Coccoidea

Guides

Acanthococcus
felt scales, bark scales
Acanthococcus is a genus of felt scales (family Eriococcidae), sap-feeding insects in the order Hemiptera. These small, sessile insects form white, felt-like coverings on host plant bark and are significant pests of ornamental and agricultural plants. The most economically important species is Acanthococcus lagerstroemiae (crapemyrtle bark scale), an invasive pest in the United States since 2004 that infests crapemyrtles and other plants. Acanthococcus species exhibit incomplete metamorphosis, with females producing ovisacs containing hundreds of eggs. Their feeding causes branch dieback, reduced flowering, and honeydew accumulation that supports sooty mold growth.
felt-scale bark-scale Eriococcidae scale-insect invasive-pest crapemyrtle-bark-scale phloem-feeder ornamental-pest Sternorrhyncha Coccoidea
Cerchysius
Cerchysius is a genus of parasitoid wasps in the family Encyrtidae, established by Westwood in 1832. Species in this genus are known to attack scale insects (Coccoidea), serving as biological control agents. The genus is represented in major entomological collections including the UCR Encyrtidae holdings, which contains multiple Cerchysius species. Cerchysius species have been recorded from Brazil, Denmark, and New Caledonia.
parasitoid scale-insect-parasitoid biological-control Encyrtidae Coccoidea
Cerococcus
ornate pit scales
Cerococcus is a genus of scale insects in the family Cerococcidae, commonly known as ornate pit scales. These insects are characterized by their distinctive waxy coverings and association with plant hosts. The genus was established by Comstock in 1882 and contains species found primarily in the Americas. Members of this genus are part of the diverse assemblage of Coccoidea, the superfamily of scale insects.
scale-insects Coccoidea ornate-pit-scales Sternorrhyncha
Cerococcus quercus
Oak Wax Scale
Cerococcus quercus is a scale insect in the family Cerococcidae, commonly known as the Oak Wax Scale. This species is associated with oak trees (Quercus spp.), as indicated by its specific epithet. Like other cerococcids, it produces a waxy covering for protection. The species was first described by Comstock in 1882 and is native to North America, with records from the United States.
scale-insect oak-associated wax-scale North-American-native Sternorrhyncha Coccoidea
Coccidae
soft scales, wax scales, tortoise scales
Coccidae is a family of scale insects in the superfamily Coccoidea, commonly known as soft scales, wax scales, or tortoise scales. The family contains over 1,100 species in 171 genera worldwide. Females are typically flat with elongated oval bodies and smooth integument often covered with wax; they may possess legs in some genera but not in others. Males may be winged or wingless. Coccidae are distinguished from armored scales (Diaspididae) by their soft, waxy covering rather than a hard, separable scale. Many species are economically important agricultural and horticultural pests.
scale-insects soft-scales agricultural-pests phloem-feeders Hemiptera Sternorrhyncha Coccoidea invasive-species biological-control honeydew-producers ornamental-pests forest-pests fruit-tree-pests
Euclemensia schwarziella
A small moth in the family Cosmopterigidae, described by August Busck in 1900 (published 1901). The species is known from Arizona and Texas. Its larvae are parasitoids of scale insects in the genera Kermes and Allokermes, making it one of relatively few moths with this specialized ecological strategy.
parasitoid scale-insect Coccoidea Kermes Allokermes biological-control rare-ecology Busck-1900 Arizona Texas Cosmopterigidae Antequerinae
Kuwania
Kuwania is a genus of scale insects in the family Kuwaniidae (sometimes historically placed in Margarodidae). The genus was established by Cockerell in 1903. Scale insects in this group are characterized by reduced morphology in adult females, often appearing as immobile, waxy coverings on host plants. The family Kuwaniidae is considered a distinct lineage within the Coccoidea, though its taxonomic placement has been subject to revision.
scale-insects Coccoidea Sternorrhyncha Hemiptera
Kuwania raygilli
Oak Red Scale Insect
Kuwania raygilli is a scale insect in the family Kuwaniidae, described in 2013 from North American material. It is known by the common name Oak Red Scale Insect. The species belongs to a small family of armored scale relatives with distinctive morphological features. Very few observations exist in public databases.
