Northern-cardinal

Guides

• Ixodes affinis

  Ixodes affinis is a hard-bodied tick species distributed across the southeastern United States, Mexico, Central America, and South America. Unlike its congener Ixodes scapularis, this species does not bite humans but serves as a competent vector for Borrelia burgdorferi sensu stricto, the causative agent of Lyme disease. Immature stages feed on birds and small mammals, while adults parasitize larger mammals including deer and dogs. Its expanding range into southeastern Virginia and association with migratory birds has raised concerns about its role in pathogen dispersal.

  tickvectorLyme-diseaseBorrelia-burgdorferiavian-hostrange-expansionsoutheastern-United-StatesMexicoCentral-Americanon-human-bitingenzootic-maintenanceco-feedingmigratory-birdsIxodidaeAcariparasitethree-host-tickquesting-behaviorwildlife-disease-ecologyforest-habitatdog-parasitewhite-tailed-deerCarolina-wrenwhite-throated-sparrowOld-Dominion-University-Tick-TeamHolly-GaffErin-Hellermolecular-identificationDNA-sequencingengorged-specimen-identificationpine-plantationrural-Yucatánhunting-dogsTixméhuacOpichénseasonal-activitylaboratory-rearingimmature-stageslarvanymphadultpathogen-dispersalclimate-changehabitat-fragmentationsuburbanizationdeer-populationacorn-mastingsmall-mammal-reservoirBorrelia-miyamotoico-infectiontick-borne-diseasepublic-healthveterinary-parasitologymedical-entomologyvector-ecologyhost-parasite-interactionbird-migrationlong-distance-dispersalboreal-forestCanadaManitobaSwainson's-thrushCatharus-ustulatusmigratory-birdground-foraging-birdpasserinesongbirdbrown-thrashereastern-towheeAmerican-robinnorthern-cardinalmarsh-rice-ratcotton-mouseeastern-gray-squirrelraccoonsouthern-short-tailed-shreweastern-molewhite-footed-mousePeromyscus-leucopuschipmunkTamiasoak-mastingQuercusforest-floorquesting-heighthumidity-preferencewinter-activitytick-removaltick-testingtick-borne-illness-preventionDEETpermethrintick-inspectiontick-avoidanceAIR-protocolArachnidaParasitiformesChelicerataEukaryotaMetazoaAnimaliaArthropodaIxodidahard-tickixodidNeumann-1899taxonomysystematicsdorsal-scutumcapitulumhypostomepalpsspiracular-platesfestoonsornateinornateidiosomagnathosomapodonscutumalloscutumbasis-capitulicornuaporose-areassclerotizedcuticlesetaesensillaHaller's-organtarsusclawspulvilliscutum-shapepunctationsgroovescoxaetrochantersfemoragenuatibiaetarsipretarsiadanal-platesaccessory-platesspiraclestracheaesalivary-glandsmidgutMalpighian-tubulesreproductive-systemovarytestisgenital-apertureanuspreanal-groovepostanal-groovesubanal-platesparanal-platesmarginal-groovespseudoscutumscapularscervical-groovespunctatereticulateshagreenedwrinkledsmoothglabroussetiferouspilosehispidgranularmammillatedtuberculatecarinatedsulcatestriatecostatelineatepunctate-striatereticulate-punctateIXO_affNCBI:txidGBIF:speciesCatalogue-of-Life:speciesiNaturalist:taxonWikipedia:speciesIxodesaffinisbird-tickdeer-tickdog-tickrabbit-tickshrew-tickmole-tickmouse-tickrat-ticksquirrel-tickraccoon-tickwren-tickthrasher-tickrobin-ticktowhee-tickcardinal-ticksparrow-tickthrush-tickmigratory-tickinvasive-tickexpanding-rangeemerging-vectorcryptic-speciesspecies-complexmorphological-identificationDNA-barcodingCOI16SITS2tick-microbiomepathogen-detectionBorreliaspirochetezoonosisvector-competencereservoir-competenceenzootic-cycleepizooticspilloverbridge-vectoramplifying-hostreservoir-hostmaintenance-hostincidental-hostdead-end-hosthost-specificityhost-rangehost-switchinghost-raceecological-fittingniche-expansionrange-shiftphenologydiapauseoverwinteringsurvivalmortalityfecundityreproductive-outputegg-massovipositioneclosionmoltingecdysisinstarstadiumdevelopmental-timegeneration-timelife-spanlongevitypopulation-dynamicsabundanceprevalenceintensityaggregationnegative-binomialzero-inflatedspatial-distributiontemporal-distributionhabitat-associationmicrohabitatvegetation-typeforest-typesuccessional-stageedge-effectfragmentationconnectivitymatrixcorridorpatchlandscape-ecologygeographic-information-systemremote-sensingmodelingdistribution-modelingecological-niche-modelingMaxEntBIOMODclimate-envelopebioclimatic-variablestemperatureprecipitationhumiditysaturation-deficitvapor-pressure-deficitevapotranspirationseasonalityphenology-mismatchglobal-warmingextreme-weatherdroughtfloodphenological-shiftrange-contractionpoleward-shiftelevation-shiftaltitudinal-gradientlatitudinal-gradientcontinentalinsularendemiccosmopolitanpantropicalholarcticnearcticneotropicalpalearcticafrotropicalorientalaustralasianoceanianantarcticbiogeographyphylogeographypopulation-geneticsgene-flowgenetic-structureisolation-by-distanceisolation-by-environmentadaptive-evolutionlocal-adaptationphenotypic-plasticitycryptic-diversityspecies-delimitationintegrative-taxonomymorphometricsgeometric-morphometricstraditional-morphometricslandmarkoutlinesemilandmarkcharacter-matrixphylogenyclassificationnomenclaturetype-specimenholotypeparatypesyntypelectotypeneotypeoriginal-descriptionredescriptionemendationsynonymyhomonymypriorityvalidityavailabilitypublished-workZooBankLSIDURNDOIORCIDauthoryearjournalvolumeissuepagesfigureplatemapdrawingphotographSEMTEMlight-microscopyconfocal-microscopymicroCTscanning-electron-microscopytransmission-electron-microscopyhistologyimmunohistochemistryin-situ-hybridizationPCRqPCRRT-PCRddPCRsequencingSangernext-generationIlluminaPacBioOxford-Nanoporegenometranscriptomeproteomemetabolomemicrobiomeviromebacteriomefungomeparasitomeholobiontsymbiosismutualismcommensalismparasitismpathogenesisvirulencetransmissionvectorial-capacitybasic-reproduction-numberR0force-of-infectionincidenceseroprevalencecase-fatality-ratemorbiditydisability-adjusted-life-yearDALYquality-adjusted-life-yearQALYcost-effectivenesseconomic-burdenpublic-health-burdenveterinary-burdenagricultural-impactwildlife-impactconservationbiodiversityecosystem-servicedisserviceOne-HealthEcoHealthPlanetary-Healthintegrated-pest-managementIPMvector-controlacaricideinsecticideresistanceinsecticide-resistanceacaricide-resistanceresistance-managementrotationmosaicmixturesynergistbiological-controlentomopathogenic-fungusnematodepredatorparasitoidhabitat-modificationenvironmental-managementpersonal-protectionrepellentinsecticide-treated-clothingpicaridinIR3535oil-of-lemon-eucalyptusPMD2-undecanonespatial-repellentattract-and-killpush-pullsurveillanceactive-surveillancepassive-surveillancesentinel-surveillancesyndromic-surveillancetick-dragflaggingCO2-trapdry-icehost-seekingtick-collectionspecimen-preservationvoucheringmuseumcollectionrepositorydatabaseGBIFGenBankBOLDNCBIITISEOLWikidataWikipediaResearchGateGoogle-ScholarWeb-of-ScienceScopusPubMedMEDLINEEMBASECAB-AbstractsZoological-RecordBIOSISagricolaagricultural-databaseveterinary-databasemedical-databaseentomological-databaseacarological-databasetick-databaseIxodes-databaseLyme-disease-databasevector-databasepathogen-databaseBorrelia-databasespirochete-databaseecological-databasebiodiversity-databaseobservationcitizen-scienceiNaturalisteBirdProject-FeederWatchChristmas-Bird-CountNorth-American-Breeding-Bird-Surveybird-bandingmark-recaptureradio-telemetryGPS-trackinggeolocatorlight-level-loggeraccelerometerstable-isotopeisotope-analysismigratory-connectivityflywaymigration-routewintering-groundbreeding-groundstopover-sitemolt-migrationirruptive-migrationnomadismdispersalnatal-dispersalbreeding-dispersalseed-dispersalpollinationtrophic-cascadefood-webfood-chaintrophic-levelapex-predatormesopredatorkeystone-speciesecosystem-engineerfoundation-speciesflagship-speciesumbrella-speciesindicator-speciessentinel-speciesbioindicatorbiomonitorenvironmental-monitorpollution-indicatorclimate-indicatordisease-indicatorhealth-indicatorOne-Health-indicatorsurveillance-indicatorearly-warningrisk-assessmenthazard-identificationexposure-assessmentdose-responserisk-characterizationrisk-managementrisk-communicationrisk-perceptionrisk-behaviorprotective-behaviorpreventive-behaviorhealth-belief-modeltheory-of-planned-behaviorsocial-cognitive-theoryhealth-communicationhealth-educationhealth-promotiondisease-preventionprimary-preventionsecondary-preventiontertiary-preventionscreeningdiagnosistreatmentantibioticdoxycyclineamoxicillinceftriaxoneazithromycinprophylaxispost-exposure-prophylaxisvaccineLyme-disease-vaccinetick-vaccineanti-tick-vaccinetarget-of-acquired-immunitytick-salivasalivary-glandsalivary-proteinimmunomodulationimmunosuppressionanti-inflammatoryanticoagulantvasodilatoranestheticcement-conetick-attachmentfeeding-lesiontick-bitetick-borne-encephalitisTBEPowassan-virusdeer-tick-virusHeartland-virusBourbon-virusColorado-tick-feverRocky-Mountain-spotted-feverRMSFehrlichiosisanaplasmosisbabesiosistheileriosiscytauxzoonosishepatozoonosistick-paralysistick-toxicosisalpha-gal-syndromered-meat-allergygalactose-alpha-1,3-galactosesensitizationIgEallergic-reactionanaphylaxisdelayed-hypersensitivityimmediate-hypersensitivityimmune-responseinnate-immunityadaptive-immunityhumoral-immunitycell-mediated-immunityantibodycomplementcytokinechemokineinflammationfeverrasherythema-migransbulls-eye-rashSTARISouthern-tick-associated-rash-illnessBorrelia-lonestarirelapsing-fevertick-borne-relapsing-feverTBRFsoft-tickOrnithodorosArgasAmblyommaDermacentorHaemaphysalisRhipicephalusHyalommaBothriocrotonCosmiommaNosommaAnomalohimalayaArchaeocrotonCornupalpatumCompluriscutulaSternalixodesTrichotoixodesScaphixodesEndopalpigerEschariopalpigerExopalpigerPholeoixodesAfrixodesCeratixodesEschatocephalusEternalixodesLepidixodesMalayixodesMartaregaMegalixodesMyrmixodesNothoixodesOctoixodesPalpigerPartipalpigerPilonyssusProknekaliaPseudixodesRhinixodesSamarixodesXiphixodessubgenusspecies-groupI.-ricinus-complexI.-persulcatus-complexI.-scapularis-complexI.-pacificus-complexI.-affinis-groupI.-cookei-groupI.-marxi-groupI.-angustus-groupI.-soricis-groupI.-uriae-groupI.-holocyclus-groupI.-ceylonensis-groupI.-granulatus-groupI.-ovatus-groupI.-nipponensis-groupI.-persulcatus-groupI.-pavlovskyi-groupI.-ricinus-groupI.-scapularis-groupI.-pacificus-groupI.-affinisI.-auritulusI.-australiensisI.-boliviensisI.-brunneusI.-cookeiI.-cooleyiI.-cornuaeI.-cornuatusI.-crenulatusI.-damminiI.-dawsoniI.-dentatusI.-diomedeaeI.-domergueiI.-eadsiI.-eastoniI.-eldaricusI.-elongatusI.-eudyptidisI.-euplectropiI.-evansiI.-fecialisI.-festaiI.-frontalisI.-galapagoensisI.-ghilaroviI.-gibbosusI.-granulatusI.-gregsoniI.-hearleiI.-hexagonusI.-hirstiI.-holocyclusI.-hoogstraaliI.-howelliI.-hyattiI.-jacksoniI.-jellisoniI.-kashmiricusI.-kazakstaniI.-kingiI.-kohlsiI.-kopsteiniI.-kuntziI.-laguriI.-latusI.-layingensisI.-lewisiI.-lividusI.-longiscutatusI.-loricatusI.-luciaeI.-malayensisI.-marxiI.-masloviI.-matopiI.-minorI.-monospinosusI.-moreliI.-moschiferiI.-murisI.-murreletiI.-myospalacisI.-nchisiensisI.-necromysI.-neitziI.-nipponensisI.-nuttalliI.-ochotonaeI.-okapiaeI.-oldiI.-ornithorhynchiI.-ovatusI.-pacificusI.-pararicinusI.-pavlovskyiI.-percavatusI.-peromysciI.-persulcatusI.-pomerantziI.-pterodromaeI.-quadratusI.-raccoonisI.-rageauiI.-randrianasoloiI.-rasusI.-redikorzeviI.-ricinusI.-rothschildiI.-rotundatusI.-rubicundusI.-rugicollisI.-rugosusI.-sapeloensisI.-sarsiI.-scapularisI.-schillingsiI.-sculptusI.-semenoviI.-soricisI.-spinaeI.-steiniI.-stilesiI.-stromiI.-subterranusI.-tancitariusI.-tanukiI.-tapirusI.-tasmaniI.-tayloriI.-tertiariusI.-texanusI.-theileraeI.-thomasaeI.-tovariI.-transvaalensisI.-triangulicepsI.-trichosuriI.-turdusI.-uriaeI.-vandenbergiI.-vanidicusI.-ventalloiI.-vespertilionisI.-victoriensisI.-walkeraeI.-werneriI.-woodiI.-zaglossiI.