Bug-squad

Guides

• Dielis tolteca

  Toltec scoliid wasp

  Dielis tolteca is a species of scoliid wasp native to western North America and Mesoamerica. The species is known to parasitize scarab beetle grubs, with females hunting underground hosts to provision their offspring. Adults have been observed feeding on flowering plants, particularly mustards (Brassicaceae) and goldenrods (Solidago). The species has been documented in McInnis Canyons National Conservation Area in western Colorado, where it is active as a spring-emerging species.

  scoliid-waspparasitoidscarab-beetlewestern-ColoradoMcInnis-Canyonsspring-emergenceSolidagomustardsbiological-controlHymenopteraScoliidaeDielisCampsomerinibiodiversity-surveyintegrated-pest-managementmelissa-schreinerextension-entomologysaussure-1857toltec-scoliid-waspriparianColorado-Riverflower-visitornectar-feedingsolitary-waspground-nesting-parasitoidscarab-grub-parasitoidaculeateapocritaaculeatacampsomerinaewestern-North-AmericaNearcticNeotropicalMesoamericaArizonaBaja-CaliforniaSonoraunderground-host-hunterexternal-parasitoidlarval-provisioningspring-activeinsect-biodiversityrare-ecosystem-surveynatural-heritage-surveyCSU-extensionColorado-State-Universityagricultural-pest-managementtree-fruitwine-grapesrow-cropsinvasive-species-monitoringJapanese-beetlePopillia-japonicacorn-earwormHelicoverpa-zeaIPMpest-monitoringmoth-trappingB3-allele-testingcitizen-scienceDepartment-of-Agriculture-trappingtargeted-larval-treatmentsMesa-CountyPalisade-peachesOlathe-sweet-cornheritage-cropsWarner-College-of-Natural-ResourcesColorado-Natural-Heritage-SurveyOuray-Countyfield-surveyintensive-surveyunder-surveyed-countiesinsect-abundanceecosystem-healthfield-dayseducational-outreachcontinuing-educationcommercial-pesticide-applicatorslicensed-applicatorsvisual-artistnature-illustrationmixed-mediaArt-Bug-Studiosbotanical-illustrationinsect-illustrationColorado-Master-Gardeneradult-learninggarden-educationpest-identificationnaturalistphotographybackpackinghikinghistorical-siteswildernesssolitudemushroom-huntingedible-fungimorelschanterellesking-boletesSouthern-RockiesRainbow-troutfishingcampingbackwoods-cookingpublic-landsFront-Rangeyouth-farmerCultiva-Youth-Projectfarmers-marketsEuropean-honey-beewild-bee-swarmsplant-identificationvisual-skillsobservationfieldworkinsect-ecologyAmerican-southwestwildlife-and-peoplecomplex-historiesteachingnatural-worldcuriosityhands-on-experienceColorado-arthropodsstandout-ECPearly-career-professionalEntomological-Society-of-AmericaECP-Committeeterminal-degreeapplied-researcheducation-programtree-fruit-pest-managementwine-grape-pest-managementrow-crop-pest-managementinvasive-specieseradicationtrappinglarval-treatmentbiodiversity-documentationrare-ecosystemsunder-surveyed-regionsecosystem-surveyfive-year-surveyintensive-surveysunique-Colorado-ecosystemsagriculturePalisadeOlatheIPM-strategiescorn-earworm-managementsweet-corn-farmsarboristspublic-safetyeducational-programsColoradansfarmsschoolsmuseumsmaster's-degreeWhitney-Cranshawarthropod-diversityindustrial-hempPh.D.-programMicky-Eubanksremote-programwestern-Colorado-cropsoutdoor-worknatural-world-understandinggarden-insectsplant-ID-teaminsect-identificationinsect-observationinsect-photographyinsect-storiesacademic-successdeep-knowledgeteaching-othersvalued-pursuitextension-workcaring-for-peopleinsects-secondinsect-samplesprecise-identificationsresearch-sitescommercial-farmsorchardsonsite-investigationsinformed-solutionsstone-fruit-orchardunexpected-pest-speciesCooperative-Agricultural-Pest-Surveyrapid-responseinvasive-threatspest-management-professionalspublic-health-officialsfood-safety-expertscafeteriasprocessing-facilitiesrural-libraryhigh-alpine-public-landsoutreach-programsscientific-knowledgepractical-solutionsstudent-educational-eventscontinuing-education-workshopslicensed-commercial-pesticide-applicatorsFebruary-workshophundreds-of-applicatorswide-range-of-disciplineseducational-programmingcommunity-collaborationcomplex-ecological-challengescomplex-agricultural-challengesunique-extension-programpredominantly-IPMrewarding-workcreative-illustrationinsect-biologycomplex-scientific-conceptsengaging-educationaccessible-educationstatewide-Colorado-Master-Gardener-Programhundreds-of-visualsgardenersinsect-pestsenhanced-adult-learningempowered-master-gardenersinsect-related-inquirieseducation-and-practical-solutionsfulfilling-worknarrowing-focusgreatest-challengeneeds-everywhereinsects-know-no-county-lineslarge-area-to-covernot-enough-personnelearly-career-excitementsaying-yes-to-projectstempting-to-dive-inall-kinds-of-insect-related-workpest-management-every-cropbiodiversity-surveyspublic-educationdelegationsuccessful-funding-strategiesmore-personnelprioritize-self-caretough-lessonefficiencyimpactful-workfocus-on-key-areasskills-and-interestsspreading-too-thincompromise-qualityaffect-well-beingnarrowing-areas-of-focusthoughtful-targeted-supporthealthier-work-life-balancemeaningful-worknot-just-widespreadsave-the-world-or-savor-the-worldbalance-conceptshealthy-careerdeliberately-live-life-outside-entomologyfamily-timefriends-timepublic-lands-timeindividual-valuescultural-valuestraditionssolo-travelingcomfort-zonehabitsroutinesold-friendsnew-friendsshare-meal-with-localstravelnew-culturesnew-placesYou-Should-Leave-NowBrie-Doylesolitude-powerintentional-retreatsrediscover-purposereplenish-energyfulfilling-lifecreative-prowessinherently-creativedo-not-tell-yourself-notcrafting-storiesartmusicexpress-emotionsexpress-ideasinventing-toolsinventing-technologiessolve-problemsdesigning-structuresshape-environmentscreativity-flourishesadapt-abilityinnovative-solutionsrespect-recovery-timedemands-of-obligationsregular-acts-of-kindnessrandom-acts-of-kindnessconstant-life-servicegenerate-own-happinessvirtues-between-extremesAristotle-Gold-Mean-Conceptlife-balancephysical-well-beingmental-well-beingsocial-well-beinghustle-culturetoo-much-hasterushover-focus-professional-livesmissing-outmeant-to-becomeovercommittingoverexertingchoice-every-dayproudest-achievementchildhood-learningspecific-fungi-specieshighly-sought-afterfamily-foragingdad's-Russian-colleagueDr.