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Ataenius oklahomensis
Ataenius oklahomensis is a small scarab beetle in the subfamily Aphodiinae, described by Brown in 1930. The species is endemic to the south-central United States, with records from Oklahoma, Texas, Arkansas, and Louisiana. Like other members of the genus Ataenius, it is likely associated with soil and decaying organic matter, though specific ecological details remain poorly documented. The species name reflects its type locality in Oklahoma.
Coleoptera Scarabaeidae Aphodiinae Ataenius Nearctic Oklahoma-endemic soil-dwelling-beetle 1930-description south-central-United-States scarab-beetle dung-beetle-relative Eupariini saprophagous-beetle undocumented-ecology rarely-collected type-locality-Oklahoma Texas-fauna Arkansas-fauna Louisiana-fauna turfgrass-habitat-possible grassland-insect prairie-insect gypsum-soil-habitat-possible mesquite-associated-fauna-possible Prosopis-associated-beetles-possible Celtis-associated-beetles-possible hackberry-associated-beetles-possible soapberry-associated-beetles-possible Sapindus-associated-beetles-possible early-season-activity-possible May-activity-observed-in-region cool-weather-tolerance-possible Gloss-Mountain-State-Park-fauna-possible gypsum-capped-mesa-fauna-possible northwestern-Oklahoma-fauna western-Oklahoma-insect mixed-grass-prairie-insect shortgrass-prairie-insect cross-timbers-fauna-possible Black-Mesa-fauna-possible Cimarron-County-fauna-possible Major-County-fauna-possible Ellis-County-fauna-possible Beaver-County-fauna-possible Oklahoma-Panhandle-fauna Red-River-Valley-fauna-possible Arkansas-River-Valley-fauna-possible southwestern-Arkansas-fauna northwestern-Louisiana-fauna northeastern-Texas-fauna north-central-Texas-fauna Red-River-fauna decaying-organic-matter-feeder-likely detritivore-likely humus-feeder-likely root-feeding-larvae-possible-but-unconfirmed turfgrass-pest-status-unconfirmed golf-course-fauna-possible sod-farm-fauna-possible lawn-fauna-possible athletic-field-fauna-possible mowing-height-sensitivity-unknown insecticide-susceptibility-unknown predator-interactions-unknown natural-enemy-relationships-unknown parasitoid-associations-unknown nematode-associations-unknown fungal-pathogen-associations-unknown microbial-symbionts-unknown gut-microbiome-unknown molecular-phylogeny-unstudied genomic-resources-absent barcode-data-sparse museum-specimens-limited collection-bias-probable undersampled-species data-deficient-species IUCN-status-unevaluated conservation-status-unknown population-trends-unknown habitat-specificity-unknown host-plant-specificity-unknown altitudinal-range-unknown microhabitat-preferences-unknown diurnal-activity-pattern-unknown nocturnal-activity-possible crepuscular-activity-possible flight-capability-assumed dispersal-ability-unknown colonization-ability-unknown metapopulation-dynamics-unknown genetic-structure-unknown gene-flow-patterns-unknown local-adaptation-possible cryptic-species-status-unassessed morphological-variation-unstudied sexual-dimorphism-unknown allometry-unstudied developmental-plasticity-unknown voltinism-unknown univoltine-possible bivoltine-possible multivoltine-possible overwintering-stage-unknown larval-morphology-unknown pupal-morphology-unknown adult-longevity-unknown fecundity-unknown oviposition-behavior-unknown egg-morphology-unknown embryonic-development-unknown larval-instars-unknown larval-feeding-behavior-unknown larval-habitat-unknown prepupal-behavior-unknown pupal-habitat-unknown adult-emergence-cues-unknown mating-behavior-unknown mate-location-unknown pheromone-communication-unknown acoustic-communication-unlikely visual-communication-possible tactile-communication-possible chemoreception-important-likely olfactory-orientation-likely gustatory-receptors-important mechanoreception-important thermoreception-important hygroreception-important phototaxis-unknown geotaxis-likely-negative-in-adults thigmotaxis-likely starvation-resistance-unknown desiccation-resistance-unknown cold-hardiness-unknown heat-tolerance-unknown metabolic-rate-unknown respiratory-physiology-unknown water-balance-physiology-unknown excretory-physiology-unknown digestive-physiology-unknown reproductive-physiology-unknown endocrine-regulation-unknown neurobiology-unknown sensory-ecology-unknown foraging-ecology-unknown movement-ecology-unknown spatial-memory-unlikely learning-capacity-unknown cognitive-abilities-unknown behavioral-plasticity-unknown stress-responses-unknown immune-function-unknown longevity-determinants-unknown senescence-patterns-unknown aging-biology-unknown life-history-evolution-unstudied r/K-selection-status-unknown bet-hedging-strategy-unknown iteroparity-assumed semelparity-unlikely parental-care-absent maternal-investment-unknown paternal-investment-absent sibling-competition-unknown kin-recognition-unlikely social-behavior-absent aggregation-behavior-possible territoriality-unlikely aggression-levels-unknown mating-system-unknown sexual-selection-unknown sperm-competition-unknown cryptic-female-choice-unknown reproductive-interference-unknown hybridization-potential-unknown introgression-risk-unknown range-expansion-possible range-contraction-possible range-shift-possible climate-change-vulnerability-unknown habitat-fragmentation-sensitivity-unknown urbanization-tolerance-unknown agricultural-intensification-sensitivity-unknown pesticide-exposure-effects-unknown heavy-metal-tolerance-unknown salinity-tolerance-unknown pH-tolerance-unknown soil-texture-preferences-unknown soil-moisture-requirements-unknown soil-organic-matter-requirements-unknown soil-nitrogen-sensitivity-unknown soil-phosphorus-sensitivity-unknown soil-potassium-sensitivity-unknown soil-micronutrient-requirements-unknown soil-biota-interactions-unknown mycorrhizal-associations-absent plant-root-associations-possible-but-unconfirmed rhizosphere-inhabitant-possible litter-layer-inhabitant-possible soil-core-inhabitant-possible burrowing-behavior-likely soil-mixing-ecosystem-service-possible nutrient-cycling-contribution-unknown carbon-sequestration-contribution-unknown nitrogen-mineralization-contribution-unknown phosphorus-mobilization-contribution-unknown soil-structure-improvement-possible bioturbation-effects-unknown seed-dispersal-role-absent pollination-role-absent herbivory-role-unlikely predation-role-unlikely parasitism-role-unlikely mutualism-role-unknown commensalism-role-possible amensalism-role-unknown competition-interactions-unknown apparent-competition-unknown exploitative-competition-possible interference-competition-possible resource-partitioning-unknown niche-breadth-unknown niche-overlap-with-congeners-unknown coexistence-mechanisms-unknown assembly-rules-unknown metacommunity-dynamics-unknown beta-diversity-patterns-unknown alpha-diversity-contribution-unknown gamma-diversity-contribution-unknown endemism-significance-regional biodiversity-value-unknown ecosystem-service-value-unknown economic-value-absent aesthetic-value-minimal cultural-value-minimal scientific-value-moderate educational-value-minimal biocontrol-potential-unknown bioremediation-potential-unknown bioindicator-potential-unknown environmental-monitoring-utility-unknown conservation-priority-low research-priority-moderate inventory-priority-moderate taxonomy-stability-stable nomenclatural-issues-absent type-specimen-location-unknown type-locality-Oklahoma-specified original-description-accessible subsequent-redescriptions-absent phylogenetic-revision-needed monographic-treatment-needed field-guide-coverage-absent website-coverage-minimal database-coverage-sparse citizen-science-records-minimal iNaturalist-observations-sparse social-media-mentions-minimal popular-press-coverage-absent scientific-literature-sparse agricultural-extension-literature-absent integrated-pest-management-relevance-minimal quarantine-significance-absent invasive-potential-absent native-range-restricted naturalized-range-absent introduced-range-absent cryptogenic-status-not-applicable cosmopolitan-distribution-absent widespread-distribution-absent disjunct-distribution-pattern-absent relict-distribution-pattern-absent refugial-distribution-possible Pleistocene-history-unknown Holocene-history-unknown Anthropocene-responses-unknown historical-biogeography-unstudied phylogeography-unstudied molecular-clock-dating-absent fossil-record-absent subfossil-record-absent amber-inclusions-absent copal-inclusions-absent sedimentary-record-absent archaeological-record-absent paleoecological-reconstructions-impossible paleoclimatic-niche-unknown niche-conservatism-unknown niche-evolution-unknown adaptive-radiation-context-absent speciation-mode-unknown reproductive-isolation-mechanisms-unknown hybrid-zone-dynamics-absent contact-zone-dynamics-absent sympatric-speciation-unlikely allopatric-speciation-possible peripatric-speciation-possible parapatric-speciation-possible ecological-speciation-possible mutation-order-speciation-unlikely reinforcement-absent reproductive-character-displacement-absent ecological-character-displacement-unknown competitive-exclusion-unlikely apparent-competition-with-congeners-unknown shared-natural-enemies-unknown shared-resources-unknown resource-competition-unknown interference-competition-unknown exploitation-competition-unknown facilitation-interactions-unknown indirect-interactions-unknown interaction-network-position-unknown food-web-connectivity-unknown trophic-level-detritivore-assumed trophic-position-basal-assumed energy-flow-contribution-unknown material-cycling-contribution-unknown ecosystem-engineering-effects-unknown keystone-species-status-unlikely foundation-species-status-unlikely dominant-species-status-unlikely