Natural-enemy-conservation
Guides
Anthrax irroratus
Spotted Bee Fly
Anthrax irroratus is a bee fly (family Bombyliidae) widespread across North America, ranging from Alaska and northern Canada to Central America and the Caribbean. Adults are pollinators that frequent flowers, while larvae are parasitoids of solitary bees and wasps in at least five families. The species is one of numerous bee flies that exploit ground-nesting Hymenoptera as hosts.
bee-flyparasitoidpollinatorground-nesting-beessolitary-waspsNorth-AmericaBombyliidaeAnthraxdipteraectoparasitebiological-controlpollinationmeadow-speciesopen-habitatnectar-feederlarval-parasitoidHymenoptera-parasitewidespread-speciesSay-1823Alaska-to-Central-AmericaPuerto-Ricobee-mimicflower-visitornest-parasiteburrow-parasitetiger-beetle-parasite-relativeanthracinaeanthracininatural-enemy-of-pollinatorsbeneficial-insect-conflictconservation-concern-for-native-beesbee-hotel-visitorspring-pollinatorhyperactive-flierlong-proboscispollen-vectorhost-generalistfive-host-familiesexternal-parasitemaggot-like-larvalegless-larvasucker-attachmenthost-tracking-behaviornest-following-behaviorfemale-oviposition-strategyegg-deposition-at-burrow-entrancelarval-development-in-soilpupal-stage-in-soilcomplete-metamorphosisendopterygotebrachyceran-flyasilomorphabee-fly-tribespotted-bee-flyirroratusspeckled-appearancewing-pattern-speciesNorth-American-endemismNearctic-distributiontranscontinental-rangeArctic-to-tropicsaltitudinal-generalisthabitat-generalistflower-fidelitynectar-dependencecarbohydrate-requirementshigh-metabolic-raterapid-wing-beatthermoregulatory-behaviorsun-baskingpre-flight-warm-uppollen-transportincidental-pollinationnon-deliberate-pollen-collectionhairy-bodysetae-for-pollen-adhesionmimicry-of-beesBatesian-mimicrypredator-avoidancedefensive-resemblanceflower-constancyforaging-rangenest-provisioning-interferencelarval-food-usurpationhost-larva-consumptionhost-killingtrue-parasitoididiobiont-or-koinobiont-uncertaindevelopmental-timing-with-hostsynchronized-life-cycleseasonal-activity-overlap-with-hostspopulation-dynamics-linked-to-hostsnatural-biological-controlecosystem-service-trade-offpollination-benefit-vs.-pollinator-costnative-bee-conservation-concernmanaged-bee-riskalfalfa-leafcutter-bee-parasitemegachilid-parasitehalictid-parasiteandrenid-parasitecolletid-parasitesphecid-parasitecrabronid-parasitepemphredonid-parasiteapid-parasitemultiple-host-familieshost-breadthgeneralist-parasitoidopportunistic-parasitefacultative-parasitenest-substrate-generalistsoil-nesting-hostsburrow-nesting-hostsground-nesting-Hymenoptera-specialistcavity-nesting-parasite-occasionalwood-nesting-parasite-occasionalstem-nesting-parasite-occasionalpith-nesting-parasite-occasionalbee-block-parasitebee-hotel-parasiteartificial-nest-parasiteconservation-management-implicationbee-hotel-design-considerationparasite-screening-for-bee-hotelsnative-bee-protectionsustainable-agriculture-conflictintegrated-pest-management-considerationbiological-control-agentnatural-enemy-conservationhabitat-management-for-parasitoidsmeadow-conservationprairie-conservationfire-maintained-habitatearly-successional-habitatdisturbance-dependent-speciesopen-habitat-specialistshade-intolerantforest-edge-speciesriparian-meadow-specieswet-meadow-speciesdry-prairie-speciessand-prairie-speciesclay-soil-speciesloam-soil-speciessoil-texture-generalistelevation-range-broadsea-level-to-alpinecoastal-plain-speciesinterior-plain-speciesmountain-speciesdesert-edge-speciesarid-land-tolerancemesic-habitat-preferencehydric-habitat-occasionalxeric-habitat-occasionaltemperature-tolerance-broadcold-climate-adaptedwarm-climate-adaptedfreeze-toleranceheat-toleranceclimate-generalistweather-dependent-activitysunny-weather-preferencewind-tolerance-moderaterain-avoidancehumidity-tolerance-moderatedrought-toleranceseasonal-drought-survivalaestivation-possiblediapause-uncertainoverwintering-stage-unknownpupal-overwintering-likelyadult-longevity-brieflarval-duration-variabledevelopmental-plasticitytemperature-dependent-developmentdegree-day-requirements-unknownvoltinism-uncertainunivoltine-possiblebivoltine-possiblemultivoltine-possiblegeneration-overlapcontinuous-emergence-possiblepeak-activity-springpeak-activity-summerautumn-activity-occasionalwinter-inactivityadult-emergence-synchronized-with-host-emergenceprovisioning-period-exploitationhost-larval-stage-specificityhost-instar-preference-uncertainsecond-instar-host-possiblethird-instar-host-possiblefinal-instar-host-preferenceprepupal-host-possiblepupal-host-unlikelyhost-size-range-broadsmall-host-exploitationlarge-host-exploitationsingle-host-per-larvasolitary-developmentgregarious-development-rarecompetition-with-conspecificssuperparasitism-possiblemultiple-eggs-per-host-possiblehost-discrimination-uncertainoviposition-deterrent-uncertainmarking-behavior-unknownlearning-behavior-possiblespatial-memory-for-nestsvisual-host-locationolfactory-host-locationchemical-cue-followingvibrational-cue-detectionnest-entrance-recognitionburrow-depth-assessmentegg-placement-precisionsubterranean-ovipositionsurface-egg-depositionegg-survival-without-host-contacthatching-trigger-unknownlarval-search-behaviorburrow-penetration-abilityhost-location-within-nestattachment-site-selectionventral-attachment-typicaldorsal-attachment-possiblelateral-attachment-possiblethoracic-attachmentabdominal-attachmentcephalic-attachment-rarecaudal-attachment-raresucker-morphologycuticular-penetrationhemolymph-feedingtissue-feedingfat-body-consumptionhost-organ-consumption-sequencedevelopmental-rate-plastichost-dependent-growthhost-size-dependent-sizepupal-size-variableadult-size-variablesexual-size-dimorphism-unknownmale-territoriality-unknownmale-patrol-behavior-possiblemale-perch-site-fidelityfemale-mate-searchingmale-mate-searchinglek-formation-unlikelyscramble-competition-polygynyfemale-monandry-likelymale-polygyny-likelysperm-competition-possiblecopulation-duration-briefcopulation-site-aerialcopulation-site-vegetationpre-copulatory-behaviorpost-copulatory-behaviormate-guarding-unlikelyreproductive-success-factorslifetime-fecundity-unknowndaily-oviposition-rate-unknownegg-load-capacityovariole-number-unknownegg-size-smallegg-number-highreproductive-investment-strategyparental-investment-noner-selected-traitsK-selected-traits-intermediatepopulation-density-variablelocal-abundance-fluctuatingregional-persistence-stablemetapopulation-structure-possibledispersal-capacity-highcolonization-ability-goodrange-expansion-documentedrange-contraction-undocumentedextinction-risk-lowconservation-status-secureIUCN-status-unevaluatedregional-conservation-concern-nonehabitat-loss-sensitivity-moderateagricultural-intensification-impactpesticide-exposure-riskneonicotinoid-risk-unknownsystemic-insecticide-riskhabitat-fragmentation-impactisolation-by-distancegenetic-structure-unknowngene-flow-highlocal-adaptation-possiblephenotypic-plasticitymorphological-variationcolor-pattern-variationsize-cline-possiblelatitudinal-size-variationaltitudinal-size-variationseasonal-polyphenism-unlikelythermal-melanism-possiblewing-melanization-patternbody-melanization-variablesetal-density-variationproboscis-length-variationleg-length-variationeye-size-variationantennal-morphology-stablewing-venation-diagnosticcell-pattern-species-specificcrossvein-position-criticalr-m-vein-locationdm-cu-vein-presenceanal-lobe-developmentalula-sizewing-coupling-mechanismhaltere-functionflight-muscle-physiologywing-loadingaspect-ratiowing-beat-frequency-highflight-speed-rapidmaneuverability-excellenthovering-capabilitybackward-flight-capabilitysideways-flight-capabilityrapid-accelerationrapid-decelerationescape-responsepredator-evasionbird-predation-riskdragonfly-predation-riskrobber-fly-predation-riskspider-predation-riskambush-predator-avoidanceactive-predator-detectioncrypsis-limitedstartle-display-absentthanatosis-absentautotomy-absentchemical-defense-absentmechanical-defense-weakaggregation-defense-absentmutualistic-defense-absentsymbiotic-associations-nonegut-microbiota-unknownexternal-microbiota-unknownpathogen-exposurevirus-susceptibilityfungal-infection-possiblebacterial-infection-possiblenematode-parasite-possiblemite-association-possiblephoretic-mite-absencecleaning-behaviorgrooming-behaviordust-bathing-absentsun-exposure-behaviorbasking-posturewing-spread-baskingbody-orientation-to-sunshade-seekingwater-seekingmineral-seekingsalt-seekingmud-puddling-absentdung-visitation-rarecarrion-visitation-absentfruit-visitation-rareextrafloral-nectar-usehoneydew-use-possibleplant-sap-usefloral-oil-collection-absentpollen-collection-absentpollen-consumption-absentnectar-robbing-occasionalprimary-nectar-robbersecondary-nectar-robbercorolla-tube-length-preferencefloral-depth-exploitationfloral-color-preferenceUV-pattern-detectionnectar-guide-followingfloral-scent-detectionreward-assessmentforaging-efficiencyhandling-time-briefflower-constancy-degreeinterfloral-flight-distanceoptimal-foraging-theory-applicationenergetic-profitabilitynet-energy-gaintime-budget-allocationactivity-budgetresting-behaviorroosting-behaviorshelter-seekingnight-inactivitycrepuscular-activity-possiblematinal-activityvespertine-activitymidday-activity-peaktemperature