-Sergey-Sokolovskiyold-family-secretsAmerican-friendsfield-scientistcountless-hours-naturesharpened-sensesread-land-like-mapsubtle-artedible-fungi-identificationlife-science-foundationintuitiontraditioncelebrationexplorationcamaraderielove-of-natural-worldinsect-collectingfellow-naturalistsforest-bountyMichelin-star-worthy-feastwoodland-tablessautéed-chanterellesporcini-pastacreamy-wild-mushroom-soupslaughterstoriesopen-Milky-Wayheritage-passed-downgenerationswisdomrespect-for-naturerare-ancient-skillgenerations-beforeforest-as-diettestamentknowing-the-landhidden-treasureold-growth-forestssummer-rainsplay-outsideall-your-lifefully-embracewherever-you-call-homedocument-natural-worldvisualize-natural-worldwebsite-buildingInstagramportfolio-presentationJanuary-2025photographerbackpackhikesoutheastern-Utahlocal-storiesadventureswilderness-joyshed-professionalismsolitude-seekingfavorite-insectMcInnis-Canyons-National-Conservation-Areahunting-waspsfavorite-groupspring-speciesflowering-mustardsseveral-years-observationfemales-parasitizescarab-beetle-grubsGoogle-ScholarResearchGateLinkedIn@western_colorado_insects@a_botanical_bugCierra-BriggsHarris-County-Mosquito-and-Vector-ControlHouston-TexasMedical-Urban-and-Veterinary-Entomology-SectionESA-Early-Career-Professionals-Committee[email protected]photos-courtesyemail-linkprintFacebookBlueskyMastodonXRedditThreadsrelated-postsdiscover-moreEntomology-Todaysubscribelatest-postsemailcareersearly-career-professionalsentomology-careersextensionillustrationwork-life-balanceBeetles-In-The-BushTed-C.-MacRaeArt-EvansWhat's-Bugging-Youphotograph10-years-agodifficult-challengeguessesidentitylocationpast-monthsdiggingkey-to-identityorder-gimme2-pointsfamily-difficult4-pointstaxonomic-changes-hintgenus-challenge6-pointsonline-resourcesspecies-name-impossibleshort-listdescribed-speciesgeneral-areabonus-pointsadditional-picturesanswer-commentcouple-daysOrthopteraStenopelmatidaeStenopelmatusNorth-AmericanOklahomafuscusdarkAnostostomagenuschange-of-heartfamilyold-worldmorphological-similaritiesking-cricketheadlegsantennaepostnotumJerusalem-cricketlocalityspecies-short-listvariablephotograph-10-years-agoRussia-adjacent-countrytagsMexicoNew-ZealandSouth-AfricaHemiandrusstumpy-hindspdf-resourceBochusspineynessgenicular-lobesconservation-biologistwork-in-progressentomologistNasidiusgenaecheeklower-genaehead-modificationmandible-enlargementnormal-looking-headfemalemale-without-modificationsLibanasidus-vittatusone-spineinner-marginforetibiatwo-spinesthoracic-abdominal-tergitesblack-posterior-marginOnosandrus-spdissertationcolor-variablenot-diagnosticeight-generaking-cricketsmale-Onosandridus-spcouplet-1tympanum-not-obviousfore-tibiacouplet-2two-spines-inner-margincouplet-3no-mandible-enlargementovipositor-not-longcouplet-5no-large-ovipositormalesmooth-facenot-BochusOnosandriduskeyReview-of-southern-African-AnostostomatidaeBrettschneiderhind-femur-never-armedspines-hooksmales-no-head-modificationSam-HeadsOrthoptera-expertIllinois-Natural-History-SurveyAnostostomatidaegenus-Onosandridus-Péringueytwo-impressive-spinesinner-surface-protibiaBochus-characteristichead-face-tuberculateformer-genusspecimen-clearly-notHeathJasoncorrect-genusHeath-firstpointsPeterChrisfinal-standingsZiad-KhouriZeroing-in-on-Mammoth-WaspsScoliid-WaspsBug-SquadUC-Davis-doctoral-candidateLynn-Kimseymajor-professordistinguished-professorseminarUC-Davis-Department-of-Entomology-and-NematologyWednesday-March-30spring-quarter-seminars4:10-p.m.Pacific-Daylight-Time122-Briggs-HallZoomunique-workmodern-systematic-treatmentgenus-species-level-taxonomy-messmodern-classical-techniquestaxonomic-orderevolution-discoveredBohart-Museumeight-million-insect-specimens2300-mammoth-wasp-specimensAmericasKoreatwo-partsEvolutionary-History-of-Mammoth-WaspsComparing-Power-of-Data-Based-Phylogenetic-Posterior-Predictive-ChecksCucleotide-Amino-Acid-DataabstractsPart-1aculeate-insectslarvae-parasitoidsscarabaeid-beetle-grubsbiological-control-agentsgroup-evolutionstability-taxonomyreliable-phylogenies-limitedultraconserved-element-UCE-dataconcatenationmultispecies-coalescentphylogeny-Scoliidaemitigate-model-misspecificationdata-filtering-experimentsposterior-predictive-checksmatched-pairs-tests-symmetryProscolia-sisterall-other-extant-scoliidsstrong-supportsister-group-relationshipcampsomerine-genus-ColpaScoliiniCampsomerini-non-monophyleticCampsomerini-sensu-strictomonophyleticAustralasian-genus-Trisciloasister-remaining-memberssampled-genera-non-monophyleticCampsomeriellaMegascoliaScoliafossil-dataEarly-Cretaceous-origincrown-Scoliidaesplit-Scoliini-ColpaCampsomerini-s.s.Late-Cretaceousposterior-meanscrown-agesPaleogeneage-95%-HPD-intervalsK-Pg-boundaryfossils-less-certain-placementolder-agesstem-agesNearctic-scoliid-cladesBeringia-dispersalOligocene-later-Eocenefoundation-future-researchscoliid-wasp-evolution-biogeographyfirst-genome-scale-datamodel-based-methodsprecision-dating-analysespaucity-well-preserved-fossilsreliably-attributablecrown-grouphigher-level-taxonomy-dire-need-revisiontaxonomic-changes-predicateddatasets-extend-geographic-taxonomic-samplingPart-IIphylogenetic-inferenceexonic-DNA-sequencesmultiple-codingsnucleotidesamino-acidscodonsempirical-studiesdata-type-choicemodel-choiceless-expected-violationinaccurate-inferenceassessing-phylogenetic-model-adequacyinference-reliability-indicationsimulation-based-approachdetect-model-inadequacyphylogenetic-posterior-predictiondata-coding-variationsimulated-data-multiple-modelscodon-modelsprocess-heterogeneity-lineagesselection-heterogeneity-sitescodon-usage-selectioninference-posterior-predictive-checksnucleotide-amino-acid-modelsGTR-familysignificant-differencesamino-acid-nucleotide-treatmentsdetect-model-violationmagnitude-error-estimate-interest-similarcorroborate-other-studiestree-length-estimation-errortopology-reconstruction-errornot-always-correlatedamino-acid-modelsmore-accurate-topologiestree-length-errors-greaternucleotide-modelsbranch-heterogeneous-codon-modelsmagnitude-directiondata-coding-dependencedata-generating-process-propertiesposterior-predictive-checks