Honeydew
Guides
Aetalionidae
Aetalionid Treehoppers
Aetalionidae is a family of treehoppers in the superfamily Membracoidea, distinguished from the more diverse Membracidae by morphological features including a front femur fused to the trochanter and a completely exposed scutellum. The family is predominantly Neotropical in distribution, with two recognized subfamilies: Biturritiinae (exclusively Neotropical) and Aetalioninae (containing the Neotropical genus Aetalion and the Paleotropical genus Darthula). Females exhibit maternal care by guarding egg masses, and many species form aggregations on tree branches where they produce honeydew and associate with ants and stingless bees.
Aphidina
Aphidina is a subtribe of aphids within the family Aphididae, established by Latreille in 1802. It encompasses numerous genera including economically significant and xerophilous species. Members exhibit diverse life cycles, with some species amenable to genetic control methods due to their holocyclic heterogenetic reproduction. The subtribe includes species with specialized host associations, particularly on Asteraceae, Chenopodiaceae, and Amaranthaceae.
Aphidoidea
aphids, plant lice, greenflies, blackflies, whiteflies
Aphidoidea is a superfamily of small sap-sucking insects within Hemiptera, commonly known as aphids. Members possess piercing-sucking mouthparts adapted for feeding on plant phloem. The superfamily includes approximately 5,000 described species across multiple families, with Aphididae being the largest and most economically significant. Many species exhibit complex life cycles involving both sexual and parthenogenetic reproduction, often with seasonal alternation between host plants.
Aphis
Aphis is a large genus of aphids in the family Aphididae containing at least 400–600 species. The genus includes numerous significant agricultural pests, notably the soybean aphid (Aphis glycines), cotton aphid (Aphis gossypii), and apple aphid (Aphis pomi). Many species form mutualistic associations with ants (myrmecophily), exchanging honeydew for protection. Species identification within the genus often requires examination of morphological characters such as siphunculi and cauda structure.
Balcanocerus
Balcanocerus is a genus of leafhoppers in the family Cicadellidae, subfamily Eurymelinae. Species in this genus are ant-attended and feed on phloem sap from host plants in Rosaceae. The genus includes both North American and European species, with documented associations with specific ant partners and host trees.
Bolbonota
seed-mimic treehopper
Bolbonota is a genus of treehoppers in the family Membracidae, tribe Membracini, distributed in the New World tropics and subtropics. Adults are characterized by their dark, globular body form with a corrugated pronotal surface and distinctive red eyes. The genus contains at least a dozen described species in Brazil alone, with many more likely awaiting description. Bolbonota species exhibit complex ontogenetic color changes: early instar nymphs are dark, late instars become mottled or streaked white, and adults revert to dark coloration.
Ceroplastes
wax scales
Ceroplastes is a genus of wax scales in the family Coccidae, comprising more than 130 described species. These insects are characterized by the conspicuous white, waxy coverings they secrete, which protect their soft bodies. Adults are largely sessile and feed on plant sap using piercing-sucking mouthparts. Many species are economically significant pests of ornamental plants, fruit trees, and shrubs. The genus exhibits parthenogenesis in many species, with females reproducing without males. Climate change has facilitated the northward expansion of several species from southern regions into previously cooler areas.
Ceroplastes cirripediformis
barnacle scale, barnacle wax scale
Ceroplastes cirripediformis, commonly known as the barnacle scale or barnacle wax scale, is a soft scale insect (Hemiptera: Coccidae) recognized for its distinctive waxy covering that resembles a small barnacle. It is a destructive invasive pest with a broad host range spanning numerous plant families. The species has expanded its range globally through human-mediated transport and climate change, with established populations across North America, South America, Europe, Asia, Africa, and Oceania. Females reproduce parthenogenetically, producing up to a thousand eggs annually without observed males in wild populations.
Cinara
giant conifer aphids, conifer aphids
Cinara is a genus of aphids comprising approximately 200-243 species, commonly known as giant conifer aphids. These insects are specialized feeders on conifers in the families Pinaceae and Cupressaceae, including pines, firs, spruces, cedars, and cypresses. Molecular studies indicate their tribe Eulachnini represents the basal group for other aphids. Several species are significant pests of cultivated Christmas trees, particularly Fraser fir production in the southeastern United States.