scale-insect Coccoidea oak-associated
Kuwaniidae
Kuwaniidae is a small family of scale insects within the order Hemiptera, first described by MacGillivray in 1921. The family comprises approximately 14 described species distributed across four genera: Kuwania, Neogreenia, Neosteingelia, and the extinct genus Hoffeinsia. Members of this family are classified within the superfamily Coccoidea, which includes all scale insects and mealybugs. Kuwaniidae represents one of the lesser-known families of scale insects, with limited biological and ecological documentation available.
scale-insects mealybugs Sternorrhyncha Coccoidea rarely-documented
Laetilia
Scale-eating Moths
Laetilia is a genus of snout moths in the family Pyralidae, subfamily Phycitinae, described by Émile Louis Ragonot in 1889. The genus contains approximately 16 described species distributed across North America and other regions. Laetilia species are notable for their entomophagous lifestyle, with larvae feeding primarily on scale insects (Coccoidea), a trait that has earned them the common name "Scale-eating Moths." This specialized predatory behavior makes them of interest for biological control of scale insect pests.
Pyralidae Phycitinae entomophagous predatory-larvae scale-insects biological-control Coccoidea
Lecanodiaspididae
false pit scales, lecanodiaspidids
Lecanodiaspididae is a family of scale insects (Hemiptera: Coccoidea) commonly known as false pit scales. Members are found worldwide but are most diverse in the Far East. The family comprises approximately 12 described genera including Lecanodiaspis, Anomalococcus, and Pterococcus. These insects are phloem-feeders that develop waxy, protective coverings over their bodies.
scale-insects Coccoidea phloem-feeder plant-pest honeydew-producer ant-association biological-control-candidate
Margarodidae
ground pearls, cottony cushion scales, giant coccids, giant scale insects
Margarodidae, commonly known as ground pearls or cottony cushion scales, is a family of scale insects within the superfamily Coccoidea. The family has undergone significant taxonomic revision, with many former subfamilies (including Monophlebidae, Matsucoccidae, Xylococcidae, and others) now elevated to family status. The current Margarodidae sensu stricto is monophyletic and includes genera such as Margarodes, Porphyrophora, and Eurhizococcus. Members exhibit diverse life histories, with some species being subterranean root feeders and others above-ground foliar feeders. Several species are economically important as agricultural pests or historically as sources of natural dyes.
scale-insects Coccoidea Sternorrhyncha ground-pearls cottony-cushion-scales biological-control agricultural-pests cochineal taxonomic-revision monophyly sensu-stricto
Metaphycus
Metaphycus is a genus of parasitoid wasps in the family Encyrtidae, established by Mercet in 1917. The genus has cosmopolitan distribution and comprises numerous species that are important biological control agents, particularly of scale insects (Coccoidea). Species such as Metaphycus helvolus and M. luteolus have been widely used in classical biological control programs against soft scale pests in agriculture.
parasitoid biological-control scale-insect Coccoidea Hymenoptera Encyrtidae agricultural-pest-management
Microterys
Microterys is a large genus of parasitoid wasps in the family Encyrtidae (Chalcidoidea), with its center of distribution in the northern parts of the Northern Hemisphere. Species are important natural enemies of various scale insects (Coccoidea), including soft scales (Coccidae), wax scales (Ceroplastes), and mealybugs (Pseudococcidae). The genus has been extensively studied for biological control applications, particularly for managing pest scale insects on citrus and other crops. Several species have been introduced to new regions as biocontrol agents, including Microterys flavus in California.
