Larval-leaf-mining
Guides
Cassidinae
tortoise and leaf-mining beetles, tortoise beetles, hispine beetles
Cassidinae is a large subfamily of leaf beetles (Chrysomelidae) comprising over 125 genera and approximately 3,000 species worldwide, with greatest diversity in the Neotropics. The subfamily encompasses two historically recognized groups: the 'cassidoids' or tortoise beetles, characterized by expanded pronotal and elytral margins that form a protective shield over the body, and the 'hispoids' or leaf-mining beetles, typically more elongated with parallel-sided bodies. Members exhibit remarkable morphological diversity, with some species displaying metallic coloration and the ability to change color through water movements within the translucent cuticle. The subfamily is notable for complex larval defensive adaptations including fecal shield construction and cycloalexic behavior, as well as the independent evolution of subsociality with paternal care in at least two lineages.