Cinara pergandei
Globose Pine Aphid
Cinara pergandei is a large aphid species in the family Aphididae, commonly known as the Globose Pine Aphid. It feeds on coniferous trees, particularly pines, and produces copious amounts of honeydew that attracts wasps and other insects. Like other Cinara species, it has piercing-sucking mouthparts adapted for tapping into phloem sap. It is part of a genus containing numerous conifer-feeding aphids, many of which are significant pests in forestry and Christmas tree production.
Cinara strobi
White Pine Aphid
Cinara strobi, the white pine aphid, is a large aphid species specialized on white pine (Pinus strobus). It is notable for exceptionally high autumn populations that produce copious honeydew, attracting stinging wasps and creating nuisance conditions. The species exhibits parthenogenetic reproduction during the growing season, with live birth of nymphs accelerating population growth. Uniquely among Cinara aphids, C. strobi harbors three bacterial endosymbionts—Buchnera aphidicola, Sodalis sp., and Serratia symbiotica—representing an intermediate stage of symbiont replacement where the latter has become metabolically incompetent yet persists.
Coccus viridis
green scale, green coffee scale, coffee green scale
Coccus viridis is a soft scale insect (Hemiptera: Coccidae) and major agricultural pest with a wide tropical and subtropical distribution. It reproduces parthenogenetically via thelytoky, with females producing 50-600 eggs that hatch within minutes to hours beneath the mother's body. The species completes its life cycle in 47-51 days at 25°C, with three nymphal instars before adulthood; first-instar nymphs actively disperse as 'crawlers' while older instars and adults become sessile. It is polyphagous, feeding on phloem sap of over 200 plant genera in 72 families, with primary economic damage to coffee, guava, and cacao crops.
Disholcaspis prehensa
clasping twig gall wasp
Disholcaspis prehensa is a gall-inducing cynipid wasp native to California that induces distinctive galls on scrub oak and leather oak. The species exhibits heterogony, alternating between sexual and asexual generations that produce two morphologically different gall types. The asexual generation forms conspicuous mushroom-shaped twig galls with clasping bases, while the sexual generation produces small, hidden bud galls.
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Dolichoderus is a genus of ants comprising 181 valid species (131 extant, 50 extinct), representing the sole member of tribe Dolichoderini and type genus of subfamily Dolichoderinae. Workers are small (approximately 4 mm), possess a thick sculptured integument, and lack a functional sting. The genus exhibits remarkable ecological diversity: most species are generalist honeydew collectors that tend aphids and scale insects, while the D. cuspidatus group in Southeast Asia practices unique nomadic aphid farming with obligate mutualist mealybugs. The genus has an extensive fossil record in Baltic amber.
Dolichoderus mariae
Mary's Odorous Ant
Dolichoderus mariae is a distinctive North American ant species named after naturalist Mary Treat, who discovered it. The species is notable for its reddish-brown and black bicolored appearance, large polydomous colonies with dozens of nests, and its specialized ecology in prairie and glade habitats where it nests among plant roots. Colonies exhibit dramatic seasonal fluctuations in nest numbers and worker populations, and engage in territorial aggression with neighboring colonies.
Dolichoderus taschenbergi
Taschenberg's Odorous Ant
Dolichoderus taschenbergi is a North American ant species in the odorous ant subfamily Dolichoderinae. Described by Mayr in 1866, it is endemic to Canada and the United States. The species exhibits sporadic dispersion patterns and has been studied for its mutualistic relationships with honeydew-producing hemipterans. Colony structure may include polygynous nesting patterns, though this requires further research.
Entylia
keel-shaped treehopper
Entylia is a genus of treehoppers in the family Membracidae, first described by Germar in 1833. The genus contains at least three described species, including Entylia carinata, which is known for its distinctive keel-shaped pronotum that resembles plant debris. These insects are notable for their mutualistic relationships with ants, which protect them from predators in exchange for honeydew. The genus is distributed in North America, with observations recorded from regions including the northeastern United States.