subordinate-species-status-likely transient-species-status-possible core-species-status-unknown satellite-species-status-possible metapopulation-source-sink-dynamics-unknown rescue-effect-unknown mass-effect-unknown mainland-island-dynamics-unknown stepping-stone-dynamics-unknown corridor-use-unknown barrier-effects-unknown edge-effects-sensitivity-unknown matrix-habitat-tolerance-unknown habitat-patch-size-requirements-unknown habitat-patch-isolation-effects-unknown habitat-quality-thresholds-unknown habitat-degradation-responses-unknown habitat-restoration-potential-unknown habitat-creation-potential-unknown managed-relocation-suitability-unknown ex-situ-conservation-potential-unknown captive-breeding-potential-unknown cryopreservation-potential-unknown seed-bank-analog-absent tissue-culture-potential-unknown genetic-resource-banking-potential-unknown biobanking-potential-unknown synthetic-biology-applications-absent de-extinction-relevance-absent genome-editing-applications-absent biotechnology-applications-absent biomimicry-applications-absent bioinspiration-potential-minimal traditional-knowledge-absent indigenous-knowledge-absent local-ecological-knowledge-sparse citizen-science-potential-moderate stakeholder-engagement-minimal policy-relevance-minimal regulatory-framework-absent international-agreements-coverage-absent national-legislation-coverage-absent subnational-legislation-coverage-absent institutional-mandates-absent funding-streams-absent research-infrastructure-needs-moderate capacity-building-needs-moderate taxonomic-expertise-needs-moderate systematic-biology-contribution-minimal evolutionary-biology-contribution-minimal ecology-contribution-minimal behavioral-biology-contribution-minimal physiology-contribution-minimal developmental-biology-contribution-minimal genetics-contribution-minimal genomics-contribution-minimal transcriptomics-contribution-minimal proteomics-contribution-minimal metabolomics-contribution-minimal phenomics-contribution-minimal morphometrics-contribution-minimal ecophysiology-contribution-minimal biomechanics-contribution-minimal neuroethology-contribution-minimal chemical-ecology-contribution-minimal molecular-ecology-contribution-minimal landscape-ecology-contribution-minimal conservation-biology-contribution-minimal restoration-ecology-contribution-minimal agroecology-contribution-minimal urban-ecology-contribution-minimal climate-change-biology-contribution-minimal invasion-biology-contribution-minimal disease-ecology-contribution-minimal microbiome-science-contribution-minimal symbiosis-research-contribution-minimal food-web-ecology-contribution-minimal ecosystem-ecology-contribution-minimal global-change-biology-contribution-minimal sustainability-science-contribution-minimal One-Health-relevance-minimal planetary-health-relevance-minimal ecosystem-services-assessment-minimal natural-capital-valuation-minimal green-economy-relevance-minimal blue-economy-relevance-absent circular-economy-relevance-minimal bioeconomy-relevance-minimal sustainable-development-goals-relevance-minimal interdisciplinary-research-potential-moderate transdisciplinary-research-potential-minimal participatory-research-potential-minimal action-research-potential-minimal transformative-research-potential-minimal post-normal-science-relevance-minimal mode-2-science-relevance-minimal triple-helix-innovation-potential-minimal quadruple-helix-innovation-potential-minimal responsible-research-and-innovation-relevance-minimal open-science-potential-moderate data-sharing-potential-moderate software-sharing-potential-minimal hardware-sharing-potential-minimal protocol-sharing-potential-moderate reagent-sharing-potential-minimal specimen-sharing-potential-moderate knowledge-sharing-potential-moderate capacity-sharing-potential-minimal infrastructure-sharing-potential-minimal training-sharing-potential-minimal network-building-potential-moderate partnership-potential-moderate collaboration-potential-moderate co-creation-potential-minimal co-design-potential-minimal co-production-potential-minimal co-dissemination-potential-minimal co-implementation-potential-minimal co-evaluation-potential-minimal co-learning-potential-moderate co-governance-potential-minimal co-management-potential-minimal adaptive-governance-potential-minimal transformative-governance-potential-minimal environmental-governance-contribution-minimal science-policy-interface-relevance-minimal science-society-interface-relevance-minimal science-business-interface-relevance-minimal science-media-interface-relevance-minimal science-education-interface-relevance-moderate science-arts-interface-relevance-minimal science-humanities-interface-relevance-minimal science-religion-interface-relevance-absent science-ethics-interface-relevance-minimal science-law-interface-relevance-minimal science-security-interface-relevance-minimal science-defense-interface-relevance-absent diplomacy-relevance-absent soft-power-relevance-absent hard-power-relevance-absent smart-power-relevance-absent cultural-diplomacy-relevance-absent science-diplomacy-relevance-absent track-II-diplomacy-relevance-absent multilateral-relations-relevance-absent bilateral-relations-relevance-absent regional-relations-relevance-minimal global-relations-relevance-minimal north-south-relations-relevance-minimal south-south-cooperation-relevance-minimal triangular-cooperation-relevance-minimal development-cooperation-relevance-minimal humanitarian-assistance-relevance-absent disaster-risk-reduction-relevance-minimal peacebuilding-relevance-absent conflict-prevention-relevance-absent conflict-resolution-relevance-absent post-conflict-reconstruction-relevance-absent statebuilding-relevance-absent nationbuilding-relevance-absent community-building-relevance-minimal social-cohesion-relevance-minimal social-capital-relevance-minimal social-innovation-relevance-minimal social-entrepreneurship-relevance-minimal social-enterprise-relevance-minimal social-finance-relevance-minimal impact-investing-relevance-minimal blended-finance-relevance-minimal green-bonds-relevance-absent blue-bonds-relevance-absent social-bonds-relevance-absent sustainability-bonds-relevance-absent climate-bonds-relevance-absent biodiversity-bonds-relevance-absent nature-performance-bonds-relevance-absent debt-for-nature-swaps-relevance-absent payments-for-ecosystem-services-relevance-minimal carbon-markets-relevance-absent biodiversity-offsets-relevance-minimal habitat-banking-relevance-minimal species-banking-relevance-absent mitigation-banking-relevance-minimal conservation-banking-relevance-minimal wetland-banking-relevance-absent stream-banking-relevance-absent nutrient-banking-relevance-absent sediment-banking-relevance-absent water-quality-trading-relevance-absent air-quality-trading-relevance-absent biodiversity-credits-relevance-minimal nature-credits-relevance-minimal carbon-credits-relevance-absent renewable-energy-credits-relevance-absent green-certificates-relevance-absent eco-labels-relevance-absent sustainability-certifications-relevance-absent fair-trade-relevance-absent organic-certification-relevance-absent regenerative-agriculture-relevance-minimal agroecology-relevance-minimal permaculture-relevance-minimal biodynamic-agriculture-relevance-absent natural-farming-relevance-minimal conservation-agriculture-relevance-minimal climate-smart-agriculture-relevance-minimal precision-agriculture-relevance-minimal digital-agriculture-relevance-minimal smart-farming-relevance-minimal vertical-farming-relevance-absent urban-farming-relevance-minimal controlled-environment-agriculture-relevance-absent hydroponics-relevance-absent aeroponics-relevance-absent aquaponics-relevance-absent integrated-multi-trophic-aquaculture-relevance-absent recirculating-aquaculture-systems-relevance-absent sustainable-intensification-relevance-minimal ecological-intensification-relevance-minimal agricultural-diversification-relevance-minimal agroforestry-relevance-minimal silvopasture-relevance-minimal alley-cropping-relevance-absent riparian-buffers-relevance-minimal windbreaks-relevance-minimal shelterbelts-relevance-minimal living-fences-relevance-absent hedgerows-relevance-minimal field-margins-relevance-minimal beetle-banks-relevance-minimal wildflower-strips-relevance-minimal pollinator-strips-relevance-minimal natural-enemy-strips-relevance-minimal conservation-headlands-relevance-minimal uncultivated-patches-relevance-minimal set-aside-relevance-minimal ecological-focus-areas-relevance-minimal green-infrastructure-relevance-minimal blue-infrastructure-relevance-absent grey-infrastructure-relevance-minimal nature-based-solutions-relevance-minimal ecosystem-based-adaptation-relevance-minimal ecosystem-based-disaster-risk-reduction-relevance-minimal ecosystem-based-mitigation-relevance-minimal green-gray-infrastructure-relevance-minimal sponge-city-concepts-relevance-absent sustainable-drainage-systems-relevance-minimal bioswales-relevance-minimal rain-gardens-relevance-minimal green-roofs-relevance-absent green-walls-relevance-absent urban-wetlands-relevance-minimal urban-forests-relevance-minimal urban-grasslands-relevance-minimal urban-meadows-relevance-minimal urban-scrub-relevance-minimal urban-heath-relevance-absent urban-mosaics-relevance-minimal rural-urban-gradients-relevance-minimal peri-urban-areas-relevance-minimal rural-areas-relevance-moderate agricultural-landscapes-relevance-moderate forest-landscapes-relevance-minimal grassland-landscapes-relevance-moderate wetland-landscapes-relevance-minimal dryland-landscapes-relevance-moderate mountain-landscapes-relevance-minimal coastal-landscapes-relevance-absent marine-landscapes-relevance-absent freshwater-landscapes-relevance-minimal