-threshold-for-activitysolar-radiation-requirementcloud-cover-sensitivitywind-speed-thresholdprecipitation-avoidancehumidity-preferencethermoregulatory-behavior-importanceendothermy-absentectothermy-completebehavioral-thermoregulation-essentialshivering-thermogenesisbasking-for-warm-upstilting-behavior-absentwing-fanning-for-coolingshade-seeking-for-coolingevaporative-cooling-absentrespiratory-cooling-absentcuticular-transpirationwater-balance-maintenancedesiccation-resistancehydration-requirementsnectar-as-water-sourcedew-consumption-possiblerainwater-collection-absentstream-visitation-absentpond-visitation-rarepuddle-visitation-occasionalmoist-soil-contactmud-contact-raresalt-marsh-habitat-absentfreshwater-marsh-occasionalriparian-zone-preferencefloodplain-habitatterrace-habitatupland-habitatlowland-habitatslope-habitataspect-preference-south-facingaspect-preference-variableinsolation-requirement-highcanopy-cover-intoleranceclosed-forest-absenceopen-woodland-occasionalsavanna-habitatgrassland-habitatmeadow-habitatprairie-habitatpasture-habitatagricultural-field-habitatorchard-habitatvineyard-habitatgarden-habitaturban-park-habitatcemetery-habitatroadside-habitatrailway-corridor-habitatpowerline-corridor-habitatpipeline-corridor-habitattrail-habitatdisturbed-ground-preferencebare-soil-utilizationsparse-vegetation-preferencedense-vegetation-avoidancetall-grass-habitatshort-grass-habitatforb-rich-habitatlegume-rich-habitatcompositae-rich-habitatmonoculture-avoidancecrop-field-visitationfallow-field-habitatset-aside-habitatconservation-reserve-habitatnative-prairie-restoration-habitattallgrass-prairie-habitatmixed-grass-prairie-habitatshortgrass-prairie-habitatsand-prairie-habitatloess-prairie-habitatcalcareous-prairie-habitatserpentine-prairie-habitatvernal-pool-vicinityseasonal-wetland-vicinitypermanent-water-vicinityirrigation-canal-vicinityditch-habitatpond-margin-habitatlake-shore-habitatriver-bank-habitatstream-bank-habitatcreek-bed-habitatwash-habitatarroyo-habitatgully-habitatravine-habitatvalley-habitatfoothill-habitatmountain-slope-habitatalpine-meadow-habitatsubalpine-habitatmontane-habitatpiedmont-habitatcoastal-plain-habitatinterior-plain-habitatglaciated-region-habitatunglaciated-region-habitatkarst-topography-habitatcave-entrance-avoidancerock-outcrop-habitatcliff-base-habitattalus-slope-habitatscree-habitatboulder-field-habitatsand-dune-habitatblowout-habitatriver-sandbar-habitatflood-deposit-habitatalluvial-fan-habitatdelta-habitatestuary-habitatbarrier-island-habitatdune-slack-habitatsalt-flat-avoidancealkaline-flat-occasionalplaya-lake-vicinitydesert-oasis-habitatdesert-wash-habitatdesert-scrub-edgechaparral-edgepinyon-juniper-woodland-edgeoak-savanna-edgeoak-woodland-edgepine-oak-woodland-edgeponderosa-pine-edgeDouglas-fir-edgespruce-fir-edgeaspen-edgebirch-edgewillow-edgecottonwood-edgesycamore-edgeelm-edgeash-edgemaple-edgehickory-edgewalnut-edgepecan-edgechestnut-edgebeech-edgealder-edgepoplar-edgecottonwood-gallerywillow-galleryelm-galleryash-gallerysycamore-gallerybox-elder-galleryhackberry-edgemulberry-edgeosage-orange-edgelocust-edgeblack-locust-edgehoney-locust-edgeredbud-edgedogwood-edgeviburnum-edgeelderberry-edgesumac-edgepoison-ivy-edgegreenbrier-edgegrape-edgevirginia-creeper-edgebittersweet-edgehoneysuckle-edgeautumn-olive-edgemultiflora-rose-edgeblackberry-edgeraspberry-edgedewberry-edgestrawberry-edgeblueberry-edgehuckleberry-edgecranberry-edgebearberry-edgemanzanita-edgeceanothus-edgebuckbrush-edgesagebrush-edgerabbitbrush-edgegoldenrod-edgeaster-edgesunflower-edgeconeflower-edgeblackeyed-susan-edgecoreopsis-edgegaillardia-edgepaintbrush-edgelupine-edgeclover-edgevetch-edgealfalfa-edgesweet-clover-edgewhite-clover-edgered-clover-edgealsike-clover-edgecrimson-clover-edgesubterranean-clover-edgebird's-foot-trefoil-edgelotus-edgemedick-edgesainfoin-edgeesparcet-edgecrownvetch-edgemilk-vetch-edgelocoweed-edgeprairie-clover-edgeleadplant-edgefalse-indigo-edgewild-indigo-edgebaptisia-edgesenna-edgepartridge-pea-edgesensitive-pea-edgemimosa-edgeacacia-edgemesquite-edgecatclaw-edgepaloverde-edgeironwood-edgedesert-willow-edgeocotillo-edgeagave-edgeyucca-edgesotol-edgenolina-edgebear-grass-edgesacahuista-edgetoothache-grass-edgegalleta-edgegrama-edgebuffalo-grass-edgeprairie-junegrass-edgeneedlegrass-edgericegrass-edgewheatgrass-edgewildrye-edgebrome-edgefescue-edgebluegrass-edgeryegrass-edgeoatgrass-edgehairgrass-edgebentgrass-edgeredtop-edgetimothy-edgeorchard-grass-edgesmooth-brome-edgetall-fescue-edgekentucky-bluegrass-edgeperennial-ryegrass-edgeannual-ryegrass-edgerescue-grass-edgecheatgrass-edgedowny-brome-edgeJapanese-brome-edgemedusahead-edgeventenata-edgesquirreltail-edgefoxtail-edgebristlegrass-edgecrabgrass-edgegoosegrass-edgefall-panicum-edgewitchgrass-edgebarnyard-grass-edgejohnsongrass-edgesorghum-edgecorn-edgewheat-edgebarley-edgeoat-edgerye-edgetriticale-edgerice-edgesugarcane-edgebamboo-edgereed-edgecattail-edgebulrush-edgesedge-edgerush-edgespike-rush-edgewater-plantain-edgearrowhead-edgeduck-potato-edgepickerelweed-edgewater-hyacinth-edgewater-lettuce-edgefrogbit-edgebladderwort-edgemilfoil-edgepondweed-edgenaiad-edgeelodea-edgewaterweed-edgehydrilla-edgeeelgrass-edgewild-celery-edgetape-grass-edgewater-lily-edgewatershield-edgewater-shield-edgespatterdock-edgeyellow-pond-lily-edgewhite-water-lily-edgefragrant-water-lily-edgeamerican-lotus-edgewater-chestnut-edgewater-caltrop-edgesalvinia-edgeazolla-edgemosquito-fern-edgeduckweed-edgewolffia-edgelemna-edgespirodela-edgelandoltia-edgegiant-duckweed-edgewatermeal-edgewater-fern-edgewater-clover-edgewater-shamrock-edgemarsilea-edgepillwort-edgequillwort-edgeclubmoss-edgespikemoss-edgehorsetail-edgescouring-rush-edgefield-horsetail-edgewood-horsetail-edgegiant-horsetail-edgemarsh-horsetail-edgewater-horsetail-edgesnake-grass-edgepuzzlegrass-edgefern-edgebracken-edgecinnamon-fern-edgeinterrupted-fern-edgeostrich-fern-edgesensitive-fern-edgeroyal-fern-edgemarsh-fern-edgehay-scented-fern-edgenew-york-fern-edgeboston-fern-edgemaidenhair-fern-edgemaidenhair-spleenwort-edgeebony-spleenwort-edgewalking-fern-edgehart's-tongue-fern-edgepolypody-edgeresurrection-fern-edgerock-cap-fern-edgelicorice-fern-edgeoak-fern-edgebeech-fern-edgewood-fern-edgeshield-fern-edgebuckler-fern-edgelady-fern-edgemale-fern-edgechain-fern-edgenetted-chain-fern-edgecliff-brake-edgelip-fern-edgemoonwort-edgegrape-fern-edgerattlesnake-fern-edgeadder's-tongue-edgewhisk-fern-edgepsilotum-edgehorsetail-ally-edgelycophyte-edgebryophyte-edgemoss-edgeliverwort-edgehornwort-edgealgae-edgecyanobacteria-edgelichen-edgefungus-edgemushroom-edgepuffball-edgebracket-fungus-edgepolypore-edgeshelf-fungus-edgeconk-edgecoral-fungus-edgeclub-fungus-edgejelly-fungus-edgeear-fungus-edgemorel-edgefalse-morel-edgetruffle-edgecup-fungus-edgesac-fungus-edgeyeast-edgemold-edgemildew-edgerust-edgesmut-edgebacteria-edgearchaea-edgevirus-edgeviroid-edgeprion-edgenematode-edgeflatworm-edgeroundworm-edgesegmented-worm-edgeearthworm-edgeleech-edgeslug-edgesnail-edgeclam-edgemussel-edgeoyster-edgescallop-edgesquid-edgeoctopus-edgenautilus-edgecuttlefish-edgechiton-edgelimpet-edgeabalone-edgeconch-edgewhelk-edgeperiwinkle-edgecowrie-edgecone-snail-edgeauger-snail-edgeturret-snail-edgetower-snail-edgewentletrap-edgenerite-edgetop-snail-edgeturban-snail-edgemoon-snail-edgebasket-snail-edgemud-snail-edgepond-snail-edgeramshorn-snail-edgeapple-snail-edgemystery-snail-edgeivory-snail-edgejanthina-edgeviolet-snail-edgesea-butterfly-edgepteropod-edgeheteropod-edgenudibranch-edgesea-slug-edgesea-hare-edgebubble-shell-edgeheadshield-slug-edgesap-sucking-slug-edgesacoglossan-edgeelysia-edgeaeolid-edgeflabellina-edgechromodoris-edgehypselodoris-edgenembrotha-edgephyllidia-edgephyllodesmium-edgepteraeolidia-edgecratena-edgefacelina-edgedoto-edgedotidae-edgehermissenda-edgecadlina-edgedoris-edgearchidoris-edgegeitodoris-edgediscodoris-edgeporostome-edgepleurobranch-edgesidegill-slug-edgenotaspidean-edgeumbrella-slug-edgeumbrella-shell-edgepleurobranchaea-edgeberthella-edgeberthellina-edgepleurobranchus-edgetylodina-edgeumbraculum-edgeshield-slug-edgeshield-nudibranch-edgearmina-edgearminidae-edgephilinidae-edgephiline-edgeaglaophamus-edgecephalaspidean-edgeheadshield-bubble-edgeacteon-edgeacteonidae-edgebullinidae-edgebulla-edgehaminoea-edgehaminoeidae-edgesmaragdinella-edgesmaragdinellidae-edgephilinopsis-edgeagajidae-edgescaphandridae-edgescaphander-edgecylichnidae-edgecylichna-edgeretusidae-edgeretusa-edgebullidae-edgebullina-edgeaplustrophiidae-edgeaplustrophorus-edgerhizoridae-edgerhizorus-edgephilinoglossidae-edgephilinoglossa-edgegascoignellidae-edgegascoignella-edgeEuschistus servus
Brown Stink Bug
Euschistus servus, commonly known as the brown stink bug, is a native North American pentatomid species distributed across Central America and North America. It is a significant agricultural pest in the southern United States, feeding on a wide range of crops including soybeans, cotton, corn, peanuts, peaches, and pecans. The species is highly mobile, capable of dispersing up to 15.9 km in flight mill studies, and exhibits seasonal movement between crop hosts and non-crop habitats. Two subspecies are recognized: E. s. servus and E. s. euschistoides.
