Honeydew

Guides

Aetalionidae
Aetalionid Treehoppers
Aetalionidae is a family of treehoppers in the superfamily Membracoidea, distinguished from the more diverse Membracidae by morphological features including a front femur fused to the trochanter and a completely exposed scutellum. The family is predominantly Neotropical in distribution, with two recognized subfamilies: Biturritiinae (exclusively Neotropical) and Aetalioninae (containing the Neotropical genus Aetalion and the Paleotropical genus Darthula). Females exhibit maternal care by guarding egg masses, and many species form aggregations on tree branches where they produce honeydew and associate with ants and stingless bees.
treehopper Membracoidea Neotropical maternal-care ant-mutualism honeydew phytophagous sap-feeder pest
Aphidina
Aphidina is a subtribe of aphids within the family Aphididae, established by Latreille in 1802. It encompasses numerous genera including economically significant and xerophilous species. Members exhibit diverse life cycles, with some species amenable to genetic control methods due to their holocyclic heterogenetic reproduction. The subtribe includes species with specialized host associations, particularly on Asteraceae, Chenopodiaceae, and Amaranthaceae.
aphid phloem-feeding honeydew xerophilous parthenogenesis holocyclic pest
Aphidoidea
aphids, plant lice, greenflies, blackflies, whiteflies
Aphidoidea is a superfamily of small sap-sucking insects within Hemiptera, commonly known as aphids. Members possess piercing-sucking mouthparts adapted for feeding on plant phloem. The superfamily includes approximately 5,000 described species across multiple families, with Aphididae being the largest and most economically significant. Many species exhibit complex life cycles involving both sexual and parthenogenetic reproduction, often with seasonal alternation between host plants.
Hemiptera Sternorrhyncha phloem-feeding parthenogenesis agricultural-pest cornicles honeydew ant-mutualism vector biological-control
Aphis
Aphis is a large genus of aphids in the family Aphididae containing at least 400–600 species. The genus includes numerous significant agricultural pests, notably the soybean aphid (Aphis glycines), cotton aphid (Aphis gossypii), and apple aphid (Aphis pomi). Many species form mutualistic associations with ants (myrmecophily), exchanging honeydew for protection. Species identification within the genus often requires examination of morphological characters such as siphunculi and cauda structure.
aphid agricultural-pest myrmecophile phloem-feeder parthenogenesis honeydew vector invasive-species
Balcanocerus
Balcanocerus is a genus of leafhoppers in the family Cicadellidae, subfamily Eurymelinae. Species in this genus are ant-attended and feed on phloem sap from host plants in Rosaceae. The genus includes both North American and European species, with documented associations with specific ant partners and host trees.
leafhopper Eurymelinae ant-attended trophobiosis Rosaceae honeydew aggregation-behavior Palaearctic Nearctic
Bolbonota
seed-mimic treehopper
Bolbonota is a genus of treehoppers in the family Membracidae, tribe Membracini, distributed in the New World tropics and subtropics. Adults are characterized by their dark, globular body form with a corrugated pronotal surface and distinctive red eyes. The genus contains at least a dozen described species in Brazil alone, with many more likely awaiting description. Bolbonota species exhibit complex ontogenetic color changes: early instar nymphs are dark, late instars become mottled or streaked white, and adults revert to dark coloration.
Membracidae treehopper ant-mutualism mimicry ontogenetic-color-change Neotropical sap-feeding Camponotus honeydew aggregation-behavior
Ceroplastes
wax scales
Ceroplastes is a genus of wax scales in the family Coccidae, comprising more than 130 described species. These insects are characterized by the conspicuous white, waxy coverings they secrete, which protect their soft bodies. Adults are largely sessile and feed on plant sap using piercing-sucking mouthparts. Many species are economically significant pests of ornamental plants, fruit trees, and shrubs. The genus exhibits parthenogenesis in many species, with females reproducing without males. Climate change has facilitated the northward expansion of several species from southern regions into previously cooler areas.
scale-insect pest horticulture agriculture parthenogenesis invasive-species climate-change sooty-mold honeydew ornamental-plants
Ceroplastes cirripediformis
barnacle scale, barnacle wax scale
Ceroplastes cirripediformis, commonly known as the barnacle scale or barnacle wax scale, is a soft scale insect (Hemiptera: Coccidae) recognized for its distinctive waxy covering that resembles a small barnacle. It is a destructive invasive pest with a broad host range spanning numerous plant families. The species has expanded its range globally through human-mediated transport and climate change, with established populations across North America, South America, Europe, Asia, Africa, and Oceania. Females reproduce parthenogenetically, producing up to a thousand eggs annually without observed males in wild populations.
invasive-species scale-insect agricultural-pest ornamental-pest parthenogenesis climate-change honeydew sooty-mold wax-covering Coccidae Hemiptera
Cinara
giant conifer aphids, conifer aphids
Cinara is a genus of aphids comprising approximately 200-243 species, commonly known as giant conifer aphids. These insects are specialized feeders on conifers in the families Pinaceae and Cupressaceae, including pines, firs, spruces, cedars, and cypresses. Molecular studies indicate their tribe Eulachnini represents the basal group for other aphids. Several species are significant pests of cultivated Christmas trees, particularly Fraser fir production in the southeastern United States.
aphids conifers forest-pests Christmas-trees honeydew phloem-feeders parthenogenesis Northern-Hemisphere
Cinara pergandei
Globose Pine Aphid
Cinara pergandei is a large aphid species in the family Aphididae, commonly known as the Globose Pine Aphid. It feeds on coniferous trees, particularly pines, and produces copious amounts of honeydew that attracts wasps and other insects. Like other Cinara species, it has piercing-sucking mouthparts adapted for tapping into phloem sap. It is part of a genus containing numerous conifer-feeding aphids, many of which are significant pests in forestry and Christmas tree production.
