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Dielis tolteca
Toltec scoliid wasp
Dielis tolteca is a species of scoliid wasp native to western North America and Mesoamerica. The species is known to parasitize scarab beetle grubs, with females hunting underground hosts to provision their offspring. Adults have been observed feeding on flowering plants, particularly mustards (Brassicaceae) and goldenrods (Solidago). The species has been documented in McInnis Canyons National Conservation Area in western Colorado, where it is active as a spring-emerging species.
scoliid-wasp parasitoid scarab-beetle western-Colorado McInnis-Canyons spring-emergence Solidago mustards biological-control Hymenoptera Scoliidae Dielis Campsomerini biodiversity-survey integrated-pest-management melissa-schreiner extension-entomology saussure-1857 toltec-scoliid-wasp riparian Colorado-River flower-visitor nectar-feeding solitary-wasp ground-nesting-parasitoid scarab-grub-parasitoid aculeate apocrita aculeata campsomerinae western-North-America Nearctic Neotropical Mesoamerica Arizona Baja-California Sonora underground-host-hunter external-parasitoid larval-provisioning spring-active insect-biodiversity rare-ecosystem-survey natural-heritage-survey CSU-extension Colorado-State-University agricultural-pest-management tree-fruit wine-grapes row-crops invasive-species-monitoring Japanese-beetle Popillia-japonica corn-earworm Helicoverpa-zea IPM pest-monitoring moth-trapping B3-allele-testing citizen-science Department-of-Agriculture-trapping targeted-larval-treatments 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insects-second insect-samples precise-identifications research-sites commercial-farms orchards onsite-investigations informed-solutions stone-fruit-orchard unexpected-pest-species Cooperative-Agricultural-Pest-Survey rapid-response invasive-threats pest-management-professionals public-health-officials food-safety-experts cafeterias processing-facilities rural-library high-alpine-public-lands outreach-programs scientific-knowledge practical-solutions student-educational-events continuing-education-workshops licensed-commercial-pesticide-applicators February-workshop hundreds-of-applicators wide-range-of-disciplines educational-programming community-collaboration complex-ecological-challenges complex-agricultural-challenges unique-extension-program predominantly-IPM rewarding-work creative-illustration insect-biology complex-scientific-concepts engaging-education accessible-education statewide-Colorado-Master-Gardener-Program hundreds-of-visuals gardeners insect-pests enhanced-adult-learning empowered-master-gardeners insect-related-inquiries education-and-practical-solutions fulfilling-work narrowing-focus greatest-challenge needs-everywhere insects-know-no-county-lines large-area-to-cover not-enough-personnel early-career-excitement saying-yes-to-projects tempting-to-dive-in all-kinds-of-insect-related-work pest-management-every-crop biodiversity-surveys public-education delegation successful-funding-strategies more-personnel prioritize-self-care tough-lesson efficiency impactful-work focus-on-key-areas skills-and-interests spreading-too-thin compromise-quality affect-well-being narrowing-areas-of-focus thoughtful-targeted-support healthier-work-life-balance meaningful-work not-just-widespread save-the-world-or-savor-the-world balance-concepts healthy-career deliberately-live-life-outside-entomology family-time friends-time public-lands-time individual-values cultural-values traditions solo-traveling comfort-zone habits routines old-friends 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Dr.-Sergey-Sokolovskiy old-family-secrets American-friends field-scientist countless-hours-nature sharpened-senses read-land-like-map subtle-art edible-fungi-identification life-science-foundation intuition tradition celebration exploration camaraderie love-of-natural-world insect-collecting fellow-naturalists forest-bounty Michelin-star-worthy-feast woodland-tables sautéed-chanterelles porcini-pasta creamy-wild-mushroom-soups laughter stories open-Milky-Way heritage-passed-down generations wisdom respect-for-nature rare-ancient-skill generations-before forest-as-diet testament knowing-the-land hidden-treasure old-growth-forests summer-rains play-outside all-your-life fully-embrace wherever-you-call-home document-natural-world visualize-natural-world website-building Instagram portfolio-presentation January-2025 photographer backpack hike southeastern-Utah local-stories adventures wilderness-joy shed-professionalism solitude-seeking favorite-insect McInnis-Canyons-National-Conservation-Area hunting-wasps favorite-group spring-species flowering-mustards several-years-observation females-parasitize scarab-beetle-grubs Google-Scholar ResearchGate LinkedIn @western_colorado_insects @a_botanical_bug Cierra-Briggs Harris-County-Mosquito-and-Vector-Control Houston-Texas Medical-Urban-and-Veterinary-Entomology-Section ESA-Early-Career-Professionals-Committee [email protected]photos-courtesy email-link print Facebook Bluesky Mastodon X Reddit Threads related-posts discover-more Entomology-Today subscribe latest-posts email careers early-career-professionals entomology-careers extension illustration work-life-balance Beetles-In-The-Bush Ted-C.