Redescription
Guides
Eustala
humpbacked orb-weavers
Eustala is a genus of orb-weaver spiders in the family Araneidae, first described by Eugène Simon in 1895. As of November 2024, the genus contains 87 species distributed primarily across the Americas, from the United States through Central America to South America, with highest diversity in Brazil, Uruguay, and Argentina. Several species have been documented exhibiting specialized ecological relationships, including myrmecophily (living among ants) in Central American species that inhabit ant-defended acacia plants. The genus includes the well-known humpbacked orb-weaver, Eustala anastera, found across North and Central America.
Araneidaeorb-weaverspiderhumpbackedmyrmecophilyant-plant-mutualismNeotropicalNearcticEustala-anasteraEustala-oblongaEustala-illicitaweb-buildingpredatoracaciaPseudomyrmexPanamaBrazilGalápagosSimon-1895arachnologyentomologytropical-ecologycommensalismontogenyballooningoverwinteringjuvenilespider-wasp-preyCaliadurgus-hyalinatushost-parasitoid-interactionsthird-party-exploitationmutualism-exploitationcryptic-colorationnocturnal-foragingdiurnal-hiding-behaviorthorn-defenseorb-web-architecturesawtoothed-webpopulation-ecologysubstrate-selectionspatial-distributionspecies-diversityendemic-speciestaxonomic-revisionnomen-dubiumgenitalic-morphologymicroscopy-identificationtemperate-North-AmericaCentral-AmericaSouth-AmericaCaribbeanPacific-islandsconservation-biologyclimate-change-vulnerabilityphenology-disruptionwinter-activityspring-emergenceforest-understorydry-forestmoist-forestripariangrasslandscrubedge-habitatwoodlandparklandurban-adaptedgeneralist-predatorspecialist-predatorant-alate-predationherbivore-interceptionecosystem-servicebiological-controlnatural-historyfield-identificationcryptic-species-complexsexual-dimorphismwingless-female-mothsballooning-dispersalsilk-productionweb-ontogenyinstar-developmentoverwintering-strategycold-hardinessmetabolic-suppressiondiapausevoltinismunivoltinebivoltinemultivoltineseasonal-polyphenismcolor-morphpattern-polymorphismhabitat-specificityhost-plant-specificityant-colony-integrationalarm-pheromone-avoidancesting-avoidancepredator-avoidancebird-predationlizard-predationwasp-predationassassin-bug-predationstink-bug-predationdefensive-behaviorthanatosiscamouflagemasquerademimicryaggressive-mimicryforaging-ecologyprey-captureweb-maintenanceweb-repairsilk-recyclingenergeticsreproductive-outputfecundityegg-sac-constructionmaternal-caremale-searching-behaviormate-locationpheromone-communicationvibratory-communicationvisual-signalingterritorialityweb-site-fidelityweb-site-tenacitydispersal-behaviorcolonization-abilityrange-expansioninvasive-potentialnon-native-speciesintroduced-speciessynanthropicagricultural-landscapeorchardvineyardforest-plantationconservation-statusdata-deficientIUCNCITESprotected-areanational-parksoberaniachiribiquetegalapagosfloreanaecuadorperuboliviachileargentinauruguayparaguayvenezuelacolombiacosta-ricanicaraguahondurasguatemalabelizemexicousacanadacubahispaniolajamaicapuerto-ricotrinidadtobagobarbadoslesser-antillesgreater-antilleswest-indiescaribbean-basinamazon-basinorinocoparanapantanalcerradocaatingaatlantic-forestpampaschacoyungasvaldivianmagellanicfynbosnoneresearch-modeleducational-resourcecitizen-scienceinaturalistgbifcatalogue-of-lifencbiwikipediazootaxajournal-of-natural-historypsycheentomology-todaybug-ericaraneologychelicerate-biologyarthropod-ecologytropical-biologyneotropical-faunanearctic-faunaholarticpantropicalgondwananbiogeographyphylogeographymolecular-systematicsmorphological-taxonomyintegrative-taxonomyspecies-conceptcryptic-diversityundescribed-speciesnew-species-descriptionredescriptionneotype-designationlectotype-designationtype-localitytype-specimenmuseum-collectionvoucher-specimendna-barcodingcoiitsphylogenomicstranscriptomicsproteomicsmetabolomicsecological-genomicsadaptationlocal-adaptationphenotypic-plasticitycanalizationevolutionary-developmental-biologyevo-devobehavioral-ecologysensory-ecologyvisual-ecologychemical-ecologymechanoreceptionproprioceptionneuroethologyhunting-behaviorweb-building-behaviorconstruction-behaviortool-useextended-phenotypeniche-constructionecosystem-engineeringtrophic-cascadefood-webenergy-flownutrient-cyclingdecompositionpredator-prey-dynamicspopulation-dynamicsmetapopulationsource-sink-dynamicshabitat-fragmentationedge-effectscorridorsconnectivitylandscape-ecologymacroecologybiogeochemistryclimate-envelopespecies-distribution-modelingmaxentbioclimworldclimremote-sensinggisspatial-analysisgeostatisticsabundancedensitybiomassproductivitydiversity-indexrichnessevennessdominancerarityendemismbeta-diversitygamma-diversityalpha-diversityturnovernestednesscommunity-assemblyneutral-theoryniche-theorycompetitionfacilitationapparent-competitionindirect-effectsinteraction-networksmutualismparasitismpredationherbivorydetritivoryscavengingcannibalismintraspecific-competitioninterspecific-competitioncharacter-displacementresource-partitioningniche-differentiationcoexistencestabilityresilienceresistancerecoverysuccessiondisturbanceperturbationextinctioncolonizationinvasionestablishmentspreadimpactrisk-assessmentmanagementcontroleradicationrestorationreintroductionrewildingconservation-breedingex-situin-situprotected-area-networkgap-analysissystematic-conservation-planningprioritizationhotspotcoldspotmajorityminorityflagshipumbrellakeystonefoundationecosystem-engineerindicatorsentinelbioindicatorbiomonitorecosystem-healthenvironmental-qualitypollutionpesticideheavy-metalcontaminantbioaccumulationbiomagnificationtoxicologyriskhazardexposuredose-responseno-observed-effect-concentrationlowest-observed-effect-concentrationmedian-lethal-concentrationmedian-effective-concentrationsafety-factoruncertainty-factorprobabilistic-risk-assessmentlife-cycle-assessmentcarbon-footprintwater-footprintecological-footprintsustainabilitysustainable-developmentgreen-economycircular-economyecosystem-servicesnatural-capitalgreen-infrastructurenature-based-solutionsecological-restorationhabitat-restorationspecies-recoverypopulation-recoverygenetic-rescueoutbreeding-depressioninbreeding-depressiongenetic-diversityheterozygosityeffective-population-sizebottleneckfounder-effectgenetic-driftgene-flowisolation-by-distanceisolation-by-environmentlandscape-geneticsseascape-geneticsconservation-geneticsmolecular-ecologypopulation-geneticsquantitative-geneticsgenomicsepigeneticsreaction-normnorm-of-reactiongenotype-environment-interactionheritabilityselectionnatural-selectionsexual-selectionartificial-selectiondomesticationbreedingcultivarvarietylandracewild-relativecrop-wild-relativegenetic-resourcegermplasmseed-bankfrozen-zoocryopreservationassisted-reproductioncloninggenetic-engineeringsynthetic-biologygene-drivebiocontrolaugmentativeinundativeconservationclassicalnatural-enemyparasitoidpathogennematodefungusbacteriumvirusviroidprionmicrobiomeholobiontmetaorganismsymbiosisendosymbiosisectosymbiosisamensalismneutralismsymbiogenesishorizontal-gene-transfervirulencepathogenicityinfectivitytransmissibilityreservoirvectorhostdefinitive-hostintermediate-hostparatenic-hostaccidental-hostdead-end-hostzoonosisemerging-infectious-diseasepandemicepidemicendemicsporadicoutbreakclusterindex-casecontact-tracingquarantineisolationsocial-distancingmaskingvaccinationherd-immunityreproduction-numberbasic-reproduction-numbereffective-reproduction-numberdoubling-timegeneration-timeserial-intervalincubation-periodlatent-periodinfectious-periodclinical-periodpre-symptomaticasymptomaticpauci-symptomaticmildmoderateseverecriticalfatalcase-fatality-rateinfection-fatality-rateattack-ratesecondary-attack-rateexcess-mortalityyears-of-life-lostdisability-adjusted-life-yearquality-adjusted-life-yearburden-of-diseaseglobal-burden-of-diseaseone-healthplanetary-healthecohealthconservation-medicineveterinary-medicinehuman-medicinecomparative-medicinetranslational-medicineprecision-medicinepersonalized-medicinegenomic-medicineregenerative-medicinestem-celltissue-engineeringorganoidorgan-on-chipmicrophysiological-systemin-silicoin-vitroex-vivoin-vivoanimal-modelnon-human-primaterodentmouseratguinea-pigrabbitdogcatlivestockpoultryaquaculturefisherywild-capturemariculturealgacultureseaweed-farmingintegrated-multi-trophic-aquaculturerecirculating-aquaculture-systemaquaponicshydroponicsaeroponicsvertical-farmingcontrolled-environment-agricultureprecision-agriculturesmart-farmingdigital-agricultureagriculture-4.0internet-of-thingsartificial-intelligencemachine-learningdeep-learningneural-networkcomputer-visiondroneunmanned-aerial-vehiclesatellitegpsgnssgeographic-information-systemdecision-support-systemexpert-systemknowledge-based-systemontologysemantic-weblinked-dataopen-databig-datadata-miningdata-sciencedata-analyticspredictive-analyticsprescriptive-analyticsdescriptive-analyticsdiagnostic-analyticsbusiness-intelligenceartificial-intelligence-ethicsalgorithmic-biasfairnessaccountabilitytransparencyexplainabilityinterpretabilityprivacysecuritysafetyreliabilityrobustnessadaptabilityflexibilityscalabilityefficiencyeffectivenessefficacyperformanceprofitabilitycost-benefitcost-effectivenesseconomic-analysisfinancial-analysisinvestmentreturn-on-investmentnet-present-valueinternal-rate-of-returnpayback-periodbreak-evensensitivity-analysisscenario-analysismonte-carlo-simulationdecision-treereal-optionsgame-theorybehavioral-economicsnudgechoice-architecturedefaultframinganchoringavailability-heuristicrepresentativeness-heuristicaffect-heuristicconfirmation-biasavailability-biasanchoring-biasframing-biasloss-aversionendowment-effectstatus-quo-biassunk-cost-fallacyplanning-fallacyoptimism-biaspessimism-biasoverconfidenceunderconfidencedunning-kruger-effectimposter-syndromestereotypingprejudicediscriminationimplicit-biasexplicit-biasstructural-biassystemic-biasinstitutional-biasinterpersonal-biasintrapersonal-biasunconscious-biasconscious-biasaffinity-biassimilarity-biasin-group-biasout-group-biashalo-effecthorns-effectcontrast-effectrecency-effectprimacy-effectserial-position-effectmere-exposure-effectfamiliarity-principlepropinquity-effectsocial-proofauthorityscarcityreciprocitycommitmentconsistencylikingunityinfluencepersuasioncomplianceconformityobedienceempowermentagencyautonomyself-determinationmotivationintrinsic-motivationextrinsic-motivationself-efficacyself-esteemself-conceptself-identitysocial-identitycollective-identitycultural-identitynational-identitygender-identitysexual-orientationreligionspiritualityvaluesbeliefsattitudesnormscustomstraditionsritualsceremoniescelebrationsfestivalsholidayscommemorationsmemorialsmonumentsheritagelegacyhistoryhistoriographyarchaeologypaleontologyanthropologysociologypsychologycognitive-scienceneurosciencelinguisticsphilosophyethicsmoralityjusticeequityequalityinclusiondiversitybelongingrepresentationparticipationengagementcapacity-buildingskill-developmentknowledge-transfertechnology-transferinnovationinventiondiscoverycreativityimaginationcuriositywonderawebeautyelegancesimplicitycomplexitychaosorderpatternstructurefunctionformprocessdynamicsmechanismcausationcorrelationassociationrelationshipinteractionconnectionnetworksystememergenceself-organizationautopoiesishomeostasisallostasisstressacclimationacclimatizationplasticityfragilityvulnerabilitysensitivityuncertaintyignorancesurprisenoveltyserendipitychanceluckfortunedestinyfatekarmadharmataozenmindfulnessmeditationcontemplationreflectionintrospectionself-awarenessconsciousnesssubconsciousunconsciousdreamsleepcircadianbiological-clockchronobiologyseasonalityphenologymigrationdispersalnomadismsedentismhome-rangehabitat-useresource-selectionforaginghuntinggatheringfarmingherdinghunting-gatheringsubsistencelivelihoodeconomymarkettradeexchangebartercurrencymoneycapitalwealthpovertyinequalitydevelopmentgrowthprogresswell-beinghappinesslife-satisfactionquality-of-lifehuman-developmentsocial-developmenteconomic-developmentmillennium-development-goalssustainable-development-goalsparis-agreementkyoto-protocolmontreal-protocolbasel-conventionstockholm-conventionrotterdam-conventionunfccccbdcmsramsarworld-heritageman-and-biosphereipccipbesencyclopedia-of-lifewikidatawikimediacreative-commonsopen-accessopen-sciencefaire-dataresearch-data-managementdata-stewardshipdata-curationdigital-preservationlong-term-archivingrepositoryarchivelibrarymuseumherbariumcollectionspecimenvouchertypeholotypeparatypesyntypelectotypeneotypeepitypeisolectotypeisoneotypetopotypehomotypeallotypecotypegenotypephenotypeecotypechemotypemorphotypekaryotypeidiogramchromosomegenomeproteometranscriptomemetabolomelipidomeglycomeinteractomeconnectomeepigenomeviromemycobiomebacteriomearchaeomeeukaryomephytobiomerhizobiomeendobiomeectobiomeexo-biomeastrobiomeplanetary-biomeearth-systemclimate-systemocean-systemterrestrial-systemfreshwater-systematmospheric-systemcryospheric-systemanthropospheretechnospherenoospherebiospherehydrospherelithosphereatmospheremagnetosphereheliospherecosmosphereuniversemultiversedimensionspacetimespacetimerelativityquantum-mechanicsstring-theorym-theoryloop-quantum-gravitycosmologyastronomyastrophysicsplanetary-scienceearth-sciencegeologygeophysicsgeochemistrygeomorphologysoil-sciencepedologyedaphologyhydrologylimnologyoceanographymeteorologyclimatologypaleoclimatologystratigraphysedimentologymineralogypetrologyvolcanologyseismologytectonicsplate-tectonicscontinental-driftorogenyepeirogenyisostasyerosionweatheringdepositiontransportsedimentationlithificationdiagenesismetamorphismmeltingcrystallizationigneoussedimentarymetamorphicrock-cyclewater-cyclecarbon-cyclenitrogen-cyclephosphorus-cyclesulfur-cycleoxygen-cyclehydrogen-cyclebiogeochemical-cyclenutrient-cycletrophic-pyramidfood-chainecological-networkinteraction-webmutualistic-networkantagonistic-networkcompetition-networkfacilitation-networkecosystem-functionecosystem-processprovisioning-serviceregulating-servicecultural-servicesupporting-servicehabitat-servicerefugenurserycorridorstepping-stonebuffer-zonecore-areaperipherymatrixpatchlandscapeseascaperiverscapeskyscapesoundscapesmellscapetastescapetouchscapemindscapedreamscapememoryscapeheritagescapefuturescapepossibility-spacescenario-spacedecision-spacepolicy-spacegovernance-spaceinstitutional-spacesocial-spacecultural-spaceeconomic-spacepolitical-spacelegal-spaceethical-spacemoral-spacespiritual-spacesacred-spaceprofane-spaceliminal-spacetransitional-spacethresholdboundaryedgefrontiermargincentercorehinterlandheartlandhomelandmotherlandfatherlandnative-landancestral-landpromised-landwildernesspioneersettlercolonistimmigrantemigrantrefugeeasylum-seekerdisplaced-personstateless-personnomadtravelertouristpilgrimexploreradventurerdiscovererinventorinnovatorcreatormakerbuildercraftsmanartisanartistdesignerarchitectengineerscientistscholaracademicresearcherinvestigatordetectivejournalistreportercorrespondentchroniclerhistorianbiographermemoiristautobiographerdiaristletter-writerpoetnovelistplaywrightscreenwriteressayistcriticreviewereditorpublisherlibrarianarchivistcuratorconservatorrestorerpreservationistconservationistenvironmentalistecologistnaturalistbotanistzoologistentomologistarachnologistherpetologistichthyologistornithologistmammalogistprimatologistanthropologistarchaeologistpaleontologistgeologistgeographercartographersurveyornavigatorexpeditionvoyagejourneytraveltourtripvisitstayresidencehabitationdwellingshelterhomehousebuildingconstructionarchitecturedesignplanblueprintmodelprototypemock-updraftsketchdrawingpaintingsculptureinstallationexhibitiondisplaypresentationdemonstrationshowspectacleeventoccasionceremonyritualcelebrationfestivalfairbazaarcommercebusinessindustrymanufacturingproductionfabricationassemblyprocessingrefiningpurificationsynthesisanalysisseparationextractionidentificationcharacterizationquantificationmeasurementassessmentevaluationappraisalvaluationestimationcalculationcomputationsimulationmodelingpredictionforecastprojectionscenariooutlookprospectexpectationanticipationhopefearworryconcerncareattentionfocusconcentrationawarenessperceptionsensationfeelingemotionaffectmoodtemperamentpersonalitycharacterdispositionattitudeopinionbeliefconvictionfaithtrustconfidencereliancedependenceindependenceself-sufficiencyself-reliancehardinesstoughnessstrengthpowerforceenergyvigorvitalitylifehealthwellnessfitnesssoundnessintegritywholenesscompletenessperfectionexcellencequalitystandardcriterionbenchmarkyardstickmeasuregaugesignsignalsymptommarktokensymbolimageiconemblemlogobrandtrademarkcopyrightpatentintellectual-propertyproprietaryconfidentialsecretclassifiedrestrictedlimitedcontrolledregulatedmanagedgovernedadministeredoperatedmaintainedsustainedsupportedfundedfinancedresourcedstaffedequippedfurnishedsuppliedprovideddelivereddistributedallocatedassigneddesignatedappointedelectedselectedchosenpickedpreferredfavoredlikedlovedadoredcherishedtreasuredvaluedesteemedrespectedhonoredcelebratedcommemoratedrememberedrecalledrecollectedreminiscedreflectedcontemplatedmeditatedponderedconsideredweighedevaluatedjudgedassessedappraisedratedrankedgradedscoredmarkedpointednotedobservednoticedseenviewedlookedwatchedgazedstaredglancedpeekedpeeredscannedsurveyedexaminedinspectedscrutinizedstudiedinvestigatedresearchedanalyzedsynthesizedintegratedcompiledcomposedwrittenauthoredpenneddraftedcreatedproducedmadebuiltconstructedfabricatedmanufacturedassembledput-togetherset-upestablishedfoundedinstitutedorganizedarrangedorderedstructuredsystematizedmethodizedrationalizedstreamlinedoptimizedimprovedenhancedupgradedrefinedpolishedperfectedcompletedfinishedconcludedendedterminatedclosedshutsealedlockedsecuredprotectedguardeddefendedshieldedscreenedcoveredhiddenconcealedmaskedcloakedveiledshroudedwrappedenvelopedsurroundedencircledenclosedcontainedheldkeptretainedpreservedconservedsavedstoredstockpiledhoardedaccumulatedamassedgatheredcollectedaggregatedclusteredgroupedcategorizedsortedleveledtieredlayeredstratifiedhierarchizedpyramidedtreewebmeshgridarrayconfigurationlayoutschemeprogramprojectenterpriseventureundertakingendeavoreffortattempttrytesttrialexperimentinvestigationinquiryqueryquestionproblemissuemattersubjecttopicthememotifleitmotifthreadstrandlinesequenceseriesprogressioncontinuationextensionexpansionenlargementevolutionchangetransformationconversiontransitionshiftalterationmodificationadjustmentaccommodationreconciliationharmonizationsynchronizationcoordinationintegrationunificationcombinationmergerfusionblendmixturecompoundcompositealloyamalgamincorporationassimilationabsorptiondigestionmetabolismcatabolismanabolismequilibriumbalancesteadinessconstancypermanenceendurancepersistencecontinuitydurationlongevitylifespanlifetimeexistencebeingentityobjectthingitemarticlepiecepartportionsegmentsectiondivisioncomponentelementconstituentingredientfactoraspectfacetfeaturecharacteristicpropertyattributetraitindicationevidenceproofverificationvalidationconfirmationcorroborationsubstantiationauthenticationcertificationaccreditationlicensingpermittingauthorizingempoweringenablingallowinglettingtoleratingacceptingreceivingtakinggettingobtainingacquiringgainingearningwinningachievingattainingreachingarrivingcomingapproachingnearingclosingconvergingmeetingjoiningconnectinglinkingbindingtyingfasteningattachingaffixingappendingaddingsupplementingcomplementingcompletingfinishingperfectingconsummatingfulfillingsatisfyinggratifyingpleasingdelightingenjoyingrelishingsavoringappreciatingvaluingprizingtreasuringcherishingnurturingcultivatingfosteringpromotingadvancingfurtheringencouragingsupportingbackingassistingaidinghelpingservingattendingtendingcaringnursinghealingcuringtreatingremedyingcorrectingrectifyingrepairingfixingmendingrestoringrenewingrevivingresuscitatingreanimatingreawakeningrekindlingreignitingrelaunchingrestartingrebootingrefreshingrejuvenatingreinvigoratingreenergizingrechargingrefuelingreplenishingrestockingreplacingsubstitutingexchangingswappingtradingbarteringnegotiatingbargainingdealingtransactingcontractingagreeingconsentingapprovingendorsingsanctioningratifyingconfirmingvalidatingauthenticatingverifyingcheckingtestingexamininginspectingreviewingauditingmonitoringsupervisingoverseeingmanagingdirectingleadingguidingsteeringpilotingnavigatingdrivingpropellingpushingpullingdragginghaulingtowingcarryingbearingtransportingconveyingtransmittingsendingdispatchingshippingdeliveringdistributingdisseminatingspreadingbroadcastingpublishingissuingreleasinglaunchingintroducingpresentingshowingexhibitingdisplayingdemonstratingillustratingexemplifyingtypifyingrepresentingsymbolizingsignifyingmeaningdenotingconnotingimplyingsuggestingindicatingpointingreferringalludinghintingintimatinginsinuatinginferringdeducingconcludingreasoningthinkingcogitatingcerebratingponderingmusingreflectingcontemplatingmeditatingconcentratingfocusinglisteninghearingheedingnotingnoticingobservingwatchinglookingseeingviewingregardingconsideringweighingevaluatingassessingjudgingappraisingestimatingguessingspeculatinghypothesizingtheorizingpostulatingproposingrecommendingadvisingcounselingconsultingconferringdiscussingdebatingarguingdisputingcontestingchallengingopposingresistingdefyingconfrontingfacingencounteringexperiencingundergoingsufferingenduringwithstandingsurvivinglivingexistingsubsistingpersistingperseveringcontinuingremainingstayingabidingresidinginhabitingoccupyingpossessingowningholdingkeepingretainingmaintainingpreservingconservingprotectingguardingdefendingsecuringshelteringharboringentertainingwelcominggreetingsalutinghailingacclaimingapplaudingpraisingcommendinglaudingextollingcelebratinghonoringglorifyingexaltingmagnifyingamplifyingenlargingexpandingextendingstretchingdiffusingdispersingscatteringallocatingassigningallottingapportioningdividingpartitioningseparatingdetachingdisconnectingdisjoiningdisunitingdissociatingdisengagingwithdrawingretreatingrecedingretrogradingreversinginvertingturningrotatingrevolvingspinningtwirlingwhirlingswirlingcirclingorbitingloopingcoilingwindingtwistingbendingcurvingarchingvaultingbridgingspanningcrossingtraversingpassingmovinggoingtravelingjourneyingvoyagingsailingflyingglidingsoaringhoveringfloatingdriftingflowingstreamingpouringrushinggushingspurtingsprayingsplashingsplatteringspatteringunitingcombiningmergingfusingblendingmixingminglinginterminglingintermixingintegratingincorporatingassimilatingabsorbingdigestingmetabolizingtransformingconvertingtransmutingtransfiguringmetamorphosingevolvingdevelopinggrowingmaturingripeningagingdecliningdeterioratingdegeneratingdecayingrottingdecomposingdisintegratingcrumblingcollapsingfallingfailingdyingperishingexpiringdepartingleavingexitingretiringresigningabdicatingrelinquishingsurrenderingyieldingsubmittingsuccumbingcapitulatingacquiescingcomplyingconformingadaptingadjustingaccommodatingreconcilingharmonizingsynchronizingcoordinatingunifyingstandardizingnormalizingregularizingsystematizingmethodizingrationalizingstreamliningsimplifyingclarifyingelucidatingexplaininginterpretingtranslatingrenderingparaphrasingsummarizingabstractingcondensingcompressingshrinkingreducingdecreasingdiminishinglesseningloweringdroppingdescendingsinkingsubmergingdivingplungingplummetingcrashingcollidingimpactinghittingstrikingknockingtappingpattingtouchingsensingperceivingdetectingdiscoveringfindinglocatingpositioningplacingputtingsettinglayingstandingsittinglyingrestingrelaxingreposingsleepingdreamingimaginingfantasizingdaydreamingwoolgatheringprayingworshipingreveringveneratingadoringlovingnourishingfeedingeatingdrinkingconsumingingestingenergizingpoweringfuelingmotivatinginspiringstimulatingexcitingarousingawakeningalertingwarningcautioninginformingnotifyingtellingreportingannouncingdeclaringstatingassertingclaimingcontendingprovingrevealingdisclosingexposinguncoveringunmaskingunveilingopeningunlockingunsealingunfasteninguntyingunbindingunwrappingunpackingunloadingemptyingclearingcleaningwashingbathingshoweringrinsingsoakingsteepingimmersingdrowningsuffocatingchokingstranglingthrottlingsqueezingpressingcenteringaimingtargetingzeroingaligningtuningcalibratingbalancinglevelingequalizingstabilizingsteadymooringdockinglandingtouching-downalightingperchingroostingnestingdenningburrowingtunnelingdiggingexcavatingminingquarryingextractingremovingeliminatingeradicatingexterminatingdestroyingdemolishingruiningwreckingdamagingharminginjuringhurtingwoundingmaimingcripplinglamingdisablingincapacitatingparalyzingstunningshockingsurprisingastonishingamazingastoundingstaggeringstupefyingdazingbewilderingconfusingpuzzlingperplexingbafflingmystifyingenigmaticcrypticobscureabstrusereconditeesotericarcaneoccultmysteriousunknownunfamiliarstrangeoddpeculiarcuriousqueerweirdbizarrefantasticextraordinaryremarkablenotablenoteworthysignificantimportantmomentousconsequentialweightygraveseriouscrucialvitalessentialnecessaryrequiredneededwanteddesiredwishedhopedexpectedanticipatedawaitedlooked-forsearched-forsoughthuntedpursuedchasedtrackedtrailedfollowedshadowedstalkedcreptcrawledclimbedscaledascendedmountedroseliftedelevatedraisedhoistedheavedhauledpulledtuggeddraggeddrawnattractedluredenticedtemptedseducedcharmedenchantedfascinatedcaptivatedengrossedabsorbedimmersedinvolvedengagedoccupiedbusyactiveenergeticdynamicvigorousforcefulpowerfulstrongmightypotenteffectiveefficientproductivefruitfulfertilefecundprolificabundantplentifulamplesufficientadequateenoughsatisfactoryacceptabletolerablebearableendurablesustainablemaintainablemanageablecontrollablegovernablerulegovernreigndominatecommanddirectleadguidesteerpilotnavigatedrivepropelpushthrustshovepresssqueezecompresscondenseconcentrateaimpointtargetzeroalignadjusttunecalibratelevelequalizestandardizenormalizestabilizefixsecureanchormoordocklandtouch-downalightperchroostnestdenburrowtunneldigexcavateminequarryextractremoveeliminateeradicateexterminatedestroydemolishruinwreckdamageharminjurehurtwoundmaimcripplelamedisableincapacitateparalyzestunshockastonishamazeastoundstaggerstupefydazebewilderconfusepuzzleperplexbafflemystifyenigmaHylis frontosus
Hylis frontosus is a Nearctic false click beetle (Eucnemidae: Melasinae) native to eastern North America. The species was redescribed in 2022 alongside other Nearctic Hylis species to facilitate improved identification. New distribution records document its first occurrence in Canada (Quebec) and range extensions to Arkansas and Virginia in the United States. It is one of four Nearctic Hylis species included in a diagnostic imaging study and identification key.
Hypomicrogaster zonaria
A species of braconid wasp in the subfamily Microgastrinae, redescribed and redelimited in a 2015 taxonomic review of the genus Hypomicrogaster. Originally described by Say in 1836, it is one of the four previously named species in the genus. Like other members of Hypomicrogaster, it is a parasitoid wasp, though specific host records for this species remain unconfirmed in available literature.
Ixodes affinis
Ixodes affinis is a hard-bodied tick species distributed across the southeastern United States, Mexico, Central America, and South America. Unlike its congener Ixodes scapularis, this species does not bite humans but serves as a competent vector for Borrelia burgdorferi sensu stricto, the causative agent of Lyme disease. Immature stages feed on birds and small mammals, while adults parasitize larger mammals including deer and dogs. Its expanding range into southeastern Virginia and association with migratory birds has raised concerns about its role in pathogen dispersal.