aphid pine conifer pest honeydew Christmas-tree forestry Hemiptera Sternorrhyncha Lachninae
Cinara strobi
White Pine Aphid
Cinara strobi, the white pine aphid, is a large aphid species specialized on white pine (Pinus strobus). It is notable for exceptionally high autumn populations that produce copious honeydew, attracting stinging wasps and creating nuisance conditions. The species exhibits parthenogenetic reproduction during the growing season, with live birth of nymphs accelerating population growth. Uniquely among Cinara aphids, C. strobi harbors three bacterial endosymbionts—Buchnera aphidicola, Sodalis sp., and Serratia symbiotica—representing an intermediate stage of symbiont replacement where the latter has become metabolically incompetent yet persists.
aphid white-pine honeydew parthenogenesis endosymbiont Serratia-symbiotica Buchnera Sodalis autumn-pest Sternorrhyncha
Coccus viridis
green scale, green coffee scale, coffee green scale
Coccus viridis is a soft scale insect (Hemiptera: Coccidae) and major agricultural pest with a wide tropical and subtropical distribution. It reproduces parthenogenetically via thelytoky, with females producing 50-600 eggs that hatch within minutes to hours beneath the mother's body. The species completes its life cycle in 47-51 days at 25°C, with three nymphal instars before adulthood; first-instar nymphs actively disperse as 'crawlers' while older instars and adults become sessile. It is polyphagous, feeding on phloem sap of over 200 plant genera in 72 families, with primary economic damage to coffee, guava, and cacao crops.
agricultural-pest coffee-pest soft-scale parthenogenesis thelytoky polyphagy ant-mutualism honeydew sooty-mold biological-control invasive-species tropical-pest phloem-feeder Hemiptera Coccidae Coccus green-scale green-coffee-scale
Disholcaspis prehensa
clasping twig gall wasp
Disholcaspis prehensa is a gall-inducing cynipid wasp native to California that induces distinctive galls on scrub oak and leather oak. The species exhibits heterogony, alternating between sexual and asexual generations that produce two morphologically different gall types. The asexual generation forms conspicuous mushroom-shaped twig galls with clasping bases, while the sexual generation produces small, hidden bud galls.
gall-wasp cynipid heterogony parthenogenesis oak-gall California-endemic ant-mutualism Disholcaspis Cynipidae Hymenoptera Quercus induced-plant-structure phloem-sap-exudation biological-mutualism seasonal-dimorphism sexual-dimorphism gall-dimorphism agamic-generation sexual-generation clasping-twig-gall bud-gall twig-gall native-California-species scrub-oak leather-oak parasitoid-defense ant-plant-interaction insect-plant-interaction plant-manipulation evolution-of-asexuality Weld-1957 Zootaxa-4859 molecular-systematics host-plant-specificity Cynipini oak-woodland chaparral insect-gall gall-morphology gall-coloration sun-exposure-effects larval-chamber emergence-timing spring-emergence late-winter-emergence gall-development oviposition-behavior egg-laying-in-buds egg-laying-in-twigs female-only-generation wingless-female winged-sexual-generation gall-size-dimorphism gall-shape-dimorphism hidden-gall exposed-gall gall-wasp-life-cycle cynipid-biology gall-wasp-taxonomy gall-wasp-identification gall-wasp-ecology insect-plant-coevolution plant-growth-manipulation hormone-mediated-gall-induction venom-gland accessory-gland secretory-structures gall-induction-mechanism gall-chemistry phloem-sap honeydew ant-attraction ant-protection parasitoid-wasp natural-enemy-defense indirect-defense mutualistic-interaction trophic-cascade community-ecology species-interaction ecosystem-engineer habitat-modification food-web insect-diversity California-biodiversity endemic-insect regional-endemism oak-specialist plant-specialist monophagous oligophagous host-range host-plant-evolution gall-wasp-radiation adaptive-radiation evolutionary-innovation reproductive-strategy asexual-reproduction thelytoky arrhenotoky sex-determination developmental-biology insect-morphology insect-anatomy female-morphology ovipositor stinger egg-injection plant-tissue-manipulation cell-proliferation plant-hormone auxin cytokinin gene-expression transcriptomics proteomics venom-composition 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protected-area natural-reserve state-park national-park wilderness-area biosphere-reserve UNESCO IUCN red-list species-assessment conservation-status data-deficient research-priority knowledge-gap future-research gall-wasp-venom gall-wasp-gland secretory-evolution gall-wasp-phylogeny Cynipidae-evolution Hymenoptera-phylogeny insect-evolution gall-evolution plant-insect-interaction-evolution coevolutionary-arms-race mutualism-evolution defense-evolution counter-defense enemy-free-space enemy-escape ecological-release adaptive-peak fitness-landscape evolutionary-constraint evolutionary-trade-off life-history-evolution reproductive-allocation fecundity survival development-time generation-time voltinism univoltine bivoltine multivoltine diapause overwintering cold-hardiness stress-tolerance desiccation-resistance starvation-resistance immune-response parasitoid-resistance pathogen-resistance symbiont microbiome gut-microbiota endosymbiont Wolbachia Rickettsia Cardinium Spiroplasma bacterial-symbiont fungal-symbiont viral-symbiont mutualistic-microbe parasitic-microbe commensal-microbe microbial-community metagenomics microbial-ecology host-microbe-interaction immune-system innate-immunity acquired-immunity humoral-immunity cellular-immunity antimicrobial-peptide reactive-oxygen-species nitric-oxide melanization encapsulation nodulation phagocytosis autophagy apoptosis programmed-cell-death stress-response heat-shock-protein oxidative-stress hypoxia anoxia metabolic-rate respiration energy-budget nutrient-allocation lipid-storage glycogen-storage protein-storage amino-acid-metabolism carbohydrate-metabolism lipid-metabolism nitrogen-metabolism excretion osmoregulation ion-transport water-balance cuticle integument sclerotization pigmentation color-pattern structural-color iridescence bioluminescence photoreceptor vision compound-eye ocellus antenna olfaction gustation chemoreception mechanoreception thermoreception hygroreception proprioception neurobiology brain 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recovery-management-management-management rehabilitation-management-management-management reconstruction-management-management-management resilience-building-management-management-management adaptive-capacity-building-management-management-management transformative-capacity-building-management-management-management
Dolichoderus
Dolichoderus is a genus of ants comprising 181 valid species (131 extant, 50 extinct), representing the sole member of tribe Dolichoderini and type genus of subfamily Dolichoderinae. Workers are small (approximately 4 mm), possess a thick sculptured integument, and lack a functional sting. The genus exhibits remarkable ecological diversity: most species are generalist honeydew collectors that tend aphids and scale insects, while the D. cuspidatus group in Southeast Asia practices unique nomadic aphid farming with obligate mutualist mealybugs. The genus has an extensive fossil record in Baltic amber.