-MacRae Art-Evans What's-Bugging-You photograph 10-years-ago difficult-challenge guesses identity location past-months digging key-to-identity order-gimme 2-points family-difficult 4-points taxonomic-changes-hint genus-challenge 6-points online-resources species-name-impossible short-list described-species general-area bonus-points additional-pictures answer-comment couple-days Orthoptera Stenopelmatidae Stenopelmatus North-American Oklahoma fuscus dark Anostostoma genus change-of-heart family old-world morphological-similarities king-cricket head legs antennae postnotum Jerusalem-cricket locality species-short-list variable photograph-10-years-ago Russia-adjacent-country tags Mexico New-Zealand South-Africa Hemiandrus stumpy-hinds pdf-resource Bochus spineyness genicular-lobes conservation-biologist work-in-progress entomologist Nasidius genae cheek lower-genae head-modification mandible-enlargement normal-looking-head female male-without-modifications Libanasidus-vittatus one-spine inner-margin foretibia two-spines thoracic-abdominal-tergites black-posterior-margin Onosandrus-sp dissertation color-variable not-diagnostic eight-genera king-crickets male-Onosandridus-sp couplet-1 tympanum-not-obvious fore-tibia couplet-2 two-spines-inner-margin couplet-3 no-mandible-enlargement ovipositor-not-long couplet-5 no-large-ovipositor male smooth-face not-Bochus Onosandridus key Review-of-southern-African-Anostostomatidae Brettschneider hind-femur-never-armed spines-hooks males-no-head-modification Sam-Heads Orthoptera-expert Illinois-Natural-History-Survey Anostostomatidae genus-Onosandridus-Péringuey two-impressive-spines inner-surface-protibia Bochus-characteristic head-face-tuberculate former-genus specimen-clearly-not Heath Jason correct-genus Heath-first points Peter Chris final-standings Ziad-Khouri Zeroing-in-on-Mammoth-Wasps Scoliid-Wasps Bug-Squad UC-Davis-doctoral-candidate Lynn-Kimsey major-professor distinguished-professor seminar UC-Davis-Department-of-Entomology-and-Nematology Wednesday-March-30 spring-quarter-seminars 4:10-p.m.Pacific-Daylight-Time 122-Briggs-Hall Zoom unique-work modern-systematic-treatment genus-species-level-taxonomy-mess modern-classical-techniques taxonomic-order evolution-discovered Bohart-Museum eight-million-insect-specimens 2300-mammoth-wasp-specimens Americas Korea two-parts Evolutionary-History-of-Mammoth-Wasps Comparing-Power-of-Data-Based-Phylogenetic-Posterior-Predictive-Checks Cucleotide-Amino-Acid-Data abstracts Part-1 aculeate-insects larvae-parasitoids scarabaeid-beetle-grubs biological-control-agents group-evolution stability-taxonomy reliable-phylogenies-limited ultraconserved-element-UCE-data concatenation multispecies-coalescent phylogeny-Scoliidae mitigate-model-misspecification data-filtering-experiments posterior-predictive-checks matched-pairs-tests-symmetry Proscolia-sister all-other-extant-scoliids strong-support sister-group-relationship campsomerine-genus-Colpa Scoliini Campsomerini-non-monophyletic Campsomerini-sensu-stricto monophyletic Australasian-genus-Trisciloa sister-remaining-members sampled-genera-non-monophyletic Campsomeriella Megascolia Scolia fossil-data Early-Cretaceous-origin crown-Scoliidae split-Scoliini-Colpa Campsomerini-s.s.Late-Cretaceous posterior-means crown-ages Paleogene age-95%-HPD-intervals K-Pg-boundary fossils-less-certain-placement older-ages stem-ages Nearctic-scoliid-clades Beringia-dispersal Oligocene-later-Eocene foundation-future-research scoliid-wasp-evolution-biogeography first-genome-scale-data model-based-methods precision-dating-analyses paucity-well-preserved-fossils reliably-attributable crown-group higher-level-taxonomy-dire-need-revision taxonomic-changes-predicated datasets-extend-geographic-taxonomic-sampling Part-II phylogenetic-inference exonic-DNA-sequences multiple-codings nucleotides amino-acids codons empirical-studies data-type-choice model-choice less-expected-violation inaccurate-inference assessing-phylogenetic-model-adequacy inference-reliability-indication simulation-based-approach detect-model-inadequacy phylogenetic-posterior-prediction data-coding-variation simulated-data-multiple-models codon-models process-heterogeneity-lineages selection-heterogeneity-sites codon-usage-selection inference-posterior-predictive-checks nucleotide-amino-acid-models GTR-family significant-differences amino-acid-nucleotide-treatments detect-model-violation magnitude-error-estimate-interest-similar corroborate-other-studies tree-length-estimation-error topology-reconstruction-error not-always-correlated amino-acid-models more-accurate-topologies tree-length-errors-greater nucleotide-models branch-heterogeneous-codon-models magnitude-direction data-coding-dependence data-generating-process-properties posterior-predictive-checks-data-filtering practical-effect-size-thresholds low-inference-reliability established-separately amino-acid-nucleotide-data caution-advised careful-model-selection data-coding-careful-selection accurate-tree-length-inference-important 2012-graduate-program doctorate-entomology dissertation-research phylogenetics-evolution-taxonomy-Scolliidae bachelor's-degree-biology 2012 Notre-Dame-University-Louaize-NDU undergraduate-research bee-pollinator-identification American-University-Beirut genetic-diversity-assessment Spartium-junceum-populations Spanish-broom rush-broom weaver's-broom insect-collection-establishment NDU-specimen-collection UC-Davis-team national-Linnaean-Games 2015-Entomological-Society-of-America-meeting Entomology-Games University-of-Florida 32-year-history-first-win video Bohart-Museum-location Room-1124-Academic-Surge-Building Crocker-Lane currently-closed-public live-petting-zoo Madagascar-hissing-cockroaches stick-insects tarantulas insect-themed-gift-shop online Shahid-Siddique assistant-professor seminar-coordinator Zoom-technical-issues [email protected]seminar-list Trisciloa-saussurei New-Guinea-native Bohart-Museum-specimen Bug-Eric Wasp-Wednesday Campsomeris December-22-2010 mystery-wasp December email-friend species-recorded-recently-Arizona Mexican-border Sabino-Canyon-Recreation-Area Fred-Heath outstanding-naturalist Introduction-to-Southern-California-Butterflies Sabino-Canyon-Volunteer-Naturalists December-14 male-specimen Campsomeris-ephippium Desert-Lavender Hyptis-emoryi conspicuous-distinctive no-image-needed email-list-message David-Lazaroff SCVN-founder camera image-permission long-antennae slender-body pseudostinger posterior-abdomen gender-reveal robust-females shorter-antennae real-stinger retractable-stinger hosts-larval-offspring