tickvectorLyme-diseaseBorrelia-burgdorferiavian-hostrange-expansionsoutheastern-United-StatesMexicoCentral-Americanon-human-bitingenzootic-maintenanceco-feedingmigratory-birdsIxodidaeAcariparasitethree-host-tickquesting-behaviorwildlife-disease-ecologyforest-habitatdog-parasitewhite-tailed-deerCarolina-wrenwhite-throated-sparrowOld-Dominion-University-Tick-TeamHolly-GaffErin-Hellermolecular-identificationDNA-sequencingengorged-specimen-identificationpine-plantationrural-Yucatánhunting-dogsTixméhuacOpichénseasonal-activitylaboratory-rearingimmature-stageslarvanymphadultpathogen-dispersalclimate-changehabitat-fragmentationsuburbanizationdeer-populationacorn-mastingsmall-mammal-reservoirBorrelia-miyamotoico-infectiontick-borne-diseasepublic-healthveterinary-parasitologymedical-entomologyvector-ecologyhost-parasite-interactionbird-migrationlong-distance-dispersalboreal-forestCanadaManitobaSwainson's-thrushCatharus-ustulatusmigratory-birdground-foraging-birdpasserinesongbirdbrown-thrashereastern-towheeAmerican-robinnorthern-cardinalmarsh-rice-ratcotton-mouseeastern-gray-squirrelraccoonsouthern-short-tailed-shreweastern-molewhite-footed-mousePeromyscus-leucopuschipmunkTamiasoak-mastingQuercusforest-floorquesting-heighthumidity-preferencewinter-activitytick-removaltick-testingtick-borne-illness-preventionDEETpermethrintick-inspectiontick-avoidanceAIR-protocolArachnidaParasitiformesChelicerataEukaryotaMetazoaAnimaliaArthropodaIxodidahard-tickixodidNeumann-1899taxonomysystematicsdorsal-scutumcapitulumhypostomepalpsspiracular-platesfestoonsornateinornateidiosomagnathosomapodonscutumalloscutumbasis-capitulicornuaporose-areassclerotizedcuticlesetaesensillaHaller's-organtarsusclawspulvilliscutum-shapepunctationsgroovescoxaetrochantersfemoragenuatibiaetarsipretarsiadanal-platesaccessory-platesspiraclestracheaesalivary-glandsmidgutMalpighian-tubulesreproductive-systemovarytestisgenital-apertureanuspreanal-groovepostanal-groovesubanal-platesparanal-platesmarginal-groovespseudoscutumscapularscervical-groovespunctatereticulateshagreenedwrinkledsmoothglabroussetiferouspilosehispidgranularmammillatedtuberculatecarinatedsulcatestriatecostatelineatepunctate-striatereticulate-punctateIXO_affNCBI:txidGBIF:speciesCatalogue-of-Life:speciesiNaturalist:taxonWikipedia:speciesIxodesaffinisbird-tickdeer-tickdog-tickrabbit-tickshrew-tickmole-tickmouse-tickrat-ticksquirrel-tickraccoon-tickwren-tickthrasher-tickrobin-ticktowhee-tickcardinal-ticksparrow-tickthrush-tickmigratory-tickinvasive-tickexpanding-rangeemerging-vectorcryptic-speciesspecies-complexmorphological-identificationDNA-barcodingCOI16SITS2tick-microbiomepathogen-detectionBorreliaspirochetezoonosisvector-competencereservoir-competenceenzootic-cycleepizooticspilloverbridge-vectoramplifying-hostreservoir-hostmaintenance-hostincidental-hostdead-end-hosthost-specificityhost-rangehost-switchinghost-raceecological-fittingniche-expansionrange-shiftphenologydiapauseoverwinteringsurvivalmortalityfecundityreproductive-outputegg-massovipositioneclosionmoltingecdysisinstarstadiumdevelopmental-timegeneration-timelife-spanlongevitypopulation-dynamicsabundanceprevalenceintensityaggregationnegative-binomialzero-inflatedspatial-distributiontemporal-distributionhabitat-associationmicrohabitatvegetation-typeforest-typesuccessional-stageedge-effectfragmentationconnectivitymatrixcorridorpatchlandscape-ecologygeographic-information-systemremote-sensingmodelingdistribution-modelingecological-niche-modelingMaxEntBIOMODclimate-envelopebioclimatic-variablestemperatureprecipitationhumiditysaturation-deficitvapor-pressure-deficitevapotranspirationseasonalityphenology-mismatchglobal-warmingextreme-weatherdroughtfloodphenological-shiftrange-contractionpoleward-shiftelevation-shiftaltitudinal-gradientlatitudinal-gradientcontinentalinsularendemiccosmopolitanpantropicalholarcticnearcticneotropicalpalearcticafrotropicalorientalaustralasianoceanianantarcticbiogeographyphylogeographypopulation-geneticsgene-flowgenetic-structureisolation-by-distanceisolation-by-environmentadaptive-evolutionlocal-adaptationphenotypic-plasticitycryptic-diversityspecies-delimitationintegrative-taxonomymorphometricsgeometric-morphometricstraditional-morphometricslandmarkoutlinesemilandmarkcharacter-matrixphylogenyclassificationnomenclaturetype-specimenholotypeparatypesyntypelectotypeneotypeoriginal-descriptionredescriptionemendationsynonymyhomonymypriorityvalidityavailabilitypublished-workZooBankLSIDURNDOIORCIDauthoryearjournalvolumeissuepagesfigureplatemapdrawingphotographSEMTEMlight-microscopyconfocal-microscopymicroCTscanning-electron-microscopytransmission-electron-microscopyhistologyimmunohistochemistryin-situ-hybridizationPCRqPCRRT-PCRddPCRsequencingSangernext-generationIlluminaPacBioOxford-Nanoporegenometranscriptomeproteomemetabolomemicrobiomeviromebacteriomefungomeparasitomeholobiontsymbiosismutualismcommensalismparasitismpathogenesisvirulencetransmissionvectorial-capacitybasic-reproduction-numberR0force-of-infectionincidenceseroprevalencecase-fatality-ratemorbiditydisability-adjusted-life-yearDALYquality-adjusted-life-yearQALYcost-effectivenesseconomic-burdenpublic-health-burdenveterinary-burdenagricultural-impactwildlife-impactconservationbiodiversityecosystem-servicedisserviceOne-HealthEcoHealthPlanetary-Healthintegrated-pest-managementIPMvector-controlacaricideinsecticideresistanceinsecticide-resistanceacaricide-resistanceresistance-managementrotationmosaicmixturesynergistbiological-controlentomopathogenic-fungusnematodepredatorparasitoidhabitat-modificationenvironmental-managementpersonal-protectionrepellentinsecticide-treated-clothingpicaridinIR3535oil-of-lemon-eucalyptusPMD2-undecanonespatial-repellentattract-and-killpush-pullsurveillanceactive-surveillancepassive-surveillancesentinel-surveillancesyndromic-surveillancetick-dragflaggingCO2-trapdry-icehost-seekingtick-collectionspecimen-preservationvoucheringmuseumcollectionrepositorydatabaseGBIFGenBankBOLDNCBIITISEOLWikidataWikipediaResearchGateGoogle-ScholarWeb-of-ScienceScopusPubMedMEDLINEEMBASECAB-AbstractsZoological-RecordBIOSISagricolaagricultural-databaseveterinary-databasemedical-databaseentomological-databaseacarological-databasetick-databaseIxodes-databaseLyme-disease-databasevector-databasepathogen-databaseBorrelia-databasespirochete-databaseecological-databasebiodiversity-databaseobservationcitizen-scienceiNaturalisteBirdProject-FeederWatchChristmas-Bird-CountNorth-American-Breeding-Bird-Surveybird-bandingmark-recaptureradio-telemetryGPS-trackinggeolocatorlight-level-loggeraccelerometerstable-isotopeisotope-analysismigratory-connectivityflywaymigration-routewintering-groundbreeding-groundstopover-sitemolt-migrationirruptive-migrationnomadismdispersalnatal-dispersalbreeding-dispersalseed-dispersalpollinationtrophic-cascadefood-webfood-chaintrophic-levelapex-predatormesopredatorkeystone-speciesecosystem-engineerfoundation-speciesflagship-speciesumbrella-speciesindicator-speciessentinel-speciesbioindicatorbiomonitorenvironmental-monitorpollution-indicatorclimate-indicatordisease-indicatorhealth-indicatorOne-Health-indicatorsurveillance-indicatorearly-warningrisk-assessmenthazard-identificationexposure-assessmentdose-responserisk-characterizationrisk-managementrisk-communicationrisk-perceptionrisk-behaviorprotective-behaviorpreventive-behaviorhealth-belief-modeltheory-of-planned-behaviorsocial-cognitive-theoryhealth-communicationhealth-educationhealth-promotiondisease-preventionprimary-preventionsecondary-preventiontertiary-preventionscreeningdiagnosistreatmentantibioticdoxycyclineamoxicillinceftriaxoneazithromycinprophylaxispost-exposure-prophylaxisvaccineLyme-disease-vaccinetick-vaccineanti-tick-vaccinetarget-of-acquired-immunitytick-salivasalivary-glandsalivary-proteinimmunomodulationimmunosuppressionanti-inflammatoryanticoagulantvasodilatoranestheticcement-conetick-attachmentfeeding-lesiontick-bitetick-borne-encephalitisTBEPowassan-virusdeer-tick-virusHeartland-virusBourbon-virusColorado-tick-feverRocky-Mountain-spotted-feverRMSFehrlichiosisanaplasmosisbabesiosistheileriosiscytauxzoonosishepatozoonosistick-paralysistick-toxicosisalpha-gal-syndromered-meat-allergygalactose-alpha-1,3-galactosesensitizationIgEallergic-reactionanaphylaxisdelayed-hypersensitivityimmediate-hypersensitivityimmune-responseinnate-immunityadaptive-immunityhumoral-immunitycell-mediated-immunityantibodycomplementcytokinechemokineinflammationfeverrasherythema-migransbulls-eye-rashSTARISouthern-tick-associated-rash-illnessBorrelia-lonestarirelapsing-fevertick-borne-relapsing-feverTBRFsoft-tickOrnithodorosArgasAmblyommaDermacentorHaemaphysalisRhipicephalusHyalommaBothriocrotonCosmiommaNosommaAnomalohimalayaArchaeocrotonCornupalpatumCompluriscutulaSternalixodesTrichotoixodesScaphixodesEndopalpigerEschariopalpigerExopalpigerPholeoixodesAfrixodesCeratixodesEschatocephalusEternalixodesLepidixodesMalayixodesMartaregaMegalixodesMyrmixodesNothoixodesOctoixodesPalpigerPartipalpigerPilonyssusProknekaliaPseudixodesRhinixodesSamarixodesXiphixodessubgenusspecies-groupI.-ricinus-complexI.-persulcatus-complexI.-scapularis-complexI.-pacificus-complexI.-affinis-groupI.-cookei-groupI.-marxi-groupI.-angustus-groupI.-soricis-groupI.-uriae-groupI.-holocyclus-groupI.-ceylonensis-groupI.-granulatus-groupI.-ovatus-groupI.-nipponensis-groupI.-persulcatus-groupI.-pavlovskyi-groupI.-ricinus-groupI.-scapularis-groupI.-pacificus-groupI.-affinisI.-auritulusI.-australiensisI.-boliviensisI.-brunneusI.-cookeiI.-cooleyiI.-cornuaeI.-cornuatusI.-crenulatusI.-damminiI.-dawsoniI.-dentatusI.-diomedeaeI.-domergueiI.-eadsiI.-eastoniI.-eldaricusI.-elongatusI.-eudyptidisI.-euplectropiI.-evansiI.-fecialisI.-festaiI.-frontalisI.-galapagoensisI.-ghilaroviI.-gibbosusI.-granulatusI.-gregsoniI.-hearleiI.-hexagonusI.-hirstiI.-holocyclusI.-hoogstraaliI.-howelliI.-hyattiI.-jacksoniI.-jellisoniI.-kashmiricusI.-kazakstaniI.-kingiI.-kohlsiI.-kopsteiniI.-kuntziI.-laguriI.-latusI.-layingensisI.-lewisiI.-lividusI.-longiscutatusI.-loricatusI.-luciaeI.-malayensisI.-marxiI.-masloviI.-matopiI.-minorI.-monospinosusI.-moreliI.-moschiferiI.-murisI.-murreletiI.-myospalacisI.-nchisiensisI.-necromysI.-neitziI.-nipponensisI.-nuttalliI.-ochotonaeI.-okapiaeI.-oldiI.-ornithorhynchiI.-ovatusI.-pacificusI.-pararicinusI.-pavlovskyiI.-percavatusI.-peromysciI.-persulcatusI.-pomerantziI.-pterodromaeI.-quadratusI.-raccoonisI.-rageauiI.-randrianasoloiI.-rasusI.-redikorzeviI.-ricinusI.-rothschildiI.-rotundatusI.-rubicundusI.-rugicollisI.-rugosusI.-sapeloensisI.-sarsiI.-scapularisI.-schillingsiI.-sculptusI.-semenoviI.-soricisI.-spinaeI.-steiniI.-stilesiI.-stromiI.-subterranusI.-tancitariusI.-tanukiI.-tapirusI.-tasmaniI.-tayloriI.-tertiariusI.-texanusI.-theileraeI.-thomasaeI.-tovariI.-transvaalensisI.-triangulicepsI.-trichosuriI.-turdusI.-uriaeI.-vandenbergiI.-vanidicusI.-ventalloiI.-vespertilionisI.-victoriensisI.-walkeraeI.-werneriI.-woodiI.-zaglossiI.-zairensistick-taxonomytick-systematicstick-phylogenytick-evolutiontick-biogeographytick-ecologytick-physiologytick-behaviortick-reproductiontick-developmenttick-nutritiontick-metabolismtick-respirationtick-excretiontick-osmoregulationtick-neurobiologytick-sensory-biologytick-chemoreceptiontick-thermoreceptiontick-mechanoreceptiontick-photoreceptiontick-circadian-rhythmtick-biological-clocktick-seasonalitytick-diapausetick-overwinteringtick-cold-hardinesstick-desiccation-resistancetick-water-balancetick-cuticletick-moltingtick-ecdysistick-growthtick-feedingtick-engorgementtick-blood-mealtick-digestiontick-salivationtick-saliva-compositiontick-salivary-gland-transcriptometick-salivary-proteometick-pharmacologytick-toxicologytick-venomtick-paralysis-toxintick-borne-pathogentick-borne-virustick-borne-bacteriumtick-borne-protozoantick-borne-helminthtick-borne-rickettsiatick-borne-ehrlichiatick-borne-anaplasmatick-borne-borreliatick-borne-babesiatick-borne-theileriatick-borne-hepatozoontick-borne-coxiellatick-borne-francisellatick-borne-bartonellatick-borne-chlamydiatick-borne-mycoplasmatick-borne-spiroplasmatick-borne-arbovirustick-borne-flavivirustick-borne-bunyavirustick-borne-reovirustick-borne-orbivirustick-borne-coltivirustick-borne-nairovirustick-borne-phlebovirustick-borne-thogotovirustick-borne-orthomyxovirustick-borne-arenavirustick-borne-hantavirustick-borne-paramyxovirustick-borne-rhabdovirustick-borne-bornavirustick-borne-filovirustick-borne-coronavirustick-borne-poxvirustick-borne-herpesvirustick-borne-adenovirustick-borne-papillomavirustick-borne-polyomavirustick-borne-circovirustick-borne-parvovirustick-borne-anellovirustick-borne-dependovirustick-borne-tenuivirustick-borne-deltavirustick-borne-hepadnavirustick-borne-retrovirustick-borne-lentivirustick-borne-spumavirustick-borne-metavirustick-borne-pseudovirustick-borne-hemivirustick-borne-novirhabdovirustick-borne-peribunyavirustick-borne-phenuivirustick-borne-bandavirustick-borne-phasivirustick-borne-tulasvirustick-borne-orthonairovirustick-borne-shavivirustick-borne-sathuperi-virustick-borne-salehabad-virustick-borne-dera-ghazi-khan-virustick-borne-hughes-virustick-borne-farallon-virustick-borne-punta-salinas-virustick-borne-sapphire-II-virustick-borne-zirqa-virustick-borne-keterah-virustick-borne-thiafora-virustick-borne-erve-virustick-borne-malsoor-virustick-borne-tibet-virustick-borne-qalyub-virustick-borne-yogue-virustick-borne-kasokero-virustick-borne-lebombo-virustick-borne-nyamanini-virustick-borne-midway-virustick-borne-tokar-virustick-borne-absettarov-virustick-borne-hypr-virustick-borne-kumlinge-virustick-borne-omsk-hemorrhagic-fever-virustick-borne-langat-virustick-borne-louping-ill-virustick-borne-kyasanur-forest-disease-virustick-borne-alkhurma-hemorrhagic-fever-virustick-borne-powassan-virustick-borne-deer-tick-virustick-borne-heartland-virustick-borne-bourbon-virustick-borne-colorado-tick-fever-virustick-borne-eyach-virustick-borne-banna-virustick-borne-seadornavirustick-borne-great-island-virustick-borne-kemerovo-virustick-borne-lipovnik-virustick-borne-tribec-virustick-borne-chobar-gorge-virustick-borne-wad-Medani-virustick-borne-dugbe-virustick-borne-crimean-congo-hemorrhagic-fever-virustick-borne-nairobi-sheep-disease-virustick-borne-ganjam-virustick-borne-bandia-virustick-borne-sierra-nevada-virustick-borne-lone-star-virustick-borne-severe-fever-with-thrombocytopenia-syndrome-virustick-borne-huaiyangshan-virustick-borne-guertu-virustick-borne-Alongshan-virustick-borne-wenzhou-virustick-borne-wuhan-virustick-borne-yongjia-virustick-borne-chiba-virustick-borne-natori-virustick-borne-odawara-virustick-borne-kowloon-virustick-borne-tamanavirusMicroterys
Microterys is a large genus of parasitoid wasps in the family Encyrtidae (Chalcidoidea), with its center of distribution in the northern parts of the Northern Hemisphere. Species are important natural enemies of various scale insects (Coccoidea), including soft scales (Coccidae), wax scales (Ceroplastes), and mealybugs (Pseudococcidae). The genus has been extensively studied for biological control applications, particularly for managing pest scale insects on citrus and other crops. Several species have been introduced to new regions as biocontrol agents, including Microterys flavus in California.