ants honeydew mutualism nomadic aphid-farming carton-nests Baltic-amber polygyny
Dolichoderus mariae
Mary's Odorous Ant
Dolichoderus mariae is a distinctive North American ant species named after naturalist Mary Treat, who discovered it. The species is notable for its reddish-brown and black bicolored appearance, large polydomous colonies with dozens of nests, and its specialized ecology in prairie and glade habitats where it nests among plant roots. Colonies exhibit dramatic seasonal fluctuations in nest numbers and worker populations, and engage in territorial aggression with neighboring colonies.
ant polydomous prairie honeydew Mary-Treat Dolichoderinae
Dolichoderus taschenbergi
Taschenberg's Odorous Ant
Dolichoderus taschenbergi is a North American ant species in the odorous ant subfamily Dolichoderinae. Described by Mayr in 1866, it is endemic to Canada and the United States. The species exhibits sporadic dispersion patterns and has been studied for its mutualistic relationships with honeydew-producing hemipterans. Colony structure may include polygynous nesting patterns, though this requires further research.
ant Dolichoderinae honeydew mutualism trophobiosis forest-ant North-America scale-insect-attendant odorous-ant
Entylia
keel-shaped treehopper
Entylia is a genus of treehoppers in the family Membracidae, first described by Germar in 1833. The genus contains at least three described species, including Entylia carinata, which is known for its distinctive keel-shaped pronotum that resembles plant debris. These insects are notable for their mutualistic relationships with ants, which protect them from predators in exchange for honeydew. The genus is distributed in North America, with observations recorded from regions including the northeastern United States.
treehopper Membracidae ant-mutualism honeydew camouflage Asteraceae phloem-feeder keel-shaped trophobiont myrmecophily North-America
Eriosoma americanum
Woolly Elm Aphid
Eriosoma americanum, commonly known as the woolly elm aphid, is a North American aphid species that alternates between two host plants: American elm (Ulmus americana) and Saskatoon (Amelanchier alnifolia). The species produces distinctive woolly or cottony wax secretions that give it its common name. Heavy infestations on American elm cause leaf curling and produce visible dark patches and cottony masses. Like other aphids, it excretes honeydew while feeding on plant sap.
aphid pest elm Saskatoon woolly North-America host-alternation honeydew leaf-curl Hemiptera Aphididae Eriosomatinae
Eulachnini
Eulachnini is a tribe of aphids within the subfamily Lachninae, comprising approximately 27 species documented in Lithuania alone. Members are specialized phloem-feeders on coniferous plants, particularly Pinaceae. Several species produce substantial honeydew, with five noted as promising honeydew producers in Lithuanian forests. Only Cinara piceae has been identified as a potential forest pest, with documented damage to firs in nursery settings.
aphid conifer forest-pest honeydew Lithuania COI-haplotype
Formica incerta
Uncertain Field Ant
Formica incerta is a common ant species of eastern North America, often the most abundant Formica in its range. It constructs underground nests with inconspicuous entrance holes and no surface mound. Workers forage diurnally for nectar and honeydew, and the species serves as the primary host for the slave-making ant Polyergus lucidus. First described by Carlo Emery in 1893, its specific epithet reflects historical taxonomic uncertainty and identification challenges.
ant Formicidae Formicinae North-America slave-making-ant-host grassland meadow prairie nectar honeydew underground-nest diurnal-forager
Formica integroides
Western Mound Ant, Vinegar Ant
Formica integroides, commonly known as the Western Mound Ant or Vinegar Ant, is a North American wood ant species in the family Formicidae. This species exhibits flexible, context-dependent foraging strategies at both colony and individual levels, adjusting its behavior based on resource characteristics and competitive pressure. It is known to utilize both terrestrial and arboreal resources, with foragers capable of defending entire trees as absolute territories. The species was first described by Wheeler in 1913.
wood-ant mound-ant foraging-behavior competition arboreal-foraging terrestrial-foraging honeydew Formica North-America
Homotomidae
fig psyllids, mulberry psyllids
Homotomidae is a family of small phloem-feeding insects in the superfamily Psylloidea, commonly known as fig or mulberry psyllids. Members feed almost exclusively on Moraceae, primarily the genus Ficus, with rare exceptions such as Triozamia on Antiaris toxicaria. The family contains approximately 70 described species across three subfamilies: Dynopsyllinae, Macrohomotominae, and Homotominae. Several species are economically significant pests of ornamental and cultivated figs, including the invasive Macrohomotoma gladiata, which has spread from Asia to Europe, North Africa, and North America.
phloem-feeding Moraceae Ficus invasive-pest urban-pest ornamental-pest biological-control Psylloidea wax-secretion honeydew sooty-mould
Liometopum apiculatum
Escamolera Ant, Velvet Tree Ant
Liometopum apiculatum is a Neotropical dolichoderine ant with significant ecological and economic importance. Colonies are large and polydomous, ranging from a few hundred to hundreds of thousands of workers. The species is notable for its distinctive carton-like nest construction and its role as a traditional food source in Mexico, where the larvae and pupae (escamoles) are harvested. It inhabits diverse arid and semi-arid vegetation types across southwestern North America.
escamoles edible-insects entomophagy mutualism honeydew semiochemicals polydomous haplometrosis conservation Mexico southwestern-USA
Melanaphis
sugarcane aphid
Melanaphis is a genus of aphids in the family Aphididae, first described by van der Goot in 1917. The genus includes species that are significant agricultural pests, particularly of sorghum and sugarcane. Melanaphis sacchari (sugarcane aphid) has become one of the most damaging insect pests of grain and forage sorghum in the United States since its first recorded occurrence on sorghum in Texas in 2013. Members of this genus are characterized by their parthenogenetic reproduction, rapid population growth, and production of honeydew.
aphid pest sorghum sugarcane agriculture honeydew parthenogenesis
Membracidae
treehoppers, thorn bugs, typical treehoppers
Membracidae, commonly called treehoppers or thorn bugs, is a family of approximately 3,200–3,500 species in over 400 genera within the order Hemiptera. The family is distinguished by extraordinary morphological diversity, particularly the pronounced enlargement and modification of the pronotum—the dorsal plate of the first thoracic segment—which can form thorns, horns, keels, or bizarre projections that often obscure the body and wings. This family represents the most diverse lineage within the superfamily Membracoidea, with its center of diversity in the New World tropics. Most species exhibit phytophagous habits, feeding on plant sap with piercing-sucking mouthparts. Many species engage in mutualistic relationships with ants, which protect them from predators in exchange for honeydew.
Hemiptera Membracoidea phytophagous ant-mutualism pronotum Neotropical-diversity maternal-care honeydew
Metcalfa
Metcalfa is a genus of planthoppers in the family Flatidae, tribe Nephesini, native to North and Central America. The genus contains at least five described species, with Metcalfa pruinosa (citrus flatid planthopper) being the most economically significant due to its invasive spread to Europe and Asia. Adults are small, ranging 5.5–8 mm in length with a waxy, often bluish or grayish appearance. The genus is characterized by forewings with parallel cross-veins along the costal margin, distinguishing it from related families.
Flatidae planthopper invasive-pest M.-pruinosa citrus-flatid-planthopper waxy-secretions honeydew sooty-mold biological-control polyphagous
Microterys
Microterys is a large genus of parasitoid wasps in the family Encyrtidae (Chalcidoidea), with its center of distribution in the northern parts of the Northern Hemisphere. Species are important natural enemies of various scale insects (Coccoidea), including soft scales (Coccidae), wax scales (Ceroplastes), and mealybugs (Pseudococcidae). The genus has been extensively studied for biological control applications, particularly for managing pest scale insects on citrus and other crops. Several species have been introduced to new regions as biocontrol agents, including Microterys flavus in California.
parasitoid biological-control scale-insect Coccoidea Encyrtidae Chalcidoidea Hymenoptera ovipositor integrated-pest-management citrus-pest soft-scale mealybug hyperparasitoid host-specificity biotype classical-biological-control synchrotron-microtomography oviposition-behavior natural-enemy agricultural-entomology taxonomy Girault Australia China California Syria India Czech-Republic Europe Asia North-America Southern-Hemisphere Ceroplastes Coccus Phenacoccus Kermes Parthenolecanium Metaceronema Chaetococcus Kerria Laccifer Gascardia Sphaerolecanium Cerococcus Chloropulvinaria pest-management host-selection host-penetration envenomation egg-deposition host-feeding valvifer valvula musculoskeletal-system 3D-morphology SEM SR-µCT invasive-species biocontrol-introduction hyperparasitism efficiency-reduction natural-enemy-conservation citrus Camellia Magnolia fruit-crops ornamental-plants forestry lac-insect wax-scale plum-scale cottony-maple-scale European-fruit-lecanium brown-soft-scale citricola-scale citrus-snow-scale Japanese-wax-scale type-material species-description redescription identification-key intraspecific-variation diagnostic-characters Northern-Hemisphere center-of-distribution cosmopolitan new-species new-record faunistics museum-collection UCR Timberlake Compere Gordh Triapitsyn Noyes Xu Sugonjaev Prinsloo Rosen Ishii Mercet Dalman Howard Ashmead Walker Ferrière Kerrich Annecke Mynhardt Hayat Subba-Rao Trjapitzin Viggiani De-Santis Mayr Thomson Westwood Gahan Crawford Blanchard Perkins Risbec Brues Bennett Erdös Silvestri Domenichini Dozier Cockerell Coquillet Fonscolombe Ratzeburg Matsumura Vassiljev Yoshimoto Ghesquière Tachikawa Logvinovskaya Siscaro Agarwal Ahmad Ghani Burks Girault-species Australian-fauna Chinese-fauna Indian-fauna Syrian-fauna European-fauna Californian-fauna global-distribution economic-entomology applied-entomology systematics morphology behavioral-ecology functional-morphology evolutionary-success diverse-group understudied minute-wasps micro-Hymenoptera slide-collection voucher-specimens