spring-2009-image south-Texas south-to-Ecuador Sunday-December-19 failed-to-find common-local-species Campsomeris-tolteca males-feeding nectar-Coreocarpus-arizonicus Little-Lemonhead Queen-butterflies Mexican-Yellows Scoliidae-family all-parasitoids parasitoid-definition heavy-spiny-legs dig-up-scarab-grub sting-brief-paralysis single-egg host-larva leave-scene host-regains-consciousness motor-skills underground-existence plant-roots-feeding wasp-egg-hatches external-parasite beetle-grub pleasure-of-hunt Sabino-Canyon Fred-Heath-outdoors David-Lazaroff-image 10:00-AM Email-ThisBlogThisShare-to-XShare-to-FacebookShare-to-Pinterest animals bugs insects nature wasps wildlife comments great-story beautiful-wasps learn-something envy-ability outside-warm-weather impending-snow bug-hunt-hope two-blogs Mosquito-Hawk same-Lemonhead-bush world-shrinks-December Tumacacori-NHP last-week-sighting photo-contact more-sightings-head's-up second-photo-species uncertain Arizona-couple-species specimen-in-hand-needed Beatty's-Guest-Ranch Miller-Canyon apple-trees-pollinating last-weekend blog-author-unable-reply working-to-resolve newer-post older-post home subscribe-post-comments atom GBIF-taxonomy-match accepted-status exact-match Animalia-Arthropoda-Insecta-Hymenoptera-Scoliidae-Dielis distribution-records NA-NT Mexico-states Baja-California-Sur Campeche Chiapas Coahuila Guanajuato Guerrero Jalisco Michoacán Morelos Oaxaca Quintana-Roo Sinaloa Tabasco Veracruz El-Salvador-San-Salvador Guatemala-Huehuetenango Honduras-Comayagua Nicaragua-Rivas Map-3 Haiti United-States Bradley-1828 Hurd-1952 Porter-1981 MacKay-1987 iNaturalist-taxon 2987-observations preferred-common-name Wikipedia-summary Solidago-plants iNaturalist-taxonomy Grasshoppers-of-Colorado Grasshoppers-of-Wyoming-and-the-West Entomology grasshopper-names genus-species common-scientific species-genus abdominalis-Chloealtis admirabilis-Syrbula agrestis-Trimerotropis alba-Hypochlora alpinus-Ceuthophilus alpinus-Melanoplus alutacea-Schistocerca angustipennis-Melanoplus apiculata-Pardalophora argentinus-Oecanthus arizonae-Melanoplus aspera-Trachyrhachys bicolor-Dactylotum bispinosus-Melanoplus bivittata-Mermiria bivittatus-Melanoplus bolli-Spharagemon borealis-Melanoplus bowditchi-Melanoplus brachyptera-Pseudopomala brevipes-Daihnia bruneri-Melanoplus brunneus-Stenobothrus californica-Trimerotropis californicus-Oecanthus campestris-Trimerotropis capito-Hippopedon carlinianus-Circotettix carolina-Dissosteira chenopodii-Aeoloplides cincta-Trimerotropis citrina-Trimerotropis clavatus-Aeropedellus collare-Spharagemon coloradus-Amphitornus complanatipes-Melanoplus confusus-Melanoplus conspersa-Arphia conspersa-Chloealtis corallipes-Xanthippus coronata-Trachyrhachys crenulata-Cordillacris curtipennis-Chorthippus cyaneipennis-Trimerotropis cyaneus-Leprus dawsonii-Melanoplus delicatula-Psoloessa deorum-Ageneotettix differentialis-Melanoplus discolor-Melanoplus dodgei-Melanoplus elliotti-Aulocara enigma-Oedaloenotus equale-Spharagemon fasciatus-Melanoplus femoratum-Aulocara femurrubrum-Melanoplus flavidus-Melanoplus foedus-Melanoplus fontana-Trimerotropis formosus-Tropidolophus fratercula-Trimerotropis fusiformis-Ceuthophilus gladstoni-Melanoplus glaucipes-Melanoplus gracile-Stethophyma gracilis-Trimerotropis haldemanii-Pardalophora haydeni-Derotmema hirtipes-Acrolophitus humile-Spharagemon inconspicua-Trimerotropis infantilis-Melanoplus keeleri-Melanoplus kennicotti-Melanoplus kiowa-Trachyrhachys lakinus-Melanoplus latifasciata-Trimerotropis longipennis-Dissosteira magna-Brachystola magnifica-Trimerotropis melanoptera-Trimerotropis montanus-Xanthippus montezuma-Syrbula nebrascensis-Phoetaliotes neglectus-Cratypedes nevadensis-Acrolophitus niveus-Oecanthus nubilum-Boopedon obscura-Opeia occidentalis-Melanoplus occipitalis-Cordillacris ocelote-Hippiscus olivacea-Campylacantha oregonensis-Melanoplus packardii-Melanoplus pallidipennis-Trimerotropis pardalinus-Metator parviceps-Cibolacris pelidna-Orphulella pellucida-Camnula picta-Mermiria pistrinaria-Trimerotropis plattei-Mestobregma ponderosus-Melanoplus pseudonietana-Arphia quadrimaculatum-Phlibostroma quadripunctatus-Oecanthus rabula-Circotettix regalis-Melanoplus robusta-Udeopsylla robustus-Leprus rufa-Heliaula sanguinipes-Melanoplus simplex-Anabrus simplex-Arphia simplex-Eritettix sordidus-Encoptolophus sparsa-Trimerotropis speciosa-Orphulella speciosus-Hesperotettix splendidus-Melanoplus spretus-Melanoplus spurcata-Dissosteira subgracilis-Encoptolophus sulcifrons-Conozoa tenuipennis-Aeoloplides texana-Conozoa texana-Mermiria texana-Psoloessa tolteca-Trimerotropis trifasciatus-Hadrotettix tristis-Melanoplus turnbulli-Aeoloplides utahensis-Ceuthophilus verruculatus-Trimerotropis virgata-Paropomala viridifasciata-Chortophaga viridis-Hesperotettix wyomingensis-Paropomala xanthoptera-Arphia yarrowii-Melanoplus not-true-grasshoppers Acrididae discussed-manual confused-with-grasshoppers top-of-page back-to-contents navigation-main-page learn-more biology distribution-maps fact-sheets key-to-stages grasshopper-development grasshopper-names-common-scientific grasshopper-names-species-genus grasshopper-names-genus-species Acrolophitus-hirtipes Acrolophitus-nevadensis Aeoloplides-chenopodii Aeoloplides-tenuipennis Aeoloplides-tumbulli Aeropedellus-clavatus Ageneotettix-deorum Amphitomus-coloradus Anabrus-simplex Arphia-conspersa Arphia-pseudonietana Arphia-simplex Arphia-xanthoptera Aulocara-elliotti Aulocara-femoratum Boopedon-nubilum Brachystola-magna Camnula-pellucida Campylacantha-olivacea Ceuthophilus-alpinus Ceuthophilus-fusiformis Ceuthophilus-utahensis Chloealtis-abdominalis Chloealtis-conspersa Chorthippus-curtipennis Chortophaga-viridifasciata Cibolacris-parviceps Circotettix-carlinianus Circotettix-rabula Conozoa-suicifrons Conozoa-texana Cordillacris-crenulata Cordillacris-occipitalis Cratypedes-neglectus Dactylotum-bicolor Daihnia-brevipes Derotmema-haydeni