Neotropical-diversity
Guides
Atractodes
Atractodes is a cosmopolitan genus of parasitoid wasps in the family Ichneumonidae, first described by Gravenhorst in 1829. The genus has been historically classified in different subfamilies, with some species groups placed in Phygadeuontinae and others in Cryptinae. Multiple species groups have been defined to accommodate morphologically similar species, particularly in the Neotropical region where recent taxonomic revisions have described numerous new species. The genus exhibits considerable diversity in the Americas, with records from North America, Central America, and South America, as well as Europe.
Cassidinae
tortoise and leaf-mining beetles, tortoise beetles, hispine beetles
Cassidinae is a large subfamily of leaf beetles (Chrysomelidae) comprising over 125 genera and approximately 3,000 species worldwide, with greatest diversity in the Neotropics. The subfamily encompasses two historically recognized groups: the 'cassidoids' or tortoise beetles, characterized by expanded pronotal and elytral margins that form a protective shield over the body, and the 'hispoids' or leaf-mining beetles, typically more elongated with parallel-sided bodies. Members exhibit remarkable morphological diversity, with some species displaying metallic coloration and the ability to change color through water movements within the translucent cuticle. The subfamily is notable for complex larval defensive adaptations including fecal shield construction and cycloalexic behavior, as well as the independent evolution of subsociality with paternal care in at least two lineages.
Chrysomelinae
broad-bodied leaf beetles, broad-shouldered leaf beetles
Chrysomelinae is a subfamily of leaf beetles (Chrysomelidae) comprising approximately 3,000 species worldwide, commonly known as broad-bodied or broad-shouldered leaf beetles. The subfamily includes the economically significant Colorado potato beetle (Leptinotarsa decemlineata), a major agricultural pest. Chrysomelinae exhibits remarkable diversity in form and coloration, with adults typically displaying convex, rounded bodies often with bright coloration and variable patterns. The subfamily is distinguished by several unique morphological features including antennae inserted on or adjacent to the anterior head edge, mandibles with large membranous prosthecae, and a single anal cell in each wing. Larvae possess six pairs of stemmata, palmate mandibles, and annular spiracles. Both life stages possess defensive glands that secrete protective chemicals.
Diabroticina
Diabroticina is a subtribe of leaf beetles (Chrysomelidae: Galerucinae: Luperini) comprising approximately six genera and numerous species, many of which are economically significant agricultural pests. The subtribe exhibits striking species diversity, particularly in the Neotropics, with feeding habits ranging from highly polyphagous to oligophagous on single host families. Adults primarily feed on pollen, while larvae are root-feeders. Ecological and behavioral data remain incomplete for most non-pest species.
Dilophonotini
Dilophonotini is a tribe of hawk moths (family Sphingidae) containing approximately 25 genera divided into two subtribes: Dilophonotina and Hemarina. The tribe includes both day-flying and crepuscular species, with Hemaris (clearwing moths) being particularly notable for their transparent wings and bee-mimicry. Members range from small to medium-sized sphingids with diverse wing patterns and body forms adapted to varied ecological niches.
Empidinae
dance flies
Empidinae, commonly called dance flies, are a subfamily of predatory empidoid flies. Males of many species form mating swarms near visual markers, often presenting captured prey to females as nuptial gifts during courtship. The subfamily exhibits diverse mating systems, including lek polygyny and sex-role reversal where females swarm and males choose mates. Many species are effective pollinators of flowering plants, particularly in alpine and subalpine ecosystems where they can achieve pollination effectiveness comparable to honeybees.
Fulcidacini
Warty Leaf Beetles
Fulcidacini is a tribe of leaf beetles (Chrysomelidae) within the subfamily Cryptocephalinae, historically treated as the distinct subfamily Chlamisinae. The tribe comprises 11 genera and approximately 400 species, with roughly 80% of species occurring in the Neotropics. Members are commonly known as warty leaf beetles due to their distinctive appearance and are characterized by case-bearing larvae that construct protective cases from fecal material.
Membracidae
treehoppers, thorn bugs, typical treehoppers
Membracidae, commonly called treehoppers or thorn bugs, is a family of approximately 3,200–3,500 species in over 400 genera within the order Hemiptera. The family is distinguished by extraordinary morphological diversity, particularly the pronounced enlargement and modification of the pronotum—the dorsal plate of the first thoracic segment—which can form thorns, horns, keels, or bizarre projections that often obscure the body and wings. This family represents the most diverse lineage within the superfamily Membracoidea, with its center of diversity in the New World tropics. Most species exhibit phytophagous habits, feeding on plant sap with piercing-sucking mouthparts. Many species engage in mutualistic relationships with ants, which protect them from predators in exchange for honeydew.