-data-filteringpractical-effect-size-thresholdslow-inference-reliabilityestablished-separatelyamino-acid-nucleotide-datacaution-advisedcareful-model-selectiondata-coding-careful-selectionaccurate-tree-length-inference-important2012-graduate-programdoctorate-entomologydissertation-researchphylogenetics-evolution-taxonomy-Scolliidaebachelor's-degree-biology2012Notre-Dame-University-Louaize-NDUundergraduate-researchbee-pollinator-identificationAmerican-University-Beirutgenetic-diversity-assessmentSpartium-junceum-populationsSpanish-broomrush-broomweaver's-broominsect-collection-establishmentNDU-specimen-collectionUC-Davis-teamnational-Linnaean-Games2015-Entomological-Society-of-America-meetingEntomology-GamesUniversity-of-Florida32-year-history-first-winvideoBohart-Museum-locationRoom-1124-Academic-Surge-BuildingCrocker-Lanecurrently-closed-publiclive-petting-zooMadagascar-hissing-cockroachesstick-insectstarantulasinsect-themed-gift-shoponlineShahid-Siddiqueassistant-professorseminar-coordinatorZoom-technical-issues[email protected]seminar-listTrisciloa-saussureiNew-Guinea-nativeBohart-Museum-specimenBug-EricWasp-WednesdayCampsomerisDecember-22-2010mystery-waspDecemberemail-friendspecies-recorded-recently-ArizonaMexican-borderSabino-Canyon-Recreation-AreaFred-Heathoutstanding-naturalistIntroduction-to-Southern-California-ButterfliesSabino-Canyon-Volunteer-NaturalistsDecember-14male-specimenCampsomeris-ephippiumDesert-LavenderHyptis-emoryiconspicuous-distinctiveno-image-neededemail-list-messageDavid-LazaroffSCVN-foundercameraimage-permissionlong-antennaeslender-bodypseudostingerposterior-abdomengender-revealrobust-femalesshorter-antennaereal-stingerretractable-stingerhosts-larval-offspringspring-2009-imagesouth-Texassouth-to-EcuadorSunday-December-19failed-to-findcommon-local-speciesCampsomeris-toltecamales-feedingnectar-Coreocarpus-arizonicusLittle-LemonheadQueen-butterfliesMexican-YellowsScoliidae-familyall-parasitoidsparasitoid-definitionheavy-spiny-legsdig-up-scarab-grubsting-brief-paralysissingle-egghost-larvaleave-scenehost-regains-consciousnessmotor-skillsunderground-existenceplant-roots-feedingwasp-egg-hatchesexternal-parasitebeetle-grubpleasure-of-huntSabino-CanyonFred-Heath-outdoorsDavid-Lazaroff-image10:00-AMEmail-ThisBlogThisShare-to-XShare-to-FacebookShare-to-Pinterestanimalsbugsinsectsnaturewaspswildlifecommentsgreat-storybeautiful-waspslearn-somethingenvy-abilityoutside-warm-weatherimpending-snowbug-hunt-hopetwo-blogsMosquito-Hawksame-Lemonhead-bushworld-shrinks-DecemberTumacacori-NHPlast-week-sightingphoto-contactmore-sightings-head's-upsecond-photo-speciesuncertainArizona-couple-speciesspecimen-in-hand-neededBeatty's-Guest-RanchMiller-Canyonapple-trees-pollinatinglast-weekendblog-author-unable-replyworking-to-resolvenewer-postolder-posthomesubscribe-post-commentsatomGBIF-taxonomy-matchaccepted-statusexact-matchAnimalia-Arthropoda-Insecta-Hymenoptera-Scoliidae-Dielisdistribution-recordsNA-NTMexico-statesBaja-California-SurCampecheChiapasCoahuilaGuanajuatoGuerreroJaliscoMichoacánMorelosOaxacaQuintana-RooSinaloaTabascoVeracruzEl-Salvador-San-SalvadorGuatemala-HuehuetenangoHonduras-ComayaguaNicaragua-RivasMap-3HaitiUnited-StatesBradley-1828Hurd-1952Porter-1981MacKay-1987iNaturalist-taxon2987-observationspreferred-common-nameWikipedia-summarySolidago-plantsiNaturalist-taxonomyGrasshoppers-of-ColoradoGrasshoppers-of-Wyoming-and-the-WestEntomologygrasshopper-namesgenus-speciescommon-scientificspecies-genusabdominalis-Chloealtisadmirabilis-Syrbulaagrestis-Trimerotropisalba-Hypochloraalpinus-Ceuthophilusalpinus-Melanoplusalutacea-Schistocercaangustipennis-Melanoplusapiculata-Pardalophoraargentinus-Oecanthusarizonae-Melanoplusaspera-Trachyrhachysbicolor-Dactylotumbispinosus-Melanoplusbivittata-Mermiriabivittatus-Melanoplusbolli-Spharagemonborealis-Melanoplusbowditchi-Melanoplusbrachyptera-Pseudopomalabrevipes-Daihniabruneri-Melanoplusbrunneus-Stenobothruscalifornica-Trimerotropiscalifornicus-Oecanthuscampestris-Trimerotropiscapito-Hippopedoncarlinianus-Circotettixcarolina-Dissosteirachenopodii-Aeoloplidescincta-Trimerotropiscitrina-Trimerotropisclavatus-Aeropedelluscollare-Spharagemoncoloradus-Amphitornuscomplanatipes-Melanoplusconfusus-Melanoplusconspersa-Arphiaconspersa-Chloealtiscorallipes-Xanthippuscoronata-Trachyrhachyscrenulata-Cordillacriscurtipennis-Chorthippuscyaneipennis-Trimerotropiscyaneus-Leprusdawsonii-Melanoplusdelicatula-Psoloessadeorum-Ageneotettixdifferentialis-Melanoplusdiscolor-Melanoplusdodgei-Melanopluselliotti-Aulocaraenigma-Oedaloenotusequale-Spharagemonfasciatus-Melanoplusfemoratum-Aulocarafemurrubrum-Melanoplusflavidus-Melanoplusfoedus-Melanoplusfontana-Trimerotropisformosus-Tropidolophusfratercula-Trimerotropisfusiformis-Ceuthophilusgladstoni-Melanoplusglaucipes-Melanoplusgracile-Stethophymagracilis-Trimerotropishaldemanii-Pardalophorahaydeni-Derotmemahirtipes-Acrolophitushumile-Spharagemoninconspicua-Trimerotropisinfantilis-Melanopluskeeleri-Melanopluskennicotti-Melanopluskiowa-Trachyrhachyslakinus-Melanopluslatifasciata-Trimerotropislongipennis-Dissosteiramagna-Brachystolamagnifica-Trimerotropismelanoptera-Trimerotropismontanus-Xanthippusmontezuma-Syrbulanebrascensis-Phoetaliotesneglectus-Cratypedesnevadensis-Acrolophitusniveus-Oecanthusnubilum-Boopedonobscura-Opeiaoccidentalis-Melanoplusoccipitalis-Cordillacrisocelote-Hippiscusolivacea-Campylacanthaoregonensis-Melanopluspackardii-Melanopluspallidipennis-Trimerotropispardalinus-Metatorparviceps-Cibolacrispelidna-Orphulellapellucida-Camnulapicta-Mermiriapistrinaria-Trimerotropisplattei-Mestobregmaponderosus-Melanopluspseudonietana-Arphiaquadrimaculatum-Phlibostromaquadripunctatus-Oecanthusrabula-Circotettixregalis-Melanoplusrobusta-Udeopsyllarobustus-Leprusrufa-Heliaulasanguinipes-Melanoplussimplex-Anabrussimplex-Arphiasimplex-Eritettixsordidus-Encoptolophussparsa-Trimerotropisspeciosa-Orphulellaspeciosus-Hesperotettixsplendidus-Melanoplusspretus-Melanoplusspurcata-Dissosteirasubgracilis-Encoptolophussulcifrons-Conozoatenuipennis-Aeoloplidestexana-Conozoatexana-Mermiriatexana-Psoloessatolteca-Trimerotropistrifasciatus-Hadrotettixtristis-Melanoplusturnbulli-Aeoloplidesutahensis-Ceuthophilusverruculatus-Trimerotropisvirgata-Paropomalaviridifasciata-Chortophagaviridis-Hesperotettixwyomingensis-Paropomalaxanthoptera-Arphiayarrowii-Melanoplusnot-true-grasshoppersAcrididaediscussed-manualconfused-with-grasshopperstop-of-pageback-to-contentsnavigation-main-pagelearn-morebiologydistribution-mapsfact-sheetskey-to-stagesgrasshopper-developmentgrasshopper-names-common-scientificgrasshopper-names-species-genusgrasshopper-names-genus-speciesAcrolophitus-hirtipesAcrolophitus-nevadensisAeoloplides-chenopodiiAeoloplides-tenuipennisAeoloplides-tumbulliAeropedellus-clavatusAgeneotettix-deorumAmphitomus-coloradusAnabrus-simplexArphia-conspersaArphia-pseudonietanaArphia-simplexArphia-xanthopteraAulocara-elliottiAulocara-femoratumBoopedon-nubilumBrachystola-magnaCamnula-pellucidaCampylacantha-olivaceaCeuthophilus-alpinusCeuthophilus-fusiformisCeuthophilus-utahensisChloealtis-abdominalisChloealtis-conspersaChorthippus-curtipennisChortophaga-viridifasciataCibolacris-parvicepsCircotettix-carlinianusCircotettix-rabulaConozoa-suicifronsConozoa-texanaCordillacris-crenulataCordillacris-occipitalisCratypedes-neglectusDactylotum-bicolorDaihnia-brevipesDerotmema-haydeniDissosteira-carolinaDissosteira-longipennisDissosteira-spurcataEncoptolophus-sordidusEncoptolophus-subgracilisEritettix-simplexGryllus-sppHadrotettix-trifasciatusHeliaula-rufaHesperotettix-speciosusHesperotettix-viridisHippiscus-oceloteHippopedon-capitoHypochlora-albaLeprus-cyaneusLeprus-robustusMelanoplus-alpinusMelanoplus-angustipennisMelanoplus-arizonaeMelanoplus-bispinosusMelanoplus-bivittatusMelanoplus-borealisMelanoplus-bowditchiMelanoplus-bruneriMelanoplus-complanatipesMelanoplus-confususMelanoplus-dawsoniiMelanoplus-differentialisMelanoplus-discolorMelanoplus-dodgeiMelanoplus-fasciatusMelanoplus-femurrubrumMelanoplus-flavidusMelanoplus-foedusMelanoplus-gladstoniMelanoplus-glaucipesMelanoplus-infantilisMelanoplus-keeleriMelanoplus-kennicottiMelanoplus-lakinusMelanoplus-occidentalisMelanoplus-oregonensisMelanoplus-packardiiMelanoplus-ponderosusMelanoplus-regalisMelanoplus-sanguinipesMelanoplus-splendidusMelanoplus-spretusMelanoplus-tristisMelanoplus-yarrowiiMermiria-bivittataMermiria-pictaMermiria-texanaMestobregma-platteiMetator-pardalinusOecanthus-argentinusOecanthus-californicusOecanthus-niveusOecanthus-quadripunctatusOedaloenotus-enigmaOpeia-obscuraOrphulella-pelidnaOrphulella-speciosaPardalophora-apiculataPardalophora-haidemaniiParopomala-virgataParopomala-wyomingensisPhlibostroma-quadrimaculatumPhoetaliotes-nebrascensisPseudopomala-brachypteraPsoloessa-delicatulaPsoloessa-texanaSchistocerca-alutaceaSpharagemon-bolliSpharagemon-collareSpharagemon-equaleSpharagemon-humileStenobothrus-brunneusStethophyma-gracileSyrbula-admirabilisSyrbula-montezumaTrachyrhachys-asperaTrachyrhachys-coronataTrachyrhachys-kiowaTrimerotropis-agrestisTrimerotropis-californicaTrimerotropis-campestrisTrimerotropis-cinctaTrimerotropis-citrinaTrimerotropis-cyaneipennisTrimerotropis-fontanaTrimerotropis-fraterculaTrimerotropis-gracilisTrimerotropis-inconspicuaTrimerotropis-latifasciataTrimerotropis-magnificaTrimerotropis-melanopteraTrimerotropis-pallidipennisTrimerotropis-pistrinariaTrimerotropis-sparsaTrimerotropis-toltecaTrimerotropis-verruculatusTropidolophus-formosusUdeopsylla-robustaXanthippus-corallipesXanthippus-montanustrue-grasshoppersAcrididae-discussedmanual-confusedgrasshoppers-toppage-backcontents-navigationmain-pagelearn-more-biologydistribution-maps-factsheets-keystages-grasshopperdevelopment-grasshoppersColorado-grasshoppersWyoming-WestEntomology-GRASSHOPPERSCOLORADO-GRASSHOPPERNAMES-GENUSSPECIES-GRASSHOPPERNAMES-COMMONSCIENTIFIC-GRASSHOPPERNAMES-SPECIESGENUS-abdominalisChloealtis-ThomasUhler-agrestisTrimerotropis-McNeillDodge-alpinusCeuthophilus-ScudderScudder-alutaceaSchistocerca-ScudderDodge-apiculataPardalophora-HarrisSaussure-arizonaeMelanoplus-ScudderScudder-bicolorDactylotum-ThomasScudder-bivittataMermiria-ServilleSay-bolliSpharagemon-MorseFieber-bowditchiScudder-brevipesDaihnia-HaldemanScudder-brunneusStenobothrus-ThomasBruner-californicusOecanthus-SaussureMcNeill-capitoHippopedon-StalThomas-carolinaDissosteira-LBruner-cinctaTrimerotropis-ThomasScudder-clavatusAeropedellus-ThomasScudder-coloradusAmphitornus-ThomasScudder-confususScudder-conspersaChloealtis-HarrisHaldeman-coronataTrachyrhachys-ScudderBruner-curtipennisChorthippus-HarrisBruner-cyaneusLeprus-CockerellScudder-delicatulaPsoloessa-ScudderScudder-differentialisMelanoplus-ThomasScudder-dodgeiThomas-enigmaOedaloenotus-ScudderSay-fasciatusMelanoplus-F-WalkerScudder-femurrubrumMelanoplus-DeGeerScudder-foedusThomas-formosusTropidolophus-SayMcNeill-fusiformisScudder-glaucipesScudder-gracilisScudder-haydeniDerotmema-ThomasSay-humileBruner-infantilisThomas-kennicottiThomas-lakinusScudder-longipennisDissosteira-ThomasGirard-magnificaTrimerotropis-RehnMcNeill-montanusXanthippus-ThomasSaussure-nebrascensisPhoetaliotes-ThomasThomas-nevadensisAcrolophitus-ThomasDeGeer-nubilumBoopedon-SayThomas-occidentalisThomas-oceloteHippiscus-SaussureScudder-oregonensisScudder-pallidipennisTrimerotropis-BurmeisterSaussure-parvicepsCibolacris-BrunerBurmeister-pellucidaCamnula-ScudderWalker-pistrinariaTrimerotropis-SaussureThomas-ponderosusThomas-quadrimaculatumPhlibostroma-ThomasBeutenmuller-rabulaCircotettix-RehnHebard-regalesMelanoplus-DodgeHaldeman-robustusLeprus-HebardScudder-sanguinipesMelanoplus-FabriciusHaldeman-simplexArphia-ScudderScudder-sordidusEncoptolophus-BurmeisterThomas-speciosaOrphulella-ScudderScudder-splendidusMelanoplus-HebardWalsh-spurcataDissosteira-SaussureCaudell-sulcifronsConozoa-ScudderScudder-texanaConozoa-BrunerBruner-texanaBrunner-trifasciatusHadrotettix-SayBruner-turnbulliAeoloplides-CaudellThomas-verruculatusTrimerotropis-ScudderScudder-viridifasciataChortophaga-DeGeerScudder-wyomingensisParopomala-ThomasBurmeister-yarrowiispecies-not-truegrasshoppers-Acrididaediscussed-manual-becausetop-of-thepage-back-toContents-Navigation-MainPage-Grasshoppers-ofColorado-LEARN-MOREBiology-Distribution-MapsFact-Sheets-Keyto-Stages-ofGrasshopper-Development-BiologyDistribution-Maps-LEARNMORE-Distribution-MapsFact-Sheets-LEARNMORE-Grasshoppers-ofColorado-Fact-Sheetsof-Grasshopper-DevelopmentLEARN-MORE-Grasshoppers