Dissosteira-carolina Dissosteira-longipennis Dissosteira-spurcata Encoptolophus-sordidus Encoptolophus-subgracilis Eritettix-simplex Gryllus-spp Hadrotettix-trifasciatus Heliaula-rufa Hesperotettix-speciosus Hesperotettix-viridis Hippiscus-ocelote Hippopedon-capito Hypochlora-alba Leprus-cyaneus Leprus-robustus Melanoplus-alpinus Melanoplus-angustipennis Melanoplus-arizonae Melanoplus-bispinosus Melanoplus-bivittatus Melanoplus-borealis Melanoplus-bowditchi Melanoplus-bruneri Melanoplus-complanatipes Melanoplus-confusus Melanoplus-dawsonii Melanoplus-differentialis Melanoplus-discolor Melanoplus-dodgei Melanoplus-fasciatus Melanoplus-femurrubrum Melanoplus-flavidus Melanoplus-foedus Melanoplus-gladstoni Melanoplus-glaucipes Melanoplus-infantilis Melanoplus-keeleri Melanoplus-kennicotti Melanoplus-lakinus Melanoplus-occidentalis Melanoplus-oregonensis Melanoplus-packardii Melanoplus-ponderosus Melanoplus-regalis Melanoplus-sanguinipes Melanoplus-splendidus Melanoplus-spretus Melanoplus-tristis Melanoplus-yarrowii Mermiria-bivittata Mermiria-picta Mermiria-texana Mestobregma-plattei Metator-pardalinus Oecanthus-argentinus Oecanthus-californicus Oecanthus-niveus Oecanthus-quadripunctatus Oedaloenotus-enigma Opeia-obscura Orphulella-pelidna Orphulella-speciosa Pardalophora-apiculata Pardalophora-haidemanii Paropomala-virgata Paropomala-wyomingensis Phlibostroma-quadrimaculatum Phoetaliotes-nebrascensis Pseudopomala-brachyptera Psoloessa-delicatula Psoloessa-texana Schistocerca-alutacea Spharagemon-bolli Spharagemon-collare Spharagemon-equale Spharagemon-humile Stenobothrus-brunneus Stethophyma-gracile Syrbula-admirabilis Syrbula-montezuma Trachyrhachys-aspera Trachyrhachys-coronata Trachyrhachys-kiowa Trimerotropis-agrestis Trimerotropis-californica Trimerotropis-campestris Trimerotropis-cincta Trimerotropis-citrina Trimerotropis-cyaneipennis Trimerotropis-fontana Trimerotropis-fratercula Trimerotropis-gracilis Trimerotropis-inconspicua Trimerotropis-latifasciata Trimerotropis-magnifica Trimerotropis-melanoptera Trimerotropis-pallidipennis Trimerotropis-pistrinaria Trimerotropis-sparsa Trimerotropis-tolteca Trimerotropis-verruculatus Tropidolophus-formosus Udeopsylla-robusta Xanthippus-corallipes Xanthippus-montanus true-grasshoppers Acrididae-discussed manual-confused grasshoppers-top page-back contents-navigation main-page learn-more-biology distribution-maps-fact sheets-key stages-grasshopper development-grasshoppers Colorado-grasshoppers Wyoming-West Entomology-GRASSHOPPERS COLORADO-GRASSHOPPER NAMES-GENUS SPECIES-GRASSHOPPER NAMES-COMMON SCIENTIFIC-GRASSHOPPER NAMES-SPECIES GENUS-abdominalis Chloealtis-Thomas Uhler-agrestis Trimerotropis-McNeill Dodge-alpinus Ceuthophilus-Scudder Scudder-alutacea Schistocerca-Scudder Dodge-apiculata Pardalophora-Harris Saussure-arizonae Melanoplus-Scudder Scudder-bicolor Dactylotum-Thomas Scudder-bivittata Mermiria-Serville Say-bolli Spharagemon-Morse Fieber-bowditchi Scudder-brevipes Daihnia-Haldeman Scudder-brunneus Stenobothrus-Thomas Bruner-californicus Oecanthus-Saussure McNeill-capito Hippopedon-Stal Thomas-carolina Dissosteira-L Bruner-cincta Trimerotropis-Thomas Scudder-clavatus Aeropedellus-Thomas Scudder-coloradus Amphitornus-Thomas Scudder-confusus Scudder-conspersa Chloealtis-Harris Haldeman-coronata Trachyrhachys-Scudder Bruner-curtipennis Chorthippus-Harris Bruner-cyaneus Leprus-Cockerell Scudder-delicatula Psoloessa-Scudder Scudder-differentialis Melanoplus-Thomas Scudder-dodgei Thomas-enigma Oedaloenotus-Scudder Say-fasciatus Melanoplus-F-Walker Scudder-femurrubrum Melanoplus-DeGeer Scudder-foedus Thomas-formosus Tropidolophus-Say McNeill-fusiformis Scudder-glaucipes Scudder-gracilis Scudder-haydeni Derotmema-Thomas Say-humile Bruner-infantilis Thomas-kennicotti Thomas-lakinus Scudder-longipennis Dissosteira-Thomas Girard-magnifica Trimerotropis-Rehn McNeill-montanus Xanthippus-Thomas Saussure-nebrascensis Phoetaliotes-Thomas Thomas-nevadensis Acrolophitus-Thomas DeGeer-nubilum Boopedon-Say Thomas-occidentalis Thomas-ocelote Hippiscus-Saussure Scudder-oregonensis Scudder-pallidipennis Trimerotropis-Burmeister Saussure-parviceps Cibolacris-Bruner Burmeister-pellucida Camnula-Scudder Walker-pistrinaria Trimerotropis-Saussure Thomas-ponderosus Thomas-quadrimaculatum Phlibostroma-Thomas Beutenmuller-rabula Circotettix-Rehn Hebard-regales Melanoplus-Dodge Haldeman-robustus Leprus-Hebard Scudder-sanguinipes Melanoplus-Fabricius Haldeman-simplex Arphia-Scudder Scudder-sordidus Encoptolophus-Burmeister Thomas-speciosa Orphulella-Scudder Scudder-splendidus Melanoplus-Hebard Walsh-spurcata Dissosteira-Saussure Caudell-sulcifrons Conozoa-Scudder Scudder-texana Conozoa-Bruner Bruner-texana Brunner-trifasciatus Hadrotettix-Say Bruner-turnbulli Aeoloplides-Caudell Thomas-verruculatus Trimerotropis-Scudder Scudder-viridifasciata Chortophaga-DeGeer Scudder-wyomingensis Paropomala-Thomas Burmeister-yarrowii species-not-true grasshoppers-Acrididae discussed-manual-because top-of-the page-back-to Contents-Navigation-Main Page-Grasshoppers-of Colorado-LEARN-MORE Biology-Distribution-Maps Fact-Sheets-Key to-Stages-of Grasshopper-Development-Biology Distribution-Maps-LEARN MORE-Distribution-Maps Fact-Sheets-LEARN MORE-Grasshoppers-of Colorado-Fact-Sheets of-Grasshopper-Development LEARN-MORE-Grasshoppers
Microterys
Microterys is a large genus of parasitoid wasps in the family Encyrtidae (Chalcidoidea), with its center of distribution in the northern parts of the Northern Hemisphere. Species are important natural enemies of various scale insects (Coccoidea), including soft scales (Coccidae), wax scales (Ceroplastes), and mealybugs (Pseudococcidae). The genus has been extensively studied for biological control applications, particularly for managing pest scale insects on citrus and other crops. Several species have been introduced to new regions as biocontrol agents, including Microterys flavus in California.