Ortheziidae
ensign scales, ortheziids
Ortheziidae is a family of scale insects (Hemiptera) commonly known as ensign scales, comprising 20 genera and 198 species. The family has a cosmopolitan distribution with species richness concentrated in the Neotropical and Nearctic regions; representation is sparse in Australasia and the Far East. Members are distinguished by their mobile adult females, six legs, and prominent waxy secretions. Several species are economically significant agricultural pests.
Oryctini
rhinoceros beetles
Oryctini is a tribe of large, robust scarab beetles within the subfamily Dynastinae, commonly referred to as "rhinoceros beetles." Members range from 25–60 mm in length and are typically dark brown to black. The tribe includes approximately 25 genera distributed worldwide, with particular diversity in the Neotropical region. Several species are economically significant pests, notably in the genus Oryctes, which damages palms and sugarcane. Males frequently possess pronounced horns or armored projections on the head and pronotum, with horn development influenced by larval nutrition.
Phengodidae
glowworm beetles, glow-worms, railroad-worms, trenecitos, bigotudos
Phengodidae is a family of bioluminescent beetles commonly known as glowworm beetles. The family contains over 250 described species distributed throughout the New World from southern Canada to Chile, with additional diversity in Western Asia through the subfamily Cydistinae. Females and larvae possess bioluminescent organs that emit yellow, green, or red light; the Brazilian railroad worm Phrixothrix hirtus is the only terrestrial organism known to produce true red light. Females are larviform and typically much larger than males, which are winged, short-lived, and often attracted to lights. The family was formerly considered to include the Old World Rhagophthalmidae, now recognized as a separate family.
Polymitarcyidae
Pale Burrowers
Polymitarcyidae is a family of burrowing mayflies comprising approximately 10 genera and more than 90 described species. The family exhibits the shortest adult lifespan among mayflies and unique biological traits including silk production by nymphs. Nymphs construct U-shaped burrows in aquatic sediments, functioning as ecosystem engineers through bioturbation. The family shows highest diversity in the Neotropical region, with Campsurinae representing the most species-rich subfamily. Adults possess atrophied legs except for the anterior legs of males.
Sarcophaginae
Typical Flesh Flies
Sarcophaginae is a diverse subfamily of flesh flies comprising at least 60 genera and over 1,800 species worldwide, with the highest diversity in the Neotropical region. These calyptrate flies are characterized by their association with decomposing organic matter, including carrion, feces, and decaying material. Many species exhibit synanthropic behavior, thriving in human-modified environments. Their life histories encompass necrophagy, coprophagy, and scavenging, with some species documented in association with eusocial wasp nests. Species identification relies heavily on male genitalia due to uniform external chaetotaxy.
Saturniidae
Emperor and Giant Silk Moths, saturniids, giant silkworm moths, royal moths
Saturniidae is a family of Lepidoptera containing approximately 2,300 described species, including some of the largest moths in the world. Adults are characterized by large lobed wings, heavy bodies covered in hair-like scales, reduced or vestigial mouthparts, and the absence of a frenulum. The family exhibits notable diversity in the Neotropics, with highest species richness in tropical and subtropical wooded regions. Many species produce silk cocoons, and several have significant economic or cultural importance.
Sciophilinae
fungus gnats
Sciophilinae is a subfamily of fungus gnats within the family Mycetophilidae, comprising at least 40 genera and 340 described species. Members are found in humid environments, often associated with decaying wood and fungi. The subfamily has a global distribution with significant diversity in Neotropical, Nearctic, and Palearctic regions. Natural history information remains limited for most species.
Sminthurididae
Sminthurididae is a family of globular springtails (Collembola: Symphypleona) distinguished by extreme sexual dimorphism, particularly in antennae structure and body size. Males possess clasper-shaped modified antennae used in courtship displays, while females retain unmodified antennae; males are also markedly smaller than females. The family comprises approximately 155 species across 12 extant genera, with notable diversity in the Neotropical region. Diagnostic morphological features include spherical short ventral tube sacs, presence of all abdominal bothriotricha (A–E), and absence of subanal appendage in females.
Syrphini
hoverflies
Syrphini is a globally distributed tribe of hoverflies (family Syrphidae) comprising approximately 37 recognized genera and hundreds of species. Members exhibit classic hoverfly morphology with compact bodies, large compound eyes, and the characteristic hovering flight behavior. The tribe is taxonomically diverse, with particularly notable patterns in the Neotropical region where species are concentrated in two large, highly diverse genera, contrasting with other zoogeographic regions where moderate-sized genera predominate. The group has been subject to recent taxonomic revision, with eight new genera and multiple subgenera described in recent decades.