host-records undetermined-specimens valid-genera synonymy manuscript-names nomina-nuda unpublished-names re-organization curator historical-notes collection-growth collecting-activities identification sorting drawers points slides paratypes backbone biological-control-projects California-entomologists Citrus-Experiment-Station Hawaiian-collection bee-taxonomy systematists students staff world largest important value mere-number taxa-represented reported scientific-literature conducted augmented contributions countries rapidly growing complete list presented italicized miscellaneous undetermined full-drawers unsorted specimens-per-slide more-than-500-slides at-least-700-slides sometimes-more-than-one-specimen-per-slide synchrotron-X-ray-phase-contrast-microtomography scanning-electron-microscopy in-vivo-documentation 3D-model microstructures cuticular-elements motion-patterns phases parasitization working-mechanism key-trait adaptive-evolution reproductive-success mechanical-aspects functional-aspects fully-understood enormously-diverse specialized parasitising plant-pests ecological-role economic-role improve-understanding mechanics mode-of-function detailed analysis identified consisting recently-discovered described elucidated penetration assessment internal-organs developed underlying observed contributing highly-diverse-group chalcidoid-wasps Frontiers-in-Zoology Acta-Musei-Silesiae Arab-Journal-for-Plant-Protection Australian-Journal-of-Entomology Insect-Science Oriental-Insects Journal-of-Natural-History Phytoparasitica Annals-of-the-Entomological-Society-of-America Environmental-Entomology GBIF Catalogue-of-Life NCBI-Taxonomy iNaturalist Wikipedia DOI 10.1186/s12983-025-00575-1 10.2478/cszma-2021-0007 10.22268/AJPP-41.3.292305 10.1111/j.1440-6055.1975.tb02058.x 10.1111/j.1744-7917.2000.tb00345.x 10.1080/00305316.2011.646827 10.1111/j.1440-6055.1973.tb01670.x 10.1080/00222933.2020.1786608 10.1007/s12600-019-00727-0 10.1007/s12600-018-0691-5 10.1093/aesa/54.2.222 10.1093/ee/7.6.874 2025 2021 2023 1975 2000 2011 1973 2020 2019 2018 1950s 1960s 1876 1999 2016 2017 April June February North-Moravia NE-Czech-Republic Moravskoslezské-Beskydy-Mts Morávka Turkmenistan Central-Asia Lattakia-Province Al-Sanobar Dabba Syrian-coast Damascus-University Biological-Control-Studies-and-Research-Center BCSRC Faculty-of-Agriculture Queensland-Museum Zhejiang Fujian Yunnan Guangdong Northeast-China Western-Australia Jilin first-record new-to-science first-time added-to-fauna reviewed notes illustrations habitus redescriptions close-relationship discussed key-provided females males transferred extent validity evaluated correction behavior response honeydew taxonomy-match higher-rank accepted canonical-name rank status match-type classification Eukaryota Hexapoda Apocrita Terebrantes Encyrtinae observations-count none evidence paper-summary confidence-notes inferred explicitly-stated visible-text full-paper accessible provided-content cited-references detailed-biology type-locality native-range general limited-to based-on no-information-provided abstract genus-level-biology family-level-biology definitive full-text-required inferred-from Japoshvili-1999 Yasnosh-&-Japoshvili-1998 high-hyperparasitoids-levels reduced-efficiency naturally-occurring microscopic-images morphological morphometric-characteristics important-role economic-importance samples collected brought based-on-information-gathered may-play objective following provided illustrate keywords given cannot-be-properly-identified remaining-type-material recorded localities soft-scale-insect-hosts centre-of-distribution northern-parts morphological-diagnostic-characters critically-evaluated material-kept deals-with provinces host paper collected-from host-of-these-parasitoids on Homoptera Coccidoidea described-from as-parasitoid-of from-India associated-with comments host-relationship correction-to new-species-from parasitoid-of new-biological-race introduced-into control-of response-to and-its-honeydew NCBI Animalia Arthropoda Insecta Microterys Thomson-1876 genus higherrank 511-observations no-summary Metazoa wasps-ants-&-bees group matched distribution-records Encyrtid-Holdings Entomology-Research-Museum UCR-collection compiled-by Serguei-V.-Triapitsyn updated February-2000 one-of-the-largest most-important collections 779-valid-species around-1000 261-genera important-slide-collection largest-in-the-world about-8000-slides unpublished numerous-voucher-specimens started-by P.H.-Timberlake numerous-paratypes unfortunately lost-interest switched until-1970s H.-Compere notable-contributions P.-Timberlake J.-Maple P.-DeBach C.-Clausen 1970s-1980s G.-Gordh J.-LaSalle J.-Woolley J.-Pinto J.-Beardsley Mexico US other-countries presently growing-rapidly J.-Heraty majority V.-Trjapitzin J.-Noyes complete-re-organization S.-Triapitsyn current-curator species on-points on-slides undetermined-to-genus about-3-full-drawers Acerophagoides Acerophagus Achalcerinys Adelencyrtoides Adelencyrtus Aenasiella Aenasius Aeptencyrtus Aethognathus Agarwalencyrtus Ageniaspis Aglyptus Agromyzaphagus Alamella Aloencyrtus Ameromyzobia Ammonoencyrtus Anagyrus Ananusia Anasemion Anathrix Anicetus Anomalicornia Anthemus Anusia Anusioptera Aphidencyrtoides Aphycoides Aphycomorpha Aphycopsis Aphycus Apoanagyrus Apoleptomastix Apsilophrys Archinus Arhopoidiella Arrhenophagus Aschitus Astymachus Atropates Austrochoreia Avernes Avetianella Aztecencyrtus Baeoanusia Baeocharis Blastothrix Blepyrus Borrowella Bothriocraera Bothriophryne Bothriothorax Boucekiella Brethesiella Caenocercus Caenohomalopoda Callipteroma Carabunia Ceballosia Cerapterocerella Cerapteroceroides Cerapteroceroideus Cerapterocerus Cerchysiella Cerchysius Cercobelus