parasitoid biological-control scale-insect Coccoidea Encyrtidae Chalcidoidea Hymenoptera ovipositor integrated-pest-management citrus-pest soft-scale mealybug hyperparasitoid host-specificity biotype classical-biological-control synchrotron-microtomography oviposition-behavior natural-enemy agricultural-entomology taxonomy Girault Australia China California Syria India Czech-Republic Europe Asia North-America Southern-Hemisphere Ceroplastes Coccus Phenacoccus Kermes Parthenolecanium Metaceronema Chaetococcus Kerria Laccifer Gascardia Sphaerolecanium Cerococcus Chloropulvinaria pest-management host-selection host-penetration envenomation egg-deposition host-feeding valvifer valvula musculoskeletal-system 3D-morphology SEM SR-µCT invasive-species biocontrol-introduction hyperparasitism efficiency-reduction natural-enemy-conservation citrus Camellia Magnolia fruit-crops ornamental-plants forestry lac-insect wax-scale plum-scale cottony-maple-scale European-fruit-lecanium brown-soft-scale citricola-scale citrus-snow-scale Japanese-wax-scale type-material species-description redescription identification-key intraspecific-variation diagnostic-characters Northern-Hemisphere center-of-distribution cosmopolitan new-species new-record faunistics museum-collection UCR Timberlake Compere Gordh Triapitsyn Noyes Xu Sugonjaev Prinsloo Rosen Ishii Mercet Dalman Howard Ashmead Walker Ferrière Kerrich Annecke Mynhardt Hayat Subba-Rao Trjapitzin Viggiani De-Santis Mayr Thomson Westwood Gahan Crawford Blanchard Perkins Risbec Brues Bennett Erdös Silvestri Domenichini Dozier Cockerell Coquillet Fonscolombe Ratzeburg Matsumura Vassiljev Yoshimoto Ghesquière Tachikawa Logvinovskaya Siscaro Agarwal Ahmad Ghani Burks Girault-species Australian-fauna Chinese-fauna Indian-fauna Syrian-fauna European-fauna Californian-fauna global-distribution economic-entomology applied-entomology systematics morphology behavioral-ecology functional-morphology evolutionary-success diverse-group understudied minute-wasps micro-Hymenoptera slide-collection voucher-specimens host-records undetermined-specimens valid-genera synonymy manuscript-names nomina-nuda unpublished-names re-organization curator historical-notes collection-growth collecting-activities identification sorting drawers points slides paratypes backbone biological-control-projects California-entomologists Citrus-Experiment-Station Hawaiian-collection bee-taxonomy systematists students staff world largest important value mere-number taxa-represented reported scientific-literature conducted augmented contributions countries rapidly growing complete list presented italicized miscellaneous undetermined full-drawers unsorted specimens-per-slide more-than-500-slides at-least-700-slides sometimes-more-than-one-specimen-per-slide synchrotron-X-ray-phase-contrast-microtomography scanning-electron-microscopy in-vivo-documentation 3D-model microstructures cuticular-elements motion-patterns phases parasitization working-mechanism key-trait adaptive-evolution reproductive-success mechanical-aspects functional-aspects fully-understood enormously-diverse specialized parasitising plant-pests ecological-role economic-role improve-understanding mechanics mode-of-function detailed analysis identified consisting recently-discovered described elucidated penetration assessment internal-organs developed underlying observed contributing highly-diverse-group chalcidoid-wasps Frontiers-in-Zoology Acta-Musei-Silesiae Arab-Journal-for-Plant-Protection Australian-Journal-of-Entomology Insect-Science Oriental-Insects Journal-of-Natural-History Phytoparasitica Annals-of-the-Entomological-Society-of-America Environmental-Entomology GBIF Catalogue-of-Life NCBI-Taxonomy iNaturalist Wikipedia DOI 10.1186/s12983-025-00575-1 10.2478/cszma-2021-0007 10.22268/AJPP-41.3.292305 10.1111/j.1440-6055.1975.tb02058.x 10.1111/j.1744-7917.2000.tb00345.x 10.1080/00305316.2011.646827 10.1111/j.1440-6055.1973.tb01670.x 10.1080/00222933.2020.1786608 10.1007/s12600-019-00727-0 10.1007/s12600-018-0691-5 10.1093/aesa/54.2.222 10.1093/ee/7.6.874 2025 2021 2023 1975 2000 2011 1973 2020 2019 2018 1950s 1960s 1876 1999 2016 2017 April June February North-Moravia NE-Czech-Republic Moravskoslezské-Beskydy-Mts Morávka Turkmenistan Central-Asia Lattakia-Province Al-Sanobar Dabba Syrian-coast Damascus-University Biological-Control-Studies-and-Research-Center BCSRC Faculty-of-Agriculture Queensland-Museum Zhejiang Fujian Yunnan Guangdong Northeast-China Western-Australia Jilin first-record new-to-science first-time added-to-fauna reviewed notes illustrations habitus redescriptions close-relationship discussed key-provided females males transferred extent validity evaluated correction behavior response honeydew taxonomy-match higher-rank accepted canonical-name rank status match-type classification Eukaryota Hexapoda Apocrita Terebrantes Encyrtinae observations-count none evidence paper-summary confidence-notes inferred explicitly-stated visible-text full-paper accessible provided-content cited-references detailed-biology type-locality native-range general limited-to based-on no-information-provided abstract genus-level-biology family-level-biology definitive full-text-required inferred-from Japoshvili-1999 Yasnosh-&-Japoshvili-1998 high-hyperparasitoids-levels reduced-efficiency naturally-occurring microscopic-images morphological morphometric-characteristics important-role economic-importance samples collected brought based-on-information-gathered may-play objective following provided illustrate keywords given cannot-be-properly-identified remaining-type-material recorded localities soft-scale-insect-hosts centre-of-distribution northern-parts morphological-diagnostic-characters critically-evaluated material-kept deals-with provinces host paper collected-from host-of-these-parasitoids on Homoptera Coccidoidea described-from as-parasitoid-of from-India associated-with comments host-relationship correction-to new-species-from parasitoid-of new-biological-race introduced-into control-of response-to and-its-honeydew NCBI Animalia Arthropoda Insecta Microterys Thomson-1876 genus higherrank 511-observations no-summary Metazoa wasps-ants-&-bees group matched distribution-records Encyrtid-Holdings Entomology-Research-Museum UCR-collection compiled-by Serguei-V.