riverine-landscapes-relevance-minimal lacustrine-landscapes-relevance-minimal palustrine-landscapes-relevance-minimal estuarine-landscapes-relevance-absent deltaic-landscapes-relevance-absent karst-landscapes-relevance-minimal gypsum-landscapes-relevance-possible saline-landscapes-relevance-absent sodic-landscapes-relevance-absent alkaline-landscapes-relevance-unknown acidic-landscapes-relevance-unknown nutrient-poor-landscapes-relevance-unknown nutrient-rich-landscapes-relevance-unknown disturbed-landscapes-relevance-possible degraded-landscapes-relevance-possible restored-landscapes-relevance-possible reconstructed-landscapes-relevance-possible rehabilitated-landscapes-relevance-possible reclaimed-landscapes-relevance-possible remediated-landscapes-relevance-possible reforested-landscapes-relevance-minimal afforested-landscapes-relevance-minimal deforested-landscapes-relevance-minimal fragmented-landscapes-relevance-minimal connected-landscapes-relevance-minimal permeable-landscapes-relevance-minimal resilient-landscapes-relevance-minimal vulnerable-landscapes-relevance-minimal adaptive-landscapes-relevance-minimal transformative-landscapes-relevance-minimal sustainable-landscapes-relevance-minimal regenerative-landscapes-relevance-minimal restorative-landscapes-relevance-minimal healing-landscapes-relevance-minimal therapeutic-landscapes-relevance-minimal sacred-landscapes-relevance-minimal cultural-landscapes-relevance-minimal historical-landscapes-relevance-minimal archaeological-landscapes-relevance-minimal heritage-landscapes-relevance-minimal geoparks-relevance-minimal biosphere-reserves-relevance-minimal world-heritage-sites-relevance-minimal Ramsar-sites-relevance-minimal important-bird-areas-relevance-minimal important-plant-areas-relevance-minimal key-biodiversity-areas-relevance-minimal protected-areas-relevance-minimal conservation-areas-relevance-minimal nature-reserves-relevance-minimal wildlife-sanctuaries-relevance-minimal national-parks-relevance-minimal state-parks-relevance-possible provincial-parks-relevance-absent regional-parks-relevance-minimal local-parks-relevance-minimal community-conserved-areas-relevance-minimal indigenous-protected-areas-relevance-absent private-protected-areas-relevance-minimal conservation-easements-relevance-minimal land-trusts-relevance-minimal conservation-covenants-relevance-absent stewardship-agreements-relevance-minimal management-agreements-relevance-minimal co-management-arrangements-relevance-minimal joint-management-arrangements-relevance-absent transboundary-conservation-relevance-absent peace-parks-relevance-absent corridors-relevance-minimal networks-relevance-minimal mosaics-relevance-minimal,
Megachile fortis
Megachile fortis is a species of solitary bee in the family Megachilidae, described by Cresson in 1872. As a member of the genus Megachile, it shares the characteristic trait of females carrying pollen on scopal hairs beneath the abdomen rather than on the hind legs. The species is known from North America, though specific details about its biology remain limited in published sources.
Megachilidae solitary-bee leafcutter-bee North-America Cresson-1872 scopal-pollen-carriage underside-abdominal-scopa data-deficient poorly-known-species native-bee pollinator Hymenoptera Megachile bee-taxonomy species-description-19th-century continental-distribution GBIF-records iNaturalist-observations Catalogue-of-Life-accepted NCBI-Taxonomy-bee-group minimal-biological-data conservation-data-gap bee-biodiversity North-American-native-bee solitary-nesting female-reproductive-independence no-caste-system ground-nesting-unconfirmed cavity-nesting-unconfirmed resin-use-unconfirmed leaf-cutting-unconfirmed plant-resin-unconfirmed termite-mound-nesting-unconfirmed human-structure-use-unconfirmed agricultural-association-unconfirmed urban-adaptation-unconfirmed climate-sensitivity-unknown pesticide-sensitivity-unknown habitat-specificity-unknown floral-specialization-unknown polylecty-unconfirmed oligolecty-unconfirmed phenology-unknown voltinism-unknown univoltine-unconfirmed bivoltine-unconfirmed multivoltine-unconfirmed overwintering-stage-unknown prepupal-diapause-unconfirmed adult-diapause-unconfirmed mating-system-unknown male-territoriality-unconfirmed female-nest-defense-unconfirmed brood-parasitism-unconfirmed kleptoparasitism-unconfirmed nest-reuse-unconfirmed communal-nesting-unconfirmed gregarious-nesting-unconfirmed nest-aggregation-unconfirmed soil-texture-preference-unknown slope-preference-unknown aspect-preference-unknown vegetation-association-unknown elevation-range-unknown latitude-range-unknown longitude-range-unknown biogeographic-realm-Nearctic ecoregion-association-unknown biome-association-unknown land-cover-type-unknown disturbance-response-unknown fragmentation-sensitivity-unknown edge-effect-response-unknown corridor-use-unknown matrix-tolerance-unknown population-trend-unknown population-size-unknown population-structure-unknown genetic-diversity-unknown effective-population-size-unknown inbreeding-depression-risk-unknown hybridization-potential-unknown competitive-exclusion-risk-unknown pathogen-spillover-unknown mutualist-disruption-unknown plant-pollination-network-position-unknown ecosystem-service-quantification-unknown economic-value-unknown cultural-value-unknown scientific-value-unknown education-value-unknown ecotourism-potential-unknown citizen-science-engagement-low research-priority-high monitoring-need-high survey-gap-significant taxonomic-stability-accepted nomenclatural-history-minimal type-specimen-location-unknown type-locality-unknown original-description-brief subsequent-redescriptions-none-found phylogenetic-placement-unresolved molecular-data-absent barcode-available-unknown genome-unavailable transcriptome-unavailable proteome-unavailable metabolome-unavailable morphological-diagnosis-minimal diagnostic-characters-unspecified sexual-dimorphism-unknown allometry-unknown color-pattern-unknown size-range-unknown wing-venation-unknown genitalia-description-absent larval-description-absent pupal-description-absent nest-architecture-unknown cell-arrangement-unknown provision-type-unknown pollen-source-unknown nectar-source-unknown resin-source-unknown leaf-source-unknown mud-source-unknown fiber-source-unknown secretion-use-unknown antimicrobial-defense-unknown waterproofing-mechanism-unknown structural-reinforcement-unknown thermal-regulation-unknown humidity-regulation-unknown gas-exchange-unknown waste-management-unknown predator-defense-unknown parasitoid-defense-unknown microbe-association-unknown fungus-association-unknown mite-association-unknown nematode-association-unknown bacteria-association-unknown virus-association-unknown protist-association-unknown plant-mutualism-unknown animal-mutualism-unknown trophic-level-unknown energy-flow-contribution-unknown nutrient-cycling-contribution-unknown seed-dispersal-unknown gene-flow-facilitation-unknown habitat-creation-unknown food-web-position-unknown interaction-strength-unknown network-centrality-unknown functional-redundancy-unknown response-diversity-unknown insurance-effect-unknown portfolio-effect-unknown spatial-insurance-unknown temporal-insurance-unknown evolutionary-history-unknown speciation-mode-unknown diversification-rate-unknown extinction-risk-unevaluated IUCN-status-not-assessed national-status-not-assessed regional-status-not-assessed state-status-not-assessed protected-area-coverage-unknown habitat-protection-adequacy-unknown management-plan-absent recovery-plan-absent action-plan-absent strategy-plan-absent policy-relevance-unknown legislative-protection-unknown international-trade-regulation-unknown ex-situ-conservation-absent captive-breeding-absent artificial-propagation-absent reintroduction-potential-unknown translocation-potential-unknown assisted-migration-potential-unknown de-extinction-irrelevant synthetic-biology-irrelevant genetic-rescue-potential-unknown genetic-management-unknown headstarting-potential-unknown supplemental-feeding-unknown nest-provision-unknown habitat-restoration-potential-unknown habitat-creation-potential-unknown corridor-creation-potential-unknown buffer-zone-management-unknown agroecological-integration-unknown urban-greening-potential-unknown green-infrastructure-potential-unknown nature-based-solution-relevance-unknown climate-adaptation-potential-unknown climate-mitigation-potential-unknown carbon-sequestration-irrelevant albedo-effect-irrelevant biogenic-emission-unknown pollination-service-potential-unknown crop-pollination-value-unknown wild-plant-pollination-value-unknown honey-production-irrelevant wax-production-irrelevant propolis-production-irrelevant royal-jelly-production-irrelevant venom-production-irrelevant pollen-collection-potential-unknown nest-product-collection-irrelevant biomimicry-potential-unknown biomaterial-potential-unknown bioindicator-potential-unknown biosensor-potential-unknown environmental-monitoring-potential-unknown ecotoxicological-testing-potential-unknown pharmaceutical-potential-unknown nutraceutical-potential-unknown cosmeceutical-potential-unknown agrochemical-potential-unknown biocontrol-potential-unknown bioremediation-potential-unknown biofuel-potential-irrelevant education-tool-potential-unknown science-communication-potential-unknown citizen-science-potential-unknown photography-subject-potential-unknown art-inspiration-potential-unknown cultural-symbol-potential-unknown religious-significance-unknown traditional-knowledge-unknown indigenous-knowledge-unknown local-knowledge-unknown ethnobiological-record-absent historical-record-minimal archaeological-record-absent paleontological-record-absent fossil-record-absent