Soft-scale

Guides

Ceroplastes rusci
Fig Wax Scale
Ceroplastes rusci, commonly known as the fig wax scale, is a soft scale insect (family Coccidae) native to the Mediterranean region. First discovered in Florida in 1994-1995, it has since spread to numerous countries across six continents and is recognized as a significant agricultural and forestry pest. The species feeds on more than 94 genera from 52 plant families, with notable impacts on figs, citrus, grapes, and ornamental plants. Climate change modeling predicts expanding suitable habitat ranges through 2100, increasing its invasive potential in temperate regions.
scale-insect agricultural-pest invasive-species phloem-feeder parthenogenetic virus-vector climate-change fig-pest citrus-pest grape-pest ornamental-pest honeydew-producer sooty-mold soft-scale Coccidae
Coccus longulus
Long Brown Scale
Coccus longulus, commonly known as the long brown scale, is a soft scale insect in the family Coccidae. It has a broad global distribution spanning Europe, Africa, Asia, the Americas, and numerous Pacific islands. As a member of the soft scales, it likely feeds on plant sap and may produce honeydew, though specific biological details remain poorly documented in accessible literature.
soft-scale Coccidae cosmopolitan-distribution sap-feeder
Coccus viridis
green scale, green coffee scale, coffee green scale
Coccus viridis is a soft scale insect (Hemiptera: Coccidae) and major agricultural pest with a wide tropical and subtropical distribution. It reproduces parthenogenetically via thelytoky, with females producing 50-600 eggs that hatch within minutes to hours beneath the mother's body. The species completes its life cycle in 47-51 days at 25°C, with three nymphal instars before adulthood; first-instar nymphs actively disperse as 'crawlers' while older instars and adults become sessile. It is polyphagous, feeding on phloem sap of over 200 plant genera in 72 families, with primary economic damage to coffee, guava, and cacao crops.
agricultural-pest coffee-pest soft-scale parthenogenesis thelytoky polyphagy ant-mutualism honeydew sooty-mold biological-control invasive-species tropical-pest phloem-feeder Hemiptera Coccidae Coccus green-scale green-coffee-scale
Eucalymnatus
tessellated scale
Eucalymnatus is a genus of soft scale insects in the family Coccidae. The best-known species, Eucalymnatus tessellatus, is native to South America and has spread to tropical and subtropical regions worldwide. Species in this genus are phytophagous and can become significant pests of ornamental plants, fruit trees, and palms in nurseries and landscapes.
scale-insect soft-scale pest ornamental-pest tropical-pest Hemiptera Coccidae phytophagous
Microterys
Microterys is a large genus of parasitoid wasps in the family Encyrtidae (Chalcidoidea), with its center of distribution in the northern parts of the Northern Hemisphere. Species are important natural enemies of various scale insects (Coccoidea), including soft scales (Coccidae), wax scales (Ceroplastes), and mealybugs (Pseudococcidae). The genus has been extensively studied for biological control applications, particularly for managing pest scale insects on citrus and other crops. Several species have been introduced to new regions as biocontrol agents, including Microterys flavus in California.