-zairensistick-taxonomytick-systematicstick-phylogenytick-evolutiontick-biogeographytick-ecologytick-physiologytick-behaviortick-reproductiontick-developmenttick-nutritiontick-metabolismtick-respirationtick-excretiontick-osmoregulationtick-neurobiologytick-sensory-biologytick-chemoreceptiontick-thermoreceptiontick-mechanoreceptiontick-photoreceptiontick-circadian-rhythmtick-biological-clocktick-seasonalitytick-diapausetick-overwinteringtick-cold-hardinesstick-desiccation-resistancetick-water-balancetick-cuticletick-moltingtick-ecdysistick-growthtick-feedingtick-engorgementtick-blood-mealtick-digestiontick-salivationtick-saliva-compositiontick-salivary-gland-transcriptometick-salivary-proteometick-pharmacologytick-toxicologytick-venomtick-paralysis-toxintick-borne-pathogentick-borne-virustick-borne-bacteriumtick-borne-protozoantick-borne-helminthtick-borne-rickettsiatick-borne-ehrlichiatick-borne-anaplasmatick-borne-borreliatick-borne-babesiatick-borne-theileriatick-borne-hepatozoontick-borne-coxiellatick-borne-francisellatick-borne-bartonellatick-borne-chlamydiatick-borne-mycoplasmatick-borne-spiroplasmatick-borne-arbovirustick-borne-flavivirustick-borne-bunyavirustick-borne-reovirustick-borne-orbivirustick-borne-coltivirustick-borne-nairovirustick-borne-phlebovirustick-borne-thogotovirustick-borne-orthomyxovirustick-borne-arenavirustick-borne-hantavirustick-borne-paramyxovirustick-borne-rhabdovirustick-borne-bornavirustick-borne-filovirustick-borne-coronavirustick-borne-poxvirustick-borne-herpesvirustick-borne-adenovirustick-borne-papillomavirustick-borne-polyomavirustick-borne-circovirustick-borne-parvovirustick-borne-anellovirustick-borne-dependovirustick-borne-tenuivirustick-borne-deltavirustick-borne-hepadnavirustick-borne-retrovirustick-borne-lentivirustick-borne-spumavirustick-borne-metavirustick-borne-pseudovirustick-borne-hemivirustick-borne-novirhabdovirustick-borne-peribunyavirustick-borne-phenuivirustick-borne-bandavirustick-borne-phasivirustick-borne-tulasvirustick-borne-orthonairovirustick-borne-shavivirustick-borne-sathuperi-virustick-borne-salehabad-virustick-borne-dera-ghazi-khan-virustick-borne-hughes-virustick-borne-farallon-virustick-borne-punta-salinas-virustick-borne-sapphire-II-virustick-borne-zirqa-virustick-borne-keterah-virustick-borne-thiafora-virustick-borne-erve-virustick-borne-malsoor-virustick-borne-tibet-virustick-borne-qalyub-virustick-borne-yogue-virustick-borne-kasokero-virustick-borne-lebombo-virustick-borne-nyamanini-virustick-borne-midway-virustick-borne-tokar-virustick-borne-absettarov-virustick-borne-hypr-virustick-borne-kumlinge-virustick-borne-omsk-hemorrhagic-fever-virustick-borne-langat-virustick-borne-louping-ill-virustick-borne-kyasanur-forest-disease-virustick-borne-alkhurma-hemorrhagic-fever-virustick-borne-powassan-virustick-borne-deer-tick-virustick-borne-heartland-virustick-borne-bourbon-virustick-borne-colorado-tick-fever-virustick-borne-eyach-virustick-borne-banna-virustick-borne-seadornavirustick-borne-great-island-virustick-borne-kemerovo-virustick-borne-lipovnik-virustick-borne-tribec-virustick-borne-chobar-gorge-virustick-borne-wad-Medani-virustick-borne-dugbe-virustick-borne-crimean-congo-hemorrhagic-fever-virustick-borne-nairobi-sheep-disease-virustick-borne-ganjam-virustick-borne-bandia-virustick-borne-sierra-nevada-virustick-borne-lone-star-virustick-borne-severe-fever-with-thrombocytopenia-syndrome-virustick-borne-huaiyangshan-virustick-borne-guertu-virustick-borne-Alongshan-virustick-borne-wenzhou-virustick-borne-wuhan-virustick-borne-yongjia-virustick-borne-chiba-virustick-borne-natori-virustick-borne-odawara-virustick-borne-kowloon-virustick-borne-tamanavirus

• Trichonephila clavata

  Jorō spider, Joro Spider, Parachute spider

  Trichonephila clavata, commonly known as the Jorō spider, is a large orb-weaving spider native to East Asia that has become established as an invasive species in the southeastern United States since approximately 2010. First confirmed in Georgia in 2014, it has expanded rapidly across multiple states through a combination of ballooning dispersal and human-mediated transport. The species is notable for its substantial size, striking coloration, and extensive golden webs, but poses minimal risk to humans due to small fangs and docile behavior. Its physiological adaptations—including higher metabolic rate, faster heart rate, and greater cold tolerance than its congener Trichonephila clavipes—suggest potential for continued northward range expansion.

