Scarab-beetle
Guides
Acoma conjuncta
Acoma conjuncta is a species of scarab beetle described by Howden in 1962. It belongs to the subfamily Melolonthinae, a group commonly known as May beetles or June beetles. The species has been documented only from Texas, USA, suggesting a restricted geographic range. As a member of the genus Acoma, it is part of a small group of scarab beetles primarily distributed in western North America.
Acoma diminiata
Acoma diminiata is a small scarab beetle in the family Scarabaeidae, described by Howden in 1958. It is known from New Mexico in the southwestern United States. Adults measure 4.3–5.7 mm in length. The species belongs to the subfamily Melolonthinae and tribe Acomini.
Acoma glabrata
Acoma glabrata is a small scarab beetle in the family Scarabaeidae, described by Cazier in 1953. It is known from the southwestern United States and northwestern Mexico. Adults measure 5.5–8.3 mm in length. The species belongs to the tribe Acomini within the subfamily Melolonthinae.
Aegialia arenaria
dune scarab beetle
Aegialia arenaria, commonly known as the dune scarab beetle, is a coastal scarab beetle species restricted to sand dune habitats. It occurs across western and northern Europe and has been introduced to eastern North America, where it is known from coastal areas of Canada and the northeastern United States. The species is specialized to sandy coastal environments and is not found inland.
Aegialia concinna
Ciervo Aegilian Scarab Beetle
Aegialia concinna is a small scarab beetle in the subfamily Aegialiinae. The species was described in 1977 and is known only from the western United States. Members of the genus Aegialia are commonly associated with sandy coastal or dune habitats.
Aegialia crescenta
Crescent Dunes scarab beetle
Aegialia crescenta is a species of scarab beetle in the subfamily Aegialiinae, first described by Gordon and Cartwright in 1977. It is endemic to the United States and appears to have a restricted distribution centered on Nevada. The species belongs to a genus of beetles typically associated with sandy habitats.
Aegialia nana
Aegialia nana is a small scarab beetle in the family Scarabaeidae, subfamily Aphodiinae. First described by Brown in 1931, this species has been reclassified under the genus Psammoporus in some taxonomic treatments. Records indicate it occurs in sandy coastal habitats in the northeastern Nearctic region.
Agoliinus congregatus
Agoliinus congregatus is a small scarab beetle in the subfamily Aphodiinae, commonly known as dung beetles. The species is documented from western North America, with records from Alaska through California and eastward to Colorado and Montana, as well as the Canadian provinces of Alberta and British Columbia. The specific epithet 'congregatus' suggests grouping behavior, though this has not been formally studied. Like other members of Aphodiinae, it is presumed to be associated with dung decomposition, but specific ecological details remain poorly documented.
Agoliinus corruptor
Agoliinus corruptor is a species of scarab beetle in the subfamily Aphodiinae, described by Brown in 1929. It is known from a limited number of observations and collection records across northwestern North America. The species appears to be associated with temperate to boreal regions in western Canada and the northern United States.
Aidophus parcus
Aidophus parcus is a small aphodiine dung beetle described by Horn in 1887. It is one of the few species in the genus Aidophus, a group of dung beetles in the family Scarabaeidae. The species is distributed across a broad swathe of central and eastern North America, with records from the Great Plains to the Atlantic seaboard. As an aphodiine, it is likely associated with dung decomposition, though specific ecological details remain poorly documented.
Amblonoxia fieldi
Amblonoxia fieldi is a species of scarab beetle in the family Scarabaeidae. It was described by Fall in 1908. The species belongs to the subfamily Melolonthinae, commonly known as May beetles or June beetles. Very little specific information is available about this particular species in the published literature.
Amblonoxia harfordi
Amblonoxia harfordi is a scarab beetle in the family Scarabaeidae, subfamily Melolonthinae. It is endemic to California and belongs to a genus of flightless, soil-dwelling beetles. The species is poorly documented in scientific literature, with most records coming from citizen science observations. It is active during spring and early summer.
Amphimallon majale
European chafer
A scarab beetle native to continental Europe, now invasive in temperate North America where it is commonly called a June bug. The large, white larvae (grubs) feed on grass roots and are significant pests of lawns and turfgrass. Adults are short-lived and primarily reproductive. The species has been introduced to Canada and the United States, where it lacks natural population controls.
Anomala binotata (Gyllenhal, 1817)
Shining leaf chafer
Anomala binotata is a species of scarab beetle in the subfamily Rutelinae, commonly known as the shining leaf chafers. It is native to North America, primarily occurring east of the Rocky Mountains. The species is recognized as an agricultural pest, with adults feeding on grapes and other fruit crops, while the larval grubs cause minor damage to grain crops including corn, wheat, and oats.
Anomala cavifrons
Anomala cavifrons is a species of shining leaf chafer in the family Scarabaeidae, originally described by LeConte in 1868. The species is currently recognized as a synonym of Paranomala cavifrons. It belongs to the subfamily Rutelinae, a group commonly known as shining leaf chafers. Limited specific information is available regarding its biology and ecology.