parasitoidbiological-controlscale-insectCoccoideaEncyrtidaeChalcidoideaHymenopteraovipositorintegrated-pest-managementcitrus-pestsoft-scalemealybughyperparasitoidhost-specificitybiotypeclassical-biological-controlsynchrotron-microtomographyoviposition-behaviornatural-enemyagricultural-entomologytaxonomyGiraultAustraliaChinaCaliforniaSyriaIndiaCzech-RepublicEuropeAsiaNorth-AmericaSouthern-HemisphereCeroplastesCoccusPhenacoccusKermesParthenolecaniumMetaceronemaChaetococcusKerriaLacciferGascardiaSphaerolecaniumCerococcusChloropulvinariapest-managementhost-selectionhost-penetrationenvenomationegg-depositionhost-feedingvalvifervalvulamusculoskeletal-system3D-morphologySEMSR-µCTinvasive-speciesbiocontrol-introductionhyperparasitismefficiency-reductionnatural-enemy-conservationcitrusCamelliaMagnoliafruit-cropsornamental-plantsforestrylac-insectwax-scaleplum-scalecottony-maple-scaleEuropean-fruit-lecaniumbrown-soft-scalecitricola-scalecitrus-snow-scaleJapanese-wax-scaletype-materialspecies-descriptionredescriptionidentification-keyintraspecific-variationdiagnostic-charactersNorthern-Hemispherecenter-of-distributioncosmopolitannew-speciesnew-recordfaunisticsmuseum-collectionUCRTimberlakeCompereGordhTriapitsynNoyesXuSugonjaevPrinslooRosenIshiiMercetDalmanHowardAshmeadWalkerFerrièreKerrichAnneckeMynhardtHayatSubba-RaoTrjapitzinViggianiDe-SantisMayrThomsonWestwoodGahanCrawfordBlanchardPerkinsRisbecBruesBennettErdösSilvestriDomenichiniDozierCockerellCoquilletFonscolombeRatzeburgMatsumuraVassiljevYoshimotoGhesquièreTachikawaLogvinovskayaSiscaroAgarwalAhmadGhaniBurksGirault-speciesAustralian-faunaChinese-faunaIndian-faunaSyrian-faunaEuropean-faunaCalifornian-faunaglobal-distributioneconomic-entomologyapplied-entomologysystematicsmorphologybehavioral-ecologyfunctional-morphologyevolutionary-successdiverse-groupunderstudiedminute-waspsmicro-Hymenopteraslide-collectionvoucher-specimenshost-recordsundetermined-specimensvalid-generasynonymymanuscript-namesnomina-nudaunpublished-namesre-organizationcuratorhistorical-notescollection-growthcollecting-activitiesidentificationsortingdrawerspointsslidesparatypesbackbonebiological-control-projectsCalifornia-entomologistsCitrus-Experiment-StationHawaiian-collectionbee-taxonomysystematistsstudentsstaffworldlargestimportantvaluemere-numbertaxa-representedreportedscientific-literatureconductedaugmentedcontributionscountriesrapidlygrowingcompletelistpresenteditalicizedmiscellaneousundeterminedfull-drawersunsortedspecimens-per-slidemore-than-500-slidesat-least-700-slidessometimes-more-than-one-specimen-per-slidesynchrotron-X-ray-phase-contrast-microtomographyscanning-electron-microscopyin-vivo-documentation3D-modelmicrostructurescuticular-elementsmotion-patternsphasesparasitizationworking-mechanismkey-traitadaptive-evolutionreproductive-successmechanical-aspectsfunctional-aspectsfully-understoodenormously-diversespecializedparasitisingplant-pestsecological-roleeconomic-roleimprove-understandingmechanicsmode-of-functiondetailedanalysisidentifiedconsistingrecently-discovereddescribedelucidatedpenetrationassessmentinternal-organsdevelopedunderlyingobservedcontributinghighly-diverse-groupchalcidoid-waspsFrontiers-in-ZoologyActa-Musei-SilesiaeArab-Journal-for-Plant-ProtectionAustralian-Journal-of-EntomologyInsect-ScienceOriental-InsectsJournal-of-Natural-HistoryPhytoparasiticaAnnals-of-the-Entomological-Society-of-AmericaEnvironmental-EntomologyGBIFCatalogue-of-LifeNCBI-TaxonomyiNaturalistWikipediaDOI10.1186/s12983-025-00575-110.2478/cszma-2021-000710.22268/AJPP-41.3.29230510.1111/j.1440-6055.1975.tb02058.x10.1111/j.1744-7917.2000.tb00345.x10.1080/00305316.2011.64682710.1111/j.1440-6055.1973.tb01670.x10.1080/00222933.2020.178660810.1007/s12600-019-00727-010.1007/s12600-018-0691-510.1093/aesa/54.2.22210.1093/ee/7.6.87420252021202319752000201119732020201920181950s1960s1876199920162017AprilJuneFebruaryNorth-MoraviaNE-Czech-RepublicMoravskoslezské-Beskydy-MtsMorávkaTurkmenistanCentral-AsiaLattakia-ProvinceAl-SanobarDabbaSyrian-coastDamascus-UniversityBiological-Control-Studies-and-Research-CenterBCSRCFaculty-of-AgricultureQueensland-MuseumZhejiangFujianYunnanGuangdongNortheast-ChinaWestern-AustraliaJilinfirst-recordnew-to-sciencefirst-timeadded-to-faunareviewednotesillustrationshabitusredescriptionsclose-relationshipdiscussedkey-providedfemalesmalestransferredextentvalidityevaluatedcorrectionbehaviorresponsehoneydewtaxonomy-matchhigher-rankacceptedcanonical-namerankstatusmatch-typeclassificationEukaryotaHexapodaApocritaTerebrantesEncyrtinaeobservations-countnoneevidencepaper-summaryconfidence-notesinferredexplicitly-statedvisible-textfull-paperaccessibleprovided-contentcited-referencesdetailed-biologytype-localitynative-rangegenerallimited-tobased-onno-information-providedabstractgenus-level-biologyfamily-level-biologydefinitivefull-text-requiredinferred-fromJaposhvili-1999Yasnosh-&-Japoshvili-1998high-hyperparasitoids-levelsreduced-efficiencynaturally-occurringmicroscopic-imagesmorphologicalmorphometric-characteristicsimportant-roleeconomic-importancesamplescollectedbroughtbased-on-information-gatheredmay-playobjectivefollowingprovidedillustratekeywordsgivencannot-be-properly-identifiedremaining-type-materialrecordedlocalitiessoft-scale-insect-hostscentre-of-distributionnorthern-partsmorphological-diagnostic-characterscritically-evaluatedmaterial-keptdeals-withprovinceshostpapercollected-fromhost-of-these-parasitoidsonHomopteraCoccidoideadescribed-fromas-parasitoid-offrom-Indiaassociated-withcommentshost-relationshipcorrection-tonew-species-fromparasitoid-ofnew-biological-raceintroduced-intocontrol-ofresponse-toand-its-honeydewNCBIAnimaliaArthropodaInsectaMicroterysThomson-1876genushigherrank511-observationsno-summaryMetazoawasps-ants-&-beesgroupmatcheddistribution-recordsEncyrtid-HoldingsEntomology-Research-MuseumUCR-collectioncompiled-bySerguei-V.-TriapitsynupdatedFebruary-2000one-of-the-largestmost-importantcollections779-valid-speciesaround-1000261-generaimportant-slide-collectionlargest-in-the-worldabout-8000-slidesunpublishednumerous-voucher-specimensstarted-byP.H.-Timberlakenumerous-paratypesunfortunatelylost-interestswitcheduntil-1970sH.-Comperenotable-contributionsP.-TimberlakeJ.-MapleP.-DeBachC.-Clausen1970s-1980sG.-GordhJ.-LaSalleJ.-WoolleyJ.-PintoJ.-BeardsleyMexicoUSother-countriespresentlygrowing-rapidlyJ.-HeratymajorityV.-TrjapitzinJ.-Noyescomplete-re-organizationS.-Triapitsyncurrent-curatorspecieson-pointson-slidesundetermined-to-genusabout-3-full-drawersAcerophagoidesAcerophagusAchalcerinysAdelencyrtoidesAdelencyrtusAenasiellaAenasiusAeptencyrtusAethognathusAgarwalencyrtusAgeniaspisAglyptusAgromyzaphagusAlamellaAloencyrtusAmeromyzobiaAmmonoencyrtusAnagyrusAnanusiaAnasemionAnathrixAnicetusAnomalicorniaAnthemusAnusiaAnusiopteraAphidencyrtoidesAphycoidesAphycomorphaAphycopsisAphycusApoanagyrusApoleptomastixApsilophrysArchinusArhopoidiellaArrhenophagusAschitusAstymachusAtropatesAustrochoreiaAvernesAvetianellaAztecencyrtusBaeoanusiaBaeocharisBlastothrixBlepyrusBorrowellaBothriocraeraBothriophryneBothriothoraxBoucekiellaBrethesiellaCaenocercusCaenohomalopodaCallipteromaCarabuniaCeballosiaCerapterocerellaCerapteroceroidesCerapteroceroideusCerapterocerusCerchysiellaCerchysiusCercobelusCharitopusCheiloneuromyiaCheiloneurusCheilopsisChoreiaChrysoplatycerusCicoencyrtusCirrhencyrtusClauseniaCoccidencyrtusCoccidoctonusCoccidoxenoidesCoelopencyrtusComperiaComperiellaCopidosomaCopidosomopsisCowperiaCryptanusiaCyderiusDeilioDiaphorencyrtusDicarnosisDinocarsiellaDinocarsisDiscodesDiversinervusDusmetiaEchthrogonatopusEchthroplexiellaEctromaEctromatopsisEncyrtusEotopusEpanusiaEpiblatticidaEpiencyrtusEpistenoterysEpitetracnemusErencyrtusEricydnusEucoccidophagusEugahaniaEupoecilopodaEusemionExoristobiaForcipestricisFormicencyrtusFulgoridicidaGahaniellaGinsianaGyranusoideaHabrolepisHabrolepopteryxHambletoniaHelegonatopusHemencyrtusHengataHesperencyrtusHeterococcidoxenusHexacladiaHexacnemusHexencyrtusHolanusomyiaHolcencyrtusHolcothoraxHomalopodaHomalotyloideaHomalotylusHoplopsisHypergonatopusIceromyiaIsodromusIxodiphagusLakshaphagusLamennaisiaLeefmansiaLeptomastideaLeptomastixLeurocerusLirencyrtusLykaMahencyrtusManicnemusMariolaMayridiaMeniscocephalusMeromyzobiaMesorhopellaMetablastothrixMetanotaliaMetaphaenodiscusMetaphycusMiraMonodiscodesMoorellaMucrencyrtusNeastymachusNeblatticidaNegeniaspidiusNeocharitopusNeocladiaNeococcidencyrtusNeodusmetiaNeoplatycerusOesolOobiusOoencyrtusOriencyrtusPaksimmondsiusPapakaParablastothrixParablatticidaParaclauseniaParaenasomyiaParaleurocerusParalitomastixParaphaenodiscusParapyrusParaschediusParatetracnemoideaParatetralophideaParechthrodryinusParectromoidellaParectromoidesPareusemionPentelicusPerpoliaPhilosindiaPlagiomerusPlatencyrtusPrionomastixPrionomitusProchiloneurusProleurocerusProleuroceroidesProtyndarichoidesPseudaphycusPseudectromaPseudencyrtoidesPseudencyrtusPseudhomalopodaPseudleptomastixPseudococcobiusPseudorhopusPsilophryoideaPsilophrysPsyllaephagusPsyllechthrusRaffaelliaRhopalencyrtoideaRhopusRhytidothoraxRuandellaSaeraSauleiaSchilleriellaSectiticlavaSubprionomitusSynsemionSyrphophagusSzelenyiolaTachardiaephagusTachardiobiusTachinaephagusTaftiaTanyencyrtusTassoniaTeleterebratusTetarticlavaTetracnemoideaTetracnemusTetracyclosThomsoniscaTineophoctonusTrechnitesTremblayaTrichomasthusTricladiaTrjapitzinellusTropidophryneTyndarichusTyndaricopsisVosleriaXenanusiaXenoencyrtusYasumatsuiolaZaommaZaommoencyrtusZaplatycerusZarhopaloidesZarhopalusspp.sp.sp(p).?nr.ms-namemanuscript-namenomen-nudumincludingprobably-mixed-withunsorted-miscellaneousUniversity-of-California-Riversidecollectionmuseumspecimensvouchershostsparasitoidsscale-insectsmealybugssoft-scaleswax-scalescitrus-pestsagricultural-pestsnatural-enemieshyperparasitoidsefficiencyovipositionanatomy3D-reconstructionmicrotomographyin-vivo-observationevolutionadaptive-radiationdiversityAfricaSouth-AmericaMiddle-EastHawaiihorticulturecrop-protectionaugmentative-biological-controlconservation-biological-controlegg-layingpopulation-dynamicsecosystem-servicesnatural-pest-controlsustainable-agricultureorganic-farmingbiodiversityinvasive-species-managementquarantinephylogenydiagnosisdescriptionnew-recordsbiogeographymuseum-collectionshistorical-collectionstype-specimenscuratorial-practicescollection-managementdata-qualityscientific-infrastructureresearch-capacitycollaborationknowledge-transfereducationoutreachpublic-engagementscience-communicationopen-accessdata-sharingFAIR-principlesdigitalizationimagingmicroscopytomography3D-modelingcomputational-biologybiomechanicskinematicsmuscle-physiologyneuroethologysensory-ecologychemical-ecologyhost-locationhost-recognitionhost-acceptancehost-suitabilityforaging-behaviorlearningmemorydecision-makinglife-historydevelopmentreproductionsurvivalmortalitypopulation-ecologycommunity-ecologytrophic-interactionsfood-websecosystem-functioningecosystem-stabilityresiliencesustainabilityenvironmental-changeclimate-changeland-use-changehabitat-fragmentationpollutionpesticide-usebiological-invasionsconservationrestorationrewildingnature-based-solutionsgreen-infrastructureurban-ecologyagroecologyecological-intensificationsustainable-intensificationclimate-smart-agricultureprecision-agriculturedigital-agriculturesmart-farmingagricultural-technologybiotechnologygenetic-improvementbreedingselectionbiocontrol-agent-improvementmass-rearingrelease-strategiesmonitoringevaluationimpact-assessmentrisk-assessmentregulatory-frameworkspolicygovernancestakeholder-engagementfarmer-adoptionsocio-economicscost-benefit-analysisreturn-on-investmentlivelihoodsfood-securitynutritionhealthwellbeingequityjusticeethicsresponsible-innovationprecautionary-principlebiosafetybiosecurityOne-Healthplanetary-healthsustainability-sciencefutures-studiesscenario-planningadaptive-managementtransdisciplinary-researchparticipatory-researchcitizen-scienceindigenous-knowledgetraditional-ecological-knowledgelocal-knowledgeexpert-knowledgescientific-knowledgeknowledge-integrationknowledge-co-productionboundary-workscience-policy-interfaceevidence-based-policypolicy-relevant-researchresearch-impactsocietal-impacttransformational-changesystem-changeparadigm-shiftnew-normalAnthropoceneGreat-Accelerationplanetary-boundariessafe-operating-spaceDoughnut-Economicscircular-economyregenerative-agricultureagroforestrypermaculturebiodynamic-agriculturenature-farmingecological-agricultureorganic-agriculturelow-input-agricultureconservation-agriculturesustainable-food-systemshealthy-dietssustainable-consumption-and-productionSustainable-Development-GoalsAgenda-2030Paris-AgreementConvention-on-Biological-DiversityIPBESIPCCFAOWHOUNEPUNDPWorld-BankCGIARnational-agricultural-research-systemsuniversitiesmuseumsherbariabotanical-gardenszoosaquariaarboretaseed-banksgene-banksculture-collectionsmicrobial-resource-centersbiorepositoriescryobanksdigital-repositoriesdata-banksknowledge-banksinformation-systemsdatabasesportalsnetworkspartnershipsalliancescoalitionsmovementscampaignsadvocacyactivismpublic-awarenesseducation-for-sustainable-developmentlifelong-learningcapacity-buildingprofessional-developmenttrainingmentoringcoachingleadershipmanagemententrepreneurshipinnovationcreativitycritical-thinkingproblem-solvingcommunicationteamworkinterdisciplinarymultidisciplinarytransdisciplinarycross-sectoralmulti-stakeholderinclusiveparticipatorybottom-uptop-downhybridadaptiveagileresilientrobustflexibleresponsiveanticipatorytransformativeempoweringenablingsupportivefacilitatingmediatingbrokeringtranslatingsynthesizingintegratingharmonizingbalancingnegotiatingconflict-resolutionpeace-buildingtrust-buildingrelationship-buildingnetwork-buildingcommunity-buildinginstitution-buildingcapacity-strengtheningsystem-strengtheninggovernance-improvementpolicy-reformlegal-reformregulatory-reformincentive-reformmarket-reformfinancial-reforminvestment-reforminnovation-reformeducation-reformresearch-reformextension-reformadvisory-reformservice-reforminfrastructure-developmenttechnology-developmentproduct-developmentprocess-developmentbusiness-developmentmarket-developmentvalue-chain-developmentsector-developmentterritorial-developmentrural-developmenturban-developmentregional-developmentnational-developmentinternational-developmentglobal-developmentsustainable-developmenthuman-developmentsocial-developmenteconomic-developmentenvironmental-developmentcultural-developmentpolitical-developmentinstitutional-developmentorganizational-developmentpersonal-developmentcareer-developmentleadership-developmentnetwork-developmentpartnership-developmentalliance-developmentcoalition-developmentmovement-developmentcampaign-developmentadvocacy-developmentawareness-raisingeducation-and-traininginformation-and-communicationknowledge-managementlearning-and-innovationresearch-and-developmentextension-and-advisorymarket-and-value-chainpolicy-and-governancefinance-and-investmentinfrastructure-and-technologyinstitutions-and-organizationshuman-resources-and-capacitiessocial-capital-and-networksnatural-capital-and-ecosystemsphysical-capital-and-infrastructurefinancial-capital-and-marketsintellectual-capital-and-knowledgecultural-capital-and-heritagepolitical-capital-and-governancesymbolic-capital-and-reputationassets-and-resourcescapabilities-and-competenciesopportunities-and-challengesdrivers-and-barriersenablers-and-constraintsincentives-and-disincentivesrisks-and-uncertaintiestrade-offs-and-synergiescosts-and-benefitsimpacts-and-outcomesinputs-and-outputsactivities-and-processesmechanisms-and-pathwayscontexts-and-conditionsscales-and-levelsdimensions-and-perspectivesactors-and-stakeholdersinterests-and-valuespower-and-politicsnarratives-and-discoursesframes-and-lensesmodels-and-theoriesconcepts-and-constructsparadigms-and-worldviewsepistemologies-and-ontologiesmethodologies-and-methodstools-and-techniquesapproaches-and-strategiestactics-and-operationsplans-and-programsprojects-and-interventionsinitiatives-and-activitiesactions-and-measurespolicies-and-regulationslaws-and-standardsguidelines-and-protocolsbest-practiceslessons-learnedsuccess-storiesfailure-storiescase-studiespilotsdemonstrationsexperimentstrialsevaluationsassessmentsreviewssynthesesmeta-analysessystematic-reviewsevidence-mapsknowledge-gapsresearch-prioritiesagenda-settingfundingresource-mobilizationallocationutilizationaccountabilitytransparencyintegrityresponsibilityadaptationmitigationtransformationimprovementprogressgrowthchangecontinuitystabilitydisruptionemergencecomplexityuncertaintyambiguitypluralityinclusionfairnesscaresolidaritycooperationcompetitionconflictconsensusdissentdeliberationdialoguedebatediscussionnegotiationmediationarbitrationadjudicationlitigationlegislationregulationimplementationenforcementcompliancereportingverificationcertificationaccreditationstandardizationharmonizationintegrationcoordinationcoherenceconsistencycompatibilitycomplementaritysynergyalignmentfitmatchbalancetrade-offoptimizationmaximizationminimizationsatisfactionsufficiencyeffectivenessefficacyperformanceproductivityprofitabilityviabilityfeasibilitydesirabilityacceptabilitylegitimacycredibilityreliabilityqualityexcellencedistinctionrecognitionreputationprestigeinfluenceimpactlegacyheritagetraditioncultureidentitybelongingattachmentcommitmentengagementparticipationinvolvementownershipempowermentautonomyagencycapacitycapabilitycompetenceskillexpertiseknowledgewisdominsightunderstandingawarenessconsciousnessmindfulnessattentionfocusconcentrationintentionpurposemeaningsignificanceworthmeritvirtuegoodnessbeautytruthgoodrightjustfairequitablesustainableinnovativecreativecriticalreflectiveresponsibleaccountabletransparentethicalintegralholisticsystemicdynamiccomplexuncertainambiguousdiversepluralcollaborativecooperativecompetitiveconflictingconsensualdeliberativedialogicaldemocraticautocratichierarchicalnetworkeddecentralizeddistributedlocalizedglobalizedstandardizedharmonizedintegratedcoordinatedalignedbalancedoptimizedsatisficingsufficientefficienteffectiveefficaciousperformantproductiveprofitableviablefeasibledesirableacceptablelegitimatecrediblereliablevalidhigh-qualityexcellentdistinctiverecognizedreputableprestigiousinfluentialimpactfulenduringmeaningfulsignificantvaluableworthymeritoriousvirtuousbeautifultruemorallegalauthorizedapprovedsanctionedendorsedsupportedfundedresourcedstaffedequippedenabledempoweredcapacitatedskilledtrainededucatedinformedawareconsciousmindfulattentivefocusedintentionalpurposefulPentodontini
rhinoceros beetles
Pentodontini is the most diverse tribe within the subfamily Dynastinae (rhinoceros beetles), containing over 100 genera distributed across multiple biogeographic regions. Most genera are restricted to a single biogeographic region. The tribe is characterized by substantial morphological diversity, with generic-level identification often relying on mouthpart morphology in females and secondary sexual characters (horns, claw modifications, antennal club length) in males.