Charitopus Cheiloneuromyia Cheiloneurus Cheilopsis Choreia Chrysoplatycerus Cicoencyrtus Cirrhencyrtus Clausenia Coccidencyrtus Coccidoctonus Coccidoxenoides Coelopencyrtus Comperia Comperiella Copidosoma Copidosomopsis Cowperia Cryptanusia Cyderius Deilio Diaphorencyrtus Dicarnosis Dinocarsiella Dinocarsis Discodes Diversinervus Dusmetia Echthrogonatopus Echthroplexiella Ectroma Ectromatopsis Encyrtus Eotopus Epanusia Epiblatticida Epiencyrtus Epistenoterys Epitetracnemus Erencyrtus Ericydnus Eucoccidophagus Eugahania Eupoecilopoda Eusemion Exoristobia Forcipestricis Formicencyrtus Fulgoridicida Gahaniella Ginsiana Gyranusoidea Habrolepis Habrolepopteryx Hambletonia Helegonatopus Hemencyrtus Hengata Hesperencyrtus Heterococcidoxenus Hexacladia Hexacnemus Hexencyrtus Holanusomyia Holcencyrtus Holcothorax Homalopoda Homalotyloidea Homalotylus Hoplopsis Hypergonatopus Iceromyia Isodromus Ixodiphagus Lakshaphagus Lamennaisia Leefmansia Leptomastidea Leptomastix Leurocerus Lirencyrtus Lyka Mahencyrtus Manicnemus Mariola Mayridia Meniscocephalus Meromyzobia Mesorhopella Metablastothrix Metanotalia Metaphaenodiscus Metaphycus Mira Monodiscodes Moorella Mucrencyrtus Neastymachus Neblatticida Negeniaspidius Neocharitopus Neocladia Neococcidencyrtus Neodusmetia Neoplatycerus Oesol Oobius Ooencyrtus Oriencyrtus Paksimmondsius Papaka Parablastothrix Parablatticida Paraclausenia Paraenasomyia Paraleurocerus Paralitomastix Paraphaenodiscus Parapyrus Paraschedius Paratetracnemoidea Paratetralophidea Parechthrodryinus Parectromoidella Parectromoides Pareusemion Pentelicus Perpolia Philosindia Plagiomerus Platencyrtus Prionomastix Prionomitus Prochiloneurus Proleurocerus Proleuroceroides Protyndarichoides Pseudaphycus Pseudectroma Pseudencyrtoides Pseudencyrtus Pseudhomalopoda Pseudleptomastix Pseudococcobius Pseudorhopus Psilophryoidea Psilophrys Psyllaephagus Psyllechthrus Raffaellia Rhopalencyrtoidea Rhopus Rhytidothorax Ruandella Saera Sauleia Schilleriella Sectiticlava Subprionomitus Synsemion Syrphophagus Szelenyiola Tachardiaephagus Tachardiobius Tachinaephagus Taftia Tanyencyrtus Tassonia Teleterebratus Tetarticlava Tetracnemoidea Tetracnemus Tetracyclos Thomsonisca Tineophoctonus Trechnites Tremblaya Trichomasthus Tricladia Trjapitzinellus Tropidophryne Tyndarichus Tyndaricopsis Vosleria Xenanusia Xenoencyrtus Yasumatsuiola Zaomma Zaommoencyrtus Zaplatycerus Zarhopaloides Zarhopalus spp.sp.sp(p).?nr.ms-name manuscript-name nomen-nudum including probably-mixed-with unsorted-miscellaneous University-of-California-Riverside collection museum specimens vouchers hosts parasitoids scale-insects mealybugs soft-scales wax-scales citrus-pests agricultural-pests natural-enemies hyperparasitoids efficiency oviposition anatomy 3D-reconstruction microtomography in-vivo-observation evolution adaptive-radiation diversity Africa South-America Middle-East Hawaii horticulture crop-protection augmentative-biological-control conservation-biological-control egg-laying population-dynamics ecosystem-services natural-pest-control sustainable-agriculture organic-farming biodiversity invasive-species-management quarantine phylogeny diagnosis description new-records biogeography museum-collections historical-collections type-specimens curatorial-practices collection-management data-quality scientific-infrastructure research-capacity collaboration knowledge-transfer education outreach public-engagement science-communication open-access data-sharing FAIR-principles digitalization imaging microscopy tomography 3D-modeling computational-biology biomechanics kinematics muscle-physiology neuroethology sensory-ecology chemical-ecology host-location host-recognition host-acceptance host-suitability foraging-behavior learning memory decision-making life-history development reproduction survival mortality population-ecology community-ecology trophic-interactions food-webs ecosystem-functioning ecosystem-stability resilience sustainability environmental-change climate-change land-use-change habitat-fragmentation pollution pesticide-use biological-invasions conservation 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sustainable innovative creative critical reflective responsible accountable transparent ethical integral holistic systemic dynamic complex uncertain ambiguous diverse plural collaborative cooperative competitive conflicting consensual deliberative dialogical democratic autocratic hierarchical networked decentralized distributed localized globalized standardized harmonized integrated coordinated aligned balanced optimized satisficing sufficient efficient effective efficacious performant productive profitable viable feasible desirable acceptable legitimate credible reliable valid high-quality excellent distinctive recognized reputable prestigious influential impactful enduring meaningful significant valuable worthy meritorious virtuous beautiful true moral legal authorized approved sanctioned endorsed supported funded resourced staffed equipped enabled empowered capacitated skilled trained educated informed aware conscious mindful attentive focused intentional purposeful
Ochetellus glaber
black household ant, black house ant, tramp ant, Copper-bellied Ant
Ochetellus glaber is a small ant species native to Australia, widely introduced globally through human commerce. Workers measure 2–3 mm, with queens reaching 5.2–5.5 mm. The species is recognized as a species complex with cryptic diversity, complicating identification and management decisions. It nests arboreally and in structures, forming conspicuous trails to exploit honeydew and food resources. Considered a household pest, it has established populations in Hawaii, Florida, New Zealand, and multiple Asian and Pacific regions.