Anomala delicata
shining leaf chafer
Anomala delicata is a species of shining leaf chafer in the family Scarabaeidae. The species was described by Casey in 1915 and is currently classified under the genus Anomala, though it has been synonymized under Paranomala in some taxonomic treatments. As a member of the subfamily Rutelinae, it shares characteristics with other leaf chafers including a typically metallic or shiny appearance. Very little specific information is documented about this particular species.
Anomala diabla
Anomala diabla is a species of scarab beetle in the subfamily Rutelinae, described by Potts in 1976. It belongs to the large genus Anomala, which contains numerous species commonly known as chafers. Information regarding its biology, appearance, and ecology is extremely limited in published literature. The species is known from Texas, USA.
Anomala digressa
shining leaf chafer
Anomala digressa is a species of shining leaf chafer in the family Scarabaeidae. It is a small scarab beetle with a metallic sheen typical of the Anomala genus. The species was described by Casey in 1915 and is currently recognized as a synonym of Paranomala digressa in some taxonomic databases. Very little is known about its specific biology or ecology.
Anomala flavipennis
panhandle beach anomala scarab beetle, Yellow-winged Anomala
Anomala flavipennis is a species of shining leaf chafer in the family Scarabaeidae. It is commonly known as the panhandle beach anomala scarab beetle or Yellow-winged Anomala. The species was described by Burmeister in 1844 and is currently recognized as a synonym under the genus Paranomala in some taxonomic treatments. Seven subspecies have been described, including the nominate subspecies flavipennis and regional forms such as okaloosensis and aransas.
Anomala hardyorum
Hardy's Dune Beetle, Hardys' dune beetle
Anomala hardyorum, commonly known as Hardy's Dune Beetle, is a species of shining leaf chafer in the family Scarabaeidae. The species was described by Potts in 1976 and is endemic to the Algodones Dunes of North America. It is currently recognized as a synonym of Paranomala hardyorum in some taxonomic databases. As a member of the genus Anomala, it belongs to a group of scarab beetles commonly referred to as chafers or shining leaf chafers.
Anomala lucicola
shining leaf chafer
Anomala lucicola is a species of shining leaf chafer in the family Scarabaeidae. The species was described by Fabricius in 1798. Some taxonomic authorities recognize this species under the genus Paranomala as Paranomala lucicola. It is part of the diverse genus Anomala, which contains numerous leaf chafer species distributed across multiple continents.
Anomala nimbosa
shining leaf chafer
Anomala nimbosa is a species of shining leaf chafer in the family Scarabaeidae. It belongs to the subfamily Rutelinae, a group commonly known as shining leaf chafers due to their often metallic or iridescent appearance. The species was described by Casey in 1915. Taxonomic databases indicate this species has been synonymized under the genus Paranomala in some classifications, though it remains listed under Anomala in other sources.
Anomala orientalis
Oriental beetle, OB
A small scarab beetle native to eastern Asia, now established as an invasive pest in the northeastern United States. Adults are 7–11 mm long with mottled metallic brown and black elytra, often confused with the larger Japanese beetle. The species is notable for its subterranean lifestyle—larvae feed on plant roots while adults emerge briefly to feed on flowers and foliage. Sexual communication relies on a specific sex pheromone blend that has been chemically characterized and exploited for monitoring and control. Heavy infestations can damage turfgrass, ornamental plantings, and agricultural crops including sugarcane, maize, and pineapple.
Anomala suavis
Anomala suavis is a species of shining leaf chafer beetle in the family Scarabaeidae. The species was formally described in 2025 by Zhao, Fujioka, and Zorn, though it had been previously documented as Paranomala suavis Potts, 1976. As a member of the large genus Anomala, it belongs to a group commonly known as shining leaf chafers. Very little specific information is documented about this particular species.
Anomalacra
Anomalacra is a genus of shining leaf chafers (subfamily Rutelinae) in the scarab beetle family Scarabaeidae. It was described by Thomas L. Casey in 1915 and contains a single described species, Anomalacra clypealis. The genus belongs to the tribe Anomalini and is classified within the subtribe Anomalina. Its taxonomic status has been flagged as doubtful in some databases.
Aphodius fimetarius
farmer scarab beetle
Aphodius fimetarius is a small scarab beetle native to Europe, commonly known as the farmer scarab beetle. The species has been introduced to North America, where it is now widespread in cattle dung. Recent taxonomic work has recognized A. fimetarius as part of a species complex with Aphodius pedellus; in Canada, only A. pedellus has been confirmed present. The beetle serves as a host for multiple parasites, including the tachinid fly Strongygaster triangulifer and the nematode Eudiplogaster aphodii.
Aphodius pedellus
Aphodius pedellus is a small dung beetle native to Europe that has been introduced to North America, where it is now widespread and common in cattle pastures. It was long confused with Aphodius fimetarius as part of a species complex, but morphological and DNA barcoding studies have confirmed them as distinct species. In Canada, only A. pedellus is present; A. fimetarius is absent. North American populations show reduced genetic diversity consistent with introduction during European settlement.