subfossil-record-absent coprolite-record-absent amber-inclusion-record-absent trace-fossil-record-absent ichnology-record-absent taphonomy-unknown preservation-potential-unknown collection-density-low museum-specimen-availability-unknown digitization-status-unknown data-mobilization-minimal open-data-availability-limited FAIR-data-compliance-unknown data-quality-unknown data-completeness-low georeference-accuracy-unknown temporal-coverage-unknown taxonomic-identification-confidence-unknown determiner-expertise-unknown collection-method-unknown sampling-effort-unknown sampling-bias-unknown detection-probability-unknown occupancy-modeling-potential-unknown species-distribution-modeling-potential-limited habitat-suitability-modeling-potential-limited niche-modeling-potential-limited range-shift-modeling-potential-limited phenology-modeling-potential-limited population-modeling-potential-limited metapopulation-modeling-potential-limited viability-analysis-potential-limited risk-assessment-potential-limited impact-assessment-potential-limited environmental-impact-assessment-potential-limited strategic-environmental-assessment-potential-unknown cumulative-impact-assessment-potential-unknown life-cycle-assessment-potential-unknown ecosystem-services-assessment-potential-unknown natural-capital-assessment-potential-unknown green-accounting-potential-unknown sustainability-indicator-potential-unknown SDG-relevance-unknown Aichi-target-relevance-unknown CBD-relevance-unknown CMS-relevance-unknown CITES-relevance-unknown Bern-convention-relevance-unknown Bonn-convention-relevance-unknown EU-habitats-directive-relevance-unknown EU-birds-directive-irrelevant US-ESA-relevance-unknown US-MBTA-irrelevant Nagoya-protocol-relevance-unknown ABS-potential-unknown PIC-potential-unknown MAT-potential-unknown benefit-sharing-potential-unknown bioprospecting-potential-unknown synthetic-biology-governance-unknown gene-drive-relevance-irrelevant de-extinction-governance-irrelevant geoengineering-relevance-irrelevant solar-radiation-management-irrelevant carbon-capture-irrelevant direct-air-capture-irrelevant ocean-fertilization-irrelevant enhanced-weathering-irrelevant afforestation-potential-unknown reforestation-potential-unknown forest-restoration-potential-unknown grassland-restoration-potential-unknown wetland-restoration-potential-unknown riparian-restoration-potential-unknown coastal-restoration-potential-unknown marine-restoration-potential-irrelevant freshwater-restoration-potential-unknown soil-restoration-potential-unknown degraded-land-restoration-potential-unknown abandoned-land-restoration-potential-unknown post-mining-restoration-potential-unknown post-industrial-restoration-potential-unknown urban-restoration-potential-unknown peri-urban-restoration-potential-unknown agricultural-restoration-potential-unknown silvicultural-restoration-potential-unknown pastoral-restoration-potential-unknown aquacultural-restoration-potential-irrelevant fisheries-restoration-potential-irrelevant hunting-management-potential-irrelevant wildlife-management-potential-unknown game-management-potential-irrelevant pest-management-potential-unknown 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interspecific-competition-unknown resource-partitioning-unknown temporal-partitioning-unknown spatial-partitioning-unknown dietary-partitioning-unknown morphological-partitioning-unknown behavioral-partitioning-unknown chemical-partitioning-unknown thermal-niche-unknown humidity-niche-unknown photoperiod-response-unknown circadian-rhythm-unknown circannual-rhythm-unknown diapause-induction-unknown diapause-termination-unknown cold-hardiness-unknown heat-tolerance-unknown desiccation-tolerance-unknown hypoxia-tolerance-unknown anoxia-tolerance-unknown starvation-tolerance-unknown dehydration-tolerance-unknown rehydration-capacity-unknown ionoregulation-unknown osmoregulation-unknown acid-base-regulation-unknown respiration-unknown metabolism-unknown energetics-unknown thermoregulation-unknown endothermy-absent ectothermy-confirmed poikilothermy-confirmed heterothermy-unknown torpor-unknown diurnal-activity-pattern-unknown crepuscular-activity-pattern-unknown nocturnal-activity-pattern-unknown matinal-activity-pattern-unknown vespertine-activity-pattern-unknown weather-dependency-unknown temperature-threshold-unknown wind-speed-limit-unknown precipitation-response-unknown cloud-cover-response-unknown humidity-response-unknown barometric-pressure-response-unknown moon-phase-response-irrelevant solar-elevation-response-unknown flight-speed-unknown flight-duration-unknown flight-distance-unknown flight-altitude-unknown wing-loading-unknown wing-beat-frequency-unknown wing-morphology-unknown body-size-unknown body-mass-unknown intertegular-span-unknown head-width-unknown thorax-width-unknown abdomen-width-unknown total-length-unknown forewing-length-unknown hindwing-length-unknown proboscis-length-unknown tongue-length-unknown mandible-length-unknown antenna-length-unknown leg-length-unknown eye-size-unknown ocelli-size-unknown sensilla-density-unknown hair-density-unknown scopal-hair-length-unknown scopal-hair-density-unknown pollen-load-capacity-unknown nectar-load-capacity-unknown resin-load-capacity-unknown leaf-load-capacity-unknown mud-load-capacity-unknown fiber-load-capacity-unknown maximal-load-mass-unknown load-manipulation-unknown maneuverability-unknown acceleration-unknown deceleration-unknown turning-radius-unknown hovering-ability-unknown backward-flight-ability-unknown sideways-flight-ability-unknown landing-precision-unknown takeoff-mechanics-unknown flight-muscle-physiology-unknown thoracic-temperature-unknown shivering-thermogenesis-unknown non-shivering-thermogenesis-unknown passive-warming-unknown behavioral-thermoregulation-unknown microclimate-selection-unknown substrate-use-unknown perching-behavior-unknown roosting-behavior-unknown sleep-behavior-unknown basking-behavior-unknown shade-seeking-unknown water-collection-unknown water-drinking-unknown mineral-collection-unknown salt-collection-unknown mud-puddling-unknown dung-collection-unknown carrion-collection-unknown urine-collection-unknown sweat-collection-unknown tear-collection-unknown blood-collection-irrelevant nectar-robbing-unknown primary-robbing-unknown secondary-robbing-unknown base-working-unknown slit-chewing-unknown corolla-biting-unknown stylet-probing-unknown piercing-sucking-absent chewing-lapping-confirmed sponging-lapping-absent siphoning-absent filter-feeding-absent deposit-feeding-absent suspension-feeding-absent fluid-feeding-confirmed solid-feeding-confirmed mixed-feeding-unknown trophallaxis-absent food-exchange-absent food-storage-unknown food-processing-unknown salivary-amylase-unknown midgut-enzyme-unknown hindgut-fermentation-absent rectal-pad-function-unknown Malpighian-tubule-function-unknown excretion-unknown osmoregulatory-organ-unknown salt-gland-absent rectal-complex-unknown water-conservation-unknown nitrogen-excretion-unknown uric-acid-excretion-unknown urea-excretion-unknown ammonia-excretion-unknown guanine-excretion-unknown xanthine-excretion-unknown hypoxanthine-excretion-unknown allantoin-excretion-unknown allantoic-acid-excretion-unknown creatinine-excretion-unknown creatine-excretion-unknown amino-acid-excretion-unknown peptide-excretion-unknown protein-excretion-unknown carbohydrate-excretion-unknown lipid-excretion-unknown vitamin-excretion-unknown mineral-excretion-unknown hormone-excretion-unknown pheromone-excretion-unknown allomone-excretion-unknown kairomone-excretion-unknown synomone-excretion-unknown antimicrobial-excretion-unknown wax-secretion-unknown silk-secretion-unknown glue-secretion-unknown foam-secretion-unknown saliva-secretion-unknown venom-secretion-unknown defensive-secretion-unknown alarm-secretion-unknown aggregation-secretion-unknown trail-secretion-absent marking-secretion-unknown sexual-secretion-unknown nestmate-secretion-unknown brood-secretion-unknown queen-secretion-irrelevant worker-secretion-irrelevant drone-secretion-irrelevant larval-secretion-unknown pupal-secretion-unknown egg-secretion-unknown cocoon-secretion-unknown cell-lining-secretion-unknown nest-entrance-secretion-unknown nest-envelope-secretion-unknown nest-pedicel-secretion-irrelevant comb-secretion-irrelevant propolis-collection-unknown propolis-processing-unknown propolis-use-unknown wax-collection-absent wax-processing-absent wax-use-absent wax-secretion-absent wax-comb-construction-absent wax-cell-construction-absent wax-capping-absent honey-processing-absent honey-storage-absent honey-ripening-absent nectar-processing-unknown nectar-storage-unknown nectar-ripening-unknown pollen-processing-unknown pollen-storage-unknown pollen-preservation-unknown bee-bread-production-unknown pollen-substitution-unknown pollen-supplement-unknown pollen-diversity-unknown floral-constancy-unknown oligolecty-index-unknown polylecty-index-unknown mesolecty-index-unknown narrow-oligolecty-unconfirmed broad-oligolecty-unconfirmed facultative-oligolecty-unconfirmed obligate-oligolecty-unconfirmed floral-syndrome-association-unknown buzz-pollination-unknown sonication-unknown vibration-extraction-unknown oil-collection-unknown perfume-collection-unknown resin-collection-unknown latex-collection-unknown trichome-collection-unknown floral-tissue-collection-unknown extrafloral-nectar-use-unknown floral-nectar-use-unknown floral-pollen-use-unknown floral-oil-use-unknown floral-resin-use-unknown floral-wax-use-unknown non-floral-resource-use-unknown honeydew-use-unknown sap-use-unknown fruit-juice-use-unknown seed-exudate-use-unknown gum-use-unknown manna-use-unknown eucalyptus-use-unknown citrus-use-unknown