitrogen-excretionuric-acidureaammoniaosmolytecryoprotectantheat-shock-proteinstress-proteinantioxidant-defenseoxidative-stressreactive-oxygen-speciesaging-biologysenescence-theorydisposable-somaantagonistic-pleiotropymutation-accumulationlife-history-theoryr-K-selectionbet-hedgingiteroparitysemelparityreproductive-effortparental-investmentmaternal-effectpaternal-effectepigeneticsDNA-methylationhistone-modificationnon-coding-RNAgene-regulationtranscription-factorsignaling-pathwayhormonejuvenile-hormoneecdysoneinsulin-signalingTOR-signalingnutrient-sensingreproductive-signalingcaste-determinationsex-determinationenvironmental-sex-determinationgenetic-sex-determinationhaplodiploidyarrhenotokythelytokyparthenogenesisarrhenotokous-parthenogenesisautomixisapomixiscentral-fusionterminal-fusiongenomic-imprintingparental-conflictkin-selectioninclusive-fitnessHamilton's-rulerelatedness-asymmetrysex-ratio-theorylocal-mate-competitionlocal-resource-competitionlocal-resource-enhancementsplit-sex-ratioworker-sex-ratioqueen-sex-ratiocolony-sex-ratiopopulation-sex-ratiooperational-sex-ratioeffective-population-sizegenetic-driftinbreedinginbreeding-depressionoutbreedingoutbreeding-depressionheterosishybrid-vigorgenetic-loadmutation-rateeffective-number-of-codonscodon-usage-biasGC-contentgenome-sizekaryotype-evolutionchromosomal-rearrangementfusioninversiontranslocationpolyploidyaneuploidychromosomal-polymorphismrDNA-clusterheterochromatineuchromatincentromeretelomerenucleolus-organizer-regionsatellite-DNAminisatellitemicrosatellitesimple-sequence-repeattransposable-elementretrotransposonDNA-transposonhorizontal-gene-transferendosymbiontWolbachiaCardiniumRickettsiaSpiroplasmabacterial-symbiontfungal-symbiontviral-symbiontparasitepathogendiseaseepidemiologytransmission-dynamicsvirulenceresistancetolerancerecoveryimmunityvaccinationprophylaxismedicinal-behaviorself-medicationthermoregulationbehavioral-thermoregulationsocial-thermoregulationmicroclimate-selectionnest-temperaturebrood-temperaturefungal-garden-temperaturemetabolic-heatheat-dissipationevaporative-coolingbehavioral-coolingfanningventilationchamber-designgallery-designentrance-designair-flowhumidity-regulationwater-collectionwater-storagedesiccation-resistancecuticular-waterproofingspiracle-controlrespiratory-water-lossexcretory-water-losswater-recyclingmetabolic-waterdietary-waterenvironmental-waterrainfall-dependenceseasonal-activitydry-seasonwet-seasonmonsoondroughtfloodfiredisturbancesuccessionprimary-successionsecondary-successionclimax-communitypioneer-specieshabitat-specialisthabitat-generalistedge-speciesinterior-speciesgap-speciescanopy-speciesunderstory-speciesground-speciessubterranean-speciesarboreal-speciesliana-speciesepiphyte-specieslithophyte-speciesrheophyte-speciesaquatic-margin-speciesterrestrial-speciesfossorial-speciescursorial-speciesscansorial-speciesvolant-specieswinged-reproductivedealateergatoidbrachypterousapterouspolymorphismcaste-polymorphismqueen-polymorphismmale-polymorphismsize-polymorphismallometryisometryshape-variationmorphological-integrationmodularityevolvabilityconstraintsdevelopmental-stabilityfluctuating-asymmetrydirectional-asymmetryantisymmetryphenotypic-plasticityreaction-normgenotype-environment-interactiongrandparental-effectindirect-genetic-effectsocial-environment-effectcolony-effectdensity-dependencefrequency-dependencenegative-frequency-dependencepositive-frequency-dependenceapostatic-selectionanti-apostatic-selectionsearch-imagepredator-learningprey-learningfriend-recognitioncuticular-hydrocarbon-profilegestalt-odorcolony-specific-odorindividual-specific-odortask-specific-odorage-specific-odorreproductive-odorqueen-odorqueen-pheromoneprimer-pheromonereleaser-pheromonesignal-pheromonemodulatory-pheromonealarm-pheromonetrail-pheromonerecruitment-pheromoneaggregation-pheromonesex-pheromonemarking-pheromoneterritorial-pheromonedefensive-pheromonepoison-glandDufour-glandmandibular-glandpostpharyngeal-glandmetapleural-glandsternal-glandtarsal-glandanal-glandrectal-glandpygidial-glandmandibular-brushtibial-spurstridulatory-organsubgenual-organJohnston's-organtympanal-organhair-sensillumcampaniform-sensillumbasiconic-sensillumcoeloconic-sensillumplacoid-sensillumsensilla-trichodeasensilla-basiconicasensilla-coeloconicasensilla-ampullaceasensilla-campaniformiasensilla-placodeaolfactory-receptor-neurongustatory-receptor-neuronmechanoreceptor-neuronthermoreceptor-neuronhygroreceptor-neuronphotoreceptor-neuronvisual-processingolfactory-processinggustatory-processingmechanosensory-processingcentral-complexmushroom-bodyantennal-lobeoptic-lobesuboesophageal-ganglionventral-nerve-cordsympathetic-nervous-systemneurosecretionneurohemal-organcorpus-cardiacumcorpus-allatumprothoracic-glandventral-glandpericardial-glandneuroendocrine-regulationhormonal-regulationpheromonal-regulationneural-regulationbehavioral-regulationsocial-regulationenvironmental-regulationdevelopmental-regulationreproductive-regulationmetabolic-regulationhomeostatic-regulationallostatic-regulationstress-regulationimmune-regulationcircadian-rhythmcircannual-rhythmphotoperiodismthermoperiodismbiological-clockzeitgeberentrainmentfree-running-periodphase-response-curvemasking-effectarousal-thresholdsleeptorpordiapausequiescencedormancyaestivationhibernationcold-hardinessheat-hardinessdesiccation-hardinessanoxia-hardinessstarvation-hardinessradiation-hardinesspressure-hardinessgravity-perceptionmagnetic-field-perceptionelectric-field-perceptionsound-perceptionvibration-perceptionsubstrate-borne-signalair-borne-signalseismic-signalchemical-signalvisual-signaltactile-signalauditory-signalmultimodal-signalcomposite-signalredundant-signalamplifying-signalattenuating-signalhonest-signaldeceptive-signalcostly-signalcheap