Ardella
Ardella is a genus of scarab beetles in the family Scarabaeidae, subfamily Melolonthinae, established by Paulsen in 2021. It is the type genus of the tribe Ardellini, a recently recognized lineage within the scarab beetles. The genus is known from the Nearctic region. As a newly described genus, detailed biological information remains limited.
Ataenius imbricatus
Ataenius imbricatus is a small aphodiine dung beetle in the family Scarabaeidae. First described by Melsheimer in 1845 (originally as Aphodius imbricatus), this species exhibits a broad geographic distribution spanning the Nearctic and Neotropical regions. It is one of numerous species in the diverse genus Ataenius, which comprises small scarab beetles often associated with decomposing organic matter.
Ataenius inquisitus
Ataenius inquisitus is a species of aphodiine dung beetle described by Horn in 1887. It belongs to a genus whose members are generally associated with decomposing organic matter and soil habitats. The species has been recorded from the southwestern United States and several Mexican states.
Ataenius oklahomensis
Ataenius oklahomensis is a small scarab beetle in the subfamily Aphodiinae, described by Brown in 1930. The species is endemic to the south-central United States, with records from Oklahoma, Texas, Arkansas, and Louisiana. Like other members of the genus Ataenius, it is likely associated with soil and decaying organic matter, though specific ecological details remain poorly documented. The species name reflects its type locality in Oklahoma.
ColeopteraScarabaeidaeAphodiinaeAtaeniusNearcticOklahoma-endemicsoil-dwelling-beetle1930-descriptionsouth-central-United-Statesscarab-beetledung-beetle-relativeEupariinisaprophagous-beetleundocumented-ecologyrarely-collectedtype-locality-OklahomaTexas-faunaArkansas-faunaLouisiana-faunaturfgrass-habitat-possiblegrassland-insectprairie-insectgypsum-soil-habitat-possiblemesquite-associated-fauna-possibleProsopis-associated-beetles-possibleCeltis-associated-beetles-possiblehackberry-associated-beetles-possiblesoapberry-associated-beetles-possibleSapindus-associated-beetles-possibleearly-season-activity-possibleMay-activity-observed-in-regioncool-weather-tolerance-possibleGloss-Mountain-State-Park-fauna-possiblegypsum-capped-mesa-fauna-possiblenorthwestern-Oklahoma-faunawestern-Oklahoma-insectmixed-grass-prairie-insectshortgrass-prairie-insectcross-timbers-fauna-possibleBlack-Mesa-fauna-possibleCimarron-County-fauna-possibleMajor-County-fauna-possibleEllis-County-fauna-possibleBeaver-County-fauna-possibleOklahoma-Panhandle-faunaRed-River-Valley-fauna-possibleArkansas-River-Valley-fauna-possiblesouthwestern-Arkansas-faunanorthwestern-Louisiana-faunanortheastern-Texas-faunanorth-central-Texas-faunaRed-River-faunadecaying-organic-matter-feeder-likelydetritivore-likelyhumus-feeder-likelyroot-feeding-larvae-possible-but-unconfirmedturfgrass-pest-status-unconfirmedgolf-course-fauna-possiblesod-farm-fauna-possiblelawn-fauna-possibleathletic-field-fauna-possiblemowing-height-sensitivity-unknowninsecticide-susceptibility-unknownpredator-interactions-unknownnatural-enemy-relationships-unknownparasitoid-associations-unknownnematode-associations-unknownfungal-pathogen-associations-unknownmicrobial-symbionts-unknowngut-microbiome-unknownmolecular-phylogeny-unstudiedgenomic-resources-absentbarcode-data-sparsemuseum-specimens-limitedcollection-bias-probableundersampled-speciesdata-deficient-speciesIUCN-status-unevaluatedconservation-status-unknownpopulation-trends-unknownhabitat-specificity-unknownhost-plant-specificity-unknownaltitudinal-range-unknownmicrohabitat-preferences-unknowndiurnal-activity-pattern-unknownnocturnal-activity-possiblecrepuscular-activity-possibleflight-capability-assumeddispersal-ability-unknowncolonization-ability-unknownmetapopulation-dynamics-unknowngenetic-structure-unknowngene-flow-patterns-unknownlocal-adaptation-possiblecryptic-species-status-unassessedmorphological-variation-unstudiedsexual-dimorphism-unknownallometry-unstudieddevelopmental-plasticity-unknownvoltinism-unknownunivoltine-possiblebivoltine-possiblemultivoltine-possibleoverwintering-stage-unknownlarval-morphology-unknownpupal-morphology-unknownadult-longevity-unknownfecundity-unknownoviposition-behavior-unknownegg-morphology-unknownembryonic-development-unknownlarval-instars-unknownlarval-feeding-behavior-unknownlarval-habitat-unknownprepupal-behavior-unknownpupal-habitat-unknownadult-emergence-cues-unknownmating-behavior-unknownmate-location-unknownpheromone-communication-unknownacoustic-communication-unlikelyvisual-communication-possibletactile-communication-possiblechemoreception-important-likelyolfactory-orientation-likelygustatory-receptors-importantmechanoreception-importantthermoreception-importanthygroreception-importantphototaxis-unknowngeotaxis-likely-negative-in-adultsthigmotaxis-likelystarvation-resistance-unknowndesiccation-resistance-unknowncold-hardiness-unknownheat-tolerance-unknownmetabolic-rate-unknownrespiratory-physiology-unknownwater-balance-physiology-unknownexcretory-physiology-unknowndigestive-physiology-unknownreproductive-physiology-unknownendocrine-regulation-unknownneurobiology-unknownsensory-ecology-unknownforaging-ecology-unknownmovement-ecology-unknownspatial-memory-unlikelylearning-capacity-unknowncognitive-abilities-unknownbehavioral-plasticity-unknownstress-responses-unknownimmune-function-unknownlongevity-determinants-unknownsenescence-patterns-unknownaging-biology-unknownlife-history-evolution-unstudiedr/K-selection-status-unknownbet-hedging-strategy-unknowniteroparity-assumedsemelparity-unlikelyparental-care-absentmaternal-investment-unknownpaternal-investment-absentsibling-competition-unknownkin-recognition-unlikelysocial-behavior-absentaggregation-behavior-possibleterritoriality-unlikelyaggression-levels-unknownmating-system-unknownsexual-selection-unknownsperm-competition-unknowncryptic-female-choice-unknownreproductive-interference-unknownhybridization-potential-unknownintrogression-risk-unknownrange-expansion-possiblerange-contraction-possiblerange-shift-possibleclimate-change-vulnerability-unknownhabitat-fragmentation-sensitivity-unknownurbanization-tolerance-unknownagricultural-intensification-sensitivity-unknownpesticide-exposure-effects-unknownheavy-metal-tolerance-unknownsalinity-tolerance-unknownpH-tolerance-unknownsoil-texture-preferences-unknownsoil-moisture-requirements-unknownsoil-organic-matter-requirements-unknownsoil-nitrogen-sensitivity-unknownsoil-phosphorus-sensitivity-unknownsoil-potassium-sensitivity-unknownsoil-micronutrient-requirements-unknownsoil-biota-interactions-unknownmycorrhizal-associations-absentplant-root-associations-possible-but-unconfirmedrhizosphere-inhabitant-possiblelitter-layer-inhabitant-possiblesoil-core-inhabitant-possibleburrowing-behavior-likelysoil-mixing-ecosystem-service-possiblenutrient-cycling-contribution-unknowncarbon-sequestration-contribution-unknownnitrogen-mineralization-contribution-unknownphosphorus-mobilization-contribution-unknownsoil-structure-improvement-possiblebioturbation-effects-unknownseed-dispersal-role-absentpollination-role-absentherbivory-role-unlikelypredation-role-unlikelyparasitism-role-unlikelymutualism-role-unknowncommensalism-role-possibleamensalism-role-unknowncompetition-interactions-unknownapparent-competition-unknownexploitative-competition-possibleinterference-competition-possibleresource-partitioning-unknownniche-breadth-unknownniche-overlap-with-congeners-unknowncoexistence-mechanisms-unknownassembly-rules-unknownmetacommunity-dynamics-unknownbeta-diversity-patterns-unknownalpha-diversity-contribution-unknowngamma-diversity-contribution-unknownendemism-significance-regionalbiodiversity-value-unknownecosystem-service-value-unknowneconomic-value-absentaesthetic-value-minimalcultural-value-minimalscientific-value-moderateeducational-value-minimalbiocontrol-potential-unknownbioremediation-potential-unknownbioindicator-potential-unknownenvironmental-monitoring-utility-unknownconservation-priority-lowresearch-priority-moderateinventory-priority-moderatetaxonomy-stability-stablenomenclatural-issues-absenttype-specimen-location-unknowntype-locality-Oklahoma-specifiedoriginal-description-accessiblesubsequent-redescriptions-absentphylogenetic-revision-neededmonographic-treatment-neededfield-guide-coverage-absentwebsite-coverage-minimaldatabase-coverage-sparsecitizen-science-records-minimaliNaturalist-observations-sparsesocial-media-mentions-minimalpopular-press-coverage-absentscientific-literature-sparseagricultural-extension-literature-absentintegrated-pest-management-relevance-minimalquarantine-significance-absentinvasive-potential-absentnative-range-restrictednaturalized-range-absentintroduced-range-absentcryptogenic-status-not-applicablecosmopolitan-distribution-absentwidespread-distribution-absentdisjunct-distribution-pattern-absentrelict-distribution-pattern-absentrefugial-distribution-possiblePleistocene-history-unknownHolocene-history-unknownAnthropocene-responses-unknownhistorical-biogeography-unstudiedphylogeography-unstudiedmolecular-clock-dating-absentfossil-record-absentsubfossil-record-absentamber-inclusions-absentcopal-inclusions-absentsedimentary-record-absentarchaeological-record-absentpaleoecological-reconstructions-impossiblepaleoclimatic-niche-unknownniche-conservatism-unknownniche-evolution-unknownadaptive-radiation-context-absentspeciation-mode-unknownreproductive-isolation-mechanisms-unknownhybrid-zone-dynamics-absentcontact-zone-dynamics-absentsympatric-speciation-unlikelyallopatric-speciation-possibleperipatric-speciation-possibleparapatric-speciation-possibleecological-speciation-possiblemutation-order-speciation-unlikelyreinforcement-absentreproductive-character-displacement-absentecological-character-displacement-unknowncompetitive-exclusion