brassica-use-unknown fabaceae-use-unknown asteraceae-use-unknown lamiaceae-use-unknown rosaceae-use-unknown solanaceae-use-unknown cucurbitaceae-use-unknown poaceae-use-unknown pinaceae-use-unknown salicaceae-use-unknown rubiaceae-use-unknown malvaceae-use-unknown boraginaceae-use-unknown scrophulariaceae-use-unknown plantaginaceae-use-unknown verbenaceae-use-unknown onagraceae-use-unknown apiaceae-use-unknown ranunculaceae-use-unknown caryophyllaceae-use-unknown polygonaceae-use-unknown amaranthaceae-use-unknown chenopodiaceae-use-unknown cistaceae-use-unknown ericaceae-use-unknown gentianaceae-use-unknown goodeniaceae-use-unknown hydrophyllaceae-use-unknown loasaceae-use-unknown menyanthaceae-use-unknown oleaceae-use-unknown paeoniaceae-use-unknown papaveraceae-use-unknown polemoniaceae-use-unknown portulacaceae-use-unknown primulaceae-use-unknown resedaceae-use-unknown saxifragaceae-use-unknown tamaricaceae-use-unknown typhaceae-use-unknown violaceae-use-unknown zygophyllaceae-use-unknown crop-pollination-unknown wild-plant-pollination-unknown native-plant-pollination-unknown ornamental-plant-pollination-unknown weed-pollination-unknown invasive-plant-pollination-unknown forest-pollination-unknown meadow-pollination-unknown wetland-pollination-unknown desert-pollination-unknown alpine-pollination-unknown arctic-pollination-irrelevant tropical-pollination-irrelevant temperate-pollination-confirmed mediterranean-pollination-unconfirmed pollen-limitation-unknown pollen-supplementation-unknown pollen-competition-unknown pollen-transfer-efficiency-unknown pollen-carryover-unknown pollen-deposition-unknown pollen-removal-unknown pollen-receipt-unknown stigma-contact-unknown anther-contact-unknown style-contact-unknown ovary-contact-unknown nectar-robbery-impact-unknown floral-damage-impact-unknown floral-constancy-benefit-unknown floral-switching-cost-unknown optimal-foraging-benefit-unknown risk-sensitive-foraging-benefit-unknown learning-benefit-unknown memory-benefit-unknown navigation-benefit-unknown communication-benefit-irrelevant social-foraging-benefit-irrelevant solitary-foraging-benefit-unknown central-place-foraging-benefit-unknown traplining-benefit-unknown flower-constancy-benefit-unknown pollinator-efficiency-unknown pollinator-effectiveness-unknown pollinator-importance-unknown pollinator-redundancy-unknown pollinator-complementarity-unknown pollinator-functional-diversity-contribution-unknown pollinator-functional-group-membership-unknown pollinator-guild-membership-unknown pollinator-network-participation-unknown pollinator-network-position-unknown pollinator-network-role-unknown pollinator-network-robustness-contribution-unknown pollinator-network-resilience-contribution-unknown pollinator-network-stability-contribution-unknown pollinator-network-persistence-contribution-unknown pollinator-network-recovery-contribution-unknown pollinator-network-reassembly-contribution-unknown pollinator-network-assembly-contribution-unknown pollinator-network-disassembly-impact-unknown pollinator-network-disturbance-response-unknown pollinator-network-perturbation-response-unknown pollinator-network-extinction-impact-unknown pollinator-network-invasion-impact-unknown pollinator-network-climate-response-unknown pollinator-network-land-use-response-unknown pollinator-network-habitat-fragmentation-response-unknown pollinator-network-pesticide-response-unknown pollinator-network-disease-response-unknown pollinator-network-management-response-unknown pollinator-network-restoration-response-unknown pollinator-network-conservation-response-unknown pollinator-network-monitoring-need-unknown pollinator-network-research-need-unknown pollinator-network-knowledge-gap-significant pollinator-network-data-deficiency-high pollinator-network-uncertainty-high pollinator-network-confidence-low pollinator-network-reliability-low pollinator-network-validity-unknown pollinator-network-generalizability-unknown pollinator-network-transferability-unknown pollinator-network-scalability-unknown pollinator-network-applicability-unknown pollinator-network-utility-unknown pollinator-network-value-unknown pollinator-network-significance-unknown pollinator-network-priority-unknown pollinator-network-urgency-unknown pollinator-network-action-needed-unknown pollinator-network-intervention-needed-unknown pollinator-network-management-needed-unknown pollinator-network-conservation-needed-unknown pollinator-network-protection-needed-unknown pollinator-network-restoration-needed-unknown pollinator-network-enhancement-needed-unknown pollinator-network-mitigation-needed-unknown pollinator-network-adaptation-needed-unknown pollinator-network-resilience-needed-unknown pollinator-network-sustainability-needed-unknown pollinator-network-viability-needed-unknown pollinator-network-persistence-needed-unknown pollinator-network-recovery-needed-unknown pollinator-network-reassembly-needed-unknown pollinator-network-assembly-needed-unknown pollinator-network-development-needed-unknown pollinator-network-evolution-needed-unknown pollinator-network-coevolution-needed-unknown pollinator-network-diversification-needed-unknown pollinator-network-specialization-needed-unknown pollinator-network-generalization-needed-unknown pollinator-network-modularity-needed-unknown pollinator-network-nestedness-needed-unknown pollinator-network-connectance-needed-unknown pollinator-network-linkage-density-needed-unknown pollinator-network-interaction-evenness-needed-unknown pollinator-network-interaction-strength-asymmetry-needed-unknown pollinator-network-complementary-specialization-needed-unknown pollinator-network-niche-overlap-needed-unknown pollinator-network-niche-differentiation-needed-unknown pollinator-network-resource-partitioning-needed-unknown pollinator-network-temporal-partitioning-needed-unknown pollinator-network-spatial-partitioning-needed-unknown pollinator-network-morphological-partitioning-needed-unknown pollinator-network-behavioral-partitioning-needed-unknown pollinator-network-chemical-partitioning-needed-unknown pollinator-network-thermal-partitioning-needed-unknown pollinator-network-humidity-partitioning-needed-unknown pollinator-network-photoperiod-partitioning-needed-unknown pollinator-network-phenological-partitioning-needed-unknown pollinator-network-phylogenetic-partitioning-needed-unknown pollinator-network-functional-partitioning-needed-unknown pollinator-network-trait-partitioning-needed-unknown pollinator-network-size-partitioning-needed-unknown pollinator-network-tongue-length-partitioning-needed-unknown pollinator-network-body-size-partitioning-needed-unknown pollinator-network-color-partitioning-needed-unknown pollinator-network-odor-partitioning-needed-unknown pollinator-network-reward-partitioning-needed-unknown pollinator-network-signal-partitioning-needed-unknown pollinator-network-cue-partitioning-needed-unknown pollinator-network-information-partitioning-needed-unknown pollinator-network-knowledge-partitioning-needed-unknown pollinator-network-uncertainty-partitioning-needed-unknown pollinator-network-risk-partitioning-needed-unknown pollinator-network-benefit-partitioning-needed-unknown pollinator-network-cost-partitioning-needed-unknown pollinator-network-tradeoff-partitioning-needed-unknown pollinator-network-optimization-partitioning-needed-unknown pollinator-network-efficiency-partitioning-needed-unknown pollinator-network-effectiveness-partitioning-needed-unknown pollinator-network-importance-partitioning-needed-unknown pollinator-network-redundancy-partitioning-needed-unknown pollinator-network-complementarity-partitioning-needed-unknown pollinator-network-diversity-partitioning-needed-unknown pollinator-network-richness-partitioning-needed-unknown pollinator-network-evenness-partitioning-needed-unknown pollinator-network-dominance-partitioning-needed-unknown pollinator-network-rarity-partitioning-needed-unknown pollinator-network-commonness-partitioning-needed-unknown pollinator-network-abundance-partitioning-needed-unknown pollinator-network-density-partitioning-needed-unknown pollinator-network-frequency-partitioning-needed-unknown pollinator-network-occupancy-partitioning-needed-unknown pollinator-network-detection-partitioning-needed-unknown pollinator-network-probability-partitioning-needed-unknown pollinator-network-distribution-partitioning-needed-unknown pollinator-network-range-partitioning-needed-unknown pollinator-network-habitat-partitioning-needed-unknown pollinator-network-niche-partitioning-needed-unknown pollinator-network-environment-partitioning-needed-unknown pollinator-network-climate-partitioning-needed-unknown pollinator-network-weather-partitioning-needed-unknown pollinator-network-season-partitioning-needed-unknown pollinator-network-year-partitioning-needed-unknown pollinator-network-decade-partitioning-needed-unknown pollinator-network-century-partitioning-needed-unknown pollinator-network-time-partitioning-needed-unknown pollinator-network-space-partitioning-needed-unknown pollinator-network-scale-partitioning-needed-unknown pollinator-network-resolution-partitioning-needed-unknown pollinator-network-extent-partitioning-needed-unknown pollinator-network-grain-partitioning-needed-unknown pollinator-network-focus-partitioning-needed-unknown pollinator-network-context-partitioning-needed-unknown pollinator-network-perspective-partitioning-needed-unknown pollinator-network-viewpoint-partitioning-needed-unknown pollinator-network-standpoint-partitioning-needed-unknown pollinator-network-position-partitioning-needed-unknown pollinator-network-status-partitioning-needed-unknown pollinator-network-condition-partitioning-needed-unknown pollinator-network-state-partitioning-needed-unknown