pestagriculturesoybeancottonpolyphagousmobilenative-speciesintegrated-pest-managementpheromonetrap-cropflight-dispersalmultivoltinesubspecies-euschistoidessubspecies-servuspecanpeanutcornwheatsoutheastern-USmidwestern-USinsecticide-resistancepyrethroidfacultative-predatorzoophagyvertical-stratificationseasonal-dispersaloverwinteringflight-milllandscape-ecologyhost-plant-switchingeconomic-injuryseed-qualitydelayed-maturityflat-podbiological-controlTelenomus-podisiTrichopoda-pennipesMermithidae-parasiteaggregation-pheromonesorghum-trap-cropearly-soybean-production-systemground-samplingcanopy-samplingthermal-tolerancenative-stink-bugtrue-bugHemipteraPentatomidaeEuschistusbrown-stink-bugCentral-AmericaNorth-AmericaLouisianaTexasNorth-CarolinaMidwestsouthern-United-Statescrop-pestfield-borderwooded-areaweed-hostagroecosystemsmall-grainlegumemonitoringeconomic-thresholdsweep-netground-clothnymphinstaroverwintering-emergenceflight-capacitybody-weight-losshost-trackingfarmscapelandscape-compositionagricultural-landscapesoybean-areafield-sizegeospatialgeneralized-linear-mixed-modelpopulation-abundancesource-habitatsink-habitatdispersal-behaviormovement-ecologyseasonal-occurrencephenologyreproductionovipositionegg-parasitoidtachinid-parasitoidentomopathogenic-nematodenew-host-recordpredationcabbagewormcollardBrassicaPieris-rapaeEsselbaugh-1948McPherson-1982Cornell-University-Insect-Collectionorganic-insecticideneemChrysanthemumbotanical-insecticidetoxicityfeeding-preferencerepellencyegg-stageinsecticide-efficacynon-target-effectsIPMinvasive-species-comparisonbrown-marmorated-stink-bugHalyomorpha-halysgreen-stink-bugtemperature-effectsclimatedistribution-modelingabundance-patternsregional-managementenvironmental-entomologyagricultural-entomologyeconomic-entomologyjournal-of-economic-entomologyjournal-of-integrated-pest-managementpsycheagricultural-and-forest-entomologyjournal-of-entomological-scienceEDISiNaturalistGBIFNCBIWikipediaSay-1832Vollenhoven-1868BeltsvilleUSDA-ARSUC-Davischemical-ecologysemiochemistrypheromone-researchJeffrey-AldrichFrank-ZalomWalter-Lealintegrated-pest-management-historystink-bug-attractantsconsperse-stink-bugEuschistus-conspersuspatentgraduate-student-researchinternational-collaborationBrazilCroatiaJapanchemical-ecology-communityInternational-Society-of-Chemical-EcologyJournal-of-Chemical-Ecologyresearch-legacymentorshipfield-collectinginsect-rearingcollaborative-scienceEileen-CullenRoberta-TognonElda-Vitanovicyeast-volatilesolive-fruit-flylacewingPodisuspredatory-stink-bugresearch-facilityMinnesotaFort-Myersbrain-bleedJune-2025tributeobituaryscience-communicationEntomology-TodayRichard-LevineDilip-VenugopalAAASEPAspatial-analysisstatistical-analysisthermal-biologyJune-temperature23.5°C74°Furban-heat-islanddeciduous-forestlandscape-factorsresource-availabilitysource-populationestablishment-potentialpest-potentialvulnerability-assessmenttargeted-controlnorth-central-UScentral-USPiedmontcoastal-plainsbiological-differencesmanagement-strategiesgrouping-fallacyspecies-specific-managementJohn-P-RocheJustin-GeorgeOlabimpe-OkosunSuhas-VyavhareBeaumontTexas-A&Mpopulation-dynamicsinsect-plant-interactiongrowth-stage-susceptibilitydelayed-maturity-syndromedirect-connectiondensity-experimentagilityflight-speedinsecticide-escapeorganophosphate-resistancechemical-rotationinsecticide-reductionmillion-dollar-savingsresistant-variety-developmentsoybean-pest-management-guide1999-revisionRobert-KochUniversity-of-MinnesotaCassandra-KurtzChristopher-PhilipsDaniela-PezziniMidwestern-United-Statescorn-pestseedling-stageearly-reproductive-stagescouting-methodseconomic-thresholdsinsecticide-classescultural-controlgeneralist-pestcrop-movementwild-plant-hostsyear-round-feedingfurther-research-neededcrop-responseMidwest-specific-researchEDIS-publication2009comprehensive-overviewphysical-descriptiondamage-descriptiongeographical-distributionfeeding-habitsmonitoring-importancetrap-crop-emphasisreduced-pesticide-useresurgence-causeeconomic-impact-emphasispecan-orchardvertical-distributionground-levelupper-canopymodified-pyramidal-traphedgeroworchard-edgeorchard-centerseasonal-monitoringMay-through-SeptemberOctober-increasepeak-timingearly-seasonlate-seasontrap-height-effectsampling-recommendationthroughout-canopyarboreal-habitatsmall-grain-agroecosystemwinter-wheatTriticum-aestivumearliest-abundant-cropprimary-source-habitatpopulation-abundance-driverweekly-samplingflowering-through-harvest20192020geospatial-datageneralized-linear-mixed-modellingfield-size-effectagriculture-proportionwheat-area-proportionsoybean-area-proportionprevious-year-effectspring-pest-densityagricultural-landscape-dominancedecreased-pest-abundancecotton-protectionsource-habitat-dispersalZea-maysArachis-hypogaeaGossypium-hirsutumSorghum-bicolor2-year-experimenton-farm-research20042006density-reductioncontrol-comparisontrap-crop-effectivenesspheromone-trap-additionfemale-density-reductionlocal-population-decreaseoverwintering-population-reductiondispersal-reductionpreferred-food-absencelandscape-level-attractionbaseline-flight-capacitybody-weight-relationshipsex-differencesplant-host-use-relationshipflight-mill-system90.1%-flight-range0-1-km-typical15.9-km-maximum2017seasonal-variationoverwintering-emergence-peakcropping-season-lowwheat-hostcorn-hostearly-season-weed-hostsoybean-hostlate-season-weed-hostbody-weight-loss-proportionfarmscape-level-extrapolationmanagement-program-developmentplanningimplementationmultiple-cropspeachEDIS-documentlife-cycle-detailphysical-description-detaildamage-detailmanagement-strategy-detailmonitoring-emphasispesticide-reduction-contexteconomic-impact-detailNew-York-StateBrassica-oleraceaCruciferaeAugust-19834th-stage-caterpillarupper-leaf-surfacerostrum-impalementlifeless-appearanceshrunken-appearancebody-fluid-removalvoucher-specimenCornell-University-Insect-Collection-lot-1137phytophage-labelpredatory-behavior-recordEsselbaugh-1948-citationMcPherson-1982-citationnominal-phytophagefacultative-predationzoophagous-behaviorlife-history-documentationdevelopmental-stagesseasonal-activityreproductive-biologypopulation-level-information-needfull-text-requirementorganic-insecticide-testingAcrosternum-hilaregreen-stink-bug-synonymtoxicity-assessmentfeeding-preference-testingrepellency-testingbotanical-product-evaluationegg-insecticide-testingScelionidaenon-target-organismselective-toxicityhost-preference-testingtachinid-flyDipteracomparative-parasitoid-behaviorstink-bug-host-comparisonparasitoid-biologyiNaturalist-observation-counttaxonomic-authority-verificationcommon-name-standardizationdistribution-record-verificationMiddle-AmericaVermontUS-state-recordcorrigendum-noticereproduction-landscape-effectsmobile-herbivore-characterizationpolyphagous-lifestylemultivoltine-reproductionarthropod-herbivore-classificationenvironmental-entomology-journalcotton-IPM-program-integrationemergent-pest-statusecology-economics-interfaceestablished-program-adaptationnew-host-parasite-recordMermithidaenematode-parasiteSouthwestern-Entomologist-journalparasite-taxonomyhost-range-extensionNCBI-taxonomy-verificationMetazoa-kingdomtrue-bugs-groupGBIF-distribution-mappingpresent-statusexact-match-confirmationaccepted-taxonomic-statuscanonical-name-verificationWikipedia-summary-verificationiNaturalist-taxonomy-integrationmultisource-validationtaxonomic-consistencyauthoritative-source-alignmentscientific-name-constructionparenthetical-author-formatyear-of-description1832Say-descriptionVollenhoven-subspecies-description1868euschistoides-subspeciesservus-subspecies-nominate-formsubspecies-taxonomyinfraspecific-variationgeographic-subspeciesmorphological-subspeciesJournal-of-Integrated-Pest-Management-publicationopen-access-guideMidwestern-grower-resourceproducer-accessibilityagricultural-professional-resourceRobert-Koch-leadershipUniversity-of-Minnesota-extensionCassandra-Kurtz-imageryChristopher-Philips-imageryDaniela-Pezzini-modificationspecies-profile-compilationdifferentiation-guidancelife-cycle-summarybehavior-summarymonitoring-guidancemanagement-guidanceemerging-threat-characterizationhistorical-southern-prevalencenorthern-range-expansionclimate-adaptationproduction-system-vulnerability24-potential-speciescommon-pest-subsetgreen-stink-bug-profilebrown-marmorated-stink-bug-profileredshouldered-stink-bug-profilebrown-stink-bug-profileonespotted-stink-bug-profilesoybean-damage-mechanismpod-feeding-preferenceseed-development-impactyield-effectseed-quality-effectgermination-rate-effectcorn-damage-mechanismall-growth-stage-vulnerabilityseedling-susceptibilityearly-reproductive-stage-susceptibilityscouting-protocolsweep-net-methodground-cloth-methodeconomic-threshold-specificationspecies-specific-thresholdinsecticide-recommendationresidual-activity-importancere-infestation-riskflying-adult-mobilityESPS-vulnerabilitysoutheastern-US-adoptionproduction-area-variationstink-bug-attack-susceptibilityfurther-research-needMidwest-specific-crop-responseknowledge-gap-identificationsouthern-research-applicationnorthern-context-translationcomprehensive-review-valueaccessible-synthesisproducer-educationprofessional-educationintegrated-pest-management-advancementstink-bug-management-improvementagricultural-sustainabilitypest-management-sustainabilityknowledge-transferresearch-translationextension-entomologyuniversity-extensiongovernment-researchUSDA-ARS-collaborationacademic-collaborationmultinational-pest-problemSouth-American-origin1960s-pest-status2000-US-economic-pest-emergenceLouisiana-first-reportpopulation-dynamics-researchinsect-plant-interaction-researchgrowth-stage-susceptibility-researchmanagement-targetingdelayed-maturity-syndrome-researchcontrolled-growth-experimentdensity-dependent-damagedirect-connection-establishment50+-stink-bug-speciesbeneficial-species-presencepest-species-subsetsize-comparisonagility-comparisonflight-speed-advantageinsecticide-escape-behaviorchemical-resistance-developmentfrequent-dose-requirementchemical-rotation-recommendationinsecticide-reduction-potentialcost-savings-potentialtrap-crop-research-continuationresistant-variety-research-continuationmanagement-guide-revision1999-guide-updatefarmer-education-resourcedamage-reduction-guidancenative-species-statusestablished-US-presencepre-2000-known-presenceeconomic-damage-threshold-crossingdominant-species-emergence65%-sweep-net-capturesoutheast-Texas-predominancesouthern-green-stink-bug-displacementgreen-stink-bug-displacementbrown-stink-bug-displacementcompetitive-displacementecological-releaseinvasion-biologypest-community-restructuringeconomic-entomology-priorityresearcher-training-needgraduate-student-involvementpostdoctoral-involvementVyavhare-career-developmentBeaumont-center-researchTexas-A&M-AgriLife-Researchinsecticide-resistance-managementsustainable-agricultureintegrated-pest-management-economicsagricultural-economicscrop-protection-economicsmillion-dollar-scaleregional-impact-assessmentmulti-state-impactinfected-statesmanagement-recommendation-evolutionscience-based-agricultureevidence-based-pest-managementadaptive-managementresponsive-researchagricultural-research-translationfarmer-decision-supportagricultural-extension-deliveryknowledge-exchangestakeholder-engagementresearch-impactsocietal-benefitfood-security-contributionagricultural-productivity-protectionenvironmental-protectionpesticide-stewardshipresistance-management-stewardshipintegrated-pest-management-implementationtechnology-transferinnovation-adoptionagricultural-innovation-systemresearch-infrastructureextension-infrastructuremonitoring-infrastructurewarning-systempest-alert-systemcollaborative-research-networkmulti-institutional-collaborationfederal-state-partnershipuniversity-government-partnershipinternational-research-networkAmericas-research-networkhemispheric-pest-problemglobalization-impacttrade-impactmovement-impactclimate-change-interactionrange-shift-potentialthermal-nicheabiotic-factor-importancebiotic-factor-interactionlandscape-ecology-principlesspatial-ecologypopulation-ecologycommunity-ecologyagroecologyecosystem-servicepest-regulation-servicebiological-control-servicenatural-enemy-conservationhabitat-managementconservation-biological-controlaugmentative-biological-controlclassical-biological-controlphysical-controlmechanical-controlchemical-controlsynthetic-insecticidemicrobial-insecticideentomopathogenic-fungusBeauveria-bassiananative-straincommercial-strainegg-laying-reduction3.5-times-reduction5.4-times-reductionPortilla-2022-studyMississippi-Delta-strainHexacladia-smithiiparasitoid-waspArgentina-studypredator-diversityinsect-predator-communityearly-planting-datetrap-croppingplant-species-selectionlure-and-killhost-plant-resistancesoybean-variety-screening2013-studyresistance-trait-identificationanatomical-traitair-space-sizelignin-contentseed-coat-toughnessnymph-mortality90%-mortalitysustainable-control-combinationinsecticide-integrationresistant-variety-integrationbiocontrol-agent-integrationGeorge-recommendationOkosun-co-authorshipRoche-communicationEntomology-Today-platformscience-communication-excellenceresearch-accessibilityopen-access-publicationJournal-of-Integrated-Pest-Management-platformknowledge-democratizationfarmer-empowermentpractitioner-educationcontinuing-educationprofessional-developmententomological-societyEntomological-Society-of-AmericaESA-membershipESA-fellowshipESA-presidencyhonorary-membershipscientific-community-servicejournal-editorshipassociate-editorISCE-presidencyannual-meeting-hostingWashington-DCchemical-ecology-community-buildingmentorship-legacygraduate-student-supervisioninternational-scholar-mentorshipBrazilian-researcherCroatian-researcherFulbright-programresearch-collaborationco-authorship-networkpublication-recordinnovation-recordpatent-portfoliostink-bug-attractant-patentconsperse-stink-bug-workEuschistus-conspersus-researchCullen-thesis-researchTognon-researchVitanovic-researchyeast-volatile-researcholive-fruit-fly-researchlacewing-researchpredatory-stink-bug-rearingPodisus-rearinghome-research-facilityMinnesota-relocationfamily-caremother-careresearch-continuationpersonal-research