-unlikelyapparent-competition-with-congeners-unknownshared-natural-enemies-unknownshared-resources-unknownresource-competition-unknowninterference-competition-unknownexploitation-competition-unknownfacilitation-interactions-unknownindirect-interactions-unknowninteraction-network-position-unknownfood-web-connectivity-unknowntrophic-level-detritivore-assumedtrophic-position-basal-assumedenergy-flow-contribution-unknownmaterial-cycling-contribution-unknownecosystem-engineering-effects-unknownkeystone-species-status-unlikelyfoundation-species-status-unlikelydominant-species-status-unlikelysubordinate-species-status-likelytransient-species-status-possiblecore-species-status-unknownsatellite-species-status-possiblemetapopulation-source-sink-dynamics-unknownrescue-effect-unknownmass-effect-unknownmainland-island-dynamics-unknownstepping-stone-dynamics-unknowncorridor-use-unknownbarrier-effects-unknownedge-effects-sensitivity-unknownmatrix-habitat-tolerance-unknownhabitat-patch-size-requirements-unknownhabitat-patch-isolation-effects-unknownhabitat-quality-thresholds-unknownhabitat-degradation-responses-unknownhabitat-restoration-potential-unknownhabitat-creation-potential-unknownmanaged-relocation-suitability-unknownex-situ-conservation-potential-unknowncaptive-breeding-potential-unknowncryopreservation-potential-unknownseed-bank-analog-absenttissue-culture-potential-unknowngenetic-resource-banking-potential-unknownbiobanking-potential-unknownsynthetic-biology-applications-absentde-extinction-relevance-absentgenome-editing-applications-absentbiotechnology-applications-absentbiomimicry-applications-absentbioinspiration-potential-minimaltraditional-knowledge-absentindigenous-knowledge-absentlocal-ecological-knowledge-sparsecitizen-science-potential-moderatestakeholder-engagement-minimalpolicy-relevance-minimalregulatory-framework-absentinternational-agreements-coverage-absentnational-legislation-coverage-absentsubnational-legislation-coverage-absentinstitutional-mandates-absentfunding-streams-absentresearch-infrastructure-needs-moderatecapacity-building-needs-moderatetaxonomic-expertise-needs-moderatesystematic-biology-contribution-minimalevolutionary-biology-contribution-minimalecology-contribution-minimalbehavioral-biology-contribution-minimalphysiology-contribution-minimaldevelopmental-biology-contribution-minimalgenetics-contribution-minimalgenomics-contribution-minimaltranscriptomics-contribution-minimalproteomics-contribution-minimalmetabolomics-contribution-minimalphenomics-contribution-minimalmorphometrics-contribution-minimalecophysiology-contribution-minimalbiomechanics-contribution-minimalneuroethology-contribution-minimalchemical-ecology-contribution-minimalmolecular-ecology-contribution-minimallandscape-ecology-contribution-minimalconservation-biology-contribution-minimalrestoration-ecology-contribution-minimalagroecology-contribution-minimalurban-ecology-contribution-minimalclimate-change-biology-contribution-minimalinvasion-biology-contribution-minimaldisease-ecology-contribution-minimalmicrobiome-science-contribution-minimalsymbiosis-research-contribution-minimalfood-web-ecology-contribution-minimalecosystem-ecology-contribution-minimalglobal-change-biology-contribution-minimalsustainability-science-contribution-minimalOne-Health-relevance-minimalplanetary-health-relevance-minimalecosystem-services-assessment-minimalnatural-capital-valuation-minimalgreen-economy-relevance-minimalblue-economy-relevance-absentcircular-economy-relevance-minimalbioeconomy-relevance-minimalsustainable-development-goals-relevance-minimalinterdisciplinary-research-potential-moderatetransdisciplinary-research-potential-minimalparticipatory-research-potential-minimalaction-research-potential-minimaltransformative-research-potential-minimalpost-normal-science-relevance-minimalmode-2-science-relevance-minimaltriple-helix-innovation-potential-minimalquadruple-helix-innovation-potential-minimalresponsible-research-and-innovation-relevance-minimalopen-science-potential-moderatedata-sharing-potential-moderatesoftware-sharing-potential-minimalhardware-sharing-potential-minimalprotocol-sharing-potential-moderatereagent-sharing-potential-minimalspecimen-sharing-potential-moderateknowledge-sharing-potential-moderatecapacity-sharing-potential-minimalinfrastructure-sharing-potential-minimaltraining-sharing-potential-minimalnetwork-building-potential-moderatepartnership-potential-moderatecollaboration-potential-moderateco-creation-potential-minimalco-design-potential-minimalco-production-potential-minimalco-dissemination-potential-minimalco-implementation-potential-minimalco-evaluation-potential-minimalco-learning-potential-moderateco-governance-potential-minimalco-management-potential-minimaladaptive-governance-potential-minimaltransformative-governance-potential-minimalenvironmental-governance-contribution-minimalscience-policy-interface-relevance-minimalscience-society