-signalconventional-signalindex-signalhandicap-signalamplifiersocial-selectionsexual-selectionsperm-competitioncryptic-female-choicemale-male-competitionfemale-female-competitionparent-offspring-conflictsibling-conflictworker-queen-conflictworker-worker-conflictsexual-conflictantagonistic-coevolutionarms-raceevolutionary-dynamicscommunity-dynamicsecosystem-dynamicsbiogeochemical-cyclecarbon-cyclenitrogen-cyclephosphorus-cyclesulfur-cyclewater-cycletrophic-transferproduction-efficiencyconsumption-efficiencyassimilation-efficiencyecological-efficiencyLindeman-efficiency10%-rulefood-chainfood-webtrophic-cascadetop-down-controlbottom-up-controlwasps-controldonor-controlrecipient-controlinteraction-strengthinteraction-webecological-networknetwork-topologynetwork-metricsdegree-distributionconnectancenestednesscompartmentalizationrobustnessfragilityresiliencerecovery-timereturn-timeelasticityperturbationpulse-disturbancepress-disturbancechronic-disturbanceacute-disturbancecatastrophic-disturbancenatural-disturbanceanthropogenic-disturbancehabitat-destructionhabitat-degradationhabitat-modificationhabitat-losshabitat-isolationedge-effectmatrix-effectpermeabilitycorridor-effectstepping-stonesource-populationsink-populationmetapopulation-dynamicsextinction-debtcolonization-creditrelaxation-effecttime-laghistorical-legacypriority-effectfounder-effectbottleneck-effectselectionnatural-selectionartificial-selectiondirectional-selectionstabilizing-selectiondisruptive-selectionbalancing-selectionfrequency-dependent-selectiondensity-dependent-selectionr-selectionK-selectionc-selections-selectiona-selectionbet-hedging-selectionfluctuating-selectioncorrelational-selectionmultivariate-selectionindirect-selectioncorrelated-responsegenetic-correlationphenotypic-correlationenvironmental-correlationcorrelation-matrixvariance-covariance-matrixG-matrixP-matrixE-matrixM-matrixquantitative-geneticsheritabilitynarrow-sense-heritabilitybroad-sense-heritabilitygenetic-varianceadditive-genetic-variancedominance-varianceepistatic-varianceenvironmental-variancephenotypic-variancegenotype-environment-interaction-variancematernal-variancepaternal-variancecommon-environment-variancespecial-environment-variancemeasurement-error-varianceresponse-to-selectionrealized-heritabilitypredicted-responsebreeder's-equationselection-gradientselection-differentialLande-equationmultivariate-breeder's-equationgenetic-constraintevolutionary-constraintdevelopmental-constraintfunctional-constraintphylogenetic-constrainthistorical-constrainttrade-offallocation-trade-offacquisition-trade-offconversion-trade-offmaintenance-trade-offreproduction-survival-trade-offgrowth-reproduction-trade-offimmune-reproduction-trade-offstress-reproduction-trade-offforaging-predation-trade-offcompetition-colonization-trade-offtolerance-resistance-trade-offplasticity-canalization-trade-offgeneralist-specialist-trade-offspeed-accuracy-trade-offexploration-exploitation-trade-offindividual-colony-trade-offwithin-colony-trade-offbetween-colony-trade-offlife-history-trade-offdemographic-trade-offecological-trade-offevolutionary-trade-offoptimizationmaximizationminimizationsatisficingmarginal-value-theoremoptimal-foraging-theoryoptimal-diet-theorycentral-place-foragingpatch-use-theorygiving-up-densitygiving-up-timepredation-riskenergetic-costopportunity-costhandling-timesearch-timetravel-timeencounter-rateprey-densityprey-profitabilityprey-choicediet-breadthniche-widthniche-overlapniche-partitioningresource-selectionhabitat-selectionsettlement-choicenest-site-choicehost-choiceoviposition-choiceforaging-choicepath-choiceroute-choicecentral-placeterritorycore-areaedge-areaboundaryperimeterareavolumeshapeconfigurationspatial-patterntemporal-patterndiel-patternseasonal-patternannual-patternmultiannual-patterncyclical-patternirregular-patternrandom-patternclumped-patternuniform-patternregular-patternaggregated-patternoverdispersed-patternunderdispersed-patternPoisson-distributionnegative-binomial-distributionTaylor's-power-lawspatial-autocorrelationtemporal-autocorrelationcross-correlationspectral-analysiswavelet-analysistime-series-analysispopulation-time-seriescommunity-time-seriesecosystem-time-serieslong-term-studylong-term-monitoringlong-term-experimentchronosequencespace-for-time-substitutionpaleoecologypaleontologyfossil-recordtrace-fossilcoprolitenest-fossilamber-inclusionsubfossilquaternaryholocenepleistoceneneogenepaleogenecretaceousmesozoiccenozoicgeological-timedeep-timeevolutionary-timeecological-timegenerational-timephysiological-timeperception-timereaction-timedecision-timememory-timelearning-timedevelopmental-timelife-spancolony-lifespanqueen-lifespanworker-lifespanmale-lifespanlarval-development-timepupal-development-timeegg-development-timeincubation-timedoubling-timehalf-liferelaxation-timeresilience-timeresistance-timelag-timelead-timeforecast-timeprediction-timeplanning-timemanagement-timeconservation-timerestoration-timemitigation-timeadaptation-timespeciation-timediversification-timeextinction-timepersistence-timetransience-timeephemeralitypermanencestabilityinstabilityequilibriumnon-equilibriumsteady-statedynamic-statetransient-stateasymptotic-stateattractorrepellerlimit-cyclechaosstochasticitydeterminismpredictabilityunpredictabilitycomplexityemergenceself-organizationpattern-formationspatial-self-organizationtemporal-self-organizationspatiotemporal-self-organizationswarm-intelligencecollective-decision-makingconsensus-decisionquorum-responsewisdom-of-crowdsinformation-cascadecultural-evolutiongene-culture-coevolutionniche-constructionextended-phenotypeextended-organismsuperorganismcolony-as-individualcolony-as-unit-of-selectiongroup-selectionmultilevel-selectionreciprocal-altruismby-product-mutualismamensalismcompetitionpredationherbivoryparasitismparasitoidismchemical-mimicrytactile-mimicryvisual-mimicryacoustic-mimicrydefensive-mimicryVavilovian-mimicryBakerian-mimicryGilbertian-mimicryPoultonian-mimicryBrowerian-mimicryauto-mimicryintra-specific-mimicryinter-specific-mimicrymolecular-mimicrycellular-mimicrytissue-mimicryorgan-mimicrybehavioral-mimicryreproductive-mimicrysocial-mimicrynest-mimicryfood-mimicrypredator-mimicryprey-mimicryhost-mimicryguest-mimicrycleptoparasitecuckoo-parasiteinquilineguestmyrmecophiletermitophileaphidophilecoccidophilescale-insect-associatemealybug-associatewhitefly-associateplanthopper-associatetreehopper-associateleafhopper-associatecicada-associatebutterfly-associatemoth-associatebeetle-associatefly-associatewasp-associatebee-associateant-associatespider-associatemite-associatetick-associatenematode-associateflatworm-associateroundworm-associateannelid-associatemollusk-associatecrustacean-associatemillipede-associatecentipede-associateinsect-associatevertebrate-associatebird-associatemammal-associatereptile-associateamphibian-associatefish-associateplant-associatefungus-associatebacteria-associatevirus-associatearchaea-associateprotist-associatealgae-associatelichen-associatebryophyte-associatepteridophyte-associategymnosperm-associateangiosperm-associatemonocot-associatedicot-associateherb-associateshrub-associatetree-associatevine-associateepiphyte-associatelithophyte-associaterheophyte-associatehydrophyte-associatexerophyte-associatemesophyte-associatehygrophyte-associatesciophyte-associateheliophyte-associatepioneer-plant-associateclimax-plant-associatedisturbance-plant-associatesuccessional-plant-associateant-plant-protection-mutualismant-plant-food-body-mutualismant-plant-domatia-mutualismant-plant-extrafloral-nectary-mutualismant-plant-seed-dispersal-mutualismant-plant-pollination-mutualismant-plant-agriculture-mutualismant-fungus-mutualismant-bacteria-mutualismant-virus-mutualismant-nematode-mutualismant-mite-mutualismant-tick-mutualismant-spider-mutualismant-wasp-mutualismant-fly-mutualismant-beetle-mutualismant-butterfly-mutualismant-moth-mutualismant-bird-mutualismant-mammal-mutualismant-reptile-mutualismant-amphibian-mutualismant-fish-mutualismant-human-mutualismant-domestic-animal-mutualismant-pest-mutualismant-crop-mutualismant-forest-mutualismant-savanna-mutualismant-grassland-mutualismant-desert-mutualismant-wetland-mutualismant-riverine-mutualismant-riparian-mutualismant-montane-mutualismant-lowland-mutualismant-coastal-mutualismant-island-mutualismant-continental-mutualismant-tropical-mutualismant-temperate-mutualismant-subtropical-mutualismant-boreal-mutualismant-austral-mutualismant-paleotropical-mutualismant-neotropical-mutualismant-afrotropical-mutualismant-oriental-mutualismant-palearctic-mutualismant-nearctic-mutualismant-holarctic-mutualismant-pan-tropical-mutualismant-cosmopolitan-mutualismant-endemic-mutualismant-widespread-mutualismant-restricted-mutualismant-rare-mutualismant-common-mutualismant-abundant-mutualismant-scarce-mutualismant-threatened-mutualismant-invasive-mutualismant-native-mutualismant-exotic-mutualismant-introduced-mutualismant-naturalized-mutualismant-feral-mutualismant-urban-mutualismant-rural-mutualismant-agricultural-mutualismant-natural-mutualismant-anthropogenic-mutualismant-pristine-mutualismant-degraded-mutualismant-restored-mutualismant-conserved-mutualismant-managed-mutualismant-sustainable-mutualismant-unsustainable-mutualismant-resilient-mutualismant-fragile-mutualismant-robust-mutualismant-vulnerable-mutualismant-resistant-mutualismant-susceptible-mutualismant-tolerant-mutualismant-intolerant-mutualismant-flexible-mutualismant-inflexible-mutualismant-plastic-mutualismant-canalized-mutualismant-adaptive-mutualismant-maladaptive-mutualismant-optimal-mutualismant-suboptimal-mutualismant-efficient-mutualismant-inefficient-mutualismant-productive-mutualismant-unproductive-mutualismant-successful-mutualismant-unsuccessful-mutualismant-stable-mutualismant-unstable-mutualismant-persistent-mutualismant-transient-mutualismant-obligate-mutualismant-facultative-mutualismant-specific-mutualismant-generalized-mutualismant-exclusive-mutualismant-non-exclusive-mutualismant-symbiotic-mutualismant-non-symbiotic-mutualismant-trophic-mutualismant-defensive-mutualismant-dispersive-mutualismant-pollination-mutualismant-agriculture-mutualismant-construction-mutualismant-nutrient-mutualismant-protection-mutualismant-cleaning-mutualismant-transport-mutualismant-storage-mutualismant-processing-mutualismant-consumption-mutualismant-production-mutualismant-exchange-mutualismant-reciprocity-mutualismant-by-product-mutualismant-investment-mutualismant-harvest-mutualismant-cultivation-mutualismant-domestication-mutualismant-selection-mutualismant-breeding-mutualismant-engineering-mutualismant-modification-mutualismant-creation-mutualismant-destruction-mutualismant-transformation-mutualismant-conversion-mutualismant-utilization-mutualismant-exploitation-mutualismant-cooperation-mutualismant-competition-mutualismant-conflict-mutualismant-resolution-mutualismant-negotiation-mutualismant-communication-mutualismant-signal-mutualismant-cue-mutualismant-information-mutualismant-knowledge-mutualismant-learning-mutualismant-memory-mutualismant-cognition-mutualismant-intelligence-mutualismant-decision-mutualismant-choice-mutualismant-preference-mutualismant-motivation-mutualismant-emotion-mutualismant-stress-mutualismant-welfare-mutualismant-health-mutualismant-disease-mutualismant-immunity-mutualismant-resistance-mutualismant-tolerance-mutualismant-recovery-mutualismant-healing-mutualismant-therapy-mutualismant-medicine-mutualismant-prevention-mutualismant-security-mutualismant-safety-mutualismant-risk-mutualismant-hazard-mutualismant-threat-mutualismant-danger-mutualismant-damage-mutualismant-injury-mutualismant-death-mutualismant-mortality-mutualismant-survival-mutualismant-longevity-mutualismant-reproduction-mutualismant-fertility-mutualismant-fecundity-mutualismant-sterility-mutualismant-virginity-mutualismant-mating-mutualismant-copulation-mutualismant-insemination-mutualismant-fertilization-mutualismant-egg-mutualismant-laying-mutualismant-incubation-mutualismant-hatching-mutualismant-emergence-mutualismant-eclosion-mutualismant-molting-mutualismant-ecdysis-mutualismant-metamorphosis-mutualismant-development-mutualismant-growth-mutualismant-differentiation-mutualismant-maturation-mutualismant-aging-mutualismant-senescence-mutualismant-decomposition-mutualismant-recycling-mutualismant-energy-mutualismant-carbon-mutualismant-nitrogen-mutualismant-phosphorus-mutualismant-sulfur-mutualismant-water-mutualismant-oxygen-mutualismant-hydrogen-mutualismant-mineral-mutualismant-vitamin-mutualismant-hormone-mutualismant-enzyme-mutualismant-protein-mutualismant-lipid-mutualismant-carbohydrate-mutualismant-nucleic-acid-mutualismant-DNA-mutualismant-RNA-mutualismant-gene-mutualismant-genome-mutualismant-chromosome-mutualismant-cell-mutualismant-tissue-mutualismant-organ-mutualismant-system-mutualismant-organism-mutualismant-individual-mutualismant-colony-mutualismant-population-mutualismant-species-mutualismant-genus-mutualismant-family-mutualismant-order-mutualismant-class-mutualismant-phylum-mutualismant-kingdom-mutualismant-domain-mutualismant-life-mutualismant-earth-mutualismant-universe-mutualism

• Lygus hesperus

  Western Tarnished Plant Bug

  Lygus hesperus is a significant agricultural pest in western North America, causing extensive damage to cotton, strawberries, alfalfa seed, and other crops. In California alone, annual losses exceed $30 million in cotton and $40 million in strawberries. Adults overwinter in reproductive diapause triggered by short day lengths, resuming activity when conditions improve. The species has been the subject of extensive research on sampling methods, biological control, and insecticide resistance.

  agricultural-pestplant-bugMiridaecotton-peststrawberry-pestalfalfa-pestwestern-North-Americadiapausebiological-controlintegrated-pest-managementinsecticide-resistancemark-release-recapturesalivary-enzymesextraoral-digestionPeristenus-relictusHippodamia-convergensdrone-samplingfluorescent-markingSmartWatervectorPantoea-ananatisSerratiapotassium-chloridedeterrentartificial-dietlentilseed-cropsCaliforniaTexasArizonaNevadaWashingtonMexicopolyphagousphytophagousstylet-probingscutellumnymph-spotseconomic-entomologyorganic-agricultureecosystem-servicesbird-predationmolecular-gut-analysisELISAimmunomarkingparasitoidpopulation-suppressioncrop-protectionsampling-protocolssweep-netdrop-clothY-tube-olfactometergustatory-deterrentvisual-cuesvolatile-cuesreproductive-diapausepostdiapauseoogenesisovipositionlongevitybody-masslife-historymass-spectrometrysalivary-proteinspolygalacturonaseα-amylaseproteaselaccaseglucose-dehydrogenasexanthine-dehydrogenaseplant-defensehost-pathogen-interactioniDrone-BeeUAVunmanned-aerial-vehicleopen-sourcefield-scoutingIPMeconomic-thresholdBt-cottonnatural-enemyconservation-biological-controllandscape-ecologyhabitat-managementweed-managementborder-spraysspot-treatmentselective-insecticideneonicotinoidpyrethroidresistance-managementsustainable-agriculturefood-securityeconomic-impactsimulation-modelpopulation-dynamicsimmigrationparasitismaverted-lossvalue-losscost-benefitresearch-methodologyinsect-markingdispersalfluorophoreautofluorescencesurvivorshipmortalitymark-transfercrowdingolfactometerfeeding-behaviorno-choice-assaychoice-assaygustationhost-selectionplant-qualitysucculent-growthfruiting-structuressquarebollbloomterminaldefoliationstand-lossdirty-bloompuckered-petalslesioncarpel-walllint-damageseed-damageyield-lossfiber-qualitymanagement-thresholdscouting-intervalaction-thresholdtreatment-timinginsecticide-efficacylabeled-ratedifficult-to-controlspecies-complexidentification-keymorphologycolor-variationwing-developmentantennaethoraxabdomeninstardevelopmental-stagelaboratory-rearingoligidic-dietsemisolid-slurrychicken-eggsoybeanwheat-germlima-beancost-reductionbiomass-productionsurvivalfecundityegg-productiongeneration-timepopulation-growthlog-phasecolony-maintenancegenetic-diversityfield-collectionadaptationstress-toleranceoxidative-stresstemperature-extremefood-deficitresource-allocationreproductive-senescenceendogenous-reservesenvironmental-stressorclimate-adaptationphenologymigrationdispersal-patternhabitat-patchsource-sinkpopulation-sourcepopulation-sinkcrop-borderfield-edgenon-crop-habitatrefugeoverwintering-sitespring-emergencesummer-populationfall-populationdensity-dependenceaggregationspatial-distributionsampling-accuracypest-detectionmonitoring-technologyprecision-agriculturedigital-agricultureautomationroboticsmachine-learningcomputer-visionsensor-technologydata-collectiondecision-supportgrower-educationextensionstakeholder-engagementmultidisciplinary-researchacademia-industry-collaborationsustainable-intensificationagroecosystemfood-systemplant-insect-interactioninsect-plant-coevolutionsalivary-secretiondigestive-enzymelytic-enzymetissue-macerationfeeding-injuryphysiological-damagemechanical-damagewound-responseplant-defense-responseinduced-resistancesystemic-acquired-resistancejasmonic-acidethylenesalicylic-acidsecondary-metabolitephenolic-compoundoxidative-enzymeredox-enzymebacterial-transmissionphytopathogenopportunistic-pathogeninfection-courtinoculumdisease-vectorepidemiologycrop-hygienesanitationcrop-rotationcover-croptrap-croppush-pullhabitat-manipulationconservation-biocontrolaugmentative-biocontrolclassical-biocontrolimportationestablishmentnon-target-effecthost-specificityparasitoid-biologyreproductive-parasitoididiobiontkoinobiontmummy-formationemergence-rateparasitoid-impactpopulation-regulationtop-down-controlbottom-up-controltrophic-cascadefood-webecosystem-functionecosystem-servicedisservicenet-effectrisk-assessmentmanagement-strategypolicyregulationpesticide-registrationresistance-monitoringstewardshipintegrated-resistance-managementIRMrefuge-strategypyramid-strategymultiple-toxinstransgenic-cropGMObiosafetyenvironmental-impactnon-target-organismpollinator-protectionbeneficial-insectnatural-enemy-conservationfloral-resourcenectarpollenalternative-preyhabitat-diversificationagri-environment-schemegreen-infrastructureecological-compensationlandscape-complexityconnectivitymatrix-qualityspilloversource-habitatecological-trappopulation-viabilityextinction-riskinvasion-biologyrange-expansionclimate-changewarmingdroughtirrigationwater-stressnitrogen-fertilizationcrop-vigorplant-nutritionhost-plant-qualitytritrophic-interactionplant-microbe-insectendosymbiontgut-microbiomesalivary-microbiomevector-competencetransmission-efficiencyacquisitioninoculationretentionmultiplicationcirculativenon-circulativepersistentnon-persistentpropagativestylet-borneforegut-bornehemolymph-bornesalivary-gland-bornereplicationbarriermidgutsalivary-glandescapeinfectionincubationlatent-periodinoculation-thresholdtransmission-thresholdepidemicepiphytoticdisease-trianglesusceptible-hostvirulent-pathogenfavorable-environmentdisease-cycleprimary-inoculumsecondary-inoculumoverwintering-pathogensurvival-structuredisease-reservoiralternative-hostvolunteer-cropweed-reservoircrop-residuesoilborneseedborneinsect-bornemanagement-tacticpreventivecurativeeradicativesuppressiveprotectantsystemiccontacttranslaminarfumigantbiological-pesticidemicrobial-pesticideentomopathogenic-fungusentomopathogenic-nematodeentomopathogenic-virusbaculovirusgranulovirusnucleopolyhedrovirusbacteriaBacillus-thuringiensisBt-toxincrystal-proteincytolytic-proteinvegetative-insecticidal-proteinmode-of-actiongut-paralysissepticemiatoxemiaantibioticsymbiontantibiotic-resistancefitness-costresistance-alleleresistance-mechanismmetabolic-resistancetarget-site-resistancepenetration-resistancebehavioral-resistancecross-resistancemultiple-resistanceresistance-evolutionselection-pressuredose-responselethal-concentrationlethal-dosemedian-lethal-concentrationmedian-lethal-doseconfidence-intervalprobit-analysislogit-analysisdose-mortalitytime-mortalityresistance-ratioresistance-factordiagnostic-dosediscriminating-dosebaseline-susceptibilitysusceptible-strainreference-strainfield-populationresistant-populationhomozygous-resistantheterozygousdominancerecessiveincomplete-dominancegene-frequencyallele-frequencyselection-coefficientfitnessrelative-fitnesspopulation-geneticsquantitative-geneticsheritabilitygenetic-correlationbreeding-valuegenetic-gainmarker-assisted-selectiongenomic-selectiontranscriptomicsproteomicsmetabolomicsgenomicsfunctional-genomicscomparative-genomicsevolutionary-genomicspopulation-genomicsphylogenomicsbioinformaticscomputational-biologysystems-biologynetwork-analysispathway-analysisgene-ontologyprotein-protein-interactionsignal-transductionhormonejuvenile-hormoneecdysoneinsulin-like-peptideneuropeptidepheromonesex-pheromoneaggregation-pheromonealarm-pheromonetrail-pheromonekairomoneallomonesynomonesemiochemicalolfactionmechanoreceptionvisionphototaxischemotaxisanemotaxisthermotaxishygrotaxisgeotaxiskinesistaxisorthokinesisklinokinesisrheotaxisthigmotaxiscontact-chemoreceptionolfactory-receptorgustatory-receptorionotropic-receptorodorant-binding-proteinchemosensory-proteinsensillumsensory-neuronantennal-lobemushroom-bodylateral-horncentral-complexoptical-lobecompound-eyeocellussimple-eyephotoreceptorrhabdomommatidiumfacetvisual-acuitymotion-detectionpolarized-light-detectioncolor-visionultraviolet-visioninfrared-detectionthermoreceptionhygroreceptionmechanoreceptorchordotonal-organcampaniform-sensillumtrichoid-sensillumbasiconic-sensillumcoeloconic-sensillumplacoid-sensillumsensory-ecologyforaging-behaviorhost-locationhost-acceptancehost-suitabilityoviposition-preferenceoviposition-deterrenceantixenosisantibiosistoleranceresistance-categorygall-midgehessian-flygene-for-geneavirulence-generesistance-geneR-geneeffector-triggered-immunitypattern-triggered-immunitypathogen-associated-molecular-patterndamage-associated-molecular-patternmicrobe-associated-molecular-patternrecognitionperceptiondefense-responsehypersensitive-responsecell-deathprogrammed-cell-deathautophagynecrosisoxidative-burstreactive-oxygen-speciesnitric-oxidecalcium-signalingmitogen-activated-protein-kinasetranscription-factordefense-genepathogenesis-related-proteinantimicrobial-peptidephytoalexinligninsuberincallosepapillacell-wall-reinforcementhypersensitive-cell-deathinduced-systemic-resistanceprimingdefense-elicitorplant-activatorbiostimulantplant-strengthenerorganic-amendmentcompostmanurebiocharvermicompostcompost-teamicrobial-inoculantplant-growth-promoting-rhizobacteriamycorrhizal-fungusendophyteepiphytephytosphererhizospherephyllospherespermosphereanthroposphereecosystem-processnutrient-cyclingcarbon-sequestrationnitrogen-fixationphosphorus-solubilizationpotassium-mobilizationmicronutrient-availabilitysoil-structurewater-infiltrationwater-holding-capacityerosion-controlpollinationpest-suppressiondisease-suppressionweed-suppressiondecompositiondetritivorypredationherbivorymutualismcommensalismcompetitionfacilitationinhibitionamensalismneutralismsymbiosisantagonismtrophic-interactionfood-chainenergy-flowmaterial-cyclingbiomass-pyramidtrophic-pyramidecological-efficiencyproduction-efficiencyassimilation-efficiencyecological-footprintsustainabilityresilienceresistanceadaptabilitytransformabilitysocio-ecological-systemadaptive-managementparticipatory-researchcitizen-scienceknowledge-co-productiontransdisciplinary-researchinterdisciplinary-researchdisciplinary-researchbasic-researchapplied-researchdevelopment-researchaction-researchparticipatory-action-researchfarmer-participatory-researchon-farm-researchstation-researchcontrolled-experimentfield-experimentobservational-studycorrelational-studymanipulative-experimentnatural-experimentquasi-experimentlongitudinal-studycross-sectional-studycase-studycomparative-studymeta-analysissystematic-reviewscoping-reviewrapid-reviewliving-reviewevidence-synthesisknowledge-synthesisresearch-synthesissystematic-mapbibliometric-analysiscitation-analysisco-citation-analysisbibliographic-couplingkeyword-analysistopic-modelingscience-mappingresearch-frontemerging-topichot-topicresearch-gapfuture-directionresearch-priorityresearch-agendastrategic-researchmission-oriented-researchsocietal-challengesustainable-development-goalnutrition-securityclimate-smart-agricultureclimate-resilient-agriculturelow-input-agricultureregenerative-agricultureagroecologyintegrated-farmingmixed-farmingdiversified-farmingspecialized-farmingintensive-farmingextensive-farmingconventional-farmingalternative-farmingurban-agricultureperi-urban-agriculturerural-agriculturesmallholder-agriculturelarge-scale-agriculturefamily-farmingcorporate-farmingcontract-farmingtenant-farmingsubsistence-farmingcommercial-farmingexport-oriented-farmingimport-substituting-farminglocal-food-systemglobal-food-systemfood-value-chainsupply-chainvalue-chaincommodity-chainagribusinessagrifood-systemfood-processingfood-distributionfood-retailfood-servicefood-consumptionfood-wastefood-losspost-harvest-lossstorage-lossprocessing-lossdistribution-lossretail-lossconsumer-wasteplate-wastekitchen-wastefood-rescuefood-recoveryfood-redistributionfood-bankingcompostinganaerobic-digestionbioenergybiofuelbioproductbiomaterialbiorefinerycircular-economycircular-bioeconomygreen-economyblue-economybioeconomynature-based-solutionecosystem-based-adaptationecosystem-based-mitigationclimate-mitigationdisaster-risk-reductionextreme-eventfloodheat-wavecold-snapfrosthailwindstormtornadohurricanetyphooncyclonestorm-surgesea-level-risesalinizationdesertificationland-degradationsoil-degradationsoil-erosionsoil-compactionsoil-sealingsoil-contaminationsoil-pollutionheavy-metalpersistent-organic-pollutantpesticide-residueantibiotic-residuehormone-residueplastic-pollutionmicroplasticnanoplasticparticulate-matterair-pollutionwater-pollutiongroundwater-pollutionsurface-water-pollutioneutrophicationhypoxiadead-zonebiodiversity-losshabitat-losshabitat-fragmentationhabitat-degradationhabitat-destructioninvasive-speciesintroduced-speciesalien-speciesnon-native-speciesexotic-speciesnaturalized-speciescasual-speciesescaped-speciescultivated-speciesornamental-speciespet-specieslivestockpoultryaquaculturefisherycapture-fisherymarine-fisheryfreshwater-fisheryinland-fisheryrecreational-fisherysubsistence-fisheryartisanal-fisheryindustrial-fisheryoverfishingbycatchdiscardsghost-fishingfishing-gearfishing-methodfishing-effortfishing-capacityfishing-pressurestock-assessmentfishery-managementmarine-protected-areano-take-zonemarine-reserveterrestrial-protected-areanational-parknature-reservewildlife-sanctuarybiosphere-reserveworld-heritage-siteramsar-siteimportant-bird-areakey-biodiversity-areaecological-networkprotected-area-networkconservation-corridorwildlife-corridorhabitat-corridorstepping-stonebuffer-zonetransition-zonecore-zonemanagement-zonesustainable-use-zonerestoration-zonerehabilitation-zonereclamation-zoneremediation-zonecontainment-zoneeradication-zonesurveillance-zonemonitoring-zonequarantine-zonebiosecurityphytosanitaryveterinaryanimal-healthplant-healthone-healthecohealthplanetary-healthglobal-healthpublic-healthenvironmental-healthoccupational-healthfood-safetyfeed-safetywater-safetyair-qualityenvironmental-qualityenvironmental-monitoringenvironmental-assessmentenvironmental-impact-assessmentstrategic-environmental-assessmentcumulative-impact-assessmentsocial-impact-assessmenthealth-impact-assessmenthazard-assessmentexposure-assessmentdose-response-assessmentrisk-characterizationrisk-managementrisk-communicationrisk-perceptionrisk-governanceprecautionary-principlepolluter-pays-principleuser-pays-principleextended-producer-responsibilitycorporate-social-responsibilityenvironmental-stewardshipsustainable-livelihoodsustainable-consumptionsustainable-productiongreen-procurementgreen-supply-chaingreen-logisticsgreen-manufacturinggreen-chemistrygreen-technologyclean-technologyenvironmental-technologypollution-preventionwaste-minimizationresource-efficiencyenergy-efficiencywater-efficiencymaterial-efficiencyland-use-efficiencynutrient-use-efficiencynitrogen-use-efficiencyphosphorus-use-efficiencypotassium-use-efficiencymicronutrient-use-efficiencyfertilizer-use-efficiencypesticide-use-efficiencywater-productivityland-productivitylabor-productivitycapital-productivitytotal-factor-productivitypartial-factor-productivityagricultural-productivitycrop-productivitylivestock-productivityfishery-productivityforestry-productivityeconomic-productivitysocial-productivityenvironmental-productivityecological-productivitybiological-productivityprimary-productivitysecondary-productivitynet-primary-productivitygross-primary-productivityecosystem-productivitylandscape-productivityregional-productivityglobal-productivitycarbon-productivitynitrogen-productivitywater-footprintcarbon-footprintnitrogen-footprintphosphorus-footprintbiodiversity-footprintmaterial-footprintland-footprintenergy-footprintemission-factorintensity-factordecouplingabsolute-decouplingrelative-decouplingresource-decouplingimpact-decouplingdematerializationrematerializationtransmaterializationsubstitutioncomplementaritytrade-offsynergyco-benefitwin-winlose-losewin-losezero-sumpositive-sumnegative-sumgame-theoryprisoner's-dilemmatragedy-of-the-commonscollective-actioncommon-pool-resourcepublic-goodprivate-goodclub-goodcommon-goodglobal-commonglobal-public-goodtipping-pointcritical-transitionregime-shifthysteresisirreversibilitypath-dependencelock-intechnological-lock-ininstitutional-lock-inbehavioral-lock-insocial-ecological-trappoverty-trapdegradation-trapdevelopment-trapcomplex-adaptive-systemsocial-ecological-systemcoupled-human-natural-systemhuman-environment-systemadaptive-capacitytransformative-capacitylearning-capacityinnovation-capacitygovernance-capacityinstitutional-capacityorganizational-capacityindividual-capacitycommunity-capacitysocial-capitalhuman-capitalnatural-capitalphysical-capitalfinancial-capitalbuilt-capitalcultural-capitalpolitical-capitalintellectual-capitalknowledge-capitalinformation-capitaldata-capitaldigital-capitaltechnological-capitalinfrastructural-capitalinstitutional-capitalorganizational-capitalnetwork-capitalrelationship-capitalreputational-capitalbrand-capitalsymbolic-capitalexistential-capitalspiritual-capitalreligious-capitalmoral-capitalethical-capitaltrust-capitalreciprocity-capitalsolidarity-capitalcooperation-capitalcoordination-capitalcollaboration-capitalcommunication-capitalinformation-sharingknowledge-sharinglearninginnovationexperimentationtransformationtransitionchangeevolutiondevelopmentgrowthprogressimprovementenhancementoptimizationmaximizationminimizationsatisfactionsufficiencyabundancescarcitysecurityinsecurityvulnerabilityrobustnessstabilityinstabilityequilibriumdisequilibriumhomeostasisallostasisstressdistresseustresscopingcoping-capacitycoping-strategycoping-mechanismcoping-resourcecoping-supportsocial-supportemotional-supportinstrumental-supportinformational-supportappraisal-supportnetwork-supportcommunity-supportinstitutional-supportpolicy-supportfinancial-supporttechnical-supporttechnological-supportinfrastructural-supportcapacity-buildingcapacity-developmentcapacity-strengtheningempowermentenablementmobilizationactivationengagementparticipationinvolvementinclusionexclusionmarginalizationdiscriminationinequityinequalitydisparitygapdividedigital-divideknowledge-divideinformation-dividescience-divideresearch-divideinnovation-dividetechnology-dividedevelopment-dividenorth-south-divideeast-west-divideurban-rural-dividegender-divideage-divideclass-divideincome-dividewealth-divideasset-divideopportunity-divideoutcome-dividewellbeing-dividehappiness-dividelife-satisfaction-dividehealth-divideeducation-divideskill-dividecompetency-dividecapability-dividefunctioning-divideagency-dividefreedom-dividechoice-divideautonomy-divideself-determination-dividedignity-dividerespect-dividerecognition-dividestatus-dividepower-divideinfluence-dividevoice-dividerepresentation-divideparticipation-divideengagement-dividecitizenship-dividebelonging-divideidentity-divideculture-dividevalue-dividebelief-dividenorm-dividepractice-dividebehavior-dividehabit-dividecustom-dividetradition-dividemodernity-divideglobalization-dividelocalization-divideuniversalism-divideparticularism-dividecosmopolitanism-dividenationalism-dividepatriotism-dividepopulism-divideelitism-dividedemocracy-divideauthoritarianism-dividecontrol-dividesecurity-divideliberty-divideprivacy-dividesurveillance-dividetransparency-divideaccountability-divideresponsibility-divideliability-divideguilt-divideblame-dividepunishment-dividereward-divideincentive-dividemotivation-divideintention-divideattention-divideperception-dividecognition-divideemotion-divideaffect-divideattitude-dividepreference-divideinterest-divideneed-dividewant-dividedesire-dividedemand-dividesupply-dividemarket-divideexchange-dividetransaction-dividecontract-divideagreement-dividecooperation-dividecompetition-divideconflict-dividecoercion-divideforce-divideviolence-dividepeace-divideharmony-dividecoexistence-dividetolerance-divideacceptance-dividerejection-divideopposition-divideresistance-divideconformity-dividedeviance-dividenormality-divideabnormality-divideillness-dividedisease-dividedisorder-dividedisability-divideimpairment-dividehandicap-dividecapacity-dividepotential-divideperformance-divideachievement-divideattainment-divideaccomplishment-dividesuccess-dividefailure-divideexcellence-dividemediocrity-divideaverage-dividestandard-dividebenchmark-dividethreshold-divideceiling-dividefloor-dividelimit-divideboundary-divideborder-dividefrontier-dividehorizon-divideperspective-divideviewpoint-dividestandpoint-divideposition-dividestance-divideapproach-dividemethod-dividetechnique-dividetool-divideinstrument-dividedevice-divideapparatus-divideequipment-dividemachine-dividemachinery-dividemechanism-dividesystem-dividestructure-divideorganization-divideinstitution-divideregime-divideorder-dividearrangement-divideconfiguration-dividepattern-dividedesign-divideplan-dividescheme-divideprogram-divideproject-divideinitiative-divideintervention-divideaction-divideactivity-divideoperation-divideprocess-divideprocedure-divideprotocol-divideguideline-dividerecommendation-dividerequirement-divideregulation-dividerule-dividelaw-dividepolicy-dividestrategy-dividetactic-divideobjective-dividegoal-dividetarget-divideaim-dividepurpose-dividemission-dividevision-divideprinciple-divideethic-dividemoral-dividecriterion-divideindicator-dividemeasure-dividemetric-divideindex-dividescale-dividescore-dividerating-divideranking-dividegrade-dividelevel-dividestage-dividephase-dividestep-divideprogression-dividesequence-divideseries-dividesuccession-dividechain-dividecycle