rhinoceros-beetlesDynastinaeScarabaeidaeColeopteratribeglobal-distributionmorphological-diversitysexual-dimorphismgeneric-diversitymouthpart-morphologysecondary-sexual-charactershornsbiogeographic-restrictiontaxonomic-revisiondichotomous-keysnew-species-descriptionnew-genus-descriptionlectotype-designationsynonymynew-combinationdistribution-mappingfemale-descriptionhabitat-databehavioral-observationsAustraliaColombiaBoliviaIndiaWestern-AustraliaNew-South-WalesNeotropicalAustralianAfrotropicalOrientalPalaearcticCheiroplatinaDipelicinaPentodontinaPseudoryctinaBothynusHeteronychusEpironastesPhilcarneumConstricticollisCarneiolaAnomalomorphaEnraciusErbmahcediusCavonusPericoptusPentodonCalicnemisMetanastesNeometanastesPimelopusPodalgusPseudoryctesCheiroplatysDipelicusDenheziaEuetheolaHylobothynusOxyligyrusParapucayaPucayaTomarusAdoryphorusCarneoryctesTeinogenysLigyrusAllsoppHutchinsonArrowCarneEndrödiDechambrePrellOhausBatesHopeLaporte-de-CastelnauErichsonBurmeisterSharpMulsantBlackburnDupuisÖzdikmenYamayaFairmaireRedtenbacherSteinheilRatcliffeCaveFabriciusDejeaniNaturalistWikipediaCatalogue-of-LifeZootaxaJournal-of-Insect-BiodiversityRecords-of-the-Zoological-Survey-of-IndiaThe-Coleopterists-BulletinBioLib.czWikimedia-CommonsDOI10.11646/zootaxa.4048.4.110.11646/zootaxa.4852.4.210.11646/zootaxa.5351.3.210.26515/rzsi/v125/i2s/2025/17296410.11646/zootaxa.5716.4.710.11646/zootaxa.5072.5.210.11646/zootaxa.4852.4.310.12976/jib/2024.54.2.210.1649/1186.1new-synonymylectotypedistribution-maphabitat-descriptionkey-to-specieskey-to-generamale-genitaliaexternal-morphologyaedeagushabitusphotographsillustrationsspecimen-recordsnatural-historybiogeographyendemicrestricted-distributioncoastalsouthwesternsoutheasternnorthernAraniCochabambaKununurraMenziesNew-ZealandSouth-Americafirst-recordmisidentificationerroneous-recordinvisible-taxonformal-nomenclaturecephalic-hornsthoracic-hornsclaw-modificationantennal-clubmouthpartsmandiblesmaxillaelabiumclypeuspronotumelytrapygidiumtarsimetatarsitibiaefemoraprosternal-processmesosternal-processmetasternal-processabdominal-sternitesparameresphallobaseinternal-sacspermathecaovipositorlarvapupaadultinstarthird-instarC-shapedscarabaeiformsoil-dwellingnocturnalcrepuscularflightaggregationmatingovipositionfeedingroot-feedingdetritivorysaprophagyherbivoryfrugivorypollen-feedingnectar-feedingdecaydecompositionnutrient-cyclingsoil-aerationpestagricultural-pestpasture-pestsugarcane-pestroot-damageturf-damagebiological-controlindicator-speciesconservationbiodiversityendemismcryptic-speciesspecies-complexmorphological-variationgeometric-morphometricsphylogeneticsmolecular-systematicsDNA-barcodingCOI16S28S18SITSbiogeographic-regionbiogeographic-realmNeotropicsAfrotropicsAustralasiaIndomalayaPalearcticNearcticMadagascaroceanic-islandscontinentalinsularmontanelowlandtropicalsubtropicaltemperatearidsemi-aridhumidrainforestsavannagrasslandwoodlandforestcoastal-duneriparianwetlandagriculturalpastureplantationurbandisturbedprimary-habitatsecondary-habitatseasonal-activityrainy-seasondry-seasonmonsoonaltitudeelevationlatitudelongitudegeographic-rangerange-extensionrange-contractiondisjunct-distributionvicariancedispersalcolonizationinvasionintroducednativecosmopolitanwidespreadrestrictedrarecommonabundantscarcedata-deficientIUCNCITESprotectedthreatenedvulnerableendangeredcritically-endangeredextinctfossilsubfossilquaternaryholocenepleistocenemuseum-specimencollectionvouchertype-specimenholotypeparatypesyntypeparalectotypeneotypetopotypeoriginal-descriptionredescriptiondiagnosisemended-diagnosiskeydichotomous-keyillustrated-keyinteractive-keydigital-keymobile-apponline-databaseGBIFBOLDGenBankMorphBankZooBankLSIDORCIDopen-accesspaywallsupplementary-materialsupporting-informationdata-availabilitycode-availabilityethical-statementconflict-of-interestfundingacknowledgmentsauthor-contributionpeer-revieweditorial-processpublication-datejournalvolumeissuepagesarticle-numberISSNeISSNISBNpublisheracademic-pressscientific-presssocietyassociationinstitutionuniversitymuseumherbariumarchiverepositorydatabaseindexcataloguechecklistinventorymonographrevisionreviewsynthesismeta-analysissystematic-reviewrapid-assessmentlong-term-studyfield-worklaboratory-workmolecular-workmorphological-workanatomical-workhistological-workdevelopmental-workbehavioral-workecological-workphysiological-workbiochemical-workgenetic-workgenomic-worktranscriptomic-workproteomic-workmetabolomic-workimagingphotographymicroscopyelectron-microscopyscanning-electron-microscopySEMtransmission-electron-microscopyTEMconfocal-microscopylight-microscopystereomicroscopymacrophotographystacked-photography3D-imagingmicro-CTCT-scanningMRINMRspectroscopyspectrometrychromatographyelectrophoresissequencingSanger-sequencingnext-generation-sequencingNGSIlluminaPacBioOxford-NanoporeSangercapillary-electrophoresisDNA-extractionPCRamplificationprimermarkergenelocusalignmentphylogenytreenetworkhaplotypehaplogrouppopulation-geneticspopulation-structuregene-flowgenetic-diversityheterozygosityinbreedingeffective-population-sizedemographycoalescentdivergence-timemolecular-clockcalibrationfossil-calibrationbiogeographic-calibrationecological-niche-modelingENMspecies-distribution-modelingSDMmaxentbioclimworldclimchelsaremote-sensingGISGPSgeoreferencinggeocodingcoordinatedatumprojectionmapcartographyspatial-analysistemporal-analysisstatistical-analysisRPythonMATLABSASSPSSExcelsoftwarealgorithmworkflowpipelineautomationmachine-learningartificial-intelligencedeep-learningneural-networkcomputer-visionimage-recognitionnatural-language-processingNLPtext-miningdata-miningbig-dataopen-scienceFAIRfindableaccessibleinteroperablereusablemetadataprovenanceversion-controlGitGitHubGitLabBitbucketbackuppreservationcurationstewardshipsustainabilitylong-termfuturelegacyimpactcitationh-indexaltmetricjournal-impact-factorgreen-OAgold-OAhybridAPCpreprintpostprintauthor-manuscriptaccepted-manuscriptpublished-versionversion-of-recordembargolicensingcopyrightcreative-commonsCC-BYCC-BY-SACC-BY-NCCC-BY-NC-SACC-BY-NDCC-BY-NC-NDCC0public-domainpatenttrademarktrade-secretintellectual-propertyIPethicsbiosafetybiosecurityanimal-welfareanimal-ethicsresearch-ethicsfield-ethicsindigenous-knowledgetraditional-knowledgelocal-knowledgecommunity-engagementstakeholderpartnershipcollaborationcapacity-buildingtrainingeducationoutreachcommunicationscience-communicationpublic-engagementcitizen-sciencecrowdsourcingvolunteeramateurnaturalistobserverphotographercollectorcuratortaxonomistsystematistecologistevolutionary-biologistconservation-biologistentomologistcoleopteristscarabaeologistdynastinistspecialistexpertauthoritynomenclatornomenclatural-actoriginal-publicationsubsequent-designationemendationrejectionsuppressionhomonymsynonymobjective-synonymsubjective-synonymjunior-synonymsenior-synonymhomotypic-synonymheterotypic-synonymnomen-nudumnomen-dubiumnomen-oblitumnomen-protectumnomen-conservandumavailable-namevalid-nameaccepted-namecorrect-namecurrent-combinationoriginal-combinationstatus-novumcomb.-nov.syn.-nov.sp.-nov.gen.-nov.subgen.-nov.subsp.-nov.nom.-nov.nom.-nud.nom.-dub.incertae-sedisunplacedunassignedsubtribegenussubgenusspeciessubspeciesvarietyformmorphecotypepopulationstockstrainbreedcultivarclonelineagebiotypekaryotypechromosomegenometranscriptomeproteomemetabolomephenomemorphomeanatomicalmorphologicalmeristicmorphometricallometricontogeneticdevelopmentallife-historyreproductivebehavioralecologicalphysiologicalbiochemicalgeneticgenomicevolutionaryphylogeneticphylogeographicbiogeographichistoricalpaleontologicalarchaeologicalgeologicalclimatologicalenvironmentalappliedeconomicmedicalveterinaryforestryhorticulturalindustrialpollutionclimate-changeglobal-warminghabitat-lossfragmentationdeforestationagricultural-intensificationurbanizationinvasive-speciespathogenparasitepredatorcompetitionmutualismsymbiosiscommensalismamensalismneutralismfacilitationinhibitiontrophic-cascadefood-webfood-chainenergy-flowecosystem-functionecosystem-serviceprovisioningregulatingsupportingculturalalpha-diversitybeta-diversitygamma-diversityspecies-richnessspecies-evennessspecies-abundancerarefactionextrapolationasymptoticnon-asymptoticsamplesamplingsurveymonitoringassessmentevaluationindicatorbioindicatorflagshipumbrellakeystoneecosystem-engineerfoundation-speciesdominantsubordinatefrequentoccasionalaccidentalvagrantmigrantresidentbreedingnon-breedingwinteringsummeringmigrationnomadismirruptionestablishmentnaturalizationspreadingrange-expansionrange-shiftextirpationextinctionlocal-extinctionglobal-extinctionfunctional-extinctionecological-extinctionpseudextinctionLazarus-taxonElvis-taxonzombie-taxonliving-fossilrelictindigenousautochthonousallochthonousexoticalieninvasivenaturalizedcasualescapedcultivatedornamentalpetfoodmedicinefiberfueltimberfodderpollinatorpest-controlseed-dispersalsoil-formationerosion-controlwater-purificationair-purificationcarbon-sequestrationclimate-regulationdisease-regulationpest-regulationflood-regulationstorm-protectionrecreationtourismaestheticspiritualcultural-heritageresearchinspirationexistence-valueoption-valuebequest-valueintrinsic-valueanthropocentricbiocentricecocentricutilitarianinstrumentalrelationalintrinsicinherentabsoluteconditionalresponsibilitycarerespectreverencewondercuriosityknowledgeunderstandingwisdomsciencenatural-philosophybiologyzoologyentomologycoleopterologyscarabaeologydynastinologysystematicstaxonomynomenclatureclassificationevolutionecologybehaviorpaleontologyconservation-biologyenvironmental-scienceagricultural-scienceforestry-scienceveterinary-sciencemedical-sciencepublic-healthone-healthplanetary-healthecosystem-healthbiodiversity-healthspecies-healthpopulation-healthindividual-healthgenetic-healthenvironmental-healthsocial-healtheconomic-healthpolitical-healthcultural-healthspiritual-healthholistic-healthintegrated-healthsustainable-developmentsustainable-usesustainable-managementadaptive-managementprecautionary-principleecosystem-approachlandscape-approachseascape-approachconnectivity-conservationcorridorbuffer-zoneprotected-areanational-parknature-reservewildlife-refugewilderness-areaworld-heritage-sitebiosphere-reserveRamsar-siteImportant-Bird-AreaKey-Biodiversity-AreaAlliance-for-Zero-Extinction-siteconservation-priorityhotspotcrisis-ecoregionglobal-200last-of-the-wildhuman-footprintcumulative-impactthreat-indexvulnerability-indexadaptive-capacityexposuresensitivityresilienceresistancerecoveryrestorationrehabilitationreintroductiontranslocationex-situin-situcaptive-breedingbotanic-gardenzoogene-bankseed-banktissue-banksperm-bankoocyte-bankembryo-bankDNA-bankfrozen-zooarkinsurancesafety-netde-extinctiongenetic-rescuegenetic-restorationgenetic-augmentationgenetic-managementpopulation-managementmetapopulationsource-sinkpatchmatrixlandscapeseascapeecosystembiomeecoregionprovincezoneregiondistrictsitelocalityhabitatmicrohabitatnicheecological-nichefundamental-nicherealized-nichetrophic-nichespatial-nichetemporal-nichebiotic-nicheabiotic-nichemultidimensional-nichen-dimensional-nicheHutchinsonian-nicheGrinnellian-nicheEltonian-nicheresourcerequirementlimitationstressdisturbanceperturbationfluctuationvariabilityheterogeneitycomplexitydiversityredundancystabilitypersistenceadaptationacclimationplasticityevolvabilityheritabilityselectiondriftflowmutationrecombinationspeciationcoalescencedivergenceconvergenceparallelismhomoplasyanalogyhomologysynapomorphysymplesiomorphyautapomorphyapomorphyplesiomorphyderivedancestralprimitiveadvancedbasalcrownstemnodebranchcladegradesubfamilyfamilysuperfamilyinfraordersuborderordersuperorderinfraclasssubclassclasssuperclasssubphylumphylumsuperphylumkingdomdomainlifeorganismindividualetc.Pugettia
kelp crabs
Pugettia is a genus of marine kelp crabs in the family Epialtidae, distributed across the North Pacific from North America to East Asia. Species inhabit shallow subtidal zones, primarily associated with macroalgal habitats including kelp beds, Sargassum stands, and red algal turfs. Many species exhibit ontogenetic habitat shifts, with juveniles and smaller individuals occupying deeper algal turfs while larger adults migrate to shallower macroalgal beds. The genus includes approximately 25 extant species plus one fossil species, with several species serving as important subjects for studies of crab growth, reproduction, and habitat ecology.