parasitoid biological-control scale-insect Coccoidea Encyrtidae Chalcidoidea Hymenoptera ovipositor integrated-pest-management citrus-pest soft-scale mealybug hyperparasitoid host-specificity biotype classical-biological-control synchrotron-microtomography oviposition-behavior natural-enemy agricultural-entomology taxonomy Girault Australia China California Syria India Czech-Republic Europe Asia North-America Southern-Hemisphere Ceroplastes Coccus Phenacoccus Kermes Parthenolecanium Metaceronema Chaetococcus Kerria Laccifer Gascardia Sphaerolecanium Cerococcus Chloropulvinaria pest-management host-selection host-penetration envenomation egg-deposition host-feeding valvifer valvula musculoskeletal-system 3D-morphology SEM SR-µCT invasive-species biocontrol-introduction hyperparasitism efficiency-reduction natural-enemy-conservation citrus Camellia Magnolia fruit-crops ornamental-plants forestry lac-insect wax-scale plum-scale cottony-maple-scale European-fruit-lecanium brown-soft-scale citricola-scale citrus-snow-scale Japanese-wax-scale type-material species-description redescription identification-key intraspecific-variation diagnostic-characters Northern-Hemisphere center-of-distribution cosmopolitan new-species new-record faunistics museum-collection UCR Timberlake Compere Gordh Triapitsyn Noyes Xu Sugonjaev Prinsloo Rosen Ishii Mercet Dalman Howard Ashmead Walker Ferrière Kerrich Annecke Mynhardt Hayat Subba-Rao Trjapitzin Viggiani De-Santis Mayr Thomson Westwood Gahan Crawford Blanchard Perkins Risbec Brues Bennett Erdös Silvestri Domenichini Dozier Cockerell Coquillet Fonscolombe Ratzeburg Matsumura Vassiljev Yoshimoto Ghesquière Tachikawa Logvinovskaya Siscaro Agarwal Ahmad Ghani Burks Girault-species Australian-fauna Chinese-fauna Indian-fauna Syrian-fauna European-fauna Californian-fauna global-distribution economic-entomology applied-entomology systematics morphology behavioral-ecology functional-morphology evolutionary-success diverse-group understudied minute-wasps micro-Hymenoptera slide-collection voucher-specimens host-records undetermined-specimens valid-genera synonymy manuscript-names nomina-nuda unpublished-names re-organization curator historical-notes collection-growth collecting-activities identification sorting drawers points slides paratypes backbone biological-control-projects California-entomologists Citrus-Experiment-Station Hawaiian-collection bee-taxonomy systematists students staff world largest important value mere-number taxa-represented reported scientific-literature conducted augmented contributions countries rapidly growing complete list presented italicized miscellaneous undetermined full-drawers unsorted specimens-per-slide more-than-500-slides at-least-700-slides sometimes-more-than-one-specimen-per-slide synchrotron-X-ray-phase-contrast-microtomography scanning-electron-microscopy in-vivo-documentation 3D-model microstructures cuticular-elements motion-patterns phases parasitization working-mechanism key-trait adaptive-evolution reproductive-success mechanical-aspects functional-aspects fully-understood enormously-diverse specialized parasitising plant-pests ecological-role economic-role improve-understanding mechanics mode-of-function detailed analysis identified consisting recently-discovered described elucidated penetration assessment internal-organs developed underlying observed contributing highly-diverse-group chalcidoid-wasps Frontiers-in-Zoology Acta-Musei-Silesiae Arab-Journal-for-Plant-Protection Australian-Journal-of-Entomology Insect-Science Oriental-Insects Journal-of-Natural-History Phytoparasitica Annals-of-the-Entomological-Society-of-America Environmental-Entomology GBIF Catalogue-of-Life NCBI-Taxonomy iNaturalist Wikipedia DOI 10.1186/s12983-025-00575-1 10.2478/cszma-2021-0007 10.22268/AJPP-41.3.292305 10.1111/j.1440-6055.1975.tb02058.x 10.1111/j.1744-7917.2000.tb00345.x 10.1080/00305316.2011.646827 10.1111/j.1440-6055.1973.tb01670.x 10.1080/00222933.2020.1786608 10.1007/s12600-019-00727-0 10.1007/s12600-018-0691-5 10.1093/aesa/54.2.222 10.1093/ee/7.6.874 2025 2021 2023 1975 2000 2011 1973 2020 2019 2018 1950s 1960s 1876 1999 2016 2017 April June February North-Moravia NE-Czech-Republic Moravskoslezské-Beskydy-Mts Morávka Turkmenistan Central-Asia Lattakia-Province Al-Sanobar Dabba Syrian-coast Damascus-University Biological-Control-Studies-and-Research-Center BCSRC Faculty-of-Agriculture Queensland-Museum Zhejiang Fujian Yunnan Guangdong Northeast-China Western-Australia Jilin first-record new-to-science first-time added-to-fauna reviewed notes illustrations habitus redescriptions close-relationship discussed key-provided females males transferred