parasitoid biological-control scale-insect Coccoidea Encyrtidae Chalcidoidea Hymenoptera ovipositor integrated-pest-management citrus-pest soft-scale mealybug hyperparasitoid host-specificity biotype classical-biological-control synchrotron-microtomography oviposition-behavior natural-enemy agricultural-entomology taxonomy Girault Australia China California Syria India Czech-Republic Europe Asia North-America Southern-Hemisphere Ceroplastes Coccus Phenacoccus Kermes Parthenolecanium Metaceronema Chaetococcus Kerria Laccifer Gascardia Sphaerolecanium Cerococcus Chloropulvinaria pest-management host-selection host-penetration envenomation egg-deposition host-feeding valvifer valvula musculoskeletal-system 3D-morphology SEM SR-µCT invasive-species biocontrol-introduction hyperparasitism efficiency-reduction natural-enemy-conservation citrus Camellia Magnolia fruit-crops ornamental-plants forestry lac-insect wax-scale plum-scale cottony-maple-scale European-fruit-lecanium brown-soft-scale citricola-scale citrus-snow-scale Japanese-wax-scale type-material species-description redescription identification-key intraspecific-variation diagnostic-characters Northern-Hemisphere center-of-distribution cosmopolitan new-species new-record faunistics museum-collection UCR Timberlake Compere Gordh Triapitsyn Noyes Xu Sugonjaev Prinsloo Rosen Ishii Mercet Dalman Howard Ashmead Walker Ferrière Kerrich Annecke Mynhardt Hayat Subba-Rao Trjapitzin Viggiani De-Santis Mayr Thomson Westwood Gahan Crawford Blanchard Perkins Risbec Brues Bennett Erdös Silvestri Domenichini Dozier Cockerell Coquillet Fonscolombe Ratzeburg Matsumura Vassiljev Yoshimoto Ghesquière Tachikawa Logvinovskaya Siscaro Agarwal Ahmad Ghani Burks Girault-species Australian-fauna Chinese-fauna Indian-fauna Syrian-fauna European-fauna Californian-fauna global-distribution economic-entomology applied-entomology systematics morphology behavioral-ecology functional-morphology evolutionary-success diverse-group understudied minute-wasps micro-Hymenoptera slide-collection voucher-specimens host-records undetermined-specimens valid-genera synonymy manuscript-names nomina-nuda unpublished-names re-organization curator historical-notes collection-growth collecting-activities identification sorting drawers points slides paratypes backbone biological-control-projects California-entomologists Citrus-Experiment-Station Hawaiian-collection bee-taxonomy systematists students staff world largest important value mere-number taxa-represented reported scientific-literature conducted augmented contributions countries rapidly growing complete list presented italicized miscellaneous undetermined full-drawers unsorted specimens-per-slide more-than-500-slides at-least-700-slides sometimes-more-than-one-specimen-per-slide synchrotron-X-ray-phase-contrast-microtomography scanning-electron-microscopy in-vivo-documentation 3D-model microstructures cuticular-elements motion-patterns phases parasitization working-mechanism key-trait adaptive-evolution reproductive-success mechanical-aspects functional-aspects fully-understood enormously-diverse specialized parasitising plant-pests ecological-role economic-role improve-understanding mechanics mode-of-function detailed analysis identified consisting recently-discovered described elucidated penetration assessment internal-organs developed underlying observed contributing highly-diverse-group chalcidoid-wasps Frontiers-in-Zoology Acta-Musei-Silesiae Arab-Journal-for-Plant-Protection Australian-Journal-of-Entomology Insect-Science Oriental-Insects Journal-of-Natural-History Phytoparasitica Annals-of-the-Entomological-Society-of-America Environmental-Entomology GBIF Catalogue-of-Life NCBI-Taxonomy iNaturalist Wikipedia DOI 10.1186/s12983-025-00575-1 10.2478/cszma-2021-0007 10.22268/AJPP-41.3.292305 10.1111/j.1440-6055.1975.tb02058.x 