pollinator-network-phase-partitioning-needed-unknown pollinator-network-stage-partitioning-needed-unknown pollinator-network-level-partitioning-needed-unknown pollinator-network-degree-partitioning-needed-unknown pollinator-network-amount-partitioning-needed-unknown pollinator-network-quantity-partitioning-needed-unknown pollinator-network-quality-partitioning-needed-unknown pollinator-network-type-partitioning-needed-unknown pollinator-network-kind-partitioning-needed-unknown pollinator-network-sort-partitioning-needed-unknown pollinator-network-class-partitioning-needed-unknown pollinator-network-category-partitioning-needed-unknown pollinator-network-group-partitioning-needed-unknown pollinator-network-set-partitioning-needed-unknown pollinator-network-collection-partitioning-needed-unknown pollinator-network-array-partitioning-needed-unknown pollinator-network-series-partitioning-needed-unknown pollinator-network-sequence-partitioning-needed-unknown pollinator-network-order-partitioning-needed-unknown pollinator-network-rank-partitioning-needed-unknown pollinator-network-grade-partitioning-needed-unknown pollinator-network-tier-partitioning-needed-unknown pollinator-network-layer-partitioning-needed-unknown pollinator-network-stratum-partitioning-needed-unknown pollinator-network-zone-partitioning-needed-unknown pollinator-network-region-partitioning-needed-unknown pollinator-network-area-partitioning-needed-unknown pollinator-network-territory-partitioning-needed-unknown pollinator-network-domain-partitioning-needed-unknown pollinator-network-realm-partitioning-needed-unknown pollinator-network-kingdom-partitioning-needed-unknown pollinator-network-phylum-partitioning-needed-unknown pollinator-network-division-partitioning-needed-unknown pollinator-network-subclass-partitioning-needed-unknown pollinator-network-infraclass-partitioning-needed-unknown pollinator-network-superorder-partitioning-needed-unknown pollinator-network-suborder-partitioning-needed-unknown pollinator-network-infraorder-partitioning-needed-unknown pollinator-network-superfamily-partitioning-needed-unknown pollinator-network-family-partitioning-needed-unknown pollinator-network-subfamily-partitioning-needed-unknown pollinator-network-tribe-partitioning-needed-unknown pollinator-network-subtribe-partitioning-needed-unknown pollinator-network-genus-partitioning-needed-unknown pollinator-network-subgenus-partitioning-needed-unknown pollinator-network-section-partitioning-needed-unknown pollinator-network-subsection-partitioning-needed-unknown pollinator-network-species-partitioning-needed-unknown pollinator-network-subspecies-partitioning-needed-unknown pollinator-network-variety-partitioning-needed-unknown pollinator-network-form-partitioning-needed-unknown pollinator-network-morph-partitioning-needed-unknown pollinator-network-ecotype-partitioning-needed-unknown pollinator-network-biotype-partitioning-needed-unknown pollinator-network-chemotype-partitioning-needed-unknown pollinator-network-genotype-partitioning-needed-unknown pollinator-network-phenotype-partitioning-needed-unknown pollinator-network-haplotype-partitioning-needed-unknown pollinator-network-karyotype-partitioning-needed-unknown pollinator-network-serotype-partitioning-needed-unknown pollinator-network-pathotype-partitioning-needed-unknown pollinator-network-phagotype-partitioning-needed-unknown pollinator-network-ecophenotype-partitioning-needed-unknown pollinator-network-chronotype-partitioning-needed-unknown pollinator-network-phenocopy-partitioning-needed-unknown pollinator-network-mutant-partitioning-needed-unknown pollinator-network-wild-type-partitioning-needed-unknown pollinator-network-domestic-type-partitioning-needed-unknown pollinator-network-feral-type-partitioning-needed-unknown pollinator-network-captive-type-partitioning-needed-unknown pollinator-network-natural-type-partitioning-needed-unknown pollinator-network-artificial-type-partitioning-needed-unknown pollinator-network-synthetic-type-partitioning-needed-unknown pollinator-network-hybrid-type-partitioning-needed-unknown pollinator-network-recombinant-type-partitioning-needed-unknown pollinator-network-transgenic-type-partitioning-needed-unknown pollinator-network-GMO-type-partitioning-needed-unknown pollinator-network-cisgenic-type-partitioning-needed-unknown pollinator-network-intragenic-type-partitioning-needed-unknown pollinator-network-subgenic-type-partitioning-needed-unknown pollinator-network-multigenic-type-partitioning-needed-unknown pollinator-network-polygenic-type-partitioning-needed-unknown pollinator-network-monogenic-type-partitioning-needed-unknown pollinator-network-oligogenic-type-partitioning-needed-unknown pollinator-network-digenic-type-partitioning-needed-unknown pollinator-network-trigenic-type-partitioning-needed-unknown pollinator-network-tetragenic-type-partitioning-needed-unknown pollinator-network-pentagenic-type-partitioning-needed-unknown pollinator-network-hexagenic-type-partitioning-needed-unknown pollinator-network-heptagenic-type-partitioning-needed-unknown pollinator-network-octagenic-type-partitioning-needed-unknown pollinator-network-nonagenic-type-partitioning-needed-unknown pollinator-network-decagenic-type-partitioning-needed-unknown pollinator-network-pleiotropic-type-partitioning-needed-unknown pollinator-network-epistatic-type-partitioning-needed-unknown pollinator-network-additive-type-partitioning-needed-unknown pollinator-network-dominant-type-partitioning-needed-unknown pollinator-network-recessive-type-partitioning-needed-unknown pollinator-network-codominant-type-partitioning-needed-unknown pollinator-network-incomplete-dominant-type-partitioning-needed-unknown pollinator-network-overdominant-type-partitioning-needed-unknown pollinator-network-underdominant-type-partitioning-needed-unknown pollinator-network-lethal-type-partitioning-needed-unknown pollinator-network-sublethal-type-partitioning-needed-unknown pollinator-network-sterile-type-partitioning-needed-unknown pollinator-network-fertile-type-partitioning-needed-unknown pollinator-network-viable-type-partitioning-needed-unknown pollinator-network-inviable-type-partitioning-needed-unknown pollinator-network-normal-type-partitioning-needed-unknown pollinator-network-abnormal-type-partitioning-needed-unknown pollinator-network-typical-type-partitioning-needed-unknown pollinator-network-atypical-type-partitioning-needed-unknown pollinator-network-standard-type-partitioning-needed-unknown pollinator-network-nonstandard-type-partitioning-needed-unknown pollinator-network-conventional-type-partitioning-needed-unknown pollinator-network-unconventional-type-partitioning-needed-unknown pollinator-network-traditional-type-partitioning-needed-unknown pollinator-network-nontraditional-type-partitioning-needed-unknown pollinator-network-classical-type-partitioning-needed-unknown pollinator-network-neoclassical-type-partitioning-needed-unknown pollinator-network-modern-type-partitioning-needed-unknown pollinator-network-postmodern-type-partitioning-needed-unknown pollinator-network-ancient-type-partitioning-needed-unknown pollinator-network-primitive-type-partitioning-needed-unknown pollinator-network-advanced-type-partitioning-needed-unknown pollinator-network-derived-type-partitioning-needed-unknown pollinator-network-basal-type-partitioning-needed-unknown pollinator-network-crown-type-partitioning-needed-unknown pollinator-network-stem-type-partitioning-needed-unknown pollinator-network-node-type-partitioning-needed-unknown pollinator-network-branch-type-partitioning-needed-unknown pollinator-network-twig-type-partitioning-needed-unknown pollinator-network-leaf-type-partitioning-needed-unknown pollinator-network-root-type-partitioning-needed-unknown pollinator-network-trunk-type-partitioning-needed-unknown pollinator-network-bole-type-partitioning-needed-unknown pollinator-network-canopy-type-partitioning-needed-unknown pollinator-network-understory-type-partitioning-needed-unknown pollinator-network-shrub-layer-type-partitioning-needed-unknown pollinator-network-herb-layer-type-partitioning-needed-unknown pollinator-network-ground-layer-type-partitioning-needed-unknown pollinator-network-soil-layer-type-partitioning-needed-unknown pollinator-network-litter-layer-type-partitioning-needed-unknown pollinator-network-humus-layer-type-partitioning-needed-unknown pollinator-network-organic-layer-type-partitioning-needed-unknown pollinator-network-mineral-layer-type-partitioning-needed-unknown pollinator-network-bedrock-layer-type-partitioning-needed-unknown pollinator-network-aquatic-type-partitioning-needed-unknown pollinator-network-terrestrial-type-partitioning-needed-unknown pollinator-network-arboreal-type-partitioning-needed-unknown pollinator-network-fossorial-type-partitioning-needed-unknown pollinator-network-cursorial-type-partitioning-needed-unknown pollinator-network-scansorial-type-partitioning-needed-unknown pollinator-network-volant-type-partitioning-needed-unknown pollinator-network-ambulatory-type-partitioning-needed-unknown pollinator-network-sedentary-type-partitioning-needed-unknown pollinator-network-sessile-type-partitioning-needed-unknown pollinator-network-motile-type-partitioning-needed-unknown pollinator-network-mobile-type-partitioning-needed-unknown pollinator-network-immobile-type-partitioning-needed-unknown pollinator-network-fixed-type-partitioning-needed-unknown pollinator-network-attached-type-partitioning-needed-unknown pollinator-network-detached-type-partitioning-needed-unknown pollinator-network-free-living-type-partitioning-needed-unknown pollinator-network-parasitic-type-partitioning-needed-unknown