-Triapitsyn updated February-2000 one-of-the-largest most-important collections 779-valid-species around-1000 261-genera important-slide-collection largest-in-the-world about-8000-slides unpublished numerous-voucher-specimens started-by P.H.-Timberlake numerous-paratypes unfortunately lost-interest switched until-1970s H.-Compere notable-contributions P.-Timberlake J.-Maple P.-DeBach C.-Clausen 1970s-1980s G.-Gordh J.-LaSalle J.-Woolley J.-Pinto J.-Beardsley Mexico US other-countries presently growing-rapidly J.-Heraty majority V.-Trjapitzin J.-Noyes complete-re-organization S.-Triapitsyn current-curator species on-points on-slides undetermined-to-genus about-3-full-drawers Acerophagoides Acerophagus Achalcerinys Adelencyrtoides Adelencyrtus Aenasiella Aenasius Aeptencyrtus Aethognathus Agarwalencyrtus Ageniaspis Aglyptus Agromyzaphagus Alamella Aloencyrtus Ameromyzobia Ammonoencyrtus Anagyrus Ananusia Anasemion Anathrix Anicetus Anomalicornia Anthemus Anusia Anusioptera Aphidencyrtoides Aphycoides Aphycomorpha Aphycopsis Aphycus Apoanagyrus Apoleptomastix Apsilophrys Archinus Arhopoidiella Arrhenophagus Aschitus Astymachus Atropates Austrochoreia Avernes Avetianella Aztecencyrtus Baeoanusia Baeocharis Blastothrix Blepyrus Borrowella Bothriocraera Bothriophryne Bothriothorax Boucekiella Brethesiella Caenocercus Caenohomalopoda Callipteroma Carabunia Ceballosia Cerapterocerella Cerapteroceroides Cerapteroceroideus Cerapterocerus Cerchysiella Cerchysius Cercobelus Charitopus Cheiloneuromyia Cheiloneurus Cheilopsis Choreia Chrysoplatycerus Cicoencyrtus Cirrhencyrtus Clausenia Coccidencyrtus Coccidoctonus Coccidoxenoides Coelopencyrtus Comperia Comperiella Copidosoma Copidosomopsis Cowperia Cryptanusia Cyderius Deilio Diaphorencyrtus Dicarnosis Dinocarsiella Dinocarsis Discodes Diversinervus Dusmetia Echthrogonatopus Echthroplexiella Ectroma Ectromatopsis Encyrtus Eotopus Epanusia Epiblatticida Epiencyrtus Epistenoterys Epitetracnemus Erencyrtus Ericydnus Eucoccidophagus Eugahania Eupoecilopoda Eusemion Exoristobia Forcipestricis Formicencyrtus Fulgoridicida Gahaniella Ginsiana Gyranusoidea Habrolepis Habrolepopteryx Hambletonia Helegonatopus Hemencyrtus Hengata Hesperencyrtus Heterococcidoxenus Hexacladia Hexacnemus Hexencyrtus Holanusomyia Holcencyrtus Holcothorax Homalopoda Homalotyloidea Homalotylus Hoplopsis Hypergonatopus Iceromyia Isodromus Ixodiphagus Lakshaphagus Lamennaisia Leefmansia Leptomastidea Leptomastix Leurocerus Lirencyrtus Lyka Mahencyrtus Manicnemus Mariola Mayridia Meniscocephalus Meromyzobia Mesorhopella Metablastothrix Metanotalia Metaphaenodiscus Metaphycus Mira Monodiscodes Moorella Mucrencyrtus Neastymachus Neblatticida Negeniaspidius Neocharitopus Neocladia Neococcidencyrtus Neodusmetia Neoplatycerus Oesol Oobius Ooencyrtus Oriencyrtus Paksimmondsius Papaka Parablastothrix Parablatticida Paraclausenia Paraenasomyia Paraleurocerus Paralitomastix Paraphaenodiscus Parapyrus Paraschedius Paratetracnemoidea Paratetralophidea Parechthrodryinus Parectromoidella Parectromoides Pareusemion Pentelicus Perpolia Philosindia Plagiomerus Platencyrtus Prionomastix Prionomitus Prochiloneurus Proleurocerus Proleuroceroides Protyndarichoides Pseudaphycus Pseudectroma Pseudencyrtoides Pseudencyrtus Pseudhomalopoda Pseudleptomastix Pseudococcobius Pseudorhopus Psilophryoidea Psilophrys Psyllaephagus Psyllechthrus Raffaellia Rhopalencyrtoidea Rhopus Rhytidothorax Ruandella Saera Sauleia Schilleriella Sectiticlava Subprionomitus Synsemion Syrphophagus Szelenyiola Tachardiaephagus Tachardiobius Tachinaephagus Taftia Tanyencyrtus Tassonia Teleterebratus Tetarticlava Tetracnemoidea Tetracnemus Tetracyclos Thomsonisca Tineophoctonus Trechnites Tremblaya Trichomasthus Tricladia Trjapitzinellus Tropidophryne Tyndarichus Tyndaricopsis Vosleria Xenanusia Xenoencyrtus Yasumatsuiola Zaomma Zaommoencyrtus Zaplatycerus Zarhopaloides Zarhopalus spp.sp.sp(p).?nr.ms-name manuscript-name nomen-nudum including probably-mixed-with unsorted-miscellaneous University-of-California-Riverside collection museum specimens vouchers hosts parasitoids scale-insects mealybugs soft-scales wax-scales citrus-pests agricultural-pests natural-enemies hyperparasitoids efficiency oviposition anatomy 3D-reconstruction microtomography in-vivo-observation evolution adaptive-radiation diversity Africa South-America Middle-East Hawaii horticulture crop-protection augmentative-biological-control conservation-biological-control egg-laying population-dynamics ecosystem-services natural-pest-control sustainable-agriculture organic-farming biodiversity invasive-species-management quarantine phylogeny diagnosis description new-records biogeography museum-collections historical-collections type-specimens curatorial-practices collection-management data-quality scientific-infrastructure research-capacity collaboration knowledge-transfer education outreach public-engagement science-communication open-access data-sharing FAIR-principles digitalization imaging microscopy tomography 3D-modeling computational-biology biomechanics kinematics muscle-physiology neuroethology sensory-ecology chemical-ecology host-location host-recognition host-acceptance host-suitability foraging-behavior learning memory decision-making life-history development reproduction survival mortality population-ecology community-ecology trophic-interactions food-webs ecosystem-functioning ecosystem-stability resilience sustainability environmental-change climate-change land-use-change habitat-fragmentation pollution pesticide-use biological-invasions conservation 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Nomotettix cristatus
crested pygmy grasshopper, crested grouse locust, northern crested grouse locust
Nomotettix cristatus is a small pygmy grasshopper in the family Tetrigidae, commonly known as the crested pygmy grasshopper or crested grouse locust. It is one of approximately 35 Nearctic species of Tetrigidae. The species exhibits three recognized subspecies with distinct geographic distributions across North America. Like other members of its family, it is characterized by an elongated pronotum that extends over the abdomen, a trait distinguishing pygmy grasshoppers from typical grasshoppers in Acrididae.