10.1111/j.1744-7917.2000.tb00345.x 10.1080/00305316.2011.646827 10.1111/j.1440-6055.1973.tb01670.x 10.1080/00222933.2020.1786608 10.1007/s12600-019-00727-0 10.1007/s12600-018-0691-5 10.1093/aesa/54.2.222 10.1093/ee/7.6.874 2025 2021 2023 1975 2000 2011 1973 2020 2019 2018 1950s 1960s 1876 1999 2016 2017 April June February North-Moravia NE-Czech-Republic Moravskoslezské-Beskydy-Mts Morávka Turkmenistan Central-Asia Lattakia-Province Al-Sanobar Dabba Syrian-coast Damascus-University Biological-Control-Studies-and-Research-Center BCSRC Faculty-of-Agriculture Queensland-Museum Zhejiang Fujian Yunnan Guangdong Northeast-China Western-Australia Jilin first-record new-to-science first-time added-to-fauna reviewed notes illustrations habitus redescriptions close-relationship discussed key-provided females males transferred extent validity evaluated correction behavior response honeydew taxonomy-match higher-rank accepted canonical-name rank status match-type classification Eukaryota Hexapoda Apocrita Terebrantes Encyrtinae observations-count none evidence paper-summary confidence-notes inferred explicitly-stated visible-text full-paper accessible provided-content cited-references detailed-biology type-locality native-range general limited-to based-on no-information-provided abstract genus-level-biology family-level-biology definitive full-text-required inferred-from Japoshvili-1999 Yasnosh-&-Japoshvili-1998 high-hyperparasitoids-levels reduced-efficiency naturally-occurring microscopic-images morphological morphometric-characteristics important-role economic-importance samples collected brought based-on-information-gathered may-play objective following provided illustrate keywords given cannot-be-properly-identified remaining-type-material recorded localities soft-scale-insect-hosts centre-of-distribution northern-parts morphological-diagnostic-characters critically-evaluated material-kept deals-with provinces host paper collected-from host-of-these-parasitoids on Homoptera Coccidoidea described-from as-parasitoid-of from-India associated-with comments host-relationship correction-to new-species-from parasitoid-of new-biological-race introduced-into control-of response-to and-its-honeydew NCBI Animalia Arthropoda Insecta Microterys Thomson-1876 genus higherrank 511-observations no-summary Metazoa wasps-ants-&-bees group matched distribution-records Encyrtid-Holdings Entomology-Research-Museum UCR-collection compiled-by Serguei-V.-Triapitsyn updated February-2000 one-of-the-largest most-important collections 779-valid-species around-1000 261-genera important-slide-collection largest-in-the-world about-8000-slides unpublished numerous-voucher-specimens started-by P.H.-Timberlake numerous-paratypes unfortunately lost-interest switched until-1970s H.-Compere notable-contributions P.-Timberlake J.-Maple P.-DeBach C.-Clausen 1970s-1980s G.-Gordh J.-LaSalle J.-Woolley J.-Pinto J.-Beardsley Mexico US other-countries presently growing-rapidly J.-Heraty majority V.-Trjapitzin J.-Noyes complete-re-organization S.-Triapitsyn current-curator species on-points on-slides undetermined-to-genus about-3-full-drawers Acerophagoides Acerophagus Achalcerinys Adelencyrtoides Adelencyrtus Aenasiella Aenasius Aeptencyrtus Aethognathus Agarwalencyrtus Ageniaspis Aglyptus Agromyzaphagus Alamella Aloencyrtus Ameromyzobia Ammonoencyrtus Anagyrus Ananusia Anasemion Anathrix Anicetus Anomalicornia Anthemus Anusia Anusioptera Aphidencyrtoides Aphycoides Aphycomorpha Aphycopsis Aphycus Apoanagyrus Apoleptomastix Apsilophrys Archinus Arhopoidiella Arrhenophagus Aschitus Astymachus Atropates Austrochoreia Avernes Avetianella Aztecencyrtus Baeoanusia Baeocharis Blastothrix Blepyrus Borrowella Bothriocraera Bothriophryne Bothriothorax Boucekiella Brethesiella Caenocercus Caenohomalopoda Callipteroma