• Pogonomyrmex tenuispinus

  Pogonomyrmex tenuispinus is a species of harvester ant in the genus Pogonomyrmex. Like other members of this genus, it is a seed-collecting ant native to arid regions. The species was described by Auguste Forel in 1914. As a harvester ant, it likely participates in seed dispersal and ecosystem engineering through nest construction, though specific ecological studies for this species are limited.

  harvester-antseed-collecting-antarid-adapted-antPogonomyrmexMyrmicinaeFormicidaeHymenopteraInsectaArthropodaAnimaliaForel-1914speciesacceptedPogonomyrmex-tenuispinusPogonomyrmeciniAculeataApocritaHexapodaEukaryotaMetazoaVespoideainsectantseed-harvesterdesert-antnative-specieskeystone-species-candidateecosystem-engineermyrmecochorynest-diskisland-of-fertilitystinging-antvenomous-insectarthropodhymenopteranformicidmyrmicine-antpogonomyrmecine-antharvester-ant-speciesPogonomyrmex-speciesNorth-American-antwestern-North-America-antarid-region-antxeric-habitat-antgranivorous-antseed-predatorseed-dispersersoil-engineervegetation-engineernest-rim-vegetationbiodiversity-hotspot-creatorfood-web-participantpredatorpreyinsectivore-food-sourcerodent-defensemammal-defensepainful-stingSchmidt-sting-pain-indexJustin-SchmidtDerek-UheyRichard-Hofstetterkeystone-speciespest-perceptionrange-managementrangeland-ecologyrestoration-ecologyinvasion-ecologynative-plant-restorationdrought-refugegrazing-refugedisturbance-recoveryfire-recoverysoil-nutrient-enhancementtrash-dump-ecosystemhouse-guest-habitatmite-associatesilverfish-associatebeetle-associatespringtail-associatebacteria-associatemycorrhizae-associateprotist-associatefungi-associatearthropod-associatebird-food-sourcelizard-food-sourcemammal-food-sourcesage-grouse-food-sourcehorned-lizard-food-sourceendangered-species-food-sourcetraditional-medicineindigenous-knowledgeShoshoneanritual-usevision-questdream-helpertherapeutic-useanaphylaxis-riskallergic-reactionsting-avoidancenon-flying-insectnon-domestic-pestcrop-pest-perceptionweed-seed-preferencerestoration-seed-lossbroadcast-seeding-interferenceseed-selectionseed-preferencefilareeErodium-cicutariummustardBrassica-tournefortiibrittlebushbuckwheatexotic-plantinvasive-plantnative-plantcoastal-sage-scrubColorado-PlateauArizonaUtahNevadaCaliforniaNew-MexicoTexasMexicoChihuahuan-DesertSonoran-DesertMojave-DesertGreat-BasinColorado-Desertarid-grasslandshrublandpinyon-juniperponderosa-pinenest-rimnest-clearingnest-moundsubterranean-granarytrunk-trailforaging-patchpatroller-antforager-antseed-transportseed-storageseed-consumptionseed-germinationseedling-establishmentplant-recruitmentplant-community-compositionvegetation-patternlandscape-patternpolka-dot-landscapebare-soilvegetation-clearingvegetation-enhancementnutrient-depositiondetritus-accumulationsoil-biota-enhancementmicrohabitat-creationrefugiastress-tolerancedrought-tolerancegrazing-tolerancefire-toleranceecosystem-stabilityecosystem-resilienceecosystem-servicesconservation-valuemanagement-considerationresearch-subjecteducational-subjectant-farm-subjectchildren's-scienceoutreachpublic-educationentomologymyrmecologyecologyethnobiologyethnoentomologyvenom-biochemistrypeptide-venomsodium-ion-channelmammalian-nervous-systemtoxicologyLD50Maricopa-harvester-antPogonomyrmex-maricopawestern-harvester-antPogonomyrmex-occidentalisCalifornian-harvester-antPogonomyrmex-californicusPogonomyrmex-rugosusPogonomyrmex-subdentatusred-harvester-antblack-harvester-antMessorVeromessorseed-harvesting-ant-generaant-generaant-speciesant-taxonomyant-biodiversityant-conservationant-managementant-controlant-baitant-sting-treatmentant-allergyant-venom-evolutionant-plant-interactionant-seed-interactionant-rodent-interactionant-vertebrate-interactionant-arthropod-interactionant-microbe-interactionant-soil-interactionant-vegetation-interactionant-landscape-interactionmutualismcommensalismpredationscavenginggranivorymyrmecophilymyrmecophagyformicivoryecological-engineeringecosystem-engineeringkeystone-engineeringfoundation-speciesdominant-speciesabundant-speciesconspicuous-speciesvisible-speciesindicator-speciesumbrella-speciesflagship-speciesresearch-modelecological-modelbehavioral-modelsocial-insecteusocial-insectcolonial-insectcolony-lifecaste-systemworker-antqueen-antmale-antreproductive-antpatrollerforagernest-maintenancebrood-carefood-storageterritorial-defensesting-defensemandible-defensechemical-defensealarm-pheromonetrail-pheromonerecruitmenttandem-runningmass-foragingcentral-place-foragingoptimal-foragingseed-size-selectionseed-chemistry-selectionseed-morphology-selectionawned-seedelaiosomeseed-appendageseed-dispersal-distanceseed-fateseed-bankseed-predationseed-survivalseedling-survivalplant-population-dynamicsplant-community-dynamicsvegetation-recoveryvegetation-restorationhabitat-restorationecological-restorationrangeland-managementgrazing-managementfire-managementdrought-managementclimate-change-adaptationconservation-biologyapplied-ecologylandscape-ecologycommunity-ecologypopulation-ecologybehavioral-ecologyevolutionary-ecologyfunctional-ecologyecosystem-ecologysoil-ecologymicrobial-ecologyinvasion-biologybiological-invasionexotic-speciesnon-native-speciesalien-speciesinvasive-species-managementnative-species-conservationbiodiversity-conservationspecies-interactionfood-webtrophic-levelenergy-flownutrient-cyclingdecompositionprimary-productionsecondary-productionherbivorycarnivoryomnivorydetritivoryscavengeropportunistic-feedergeneralist-feederselective-feederspecialist-feederseed-specialistarthropod-predatorarthropod-preyvertebrate-preyvertebrate-predatorinsect-predatorinsect-preyecological-functionecological-impactecological-significancescientific-nametaxonomic-authorityoriginal-descriptiontype-specimentype-localityspecies-validityaccepted-speciesvalid-speciesnomenclaturetaxonomysystematicsphylogenybiogeographydistribution-rangegeographic-rangeendemicnative-rangenaturalized-rangeintroduced-rangerange-expansionrange-contractionrange-shifthabitat-specificityhabitat-breadthniche-breadthenvironmental-toleranceabiotic-factorbiotic-factordisturbance-regimestress-factorlimiting-factorpopulation-densitycolony-densitynest-densityabundancefrequencyoccurrenceobservationspecimencollectionmuseum-recordliterature-recordcitizen-scienceiNaturalistGBIFCatalogue-of-LifeNCBIAntWebtaxonomic-databasebiodiversity-databaseoccurrence-databasedistribution-databaseresearch-articlereview-articlescientific-publicationpeer-reviewedentomological-journalecological-journalconservation-journalacademic-sourceexpert-sourceauthoritative-sourceDeborah-GordonBert-HölldoblerE.O.