pygmy-grasshopper groundhopper Tetrigidae Nearctic moist-habitat pronotum cristate North-America leaf-litter aquatic-margin Orthoptera Caelifera jumping-insect small-grasshopper crested-pronotum subspecies-variation Alberta Connecticut Florida Georgia Illinois lake-shore pond-edge stream-margin ground-dwelling Scudder-1862 Batrachidea-cristata Nomotettix-cristatus-cristatus Nomotettix-cristatus-compressus Nomotettix-cristatus-floridanus Tetriginae Acrididea Tetrigoidea Hexapoda Pterygota insect arthropod animal eukaryote metazoa animalia arthropoda insecta nomotettix cristatus crested pygmy grasshopper grouse-locust northern-crested-grouse-locust crested-grouse-locust crested-pygmy-grasshopper iNaturalist GBIF Catalogue-of-Life NCBI-Taxonomy Wikipedia Skejo Tumbrinck Devriese Hochkirch Morse-1895 Hancock-1902 Scudder-1863 Batrachidea cristata compressus floridanus Nearctic-region 35-species 2000-species 230-million-years dinosaur-contemporaries water-dependent-lifecycle Peruća-lake Croatia Balkans Europe systematics taxonomy faunistics natural-history biodiversity conservation endangered imperiled ZooKeys research amateur-naturalist citizen-science observation 53-observations rare neglected-diversity tropics Madagascar Australia New-Guinea Borneo Brazil Atlantic-Forest leaf-like-morphology spines warts undulations horns bizarre grotesque Hancock-1907 Genera-Insectorum Acididae pronotal-projection Holocerus Notocerus Holocerus-lucifer Holocerus-devriesei Notocerus-formidabilis Metopomystrum-muriciense Ophiotettix-pulcherrima Paraselina-brunneri Selivinga-tribulata Criotettix-bispinosus Tetrix-subulata Tettigidea-lateralis Tetrix-undulata Tetrix-tenuicornis Paratettix-meridionalis Paratettix-mexicanus Tetrix-depressa Tetrix-arenosa Tetrix-bipunctata Tetrix-japonica Paratettix-aztecus Hyperyboella-orphania Scelimena-producta Eurymorphopus-bolivariensis Discotettix-belzebuth Tetrix-ceperoi Tetrix-transsylvanica Arulenus Rhacocleis-buchichii Barbitistes-kaltenbachi Adžić Deranja Pavlović Fran-Rebrina Ivan-Budinski Vrlika HPM Hrvatski-Prirodoslovni-Muzej Zagreb Kitonić Miko Franjević Medak Mathieu Silva Pereira De-Domenico Sperber Connors Hendriksen Lambert Chong McMaster Monaghan Rentz Richter Rose Telnov Barclay Pauwels Entomological-Society-of-Latvia Riga Latvia Wallacea Volume-III biogeography nature-conservation Trinity-Audio guest-blog Josip-Skejo Orthoptera-of-Croatia Balkan-endemic endemic-species natural-history-museum museum-collections field-research career-determination serendipitous-discovery new-species systematic-research understudied-group regular-publishing 2021 2017 2020 2014 2016 2013 2012 2011 primary-school high-school biology-student snakes Uropeltidae Scolecophidia shield-tailed-snakes blind-snakes grasshoppers crickets Croatian-endemic first-encounter water-dependency lifecycle leading-European-orthopterists rare-species new-Arulenus-species understudied publishing Tetrigidae-research encyclopedia Elsevier digital-taxonomy open-access innovations journal growth trending social-media Flickr unidentified-rare-pygmy-grasshoppers students amateurs new-records not-rare Tribulation-helmed-groundhopper Kuranda Tully-Range Redlynch Kingfisher-park Speewah selected-awesome-places selected-amazing-taxa rainforests humid-forests extremely-rare weird-looking largest most-colourful ZooKeys-studies photographs identified-to-species-level variability pronotal-projection-morphology holotype Maroantsentra Antongil-Bay Tamatave Andasibe Vohimana Ranomafana Analamazaotra Traveler's-Palm Ravenala-madagascariensis natural-habitat skilful-flier flight large-back-spines Formidable-Pygmy-Grasshopper Sava-region true-colours small interesting Pygmy-unicorns Southern-America Murici description male-holotype head sternum frontal-view dorsal-view lateral-view sternomentum scale-bars Giraffehoppers unique antennal-shape face-coloration visual-animals courtship mating-pair Yapen-Island Cenderawasih-Bay W-New-Guinea young-entomologists decision study-pygmy-grasshoppers direction primary-and-high-school compare-data first-idea familiar-animals field order-Orthoptera friend fellow-student first-systematic-research two-Croatian-endemic-species first-years 2011-2012 never-saw single-pygmy-grasshopper BIOM Sinj Tetrigidae-location around-water first-time-ever lifecycle-water-dependent researching contacting Hendrik-Devriese Axel-Hochkirch Josef-Tumbrinck Croatian-Natural-History-Museum Skejo-et-al.-2014 Skejo-&-Caballero-2016 regularly-publishing Adžić-et-al.-2021 Endangered-Pygmy-Grasshoppers Encyclopaedia-of-Conservation Mathieu-et-al.-2021 ZooKeys-1042 Silva-et-al.-2017 comments recent-changes Metopomystrum Cleostratini Miriatrini ZooKeys-702 Skejo-et-al.-2020 online-social-media Anaselina Paraselina Selivinga rare-Australian-pygmy-grasshoppers ZooKeys-948 Malagasy-devils north south ZooKeys-957 Tumbrinck-&-Skejo-2017 taxonomic-and-biogeographic-revision New-Guinean-genus Ophiotettix-Walker-1871 Metrodorinae Ophiotettigini-trib.-nov.33-new-species nature's-creations encouraged U.S.