Carabunia Ceballosia Cerapterocerella Cerapteroceroides Cerapteroceroideus Cerapterocerus Cerchysiella Cerchysius Cercobelus Charitopus Cheiloneuromyia Cheiloneurus Cheilopsis Choreia Chrysoplatycerus Cicoencyrtus Cirrhencyrtus Clausenia Coccidencyrtus Coccidoctonus Coccidoxenoides Coelopencyrtus Comperia Comperiella Copidosoma Copidosomopsis Cowperia Cryptanusia Cyderius Deilio Diaphorencyrtus Dicarnosis Dinocarsiella Dinocarsis Discodes Diversinervus Dusmetia Echthrogonatopus Echthroplexiella Ectroma Ectromatopsis Encyrtus Eotopus Epanusia Epiblatticida Epiencyrtus Epistenoterys Epitetracnemus Erencyrtus Ericydnus Eucoccidophagus Eugahania Eupoecilopoda Eusemion Exoristobia Forcipestricis Formicencyrtus Fulgoridicida Gahaniella Ginsiana Gyranusoidea Habrolepis Habrolepopteryx Hambletonia Helegonatopus Hemencyrtus Hengata Hesperencyrtus Heterococcidoxenus Hexacladia Hexacnemus Hexencyrtus Holanusomyia Holcencyrtus Holcothorax Homalopoda Homalotyloidea Homalotylus Hoplopsis Hypergonatopus Iceromyia Isodromus Ixodiphagus Lakshaphagus Lamennaisia Leefmansia Leptomastidea Leptomastix Leurocerus Lirencyrtus Lyka Mahencyrtus Manicnemus Mariola Mayridia Meniscocephalus Meromyzobia Mesorhopella Metablastothrix Metanotalia Metaphaenodiscus Metaphycus Mira Monodiscodes Moorella Mucrencyrtus Neastymachus Neblatticida Negeniaspidius Neocharitopus Neocladia Neococcidencyrtus Neodusmetia Neoplatycerus Oesol Oobius Ooencyrtus Oriencyrtus Paksimmondsius Papaka Parablastothrix Parablatticida Paraclausenia Paraenasomyia Paraleurocerus Paralitomastix Paraphaenodiscus Parapyrus Paraschedius Paratetracnemoidea Paratetralophidea Parechthrodryinus Parectromoidella Parectromoides Pareusemion Pentelicus Perpolia Philosindia Plagiomerus Platencyrtus Prionomastix Prionomitus Prochiloneurus Proleurocerus Proleuroceroides Protyndarichoides Pseudaphycus Pseudectroma Pseudencyrtoides Pseudencyrtus Pseudhomalopoda Pseudleptomastix Pseudococcobius Pseudorhopus Psilophryoidea Psilophrys Psyllaephagus Psyllechthrus Raffaellia Rhopalencyrtoidea Rhopus Rhytidothorax Ruandella Saera Sauleia Schilleriella Sectiticlava Subprionomitus Synsemion Syrphophagus Szelenyiola Tachardiaephagus Tachardiobius Tachinaephagus Taftia Tanyencyrtus Tassonia Teleterebratus Tetarticlava Tetracnemoidea Tetracnemus Tetracyclos Thomsonisca Tineophoctonus Trechnites Tremblaya Trichomasthus Tricladia Trjapitzinellus Tropidophryne Tyndarichus Tyndaricopsis Vosleria Xenanusia Xenoencyrtus Yasumatsuiola Zaomma Zaommoencyrtus Zaplatycerus Zarhopaloides Zarhopalus spp.sp.sp(p).?nr.ms-name manuscript-name nomen-nudum including probably-mixed-with unsorted-miscellaneous University-of-California-Riverside collection museum specimens vouchers hosts parasitoids scale-insects mealybugs soft-scales wax-scales citrus-pests agricultural-pests natural-enemies hyperparasitoids efficiency oviposition anatomy 3D-reconstruction microtomography in-vivo-observation evolution adaptive-radiation diversity Africa South-America Middle-East Hawaii horticulture crop-protection augmentative-biological-control conservation-biological-control egg-laying population-dynamics ecosystem-services natural-pest-control sustainable-agriculture organic-farming biodiversity 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Neolecanium
Neolecanium is a genus of soft scale insects (family Coccidae) established by Parrott in 1901. The genus includes species that are pests of woody plants, notably magnolia scale (Neolecanium cornuparvum), which is monitored by the USA National Phenology Network for phenology-based pest management. Like other soft scales, members of this genus possess a waxy protective covering and feed on plant phloem. The genus is included in the Pheno Forecast program, which uses heat accumulation models to predict insect development and optimize control timing.
soft-scale pest woody-plants phenology integrated-pest-management North-America
Parthenolecanium
soft scales, lecanium scales
Parthenolecanium is a genus of soft scale insects in the family Coccidae. These hemipterans are dome-shaped, sap-feeding pests of woody plants including oaks, fruit trees, and grapevines. Several species are significant urban tree pests, notably P. quercifex (oak lecanium scale) and P. corni (European fruit lecanium scale). The genus exhibits variation in reproductive mode: some species reproduce by parthenogenesis, while P. corni reproduces sexually (gamogenesis). Many species have been documented as vectors of grapevine viruses.
scale-insect pest urban-entomology agricultural-pest vector vineyard-pest tree-pest soft-scale Hemiptera Coccidae
Parthenolecanium quercifex
oak lecanium scale
Parthenolecanium quercifex, commonly known as the oak lecanium scale, is a native soft scale insect in the family Coccidae. Adult females are hemispherical, 4–7 mm long, and light to dark brown or gray with lateral humps. The species is prevalent in the eastern United States, primarily infesting oak trees but occasionally found on hickory and birch. It has one annual generation, with eggs laid in late May to June and crawlers hatching in June to early July. Heavy infestations can cause stunted foliage, chlorosis, twig death, and dieback, while moderate populations may support beneficial natural enemies.
scale-insect urban-pest oak-pest soft-scale phloem-feeder honeydew-producer North-America Hemiptera Coccidae
Phalacrococcus
Phalacrococcus is a genus of soft scale insects (Hemiptera: Coccidae) established in 2010. The genus contains at least one exotic species, Phalacrococcus howertoni, which has been documented in Florida. As a member of the Coccidae family, these insects are sessile, sap-feeding pests that produce protective waxy coverings.
Coccidae soft-scale exotic Florida Hemiptera scale-insect
Phalacrococcus howertoni
Croton Scale
Phalacrococcus howertoni is a soft scale insect in the family Coccidae, described by Hodges and Hodgson in 2010. The species has been documented in Florida and is known by the common name Croton Scale. As a member of the Coccidae, it possesses the protective waxy covering characteristic of soft scales.
Coccidae soft-scale exotic-species Florida Hemiptera Croton-Scale
Prococcus acutissimus
banana-shaped scale
Prococcus acutissimus is a soft scale insect (family Coccidae) originally described from Sri Lanka in 1896. It is commonly known as the banana-shaped scale, reflecting its distinctive body form. The species has become widely distributed across tropical and subtropical regions, with records spanning the Indo-Pacific, Africa, the Caribbean, and North America. It is associated with diverse host plants and has been observed on agricultural crops, though specific economic impacts remain poorly documented.
Coccidae soft-scale banana-pest tropical-agriculture Indo-Pacific invasive-potential poorly-studied-pest
Pseudokermes
Pseudokermes is a genus of scale insects in the family Coccidae, first described by Cockerell in 1895. These insects are soft scales that feed on plant sap. The genus is part of the diverse Coccidae family, which contains many economically significant agricultural pests. Species within Pseudokermes are relatively poorly documented compared to other coccid genera.
scale-insect Coccidae soft-scale Sternorrhyncha sap-feeder
Pulvinaria acericola
Cottony Maple Leaf Scale
Pulvinaria acericola, commonly known as the Cottony Maple Leaf Scale, is a soft scale insect (family Coccidae) native to North America. Like other Pulvinaria species, the adult female produces a conspicuous white, cottony ovisac that houses eggs. The species is associated with maple and other woody plants, where nymphs and adults feed on phloem sap using piercing-sucking mouthparts. Heavy infestations can lead to honeydew accumulation and subsequent sooty mold growth.
scale-insect Coccidae soft-scale maple-pest ornamental-pest North-America
Pulvinaria floccifera
Cottony Camellia Scale, Cottony Yew Scale, Cottony Taxus Scale
Pulvinaria floccifera is a soft scale insect (Coccidae) known for producing conspicuous white, cottony ovisacs that cover eggs. It is a significant pest of woody ornamentals including camellia, holly, yew, and tea plants. The species has a broad global distribution spanning temperate and subtropical regions. Females are sessile and wingless, while males develop wings for mating. Heavy infestations produce honeydew that supports sooty mold growth, reducing plant vigor and aesthetic value.