ant tramp-species household-pest species-complex invasive Australia honeydew arboreal Dolichoderinae
Oncometopia
sharpshooters
Oncometopia is a genus of sharpshooters (leafhoppers in the family Cicadellidae) found in North and South America, erected by Carl Stål in 1869. Species range from 9.4 to 15.0 mm in length and are characterized by distinctive morphological features including a moderately produced head, coriaceous forewings with apical membranes, and specialized hindleg tarsomere proportions. Several species are economically important as vectors of plant pathogens, particularly Xylella fastidiosa, which causes diseases such as phony peach disease and citrus variegated chlorosis.
sharpshooter leafhopper Cicadellidae Proconiini xylem-feeder vector Xylella-fastidiosa brochosomes honeydew pest North-America South-America citrus-pest disease-vector agricultural-pest
Oncometopia hamiltoni
Oncometopia hamiltoni is a species of sharpshooter leafhopper in the family Cicadellidae, described by Rakitov in 2016. It belongs to the tribe Proconiini, a group of large, active leafhoppers known for their xylem-feeding habits and distinctive "raining" behavior caused by forceful excretion of excess fluid. The species is part of a genus containing several economically significant sharpshooters that have been studied for their role in plant disease transmission and their interactions with egg parasitoids.
leafhopper sharpshooter Proconiini xylem-feeder brochosomes honeydew egg-parasitoid-host
Oncometopia orbona
Broad-headed Sharpshooter
Oncometopia orbona, commonly known as the broad-headed sharpshooter, is a large leafhopper in the family Cicadellidae. It feeds on xylem sap from plants, processing large volumes of nutrient-poor fluid and excreting the excess as honeydew—a behavior that creates the phenomenon of 'raining trees.' The species possesses distinctive white chalky brochosomes on its sides, used to powder egg-laying sites. It is a strong flier with excellent eyesight and serves as a host for egg parasitoids in the family Mymaridae.
sharpshooter leafhopper xylem-feeder honeydew brochosomes raining-trees Cicadellidae Proconiini egg-parasitoid-host urban-pest-nuisance
Panaphis juglandis
Walnut Aphid, Large Walnut Aphid
Panaphis juglandis, the large walnut aphid, is a phloem-feeding pest of walnut trees (Juglans regia). It colonizes exclusively the upper (adaxial) surface of walnut compound leaves, forming dense colonies along the main vascular veins of the outer leaflets. Native to the Old World (Central Asia and southeastern Europe), it has spread across Europe and was introduced to California in the early 20th century. The species reduces tree vigor, nut yield, and quality, and excretes honeydew that promotes sooty mold growth. It forms mutualistic relationships with ant species that tend and protect colonies from predators.
aphid pest walnut Juglans phloem-feeder honeydew mutualism ants biological-control invasive-species California Europe agriculture urban-ecology
Petrusa
Petrusa is a genus of planthoppers in the family Flatidae, subfamily Flatinae. Members of this genus are found in the Neotropical region, with Petrusa epilepsis being a notable species associated with seagrape and other host plants in the Caribbean. The genus is characterized by the typical flatid morphology with wings held tent-like over the body and parallel cross-veins along the costal margin of the forewings.
planthopper flatid Hemiptera Neotropical pest-potential seagrape honeydew sooty-mold host-plant-generalist Caribbean
Polistes dominula
European Paper Wasp
Polistes dominula is a highly successful invasive social wasp native to Eurasia that has established populations across North America, South America, New Zealand, South Africa, and other regions. First detected in North America near Boston in 1978, it has become one of the most abundant wasps on the continent. The species builds small, exposed paper nests in protected locations and preys primarily on live insects, particularly caterpillars. Unlike yellowjackets, it does not scavenge for meat or sugar. Its rapid spread has been attributed to ecological flexibility, superior competitive ability, and tolerance of human-altered environments.
invasive-species social-wasp paper-wasp predator pollinator urban-ecology eusocial nest-construction biological-control thermal-biology spatial-learning vibrational-communication aposematism honest-signal overwintering diapause Polistes Vespidae Hymenoptera North-America Europe New-Zealand South-Africa South-America monarch-butterfly caterpillar-predation nest-architecture gyne foundress dominance-hierarchy climate-change-impact energetics metabolic-rate microclimate suburban-habitat anthropogenic ornamental-pest vineyard-pest fruit-damage stinging-insect venom poison-gland warning-coloration Zahavian-signal Strepsiptera parasitoid Xenos behavioral-plasticity foraging learning memory thermoregulation ectothermy endothermy solar-radiation nest-temperature brood-development colony-survival multiple-founding single-founding season-length latitude temperate Mediterranean hibernacle overwintering-costs climate-warming range-expansion competition native-species-displacement biodiversity-impact conservation management control integrated-pest-management IPM identification antennae-color facial-markings caste reproduction male female sexual-dimorphism antennae abdomen wing mandible saliva paper comb cell pedicel trophallaxis larva pupa adult worker queen subordinate dominant aggression defense threat-display sting toxicity brightness coloration predator-avoidance prey caterpillar Lepidoptera butterfly moth garden agriculture vineyard orchard urban suburban rural forest habitat-preference temperature metabolism quiescence torpor survival fitness colony nest population abundance distribution range expansion invasion establishment introduction biotic-interaction trophic-cascade herbivory-suppression plant-fitness milkweed Danaus-plexippus monarch Nelson Tasman ecosystem-impact conservation-management biological-invasion ecology behavior physiology morphology taxonomy systematics nomenclature gender-agreement Latin Christ 1791 Vespa-dominula basionym 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Publilia
Publilia is a genus of treehoppers (Membracidae) native to North America, with species occurring primarily in the eastern and central United States. The genus is notable for forming dense aggregations on host plants, particularly goldenrods (Solidago spp.), where nymphs and adults are tended by ants in exchange for honeydew. This mutualistic relationship has made Publilia a model system for studying ant-hemipteran interactions and their ecological consequences for host plants. The genus is distinguished from related treehoppers by its relatively unadorned pronotum compared to the elaborate projections seen in many Neotropical membracids.