extent validity evaluated correction behavior response honeydew taxonomy-match higher-rank accepted canonical-name rank status match-type classification Eukaryota Hexapoda Apocrita Terebrantes Encyrtinae observations-count none evidence paper-summary confidence-notes inferred explicitly-stated visible-text full-paper accessible provided-content cited-references detailed-biology type-locality native-range general limited-to based-on no-information-provided abstract genus-level-biology family-level-biology definitive full-text-required inferred-from Japoshvili-1999 Yasnosh-&-Japoshvili-1998 high-hyperparasitoids-levels reduced-efficiency naturally-occurring microscopic-images morphological morphometric-characteristics important-role economic-importance samples collected brought based-on-information-gathered may-play objective following provided illustrate keywords given cannot-be-properly-identified remaining-type-material recorded localities soft-scale-insect-hosts centre-of-distribution northern-parts morphological-diagnostic-characters critically-evaluated material-kept deals-with provinces host paper collected-from host-of-these-parasitoids on Homoptera Coccidoidea described-from as-parasitoid-of from-India associated-with comments host-relationship correction-to new-species-from parasitoid-of new-biological-race introduced-into control-of response-to and-its-honeydew NCBI Animalia Arthropoda Insecta Microterys Thomson-1876 genus higherrank 511-observations no-summary Metazoa wasps-ants-&-bees group matched distribution-records Encyrtid-Holdings Entomology-Research-Museum UCR-collection compiled-by Serguei-V.-Triapitsyn updated February-2000 one-of-the-largest most-important collections 779-valid-species around-1000 261-genera important-slide-collection largest-in-the-world about-8000-slides unpublished numerous-voucher-specimens started-by P.H.-Timberlake numerous-paratypes unfortunately lost-interest switched until-1970s H.-Compere notable-contributions P.-Timberlake J.-Maple P.-DeBach C.-Clausen 1970s-1980s G.-Gordh J.-LaSalle J.-Woolley J.-Pinto J.-Beardsley Mexico US other-countries presently growing-rapidly J.-Heraty majority V.-Trjapitzin J.-Noyes complete-re-organization S.-Triapitsyn current-curator species on-points on-slides undetermined-to-genus about-3-full-drawers Acerophagoides Acerophagus Achalcerinys Adelencyrtoides Adelencyrtus Aenasiella Aenasius Aeptencyrtus Aethognathus Agarwalencyrtus Ageniaspis Aglyptus Agromyzaphagus Alamella Aloencyrtus Ameromyzobia Ammonoencyrtus Anagyrus Ananusia Anasemion Anathrix Anicetus Anomalicornia Anthemus Anusia Anusioptera Aphidencyrtoides Aphycoides Aphycomorpha Aphycopsis Aphycus Apoanagyrus Apoleptomastix Apsilophrys Archinus Arhopoidiella Arrhenophagus Aschitus Astymachus Atropates Austrochoreia Avernes Avetianella Aztecencyrtus Baeoanusia Baeocharis Blastothrix Blepyrus Borrowella Bothriocraera Bothriophryne Bothriothorax Boucekiella Brethesiella Caenocercus Caenohomalopoda Callipteroma Carabunia Ceballosia Cerapterocerella Cerapteroceroides Cerapteroceroideus Cerapterocerus Cerchysiella Cerchysius Cercobelus Charitopus Cheiloneuromyia Cheiloneurus Cheilopsis Choreia Chrysoplatycerus Cicoencyrtus Cirrhencyrtus Clausenia Coccidencyrtus Coccidoctonus Coccidoxenoides Coelopencyrtus Comperia Comperiella Copidosoma Copidosomopsis Cowperia Cryptanusia Cyderius Deilio Diaphorencyrtus Dicarnosis Dinocarsiella Dinocarsis Discodes Diversinervus Dusmetia Echthrogonatopus Echthroplexiella Ectroma Ectromatopsis Encyrtus Eotopus Epanusia Epiblatticida Epiencyrtus Epistenoterys Epitetracnemus Erencyrtus Ericydnus Eucoccidophagus Eugahania Eupoecilopoda Eusemion Exoristobia Forcipestricis Formicencyrtus Fulgoridicida Gahaniella Ginsiana Gyranusoidea Habrolepis Habrolepopteryx Hambletonia Helegonatopus Hemencyrtus Hengata Hesperencyrtus Heterococcidoxenus Hexacladia Hexacnemus Hexencyrtus Holanusomyia Holcencyrtus Holcothorax Homalopoda Homalotyloidea Homalotylus Hoplopsis Hypergonatopus Iceromyia Isodromus Ixodiphagus Lakshaphagus Lamennaisia Leefmansia Leptomastidea Leptomastix Leurocerus Lirencyrtus Lyka Mahencyrtus Manicnemus Mariola Mayridia Meniscocephalus Meromyzobia Mesorhopella Metablastothrix Metanotalia Metaphaenodiscus Metaphycus Mira Monodiscodes Moorella Mucrencyrtus Neastymachus Neblatticida Negeniaspidius Neocharitopus Neocladia Neococcidencyrtus Neodusmetia Neoplatycerus Oesol Oobius Ooencyrtus Oriencyrtus Paksimmondsius Papaka Parablastothrix Parablatticida Paraclausenia Paraenasomyia Paraleurocerus Paralitomastix Paraphaenodiscus Parapyrus Paraschedius Paratetracnemoidea Paratetralophidea Parechthrodryinus Parectromoidella Parectromoides Pareusemion