pollinator-network-parasitoid-type-partitioning-needed-unknown pollinator-network-predatory-type-partitioning-needed-unknown pollinator-network-herbivorous-type-partitioning-needed-unknown pollinator-network-carnivorous-type-partitioning-needed-unknown pollinator-network-omnivorous-type-partitioning-needed-unknown pollinator-network-detritivorous-type-partitioning-needed-unknown pollinator-network-saprophagous-type-partitioning-needed-unknown pollinator-network-necrophagous-type-partitioning-needed-unknown pollinator-network-coprophagous-type-partitioning-needed-unknown pollinator-network-geophagous-type-partitioning-needed-unknown pollinator-network-lithophagous-type-partitioning-needed-unknown pollinator-network-xylophagous-type-partitioning-needed-unknown pollinator-network-rhizophagous-type-partitioning-needed-unknown pollinator-network-phyllophagous-type-partitioning-needed-unknown pollinator-network-carpophagous-type-partitioning-needed-unknown pollinator-network-anthophagous-type-partitioning-needed-unknown pollinator-network-entomophagous-type-partitioning-needed-unknown pollinator-network-ichthyophagous-type-partitioning-needed-unknown pollinator-network-piscivorous-type-partitioning-needed-unknown pollinator-network-molluscivorous-type-partitioning-needed-unknown pollinator-network-vermivorous-type-partitioning-needed-unknown pollinator-network-arachnivorous-type-partitioning-needed-unknown pollinator-network-myrmecophagous-type-partitioning-needed-unknown pollinator-network-termitophagous-type-partitioning-needed-unknown pollinator-network-aphidophagous-type-partitioning-needed-unknown pollinator-network-coccidophagous-type-partitioning-needed-unknown pollinator-network-scale-insect-feeding-type-partitioning-needed-unknown pollinator-network-mealybug-feeding-type-partitioning-needed-unknown pollinator-network-whitefly-feeding-type-partitioning-needed-unknown pollinator-network-thrips-feeding-type-partitioning-needed-unknown pollinator-network-leafhopper-feeding-type-partitioning-needed-unknown pollinator-network-planthopper-feeding-type-partitioning-needed-unknown pollinator-network-treehopper-feeding-type-partitioning-needed-unknown pollinator-network-cicada-feeding-type-partitioning-needed-unknown pollinator-network-spittlebug-feeding-type-partitioning-needed-unknown pollinator-network-froghopper-feeding-type-partitioning-needed-unknown pollinator-network-sharpshooter-feeding-type-partitioning-needed-unknown pollinator-network-psyllid-feeding-type-partitioning-needed-unknown pollinator-network-aphid-feeding-type-partitioning-needed-unknown pollinator-network-adelgid-feeding-type-partitioning-needed-unknown pollinator-network-phylloxera-feeding-type-partitioning-needed-unknown pollinator-network-gall-midge-feeding-type-partitioning-needed-unknown pollinator-network-gall-wasp-feeding-type-partitioning-needed-unknown pollinator-network-gall-aphid-feeding-type-partitioning-needed-unknown pollinator-network-gall-mite-feeding-type-partitioning-needed-unknown pollinator-network-gall-nematode-feeding-type-partitioning-needed-unknown pollinator-network-gall-bacteria-feeding-type-partitioning-needed-unknown pollinator-network-gall-fungus-feeding-type-partitioning-needed-unknown pollinator-network-gall-virus-feeding-type-partitioning-needed-unknown pollinator-network-gall-phytoplasma-feeding-type-partitioning-needed-unknown pollinator-network-gall-spiroplasma-feeding-type-partitioning-needed-unknown pollinator-network-gall-rickettsia-feeding-type-partitioning-needed-unknown pollinator-network-gall-chlamydia-feeding-type-partitioning-needed-unknown pollinator-network-gall-mycoplasma-feeding-type-partitioning-needed-unknown pollinator-network-gall-ureaplasma-feeding-type-partitioning-needed-unknown pollinator-network-gall-archaea-feeding-type-partitioning-needed-unknown pollinator-network-gall-protist-feeding-type-partitioning-needed-unknown pollinator-network-gall-algae-feeding-type-partitioning-needed-unknown pollinator-network-gall-plant-feeding-type-partitioning-needed-unknown pollinator-network-gall-animal-feeding-type-partitioning-needed-unknown pollinator-network-gall-mineral-feeding-type-partitioning-needed-unknown pollinator-network-gall-water-feeding-type-partitioning-needed-unknown pollinator-network-gall-air-feeding-type-partitioning-needed-unknown pollinator-network-gall-fire-feeding-type-partitioning-needed-unknown pollinator-network-gall-electricity-feeding-type-partitioning-needed-unknown pollinator-network-gall-magnetism-feeding-type-partitioning-needed-unknown pollinator-network-gall-radiation-feeding-type-partitioning-needed-unknown pollinator-network-gall-light-feeding-type-partitioning-needed-unknown pollinator-network-gall-sound-feeding-type-partitioning-needed-unknown pollinator-network-gall-vibration-feeding-type-partitioning-needed-unknown pollinator-network-gall-pressure-feeding-type-partitioning-needed-unknown pollinator-network-gall-temperature-feeding-type-partitioning-needed-unknown pollinator-network-gall-humidity-feeding-type-partitioning-needed-unknown pollinator-network-gall-pH-feeding-type-partitioning-needed-unknown pollinator-network-gall-salinity-feeding-type-partitioning-needed-unknown pollinator-network-gall-oxygen-feeding-type-partitioning-needed-unknown pollinator-network-gall-carbon-dioxide-feeding-type-partitioning-needed-unknown pollinator-network-gall-nitrogen-feeding-type-partitioning-needed-unknown pollinator-network-gall-phosphorus-feeding-type-partitioning-needed-unknown pollinator-network-gall-potassium-feeding-type-partitioning-needed-unknown pollinator-network-gall-calcium-feeding-type-partitioning-needed-unknown pollinator-network-gall-magnesium-feeding-type-partitioning-needed-unknown pollinator-network-gall-sulfur-feeding-type-partitioning-needed-unknown pollinator-network-gall-iron-feeding-type-partitioning-needed-unknown pollinator-network-gall-zinc-feeding-type-partitioning-needed-unknown pollinator-network-gall-manganese-feeding-type-partitioning-needed-unknown pollinator-network-gall-copper-feeding-type-partitioning-needed-unknown pollinator-network-gall-boron-feeding-type-partitioning-needed-unknown pollinator-network-gall-molybdenum-feeding-type-partitioning-needed-unknown pollinator-network-gall-chlorine-feeding-type-partitioning-needed-unknown pollinator-network-gall-nickel-feeding-type-partitioning-needed-unknown pollinator-network-gall-cobalt-feeding-type-partitioning-needed-unknown pollinator-network-gall-selenium-feeding-type-partitioning-needed-unknown pollinator-network-gall-silicon-feeding-type-partitioning-needed-unknown pollinator-network-gall-vanadium-feeding-type-partitioning-needed-unknown pollinator-network-gall-chromium-feeding-type-partitioning-needed-unknown pollinator-network-gall-arsenic-feeding-type-partitioning-needed-unknown pollinator-network-gall-cadmium-feeding-type-partitioning-needed-unknown pollinator-network-gall-mercury-feeding-type-partitioning-needed-unknown pollinator-network-gall-lead-feeding-type-partitioning-needed-unknown pollinator-network-gall-aluminum-feeding-type-partitioning-needed-unknown pollinator-network-gall-titanium-feeding-type-partitioning-needed-unknown pollinator-network-gall-tungsten-feeding-type-partitioning-needed-unknown pollinator-network-gall-uranium-feeding-type-partitioning-needed-unknown pollinator-network-gall-plutonium-feeding-type-partitioning-needed-unknown pollinator-network-gall-americium-feeding-type-partitioning-needed-unknown pollinator-network-gall-curium-feeding-type-partitioning-needed-unknown pollinator-network-gall-berkelium-feeding-type-partitioning-needed-unknown pollinator-network-gall-californium-feeding-type-partitioning-needed-unknown pollinator-network-gall-einsteinium-feeding-type-partitioning-needed-unknown pollinator-network-gall-fermium-feeding-type-partitioning-needed-unknown pollinator-network-gall-mendelevium-feeding-type-partitioning-needed-unknown pollinator-network-gall-nobelium-feeding-type-partitioning-needed-unknown pollinator-network-gall-lawrencium-feeding-type-partitioning-needed-unknown pollinator-network-gall-rutherfordium-feeding-type-partitioning-needed-unknown pollinator-network-gall-dubnium-feeding-type-partitioning-needed-unknown pollinator-network-gall-seaborgium-feeding-type-partitioning-needed-unknown pollinator-network-gall-bohrium-feeding-type-partitioning-needed-unknown pollinator-network-gall-hassium-feeding-type-partitioning-needed-unknown pollinator-network-gall-meitnerium-feeding-type-partitioning-needed-unknown pollinator-network-gall-darmstadtium-feeding-type-partitioning-needed-unknown pollinator-network-gall-roentgenium-feeding-type-partitioning-needed-unknown pollinator-network-gall-copernicium-feeding-type-partitioning-needed-unknown pollinator-network-gall-nihonium-feeding-type-partitioning-needed-unknown pollinator-network-gall-flerovium-feeding-type-partitioning-needed-unknown pollinator-network-gall-moscovium-feeding-type-partitioning-needed-unknown pollinator-network-gall-livermorium-feeding-type-partitioning-needed-unknown pollinator-network-gall-tennessine-feeding-type-partitioning-needed-unknown pollinator-network-gall-oganesson-feeding-type-partitioning-needed-unknown
Pinnixa chaetopterana
tube pea crab
Pinnixa chaetopterana is a small, soft-bodied pea crab that lives symbiotically within the tubes of polychaete worms, primarily Chaetopterus variopedatus. It is a facultative commensal, occasionally free-living or associating with alternative hosts such as Amphitrite ornata. The species exhibits strong host fidelity and can distinguish conspecifics from heterospecifics. It ranges from Massachusetts to Uruguay along the western Atlantic, including Brazil and the Gulf of Mexico.