  invasive-speciesorb-weaverballooningurban-wildlifebiological-controlcold-tolerancestartle-responsecannibalismkleptoparasitismJapanese-folkloreNephilidaeTrichonephilaspider-silkrange-expansionphysiological-adaptationurban-ecologyagricultural-pest-predatoriNaturalistGeorgiaMarylandEast-AsiaUnited-Statesspider-bitevenomdocilenon-aggressiveweb-architectureegg-sacoverwinteringsexual-dimorphismmetabolic-rateheart-ratefreeze-toleranceroad-ecologyprey-capturecompetitionnative-spider-impactspublic-perceptionmedia-sensationalismmanagementpesticide-alternativesbroom-removalornamentalstructural-pestornithologycardinalweb-strengthtrophic-interactionsDNA-metabarcodingdiet-analysisspecies-distribution-modelphysiological-entomologybehavioral-ecologyarachnologyballooning-dispersalhuman-mediated-dispersalhitchhikercargo-stowawayport-of-entryclimate-changewarmingnorthward-expansionDMVHoward-CountyElkridgeIlchesterWest-Elkridgebeech-forestsecondary-forestsuburbanroadsidetelephone-polestreetlampforest-edgedisturbance-toleranceprey-detectionvibrational-noiseauditory-disturbancemotionlessfreezingshynessHannibal-Lecterstink-buglanternflyevolutionary-historyancient-associationecosystem-servicebiocontrolnatural-enemyinvasive-pestbrown-marmorated-stink-bugspotted-lanternflyHalyomorpha-halysLycorma-delicatulacompetitive-exclusiondominant-orb-weaverabundantpopulation-growthrapid-spreadrange-span120,000-km2physiological-plasticitymetabolismtwice-as-high77%-higher-heart-rate74%-freeze-survivaldevelopment-rateseasonal-windowlife-cycle-completionthermal-limitationcongener-comparisonnatural-dispersalhuman-transportiNaturalist-recordcitizen-scienceconfirmed-sightingbreeding-populationmale-and-femalegravid-femaleegg-casestowawaycargo-containerport-introductionlocal-portvehicle-transporthitchhikingaccelerated-arrivalseveral-yearsnatural-meansprognosticationpredictionmodelsuitable-habitatabundant-suitable-habitateastern-North-Americawestern-North-Americapotential-rangeinvasive-rangepotential-impactenvironmental-impactecological-impactecosystem-impactmonitorcautiousevidence-basedreasonable-journalismsensationalismexaggerationuncritical-acceptanceharmlesscarefully-monitorednext-stepsresearch-opportunitycall-for-researchknowledge-gaplittle-knownnovel-ecosystemspreadingrapidly-expandinginvasive-predatorinvasive-orb-weavernon-nativeintroducedestablishedcolonizedthrivingdoing-finehappymerely-20-minutes-awayroad-tripvisitaccess-spreadbeech-leaf-diseaseBLDstate-forestremarkable-colonyamongst-beech-treesconfirmed-prognosticationDavis-and-Frickescape-relative-warmthexpand-range-northwardeastern-seaboardmerely-20-minutes-away-from-homeresist-opportunityamazing-predatorstales-of-arrivalprevious-episodes20222024reduce-angstlarge-non-native-spiderestablishing-in-DMVfacts-presentedpast-episodevenomous-terrible-painfulNahexpert-Rick-Hoebekerisks-smallpuny-fangsunlikely-pierce-skinvideocompletely-non-aggressiveeastern-Howard-Countyhaphazard-webslittered-with-remainsformer-victimsleavesshed-exoskeletonsmuch-larger-femaledwarfs-matepositioned-just-abovestrand-of-silkspinneretsunderside-of-abdomennear-red-moundrelative-to-handlarge-and-docilewait-and-seewhat-Jorō-means-to-ecosystemshelp-other-spidersput-a-beat-down-on-invasive-pestsstink-bugsspotted-lanternfliespassive-huntersenormous-webslarger-than-a-meter-in-diametercapture-prey-snared-in-silkarachnophobes-scaryarachnophiles-beautifulimportant-ecosystem-servicescrop-pestsnative-rangeAsiahaving-an-old-friend-over-for-dinnerreunitefirst-timelarge-spidersjuicy-prey-itemsfeathered-and-non-feathered-reptilesposes-no-known-threatdifficult-to-predict-impactnon-native-speciesexperts-suggestbeyond-scary-miengive-indigenous-large-orb-weavers-run-for-their-moneyblack-and-yellow-garden-spidermarbled-orb-weaverspotted-orb-weaverother-parts-of-worldmost-abundant-and-dominant-orb-weaverwhat-will-it-meanonly-time-will-tellfinal-tidbitJorō-shapeshifterJorō-gumobeautiful-womanseduces-menbinds-with-silkdevoursbad-dateacknowledgementsRick-Hoebekeidentifying-JorōinsightsDavid-CoyleBob-Bellingergreat-imagesknowledgestudiesVeni-vidi-viciNelsen-et-al.Physiological-evaluationNephila-clavata-newly-recordedHoebeke-et-al.Life-Cycle-Habitat-and-VariationMooreStartle-responsesDavis-and-AneraoHow-Urban-Tolerant-Are-TheyDavis-et-al.inspirationdetailsstarsDave-ClementMiri-TalabacMaddie-Potterhooked-up-with-colonydifferentiateresources.ipmcenters.orglinkJorō-guruDavid-Coyle's-takeYouTubecalm-your-fearsJournal-of-Economic-EntomologyJournal-of-Medical-Entomologychemical-management-strategieseffective-management-methodsspider-control-productssafe-and-effectiveunlikely-to-bitegenerally-harmlessbig-spiderscolorful-spidersinvasive-spidersin-your-face-spiderswebs-on-homeslandscape-plantsstructuresmiddle-of-where-people-live-work-and-playnorthern-Georgia-2014presumed-arrived-few-years-prior15-yearsincreased-dramaticallymuch-of-Southeastdisjunct-populationsnortheastern-U.S.firestorm-of-interestrapidly-expanding-rangesoutheastern-statesmany-countiesCarolinasTennesseeOklahomamore-than-a-centuryparts-of-Floridararely-Pennsylvaniatropicsthermal-limitationshigher-metabolismfaster-heart-ratebetter-ability-to-tolerate-freezingcomplete-life-cycle-rapidlychilly-temperaturessuite-of-adaptationsexpand-northwardwarm-loving-cousinfindings-support-potentialmuch-yet-to-be-learnedsuccessfully-survive-northern-wintersnation-and-world-warmsouthern-species-expandhigher-latitudes-and-altitudesrapidly-range-expansion-will-occurtypical-mode-of-dispersalaerial-dispersalballooning-on-strands-of-silkCharlotte's-babiesCharlotte's-Webspectacular-monikerparachute-spiderrain-down-from-airplaneslong-distance-transithuman-assistancegood-hitchhikersinseminated-gravid-femaleegg-case-stowawayarrival-in-DMVseveral-years-by-natural-meansnew-introduction-at-local-porthuman-assist-in-vehiclesaccelerate-arrivalterrible-and-painfulsmall-fangsvisited-cousingolden-silk-spiderCosta-Ricaexceeding-meter-diameterindigenous-large-orb-weaversrun-for-their-moneyminimal-direct-impactfirestormAndrew-DavisBenjamin-FrickUniversity-of-Georgiapast-weeknative-to-eastern-AsiaJapanChinaKoreaTaiwanknown-in-Georgia-since-2013spreading-rapidlyjoined-its-cousinTrichonephila-clavipesestablished-over-160-yearsbetter-tolerate-freezingsurvive-northern-wintersrapidly-range-expansiontypical-modelong-distancenew-introductionhuman-assistarachnophobesarachnophilesbrown-marmorated-stink-bugsdifficult-to-predictscary-mienindigenous-orb-weaversMary-NouriSarah-MorganWAMUrelatively-harmlessspider-seasonfallleaves-changing-colorcool-crisp-autumn-airpumpkin-spiceextension-entomologistSoutheastnot-just-spiders20-feet-widehomespeople-live-work-and-play15-years-or-soinflux-of-callspublicentomology-extension-programsmedia-frenzytwo-questionsget-rid-of-Jorō-spidersare-Jorō-spiders-dangerousClemson-UniversitySouthern-Adventist-UniversityWashington-CollegeUnion-Collegetwo-new-studiestest-methods-for-eliminatinghow-likely-to-biteharmscoured-internetrecommended-treatment-optionsspider-specific-management-recommendationsmedically-relevantwidow-spidersbrown-reclusesdo-not-enter-homesrelatively-newrecommended-productslabelled-spider-control-productstested-productsseveral-products-eliminatenot-legal-pesticidesmachine-lubricantnot-recommendedcommercially-availablestrongly-encouragelabelled-productseasiest-and-cheapestwithout-chemicalsbroom-or-stickwalked-into-spider-webrode-biketook-spider-to-facenot-great-feelingwonder-if-biteygive-bitten-superpowersunique-studiesevaluated-behaviorsreact-to-peoplerather-drop-off-webhang-around-and-bitereally-provokevolunteered-to-get-bittenyes-you-read-correctlyakin-to-mosquito-bitelittle-rednesslittle-swellinglargely-gone-next-dayno-acquired-superpowerstake-home-messagetwofoldsimplest-and-most-effectivewithout-pesticidesfeel-you-need-pesticidesproduct-labeled-for-spider-controltry-not-to-freak-outdoesn't-want-to-be-on-youodds-of-getting-bitten-extremely-lowenjoy-glorious-fall-weatherdrink-pumpkin-spiced-beveragewatch-where-you-walkKeep-calm-and-carry-onbites-unlikelycause-minimal-discomfortassociate-professorDepartment-of-Forestry-and-Environmental-Conservationsocialsemailextensionfeaturedpest-managementEntomology-Todaylatest-postsemail-subscriptionMarch-2022wonderedmake-its-way-to-DMVSeptember-2022two-observationsthree-years40-sightingshappy-and-doing-just-fineseveral-Howard-County-locationsweek-or-so-agoteam-of-scientistsUniversitymissionkilling-ultra-valuable-beech-treesthriving-amongst-beech-treesvisit-amazing-predatorsmature-femalesboth-Trichonephila-speciesthree-locally-common-orb-weaving-speciestime-spent-immobilemild-disturbancebrief-puff-of-airair-puff-response-datafive-other-North-American-speciespublished-literature453-observationsfreezing-behavior10-spider-speciesremained-immobile-under-a-minuteboth-Trichonephila-spidersover-an-hourunprecedentedshyest-ever-documentedtolerate-urban-environmentsremaining-motionlessfleeingnonsexual-cannibalismmating-activityfemales-cannibalize-malesfood-availabilityterritorial-aggressionSoutheastern-United-Statesexpanding-rangeshybehavioral-reactionsdescriptive-observationsphoto-documentationanecdotal-observationscontrolled-pairingslabnatural-websfieldone-female-biting-and-killing-anothershort-fightsimilar-sizecontainer25-trialsfights-ensued-40%different-sizes27-trialsfights-happened-18%larger-females-not-always-aggressor52-lab-trialssix-bouts9%direct-killingfield-trialsempty-web14-trialsone-fight7%aggressor-killing-and-wrappingsilkaway-from-any-webnot-necessarily-territorialityintraspecific-aggressionprovoked-or-stresseddeserves-more-studynewly-invasivebiologyphysiologynew-rangeclosely-related-speciessuccessfully-established160-yearsinvestigationcompareonline-recordsiNaturalist.orgseasonal-distributionstimingfemalesphysiological-traitsenvironmental-tolerancemetabolic-ratescardiac-functioningcold-exposuresurvivalbrief-below-freezing-temperatureshorter-seasoncomplete-lifecyclenarrow-periodsuitable-weatherinherently-higher-metabolismlow-temperaturesurvive-better74%-compared-to-50%brief-freezecolder-climatic-regionsoutheastern-USAuseful-informationplanningurban-landscapesbusy-roadsfrequent-disturbancesauditory-and-vibrationalphysiological-or-behavioral-changesnortheast-Georgiavarying-levels-of-road-trafficprey-capture-behaviorsimulated-preytuning-fork128-hz-frequencytouched-to-webattacked357-total-trials20-different-roadsattacked-59%high-variabilitysome-roadsides-over-80%others-less-than-30%small-but-significant-negative-correlationdaily-road-trafficspider-attack-ratesmoderate--to-heavy-traffic-roadsslightly-less-likely-to-attack51%-vs-65%able-to-live-near-roadscost-in-terms-of-prey-capturedid-not-weigh-lesscompensate-for-disturbanceaccumulating-evidencehuman-dominated-landscapesaid-spreadintroduced-rangeEriostethus-rufuspolysphinctine-ectoparasitoidaraneid-spidersNeoscona-spp.endemic-to-JapanYamagata-Prefecture38º46'-Nnorthern-Japannorthernmost-recordgenusidentification-confirmedmorphologyDNA-barcodingcocoonlarge-modified-webuniqueinverted-triangleover-50-cmhanging-from-ill-defined-partno-organized-structurehost-spidercircumstantial-evidencesstructure-of-modified-webpre-existing-web-partly-reusedorb-web-completely-removedsustaining-threads-newly-mooredparasitises-host-spiderstwo-subfamiliesunusual-for-groupinvasion-potentialknown-invasive-rangereviewdispersal-abilitiespotential-impactsmanagement-strategiesOctober-2022at-least-120,000-km2South-CarolinaNorth-CarolinaAlabamaWest-Virginiapattern-of-spreadnatural-dispersal-mechanismshuman-mediated-transportlarge-bodied-orb-weaversflying-insectssmall-animalsthirteen-co-occurring-spider-speciesmonitored-for-competitionresourcesweb-building-sitesnatural-and-urban-habitatsmanagement-options-limitedadvise-journalists-and-expertsagainst-exaggerating-potential-environmental-impactecologically-harmlessrapid-spread-carefully-monitoredcautious-evidence-based-approachdetermining-next-stepsthree-material-typesweb-captureintroduced-speciesalter-established-trophic-interactionsmolecular-analysisresolve-changescommunity-structureforaging-linkseast-coastfecal-samplesprey-remains-from-websdissected-gutscompare-diet-compositionarthropod-targeted-COI-primersIllumina-MiSeqhighest-diversityhighest-richnesshighest-proportion-prey-readsrecovery-of-prey-readsdissected-gut-contentlowoverwhelmed-by-Jorō-spider-DNAhigh-proportions-Jorō-spider-readshigher-prey-diversity-and-richnessdetected-prey-DNAseveral-days-after-captureinitial-gut-retention-time-estimatesfirst-glimpsecomplexity-of-trophic-associationsintroduced-web-building-spiderviable-materialsource-of-prey-DNAestimates-of-biodiversityJammu-and-Kashmirnorthernmost-shiftingdocument-presencebehaviorhabitat-adaptationcolder-temperature-environmentGoha-TehsilDoda-district1556.74-metersroad-bridgenearby-vegetationbrightly-colored-abdomensblackgreenyellowredlong-legsyellow-and-black-bandingspredominantly-seensmallerfewerunique-behaviorsremaining-motionless-after-disturbances5-10-meters-of-roadsadaptation-to-human-settlementsfield-observationsimage-documentationexpert-confirmationthrive-in-colder-conditions15-20-degrees-CelsiusOctober-Novemberno-spider-foundhigher-elevationssignificantly-lower-temperaturesadaptation-limited-to-milder-cold-regionsnew-insightadaptationneed-for-further-researchnon-native-habitatseggsoverwinterharsh-weather-conditionsno-protectioncompact-silk-casemilky-coatingchorionic-microspheresCMfine-structural-characteristicsecological-importanceuneven-size-distributionouter-eggmassevenly-coveredsingle-layerdiameter-2.3-µmshort-papillary-projectionssegmental-adhesionmucous-componentsinsoluble-in-waterpartially-soluble-in-absolute-ethanolspherical-structureHFIPstrong-organic-solventnot-derived-from-vitellogenic-or-choriogenetic-processesCM-adhesive-coatingsovipositional-processequivalent-to-cocoon-silkprotective-functionssilken-eggcaseNorthern-CardinalCardinalis-cardinalisperchessteals-foodsoutheast-of-the-United-Statesquestionsnative-fauna-affectedconsuming-prey-itemsexamplenative-species-deriving-benefitmanner-of-kleptoparasitismperched-directly-on-websupported-its-weight42-48-gfirst-documented-casespider-web-supporting-perching-birdmeasurementsforce-gaugetypical-webssupport-masses-up-to-70-gsmall-but-growing-body-of-knowledgenon-native-spiderspecies-distribution-modelsecological-niche-modelingpredict-areasestablish-if-introducedthree-distinct-genetic-lineagesmitochondrial-COIgenome-wide-SNPsKorean-Peninsula