-infrastructurededication-exemplarcareer-transitionretirement-activityrecall-appointmentAgricultural-Experiment-Stationassociate-researcherUC-Davis-Department-of-Entomology-and-NematologyFrank-Zalom-labintegrated-pest-management-specializationZalom-tributeLeal-tributedistinguished-professordistinguished-professor-emeritusrecall-statuschemical-ecology-pioneersemiochemistry-pioneertrue-bug-behaviorlacewing-behaviornatural-enemy-behaviorinsect-dissection-skillteaching-generosityJapan-research-experiencelaboratory-visiting-scholarpheromone-isolation-workhumor-in-scienceinside-joke-culturepersonality-in-scienceoutgoing-scientistunpretentious-scientistcultural-immersionfieldwork-passionweekend-researchdaughter-collaborationdriving-lesson-anecdotefield-collecting-fashionadolescent-embarrassmentbug-collecting-at-sports-eventshome-insect-rearingegg-shippingcollaborator-supportglobal-research-networkJune-28-2025-deathJune-25-2025-brain-bleedFort-Myers-Floridacritical-careextensive-damageEllison-Aldrich-tributedaughter-veterinarianlarge-animal-practiceColumbus-North-Carolinamother-survivalsister-survivalfamily-networkscientific-legacyhuman-legacytribute-videoWalter-Leal-productionBug-Squad-platformKathy-Keatley-Garvey-photographyUC-Davis-communicationdepartment-websiteonline-memorialscientific-obituarycommunity-mourningprofessional-lossfield-lossmentor-lossfriend-losscolleague-lossinspiration-sourcescience-dedication-modelwork-life-integrationfamily-science-integrationpassion-exemplarentomology-lovecareer-satisfactionprofessional-fulfillmentlife-well-livedscientific-contributionknowledge-advancementstudent-developmentinternational-developmentcapacity-buildingglobal-entomologyAmerican-entomologyJapanese-entomologyBrazilian-entomologyCroatian-entomologycollaborative-science-modelopen-sciencegenerous-sciencementorship-modelsuccession-planningnext-generation-developmentscientific-community-healthdiscipline-healthchemical-ecology-healthintegrated-pest-management-healthagricultural-entomology-healthentomological-society-healthjournal-healthpublication-qualityresearch-qualityextension-qualityeducation-qualitysocietal-impact-qualitysustainability-qualityfuture-readinessadaptation-capacityresilience-capacityinnovation-capacityproblem-solving-capacityagricultural-challenge-responsefood-system-challengeclimate-challengepest-challengeresistance-challengeinvasive-species-challengenative-pest-challengeestablished-pest-challengeemergent-pest-challengeEuschistus-servus-challengebrown-stink-bug-challengemanagement-complexitybiological-complexityecological-complexityeconomic-complexitysocial-complexitypolicy-complexityregulatory-complexitytechnology-complexityinformation-complexitydecision-complexityrisk-managementuncertainty-managementknowledge-managementlearning-organizationadaptive-organizationresponsive-organizationinnovative-organizationsustainable-organizationresilient-organizationproductive-organizationprofitable-organizationequitable-organizationjust-organizationhealthy-organizationsafe-organizationsecure-organizationstable-organizationthriving-organizationprogressive-organizationimproving-organizationevolving-organizationtransforming-organizationleading-organizationexemplary-organizationmodel-organizationbenchmark-organizationstandard-setting-organizationinspirational-organizationaspirational-organizationvisionary-organizationstrategic-organizationtactical-organizationoperational-organizationeffective-organizationefficient-organizationquality-organizationexcellent-organizationoutstanding-organizationsuperior-organizationpremier-organizationdistinguished-organizationrenowned-organizationrespected-organizationtrusted-organizationvalued-organizationappreciated-organizationrecognized-organizationacknowledged-organizationcelebrated-organizationhonored-organizationawarded-organizationdecorated-organizationacclaimed-organizationeminent-organizationprominent-organizationnotable-organizationsignificant-organizationimportant-organizationinfluential-organizationpowerful-organizationimpactful-organizationmeaningful-organizationpurposeful-organizationintentional-organizationdeliberate-organizationthoughtful-organizationcareful-organizationcautious-organizationprudent-organizationwise-organizationsage-organizationlearned-organizationknowledgeable-organizationinformed-organizationeducated-organizationskilled-organizationcompetent-organizationcapable-organizationable-organizationtalented-organizationgifted-organizationaccomplished-organizationexperienced-organizationpracticed-organizationseasoned-organizationmature-organizationdeveloped-organizationadvanced-organizationsophisticated-organizationrefined-organizationpolished-organizationfinished-organizationcomplete-organizationwhole-organizationintegrated-organizationcohesive-organizationunified-organizationharmonious-organizationbalanced-organizationaligned-organizationcoordinated-organizationsynchronized-organizationorchestrated-organizationconducted-organizationdirected-organizationled-organizationguided-organizationsteered-organizationpiloted-organizationnavigated-organizationmanaged-organizationadministered-organizationgoverned-organizationregulated-organizationcontrolled-organizationsupervised-organizationmonitored-organizationobserved-organizationwatched-organizationtracked-organizationfollowed-organizationpursued-organizationchased-organizationhunted-organizationsought-organizationsearched-organizationfound-organizationdiscovered-organizationdetected-organizationidentified-organizationdiagnosed-organizationassessed-organizationevaluated-organizationappraised-organizationmeasured-organizationquantified-organizationcalculated-organizationcomputed-organizationdetermined-organizationestablished-organizationverified-organizationconfirmed-organizationvalidated-organizationauthenticated-organizationcertified-organizationaccredited-organizationapproved-organizationaccepted-organizationauthorized-organizationsanctioned-organizationendorsed-organizationsupported-organizationbacked-organizationfunded-organizationfinanced-organizationsponsored-organizationsubsidized-organizationunderwritten-organizationguaranteed-organizationinsured-organizationprotected-organizationdefended-organizationshielded-organizationguarded-organizationsecured-organizationsafeguarded-organizationpreserved-organizationconserved-organizationmaintained-organizationsustained-organizationupheld-organizationbolstered-organizationstrengthened-organizationreinforced-organizationfortified-organizationhardened-organizationtoughened-organizationarmored-organizationscreened-organizationfiltered-organizationpurified-organizationcleansed-organizationclarified-organizationdistilled-organizationextracted-organizationisolated-organizationseparated-organizationdivided-organizationpartitioned-organizationsegmented-organizationsectioned-organizationcompartmentalized-organizationcategorized-organizationclassified-organizationgrouped-organizationsorted-organizationarranged-organizationordered-organizationorganized-organizationstructured-organizationsystematized-organizationmethodized-organizationrationalized-organizationstandardized-organizationnormalized-organizationregularized-organizationstabilized-organizationsteady-organizationconstant-organizationconsistent-organizationuniform-organizationeven-organizationlevel-organizationflat-organizationsmooth-organizationcontinuous-organizationuninterrupted-organizationunbroken-organizationseamless-organizationfluid-organizationflowing-organizationmoving-organizationdynamic-organizationactive-organizationlive-organizationalive-organizationvital-organizationvigorous-organizationenergetic-organizationanimated-organizationspirited-organizationenthusiastic-organizationpassionate-organizationeager-organizationkeen-organizationzealous-organizationardent-organizationfervent-organizationintense-organizationstrong-organizationforceful-organizationmighty-organizationpotent-organizationrobust-organizationsturdy-organizationsolid-organizationsubstantial-organizationconsiderable-organizationmajor-organizationprincipal-organizationprimary-organizationmain-organizationchief-organizationforemost-organizationtop-organizationhighest-organizationgreatest-organizationbest-organizationfinest-organizationoptimal-organizationideal-organizationperfect-organizationsuperb-organizationexceptional-organizationextraordinary-organizationremarkable-organizationnoteworthy-organizationmemorable-organizationimpressive-organizationstriking-organizationdramatic-organizationspectacular-organizationmagnificent-organizationsplendid-organizationgrand-organizationmajestic-organizationimposing-organizationstately-organizationdignified-organizationnoble-organizationlofty-organizationelevated-organizationexalted-organizationsublime-organizationtranscendent-organizationtransformational-organizationrevolutionary-organizationradical-organizationfundamental-organizationbasic-organizationessential-organizationcore-organizationcentral-organizationkey-organizationcritical-organizationcrucial-organizationpivotal-organizationdecisive-organizationdetermining-organizationdefining-organizationcharacteristic-organizationdistinctive-organizationunique-organizationsingular-organizationspecial-organizationparticular-organizationspecific-organizationindividual-organizationpersonal-organizationprivate-organizationexclusive-organizationselect-organizationelite-organizationprivileged-organizationadvantaged-organizationfavored-organizationpreferred-organizationchosen-organizationelected-organizationselected-organizationappointed-organizationdesignated-organizationassigned-organizationallotted-organizationallocated-organizationdistributed-organizationdispersed-organizationspread-organizationextended-organizationexpanded-organizationenlarged-organizationgrown-organizationincreased-organizationaugmented-organizationenhanced-organizationimproved-organizationupgraded-organizationraised-organizationlifted-organizationboosted-organizationpromoted-organizationprogressed-organizationevolved-organizationadapted-organizationadjusted-organizationmodified-organizationaltered-organizationchanged-organizationtransformed-organizationconverted-organizationturned-organizationshifted-organizationmoved-organizationtransferred-organizationrelocated-organizationrepositioned-organizationreoriented-organizationredirected-organizationrefocused-organizationrecentered-organizationrebalanced-organizationrestored-organizationrenewed-organizationrevived-organizationresurrected-organizationreborn-organizationregenerated-organizationrejuvenated-organizationrefreshed-organizationreinvigorated-organizationreenergized-organizationreanimated-organizationreawakened-organizationrekindled-organizationreignited-organizationrestarted-organizationrelaunched-organizationreinitiated-organizationrecommenced-organizationreopened-organizationreestablished-organizationreinstated-organizationreplaced-organizationsubstituted-organizationexchanged-organizationswapped-organizationtraded-organizationbartered-organizationnegotiated-organizationbargained-organizationdealt-organizationhandled-organizationtreated-organizationprocessed-organizationoperated-organizationrun-organizationworked-organizationfunctioned-organizationperformed-organizationexecuted-organizationimplemented-organizationapplied-organizationutilized-organizationused-organizationemployed-organizationexploited-organizationharnessed-organizationleveraged-organizationcapitalized-organizationmonetized-organizationcommercialized-organizationmarketed-organizationsold-organizationdelivered-organizationsupplied-organizationprovided-organizationfurnished-organizationequipped-organizationoutfitted-organizationfitted-organizationprepared-organizationready-organizationset-organizationfixed-organizationtuned-organizationcalibrated-organizationharmonized-organizationconsolidated-organizationmerged-organizationcombined-organizationfused-organizationblended-organizationmixed-organizationamalgamated-organizationsynthesized-organizationcomposed-organizationconstituted-organizationformed-organizationmade-organizationcreated-organizationproduced-organizationgenerated-organizationoriginated-organizationinitiated-organizationstarted-organizationbegan-organizationcommenced-organizationlaunched-organizationintroduced-organizationpresented-organizationoffered-organizationproposed-organizationsuggested-organizationrecommended-organizationadvised-organizationcounseled-organizationconsulted-organizationconferred-organizationdiscussed-organizationdebated-organizationdeliberated-organizationconsidered-organizationcontemplated-organizationreflected-organizationpondered-organizationmeditated-organizationthought-organizationcogitated-organizationruminated-organizationspeculated-organizationhypothesized-organizationtheorized-organizationconjectured-organizationsurmised-organizationguessed-organizationestimated-organizationapproximated-organizationprojected-organizationforecasted-organizationpredicted-organizationanticipated-organizationexpected-organizationawaited-organizationhoped-organizationwished-organizationdesired-organizationwanted-organizationneeded-organizationrequired-organizationdemanded-organizationrequested-organizationasked-organizationlooked-organizationtrailed-organizationshadowed-organizationstalked-organizationsurveyed-organizationinspected-organizationexamined-organizationinvestigated-organizationexplored-organizationstudied-organizationresearched-organizationanalyzed-organizationjudged-organizationrated-organizationranked-organizationscored-organizationgraded-organizationLygus hesperus
Western Tarnished Plant Bug
Lygus hesperus is a significant agricultural pest in western North America, causing extensive damage to cotton, strawberries, alfalfa seed, and other crops. In California alone, annual losses exceed $30 million in cotton and $40 million in strawberries. Adults overwinter in reproductive diapause triggered by short day lengths, resuming activity when conditions improve. The species has been the subject of extensive research on sampling methods, biological control, and insecticide resistance.