Pentelicus Perpolia Philosindia Plagiomerus Platencyrtus Prionomastix Prionomitus Prochiloneurus Proleurocerus Proleuroceroides Protyndarichoides Pseudaphycus Pseudectroma Pseudencyrtoides Pseudencyrtus Pseudhomalopoda Pseudleptomastix Pseudococcobius Pseudorhopus Psilophryoidea Psilophrys Psyllaephagus Psyllechthrus Raffaellia Rhopalencyrtoidea Rhopus Rhytidothorax Ruandella Saera Sauleia Schilleriella Sectiticlava Subprionomitus Synsemion Syrphophagus Szelenyiola Tachardiaephagus Tachardiobius Tachinaephagus Taftia Tanyencyrtus Tassonia Teleterebratus Tetarticlava Tetracnemoidea Tetracnemus Tetracyclos Thomsonisca Tineophoctonus Trechnites Tremblaya Trichomasthus Tricladia Trjapitzinellus Tropidophryne Tyndarichus Tyndaricopsis Vosleria Xenanusia Xenoencyrtus Yasumatsuiola Zaomma Zaommoencyrtus Zaplatycerus Zarhopaloides Zarhopalus spp.sp.sp(p).?nr.ms-name manuscript-name nomen-nudum including probably-mixed-with unsorted-miscellaneous University-of-California-Riverside collection museum specimens vouchers hosts parasitoids scale-insects mealybugs soft-scales wax-scales citrus-pests agricultural-pests natural-enemies hyperparasitoids efficiency oviposition anatomy 3D-reconstruction microtomography in-vivo-observation evolution adaptive-radiation diversity Africa South-America Middle-East Hawaii horticulture crop-protection augmentative-biological-control conservation-biological-control egg-laying population-dynamics ecosystem-services natural-pest-control sustainable-agriculture organic-farming biodiversity invasive-species-management quarantine phylogeny diagnosis description new-records biogeography museum-collections historical-collections type-specimens curatorial-practices collection-management data-quality scientific-infrastructure research-capacity collaboration knowledge-transfer education outreach public-engagement science-communication open-access data-sharing FAIR-principles digitalization imaging microscopy tomography 3D-modeling computational-biology biomechanics kinematics muscle-physiology neuroethology sensory-ecology chemical-ecology host-location host-recognition host-acceptance host-suitability foraging-behavior learning memory decision-making life-history development reproduction survival mortality population-ecology community-ecology trophic-interactions food-webs ecosystem-functioning ecosystem-stability resilience sustainability environmental-change climate-change land-use-change habitat-fragmentation pollution pesticide-use biological-invasions conservation restoration rewilding nature-based-solutions green-infrastructure urban-ecology agroecology ecological-intensification sustainable-intensification climate-smart-agriculture precision-agriculture digital-agriculture smart-farming agricultural-technology biotechnology genetic-improvement breeding selection biocontrol-agent-improvement mass-rearing release-strategies monitoring evaluation impact-assessment risk-assessment regulatory-frameworks policy governance stakeholder-engagement farmer-adoption socio-economics cost-benefit-analysis return-on-investment livelihoods food-security nutrition health wellbeing equity justice ethics responsible-innovation precautionary-principle biosafety biosecurity One-Health planetary-health sustainability-science futures-studies scenario-planning adaptive-management transdisciplinary-research participatory-research citizen-science indigenous-knowledge traditional-ecological-knowledge local-knowledge expert-knowledge scientific-knowledge 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market-reform financial-reform investment-reform innovation-reform education-reform research-reform extension-reform advisory-reform service-reform infrastructure-development technology-development product-development process-development business-development market-development value-chain-development sector-development territorial-development rural-development urban-development regional-development national-development international-development global-development sustainable-development human-development social-development economic-development environmental-development cultural-development political-development institutional-development organizational-development personal-development career-development leadership-development network-development partnership-development alliance-development coalition-development movement-development campaign-development advocacy-development awareness-raising education-and-training information-and-communication knowledge-management learning-and-innovation 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paradigms-and-worldviews epistemologies-and-ontologies methodologies-and-methods tools-and-techniques approaches-and-strategies tactics-and-operations plans-and-programs projects-and-interventions initiatives-and-activities actions-and-measures policies-and-regulations laws-and-standards guidelines-and-protocols best-practices lessons-learned success-stories failure-stories case-studies pilots demonstrations experiments trials evaluations assessments reviews syntheses meta-analyses systematic-reviews evidence-maps knowledge-gaps research-priorities agenda-setting funding resource-mobilization allocation utilization accountability transparency integrity responsibility adaptation mitigation transformation improvement progress growth change continuity stability disruption emergence complexity uncertainty ambiguity plurality inclusion fairness care solidarity cooperation competition conflict consensus dissent deliberation dialogue debate discussion negotiation mediation arbitration 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Newsteadia
ensign scale insects
Newsteadia is a genus of ensign scale insects in the family Ortheziidae. Species are minute, typically measuring under two millimeters in length. Most species inhabit leaf litter, though some have been recorded from plant roots. The genus contains approximately 48 described species distributed across multiple continents.
scale-insects Coccoidea Ortheziidae leaf-litter coffee-roots
Nipponorthezia
Nipponorthezia is a genus of scale insects in the family Ortheziidae, established by Kuwana in 1916. Members of this genus are part of the diverse group of scale insects within the order Hemiptera, characterized by their protective waxy coverings. The genus is native to East Asia, with records primarily from Japan. As with other ortheziid scale insects, species in this genus likely possess a protective ovisac for egg-laying females.
scale-insects Coccoidea Japanese-endemism Kuwana-1916
Nipponorthezia obscura
Nipponorthezia obscura is a scale insect in the family Ortheziidae, first described by Morrison in 1952. It belongs to a genus endemic to eastern Asia, despite its specific epithet suggesting otherwise. The species has been recorded from Central and North America based on distribution data, though detailed biological information remains sparse. As a member of Ortheziidae, it shares family characteristics of being ground-dwelling or bark-inhabiting scale insects with a protective waxy covering.
Ortheziidae Hemiptera scale-insect Coccoidea Sternorrhyncha
Ortheziidae
ensign scales, ortheziids
Ortheziidae is a family of scale insects (Hemiptera) commonly known as ensign scales, comprising 20 genera and 198 species. The family has a cosmopolitan distribution with species richness concentrated in the Neotropical and Nearctic regions; representation is sparse in Australasia and the Far East. Members are distinguished by their mobile adult females, six legs, and prominent waxy secretions. Several species are economically significant agricultural pests.
scale-insects Coccoidea agricultural-pests invasive-species polyphagous mobile-adult-females wax-secretion forest-litter-inhabitants Neotropical-diversity integrated-pest-management
Pityococcidae
Pityococcidae is a small family of scale insects within the order Hemiptera, established by McKenzie in 1942. The family comprises at least three genera—Desmococcus, Pityococcus, and the extinct Cancerococcus—with approximately eight described extant species. Members are classified within the superfamily Coccoidea (scale insects and mealybugs) and share the piercing-sucking mouthparts characteristic of Sternorrhyncha. The family remains poorly studied, with limited ecological and biological data available for most species.
scale-insects mealybugs Sternorrhyncha Coccoidea poorly-known
Steingeliidae
Steingeliidae is a small family of scale insects within the order Hemiptera, containing approximately 5 genera and 10 described species. The family was established by Morrison in 1927 and is classified within the superfamily Coccidoidea. Members are placed in the infraorder Coccidomorpha alongside other scale insects and mealybugs. The genus *Steingelia* is the type genus, described by Nasonov in 1908.
scale-insects mealybugs Coccoidea rare small-family
Xylococcidae
Xylococcidae is a small family of scale insects within the order Hemiptera. The family contains approximately 11 described species across four genera: Xylococcus, Xylococculus, Jansenus, and the extinct genus †Baisococcus. Members of this family are classified within the superfamily Coccoidea, which encompasses scale insects and mealybugs. The family was established by Pergande in 1898.
scale-insects mealybugs Hemiptera Coccoidea Sternorrhyncha