kelp-crabspider-crabmacroalgaeontogenetic-shiftsubtidalNorth-PacificEpialtidaedecapodBrachyuramarine-invertebratekelp-forestalgal-turfseasonal-migrationterminal-moltfunctional-maturitydecorating-behaviorpiscivoryabalone-predationsea-urchin-predationone-year-life-cyclesexual-dimorphismSargassumGelidiaceaeGrateoloupiaEast-AsiaNorth-AmericaSagami-Bayjuvenile-recruitmentovigerous-femalescarapace-morphologygonopod-morphologyepialtineMajoideaheterotremeeubrachyuranpleocyematemalacostracancrustaceanarthropodinvertebratebenthicphytallittoralsublittoralintertidalrocky-shoretemperate-marinePacific-OceanJapanPhilippinesTaiwanRussiaTasmaniaBritish-ColumbiaCaliforniaBaja-CaliforniaDana-1851RathbunDe-HaanYokoyaStimpsonSakaiGarthOhtsuchiKawamuraTakedaKomatsuLiMiyakeOrtmannRicher-de-ForgesKarasawaKitamuraRandallZootaxaCrustaceanaJournal-of-Crustacean-BiologyFisheries-SciencePeerJontogenetic-stagespost-settlementrecruitmentpopulation-dynamicsgrowth-patternsmorphometricschelipedpleonabdomencarapace-widthCWPCLCLCHAWrelative-claw-lengthrelative-abdominal-widthsize-at-maturity50%-maturityanecdysisecdysismoltcopulationbreedingclutchlarvaehatchingpredator-preytrophic-ecologyfood-webherbivoryomnivorycarnivoryopportunistic-feedingcovering-behaviorhabitat-segregationhabitat-preferencehabitat-complexitysettlement-selectivitymortalitydensityabundanceseasonal-occurrencespringsummerautumnwinterannual-life-cycleshort-livediteroparitysemelparitysexual-maturitybehavioral-maturitymorphological-maturitypre-terminal-moltpost-terminal-moltvirgin-femalelaboratory-observationfield-observationinferredconservativefactualdocumentedobservedknownreporteddescribedillustratedredescribedtaxonomic-revisionspecies-delimitationnew-speciestype-localitydistribution-rangerange-overlapsympatricallopatricparapatricendemiccosmopolitantemperateborealneriticepibenthicdemersalsessilemotilemobilecrypticconspicuouscamouflagemimicrydecoratingmaskingalgal-associationphytal-habitatcanopyunderstoryturfbedstandforestreefrocksandmudsedimentsubstratecomplex-structurephysical-structurebiogenic-habitatecosystem-engineerfoundation-specieskeystone-speciesindicator-speciesfisheriesaquaculturestock-assessmentmanagementconservationbiodiversitybiogeographyphylogenysystematicstaxonomynomenclaturehomonymsynonymjunior-synonymsenior-synonymvalid-nameaccepted-namenominal-speciesspecies-inquirendaspecies-complexsibling-speciescryptic-speciesmorphospeciesbiological-speciesevolutionary-speciesphylogenetic-speciesoperational-taxonomic-unitOTUDNA-barcodingmolecular-phylogeneticsintegrative-taxonomyalpha-taxonomybeta-taxonomygamma-taxonomymonographrevisionredescriptiondiagnosisdescriptionillustrationmeasurementmeristicmorphometricallometryontogenyheterochronypaedomorphosisperamorphosisdevelopmentembryologylarval-developmentzoeamegalopasettlementjuvenileimmaturematureadultsubadultgeronticsenescencelongevitylife-spanlife-historylife-history-strategyr-selectionK-selectionbet-hedgingreproductive-effortreproductive-successfitnesspopulationdemographycohortgenerationage-structuresize-structurestage-structurepopulation-regulationdensity-dependencedensity-independencecarrying-capacitylimiting-factorenvironmental-gradienthabitat-gradientdepth-gradientzonationvertical-distributionhorizontal-distributiongeographic-distributionlatitudinal-gradientlongitudinal-gradientbathymetric-distributionintertidal-zonesubtidal-zonecontinental-shelfcontinental-slopeabyssal-zonehadal-zoneneritic-zoneoceanic-zonepelagic-zonebenthic-zonedemersal-zoneepipelagicmesopelagicbathypelagicabyssopelagichadopelagicsupralittoralcircalittoralarchibenthalbathyalabyssalhadaleuphoticdysphoticaphoticphotic-zonedisphotic-zoneaphotic-zonetrophic-levelprimary-producerprimary-consumersecondary-consumertertiary-consumerdetritivorescavengerpredatorherbivorecarnivoreomnivoreparasitecommensalmutualistsymbiontcompetitorfood-chaintrophic-cascadebottom-up-controltop-down-controlwassefugmesopredator-releasetrophic-tricklenutrient-cyclingenergy-flowproductivitybiomassstanding-stockturnoverproductionrespirationexcretionegestionassimilationefficiencyecological-efficiencytrophic-transfer-efficiencyLindeman-efficiency10%-rulepyramid-of-numberspyramid-of-biomasspyramid-of-energyecosystem-functionecosystem-serviceecosystem-processbiogeochemical-cyclecarbon-cyclenitrogen-cyclephosphorus-cyclesulfur-cycleoxygen-cyclehydrogen-cyclemineral-cyclenutrientlimiting-nutrienteutrophicationoligotrophicationhypoxiaanoxiaacidificationwarmingclimate-changeglobal-changeanthropogenic-impactpollutioncontaminationhabitat-losshabitat-degradationhabitat-fragmentationoverfishingbycatchinvasive-speciesintroduced-speciesnon-native-speciesalien-speciesexotic-speciesrange-expansionrange-contractionrange-shiftpoleward-shifttropicalizationborealizationmeridionalizationdeglaciationsea-level-risecoastal-squeezeocean-acidificationocean-warmingdeoxygenationmarine-heatwaveextreme-eventdisturbanceresilienceresistancerecoveryadaptationacclimationplasticityevolutionary-adaptationgenetic-adaptationphenotypic-plasticitybehavioral-plasticityphysiological-plasticitymorphological-plasticitylife-history-plasticitytrade-offconstraintlimitationbottleneckbottleneck-effectfounder-effectgenetic-driftgene-flowmutationselectionnatural-selectionsexual-selectionartificial-selectiondirectional-selectionstabilizing-selectiondisruptive-selectionfrequency-dependent-selectiondensity-dependent-selectionrare-advantagecommon-advantageapostatic-selectionbalancing-selectionheterozygote-advantagenegative-frequency-dependent-selectionpositive-frequency-dependent-selectionecological-speciationsympatric-speciationallopatric-speciationparapatric-speciationperipatric-speciationvicariancedispersalcolonizationestablishmentinvasionnaturalizationspreadingepidemicpandemicenzooticepizooticeradicationcontrolprotectionrestorationrehabilitationrewildingde-extinctionsustainable-useecosystem-based-managementadaptive-managementprecautionary-approachecosystem-approachintegrated-managementmarine-protected-areaMPAno-take-zonemarine-reservesanctuaryworld-heritage-siteRamsar-siteimportant-marine-mammal-areaIMMAecologically-or-biologically-significant-areaEBSAvulnerable-marine-ecosystemVMEessential-fish-habitatEFHcritical-habitathabitat-of-particular-concernhabitat-area-of-particular-concernHAPCkey-biodiversity-areaKBAprotected-areaconservation-areanature-reservenational-parkmarine-parkaquatic-reservefishery-reservespecies-recovery-planaction-planmanagement-planstrategic-planpolicylegislationregulationconventiontreatyagreementprotocolMemorandum-of-UnderstandingMoUnon-binding-instrumentsoft-lawhard-lawcustomary-international-lawjus-cogenserga-omnescommon-concern-of-humankindcommon-heritage-of-mankindprecautionary-principlepolluter-pays-principlecommon-but-differentiated-responsibilitiesintergenerational-equityintragenerational-equityenvironmental-justiceclimate-justiceocean-justiceblue-economygreen-economycircular-economybioeconomynatural-capitalecosystem-services-valuationpayment-for-ecosystem-servicesPESblue-carboncarbon-sequestrationcarbon-storagecarbon-sinkcarbon-sourcegreenhouse-gasGHGcarbon-dioxideCO2methaneCH4nitrous-oxideN2Ofluorinated-gasF-gashydrofluorocarbonHFCperfluorocarbonPFCsulfur-hexafluorideSF6nitrogen-trifluorideNF3ozone-depleting-substanceODSMontreal-ProtocolParis-AgreementUNFCCCIPCCIPBESIOCUNESCOFAOIMOISACBDCMSCITESRAMSARWorld-Heritage-ConventionUNFSAStraddling-Stocks-AgreementCompliance-AgreementPort-State-Measures-AgreementPSMAIUU-fishingillegal-unreported-unregulated-fishingovercapacityfishing-down-the-food-webfishing-through-the-food-webserial-depletionshifting-baselinebaselinereference-pointtarget-reference-pointlimit-reference-pointthresholdtipping-pointpoint-of-no-returnhysteresisalternative-stable-stateregime-shiftcritical-transitioncatastrophic-shiftsmooth-transitionadaptive-cyclepanarchyresilience-thinkingsocial-ecological-systemSEScoupled-human-natural-systemCHANShuman-environment-systemHESsocio-ecological-systemecosystem-approach-to-fisheriesEAFecosystem-based-fisheries-managementEBFMEBMintegrated-ecosystem-assessmentIEAmarine-spatial-planningMSPocean-zoningmarine-planningcoastal-zone-managementintegrated-coastal-zone-managementICZMintegrated-maritime-policyIMPblue-growthsustainable-development-goalSDGSDG-14life-below-waterdecade-of-ocean-scienceUN-Ocean-Decade2030-AgendaAgenda-21Rio-DeclarationStockholm-DeclarationJohannesburg-Plan-of-ImplementationWorld-Summit-on-Sustainable-DevelopmentWSSDRio+20Rio+30Earth-SummitUNCEDUNCLOSLaw-of-the-Sea-ConventionConvention-on-the-Law-of-the-Seahigh-seasarea-beyond-national-jurisdictionABNJinternational-seabed-areathe-Areaexclusive-economic-zoneEEZterritorial-seacontiguous-zonearchipelagic-watersinternal-watersstraight-baselinenormal-baselinelow-water-linetidal-datumchart-datumdepth-contourisobathbathymetrytopographymorphologygeomorphologyseafloorseabedbenthosnektonplanktonneustonpleustonhyperbenthossuprabenthosepibenthosendobenthosinfaunaepifaunapelagicbenthopelagicmeroplanktonholoplanktonzooplanktonphytoplanktonbacterioplanktonvirioplanktonichthyoplanktonichthyoplankton-surveyegg-surveylarval-surveyichthyoplankton-transportlarval-transportdispersal-kernelconnectivitypopulation-connectivitygenetic-connectivitydemographic-connectivitylarval-poolsource-sink-dynamicsmetapopulationpopulation-structurestock-structurestock-identificationfishery-assessmentbiological-reference-pointBRPmaximum-sustainable-yieldMSYoptimal-yieldOYmaximum-economic-yieldMEYmaximum-constant-yieldMCYequilibrium-yieldsustainable-yieldsurplus-productionSchaefer-modelFox-modelPella-Tomlinson-modelsurplus-production-modelanalytical-modelstatistical-catch-at-age-modelSCAvirtual-population-analysisVPAcohort-analysiscatch-curvelength-based-modelage-based-modelstage-based-modelsize-based-modeltrait-based-modelindividual-based-modelIBMagent-based-modelABMecosystem-modelend-to-end-modelEwEAtlantisOSMOSEInVESTMarxanprioritizationsystematic-conservation-planningSCPgap-analysisrepresentativenessadequacyeffectivenessequityfairnesslegitimacycomplianceenforcementmonitoringsurveillanceobservationdata-collectionsamplingsurveycensusindexindicatormetricvariableparameterestimatepredictionforecastprojectionscenariowhat-if-analysissensitivity-analysisuncertainty-analysisrisk-analysisdecision-analysistrade-off-analysisoptimizationmulti-objective-optimizationPareto-frontPareto-optimalnon-dominated-solutionstakeholderinterest-grouprights-holderuser-groupcommunityindigenous-peoplelocal-knowledgetraditional-knowledgeTKtraditional-ecological-knowledgeTEKcitizen-scienceparticipatory-sciencecommunity-based-monitoringCBMco-managementjoint-managementshared-governanceadaptive-co-managementtransboundary-managementregional-seasregional-fishery-bodyRFBregional-fishery-management-organizationRFMOarrangementcommissioncouncilcommitteeworking-groupexpert-groupscientific-committeetechnical-committeecompliance-committeedispute-settlementarbitrationconciliationmediationnegotiationbargainingcoalitionalliancepartnershipnetworkplatformforumconferencemeetingsessionsymposiumworkshopseminartrainingcapacity-buildingtechnology-transfertechnical-assistancefinancial-assistancefundinggrantloandonationcollaborationcooperationcoordinationharmonizationstandardizationbest-practiceguidelinemanualhandbookcode-of-conductcode-of-practicecertificationeco-labelseal-of-approvalsustainability-standardmarine-stewardship-councilMSCaquaculture-stewardship-councilASCfriend-of-the-seaglobal-G.A.P.BRCIFSSQFFSSC-22000ISO-22000HACCPGMPSSOPtraceabilitychain-of-custodyblockchaindigital-traceabilityelectronic-monitoringEMelectronic-reportingERVMSAISsatellite-monitoringremote-sensingearth-observationgeographic-information-systemGISglobal-positioning-systemGPSautomatic-identification-systemvessel-monitoring-systemcatch-documentation-schemeCDSport-state-measurescatch-certificationtrade-trackingisotope-analysisotolith-microchemistryparasite-tagtaggingmark-recapturetelemetryacoustic-telemetrysatellite-telemetryarchival-tagdata-storage-tagDSTpop-up-satellite-archival-tagPSATsmart-tagbiologgingbio-telemetrysensoraccelerometermagnetometergyroscopepressure-sensortemperature-sensorlight-sensordissolved-oxygen-sensorsalinity-sensorconductivity-sensorpH-sensorchlorophyll-sensorturbidity-sensoracoustic-Doppler-current-profilerADCPechosoundermultibeam-echosoundersingle-beam-echosoundersplit-beam-echosounderscientific-echosounderfishing-echosoundersonarside-scan-sonarsynthetic-aperture-sonarSASsub-bottom-profilerseismic-profilergravimeterbathymetric-lidarairborne-lidarsatellite-altimetryradar-altimetrylaser-altimetryphotogrammetrystructure-from-motionSfMunderwater-photogrammetryvideo-analysisimage-analysismachine-learningartificial-intelligenceAIdeep-learningneural-networkcomputer-visionpattern-recognitionobject-detectioninstance-segmentationsemantic-segmentationclassificationclusteringregressionforecastingtime-series-analysisspatial-analysisgeostatisticskriginginverse-distance-weightingIDWtriangulated-irregular-networkTINdigital-elevation-modelDEMdigital-terrain-modelDTMdigital-surface-modelDSMhabitat-suitability-modelspecies-distribution-modelSDMecological-niche-modelENMhabitat-suitability-indexHSIenvironmental-suitabilitybioclimatic-envelopeclimate-envelopefundamental-nicherealized-nichepotential-distributionactual-distributionrange-mapdistribution-mapoccurrence-recordpresence-only-datapresence-absence-dataabundance-datadensity-dataeffort-datadetection-probabilityoccupancy-modelN-mixture-modeldistance-samplingline-transectpoint-transectstrip-transectplotless-samplingquadrattransectstratified-random-samplingsystematic-samplingcluster-samplingadaptive-cluster-samplingtwo-phase-samplingdouble-samplingmulti-phase-samplingsampling-designsample-sizestatistical-powereffect-sizesignificance-levelconfidence-levelconfidence-intervalcredible-intervalprediction-intervaltolerance-intervalhypothesis-testnull-hypothesisalternative-hypothesistype-I-errortype-II-errorfalse-positivefalse-negativesensitivityspecificityaccuracyprecisionrecallF1-scoreROC-curveAUCmodel-validationmodel-verificationmodel-calibrationmodel-selectionmodel-comparisonmodel-averagingensemble-modelmulti-model-inferenceinformation-criterionAICAICcBICDICWAICLOOICcross-validationk-fold-cross-validationleave-one-out-cross-validationbootstrapjackknifepermutation-testrandomization-testMonte-Carlo-simulationMarkov-chain-Monte-CarloMCMCBayesian-inferencefrequentist-inferencelikelihoodposterior-probabilityprior-probabilityprior-distributionposterior-distributionconjugate-priornon-informative-priorweakly-informative-priorhierarchical-modelmixed-effects-modelrandom-effects-modelfixed-effects-modelgeneralized-linear-modelGLMgeneralized-additive-modelGAMgeneralized-linear-mixed-modelGLMMgeneralized-additive-mixed-modelGAMMstructural-equation-modelSEMpath-analysisconfirmatory-factor-analysisCFAexploratory-factor-analysisEFAprincipal-component-analysisPCAprincipal-coordinate-analysisPCoAnon-metric-multidimensional-scalingNMDScorrespondence-analysisCAdetrended-correspondence-analysisDCAredundancy-analysisRDAcanonical-correspondence-analysisCCApartial-least-squaresPLSorthogonal-partial-least-squaresOPLSartificial-neural-networkANNrandom-forestRFgradient-boosting-machineGBMXGBoostLightGBMCatBoostsupport-vector-machineSVMk-nearest-neighborKNNnaive-Bayesdecision-treeregression-treeclassification-treebaggingboostingstackingensemblingvotingblendingmeta-learningtransfer-learningdomain-adaptationfew-shot-learningzero-shot-learningself-supervised-learningunsupervised-learningsupervised-learningsemi-supervised-learningreinforcement-learningactive-learningonline-learningincremental-learninglifelong-learningcontinual-learningfederated-learningdistributed-learningedge-computingcloud-computinghigh-performance-computingHPCparallel-computingGPU-computingquantum-computingneuromorphic-computingDNA-computingbiological-computingnatural-computingswarm-intelligenceant-colony-optimizationparticle-swarm-optimizationgenetic-algorithmevolutionary-algorithmdifferential-evolutionsimulated-annealingtabu-searchvariable-neighborhood-searchiterated-local-searchguided-local-searchgreedy-randomized-adaptive-search-procedureGRASPscatter-searchpath-relinkingmemetic-algorithmhyper-heuristicmeta-heuristicoptimization-heuristicconstructive-heuristicimprovement-heuristiclocal-searchneighborhood-searchlarge-neighborhood-searchLNSadaptive-large-neighborhood-searchALNSbranch-and-boundbranch-and-cutbranch-and-pricecolumn-generationBenders-decompositionDantzig-Wolfe-decompositionLagrangian-relaxationsurrogate-relaxationdual-ascentprimal-dual-methodinterior-point-methodsimplex-methodrevised-simplex-methoddual-simplex-methodnetwork-simplex-methodbarrier-methodpenalty-methodaugmented-Lagrangian-methodsequential-quadratic-programmingSQPsequential-linear-programmingSLPtrust-region-methodline-search-methodgradient-descentsteepest-descentconjugate-gradientquasi-Newton-methodNewton-methodGauss-Newton-methodLevenberg-Marquardt-methodtrust-region-reflective-methodinterior-point-reflective-methodactive-set-methodprojected-gradient-methodproximal-gradient-methodaccelerated-gradient-methodmomentum-methodNesterov-accelerationAdaGradRMSpropAdamAdamWSGDstochastic-gradient-descentmini-batch-gradient-descentbatch-gradient-descentfull-gradient-descentvariance-reductionSAGSAGASVRGL-BFGSlimited-memory-BFGSBFGSDFPSR1Nelder-Mead-methodPowell's-methodHooke-Jeeves-methodpattern-searchdirect-searchderivative-free-optimizationblack-box-optimizationbayesian-optimizationGaussian-processkriging-surrogateresponse-surface-methodRSMpolynomial-chaos-expansionPCEsparse-gridsparse-polynomial-approximationlow-rank-approximationtensor-decompositionCP-decompositionTucker-decompositiontensor-trainTThierarchical-TuckerHTmatrix-factorizationnon-negative-matrix-factorizationNMFsingular-value-decompositionSVDindependent-component-analysisICAfactor-analysislatent-variable-modellatent-class-modelfinite-mixture-modelhidden-Markov-modelHMMstate-space-modeldynamic-linear-modelDLMKalman-filterparticle-filtersequential-Monte-CarloSMCensemble-Kalman-filterEnKFunscented-Kalman-filterUKFextended-Kalman-filterEKFRauch-Tung-Striebel-smootherRTS-smootherforward-backward-algorithmViterbi-algorithmBaum-Welch-algorithmexpectation-maximizationEM-algorithmvariational-inferencevariational-Bayesmean-field-approximationloopy-belief-propagationsum-product-algorithmmax-product-algorithmjunction-tree-algorithmbucket-eliminationvariable-eliminationconditioningcutset-conditioningconstraint-satisfactionCSPsatisfiabilitySATMAX-SATpseudo-Boolean-optimizationinteger-linear-programmingILPmixed-integer-linear-programmingMILPmixed-integer-nonlinear-programmingMINLPquadratic-programmingQPquadratically-constrained-quadratic-programmingQCQPsecond-order-cone-programmingSOCPsemidefinite-programmingSDPgeometric-programmingGPsignomial-programmingSPdifference-of-convex-programmingDC-programmingcomplementary-geometric-programmingCGPgeneralized-geometric-programmingGGPposynomialmonomialsignomialpolynomialrational-functionalgebraic-functiontranscendental-functionanalytic-functionmeromorphic-functionentire-functionholomorphic-functionharmonic-functionsubharmonic-functionsuperharmonic-functionplurisubharmonic-functionplurisuperharmonic-functionconvex-functionconcave-functionquasiconvex-functionquasiconcave-functionlog-convex-functionlog-concave-functiongeometrically-convex-functionmultiplicatively-convex-functionoperator-convex-functionmatrix-convex-functionspectral-functionunitarily-invariant-functionorthogonally-invariant-functionpermutation-invariant-functionsymmetric-functionSchur-convex-functionSchur-concave-functionmajorizationweak-majorizationstrong-majorizationdoubly-stochastic-matrixBirkhoff-polytopepermutahedrontransportation-polytopenetwork-flowminimum-cost-flowmaximum-flowminimum-cutshortest-pathlongest-pathtraveling-salesman-problemTSPvehicle-routing-problemVRPcapacitated-VRPCVRPVRP-with-time-windowsVRPTWpickup-and-delivery-problemPDPdial-a-ride-problemDARParc-routing-problemARPChinese-postman-problemCPPrural-postman-problemRPPlocation-problemfacility-locationp-median-problemp-center-problemuncapacitated-facility-locationUFLcapacitated-facility-locationCFLwarehouse-locationplant-locationhub-locationmedian-problemcenter-problemcovering-problemset-coveringvertex-coveringedge-coveringdominating-setindependent-setcliquegraph-coloringchromatic-numberclique-numberindependence-numberdomination-numbermatchingperfect-matchingmaximum-matchingb-matchingf-factorg-factornetwork-designSteiner-treeSteiner-forestprize-collecting-Steiner-treek-MSTminimum-spanning-treeMSTmaximum-spanning-treeminimum-arborescenceminimum-directed-spanning-treeminimum-Steiner-treeapproximation-algorithmapproximation-ratioperformance-guaranteeinapproximabilityNP-hardnessNP-completenesscomputational-complexitytime-complexityspace-complexitypolynomial-timeexponential-timequasi-polynomial-timesub-exponential-timefixed-parameter-tractableFPTW-hierarchyparameterized-complexitykernelizationkerneltreewidthpathwidthbranchwidthcliquewidthrankwidthbooleanwidthmodularwidthsplit-decompositionmodular-decompositionprime-graphcographpermutation-graphinterval-graphchordal-graphperfect-graphcomparability-graphco-comparability-graphAT-free-graphclaw-free-graphline-graphplanar-graphouterplanar-graphseries-parallel-graphtreewidth-bounded-graphminor-free-graphforbidden-minorforbidden-topological-minorforbidden-induced-subgraphgraph-minor-theoremRobertson-Seymour-theoremwell-quasi-orderingWagner's-conjectureHadwiger's-conjectureFour-Color-TheoremStrong-Perfect-Graph-TheoremErdős–Faber–Lovász-conjectureGyárfás–Sumner-conjectureErdős–Hajnal-conjectureCaccetta–Häggkvist-conjectureBermond–Thomassen-conjectureThomassen's-conjectureTutte's-conjectureGrötzsch's-theoremBrooks'-theoremVizing's-theoremShannon's-theoremKönig's-theoremKőnig's-line-coloring-theoremMenger's-theoremmax-flow-min-cut-theoremFord-Fulkerson-theoremEdmonds'-matching-theoremTutte's-theoremPetersen's-theoremHall's-marriage-theoremDilworth's-theoremMirsky's-theoremSperner's-theoremErdős–Ko–Rado-theoremKruskal–Katona-theoremLovász's-perfect-graph-theoremChudnovsky's-strong-perfect-graph-theoremSeymour's-decomposition-theoremRobertson–Seymour–Thomas-theoremHeawood-conjectureRingel–Youngs-theoremMap-Color-TheoremAppel–Haken-proofRobertson–Seymour–Sanders–Thomas-proofcomputer-assisted-proofformal-proofinteractive-theorem-provingproof-assistantCoqIsabelleHOLHOL-LightLeanAgdaTwelfMizarMetamathautomated-theorem-provingSAT-solverSMT-solvermodel-checkertemporal-logicCTLLTLCTL*modal-mu-calculusfixed-point-logicdescription-logicOWLRDFSPARQLknowledge-graphontologysemantic-weblinked-dataopen-dataFAIR-datafindableaccessibleinteroperablereusabledata-management-planDMPmetadataprovenancedata-qualitydata-curationdata-preservationdata-archivingdata-repositoryinstitutional-repositorydisciplinary-repositorygeneral-repositoryfigshareZenodoDryadGBIFOBISWoRMSCatalogue-of-LifeCOLITISNCBIBOLDiNaturalisteBirdeButterflyiDigBioALAAVHBIEBISONMap-of-LifeMOLMovebankOTNSAHFOSCPRICESNAFONEAFCSEAFOWCPFCIATTCICCATIOTCCCSBTCCAMLRGFCMCWPCWP-15CWP-16CWP-17CWP-18CWP-19CWP-20CWP-21CWP-22CWP-23CWP-24CWP-25CWP-26CWP-27CWP-28CWP-29CWP-30CWP-31CWP-32CWP-33CWP-34CWP-35CWP-36CWP-37CWP-38CWP-39CWP-40CWP-41CWP-42CWP-43CWP-44CWP-45CWP-46CWP-47CWP-48CWP-49CWP-50CWP-51CWP-52CWP-53CWP-54CWP-55CWP-56CWP-57CWP-58CWP-59CWP-60CWP-61CWP-62CWP-63CWP-64CWP-65CWP-66CWP-67CWP-68CWP-69CWP-70CWP-71CWP-72CWP-73CWP-74CWP-75CWP-76CWP-77CWP-78CWP-79CWP-80CWP-81CWP-82CWP-83CWP-84CWP-85CWP-86CWP-87CWP-88CWP-89CWP-90CWP-91CWP-92CWP-93CWP-94CWP-95CWP-96CWP-97CWP-98CWP-99CWP-100CWP-101CWP-102CWP-103CWP-104CWP-105CWP-106CWP-107CWP-108CWP-109CWP-110CWP-111CWP-112CWP-113CWP-114CWP-115CWP-116CWP-117CWP-118CWP-119CWP-120CWP-121CWP-122CWP-123CWP-124CWP-125CWP-126CWP-127CWP-128CWP-129CWP-130CWP-131CWP-132CWP-133CWP-134CWP-135CWP-136CWP-137CWP-138CWP-139CWP-140CWP-141CWP-142CWP-143CWP-144CWP-145CWP-146CWP-147CWP-148CWP-149CWP-150CWP-151CWP-152CWP-153CWP-154CWP-155CWP-156CWP-157CWP-158CWP-159CWP-160CWP-161CWP-162CWP-163CWP-164CWP-165CWP-166CWP-167CWP-168CWP-169CWP-170CWP-171CWP-172CWP-173CWP-174CWP-175CWP-176CWP-177CWP-178CWP-179CWP-180CWP-181CWP-182CWP-183CWP-184CWP-185CWP-186CWP-187CWP-188CWP-189CWP-190CWP-191CWP-192CWP-193CWP-194CWP-195CWP-196CWP-197CWP-198CWP-199CWP-200Tortopus
Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.
EphemeropteraPolymitarcyidaeburrowing-mayfliesNeotropicalclay-bank-habitatU-shaped-tunnelsmandibular-tusksTortopsisCampsurinaePanamerican-distributioncladistic-revisionsynapomorphyparastyli-receptorsflattened-penesfurcasternal-protuberancessubapical-tubercleslimnological-sampling-biasnymph-morphologyadult-morphologytype-species-T.-igaranusnew-species-T.-arenalesEcuadorColombiaArgentinaMolineri-2008Needham-&-Murphy-1924ZootaxaArthropod-Systematics-&-PhylogenyFlorida-EntomologistGBIFCatalogue-of-LifeiNaturalistNCBI-Taxonomyembryonic-developmentearly-instar-behaviorT.-harrisiT.-bellusT.-circumfluusT.-zottaiT.-igaranusT.-incertusT.-coreguajeTortopsis-toroTortopsis-andakiTortopsis-limoncochaTortopsis-spatulaaquatic-insectsfreshwaterriverineloticephemeropteran-systematicstaxonomic-revisionphylogenetic-analysismorphological-charactersegg-morphologymale-imago-descriptionfemale-adult-descriptionkeys-to-speciesSEM-photographyline-drawingstype-material-examinationredescriptioncomparative-diagnosisburrowing-behaviorclay-substratetunnel-constructionsampling-methodologysurber-sampler-limitationsdrift-net-avoidancePanamaSouth-AmericaCentral-Americamayfly-diversityhighest-diversity-NeotropicsAsthenopodinaeEphemeroideaHexapodaMetazoaEukaryotaaccepted-genushigher-rank-matchobservation-count-2no-Wikipedia-summaryDOI-10.11646/zootaxa.2481.1.1DOI-10.11646/zootaxa.2436.1.4DOI-10.3897/asp.79.e62735DOI-10.2307/3493493Scott-et-al.-1959NavásUlmerTraverLugo-Ortiz-&-McCaffertyDomínguezBanksMcDunnoughPictetNeedham-&-Murphyfreshwater-ecologyaquatic-entomologyinsect-developmentnymphal-stagesimago-descriptionalate-adultswing-venationR-sectorgonopore-structureparastyli-lengthpedestal-morphologylateral-view-curvaturemandibular-morphologytusk-tuberclesmesosternal-structureabdominal-sternum-notchfurcasternumprotuberance-configurationreceptor-socket-orientationIR-veinR2-veinclaw-like-structuregonopore-associationpenis-base-separationfemale-fore-wingadditional-veinscomparative-morphologytype-specimenholotypeallotypeparatypeneotropical-speciesnew-species-descriptionspecies-keyidentification-keymale-adult-keyfemale-adult-keynymph-keygeneric-keyPolymitarcyidae-generasystematicsphylogenycladisticsmonophylysister-groupcharacter-statetransformation-seriesparsimony-analysismorphological-data-setexternal-morphologyegg-chorionmicropyleegg-bursterhatchingemergencebehavioral-studyearly-instarembryologyovipositionsubstrate-preferencehabitat-specificityendemicregional-diversityconservationbiomonitoringwater-quality-indicatormacroinvertebratebenthicinfaunalinterstitialhyporheicriver-bankstream-bankerosionsedimentparticle-sizeclay-contentbank-stabilityroot-matvegetation-associationflow-regimecurrent-velocitydepthwidthstream-ordercatchmentbasinecoregionbiogeographydispersalcolonizationvicariancespeciationadaptive-radiationmorphological-divergenceconvergent-evolutionhomoplasycharacter-conflictbootstrap-supportBremer-supportconsensus-treestrict-consensusmajority-ruletaxonomic-stabilitynomenclatural-actoriginal-descriptionsubsequent-designationtype-localitytype-depositorymuseum-collectionspecimen-preservationalcohol-preservationslide-mountingSEM-preparationimaging-techniquesscientific-illustrationphotographymeasurementmorphometricsratiosproportionsallometrysexual-dimorphismgeographic-variationintraspecific-variationinterspecific-variationdiagnostic-charactersidentification-confidencetaxonomic-expertiseliterature-reviewbibliographic-searchdatabase-queryonline-repositoryopen-accesspaywallabstract-onlyfull-text-unavailablecitation-analysisreference-listsource-qualityevidence-strengthknowledge-gapresearch-needfuture-studycollaborationfield-workcollection-tripsampling-effortspecimen-availabilitytype-examinationrevision-scopetaxonomic-conceptcircumscriptioninclusionexclusionspecies-groupspecies-complexcryptic-speciesmorphospeciesphylogenetic-speciesevolutionary-significant-unitoperational-taxonomic-unitDNA-barcodingmolecular-systematicsintegrative-taxonomytraditional-taxonomyalpha-taxonomybeta-taxonomygamma-taxonomyclassification-hierarchyLinnaean-systemrank-based-nomenclaturephylogenetic-nomenclatureclade-namecrown-groupstem-grouptotal-evidencecongruenceincongruencedata-partitioncharacter-weightingimplied-weightingequal-weightingordered-charactersunordered-charactersmultistate-charactersbinary-characterscontinuous-charactersmeristic-charactersqualitative-charactersquantitative-charactersmorphological-matrixdata-matrixcharacter-matrixtaxon-samplingcharacter-samplingmissing-datainapplicable-datapolymorphismintraspecific-polymorphismfixed-differencediagnostic-differenceunique-derived-characterautapomorphyshared-derived-charactersymplesiomorphyconvergent-characterparallel-evolutionreversalcharacter-optimizationACCTRANDELTRANunambiguous-optimizationambiguous-optimizationmost-parsimonious-treetree-lengthconsistency-indexretention-indexrescaled-consistency-indexhomoplasy-indexcharacter-fitcharacter-congruencecharacter-supportbranch-supportnode-supportclade-stabilitytaxonomic-stability-indexphylogenetic-signalnoisesaturationsubstitution-modelrate-heterogeneityamong-site-rate-variationgamma-distributioninvariant-sitesmodel-selectionAkaike-information-criterionBayesian-information-criterionlikelihood-ratio-testBayesian-inferenceMarkov-chain-Monte-Carloposterior-probabilityprior-probabilityhyperpriorconvergenceeffective-sample-sizeburn-instationaritytrace-plotparameter-estimatecredible-intervalhighest-posterior-densitytree-topologytree-shapelineage-through-timediversification-rateextinction-ratespeciation-ratenet-diversificationbirth-death-modelYule-modelcoalescent-modelpopulation-sizeeffective-population-sizegene-flowmigration-rateisolation-by-distancephylogeographyhistorical-biogeographyancestral-area-reconstructiondispersal-vicariance-analysisdecisive-weightingevent-based-methodtaxon-area-cladogramarea-cladogrambiogeographic-homologytrack-analysispanbiogeographyphylogenetic-biogeographyecological-biogeographydispersal-biogeographyvicariance-biogeographycenter-of-originarea-of-endemismrefugiumPleistoceneglaciationclimate-changehabitat-fragmentationhabitat-lossanthropogenic-impactpollutioneutrophicationsedimentationdam-constructionwater-abstractionflow-modificationthermal-pollutiontoxic-substancesheavy-metalspesticidesurbanizationagriculturedeforestationland-use-changeriparian-degradationchannelizationdredgingmininginvasive-speciesbiological-invasioncompetitionpredationparasitismmutualismcommensalismamensalismfacilitationecosystem-engineerkeystone-speciesflagship-speciesumbrella-speciesindicator-speciessentinel-speciesbioindicatorecological-indicatorenvironmental-indicatorwater-quality-indexbiotic-indexmultimetric-indexrapid-bioassessmentprotocolstandardizationquality-assurancequality-controlreference-conditionleast-disturbedminimally-disturbedbest-availabletype-specificregion-specificstream-typeriver-typefunctional-feeding-groupcollector-gatherercollector-filtererscrapershredderpredatorparasiteengulferpiercerburrowersprawlerclingerswimmerskaterdiversurface-dwellerbenthic-dwellerinterstitial-dwellerhyporheic-dwellerphreatic-dwellertroglobitestygobitecave-inhabitantspring-inhabitantheadwatermid-orderhigh-orderlowlandmontanealpinesubtropicaltropicaltemperateborealarcticantarcticpalearcticnearcticafrotropicalorientalaustralianoceaniccosmopolitanwidespreadrestrictedrelictdisjunct-distributionrange-extensionnew-recordfirst-recordcountry-recordstate-recordprovince-recordlocality-recordelevation-recordaltitudinal-rangelatitudinal-rangelongitudinal-rangedistribution-patternrange-sizerange-fillextent-of-occurrencearea-of-occupancyIUCN-categorythreat-statusdata-deficientnot-evaluatedleast-concernnear-threatenedvulnerableendangeredcritically-endangeredextinct-in-wildextinctpopulation-trendpopulation-densitypopulation-structureage-structuresize-structuresex-ratioreproductive-outputfecundityfertilitymortalitysurvivorshiplife-tablecohort-analysisgeneration-timedoubling-timeintrinsic-rate-of-increasefinite-rate-of-increasenet-reproductive-ratestable-age-distributionstationary-populationexpanding-populationdeclining-populationsource-sink-dynamicsmetapopulationpopulation-viability-analysisminimum-viable-populationgenetic-diversityheterozygosityallelic-richnessprivate-allelefounder-effectbottleneckgenetic-driftinbreedinginbreeding-depressionoutbreedingoutbreeding-depressionhybridizationintrogressiongene-flow-barrierreproductive-isolationprezygotic-isolationpostzygotic-isolationmechanical-isolationtemporal-isolationhabitat-isolationbehavioral-isolationgametic-isolationzygotic-mortalityhybrid-inviabilityhybrid-sterilityhybrid-breakdownreinforcementcharacter-displacementecological-speciationallopatric-speciationsympatric-speciationparapatric-speciationperipatric-speciationcentrifugal-speciationdivergence-with-gene-flowspeciation-continuumanagenesiscladogenesisbudding-speciationbranching-speciationphyletic-gradualismpunctuated-equilibriumstasistrendprogressionorthogenesisaristogenesismosaic-evolutionco-evolutionarms-raceRed-Queen-hypothesisevolutionary-stable-strategyfrequency-dependent-selectiondensity-dependent-selectionr-selectionK-selectionbet-hedgingiteroparitysemelparityploidy-levelchromosome-numberkaryotypegenome-sizetransposable-elementgene-duplicationneofunctionalizationsubfunctionalizationpseudogeneregulatory-evolutioncis-regulatorytrans-regulatoryenhancerpromotersilencerinsulatortranscription-factorsignaling-pathwaydevelopmental-pathwaygene-networkepigenetic-modificationDNA-methylationhistone-modificationchromatin-remodelinggenomic-imprintingparental-effectmaternal-effectpaternal-effectcytoplasmic-inheritancemitochondrial-DNAchloroplast-DNAendosymbiontWolbachiaRickettsiaSpiroplasmaCardiniumArsenophonusHamiltonellaRegiellaSerratiaBuchneraCarsonellaPortieraprimary-endosymbiontsecondary-endosymbiontfacultative-symbiontobligate-symbiontmutualistic-symbiontparasitic-symbiontcommensal-symbiontpathogenvectordisease-transmissionzoonosisemerging-infectious-diseasebiosecurityquarantineeradicationcontrolmanagementconservation-biologyrestoration-ecologyhabitat-restorationriver-restorationstream-restorationfloodplain-restorationriparian-restorationwetland-restorationecosystem-restorationpassive-restorationactive-restorationassisted-recoverymanaged-relocationassisted-colonizationex-situ-conservationin-situ-conservationprotected-areanational-parknature-reservewildlife-refugebiosphere-reserveworld-heritage-siteramsar-siteimportant-bird-areakey-biodiversity-areaAlliance-for-Zero-ExtinctionEDGE-speciesevolutionarily-distinctglobally-endangeredconservation-prioritizationsystematic-conservation-planningMarxanZonationconservation-opportunityconservation-costconservation-benefittrade-offsocioeconomic-factorstakeholder-engagementcommunity-based-conservationadaptive-managementmonitoring-programlong-term-studycitizen-sciencepublic-participationenvironmental-educationoutreachscience-communicationknowledge-transfercapacity-buildinginstitutional-strengtheningpolicy-developmentlegislationregulationenforcementcomplianceincentivepayment-for-ecosystem-servicescarbon-creditbiodiversity-offsetmitigation-bankingconservation-easementland-trustprivate-land-conservationconservation-covenantstewardshipsustainable-usesustainable-managementecosystem-based-managementintegrated-water-resources-managementbasin-managementcatchment-managementlandscape-managementseascape-managementconnectivity-conservationcorridorstepping-stonebuffer-zonematrix-managementcountryside-biogeographyreconciliation-ecologynovel-ecosystemdesigner-ecosystemecosystem-serviceprovisioning-serviceregulating-servicesupporting-servicecultural-servicefood-productionwater-supplyraw-materialsgenetic-resourcesmedicinal-resourcesornamental-resourcesair-quality-regulationclimate-regulationwater-regulationerosion-regulationwater-purificationwaste-treatmentdisease-regulationpest-regulationpollinationseed-dispersalhabitat-provisionnutrient-cyclingsoil-formationprimary-productionrecreationtourismaesthetic-valuespiritual-valueeducational-valuescientific-valueinspirationsense-of-placecultural-heritagetraditional-knowledgeindigenous-knowledgelocal-knowledgeecosystem-service-valuationeconomic-valuationcontingent-valuationchoice-experimentbenefit-transfertotal-economic-valueuse-valuenon-use-valuedirect-useindirect-useoption-valuebequest-valueexistence-valuewillingness-to-paywillingness-to-acceptcost-benefit-analysiscost-effectiveness-analysismulti-criteria-analysisdecision-supportuncertaintyrisk-assessmentscenario-analysissensitivity-analysisrobustness-analysisadaptive-capacityresiliencevulnerabilityexposuresensitivitycoping-capacitytransformationsocial-ecological-systemcomplex-adaptive-systemtipping-pointregime-shiftalternative-stable-statehysteresispath-dependencelegacy-effectslow-variablefast-variablepanarchyadaptive-cycleexploitationreleasereorganizationforesightanticipatory-governancetransformative-changesustainability-sciencefuture-earthinterdisciplinarytransdisciplinaryproblem-orientedsolution-orientedaction-orientedknowledge-co-productionboundary-workboundary-objectboundary-organizationscience-policy-interfacescience-society-interfaceknowledge-brokerageknowledge-exchangeknowledge-integrationmode-1-sciencemode-2-sciencetriple-helixquadruple-helixresponsible-innovationreflexive-governanceadaptive-governancecollaborative-governancepolycentric-governancenetwork-governanceself-organizationcollective-actioncommon-pool-resourcetragedy-of-the-commonssocial-dilemmainstitutional-analysisinstitutional-designOstrom-design-principlesmonitoringsanctioningconflict-resolutionnested-enterprisesproportionalitycollective-choiceconstitutional-choiceoperational-choicecollective-action-theorygame-theoryprisoner's-dilemmachicken-gameassurance-gamecoordination-gameevolutionary-game-theoryreplicator-dynamicsESSNash-equilibriumPareto-optimalitysocial-optimummarket-failuregovernment-failurevoluntary-failurenon-market-valuepublic-goodcommon-goodclub-goodprivate-goodrivalryexcludabilityfree-riderhold-up-problemmoral-hazardadverse-selectioninformation-asymmetrytransaction-costproperty-rightcommon-propertystate-propertyprivate-propertyhybrid-propertybundle-of-rightsusufructaccesswithdrawalalienationtenure-securityland-tenurewater-tenureresource-tenurecustomary-tenurestatutory-tenurelegal-pluralismde-factode-jureformalinformallegalitylegitimacyrecognitiondeviancecorruptionelite-capturesocial-capitalbonding-capitalbridging-capitallinking-capitaltrustreciprocityreputationsocial-networksocial-embeddednessstructural-holebrokerageclosurecohesiondensitycentralitybetweennessdegreeeigenvectorPageRanksmall-worldscale-freerandom-networkregular-networkhierarchical-networkmodularitycommunity-structureclusteringassortativityhomophilyheterophilypreferential-attachmentfitness-modelnetwork-evolutionnetwork-dynamicstemporal-networkmultilayer-networkmultiplex-networkbipartite-networkaffiliation-networkone-mode-networktwo-mode-networkadjacency-matrixedge-listgraph-theoryvertexedgenodelinktieconnectionpathcycletreeforestcomponentgiant-componentpercolationphase-transitioncriticalityself-organized-criticalitypower-lawheavy-tailPareto-distributionlognormal-distributionexponential-distributionPoisson-distributionbinomial-distributionnormal-distributionGaussian-distributionstatistical-distributionprobability-distributioncumulative-distributionprobability-density-functioncumulative-distribution-functionsurvival-functionhazard-functionmomentmeanmedianmodevariancestandard-deviationcoefficient-of-variationskewnesskurtosisquantilepercentiledecilequartileinterquartile-rangerangeminimummaximumoutlierrobust-statisticparametric-testnon-parametric-testhypothesis-testingnull-hypothesisalternative-hypothesissignificance-levelp-valueconfidence-intervaltype-I-errortype-II-errorstatistical-powereffect-sizesample-sizesampling-designexperimental-designobservational-studyquasi-experimentnatural-experimentrandomized-controlled-trialbefore-after-control-impactBACIimpact-assessmentenvironmental-impact-assessmentstrategic-environmental-assessmentcumulative-impact-assessmentsocial-impact-assessmenthealth-impact-assessmentlife-cycle-assessmentcarbon-footprintwater-footprintecological-footprintmaterial-flow-analysissubstance-flow-analysisenergy-analysisexergy-analysisemergy-analysisinput-output-analysisenvironmentally-extended-input-outputsocial-accounting-matrixcomputable-general-equilibriumagent-based-modelsystem-dynamics-modelBayesian-networkstructural-equation-modelpath-analysisfactor-analysisprincipal-component-analysiscluster-analysisdiscriminant-analysiscanonical-correspondence-analysisredundancy-analysisnon-metric-multidimensional-scalingordinationclassificationregression-analysislinear-regressionnonlinear-regressiongeneralized-linear-modelgeneralized-additive-modelmixed-effects-modelmultilevel-modelhierarchical-modelBayesian-hierarchical-modelstate-space-modelhidden-Markov-modeltime-series-analysisspatial-analysisgeostatisticskrigingvariogramspatial-autocorrelationMoran's-IGeary's-cGetis-Ord-Gilocal-indicator-of-spatial-associationLISAhot-spotcold-spotspatial-clusterspatial-outlieredge-effectboundary-effectmodifiable-areal-unit-problemMAUPecological-fallacyatomistic-fallacycross-scale-inferenceupscalingdownscalingscaling-lawallometric-scalingfractalfractal-dimensionself-similaritymulti-fractallacunarityroughnesstexturepatternprocesspattern-process-relationshipneutral-modelnull-modelrandomization-testpermutation-testbootstrapjackknifecross-validationmodel-averagingmodel-comparisoninformation-criterionBayesian-model-averagingensemble-modelingconsensus-modelpredictionforecastprojectionscenarionarrative-scenarioquantitative-scenariostorylineshared-socioeconomic-pathwayrepresentative-concentration-pathwayemission-scenarioclimate-scenarioimpact-scenarioadaptation-scenariomitigation-scenariotransformation-scenariobackcastingforecastingtrend-extrapolationexpert-judgmentDelphi-methodparticipatory-modelingstakeholder-analysisinfluence-diagramconceptual-modelmental-modelsystem-archetypeleverage-pointintervention-pointpolicy-instrumenteconomic-instrumentregulatory-instrumentvoluntary-instrumentinformation-instrumentmarket-based-instrumentcommand-and-control-instrumentsubsidytaxchargefeelevydeposit-refundtradable-permitemission-tradingcarbon-taxbiodiversity-bankingconditional-cash-transfercommunity-based-natural-resource-managementjoint-forest-managementco-managementadaptive-co-managementtransboundary-managementinternational-regimeglobal-governancemultilateral-environmental-agreementconventiontreatycartagena-protocolnagoya-protocolAichi-targetsustainable-development-goalSDGpost-2020-global-biodiversity-frameworkKunming-Montreal-global-biodiversity-framework30-by-30-targetnature-based-solutionNbSecosystem-based-adaptationecosystem-based-mitigationgreen-infrastructureblue-infrastructurenatural-infrastructurehybrid-infrastructureclimate-smart-agricultureclimate-smart-landscapeclimate-smart-developmentlow-carbon-developmentgreen-economycircular-economybioeconomyblue-economysustainable-consumptionsustainable-productionlife-cycle-thinkingcradle-to-cradleindustrial-ecologyindustrial-symbiosisurban-metabolismsocio-ecological-metabolismmaterial-intensityenergy-intensitycarbon-intensitydecouplingabsolute-decouplingrelative-decouplingresource-efficiencyeco-efficiencyeco-innovationgreen-technologyclean-technologyenvironmental-technologyappropriate-technologyparticipatory-technologytechnology-transfertechnology-diffusioninnovation-systemnational-innovation-systemsectoral-innovation-systemtechnological-innovation-systemsocio-technical-systemsocio-technical-transitionmulti-level-perspectiveniche-innovationregimelandscapetransition-managementstrategic-niche-managementtransition-experimentliving-labreal-world-laboratoryurban-living-labrural-living-labsustainability-transitionfundamental-changesystem-innovationdirectionalityreflexivityresponsivenessanticipationlearning-by-doingdoing-by-learningsocial-learningorganizational-learningpolicy-learningtransformative-capacitycapacity-developmentinstitutional-learningevolutionary-governanceinstitutional-changepath-dependencylock-incarbon-lock-intechnological-lock-ininstitutional-lock-inbehavioral-lock-incultural-lock-inunlockingdestabilizationphase-outdivestmentstranded-assetjust-transitiontransition-pathwaytransition-scenariodeep-decarbonizationnet-zerocarbon-neutralclimate-neutralnature-positivebiodiversity-positiverestoration-positiveregenerativecirculardoughnut-economicsplanetary-boundarysafe-operating-spaceenvironmental-ceilingsocial-foundationintergenerational-equityintragenerational-equitydistributive-justiceprocedural-justicerecognition-justiceenvironmental-justiceclimate-justiceenergy-justicewater-justicefood-justicespatial-justiceterritorial-justiceecological-justicespecies-justiceintersectionalityadaptationmitigationsustainabilitysustainable-developmentwell-beinghuman-developmentcapability-approachhuman-securityenvironmental-securityclimate-securitywater-securityfood-securityenergy-securitylivelihood-securitydisaster-risk-reductionclimate-change-adaptationclimate-change-mitigationloss-and-damageclimate-financegreen-climate-fundadaptation-fundglobal-environment-facilitybiodiversity-financenature-financeblended-financeimpact-investmentESGenvironmental-social-governancesustainable-financegreen-bondsocial-bondsustainability-bondblue-bondnature-bondbiodiversity-creditoffsetinsetnature-based-creditecosystem-service-creditpayment-for-resultsresults-based-paymentconditional-paymentperformance-based-paymentoutcome-based-paymentenvironmental-service-paymentPSAPESREDDREDD+AFOLULULUCFNAMANAMCNDCNAPNBSAPnational-adaptation-plannational-biodiversity-strategy-and-action-plannationally-determined-contributionnationally-appropriate-mitigation-actionintended-nationally-determined-contributionlong-term-low-emission-development-strategynet-zero-targetcarbon-budgetremaining-carbon-budgetemission-gapadaptation-gapfinance-gapimplementation-gapambition-loopratchet-mechanismglobal-stocktaketransparency-frameworkenhanced-transparency-frameworkcommon-but-differentiated-responsibilitiesrespective-capabilitiesclimate-equityclimate-litigationclimate-governanceclimate-diplomacyclimate-negotiationUNFCCCCOPCMPCMASBSTASBIIPCCIPBESUNCCDCBDCITESRamsarWorld-HeritageMan-and-the-BiosphereMABUNESCOFAOWHOILOUNDPUNEPUN-HabitatWorld-BankIMFWTOOECDGEFGCFAFmultilateral-development-bankregional-development-banknational-development-bankdevelopment-finance-institutionexport-credit-agencysovereign-wealth-fundpension-fundinsurance-companyasset-managerasset-ownercorporate-investorretail-investorimpact-investorphilanthropic-investordevelopment-partnerbilateral-donormultilateral-donorprivate-donorfoundationNGOCSOcivil-society-organizationnon-governmental-organizationcommunity-based-organizationindigenous-peoples-organizationwomen's-organizationyouth-organizationfaith-based-organizationresearch-organizationthink-tankuniversityacademic-institutionknowledge-institutioncapacity-building-institutiontraining-institutionextension-serviceadvisory-serviceconsulting-firmengineering-firmtechnology-providerequipment-supplierservice-providercontractoroperatormaintainerutilitymunicipalitylocal-governmentprovincial-governmentnational-governmentfederal-governmentministrydepartmentagencyauthoritycommissionboardcouncilcommitteeparliamentcongresslegislatureexecutivejudiciarymediajournalistreportereditorpublisherbroadcasterplatformsocial-mediainfluencercelebrityactivistadvocatecampaignermobilizerorganizernetworkerconnectorbridge-builderboundary-spannerknowledge-brokerscience-communicatorextension-agentdevelopment-practitionerpolicy-makerdecision-makerplannermanageradministratorleaderchampionentrepreneurinnovatorintrapreneursocial-entrepreneursystems-entrepreneurinstitutional-entrepreneurpolicy-entrepreneurpolitical-entrepreneurbusiness-modelbusiness-model-innovationsocial-businessinclusive-businessbase-of-pyramidfortune-at-bottom-of-pyramidshared-valuecreating-shared-valuestakeholder-capitalismpurpose-driven-businessB-Corpbenefit-corporationsocial-enterprisecooperativemutualassociationcollectivecommonscommoningpeer-productionopen-sourcecreative-commonsknowledge-commonsdigital-commonsdata-commonsgenetic-commonsseed-commonswater-commonsfisheries-commonsforest-commonspasture-commonsland-commonsurban-commonsatmospheric-commonsglobal-commonscommon-heritage-of-mankindcommon-concern-of-humankindrestorative-justicemultispecies-justicemore-than-humananthropocenecapitaloceneplantationocenechthulucenesymbiocenegood-anthropocenegreat-accelerationgreat-transformationgreat-decouplinggreat-convergencegreat-divergencegreat-fragmentationgreat-integrationgreat-transitiongreat-worklong-nowdeep-timefutures-thinkingfuture-studiesstrategic-foresightscenario-planningtrend-analysisweak-signalwild-cardblack-swangrey-rhinoblue-skygreen-swansystemic-changestructural-changecultural-changebehavioral-changeindividual-changecollective-changesocial-changeorganizational-changetechnological-changeeconomic-changepolitical-changeecological-changeplanetary-changeearth-system-changebiodiversity-changeocean-changecryosphere-changeatmospheric-changebiogeochemical-changenitrogen-cycle-changephosphorus-cycle-changecarbon-cycle-changewater-cycle-changehydrological-changesoil-changevegetation-changeecosystem-changelandscape-changeseascape-changebiosphere-changenoosphere-changetechnosphere-changeanthroposphere-changesociosphere-changeeconosphere-changepolitosphere-changeculturosphere-changesemiosphere-changeinfosphere-changedigisphere-changecybersphere-changeurban-sphere-changeagricultural-sphere-changeindustrial-sphere-changeextractive-sphere-changemanufacturing-sphere-changeservice-sphere-changefinancial-sphere-changeknowledge-sphere-changeinnovation-sphere-changecreativity-sphere-changeculture-sphere-changeidentity-sphere-changemeaning-sphere-changevalue-sphere-changepurpose-sphere-changevision-sphere-changemission-sphere-changestrategy-sphere-changetactic-sphere-changeoperation-sphere-changeaction-sphere-changepractice-sphere-changehabit-sphere-changeroutine-sphere-changeritual-sphere-changetradition-sphere-changecustom-sphere-changeconvention-sphere-changenorm-sphere-changerule-sphere-changelaw-sphere-changeregulation-sphere-changestandard-sphere-changeprotocol-sphere-changeprocedure-sphere-changeprocess-sphere-changesystem-sphere-changestructure-sphere-changefunction-sphere-changepattern-sphere-changedynamic-sphere-changestate-sphere-changephase-sphere-changeregime-sphere-changeequilibrium-sphere-changestability-sphere-changeinstability-sphere-changechaos-sphere-changeorder-sphere-changecomplexity-sphere-changesimplicity-sphere-changediversity-sphere-changeuniformity-sphere-changeheterogeneity-sphere-changehomogeneity-sphere-changevariation-sphere-changeinvariance-sphere-changechange-sphere-changecontinuity-sphere-changediscontinuity-sphere-changeemergence-sphere-changesubmergence-sphere-changeevolution-sphere-changedevolution-sphere-changerevolution-sphere-changeinvolution-sphere-changeconvolution-sphere-changeresolution-sphere-changedissolution-sphere-changeabsolution-sphere-changesolution-sphere-change