scale-insect soft-scale pest ornamental tea-pest Coccidae Sternorrhyncha honeydew sooty-mold biological-control-host
Pulvinaria innumerabilis
Cottony Maple Scale
A soft scale insect (family Coccidae) commonly known as the cottony maple scale. Adult females are small, flattened, and brown, approximately 3 mm in length. Mature females produce conspicuous white, cottony ovisacs containing up to 1,500 eggs. The species is frequently found on maple trees, particularly silver maple (Acer saccharinum), but has been recorded on numerous other woody plants. Heavy infestations can cause aesthetic damage and minor physiological stress to host plants, though established trees typically tolerate infestations well.
soft-scale maple-pest ornamental-pest phloem-feeder honeydew-producer univoltine temperate North-American-native
Saissetia coffeae
hemispherical scale, helmet scale, coffee brown scale
Saissetia coffeae is a polyphagous soft scale insect (Coccidae) with a cosmopolitan tropical and subtropical distribution. Adults are sessile, hemispherical, and helmet-shaped, ranging 2–4.5 mm in diameter depending on host plant. The species reproduces parthenogenetically; all individuals are female. It is a significant agricultural pest of coffee, citrus, avocado, fig, guava, and numerous ornamental plants, causing damage through phloem sap feeding and honeydew secretion that promotes sooty mold growth. Multiple parasitoid wasps and predatory insects are associated with natural population regulation.
scale-insect soft-scale agricultural-pest polyphagous parthenogenetic biological-control honeydew-producer sooty-mold ant-mutualism coffee-pest citrus-pest tropical-pest
Toumeyella liriodendri
tuliptree scale
Toumeyella liriodendri, the tuliptree scale, is a soft scale insect (Coccidae) native to North America and specialized on Liriodendron tulipifera (tuliptree or yellow-poplar). This univoltine species overwinters as second-instar nymphs on twigs, matures to adults in spring, and produces eggs that hatch into crawlers in early summer. First-instar nymphs feed on leaf undersides before migrating back to twigs in autumn. Males are unknown; reproduction is presumed parthenogenetic. Heavy infestations cause branch dieback, reduced growth, and honeydew production that supports sooty mold growth. The species has been the subject of recent genomic research, with a 536Mb genome assembly revealing it as an early-diverging soft scale with 17 chromosomes.
scale-insect soft-scale Coccidae Liriodendron tuliptree yellow-poplar univoltine parthenogenesis honeydew sooty-mold forest-pest ornamental-pest genome chromosome-evolution phloem-feeder ant-tended North-America Maryland Hemiptera Sternorrhyncha
Toumeyella mirabilis
Toumeyella mirabilis is a soft scale insect in the family Coccidae, described by Cockerell in 1895. As a member of the genus Toumeyella, it shares characteristics with other soft scales that feed on plant vascular fluids and produce honeydew. The species occurs in North and Middle America, with records from the United States and Mexico. Like related species in the genus, it likely exhibits typical soft scale biology with sedentary adult females and mobile crawler nymphs.
Coccidae Hemiptera Sternorrhyncha scale-insect soft-scale North-America Mexico Cockerell-1895
Toumeyella pini
Striped Pine Scale
Toumeyella pini, commonly known as the Striped Pine Scale, is a soft scale insect native to North America. It is a significant pest of pine trees, particularly two- and three-needle pine species. The species exhibits pronounced sexual dimorphism in both morphology and feeding location: females are larger, highly convex, and develop on terminal twigs, while males are smaller, more elongate, and develop on needles. The species secretes copious honeydew, which attracts ants, wasps, bees, and flies, and can lead to sooty mold growth that inhibits photosynthesis. In southern Georgia, it completes three generations per year, with crawler activity peaking in May, mid-July to early August, and late October to early November.
scale-insect soft-scale pine-pest honeydew-producer Sternorrhyncha Coccidae North-America forest-pest ornamental-pest