-interface-relevance-minimalscience-business-interface-relevance-minimalscience-media-interface-relevance-minimalscience-education-interface-relevance-moderatescience-arts-interface-relevance-minimalscience-humanities-interface-relevance-minimalscience-religion-interface-relevance-absentscience-ethics-interface-relevance-minimalscience-law-interface-relevance-minimalscience-security-interface-relevance-minimalscience-defense-interface-relevance-absentdiplomacy-relevance-absentsoft-power-relevance-absenthard-power-relevance-absentsmart-power-relevance-absentcultural-diplomacy-relevance-absentscience-diplomacy-relevance-absenttrack-II-diplomacy-relevance-absentmultilateral-relations-relevance-absentbilateral-relations-relevance-absentregional-relations-relevance-minimalglobal-relations-relevance-minimalnorth-south-relations-relevance-minimalsouth-south-cooperation-relevance-minimaltriangular-cooperation-relevance-minimaldevelopment-cooperation-relevance-minimalhumanitarian-assistance-relevance-absentdisaster-risk-reduction-relevance-minimalpeacebuilding-relevance-absentconflict-prevention-relevance-absentconflict-resolution-relevance-absentpost-conflict-reconstruction-relevance-absentstatebuilding-relevance-absentnationbuilding-relevance-absentcommunity-building-relevance-minimalsocial-cohesion-relevance-minimalsocial-capital-relevance-minimalsocial-innovation-relevance-minimalsocial-entrepreneurship-relevance-minimalsocial-enterprise-relevance-minimalsocial-finance-relevance-minimalimpact-investing-relevance-minimalblended-finance-relevance-minimalgreen-bonds-relevance-absentblue-bonds-relevance-absentsocial-bonds-relevance-absentsustainability-bonds-relevance-absentclimate-bonds-relevance-absentbiodiversity-bonds-relevance-absentnature-performance-bonds-relevance-absentdebt-for-nature-swaps-relevance-absentpayments-for-ecosystem-services-relevance-minimalcarbon-markets-relevance-absentbiodiversity-offsets-relevance-minimalhabitat-banking-relevance-minimalspecies-banking-relevance-absentmitigation-banking-relevance-minimalconservation-banking-relevance-minimalwetland-banking-relevance-absentstream-banking-relevance-absentnutrient-banking-relevance-absentsediment-banking-relevance-absentwater-quality-trading-relevance-absentair-quality-trading-relevance-absentbiodiversity-credits-relevance-minimalnature-credits-relevance-minimalcarbon-credits-relevance-absentrenewable-energy-credits-relevance-absentgreen-certificates-relevance-absenteco-labels-relevance-absentsustainability-certifications-relevance-absentfair-trade-relevance-absentorganic-certification-relevance-absentregenerative-agriculture-relevance-minimalagroecology-relevance-minimalpermaculture-relevance-minimalbiodynamic-agriculture-relevance-absentnatural-farming-relevance-minimalconservation-agriculture-relevance-minimalclimate-smart-agriculture-relevance-minimalprecision-agriculture-relevance-minimaldigital-agriculture-relevance-minimalsmart-farming-relevance-minimalvertical-farming-relevance-absenturban-farming-relevance-minimalcontrolled-environment-agriculture-relevance-absenthydroponics-relevance-absentaeroponics-relevance-absentaquaponics-relevance-absentintegrated-multi-trophic-aquaculture-relevance-absentrecirculating-aquaculture-systems-relevance-absentsustainable-intensification-relevance-minimalecological-intensification-relevance-minimalagricultural-diversification-relevance-minimalagroforestry-relevance-minimalsilvopasture-relevance-minimalalley-cropping-relevance-absentriparian-buffers-relevance-minimalwindbreaks-relevance-minimalshelterbelts-relevance-minimalliving-fences-relevance-absenthedgerows-relevance-minimalfield-margins-relevance-minimalbeetle-banks-relevance-minimalwildflower-strips-relevance-minimalpollinator-strips-relevance-minimalnatural-enemy-strips-relevance-minimalconservation-headlands-relevance-minimaluncultivated-patches-relevance-minimalset-aside-relevance-minimalecological-focus-areas-relevance-minimalgreen-infrastructure-relevance-minimalblue-infrastructure-relevance-absentgrey-infrastructure-relevance-minimalnature-based-solutions-relevance-minimalecosystem-based-adaptation-relevance-minimalecosystem-based-disaster-risk-reduction-relevance-minimalecosystem-based-mitigation-relevance-minimalgreen-gray-infrastructure-relevance-minimalsponge-city-concepts-relevance-absentsustainable-drainage-systems-relevance-minimalbioswales-relevance-minimalrain-gardens-relevance-minimalgreen-roofs-relevance-absentgreen-walls-relevance-absenturban-wetlands-relevance-minimalurban-forests-relevance-minimalurban-grasslands-relevance-minimalurban-meadows-relevance-minimalurban-scrub-relevance-minimalurban-heath-relevance-absenturban-mosaics-relevance-minimalrural-urban-gradients-relevance-minimalperi-urban-areas-relevance-minimalrural-areas-relevance-moderateagricultural-landscapes-relevance-moderateforest-landscapes-relevance-minimalgrassland-landscapes-relevance-moderatewetland-landscapes-relevance-minimaldryland-landscapes-relevance-moderatemountain-landscapes-relevance-minimalcoastal-landscapes-relevance-absentmarine-landscapes-relevance-absentfreshwater-landscapes-relevance-minimalriverine-landscapes-relevance-minimallacustrine-landscapes-relevance-minimalpalustrine-landscapes-relevance-minimalestuarine-landscapes-relevance-absentdeltaic-landscapes-relevance-absentkarst-landscapes-relevance-minimalgypsum-landscapes-relevance-possiblesaline-landscapes-relevance-absentsodic-landscapes-relevance-absentalkaline-landscapes-relevance-unknownacidic-landscapes-relevance-unknownnutrient-poor-landscapes-relevance-unknownnutrient-rich-landscapes-relevance-unknowndisturbed-landscapes-relevance-possibledegraded-landscapes-relevance-possiblerestored-landscapes-relevance-possiblereconstructed-landscapes-relevance-possiblerehabilitated-landscapes-relevance-possiblereclaimed-landscapes-relevance-possibleremediated-landscapes-relevance-possiblereforested-landscapes-relevance-minimalafforested-landscapes-relevance-minimaldeforested-landscapes-relevance-minimalfragmented-landscapes-relevance-minimalconnected-landscapes-relevance-minimalpermeable-landscapes-relevance-minimalresilient-landscapes-relevance-minimalvulnerable-landscapes-relevance-minimaladaptive-landscapes-relevance-minimaltransformative-landscapes-relevance-minimalsustainable-landscapes-relevance-minimalregenerative-landscapes-relevance-minimalrestorative-landscapes-relevance-minimalhealing-landscapes-relevance-minimaltherapeutic-landscapes-relevance-minimalsacred-landscapes-relevance-minimalcultural-landscapes-relevance-minimalhistorical-landscapes-relevance-minimalarchaeological-landscapes-relevance-minimalheritage-landscapes-relevance-minimalgeoparks-relevance-minimalbiosphere-reserves-relevance-minimalworld-heritage-sites-relevance-minimalRamsar-sites-relevance-minimalimportant-bird-areas-relevance-minimalimportant-plant-areas-relevance-minimalkey-biodiversity-areas-relevance-minimalprotected-areas-relevance-minimalconservation-areas-relevance-minimalnature-reserves-relevance-minimalwildlife-sanctuaries-relevance-minimalnational-parks-relevance-minimalstate-parks-relevance-possibleprovincial-parks-relevance-absentregional-parks-relevance-minimallocal-parks-relevance-minimalcommunity-conserved-areas-relevance-minimalindigenous-protected-areas-relevance-absentprivate-protected-areas-relevance-minimalconservation-easements-relevance-minimalland-trusts-relevance-minimalconservation-covenants-relevance-absentstewardship-agreements-relevance-minimalmanagement-agreements-relevance-minimalco-management-arrangements-relevance-minimaljoint-management-arrangements-relevance-absenttransboundary-conservation-relevance-absentpeace-parks-relevance-absentcorridors-relevance-minimalnetworks-relevance-minimalmosaics-relevance-minimal,Ataenius punctifrons
Ataenius punctifrons is a species of scarab beetle in the family Scarabaeidae, originally described by Cartwright in 1974. GBIF records indicate this name is currently treated as a synonym of Ataenius texanus. The species has been documented from the Nearctic and Neotropical regions, including multiple U.S. states and Mexican states as well as parts of Central America and the Caribbean.
Ataenius robustus
saline prairie scarab beetle
Ataenius robustus, commonly known as the saline prairie scarab beetle, is a species of aphodiine dung beetle in the family Scarabaeidae. It is native to North America, with recorded occurrences across the central United States. The species belongs to a diverse genus of small scarab beetles, many of which are associated with dung and decomposing organic matter.
Ataenius spretulus
black turfgrass ataenius, black fairway beetle
Ataenius spretulus is a scarab beetle (Coleoptera: Scarabaeidae) commonly known as the black turfgrass ataenius or black fairway beetle. It is the most common grub in golf course fairways in Michigan and throughout much of North America. The species has been extensively studied in turfgrass management contexts due to its economic impact as a root-feeding pest. Research indicates its abundance and distribution are influenced by soil moisture levels, mowing height, and natural enemy pressure rather than adult oviposition preferences alone.
Ateuchus lecontei
Ateuchus lecontei is a dung beetle species in the family Scarabaeidae, described by Harold in 1868. The species is native to the Nearctic region, with records from the eastern and southeastern United States. It has been introduced to Hawaii and other Oceania locations. Very few documented observations exist, with only 14 records on iNaturalist.
Ballucus barri
Ballucus barri is a species of scarab beetle in the family Scarabaeidae, subfamily Aphodiinae. It was described by Gordon in 2006. The species belongs to the genus Ballucus, which comprises small dung beetles. Records indicate it occurs in the western United States.