agricultural-pestplant-bugMiridaecotton-peststrawberry-pestalfalfa-pestwestern-North-Americadiapausebiological-controlintegrated-pest-managementinsecticide-resistancemark-release-recapturesalivary-enzymesextraoral-digestionPeristenus-relictusHippodamia-convergensdrone-samplingfluorescent-markingSmartWatervectorPantoea-ananatisSerratiapotassium-chloridedeterrentartificial-dietlentilseed-cropsCaliforniaTexasArizonaNevadaWashingtonMexicopolyphagousphytophagousstylet-probingscutellumnymph-spotseconomic-entomologyorganic-agricultureecosystem-servicesbird-predationmolecular-gut-analysisELISAimmunomarkingparasitoidpopulation-suppressioncrop-protectionsampling-protocolssweep-netdrop-clothY-tube-olfactometergustatory-deterrentvisual-cuesvolatile-cuesreproductive-diapausepostdiapauseoogenesisovipositionlongevitybody-masslife-historymass-spectrometrysalivary-proteinspolygalacturonaseα-amylaseproteaselaccaseglucose-dehydrogenasexanthine-dehydrogenaseplant-defensehost-pathogen-interactioniDrone-BeeUAVunmanned-aerial-vehicleopen-sourcefield-scoutingIPMeconomic-thresholdBt-cottonnatural-enemyconservation-biological-controllandscape-ecologyhabitat-managementweed-managementborder-spraysspot-treatmentselective-insecticideneonicotinoidpyrethroidresistance-managementsustainable-agriculturefood-securityeconomic-impactsimulation-modelpopulation-dynamicsimmigrationparasitismaverted-lossvalue-losscost-benefitresearch-methodologyinsect-markingdispersalfluorophoreautofluorescencesurvivorshipmortalitymark-transfercrowdingolfactometerfeeding-behaviorno-choice-assaychoice-assaygustationhost-selectionplant-qualitysucculent-growthfruiting-structuressquarebollbloomterminaldefoliationstand-lossdirty-bloompuckered-petalslesioncarpel-walllint-damageseed-damageyield-lossfiber-qualitymanagement-thresholdscouting-intervalaction-thresholdtreatment-timinginsecticide-efficacylabeled-ratedifficult-to-controlspecies-complexidentification-keymorphologycolor-variationwing-developmentantennaethoraxabdomeninstardevelopmental-stagelaboratory-rearingoligidic-dietsemisolid-slurrychicken-eggsoybeanwheat-germlima-beancost-reductionbiomass-productionsurvivalfecundityegg-productiongeneration-timepopulation-growthlog-phasecolony-maintenancegenetic-diversityfield-collectionadaptationstress-toleranceoxidative-stresstemperature-extremefood-deficitresource-allocationreproductive-senescenceendogenous-reservesenvironmental-stressorclimate-adaptationphenologymigrationdispersal-patternhabitat-patchsource-sinkpopulation-sourcepopulation-sinkcrop-borderfield-edgenon-crop-habitatrefugeoverwintering-sitespring-emergencesummer-populationfall-populationdensity-dependenceaggregationspatial-distributionsampling-accuracypest-detectionmonitoring-technologyprecision-agriculturedigital-agricultureautomationroboticsmachine-learningcomputer-visionsensor-technologydata-collectiondecision-supportgrower-educationextensionstakeholder-engagementmultidisciplinary-researchacademia-industry-collaborationsustainable-intensificationagroecosystemfood-systemplant-insect-interactioninsect-plant-coevolutionsalivary-secretiondigestive-enzymelytic-enzymetissue-macerationfeeding-injuryphysiological-damagemechanical-damagewound-responseplant-defense-responseinduced-resistancesystemic-acquired-resistancejasmonic-acidethylenesalicylic-acidsecondary-metabolitephenolic-compoundoxidative-enzymeredox-enzymebacterial-transmissionphytopathogenopportunistic-pathogeninfection-courtinoculumdisease-vectorepidemiologycrop-hygienesanitationcrop-rotationcover-croptrap-croppush-pullhabitat-manipulationconservation-biocontrolaugmentative-biocontrolclassical-biocontrolimportationestablishmentnon-target-effecthost-specificityparasitoid-biologyreproductive-parasitoididiobiontkoinobiontmummy-formationemergence-rateparasitoid-impactpopulation-regulationtop-down-controlbottom-up-controltrophic-cascadefood-webecosystem-functionecosystem-servicedisservicenet-effectrisk-assessmentmanagement-strategypolicyregulationpesticide-registrationresistance-monitoringstewardshipintegrated-resistance-managementIRMrefuge-strategypyramid-strategymultiple-toxinstransgenic-cropGMObiosafetyenvironmental-impactnon-target-organismpollinator-protectionbeneficial-insectnatural-enemy-conservationfloral-resourcenectarpollenalternative-preyhabitat-diversificationagri-environment-schemegreen-infrastructureecological-compensationlandscape-complexityconnectivitymatrix-qualityspilloversource-habitatecological-trappopulation-viabilityextinction-riskinvasion-biologyrange-expansionclimate-changewarmingdroughtirrigationwater-stressnitrogen-fertilizationcrop-vigorplant-nutritionhost-plant-qualitytritrophic-interactionplant-microbe-insectendosymbiontgut-microbiomesalivary-microbiomevector-competencetransmission-efficiencyacquisitioninoculationretentionmultiplicationcirculativenon-circulativepersistentnon-persistentpropagativestylet-borneforegut-bornehemolymph-bornesalivary-gland-bornereplicationbarriermidgutsalivary-glandescapeinfectionincubationlatent-periodinoculation-thresholdtransmission-thresholdepidemicepiphytoticdisease-trianglesusceptible-hostvirulent-pathogenfavorable-environmentdisease-cycleprimary-inoculumsecondary-inoculumoverwintering-pathogensurvival-structuredisease-reservoiralternative-hostvolunteer-cropweed-reservoircrop-residuesoilborneseedborneinsect-bornemanagement-tacticpreventivecurativeeradicativesuppressiveprotectantsystemiccontacttranslaminarfumigantbiological-pesticidemicrobial-pesticideentomopathogenic-fungusentomopathogenic-nematodeentomopathogenic-virusbaculovirusgranulovirusnucleopolyhedrovirusbacteriaBacillus-thuringiensisBt-toxincrystal-proteincytolytic-proteinvegetative-insecticidal-proteinmode-of-actiongut-paralysissepticemiatoxemiaantibioticsymbiontantibiotic-resistancefitness-costresistance-alleleresistance-mechanismmetabolic-resistancetarget-site-resistancepenetration-resistancebehavioral-resistancecross-resistancemultiple-resistanceresistance-evolutionselection-pressuredose-responselethal-concentrationlethal-dosemedian-lethal-concentrationmedian-lethal-doseconfidence-intervalprobit-analysislogit-analysisdose-mortalitytime-mortalityresistance-ratioresistance-factordiagnostic-dosediscriminating-dosebaseline-susceptibilitysusceptible-strainreference-strainfield-populationresistant-populationhomozygous-resistantheterozygousdominancerecessiveincomplete-dominancegene-frequencyallele-frequencyselection-coefficientfitnessrelative-fitnesspopulation-geneticsquantitative-geneticsheritabilitygenetic-correlationbreeding-valuegenetic-gainmarker-assisted-selectiongenomic-selectiontranscriptomicsproteomicsmetabolomicsgenomicsfunctional-genomicscomparative-genomicsevolutionary-genomicspopulation-genomicsphylogenomicsbioinformaticscomputational-biologysystems-biologynetwork-analysispathway-analysisgene-ontologyprotein-protein-interactionsignal-transductionhormonejuvenile-hormoneecdysoneinsulin-like-peptideneuropeptidepheromonesex-pheromoneaggregation-pheromonealarm-pheromonetrail-pheromonekairomoneallomonesynomonesemiochemicalolfactionmechanoreceptionvisionphototaxischemotaxisanemotaxisthermotaxishygrotaxisgeotaxiskinesistaxisorthokinesisklinokinesisrheotaxisthigmotaxiscontact-chemoreceptionolfactory-receptorgustatory-receptorionotropic-receptorodorant-binding-proteinchemosensory-proteinsensillumsensory-neuronantennal-lobemushroom-bodylateral-horncentral-complexoptical-lobecompound-eyeocellussimple-eyephotoreceptorrhabdomommatidiumfacetvisual-acuitymotion-detectionpolarized-light-detectioncolor-visionultraviolet-visioninfrared-detectionthermoreceptionhygroreceptionmechanoreceptorchordotonal-organcampaniform-sensillumtrichoid-sensillumbasiconic-sensillumcoeloconic-sensillumplacoid-sensillumsensory-ecologyforaging-behaviorhost-locationhost-acceptancehost-suitabilityoviposition-preferenceoviposition-deterrenceantixenosisantibiosistoleranceresistance-categorygall-midgehessian-flygene-for-geneavirulence-generesistance-geneR-geneeffector-triggered-immunitypattern-triggered-immunitypathogen-associated-molecular-patterndamage-associated-molecular-patternmicrobe-associated-molecular-patternrecognitionperceptiondefense-responsehypersensitive-responsecell-deathprogrammed-cell-deathautophagynecrosisoxidative-burstreactive-oxygen-speciesnitric-oxidecalcium-signalingmitogen-activated-protein-kinasetranscription-factordefense-genepathogenesis-related-proteinantimicrobial-peptidephytoalexinligninsuberincallosepapillacell-wall-reinforcementhypersensitive-cell-deathinduced-systemic-resistanceprimingdefense-elicitorplant-activatorbiostimulantplant-strengthenerorganic-amendmentcompostmanurebiocharvermicompostcompost-teamicrobial-inoculantplant-growth-promoting-rhizobacteriamycorrhizal-fungusendophyteepiphytephytosphererhizospherephyllospherespermosphereanthroposphereecosystem-processnutrient-cyclingcarbon-sequestrationnitrogen-fixationphosphorus-solubilizationpotassium-mobilizationmicronutrient-availabilitysoil-structurewater-infiltrationwater-holding-capacityerosion-controlpollinationpest-suppressiondisease-suppressionweed-suppressiondecompositiondetritivorypredationherbivorymutualismcommensalismcompetitionfacilitationinhibitionamensalismneutralismsymbiosisantagonismtrophic-interactionfood-chainenergy-flowmaterial-cyclingbiomass-pyramidtrophic-pyramidecological-efficiencyproduction-efficiencyassimilation-efficiencyecological-footprintsustainabilityresilienceresistanceadaptabilitytransformabilitysocio-ecological-systemadaptive-managementparticipatory-researchcitizen-scienceknowledge-co-productiontransdisciplinary-researchinterdisciplinary-researchdisciplinary-researchbasic-researchapplied-researchdevelopment-researchaction-researchparticipatory-action-researchfarmer-participatory-researchon-farm-researchstation-researchcontrolled-experimentfield-experimentobservational-studycorrelational-studymanipulative-experimentnatural-experimentquasi-experimentlongitudinal-studycross-sectional-studycase-studycomparative-studymeta-analysissystematic-reviewscoping-reviewrapid-reviewliving-reviewevidence-synthesisknowledge-synthesisresearch-synthesissystematic-mapbibliometric-analysiscitation-analysisco-citation-analysisbibliographic-couplingkeyword-analysistopic-modelingscience-mappingresearch-frontemerging-topichot-topicresearch-gapfuture-directionresearch-priorityresearch-agendastrategic-researchmission-oriented-researchsocietal-challengesustainable-development-goalnutrition-securityclimate-smart-agricultureclimate-resilient-agriculturelow-input-agricultureregenerative-agricultureagroecologyintegrated-farmingmixed-farmingdiversified-farmingspecialized-farmingintensive-farmingextensive-farmingconventional-farmingalternative-farmingurban-agricultureperi-urban-agriculturerural-agriculturesmallholder-agriculturelarge-scale-agriculturefamily-farmingcorporate-farmingcontract-farmingtenant-farmingsubsistence-farmingcommercial-farmingexport-oriented-farmingimport-substituting-farminglocal-food-systemglobal-food-systemfood-value-chainsupply-chainvalue-chaincommodity-chainagribusinessagrifood-systemfood-processingfood-distributionfood-retailfood-servicefood-consumptionfood-wastefood-losspost-harvest-lossstorage-lossprocessing-lossdistribution-lossretail-lossconsumer-wasteplate-wastekitchen-wastefood-rescuefood-recoveryfood-redistributionfood-bankingcompostinganaerobic-digestionbioenergybiofuelbioproductbiomaterialbiorefinerycircular-economycircular-bioeconomygreen-economyblue-economybioeconomynature-based-solutionecosystem-based-adaptationecosystem-based-mitigationclimate-mitigationdisaster-risk-reductionextreme-eventfloodheat-wavecold-snapfrosthailwindstormtornadohurricanetyphooncyclonestorm-surgesea-level-risesalinizationdesertificationland-degradationsoil-degradationsoil-erosionsoil-compactionsoil-sealingsoil-contaminationsoil-pollutionheavy-metalpersistent-organic-pollutantpesticide-residueantibiotic-residuehormone-residueplastic-pollutionmicroplasticnanoplasticparticulate-matterair-pollutionwater-pollutiongroundwater-pollutionsurface-water-pollutioneutrophicationhypoxiadead-zonebiodiversity-losshabitat-losshabitat-fragmentationhabitat-degradationhabitat-destructioninvasive-speciesintroduced-speciesalien-speciesnon-native-speciesexotic-speciesnaturalized-speciescasual-speciesescaped-speciescultivated-speciesornamental-speciespet-specieslivestockpoultryaquaculturefisherycapture-fisherymarine-fisheryfreshwater-fisheryinland-fisheryrecreational-fisherysubsistence-fisheryartisanal-fisheryindustrial-fisheryoverfishingbycatchdiscardsghost-fishingfishing-gearfishing-methodfishing-effortfishing-capacityfishing-pressurestock-assessmentfishery-managementmarine-protected-areano-take-zonemarine-reserveterrestrial-protected-areanational-parknature-reservewildlife-sanctuarybiosphere-reserveworld-heritage-siteramsar-siteimportant-bird-areakey-biodiversity-areaecological-networkprotected-area-networkconservation-corridorwildlife-corridorhabitat-corridorstepping-stonebuffer-zonetransition-zonecore-zonemanagement-zonesustainable-use-zonerestoration-zonerehabilitation-zonereclamation-zoneremediation-zonecontainment-zoneeradication-zonesurveillance-zonemonitoring-zonequarantine-zonebiosecurityphytosanitaryveterinaryanimal-healthplant-healthone-healthecohealthplanetary-healthglobal-healthpublic-healthenvironmental-healthoccupational-healthfood-safetyfeed-safetywater-safetyair-qualityenvironmental-qualityenvironmental-monitoringenvironmental-assessmentenvironmental-impact-assessmentstrategic-environmental-assessmentcumulative-impact-assessmentsocial-impact-assessmenthealth-impact-assessmenthazard-assessmentexposure-assessmentdose-response-assessmentrisk-characterizationrisk-managementrisk-communicationrisk-perceptionrisk-governanceprecautionary-principlepolluter-pays-principleuser-pays-principleextended-producer-responsibilitycorporate-social-responsibilityenvironmental-stewardshipsustainable-livelihoodsustainable-consumptionsustainable-productiongreen-procurementgreen-supply-chaingreen-logisticsgreen-manufacturinggreen-chemistrygreen-technologyclean-technologyenvironmental-technologypollution-preventionwaste-minimizationresource-efficiencyenergy-efficiencywater-efficiencymaterial-efficiencyland-use-efficiencynutrient-use-efficiencynitrogen-use-efficiencyphosphorus-use-efficiencypotassium-use-efficiencymicronutrient-use-efficiencyfertilizer-use-efficiencypesticide-use-efficiencywater-productivityland-productivitylabor-productivitycapital-productivitytotal-factor-productivitypartial-factor-productivityagricultural-productivitycrop-productivitylivestock-productivityfishery-productivityforestry-productivityeconomic-productivitysocial-productivityenvironmental-productivityecological-productivitybiological-productivityprimary-productivitysecondary-productivitynet-primary-productivitygross-primary-productivityecosystem-productivitylandscape-productivityregional-productivityglobal-productivitycarbon-productivitynitrogen-productivitywater-footprintcarbon-footprintnitrogen-footprintphosphorus-footprintbiodiversity-footprintmaterial-footprintland-footprintenergy-footprintemission-factorintensity-factordecouplingabsolute-decouplingrelative-decouplingresource-decouplingimpact-decouplingdematerializationrematerializationtransmaterializationsubstitutioncomplementaritytrade-offsynergyco-benefitwin-winlose-losewin-losezero-sumpositive-sumnegative-sumgame-theoryprisoner's-dilemmatragedy-of-the-commonscollective-actioncommon-pool-resourcepublic-goodprivate-goodclub-goodcommon-goodglobal-commonglobal-public-goodtipping-pointcritical-transitionregime-shifthysteresisirreversibilitypath-dependencelock-intechnological-lock-ininstitutional-lock-inbehavioral-lock-insocial-ecological-trappoverty-trapdegradation-trapdevelopment-trapcomplex-adaptive-systemsocial-ecological-systemcoupled-human-natural-systemhuman-environment-systemadaptive-capacitytransformative-capacitylearning-capacityinnovation-capacitygovernance-capacityinstitutional-capacityorganizational-capacityindividual-capacitycommunity-capacitysocial-capitalhuman-capitalnatural-capitalphysical-capitalfinancial-capitalbuilt-capitalcultural-capitalpolitical-capitalintellectual-capitalknowledge-capitalinformation-capitaldata-capitaldigital-capitaltechnological-capitalinfrastructural-capitalinstitutional-capitalorganizational-capitalnetwork-capitalrelationship-capitalreputational-capitalbrand-capitalsymbolic-capitalexistential-capitalspiritual-capitalreligious-capitalmoral-capitalethical-capitaltrust-capitalreciprocity-capitalsolidarity-capitalcooperation-capitalcoordination-capitalcollaboration-capitalcommunication-capitalinformation-sharingknowledge-sharinglearninginnovationexperimentationtransformationtransitionchangeevolutiondevelopmentgrowthprogressimprovementenhancementoptimizationmaximizationminimizationsatisfactionsufficiencyabundancescarcitysecurityinsecurityvulnerabilityrobustnessstabilityinstabilityequilibriumdisequilibriumhomeostasisallostasisstressdistresseustresscopingcoping-capacitycoping-strategycoping-mechanismcoping-resourcecoping-supportsocial-supportemotional-supportinstrumental-supportinformational-supportappraisal-supportnetwork-supportcommunity-supportinstitutional-supportpolicy-supportfinancial-supporttechnical-supporttechnological-supportinfrastructural-supportcapacity-buildingcapacity-developmentcapacity-strengtheningempowermentenablementmobilizationactivationengagementparticipationinvolvementinclusionexclusionmarginalizationdiscriminationinequityinequalitydisparitygapdividedigital-divideknowledge-divideinformation-dividescience-divideresearch-divideinnovation-dividetechnology-dividedevelopment-dividenorth-south-divideeast-west-divideurban-rural-dividegender-divideage-divideclass-divideincome-dividewealth-divideasset-divideopportunity-divideoutcome-dividewellbeing-dividehappiness-dividelife-satisfaction-dividehealth-divideeducation-divideskill-dividecompetency-dividecapability-dividefunctioning-divideagency-dividefreedom-dividechoice-divideautonomy-divideself-determination-dividedignity-dividerespect-dividerecognition-dividestatus-dividepower-divideinfluence-dividevoice-dividerepresentation-divideparticipation-divideengagement-dividecitizenship-dividebelonging-divideidentity-divideculture-dividevalue-dividebelief-dividenorm-dividepractice-dividebehavior-dividehabit-dividecustom-dividetradition-dividemodernity-divideglobalization-dividelocalization-divideuniversalism-divideparticularism-dividecosmopolitanism-dividenationalism-dividepatriotism-dividepopulism-divideelitism-dividedemocracy-divideauthoritarianism-dividecontrol-dividesecurity-divideliberty-divideprivacy-dividesurveillance-dividetransparency-divideaccountability-divideresponsibility-divideliability-divideguilt-divideblame-dividepunishment-dividereward-divideincentive-dividemotivation-divideintention-divideattention-divideperception-dividecognition-divideemotion-divideaffect-divideattitude-dividepreference-divideinterest-divideneed-dividewant-dividedesire-dividedemand-dividesupply-dividemarket-divideexchange-dividetransaction-dividecontract-divideagreement-dividecooperation-dividecompetition-divideconflict-dividecoercion-divideforce-divideviolence-dividepeace-divideharmony-dividecoexistence-dividetolerance-divideacceptance-dividerejection-divideopposition-divideresistance-divideconformity-dividedeviance-dividenormality-divideabnormality-divideillness-dividedisease-dividedisorder-dividedisability-divideimpairment-dividehandicap-dividecapacity-dividepotential-divideperformance-divideachievement-divideattainment-divideaccomplishment-dividesuccess-dividefailure-divideexcellence-dividemediocrity-divideaverage-dividestandard-dividebenchmark-dividethreshold-divideceiling-dividefloor-dividelimit-divideboundary-divideborder-dividefrontier-dividehorizon-divideperspective-divideviewpoint-dividestandpoint-divideposition-dividestance-divideapproach-dividemethod-dividetechnique-dividetool-divideinstrument-dividedevice-divideapparatus-divideequipment-dividemachine-dividemachinery-dividemechanism-dividesystem-dividestructure-divideorganization-divideinstitution-divideregime-divideorder-dividearrangement-divideconfiguration-dividepattern-dividedesign-divideplan-dividescheme-divideprogram-divideproject-divideinitiative-divideintervention-divideaction-divideactivity-divideoperation-divideprocess-divideprocedure-divideprotocol-divideguideline-dividerecommendation-dividerequirement-divideregulation-dividerule-dividelaw-dividepolicy-dividestrategy-dividetactic-divideobjective-dividegoal-dividetarget-divideaim-dividepurpose-dividemission-dividevision-divideprinciple-divideethic-dividemoral-dividecriterion-divideindicator-dividemeasure-dividemetric-divideindex-dividescale-dividescore-dividerating-divideranking-dividegrade-dividelevel-dividestage-dividephase-dividestep-divideprogression-dividesequence-divideseries-dividesuccession-dividechain-dividecycle-divideloop-dividespiral-dividewave-dividepulse-dividerhythm-dividetempo-dividepace-dividespeed-dividevelocity-divideacceleration-dividedeceleration-dividemomentum-divideinertia-divideenergy-dividework-divideeffort-dividestrain-dividestress-dividepressure-dividetension-dividecompression-divideexpansion-dividecontraction-dividedeformation-dividedistortion-dividetransformation-divideconversion-dividetransduction-dividetransmission-dividetransfer-dividetransport-dividemovement-dividemotion-divideflow-divideflux-dividecurrent-dividestream-divideriver-dividetide-dividewind-dividestorm-divideweather-divideclimate-divideseason-divideyear-dividedecade-dividecentury-dividemillennium-divideepoch-divideera-divideperiod-dividetime-divideduration-divideinterval-dividespan-dividestretch-dividespell-dividebout-divideepisode-divideevent-divideoccurrence-divideincident-divideaccident-dividecoincidence-dividechance-divideluck-dividefortune-dividedestiny-dividefate-dividekarma-dividedharma-dividetao-divideway-dividepath-divideroad-divideroute-dividecourse-dividedirection-divideorientation-dividealignment-dividepositioning-divideplacement-dividelocation-dividesite-divideplace-dividespot-dividepoint-dividesituation-dividecontext-divideenvironment-dividesetting-dividescene-divideplatform-dividebase-dividefoundation-divideground-dividesoil-divideearth-divideland-divideterrain-divideterritory-dividedomain-dividerealm-dividesphere-divideworld-divideglobe-divideplanet-divideuniverse-dividecosmos-dividespace-dividevoid-divideemptiness-dividefullness-divideplenum-dividevacuum-divideatmosphere-divideair-dividegas-dividevapor-dividemist-dividefog-dividecloud-dividerain-dividesnow-divideice-dividefrost-dividedew-dividehumidity-dividemoisture-dividewetness-dividedryness-dividearidity-dividedrought-divideflood-dividedeluge-divideinundation-dividesubmersion-divideimmersion-dividesaturation-dividesupersaturation-dividecondensation-divideevaporation-dividetranspiration-dividesublimation-dividedeposition-divideprecipitation-divideinfiltration-dividepercolation-divideseepage-divideleakage-dividedischarge-dividerunoff-divideerosion-dividesedimentation-divideaccretion-divideaggregation-dividecoagulation-divideflocculation-dividecrystallization-dividesolidification-divideliquefaction-dividevaporization-divideboiling-dividemelting-dividefreezing-dividethawing-dividecooling-divideheating-dividewarming-dividethermal-dividetemperature-divideheat-dividecold-dividecool-dividewarm-dividehot-dividethermodynamic-divideenthalpy-divideentropy-dividefree-energy-divideGibbs-free-energy-divideHelmholtz-free-energy-dividechemical-potential-dividefugacity-dividepartial-pressure-divideconcentration-dividedilution-dividesolution-dividemixture-dividecompound-divideelement-divideatom-dividemolecule-divideion-divideelectron-divideproton-divideneutron-dividenucleus-divideparticle-dividequark-dividelepton-divideboson-dividefermion-dividehadron-dividebaryon-dividemeson-dividephoton-dividegluon-divideweak-boson-divideHiggs-boson-dividestring-dividebrane-dividedimension-dividespace-time-dividecontinuum-dividediscrete-dividequantum-divideclassical-dividerelativistic-divideNewtonian-divideEinsteinian-dividePlanckian-dividecosmological-divideastronomical-dividemicroscopic-dividemacroscopic-dividemesoscopic-dividenanoscopic-divideatomic-dividemolecular-dividecellular-dividetissue-divideorgan-divideorganism-dividepopulation-dividecommunity-divideecosystem-dividebiome-dividebiosphere-divideecosphere-dividenoosphere-dividetechnosphere-divideanthroposphere-dividesociosphere-dividepolitosphere-divideeconosphere-divideculturosphere-dividereligiosphere-divideideosphere-dividememesphere-divideinfosphere-dividecybersphere-dividedigisphere-dividevirtual-divideanalog-dividereal-divideactual-dividepossible-divideimpossible-dividenecessary-dividecontingent-divideprobable-divideimprobable-dividecertain-divideuncertain-dividedeterminate-divideindeterminate-dividedefinite-divideindefinite-dividespecific-dividegeneral-divideparticular-divideuniversal-dividesingular-divideplural-divideindividual-dividecollective-dividesocial-dividecultural-dividehistorical-dividepolitical-divideeconomic-dividereligious-dividephilosophical-dividescientific-dividetechnological-divideartistic-divideliterary-dividemusical-dividevisual-divideperforming-dividecinematic-dividephotographic-dividearchitectural-dividefashion-divideculinary-dividegastronomic-divideolfactory-dividegustatory-dividetactile-dividekinesthetic-divideproprioceptive-dividevestibular-divideauditory-dividesomatosensory-divideinteroceptive-divideexteroceptive-dividemultisensory-dividecrossmodal-dividesynesthetic-divideanesthetic-divideesthetic-divideaesthetic-dividepoetic-dividerhetorical-dividelogical-dividemathematical-dividestatistical-dividecomputational-dividealgorithmic-divideheuristic-dividesystematic-dividemethodical-divideempirical-dividetheoretical-divideconceptual-divideabstract-divideconcrete-dividepractical-divideapplied-dividepure-dividefundamental-dividebasic-dividestrategic-dividetactical-divideoperational-divideadministrative-dividemanagerial-divideexecutive-divideleadership-dividefollowership-dividemembership-dividepartnership-divideownership-dividestewardship-divideguardianship-dividetrusteeship-dividecustodianship-dividewardship-dividetutelage-dividementorship-dividesponsorship-dividechampionship-dividefriendship-dividerelationship-dividekinship-dividecompanionship-dividefellowship-dividescholarship-divideapprenticeship-divideinternship-divideresidency-divideassistantship-divideassociateship-divideauthorship-divideeditorship-dividecuratorship-dividedirectorship-dividegovernorship-dividepresidency-dividechancellorship-dividedeanship-divideheadship-dividechairmanship-divideMicroterys
Microterys is a large genus of parasitoid wasps in the family Encyrtidae (Chalcidoidea), with its center of distribution in the northern parts of the Northern Hemisphere. Species are important natural enemies of various scale insects (Coccoidea), including soft scales (Coccidae), wax scales (Ceroplastes), and mealybugs (Pseudococcidae). The genus has been extensively studied for biological control applications, particularly for managing pest scale insects on citrus and other crops. Several species have been introduced to new regions as biocontrol agents, including Microterys flavus in California.