treehopper membracidae ant-mutualism myrmecophily hemiptera solidago goldenrod aggregation-behavior honeydew nearctic
Publilia porrecta
Publilia porrecta is a Nearctic treehopper species in the family Membracidae, tribe Polyglyptini. The genus Publilia is known for ant-tended mutualistic associations, with multiple ant species frequently visiting aggregations. Like other members of its tribe, this species exhibits subsocial behaviors and is associated with herbaceous host plants. The species has been documented in western North America, including Utah and Mexico.
treehopper ant-mutualism Nearctic Membracidae Polyglyptini honeydew
Pulvinaria floccifera
Cottony Camellia Scale, Cottony Yew Scale, Cottony Taxus Scale
Pulvinaria floccifera is a soft scale insect (Coccidae) known for producing conspicuous white, cottony ovisacs that cover eggs. It is a significant pest of woody ornamentals including camellia, holly, yew, and tea plants. The species has a broad global distribution spanning temperate and subtropical regions. Females are sessile and wingless, while males develop wings for mating. Heavy infestations produce honeydew that supports sooty mold growth, reducing plant vigor and aesthetic value.
scale-insect soft-scale pest ornamental tea-pest Coccidae Sternorrhyncha honeydew sooty-mold biological-control-host
Sarucallis
crape myrtle aphid
Sarucallis is a monotypic genus of aphids in the family Aphididae. The sole described species, Sarucallis kahawaluokalani, is commonly known as the crape myrtle aphid. Adults measure 1.2 to 1.8 millimeters in body length. The genus is native to Asia and has become widely distributed globally through the ornamental plant trade, tracking the spread of its host plant, crape myrtle (Lagerstroemia spp.).
aphid pest ornamental invasive monophagous honeydew sooty-mold integrated-pest-management urban-ecology
Sternorrhyncha
plant-parasitic hemipterans
Sternorrhyncha is a suborder of Hemiptera containing aphids, whiteflies, scale insects, and psyllids—groups formerly classified in the obsolete order 'Homoptera'. The name refers to the rearward position of the mouthparts relative to the head. All members are obligate plant-feeders with piercing-sucking mouthparts adapted for extracting phloem or xylem sap. Many species exhibit highly modified life cycles including parthenogenesis, sexual dimorphism, flightless morphs, and eusociality. The group includes numerous economically important agricultural and horticultural pests.
Hemiptera plant-feeding sap-sucking agricultural-pests honeydew parthenogenesis sexual-dimorphism invasive-species biological-control
Tapinoma
Tapinoma is a moderately diverse genus of ants in the subfamily Dolichoderinae, comprising approximately 87 described species (81 extant, 6 fossil) with a crown age estimated at 34–49 million years. Species occur worldwide in tropical and temperate regions, exhibiting generalized foraging behavior and flexible nesting habits. Several species, including T. sessile and T. melanocephalum, are notable as household pests and invasive species associated with human disturbance.
ants Dolichoderinae invasive-species household-pest honeydew trophobiosis urban-adaptation tramp-species polygyny unicoloniality
Toumeyella liriodendri
tuliptree scale
Toumeyella liriodendri, the tuliptree scale, is a soft scale insect (Coccidae) native to North America and specialized on Liriodendron tulipifera (tuliptree or yellow-poplar). This univoltine species overwinters as second-instar nymphs on twigs, matures to adults in spring, and produces eggs that hatch into crawlers in early summer. First-instar nymphs feed on leaf undersides before migrating back to twigs in autumn. Males are unknown; reproduction is presumed parthenogenetic. Heavy infestations cause branch dieback, reduced growth, and honeydew production that supports sooty mold growth. The species has been the subject of recent genomic research, with a 536Mb genome assembly revealing it as an early-diverging soft scale with 17 chromosomes.
scale-insect soft-scale Coccidae Liriodendron tuliptree yellow-poplar univoltine parthenogenesis honeydew sooty-mold forest-pest ornamental-pest genome chromosome-evolution phloem-feeder ant-tended North-America Maryland Hemiptera Sternorrhyncha
Uroleucon taraxaci
Bronze-brown Dandelion Aphid
Uroleucon taraxaci is a species of aphid in the family Aphididae, commonly known as the Bronze-brown Dandelion Aphid. As a member of the genus Uroleucon, it shares characteristics with related species that exhibit parthenogenetic reproduction during spring and summer, forming all-female colonies. The species is associated with plants in the Asteraceae family, particularly dandelion (Taraxacum). Like other Uroleucon aphids, it produces honeydew that attracts predators and parasitoids. The species has been documented across parts of Europe and Asia.
aphid Hemiptera Sternorrhyncha Aphididae Uroleucon dandelion Taraxacum parthenogenesis collective-defense honeydew biological-control predator-prey Asteraceae Bronze-brown-Dandelion-Aphid