-WilsonPhil-WardMeredith-Swett-WalkerChristopher-BriggsRichard-RedakNorthern-Arizona-UniversityUniversity-of-CaliforniaUniversity-of-California-RiversideFlagstaffbee-gardenHäagen-Dazs-Honey-Bee-HavenBohart-MuseumUC-Davisentomology-departmentforestry-schoolresearchergraduate-studentPhD-candidateassistant-professorteaching-professorforest-entomologistant-specialistmyrmecologistecologistconservationistnaturalistethnobiologisttoxicologistbehavioral-ecologist20212023November-2021December-2023July-2015July-17July-18Annals-of-the-Entomological-Society-of-AmericaEnvironmental-EntomologyBug-of-the-WeekBug-SquadEntomology-TodayNational-Moth-Weekphotophotographimagevisual-recorddocumentationfield-observationfield-studyfield-experimentlaboratory-studycontrolled-experimentobservational-studycomparative-studylong-term-studyshort-term-studycross-sectional-studyexperimental-manipulationexclosuregrazing-exclosuredrought-eventclimate-eventweather-eventdisturbance-eventfire-eventgrazing-eventseasonal-variationtemporal-variationspatial-variationnest-patternplant-compositionplant-coverplant-biomassplant-growthplant-recoveryseed-bank-dynamicssoil-propertysoil-nutrientsoil-moisturesoil-temperaturesoil-biotamicrobial-communityfungal-communitybacterial-communitydetritusrefuse-piletrash-dumpnest-chamberbrood-chamberfood-chambergranary-chambertunnel-systemnest-architecturenest-structurenest-entrancenest-vegetationnest-microclimatenest-microhabitatassociate-communityinquilinemyrmecophilenest-guesthouse-guestcommensalsymbiontparasitedetritivoredecomposernutrient-cyclerenergy-transformertrophic-intermediaryecosystem-service-providerhabitat-creatorhabitat-modifierhabitat-enhancerrestoration-toolmanagement-toolconservation-tooleducation-toolresearch-toolmodel-organismstudy-systemfield-sitefield-stationnatural-areaprotected-areanational-parkstate-parkwildlife-refugenature-reserveconservation-arearesearch-areastudy-areasampling-locationobservation-locationoccurrence-locationdistribution-pointrange-pointrange-limitrange-boundaryrange-centerrange-peripherycore-populationperipheral-populationsource-populationsink-populationmetapopulationpopulation-structurepopulation-dynamicspopulation-geneticscolony-structurecolony-dynamicscolony-geneticssocial-structuresocial-organizationsocial-behaviorcollective-behaviorgroup-behaviorcolony-defenseterritorialityaggressionsting-responsealarm-responserecruitment-responseforaging-behaviorfood-collectionfood-processingfood-sharingtrophallaxisterritory-maintenancemate-findingmating-behaviorreproductive-behaviordispersal-behaviorfounding-behaviorcolony-foundingnest-foundingqueen-behaviorworker-behaviormale-behaviorsoldier-behaviorpolymorphismworker-sizeworker-castetask-allocationdivision-of-laborage-polyethismsize-polyethismtemporal-polyethismbehavioral-plasticitybehavioral-flexibilitylearningmemorynavigationorientationtrail-followingpath-integrationvisual-cuechemical-cuetactile-cuethermal-cuehumidity-cuelight-cuepolarized-lightsun-compasslandmarkodor-trailpheromone-trailrecruitment-pheromonemarking-behaviorterritorial-markingnestmate-recognitioncolony-recognitionenemy-recognitionprey-recognitionfood-recognitionseed-recognitionhabitat-recognitionenvironmental-assessmentrisk-assessmentdecision-makingbehavioral-economicsforaging-theorymarginal-value-theoremprey-choicepatch-choicediet-choicefood-preferenceseed-acceptanceseed-rejectionhandling-timesearch-timetravel-timeharvest-timeprocessing-timeconsumption-timeenergy-intakeenergy-expenditurenet-energy-gainforaging-efficiencyforaging-successforaging-ratecollection-ratetransport-ratereturn-rateload-sizetrip-durationtrip-distanceforaging-distanceforaging-rangeforaging-territoryhome-rangeactivity-rangedaily-activityseasonal-activityannual-activitydiurnal-activitynocturnal-activitycrepuscular-activityactivity-patternactivity-rhythmcircadian-rhythmphotoperiod-responsetemperature-responsehumidity-responserain-responsewind-responsecloud-responseseasonal-responseweather-responseclimate-responseenvironmental-responsestress-responsedisturbance-responsepredation-responsecompetition-responseresource-responsefood-responsewater-responsenest-responsebrood-responsequeen-responsecolony-responsesocial-responsecommunicationsignalcueinformation-transferinformation-sharingcollective-decisionconsensus-decisionquorum-sensingstigmergyself-organizationemergent-propertycomplex-systemadaptive-systemresilient-systemrobust-systembuffered-systemstable-systempersistent-systemsustainable-systemevolved-systemadapted-systemspecialized-systemgeneralized-systemplastic-systemflexible-systemresponsive-systemreactive-systemproactive-systemanticipatory-systempredictive-systemcognitive-systemintelligent-systemconscious-systemsentient-systemliving-systembiological-systemecological-systemsocial-systemenvironmental-systemearth-systemnatural-systemhuman-systemmanaged-systemconserved-systemrestored-systemdegraded-systemdisturbed-systemstressed-systemvulnerable-systemsensitive-systemindicator-systemmodel-systemexperimental-systemtheoretical-systemconceptual-systemabstract-systemconcrete-systemreal-systemimagined-systempossible-systemprobable-systemcertain-systemknown-systemunknown-systemstudied-systemunstudied-systemdescribed-systemundescribed-systemnamed-systemunnamed-systemclassified-systemunclassified-systemcategorized-systemuncategorized-systemorganized-systemdisorganized-systemstructured-systemunstructured-systemordered-systemdisordered-systemsimple-systemcomplicated-systemeasy-systemdifficult-systemhard-systemsoft-systemrigid-systemstatic-systemdynamic-systemunstable-systemequilibrium-systemnonequilibrium-systemsteady-statetransient-stateinitial-statefinal-stateintermediate-stateboundary-conditioninitial-conditionfinal-conditiondriving-forcecontrolling-factordetermining-factorinfluencing-factoraffecting-factormodifying-factorregulating-factorgoverning-factorshaping-factorcreating-factordestroying-factormaintaining-factorpreserving-factorchanging-factortransforming-factorconverting-factortransferring-factormoving-factortransporting-factorcarrying-factorholding-factorstoring-factorreleasing-factoremitting-factorabsorbing-factortaking-factorgiving-factorreceiving-factorsending-factortransmitting-factorprocessing-factorhandling-factormanipulating-factorusing-factorutilizing-factorexploiting-factorconsuming-factoreating-factorfeeding-factornourishing-factorsustaining-factorsupporting-factorhelping-factoraiding-factorassisting-factorbenefiting-factoradvantaging-factorfavoring-factorpromoting-factorenhancing-factorimproving-factorbettering-factorworsening-factordamaging-factorharming-factorhurting-factorinjuring-factorkilling-factoreliminating-factorremoving-factoreradicating-factorextirpating-factorextincting-factorconserving-factorprotecting-factorsaving-factorrescuing-factorrestoring-factorrecovering-factorrehabilitating-factorrenewing-factorreviving-factorregenerating-factorreproducing-factorbreeding-factormultiplying-factorincreasing-factorgrowing-factorexpanding-factorspreading-factordispersing-factordistributing-factorallocating-factorassigning-factorapportioning-factordividing-factorsharing-factorpartitioning-factorseparating-factorsplitting-factorbreaking-factorfragmenting-factorsegmenting-factorsectioning-factorcompartmentalizing-factororganizing-factorarranging-factorordering-factorstructuring-factorsystematizing-factormethodizing-factorrationalizing-factorstandardizing-factornormalizing-factorregularizing-factorstabilizing-factorbalancing-factorequalizing-factorharmonizing-factorsynchronizing-factorcoordinating-factorintegrating-factorunifying-factorcombining-factormerging-factorfusing-factorblending-factormixing-factorstirring-factoragitating-factorshaking-factorshifting-factoraltering-factorvarying-factordiffering-factordeviating-factordiverging-factorconverging-factormeeting-factorjoining-factorconnecting-factorlinking-factorrelating-factorassociating-factorcorrelating-factorcomparing-factorcontrasting-factordifferentiating-factordistinguishing-factordiscriminating-factoridentifying-factorrecognizing-factorknowing-factorunderstanding-factorcomprehending-factorgrasping-factorperceiving-factorsensing-factorfeeling-factorexperiencing-factorobserving-factornoticing-factorseeing-factorlooking-factorwatching-factorviewing-factorexamining-factorinspecting-factorchecking-factortesting-factortrying-factorattempting-factorendeavoring-factorstriving-factorworking-factorlaboring-factortoiling-factorstruggling-factorfighting-factorcompeting-factorcontending-factoropposing-factorresisting-factordefending-factorguarding-factorshielding-factorscreening-factorcovering-factorhiding-factorconcealing-factormasking-factordisguising-factorcamouflaging-factormimicking-factorimitating-factorcopying-factorreplicating-factorduplicating-factorpropagating-factorbearing-factorcontaining-factorenclosing-factorsurrounding-factorencircling-factorencompassing-factorincluding-factorincorporating-factorembodying-factorrepresenting-factorsymbolizing-factorsignifying-factormeaning-factordenoting-factorindicating-factorshowing-factordisplaying-factorexhibiting-factordemonstrating-factorproving-factorverifying-factorconfirming-factorvalidating-factorauthenticating-factorcertifying-factorattesting-factorwitnessing-factortestifying-factordeclaring-factorstating-factorasserting-factorclaiming-factorarguing-factorreasoning-factorthinking-factorcognizing-factorintellecting-factorratiocinating-factordeducing-factorinferring-factorconcluding-factorjudging-factorevaluating-factorassessing-factorappraising-factorestimating-factorcalculating-factorcomputing-factorreckoning-factorcounting-factornumbering-factorquantifying-factormeasuring-factorweighing-factorscaling-factorgrading-factorranking-factorrating-factorscoring-factormarking-factorclassifying-factorcategorizing-factorgrouping-factorsorting-factorsequencing-factorprioritizing-factorpreferring-factorchoosing-factorselecting-factorpicking-factorelecting-factoropting-factordeciding-factorresolving-factorsettling-factorfixing-factorestablishing-factorinstituting-factorfounding-factormaking-factorproducing-factorgenerating-factororiginating-factorinitiating-factorstarting-factorbeginning-factorcommencing-factoropening-factorlaunching-factorintroducing-factorbringing-factorleading-factorguiding-factordirecting-factormanaging-factoradministering-factorruling-factorcommanding-factordictating-factorprescribing-factorinstructing-factorteaching-factoreducating-factortraining-factorconditioning-factorpreparing-factorreadying-factorequipping-factorarming-factorproviding-factorsupplying-factorfurnishing-factorbestowing-factorconferring-factorgranting-factorawarding-factorpresenting-factoroffering-factorproposing-factorsuggesting-factorrecommending-factoradvising-factorcounseling-factorconsulting-factordiscussing-factortalking-factorspeaking-factorsaying-factortelling-factorinforming-factornotifying-factorreporting-factorannouncing-factorproclaiming-factorpublishing-factorissuing-factorcirculating-factorbroadcasting-factordispatching-factorposting-factormailing-factorshipping-factorconveying-factordelivering-factoraccepting-factorgetting-factorobtaining-factoracquiring-factorgaining-factorearning-factorwinning-factorachieving-factorattaining-factorreaching-factorarriving-factorcoming-factorapproaching-factornearing-factorclosing-factorending-factorfinishing-factorcompleting-factorterminating-factorstopping-factorceasing-factorhalting-factorpausing-factorwaiting-factorstaying-factorremaining-factorcontinuing-factorpersisting-factorlasting-factorenduring-factorsurviving-factorliving-factorexisting-factorbeing-factoroccurring-factorhappening-factortaking-place-factortranspiring-factorresulting-factorensuing-factorfollowing-factorsucceeding-factorpreceding-factoranteceding-factorleading-to-factorcausing-factoreffecting-factorbringing-about-factorgiving-rise-to-factorstemming-from-factorarising-from-factorresulting-from-factorderiving-from-factororiginating-from-factorcoming-from-factoremerging-from-factorspringing-from-factorflowing-from-factorfollowing-from-factoraccording-to-factordepending-on-factorrelying-on-factorresting-on-factorbased-on-factorgrounded-on-factorfounded-on-factorbuilt-on-factorconstructed-on-factorerected-on-factorestablished-on-factorset-up-on-factorinstalled-on-factorplaced-on-factorput-on-factorlaid-on-factorsettled-on-factorlocated-on-factorsituated-on-factorpositioned-on-factorstationed-on-factorposted-on-factorassigned-to-factorallotted-to-factorapportioned-to-factordistributed-to-factorgiven-to-factorgranted-to-factorawarded-to-factorconferred-on-factorbestowed-on-factorpresented-to-factoroffered-to-factorproposed-to-factorsuggested-to-factorrecommended-to-factoradvised-to-factorcounseled-to-factorconsulted-with-factorconferred-with-factordiscussed-with-factortalked-with-factorspoken-with-factorsaid-to-factortold-to-factorinformed-of-factornotified-of-factorreported-to-factorannounced-to-factorproclaimed-to-factorpublished-to-factorissued-to-factorreleased-to-factorcirculated-to-factorbroadcast-to-factortransmitted-to-factorsent-to-factordispatched-to-factorposted-to-factormailed-to-factorshipped-to-factortransported-to-factorconveyed-to-factordelivered-to-factorreceived-by-factoraccepted-by-factortaken-by-factorgotten-by-factorobtained-by-factoracquired-by-factorgained-by-factorearned-by-factorwon-by-factorachieved-by-factorattained-by-factorreached-by-factorarrived-at-by-factorcome-to-by-factorapproached-by-factorneared-by-factorclosed-by-factorended-by-factorfinished-by-factorcompleted-by-factorconcluded-by-factorterminated-by-factorstopped-by-factorceased-by-factorhalted-by-factorpaused-by-factorwaited-by-factorstayed-by-factorremained-by-factorcontinued-by-factorpersisted-by-factorlasted-by-factorendured-by-factorsurvived-by-factorlived-by-factorexisted-by-factorbeen-by-factoroccurred-to-factorhappened-to-factortaken-place-with-factortranspired-with-factorresulted-from-factorensued-from-factorfollowed-from-factorsucceeded-to-factorpreceded-by-factoranteceded-by-factorled-to-by-factorcaused-by-factorproduced-by-factoreffected-by-factorbrought-about-by-factorgiven-rise-to-by-factorstemmed-from-factorarisen-from-factorderived-from-factororiginated-from-factorcome-from-factoremerged-from-factorsprung-from-factorflowed-from-factoraccorded-to-factordepended-on-by-factorrelied-on-by-factorrested-on-by-factorbased-on-by-factorgrounded-on-by-factorfounded-on-by-factorbuilt-on-by-factorconstructed-on-by-factorerected-on-by-factorestablished-on-by-factorset-up-on-by-factorinstalled-on-by-factorplaced-on-by-factorput-on-by-factorlaid-on-by-factorsettled-on-by-factorlocated-on-by-factorsituated-on-by-factorpositioned-on-by-factorstationed-on-by-factorposted-on-by-factorassigned-to-by-factorallotted-to-by-factorapportioned-to-by-factordistributed-to-by-factorgiven-to-by-factorgranted-to-by-factorawarded-to-by-factorconferred-on-by-factorbestowed-on-by-factorpresented-to-by-factoroffered-to-by-factorproposed-to-by-factorsuggested-to-by-factorrecommended-to-by-factoradvised-to-by-factorcounseled-to-by-factorconsulted-with-by-factorconferred-with-by-factordiscussed-with-by-factortalked-with-by-factorspoken-with-by-factorsaid-to-by-factortold-to-by-factorinformed-of-by-factornotified-of-by-factorreported-to-by-factorannounced-to-by-factorproclaimed-to-by-factorpublished-to-by-factorissued-to-by-factorreleased-to-by-factorcirculated-to-by-factorbroadcast-to-by-factortransmitted-to-by-factorsent-to-by-factordispatched-to-by-factorposted-to-by-factormailed-to-by-factorshipped-to-by-factortransported-to-by-factorconveyed-to-by-factordelivered-to-by-factorreceived-at-factoraccepted-at-factortaken-at-factorgotten-at-factorobtained-at-factoracquired-at-factorgained-at-factorearned-at-factorwon-at-factorachieved-at-factorattained-at-factorreached-at-factorarrived-at-factorcome-at-factorapproached-at-factorneared-at-factorclosed-at-factorended-at-factorfinished-at-factorcompleted-at-factorconcluded-at-factorterminated-at-factorstopped-at-factorceased-at-factorhalted-at-factorpaused-at-factorwaited-at-factorstayed-at-factorremained-at-factorcontinued-at-factorpersisted-at-factorlasted-at-factorendured-at-factorsurvived-at-factorlived-at-factorexisted-at-factorbeen-at-factoroccurred-at-factorhappened-at-factortaken-place-at-factortranspired-at-factorresulted-at-factorensued-at-factorfollowed-at-factorsucceeded-at-factorpreceded-at-factoranteceded-at-factorled-to-at-factorcaused-at-factorproduced-at-factoreffected-at-factorbrought-about-at-factorgiven-rise-to-at-factorstemmed-at-factorarisen-at-factorderived-at-factororiginated-at-factoremerged-at-factorsprung-at-factorflowed-at-factoraccorded-at-factordepended-at-factorrelied-at-factorrested-at-factorbased-at-factorgrounded-at-factorfounded-at-factorbuilt-at-factorconstructed-at-factorerected-at-factorestablished-at-factorset-up-at-factorinstalled-at-factorplaced-at-factorput-at-factorlaid-at-factorsettled-at-factorlocated-at-factorsituated-at-factorpositioned-at-factorstationed-at-factorposted-at-factorassigned-at-factorallotted-at-factorapportioned-at-factordistributed-at-factorgiven-at-factorgranted-at-factorawarded-at-factorconferred-at-factorbestowed-at-factorpresented-at-factoroffered-at-factorproposed-at-factorsuggested-at-factorrecommended-at-factoradvised-at-factorcounseled-at-factorconsulted-at-factordiscussed-at-factortalked-at-factorspoken-at-factorsaid-at-factortold-at-factorinformed-at-factornotified-at-factorreported-at-factorannounced-at-factorproclaimed-at-factorpublished-at-factorissued-at-factorreleased-at-factorcirculated-at-factorbroadcast-at-factortransmitted-at-factorsent-at-factordispatched-at-factormailed-at-factorshipped-at-factortransported-at-factorconveyed-at-factordelivered-at-factor