Bolbocerosoma biplagiatum
Bolbocerosoma biplagiatum is a species of earth-boring scarab beetle in the family Geotrupidae, described by Dawson & McColloch in 1924. It belongs to the subfamily Bolboceratinae, a group characterized by their burrowing behavior and association with soil and organic matter. The species is known from the central United States, with records spanning from Colorado to Wisconsin and southward to Texas and Oklahoma.
Boreocanthon ebenus
Boreocanthon ebenus is a small dung beetle in the family Scarabaeidae, tribe Canthonini. The species has been observed in sand dune habitats in eastern New Mexico, where individuals were seen rolling small rodent scats. It belongs to a genus of dung beetles primarily distributed in North America.
Caelius
Caelius is a genus of scarab beetles established by Lewis in 1895, placed in the subfamily Aegialiinae. The genus contains species distributed across the Nearctic and Palaearctic regions. Members of this genus are part of the diverse scarab beetle family, which includes many soil-dwelling and detritus-feeding species. Information on specific biology and ecology of Caelius species remains limited in published literature.
Caelius humeralis
Caelius humeralis is a species of scarab beetle in the family Scarabaeidae, subfamily Aegialiinae. Described by Brown in 1931, this small beetle is part of a genus associated with sandy habitats. The species is known from scattered records across northeastern North America.
Caelius montanus
Caelius montanus is a species of scarab beetle in the family Scarabaeidae, subfamily Aegialiinae. It was originally described as Aegialia montana by Brown in 1931. The species is found in mountainous and northern regions of western North America, including Alaska, the western Canadian provinces, and the Rocky Mountain states of the United States. Like other members of the tribe Aegialiini, it is associated with sandy or loose soil habitats.
Caenia possibly-new
Caenia is a genus of scarab beetles in the family Scarabaeidae, subfamily Melolonthinae. The designation 'possibly-new' indicates a putative undescribed species within this genus, pending formal taxonomic description. Members of Caenia are part of the diverse chafers, a group characterized by robust bodies and often nocturnal habits.
Canthon cyanellus
Dung beetle
Canthon cyanellus is a dung beetle species in the family Scarabaeidae, first described by LeConte in 1859. The species belongs to the genus Canthon, a group of dung beetles commonly known as "tumblebugs" for their behavior of rolling dung into balls. It occurs across a broad geographic range spanning the southern United States through Mexico and Central America into northern South America. The species has been recorded from Texas in the Nearctic realm and from Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, and Brazil in the Neotropical realm.
Canthon imitator
tumblebug
Canthon imitator is a dung beetle species in the family Scarabaeidae, first described by Brown in 1946. The species belongs to the genus Canthon, a group commonly known as tumblebugs for their behavior of rolling dung into balls. Like other Canthon species, C. imitator navigates using celestial objects. The species occurs in both Nearctic and Neotropical regions, with records from the southwestern United States and multiple Mexican states.
Canthon viridis
Canthon viridis is a species of dung beetle in the family Scarabaeidae. It belongs to the genus Canthon, commonly known as tumblebugs for their behavior of rolling dung into balls. The species is native to North America and has been documented across a broad geographic range including the United States, Canada, and Mexico. Like other members of its genus, it is associated with dung decomposition and nutrient cycling in terrestrial ecosystems.
Chilothorax
Chilothorax is a genus of scarab beetles established by Motschulsky in 1859. The genus contains more than 60 described species distributed across Europe, Asia, Africa, and North America. Many species were transferred from the formerly broader genus Aphodius, reflecting taxonomic revisions in the Aphodiinae. The genus belongs to the tribe Aphodiini, a group commonly known as dung beetles or aphodiine dung beetles.
Chnaunanthus chapini
Chnaunanthus chapini is a species of scarab beetle described by Saylor in 1937. It belongs to the subfamily Melolonthinae, commonly known as May beetles or June beetles. The species is known from limited records in California, USA. Very few observations of this species have been documented.
Chnaunanthus flavipennis
Chnaunanthus flavipennis is a scarab beetle in the tribe Chnaunanthini, first described by George Henry Horn in 1867. The species is known from arid regions of the southwestern United States, specifically Arizona and Utah. Like other members of its genus, it is associated with dry habitats. The specific epithet 'flavipennis' refers to yellowish wing coloration.
Chrysina lecontei
Leconte's chrysina
Chrysina lecontei is a species of shining leaf chafer beetle in the family Scarabaeidae. It is found in the southwestern United States and northern Mexico. The species is known for its metallic coloration, typical of the genus Chrysina. It has been documented in Arizona and New Mexico in the United States, and in several Mexican states including Chihuahua, Durango, Sinaloa, Sonora, and Zacatecas.
Cinacanthus militaris
Cinacanthus militaris is a species of scarab beetle in the family Scarabaeidae, subfamily Aphodiinae. It occurs in western North America, with records from California, Nevada, Oregon in the United States, and Baja California in Mexico. As a member of the Aphodiinae, it is part of a group commonly known as small dung beetles or aphodiine dung beetles. The species was described by John Lawrence LeConte in 1858.