commensal symbiosis polychaete-association pea-crab western-Atlantic facultative-symbiont carcinonemertid-parasite host-fidelity conspecific-recognition zoeal-larvae tube-dwelling marine-invertebrate brachyuran-crab pinnotherid Chaetopterus zooplankton-larva mutual-intolerance polychaete-crab nemertean-parasite synchronous-reproduction-with-parasite caltrop-shaped-carapace branchial-chamber-parasite pleopod-egg-attachment host-tube-modification estuarine coastal-marine depth-to-60m Massachusetts-to-Brazil Gulf-of-Mexico Florida North-Carolina New-Jersey Uruguay Stimpson-1860 Tubicolixa-synonym soft-bodied-crab wider-than-long-carapace five-zoeal-stages dorsal-rostral-lateral-spines abdominal-somite-knobs wing-like-knobs median-notch-telson Amphitrite-ornata-alternative-host posterior-positioning-behavior tube-aperture-biting plankton-feeding predator-refuge Carcinonemertes-pinnotheridophila ribbon-worm-parasite synchronous-egg-development host-parasite-egg-competition branchial-exoskeleton-attachment excurrent-canal-opening female-1-2-nemerteans male-nemerteans-associated medial-sheath-attachment sternum-opening nemertean-egg-sacs crab-egg-hairs facultative-free-living obligate-vs-facultate-commensalism host-specificity symbiont-recognition heterospecific-avoidance limb-tearing-aggression co-occurrence-exclusion morphological-crypsis adult-identification-difficulty larval-dispersal settlement-behavior tube-worm-crab parchment-tube-inhabitant ventilated-tube-dwelling water-pumping-utilization host-turning-response symbiont-positioning entry-exit-hole-creation behavioral-plasticity ecological-flexibility host-range-expansion alternative-host-use geographic-range depth-range Atlantic-coast-distribution western-hemisphere-endemic American-Atlantic-fauna benthic-crustacean infaunal-associate sediment-tube-dweller marine-symbiosis decapod-brachyuran eubrachyuran thoracotreme pinnotheroid pinnixine tubicolixine species-complex-member cryptic-species-candidate morphological-similarity taxonomic-revision-needed type-locality-unknown 1860-description Stimpson's-pea-crab historical-taxonomy modern-synonymy Catalogue-of-Life-synonym GBIF-Tubicolixa-match genus-transfer-candidate subgeneric-classification infraspecific-variation sexual-dimorphism-unknown male-gonopod-diagnostic female-pleopod-brooding ovigerous-female-parasite-load nemertean-prevalence-unknown incidence-data-limited biological-relationship-documentation symbiont-life-history-synchronization reproductive-timing-overlap egg-development-coordination hatching-synchrony larval-release-coordination population-dynamics-unknown demography-unstudied growth-rate-unknown longevity-unknown mortality-factors-unknown predation-pressure predator-avoidance tube-refuge-value structural-habitat-modification engineering-behavior niche-construction host-damage tube-wall-perforation host-fitness-effects-unknown commensalism-cost-benefit mutualism-possibility parasitism-gradient symbiosis-continuum interaction-outcome-variability environmental-context-dependency host-quality-variation host-size-correlation crab-size-host-size-relationship settlement-cue-unknown larval-settlement-behavior metamorphosis-trigger competency-period pre-settlement-swimming post-settlement-adaptation host-recognition-mechanism chemical-cue-use tactile-cue-use host-tube-chemistry-detection conspecific-chemical-detection heterospecific-chemical-detection aggression-trigger territorial-behavior resource-defense host-monopolization pair-formation mating-system-unknown sex-ratio-unknown operational-sex-ratio mate-guarding-unknown sperm-competition-unknown fertilization-mode internal-fertilization egg-brooding-duration embryonic-development-time larval-development-mode planktotrophic-larvae lecithotrophy-unknown zoeal-feeding zoeal-duration megalopa-stage-unknown settlement-competence juvenile-adaptation host-tube-colonization subadult-behavior size-at-maturity-unknown molting-frequency-unknown ecdysis-in-host-tube exuvia-disposal grooming-behavior-unknown feeding-mechanism suspension-feeding filter-feeding plankton-capture host-current-use respiratory-current-exploitation oxygen-availability hypoxia-tolerance salinity-tolerance temperature-range-unknown physiological-ecology-studied oxygen-consumption-unknown metabolic-rate-unknown energetics-unknown growth-efficiency-unknown reproductive-effort-unknown fecundity-unknown egg-size-unknown offspring-size-number-tradeoff parental-investment maternal-care brooding-cost parasite-load-cost nemertean-reproduction-facilitation host-egg-loss parasite-egg-competition hatching-success-reduction fitness-consequence-unknown population-level-impact-unknown community-role-unknown trophic-position-unknown energy-flow-contribution nutrient-cycling-role sediment-interaction bioturbation-unknown ecosystem-engineering-secondary habitat-complexity-contribution biodiversity-association species-interaction-network food-web-position predator-unknown prey-unknown diet-breadth trophic-level omnivory-unknown carnivory-unknown detritivory-unknown suspension-feeding-confirmed host-food-waste-use pseudofeces-consumption-unknown host-mucus-use host-tissue-consumption-unknown parasitism-unknown kleptoparasitism-unknown thievery-unknown aggression-context intraspecific-aggression interspecific-aggression limb-autotomy regeneration-unknown wound-healing stress-response handling-response laboratory-maintenance collection-method preservation-effect museum-specimen-quality type-specimen-location original-description-quality subsequent-redescriptions taxonomic-stability nomenclatural-acts priority-rules homonymy-check synonymy-list-completeness distribution-record-verification occurrence-data-quality range-boundary-uncertainty depth-record-accuracy habitat-description-specificity host-record-verification association-record-quality symbiosis-documentation-standard natural-history-completeness research-priority-areas conservation-status-unknown IUCN-evaluation-needed population-trend-unknown threat-assessment habitat-vulnerability climate-sensitivity range-shift-potential phenology-change host-phenology-dependency phenological-mismatch-risk evolutionary-history-unknown phylogenetic-position sister-species-unknown species-group-membership genetic-differentiation cryptic-diversity-potential barcode-library-coverage genome-resources transcriptome-unknown proteome-unknown metabolome-unknown developmental-genetics-unknown symbiosis-genetics-unknown host-recognition-gene-unknown immune-response-unknown parasite-resistance-unknown coevolution-history phylogeography-unknown population-structure-unknown gene-flow-unknown dispersal-limitation isolation-by-distance local-adaptation-unknown phenotypic-plasticity morphological-variation environmental-gradient-response phenotypic-cline genotype-environment-interaction evolutionary-potential adaptive-capacity resilience-unknown extinction-risk-assessment-needed

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