Intraspecific-aggression

Guides

• Pinnixa chaetopterana

  tube pea crab

  Pinnixa chaetopterana is a small, soft-bodied pea crab that lives symbiotically within the tubes of polychaete worms, primarily Chaetopterus variopedatus. It is a facultative commensal, occasionally free-living or associating with alternative hosts such as Amphitrite ornata. The species exhibits strong host fidelity and can distinguish conspecifics from heterospecifics. It ranges from Massachusetts to Uruguay along the western Atlantic, including Brazil and the Gulf of Mexico.

  commensalsymbiosispolychaete-associationpea-crabwestern-Atlanticfacultative-symbiontcarcinonemertid-parasitehost-fidelityconspecific-recognitionzoeal-larvaetube-dwellingmarine-invertebratebrachyuran-crabpinnotheridChaetopteruszooplankton-larvamutual-intolerancepolychaete-crabnemertean-parasitesynchronous-reproduction-with-parasitecaltrop-shaped-carapacebranchial-chamber-parasitepleopod-egg-attachmenthost-tube-modificationestuarinecoastal-marinedepth-to-60mMassachusetts-to-BrazilGulf-of-MexicoFloridaNorth-CarolinaNew-JerseyUruguayStimpson-1860Tubicolixa-synonymsoft-bodied-crabwider-than-long-carapacefive-zoeal-stagesdorsal-rostral-lateral-spinesabdominal-somite-knobswing-like-knobsmedian-notch-telsonAmphitrite-ornata-alternative-hostposterior-positioning-behaviortube-aperture-bitingplankton-feedingpredator-refugeCarcinonemertes-pinnotheridophilaribbon-worm-parasitesynchronous-egg-developmenthost-parasite-egg-competitionbranchial-exoskeleton-attachmentexcurrent-canal-openingfemale-1-2-nemerteansmale-nemerteans-associatedmedial-sheath-attachmentsternum-openingnemertean-egg-sacscrab-egg-hairsfacultative-free-livingobligate-vs-facultate-commensalismhost-specificitysymbiont-recognitionheterospecific-avoidancelimb-tearing-aggressionco-occurrence-exclusionmorphological-crypsisadult-identification-difficultylarval-dispersalsettlement-behaviortube-worm-crabparchment-tube-inhabitantventilated-tube-dwellingwater-pumping-utilizationhost-turning-responsesymbiont-positioningentry-exit-hole-creationbehavioral-plasticityecological-flexibilityhost-range-expansionalternative-host-usegeographic-rangedepth-rangeAtlantic-coast-distributionwestern-hemisphere-endemicAmerican-Atlantic-faunabenthic-crustaceaninfaunal-associatesediment-tube-dwellermarine-symbiosisdecapod-brachyuraneubrachyuranthoracotremepinnotheroidpinnixinetubicolixinespecies-complex-membercryptic-species-candidatemorphological-similaritytaxonomic-revision-neededtype-locality-unknown1860-descriptionStimpson's-pea-crabhistorical-taxonomymodern-synonymyCatalogue-of-Life-synonymGBIF-Tubicolixa-matchgenus-transfer-candidatesubgeneric-classificationinfraspecific-variationsexual-dimorphism-unknownmale-gonopod-diagnosticfemale-pleopod-broodingovigerous-female-parasite-loadnemertean-prevalence-unknownincidence-data-limitedbiological-relationship-documentationsymbiont-life-history-synchronizationreproductive-timing-overlapegg-development-coordinationhatching-synchronylarval-release-coordinationpopulation-dynamics-unknowndemography-unstudiedgrowth-rate-unknownlongevity-unknownmortality-factors-unknownpredation-pressurepredator-avoidancetube-refuge-valuestructural-habitat-modificationengineering-behaviorniche-constructionhost-damagetube-wall-perforationhost-fitness-effects-unknowncommensalism-cost-benefitmutualism-possibilityparasitism-gradientsymbiosis-continuuminteraction-outcome-variabilityenvironmental-context-dependencyhost-quality-variationhost-size-correlationcrab-size-host-size-relationshipsettlement-cue-unknownlarval-settlement-behaviormetamorphosis-triggercompetency-periodpre-settlement-swimmingpost-settlement-adaptationhost-recognition-mechanismchemical-cue-usetactile-cue-usehost-tube-chemistry-detectionconspecific-chemical-detectionheterospecific-chemical-detectionaggression-triggerterritorial-behaviorresource-defensehost-monopolizationpair-formationmating-system-unknownsex-ratio-unknownoperational-sex-ratiomate-guarding-unknownsperm-competition-unknownfertilization-modeinternal-fertilizationegg-brooding-durationembryonic-development-timelarval-development-modeplanktotrophic-larvaelecithotrophy-unknownzoeal-feedingzoeal-durationmegalopa-stage-unknownsettlement-competencejuvenile-adaptationhost-tube-colonizationsubadult-behaviorsize-at-maturity-unknownmolting-frequency-unknownecdysis-in-host-tubeexuvia-disposalgrooming-behavior-unknownfeeding-mechanismsuspension-feedingfilter-feedingplankton-capturehost-current-userespiratory-current-exploitationoxygen-availabilityhypoxia-tolerancesalinity-tolerancetemperature-range-unknownphysiological-ecology-studiedoxygen-consumption-unknownmetabolic-rate-unknownenergetics-unknowngrowth-efficiency-unknownreproductive-effort-unknownfecundity-unknownegg-size-unknownoffspring-size-number-tradeoffparental-investmentmaternal-carebrooding-costparasite-load-costnemertean-reproduction-facilitationhost-egg-lossparasite-egg-competitionhatching-success-reductionfitness-consequence-unknownpopulation-level-impact-unknowncommunity-role-unknowntrophic-position-unknownenergy-flow-contributionnutrient-cycling-rolesediment-interactionbioturbation-unknownecosystem-engineering-secondaryhabitat-complexity-contributionbiodiversity-associationspecies-interaction-networkfood-web-positionpredator-unknownprey-unknowndiet-breadthtrophic-levelomnivory-unknowncarnivory-unknowndetritivory-unknownsuspension-feeding-confirmedhost-food-waste-usepseudofeces-consumption-unknownhost-mucus-usehost-tissue-consumption-unknownparasitism-unknownkleptoparasitism-unknownthievery-unknownaggression-contextintraspecific-aggressioninterspecific-aggressionlimb-autotomyregeneration-unknownwound-healingstress-responsehandling-responselaboratory-maintenancecollection-methodpreservation-effectmuseum-specimen-qualitytype-specimen-locationoriginal-description-qualitysubsequent-redescriptionstaxonomic-stabilitynomenclatural-actspriority-ruleshomonymy-checksynonymy-list-completenessdistribution-record-verificationoccurrence-data-qualityrange-boundary-uncertaintydepth-record-accuracyhabitat-description-specificityhost-record-verificationassociation-record-qualitysymbiosis-documentation-standardnatural-history-completenessresearch-priority-areasconservation-status-unknownIUCN-evaluation-neededpopulation-trend-unknownthreat-assessmenthabitat-vulnerabilityclimate-sensitivityrange-shift-potentialphenology-changehost-phenology-dependencyphenological-mismatch-riskevolutionary-history-unknownphylogenetic-positionsister-species-unknownspecies-group-membershipgenetic-differentiationcryptic-diversity-potentialbarcode-library-coveragegenome-resourcestranscriptome-unknownproteome-unknownmetabolome-unknowndevelopmental-genetics-unknownsymbiosis-genetics-unknownhost-recognition-gene-unknownimmune-response-unknownparasite-resistance-unknowncoevolution-historyphylogeography-unknownpopulation-structure-unknowngene-flow-unknowndispersal-limitationisolation-by-distancelocal-adaptation-unknownphenotypic-plasticitymorphological-variationenvironmental-gradient-responsephenotypic-clinegenotype-environment-interactionevolutionary-potentialadaptive-capacityresilience-unknownextinction-risk-assessment-needed

• Trichonephila clavata

  Jorō spider, Joro Spider, Parachute spider

  Trichonephila clavata, commonly known as the Jorō spider, is a large orb-weaving spider native to East Asia that has become established as an invasive species in the southeastern United States since approximately 2010. First confirmed in Georgia in 2014, it has expanded rapidly across multiple states through a combination of ballooning dispersal and human-mediated transport. The species is notable for its substantial size, striking coloration, and extensive golden webs, but poses minimal risk to humans due to small fangs and docile behavior. Its physiological adaptations—including higher metabolic rate, faster heart rate, and greater cold tolerance than its congener Trichonephila clavipes—suggest potential for continued northward range expansion.

  invasive-speciesorb-weaverballooningurban-wildlifebiological-controlcold-tolerancestartle-responsecannibalismkleptoparasitismJapanese-folkloreNephilidaeTrichonephilaspider-silkrange-expansionphysiological-adaptationurban-ecologyagricultural-pest-predatoriNaturalistGeorgiaMarylandEast-AsiaUnited-Statesspider-bitevenomdocilenon-aggressiveweb-architectureegg-sacoverwinteringsexual-dimorphismmetabolic-rateheart-ratefreeze-toleranceroad-ecologyprey-capturecompetitionnative-spider-impactspublic-perceptionmedia-sensationalismmanagementpesticide-alternativesbroom-removalornamentalstructural-pestornithologycardinalweb-strengthtrophic-interactionsDNA-metabarcodingdiet-analysisspecies-distribution-modelphysiological-entomologybehavioral-ecologyarachnologyballooning-dispersalhuman-mediated-dispersalhitchhikercargo-stowawayport-of-entryclimate-changewarmingnorthward-expansionDMVHoward-CountyElkridgeIlchesterWest-Elkridgebeech-forestsecondary-forestsuburbanroadsidetelephone-polestreetlampforest-edgedisturbance-toleranceprey-detectionvibrational-noiseauditory-disturbancemotionlessfreezingshynessHannibal-Lecterstink-buglanternflyevolutionary-historyancient-associationecosystem-servicebiocontrolnatural-enemyinvasive-pestbrown-marmorated-stink-bugspotted-lanternflyHalyomorpha-halysLycorma-delicatulacompetitive-exclusiondominant-orb-weaverabundantpopulation-growthrapid-spreadrange-span120,000-km2physiological-plasticitymetabolismtwice-as-high77%-higher-heart-rate74%-freeze-survivaldevelopment-rateseasonal-windowlife-cycle-completionthermal-limitationcongener-comparisonnatural-dispersalhuman-transportiNaturalist-recordcitizen-scienceconfirmed-sightingbreeding-populationmale-and-femalegravid-femaleegg-casestowawaycargo-containerport-introductionlocal-portvehicle-transporthitchhikingaccelerated-arrivalseveral-yearsnatural-meansprognosticationpredictionmodelsuitable-habitatabundant-suitable-habitateastern-North-Americawestern-North-Americapotential-rangeinvasive-rangepotential-impactenvironmental-impactecological-impactecosystem-impactmonitorcautiousevidence-basedreasonable-journalismsensationalismexaggerationuncritical-acceptanceharmlesscarefully-monitorednext-stepsresearch-opportunitycall-for-researchknowledge-gaplittle-knownnovel-ecosystemspreadingrapidly-expandinginvasive-predatorinvasive-orb-weavernon-nativeintroducedestablishedcolonizedthrivingdoing-finehappymerely-20-minutes-awayroad-tripvisitaccess-spreadbeech-leaf-diseaseBLDstate-forestremarkable-colonyamongst-beech-treesconfirmed-prognosticationDavis-and-Frickescape-relative-warmthexpand-range-northwardeastern-seaboardmerely-20-minutes-away-from-homeresist-opportunityamazing-predatorstales-of-arrivalprevious-episodes20222024reduce-angstlarge-non-native-spiderestablishing-in-DMVfacts-presentedpast-episodevenomous-terrible-painfulNahexpert-Rick-Hoebekerisks-smallpuny-fangsunlikely-pierce-skinvideocompletely-non-aggressiveeastern-Howard-Countyhaphazard-webslittered-with-remainsformer-victimsleavesshed-exoskeletonsmuch-larger-femaledwarfs-matepositioned-just-abovestrand-of-silkspinneretsunderside-of-abdomennear-red-moundrelative-to-handlarge-and-docilewait-and-seewhat-Jorō-means-to-ecosystemshelp-other-spidersput-a-beat-down-on-invasive-pestsstink-bugsspotted-lanternfliespassive-huntersenormous-webslarger-than-a-meter-in-diametercapture-prey-snared-in-silkarachnophobes-scaryarachnophiles-beautifulimportant-ecosystem-servicescrop-pestsnative-rangeAsiahaving-an-old-friend-over-for-dinnerreunitefirst-timelarge-spidersjuicy-prey-itemsfeathered-and-non-feathered-reptilesposes-no-known-threatdifficult-to-predict-impactnon-native-speciesexperts-suggestbeyond-scary-miengive-indigenous-large-orb-weavers-run-for-their-moneyblack-and-yellow-garden-spidermarbled-orb-weaverspotted-orb-weaverother-parts-of-worldmost-abundant-and-dominant-orb-weaverwhat-will-it-meanonly-time-will-tellfinal-tidbitJorō-shapeshifterJorō-gumobeautiful-womanseduces-menbinds-with-silkdevoursbad-dateacknowledgementsRick-Hoebekeidentifying-JorōinsightsDavid-CoyleBob-Bellingergreat-imagesknowledgestudiesVeni-vidi-viciNelsen-et-al.Physiological-evaluationNephila-clavata-newly-recordedHoebeke-et-al.Life-Cycle-Habitat-and-VariationMooreStartle-responsesDavis-and-AneraoHow-Urban-Tolerant-Are-TheyDavis-et-al.inspirationdetailsstarsDave-ClementMiri-TalabacMaddie-Potterhooked-up-with-colonydifferentiateresources.ipmcenters.orglinkJorō-guruDavid-Coyle's-takeYouTubecalm-your-fearsJournal-of-Economic-EntomologyJournal-of-Medical-Entomologychemical-management-strategieseffective-management-methodsspider-control-productssafe-and-effectiveunlikely-to-bitegenerally-harmlessbig-spiderscolorful-spidersinvasive-spidersin-your-face-spiderswebs-on-homeslandscape-plantsstructuresmiddle-of-where-people-live-work-and-playnorthern-Georgia-2014presumed-arrived-few-years-prior15-yearsincreased-dramaticallymuch-of-Southeastdisjunct-populationsnortheastern-U.S.firestorm-of-interestrapidly-expanding-rangesoutheastern-statesmany-countiesCarolinasTennesseeOklahomamore-than-a-centuryparts-of-Floridararely-Pennsylvaniatropicsthermal-limitationshigher-metabolismfaster-heart-ratebetter-ability-to-tolerate-freezingcomplete-life-cycle-rapidlychilly-temperaturessuite-of-adaptationsexpand-northwardwarm-loving-cousinfindings-support-potentialmuch-yet-to-be-learnedsuccessfully-survive-northern-wintersnation-and-world-warmsouthern-species-expandhigher-latitudes-and-altitudesrapidly-range-expansion-will-occurtypical-mode-of-dispersalaerial-dispersalballooning-on-strands-of-silkCharlotte's-babiesCharlotte's-Webspectacular-monikerparachute-spiderrain-down-from-airplaneslong-distance-transithuman-assistancegood-hitchhikersinseminated-gravid-femaleegg-case-stowawayarrival-in-DMVseveral-years-by-natural-meansnew-introduction-at-local-porthuman-assist-in-vehiclesaccelerate-arrivalterrible-and-painfulsmall-fangsvisited-cousingolden-silk-spiderCosta-Ricaexceeding-meter-diameterindigenous-large-orb-weaversrun-for-their-moneyminimal-direct-impactfirestormAndrew-DavisBenjamin-FrickUniversity-of-Georgiapast-weeknative-to-eastern-AsiaJapanChinaKoreaTaiwanknown-in-Georgia-since-2013spreading-rapidlyjoined-its-cousinTrichonephila-clavipesestablished-over-160-yearsbetter-tolerate-freezingsurvive-northern-wintersrapidly-range-expansiontypical-modelong-distancenew-introductionhuman-assistarachnophobesarachnophilesbrown-marmorated-stink-bugsdifficult-to-predictscary-mienindigenous-orb-weaversMary-NouriSarah-MorganWAMUrelatively-harmlessspider-seasonfallleaves-changing-colorcool-crisp-autumn-airpumpkin-spiceextension-entomologistSoutheastnot-just-spiders20-feet-widehomespeople-live-work-and-play15-years-or-soinflux-of-callspublicentomology-extension-programsmedia-frenzytwo-questionsget-rid-of-Jorō-spidersare-Jorō-spiders-dangerousClemson-UniversitySouthern-Adventist-UniversityWashington-CollegeUnion-Collegetwo-new-studiestest-methods-for-eliminatinghow-likely-to-biteharmscoured-internetrecommended-treatment-optionsspider-specific-management-recommendationsmedically-relevantwidow-spidersbrown-reclusesdo-not-enter-homesrelatively-newrecommended-productslabelled-spider-control-productstested-productsseveral-products-eliminatenot-legal-pesticidesmachine-lubricantnot-recommendedcommercially-availablestrongly-encouragelabelled-productseasiest-and-cheapestwithout-chemicalsbroom-or-stickwalked-into-spider-webrode-biketook-spider-to-facenot-great-feelingwonder-if-biteygive-bitten-superpowersunique-studiesevaluated-behaviorsreact-to-peoplerather-drop-off-webhang-around-and-bitereally-provokevolunteered-to-get-bittenyes-you-read-correctlyakin-to-mosquito-bitelittle-rednesslittle-swellinglargely-gone-next-dayno-acquired-superpowerstake-home-messagetwofoldsimplest-and-most-effectivewithout-pesticidesfeel-you-need-pesticidesproduct-labeled-for-spider-controltry-not-to-freak-outdoesn't-want-to-be-on-youodds-of-getting-bitten-extremely-lowenjoy-glorious-fall-weatherdrink-pumpkin-spiced-beveragewatch-where-you-walkKeep-calm-and-carry-onbites-unlikelycause-minimal-discomfortassociate-professorDepartment-of-Forestry-and-Environmental-Conservationsocialsemailextensionfeaturedpest-managementEntomology-Todaylatest-postsemail-subscriptionMarch-2022wonderedmake-its-way-to-DMVSeptember-2022two-observationsthree-years40-sightingshappy-and-doing-just-fineseveral-Howard-County-locationsweek-or-so-agoteam-of-scientistsUniversitymissionkilling-ultra-valuable-beech-treesthriving-amongst-beech-treesvisit-amazing-predatorsmature-femalesboth-Trichonephila-speciesthree-locally-common-orb-weaving-speciestime-spent-immobilemild-disturbancebrief-puff-of-airair-puff-response-datafive-other-North-American-speciespublished-literature453-observationsfreezing-behavior10-spider-speciesremained-immobile-under-a-minuteboth-Trichonephila-spidersover-an-hourunprecedentedshyest-ever-documentedtolerate-urban-environmentsremaining-motionlessfleeingnonsexual-cannibalismmating-activityfemales-cannibalize-malesfood-availabilityterritorial-aggressionSoutheastern-United-Statesexpanding-rangeshybehavioral-reactionsdescriptive-observationsphoto-documentationanecdotal-observationscontrolled-pairingslabnatural-websfieldone-female-biting-and-killing-anothershort-fightsimilar-sizecontainer25-trialsfights-ensued-40%different-sizes27-trialsfights-happened-18%larger-females-not-always-aggressor52-lab-trialssix-bouts9%direct-killingfield-trialsempty-web14-trialsone-fight7%aggressor-killing-and-wrappingsilkaway-from-any-webnot-necessarily-territorialityintraspecific-aggressionprovoked-or-stresseddeserves-more-studynewly-invasivebiologyphysiologynew-rangeclosely-related-speciessuccessfully-established160-yearsinvestigationcompareonline-recordsiNaturalist.orgseasonal-distributionstimingfemalesphysiological-traitsenvironmental-tolerancemetabolic-ratescardiac-functioningcold-exposuresurvivalbrief-below-freezing-temperatureshorter-seasoncomplete-lifecyclenarrow-periodsuitable-weatherinherently-higher-metabolismlow-temperaturesurvive-better74%-comp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