-species spectacular closed Halloween orange-and-black Argus-tortoise-beetle Chelymorpha-cassidea oleander-caterpillar Syntomeida-epilais wheel-bug Arilus-cristatus morning-glory Convolvulaceae sweet-potato toxic-compounds alkaloids nerve-poisons monarch-butterfly milkweed-leaf-beetle oleander-aphid predator-protection polka-dot-wasp-moth vermillion-wings iridescent-blue-body white-spots wasp-mimicry ultrasonic-song mate-attraction egg-deposition oleander-leaf caterpillars cluster-feeding cardiac-glycosides heart-poisons dogbane-beetles natural-enemies barrel-shaped-eggs pale-orange exoskeleton-hardening orange-black-orange antennae-pale-orange body-and-legs-jet-black rear-end-brilliant-orange stout-beak proboscis impalement digestive-enzymes liquefaction medieval-torture-wheel Hemiptera true-bugs sucking-mouthparts incomplete-metamorphosis egg-nymph-adult plant-feeders harlequin-bugs squash-bugs stink-bugs fierce-predators pest-control Reduviidae assassin-bugs Zelus-longipes milkweed-assassin-bug sticky-forelegs prey-snaring proteolytic-enzymes predigestion muscular-pump black-wing-buds goldenrods meadow-plants Pselliopus-barberi orange-assassin-bug black-stripes stealth leafhopper brown-marmorated-stink-bug Halyomorpha-halys biological-control-agent native-protein-sources autumn-egg-laying spring-hatching May-June red-abdomens orange-antennae sawfly-larvae beetles other-bugs painful-poke caution handling pet-keeping Mad-Magazine Spy-vs-Spy pointy-nosed-secret-agents entomological-equivalent Bug-vs-Bug Heteroptera clan awesome-predator native-to-North-America record-numbers nursery function-of-wheel speculation cautious-approach embrace liquefied-meal protein-for-growth egg-production cluster-size tree-bark magnificent-creatures wide-variety scores stealthy-stalking assassination complementary-outbreak benefit naturally-occurring-predators parasites stem-onslaught no-interest-in-humans retreat memorable-painful-poke firsthand-experience clever-assassin USDA-NIFA-SCRI-Award research-support action-video website predator-prey-relationship natural-enemy ultimate-comeuppance Chinese-praying-mantis yellow-and-black-garden-spider arachnid close-encounter invasive-pest Asia Allentown-PA 1990s millions-of-dollars-damage crops home-invasion 33-states winter-refuge rising-crescendo concern farm-fields landscapes creature's-bane bounty awesome-six-legged-predator medieval-torture-device source-of-speculation little-fact tasty-morsel mark hapless-victim slurp protein-for-development autumn well-fed-female several-to-more-than-one-hundred sycamore-tree attendance irony nocent triggered-outbreak remain-to-be-seen word-of-caution Assasi pet laboratory learned-firsthand try-not-to-handle victim Shrewsbury Martinson Sargent fascination inspiration archive creatures trees UF extension integrated-pest-management NC-State AG295 html tortoise-beetles IFAS orn ole-cpillar UKY Agriculture CritterFiles casefile insects bugs assassin suggested-common-name scientific-name Linnaeus Megha-Kalsi Dakshina-R.-Seal publication preparation happy-safe-Halloween eat-stink-bug part-3 meets-wheel-bug mid-Atlantic-region inimitable-pest apples vegetables soybeans homes discovery damage distress homeowners wicked-winter previous-episodes give-stink-bugs Baby-boomers outwit undo sucking-insects Mother-Nature liquefied-tissues egg-conversion clusters spring bright-red-abdomens lucky half-dozen stealthily-stalked assassinating stinky-marmorated-cousins no-particular-interest stalking-humans too-large taste-bad firsthand beware handling-directly Ashley Nancy Chris Ryan Erik Caroline wrangling death-of-stink-bug Part-2 vandalizing-vegetables sullying-soybeans invading-homes wide-variety-crops curious-reunion arachnid-kind stalking capturing assassinating-stink-bugs game gorgeous-wheel-bug-nymphs cluster-near-eggs venturing-off first-victim true-bug-clan bed-bugs leaf-footed-bugs nefarious-brown-marmorated-stink-bug predicted struggle protect-crops prevent-stinky-invaders overlook eating-mechanism business-end powerful-beak front-legs cautiously-approaches embraces impales pumps-strong-digestive-enzymes liquefy-body-tissues slurps larvae tens-to-more-than-one-hundred plant-hoppers aphids increase-in-populations stink-bug-laden-nursery curious-irony outbreak beneficial-wheel-bugs iNaturalist-taxon rank preferred-common-name observations-count 399 Wikipedia-summary kingdom phylum class order family genus GBIF-taxonomy-match matched-scientific-name canonical-name status accepted match-type exact distribution-records authorship specific-epithet classification eukaryota nomotettix-cristatus authority basionym group grasshoppers-crickets-katydids species instructions fill-all-fields available-knowledge field-cannot-be-supported return-null NOT-repeat-information each-section-focused useful-detail schema taxon-record entomology-guide accurate conservative informative factual-correctness completeness clarity verbosity usefulness CRITICAL-RULES information-not-clearly-supported NOT-infer-species-level-traits higher-taxa explicitly-justified NOT-repeat-same-information multiple-fields unique-non-overlapping-content vague-generalizations like-most-insects typically-feeds-on-plants cautious-language has-been-observed is-known-to NOT-fabricate behaviors diet life-cycle-details host-relationships FIELD-INTENT summary high-level-overview 3-5-sentences appearance physical-description-only identification distinguish-from-similar-taxa habitat environment-conditions distribution geographic-range-only seasonality timing-of-activity feeding-habits null-if-unknown developmental-stages behavior notable-actions-or-habits ecologicalRole role-in-ecosystem humanRelevance interaction-with-humans similarTaxa must-include-reason misconceptions only-if-meaningful extraDetails important-additional-context STYLE-RULES clear-direct-sentences avoid-fluff-filler-language repeating-taxonomy-in-prose overly-technical-jargon concrete-statements abstract-descriptions QUALITY-RULES completeness-high most-fields-well-supported completeness-medium partial-but-reliable completeness-low sparse-data hasInferredContent true-ONLY-if-generalization-used otherwise-false OUTPUT-FORMAT strictly-match-JSON-schema no-extra-fields no-commentary-outside-JSON water-associated three-subspecies N.-c.-cristatus N.-c.-compressus N.-c.-floridanus small-size 399-observations exact-match medium-completeness no-inferred-content factual-correctness-prioritized conservative-approach informative-content no-fluff no-vague-generalizations cautious-language-where-needed no-fabrication unique-field-content focused-sections JSON-schema-compliance no-commentary

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