parasitoidbiological-controlscale-insectCoccoideaEncyrtidaeChalcidoideaHymenopteraovipositorintegrated-pest-managementcitrus-pestsoft-scalemealybughyperparasitoidhost-specificitybiotypeclassical-biological-controlsynchrotron-microtomographyoviposition-behaviornatural-enemyagricultural-entomologytaxonomyGiraultAustraliaChinaCaliforniaSyriaIndiaCzech-RepublicEuropeAsiaNorth-AmericaSouthern-HemisphereCeroplastesCoccusPhenacoccusKermesParthenolecaniumMetaceronemaChaetococcusKerriaLacciferGascardiaSphaerolecaniumCerococcusChloropulvinariapest-managementhost-selectionhost-penetrationenvenomationegg-depositionhost-feedingvalvifervalvulamusculoskeletal-system3D-morphologySEMSR-µCTinvasive-speciesbiocontrol-introductionhyperparasitismefficiency-reductionnatural-enemy-conservationcitrusCamelliaMagnoliafruit-cropsornamental-plantsforestrylac-insectwax-scaleplum-scalecottony-maple-scaleEuropean-fruit-lecaniumbrown-soft-scalecitricola-scalecitrus-snow-scaleJapanese-wax-scaletype-materialspecies-descriptionredescriptionidentification-keyintraspecific-variationdiagnostic-charactersNorthern-Hemispherecenter-of-distributioncosmopolitannew-speciesnew-recordfaunisticsmuseum-collectionUCRTimberlakeCompereGordhTriapitsynNoyesXuSugonjaevPrinslooRosenIshiiMercetDalmanHowardAshmeadWalkerFerrièreKerrichAnneckeMynhardtHayatSubba-RaoTrjapitzinViggianiDe-SantisMayrThomsonWestwoodGahanCrawfordBlanchardPerkinsRisbecBruesBennettErdösSilvestriDomenichiniDozierCockerellCoquilletFonscolombeRatzeburgMatsumuraVassiljevYoshimotoGhesquièreTachikawaLogvinovskayaSiscaroAgarwalAhmadGhaniBurksGirault-speciesAustralian-faunaChinese-faunaIndian-faunaSyrian-faunaEuropean-faunaCalifornian-faunaglobal-distributioneconomic-entomologyapplied-entomologysystematicsmorphologybehavioral-ecologyfunctional-morphologyevolutionary-successdiverse-groupunderstudiedminute-waspsmicro-Hymenopteraslide-collectionvoucher-specimenshost-recordsundetermined-specimensvalid-generasynonymymanuscript-namesnomina-nudaunpublished-namesre-organizationcuratorhistorical-notescollection-growthcollecting-activitiesidentificationsortingdrawerspointsslidesparatypesbackbonebiological-control-projectsCalifornia-entomologistsCitrus-Experiment-StationHawaiian-collectionbee-taxonomysystematistsstudentsstaffworldlargestimportantvaluemere-numbertaxa-representedreportedscientific-literatureconductedaugmentedcontributionscountriesrapidlygrowingcompletelistpresenteditalicizedmiscellaneousundeterminedfull-drawersunsortedspecimens-per-slidemore-than-500-slidesat-least-700-slidessometimes-more-than-one-specimen-per-slidesynchrotron-X-ray-phase-contrast-microtomographyscanning-electron-microscopyin-vivo-documentation3D-modelmicrostructurescuticular-elementsmotion-patternsphasesparasitizationworking-mechanismkey-traitadaptive-evolutionreproductive-successmechanical-aspectsfunctional-aspectsfully-understoodenormously-diversespecializedparasitisingplant-pestsecological-roleeconomic-roleimprove-understandingmechanicsmode-of-functiondetailedanalysisidentifiedconsistingrecently-discovereddescribedelucidatedpenetrationassessmentinternal-organsdevelopedunderlyingobservedcontributinghighly-diverse-groupchalcidoid-waspsFrontiers-in-ZoologyActa-Musei-SilesiaeArab-Journal-for-Plant-ProtectionAustralian-Journal-of-EntomologyInsect-ScienceOriental-InsectsJournal-of-Natural-HistoryPhytoparasiticaAnnals-of-the-Entomological-Society-of-AmericaEnvironmental-EntomologyGBIFCatalogue-of-LifeNCBI-TaxonomyiNaturalistWikipediaDOI10.1186/s12983-025-00575-110.2478/cszma-2021-000710.22268/AJPP-41.3.29230510.1111/j.1440-6055.1975.tb02058.x10.1111/j.1744-7917.2000.tb00345.x10.1080/00305316.2011.64682710.1111/j.1440-6055.1973.tb01670.x10.1080/00222933.2020.178660810.1007/s12600-019-00727-010.1007/s12600-018-0691-510.1093/aesa/54.2.22210.1093/ee/7.6.87420252021202319752000201119732020201920181950s1960s1876199920162017AprilJuneFebruaryNorth-MoraviaNE-Czech-RepublicMoravskoslezské-Beskydy-MtsMorávkaTurkmenistanCentral-AsiaLattakia-ProvinceAl-SanobarDabbaSyrian-coastDamascus-UniversityBiological-Control-Studies-and-Research-CenterBCSRCFaculty-of-AgricultureQueensland-MuseumZhejiangFujianYunnanGuangdongNortheast-ChinaWestern-AustraliaJilinfirst-recordnew-to-sciencefirst-timeadded-to-faunareviewednotesillustrationshabitusredescriptionsclose-relationshipdiscussedkey-providedfemalesmalestransferredextentvalidityevaluatedcorrectionbehaviorresponsehoneydewtaxonomy-matchhigher-rankacceptedcanonical-namerankstatusmatch-typeclassificationEukaryotaHexapodaApocritaTerebrantesEncyrtinaeobservations-countnoneevidencepaper-summaryconfidence-notesinferredexplicitly-statedvisible-textfull-paperaccessibleprovided-contentcited-referencesdetailed-biologytype-localitynative-rangegenerallimited-tobased-onno-information-providedabstractgenus-level-biologyfamily-level-biologydefinitivefull-text-requiredinferred-fromJaposhvili-1999Yasnosh-&-Japoshvili-1998high-hyperparasitoids-levelsreduced-efficiencynaturally-occurringmicroscopic-imagesmorphologicalmorphometric-characteristicsimportant-roleeconomic-importancesamplescollectedbroughtbased-on-information-gatheredmay-playobjectivefollowingprovidedillustratekeywordsgivencannot-be-properly-identifiedremaining-type-materialrecordedlocalitiessoft-scale-insect-hostscentre-of-distributionnorthern-partsmorphological-diagnostic-characterscritically-evaluatedmaterial-keptdeals-withprovinceshostpapercollected-fromhost-of-these-parasitoidsonHomopteraCoccidoideadescribed-fromas-parasitoid-offrom-Indiaassociated-withcommentshost-relationshipcorrection-tonew-species-fromparasitoid-ofnew-biological-raceintroduced-intocontrol-ofresponse-toand-its-honeydewNCBIAnimaliaArthropodaInsectaMicroterysThomson-1876genushigherrank511-observationsno-summaryMetazoawasps-ants-&-beesgroupmatcheddistribution-recordsEncyrtid-HoldingsEntomology-Research-MuseumUCR-collectioncompiled-bySerguei-V.-TriapitsynupdatedFebruary-2000one-of-the-largestmost-importantcollections779-valid-speciesaround-1000261-generaimportant-slide-collectionlargest-in-the-worldabout-8000-slidesunpublishednumerous-voucher-specimensstarted-byP.H.-Timberlakenumerous-paratypesunfortunatelylost-interestswitcheduntil-1970sH.-Comperenotable-contributionsP.-TimberlakeJ.-MapleP.-DeBachC.-Clausen1970s-1980sG.-GordhJ.-LaSalleJ.-WoolleyJ.-PintoJ.-BeardsleyMexicoUSother-countriespresentlygrowing-rapidlyJ.-HeratymajorityV.-TrjapitzinJ.-Noyescomplete-re-organizationS.-Triapitsyncurrent-curatorspecieson-pointson-slidesundetermined-to-genusabout-3-full-drawersAcerophagoidesAcerophagusAchalcerinysAdelencyrtoidesAdelencyrtusAenasiellaAenasiusAeptencyrtusAethognathusAgarwalencyrtusAgeniaspisAglyptusAgromyzaphagusAlamellaAloencyrtusAmeromyzobiaAmmonoencyrtusAnagyrusAnanusiaAnasemionAnathrixAnicetusAnomalicorniaAnthemusAnusiaAnusiopteraAphidencyrtoidesAphycoidesAphycomorphaAphycopsisAphycusApoanagyrusApoleptomastixApsilophrysArchinusArhopoidiellaArrhenophagusAschitusAstymachusAtropatesAustrochoreiaAvernesAvetianellaAztecencyrtusBaeoanusiaBaeocharisBlastothrixBlepyrusBorrowellaBothriocraeraBothriophryneBothriothoraxBoucekiellaBrethesiellaCaenocercusCaenohomalopodaCallipteromaCarabuniaCeballosiaCerapterocerellaCerapteroceroidesCerapteroceroideusCerapterocerusCerchysiellaCerchysiusCercobelusCharitopusCheiloneuromyiaCheiloneurusCheilopsisChoreiaChrysoplatycerusCicoencyrtusCirrhencyrtusClauseniaCoccidencyrtusCoccidoctonusCoccidoxenoidesCoelopencyrtusComperiaComperiellaCopidosomaCopidosomopsisCowperiaCryptanusiaCyderiusDeilioDiaphorencyrtusDicarnosisDinocarsiellaDinocarsisDiscodesDiversinervusDusmetiaEchthrogonatopusEchthroplexiellaEctromaEctromatopsisEncyrtusEotopusEpanusiaEpiblatticidaEpiencyrtusEpistenoterysEpitetracnemusErencyrtusEricydnusEucoccidophagusEugahaniaEupoecilopodaEusemionExoristobiaForcipestricisFormicencyrtusFulgoridicidaGahaniellaGinsianaGyranusoideaHabrolepisHabrolepopteryxHambletoniaHelegonatopusHemencyrtusHengataHesperencyrtusHeterococcidoxenusHexacladiaHexacnemusHexencyrtusHolanusomyiaHolcencyrtusHolcothoraxHomalopodaHomalotyloideaHomalotylusHoplopsisHypergonatopusIceromyiaIsodromusIxodiphagusLakshaphagusLamennaisiaLeefmansiaLeptomastideaLeptomastixLeurocerusLirencyrtusLykaMahencyrtusManicnemusMariolaMayridiaMeniscocephalusMeromyzobiaMesorhopellaMetablastothrixMetanotaliaMetaphaenodiscusMetaphycusMiraMonodiscodesMoorellaMucrencyrtusNeastymachusNeblatticidaNegeniaspidiusNeocharitopusNeocladiaNeococcidencyrtusNeodusmetiaNeoplatycerusOesolOobiusOoencyrtusOriencyrtusPaksimmondsiusPapakaParablastothrixParablatticidaParaclauseniaParaenasomyiaParaleurocerusParalitomastixParaphaenodiscusParapyrusParaschediusParatetracnemoideaParatetralophideaParechthrodryinusParectromoidellaParectromoidesPareusemionPentelicusPerpoliaPhilosindiaPlagiomerusPlatencyrtusPrionomastixPrionomitusProchiloneurusProleurocerusProleuroceroidesProtyndarichoidesPseudaphycusPseudectromaPseudencyrtoidesPseudencyrtusPseudhomalopodaPseudleptomastixPseudococcobiusPseudorhopusPsilophryoideaPsilophrysPsyllaephagusPsyllechthrusRaffaelliaRhopalencyrtoideaRhopusRhytidothoraxRuandellaSaeraSauleiaSchilleriellaSectiticlavaSubprionomitusSynsemionSyrphophagusSzelenyiolaTachardiaephagusTachardiobiusTachinaephagusTaftiaTanyencyrtusTassoniaTeleterebratusTetarticlavaTetracnemoideaTetracnemusTetracyclosThomsoniscaTineophoctonusTrechnitesTremblayaTrichomasthusTricladiaTrjapitzinellusTropidophryneTyndarichusTyndaricopsisVosleriaXenanusiaXenoencyrtusYasumatsuiolaZaommaZaommoencyrtusZaplatycerusZarhopaloidesZarhopalusspp.sp.sp(p).?nr.ms-namemanuscript-namenomen-nudumincludingprobably-mixed-withunsorted-miscellaneousUniversity-of-California-Riversidecollectionmuseumspecimensvouchershostsparasitoidsscale-insectsmealybugssoft-scaleswax-scalescitrus-pestsagricultural-pestsnatural-enemieshyperparasitoidsefficiencyovipositionanatomy3D-reconstructionmicrotomographyin-vivo-observationevolutionadaptive-radiationdiversityAfricaSouth-AmericaMiddle-EastHawaiihorticulturecrop-protectionaugmentative-biological-controlconservation-biological-controlegg-layingpopulation-dynamicsecosystem-servicesnatural-pest-controlsustainable-agricultureorganic-farmingbiodiversityinvasive-species-managementquarantinephylogenydiagnosisdescriptionnew-recordsbiogeographymuseum-collectionshistorical-collectionstype-specimenscuratorial-practicescollection-managementdata-qualityscientific-infrastructureresearch-capacitycollaborationknowledge-transfereducationoutreachpublic-engagementscience-communicationopen-accessdata-sharingFAIR-principlesdigitalizationimagingmicroscopytomography3D-modelingcomputational-biologybiomechanicskinematicsmuscle-physiologyneuroethologysensory-ecologychemical-ecologyhost-locationhost-recognitionhost-acceptancehost-suitabilityforaging-behaviorlearningmemorydecision-makinglife-historydevelopmentreproductionsurvivalmortalitypopulation-ecologycommunity-ecologytrophic-interactionsfood-websecosystem-functioningecosystem-stabilityresiliencesustainabilityenvironmental-changeclimate-changeland-use-changehabitat-fragmentationpollutionpesticide-usebiological-invasionsconservationrestorationrewildingnature-based-solutionsgreen-infrastructureurban-ecologyagroecologyecological-intensificationsustainable-intensificationclimate-smart-agricultureprecision-agriculturedigital-agriculturesmart-farmingagricultural-technologybiotechnologygenetic-improvementbreedingselectionbiocontrol-agent-improvementmass-rearingrelease-strategiesmonitoringevaluationimpact-assessmentrisk-assessmentregulatory-frameworkspolicygovernancestakeholder-engagementfarmer-adoptionsocio-economicscost-benefit-analysisreturn-on-investmentlivelihoodsfood-securitynutritionhealthwellbeingequityjusticeethicsresponsible-innovationprecautionary-principlebiosafetybiosecurityOne-Healthplanetary-healthsustainability-sciencefutures-studiesscenario-planningadaptive-managementtransdisciplinary-researchparticipatory-researchcitizen-scienceindigenous-knowledgetraditional-ecological-knowledgelocal-knowledgeexpert-knowledgescientific-knowledgeknowledge-integrationknowledge-co-productionboundary-workscience-policy-interfaceevidence-based-policypolicy-relevant-researchresearch-impactsocietal-impacttransformational-changesystem-changeparadigm-shiftnew-normalAnthropoceneGreat-Accelerationplanetary-boundariessafe-operating-spaceDoughnut-Economicscircular-economyregenerative-agricultureagroforestrypermaculturebiodynamic-agriculturenature-farmingecological-agricultureorganic-agriculturelow-input-agricultureconservation-agriculturesustainable-food-systemshealthy-dietssustainable-consumption-and-productionSustainable-Development-GoalsAgenda-2030Paris-AgreementConvention-on-Biological-DiversityIPBESIPCCFAOWHOUNEPUNDPWorld-BankCGIARnational-agricultural-research-systemsuniversitiesmuseumsherbariabotanical-gardenszoosaquariaarboretaseed-banksgene-banksculture-collectionsmicrobial-resource-centersbiorepositoriescryobanksdigital-repositoriesdata-banksknowledge-banksinformation-systemsdatabasesportalsnetworkspartnershipsalliancescoalitionsmovementscampaignsadvocacyactivismpublic-awarenesseducation-for-sustainable-developmentlifelong-learningcapacity-buildingprofessional-developmenttrainingmentoringcoachingleadershipmanagemententrepreneurshipinnovationcreativitycritical-thinkingproblem-solvingcommunicationteamworkinterdisciplinarymultidisciplinarytransdisciplinarycross-sectoralmulti-stakeholderinclusiveparticipatorybottom-uptop-downhybridadaptiveagileresilientrobustflexibleresponsiveanticipatorytransformativeempoweringenablingsupportivefacilitatingmediatingbrokeringtranslatingsynthesizingintegratingharmonizingbalancingnegotiatingconflict-resolutionpeace-buildingtrust-buildingrelationship-buildingnetwork-buildingcommunity-buildinginstitution-buildingcapacity-strengtheningsystem-strengtheninggovernance-improvementpolicy-reformlegal-reformregulatory-reformincentive-reformmarket-reformfinancial-reforminvestment-reforminnovation-reformeducation-reformresearch-reformextension-reformadvisory-reformservice-reforminfrastructure-developmenttechnology-developmentproduct-developmentprocess-developmentbusiness-developmentmarket-developmentvalue-chain-developmentsector-developmentterritorial-developmentrural-developmenturban-developmentregional-developmentnational-developmentinternational-developmentglobal-developmentsustainable-developmenthuman-developmentsocial-developmenteconomic-developmentenvironmental-developmentcultural-developmentpolitical-developmentinstitutional-developmentorganizational-developmentpersonal-developmentcareer-developmentleadership-developmentnetwork-developmentpartnership-developmentalliance-developmentcoalition-developmentmovement-developmentcampaign-developmentadvocacy-developmentawareness-raisingeducation-and-traininginformation-and-communicationknowledge-managementlearning-and-innovationresearch-and-developmentextension-and-advisorymarket-and-value-chainpolicy-and-governancefinance-and-investmentinfrastructure-and-technologyinstitutions-and-organizationshuman-resources-and-capacitiessocial-capital-and-networksnatural-capital-and-ecosystemsphysical-capital-and-infrastructurefinancial-capital-and-marketsintellectual-capital-and-knowledgecultural-capital-and-heritagepolitical-capital-and-governancesymbolic-capital-and-reputationassets-and-resourcescapabilities-and-competenciesopportunities-and-challengesdrivers-and-barriersenablers-and-constraintsincentives-and-disincentivesrisks-and-uncertaintiestrade-offs-and-synergiescosts-and-benefitsimpacts-and-outcomesinputs-and-outputsactivities-and-processesmechanisms-and-pathwayscontexts-and-conditionsscales-and-levelsdimensions-and-perspectivesactors-and-stakeholdersinterests-and-valuespower-and-politicsnarratives-and-discoursesframes-and-lensesmodels-and-theoriesconcepts-and-constructsparadigms-and-worldviewsepistemologies-and-ontologiesmethodologies-and-methodstools-and-techniquesapproaches-and-strategiestactics-and-operationsplans-and-programsprojects-and-interventionsinitiatives-and-activitiesactions-and-measurespolicies-and-regulationslaws-and-standardsguidelines-and-protocolsbest-practiceslessons-learnedsuccess-storiesfailure-storiescase-studiespilotsdemonstrationsexperimentstrialsevaluationsassessmentsreviewssynthesesmeta-analysessystematic-reviewsevidence-mapsknowledge-gapsresearch-prioritiesagenda-settingfundingresource-mobilizationallocationutilizationaccountabilitytransparencyintegrityresponsibilityadaptationmitigationtransformationimprovementprogressgrowthchangecontinuitystabilitydisruptionemergencecomplexityuncertaintyambiguitypluralityinclusionfairnesscaresolidaritycooperationcompetitionconflictconsensusdissentdeliberationdialoguedebatediscussionnegotiationmediationarbitrationadjudicationlitigationlegislationregulationimplementationenforcementcompliancereportingverificationcertificationaccreditationstandardizationharmonizationintegrationcoordinationcoherenceconsistencycompatibilitycomplementaritysynergyalignmentfitmatchbalancetrade-offoptimizationmaximizationminimizationsatisfactionsufficiencyeffectivenessefficacyperformanceproductivityprofitabilityviabilityfeasibilitydesirabilityacceptabilitylegitimacycredibilityreliabilityqualityexcellencedistinctionrecognitionreputationprestigeinfluenceimpactlegacyheritagetraditioncultureidentitybelongingattachmentcommitmentengagementparticipationinvolvementownershipempowermentautonomyagencycapacitycapabilitycompetenceskillexpertiseknowledgewisdominsightunderstandingawarenessconsciousnessmindfulnessattentionfocusconcentrationintentionpurposemeaningsignificanceworthmeritvirtuegoodnessbeautytruthgoodrightjustfairequitablesustainableinnovativecreativecriticalreflectiveresponsibleaccountabletransparentethicalintegralholisticsystemicdynamiccomplexuncertainambiguousdiversepluralcollaborativecooperativecompetitiveconflictingconsensualdeliberativedialogicaldemocraticautocratichierarchicalnetworkeddecentralizeddistributedlocalizedglobalizedstandardizedharmonizedintegratedcoordinatedalignedbalancedoptimizedsatisficingsufficientefficienteffectiveefficaciousperformantproductiveprofitableviablefeasibledesirableacceptablelegitimatecrediblereliablevalidhigh-qualityexcellentdistinctiverecognizedreputableprestigiousinfluentialimpactfulenduringmeaningfulsignificantvaluableworthymeritoriousvirtuousbeautifultruemorallegalauthorizedapprovedsanctionedendorsedsupportedfundedresourcedstaffedequippedenabledempoweredcapacitatedskilledtrainededucatedinformedawareconsciousmindfulattentivefocusedintentionalpurposeful