-divideloop-dividespiral-dividewave-dividepulse-dividerhythm-dividetempo-dividepace-dividespeed-dividevelocity-divideacceleration-dividedeceleration-dividemomentum-divideinertia-divideenergy-dividework-divideeffort-dividestrain-dividestress-dividepressure-dividetension-dividecompression-divideexpansion-dividecontraction-dividedeformation-dividedistortion-dividetransformation-divideconversion-dividetransduction-dividetransmission-dividetransfer-dividetransport-dividemovement-dividemotion-divideflow-divideflux-dividecurrent-dividestream-divideriver-dividetide-dividewind-dividestorm-divideweather-divideclimate-divideseason-divideyear-dividedecade-dividecentury-dividemillennium-divideepoch-divideera-divideperiod-dividetime-divideduration-divideinterval-dividespan-dividestretch-dividespell-dividebout-divideepisode-divideevent-divideoccurrence-divideincident-divideaccident-dividecoincidence-dividechance-divideluck-dividefortune-dividedestiny-dividefate-dividekarma-dividedharma-dividetao-divideway-dividepath-divideroad-divideroute-dividecourse-dividedirection-divideorientation-dividealignment-dividepositioning-divideplacement-dividelocation-dividesite-divideplace-dividespot-dividepoint-dividesituation-dividecontext-divideenvironment-dividesetting-dividescene-divideplatform-dividebase-dividefoundation-divideground-dividesoil-divideearth-divideland-divideterrain-divideterritory-dividedomain-dividerealm-dividesphere-divideworld-divideglobe-divideplanet-divideuniverse-dividecosmos-dividespace-dividevoid-divideemptiness-dividefullness-divideplenum-dividevacuum-divideatmosphere-divideair-dividegas-dividevapor-dividemist-dividefog-dividecloud-dividerain-dividesnow-divideice-dividefrost-dividedew-dividehumidity-dividemoisture-dividewetness-dividedryness-dividearidity-dividedrought-divideflood-dividedeluge-divideinundation-dividesubmersion-divideimmersion-dividesaturation-dividesupersaturation-dividecondensation-divideevaporation-dividetranspiration-dividesublimation-dividedeposition-divideprecipitation-divideinfiltration-dividepercolation-divideseepage-divideleakage-dividedischarge-dividerunoff-divideerosion-dividesedimentation-divideaccretion-divideaggregation-dividecoagulation-divideflocculation-dividecrystallization-dividesolidification-divideliquefaction-dividevaporization-divideboiling-dividemelting-dividefreezing-dividethawing-dividecooling-divideheating-dividewarming-dividethermal-dividetemperature-divideheat-dividecold-dividecool-dividewarm-dividehot-dividethermodynamic-divideenthalpy-divideentropy-dividefree-energy-divideGibbs-free-energy-divideHelmholtz-free-energy-dividechemical-potential-dividefugacity-dividepartial-pressure-divideconcentration-dividedilution-dividesolution-dividemixture-dividecompound-divideelement-divideatom-dividemolecule-divideion-divideelectron-divideproton-divideneutron-dividenucleus-divideparticle-dividequark-dividelepton-divideboson-dividefermion-dividehadron-dividebaryon-dividemeson-dividephoton-dividegluon-divideweak-boson-divideHiggs-boson-dividestring-dividebrane-dividedimension-dividespace-time-dividecontinuum-dividediscrete-dividequantum-divideclassical-dividerelativistic-divideNewtonian-divideEinsteinian-dividePlanckian-dividecosmological-divideastronomical-dividemicroscopic-dividemacroscopic-dividemesoscopic-dividenanoscopic-divideatomic-dividemolecular-dividecellular-dividetissue-divideorgan-divideorganism-dividepopulation-dividecommunity-divideecosystem-dividebiome-dividebiosphere-divideecosphere-dividenoosphere-dividetechnosphere-divideanthroposphere-dividesociosphere-dividepolitosphere-divideeconosphere-divideculturosphere-dividereligiosphere-divideideosphere-dividememesphere-divideinfosphere-dividecybersphere-dividedigisphere-dividevirtual-divideanalog-dividereal-divideactual-dividepossible-divideimpossible-dividenecessary-dividecontingent-divideprobable-divideimprobable-dividecertain-divideuncertain-dividedeterminate-divideindeterminate-dividedefinite-divideindefinite-dividespecific-dividegeneral-divideparticular-divideuniversal-dividesingular-divideplural-divideindividual-dividecollective-dividesocial-dividecultural-dividehistorical-dividepolitical-divideeconomic-dividereligious-dividephilosophical-dividescientific-dividetechnological-divideartistic-divideliterary-dividemusical-dividevisual-divideperforming-dividecinematic-dividephotographic-dividearchitectural-dividefashion-divideculinary-dividegastronomic-divideolfactory-dividegustatory-dividetactile-dividekinesthetic-divideproprioceptive-dividevestibular-divideauditory-dividesomatosensory-divideinteroceptive-divideexteroceptive-dividemultisensory-dividecrossmodal-dividesynesthetic-divideanesthetic-divideesthetic-divideaesthetic-dividepoetic-dividerhetorical-dividelogical-dividemathematical-dividestatistical-dividecomputational-dividealgorithmic-divideheuristic-dividesystematic-dividemethodical-divideempirical-dividetheoretical-divideconceptual-divideabstract-divideconcrete-dividepractical-divideapplied-dividepure-dividefundamental-dividebasic-dividestrategic-dividetactical-divideoperational-divideadministrative-dividemanagerial-divideexecutive-divideleadership-dividefollowership-dividemembership-dividepartnership-divideownership-dividestewardship-divideguardianship-dividetrusteeship-dividecustodianship-dividewardship-dividetutelage-dividementorship-dividesponsorship-dividechampionship-dividefriendship-dividerelationship-dividekinship-dividecompanionship-dividefellowship-dividescholarship-divideapprenticeship-divideinternship-divideresidency-divideassistantship-divideassociateship-divideauthorship-divideeditorship-dividecuratorship-dividedirectorship-dividegovernorship-dividepresidency-dividechancellorship-dividedeanship-divideheadship-dividechairmanship-divide