Eriosoma americanum
Woolly Elm Aphid
Eriosoma americanum, commonly known as the woolly elm aphid, is a North American aphid species that alternates between two host plants: American elm (Ulmus americana) and Saskatoon (Amelanchier alnifolia). The species produces distinctive woolly or cottony wax secretions that give it its common name. Heavy infestations on American elm cause leaf curling and produce visible dark patches and cottony masses. Like other aphids, it excretes honeydew while feeding on plant sap.
Eulachnini
Eulachnini is a tribe of aphids within the subfamily Lachninae, comprising approximately 27 species documented in Lithuania alone. Members are specialized phloem-feeders on coniferous plants, particularly Pinaceae. Several species produce substantial honeydew, with five noted as promising honeydew producers in Lithuanian forests. Only Cinara piceae has been identified as a potential forest pest, with documented damage to firs in nursery settings.
Formica incerta
Uncertain Field Ant
Formica incerta is a common ant species of eastern North America, often the most abundant Formica in its range. It constructs underground nests with inconspicuous entrance holes and no surface mound. Workers forage diurnally for nectar and honeydew, and the species serves as the primary host for the slave-making ant Polyergus lucidus. First described by Carlo Emery in 1893, its specific epithet reflects historical taxonomic uncertainty and identification challenges.
Formica integroides
Western Mound Ant, Vinegar Ant
Formica integroides, commonly known as the Western Mound Ant or Vinegar Ant, is a North American wood ant species in the family Formicidae. This species exhibits flexible, context-dependent foraging strategies at both colony and individual levels, adjusting its behavior based on resource characteristics and competitive pressure. It is known to utilize both terrestrial and arboreal resources, with foragers capable of defending entire trees as absolute territories. The species was first described by Wheeler in 1913.
Homotomidae
fig psyllids, mulberry psyllids
Homotomidae is a family of small phloem-feeding insects in the superfamily Psylloidea, commonly known as fig or mulberry psyllids. Members feed almost exclusively on Moraceae, primarily the genus Ficus, with rare exceptions such as Triozamia on Antiaris toxicaria. The family contains approximately 70 described species across three subfamilies: Dynopsyllinae, Macrohomotominae, and Homotominae. Several species are economically significant pests of ornamental and cultivated figs, including the invasive Macrohomotoma gladiata, which has spread from Asia to Europe, North Africa, and North America.
Liometopum apiculatum
Escamolera Ant, Velvet Tree Ant
Liometopum apiculatum is a Neotropical dolichoderine ant with significant ecological and economic importance. Colonies are large and polydomous, ranging from a few hundred to hundreds of thousands of workers. The species is notable for its distinctive carton-like nest construction and its role as a traditional food source in Mexico, where the larvae and pupae (escamoles) are harvested. It inhabits diverse arid and semi-arid vegetation types across southwestern North America.
Melanaphis
sugarcane aphid
Melanaphis is a genus of aphids in the family Aphididae, first described by van der Goot in 1917. The genus includes species that are significant agricultural pests, particularly of sorghum and sugarcane. Melanaphis sacchari (sugarcane aphid) has become one of the most damaging insect pests of grain and forage sorghum in the United States since its first recorded occurrence on sorghum in Texas in 2013. Members of this genus are characterized by their parthenogenetic reproduction, rapid population growth, and production of honeydew.
Membracidae
treehoppers, thorn bugs, typical treehoppers
Membracidae, commonly called treehoppers or thorn bugs, is a family of approximately 3,200–3,500 species in over 400 genera within the order Hemiptera. The family is distinguished by extraordinary morphological diversity, particularly the pronounced enlargement and modification of the pronotum—the dorsal plate of the first thoracic segment—which can form thorns, horns, keels, or bizarre projections that often obscure the body and wings. This family represents the most diverse lineage within the superfamily Membracoidea, with its center of diversity in the New World tropics. Most species exhibit phytophagous habits, feeding on plant sap with piercing-sucking mouthparts. Many species engage in mutualistic relationships with ants, which protect them from predators in exchange for honeydew.
Metcalfa
Metcalfa is a genus of planthoppers in the family Flatidae, tribe Nephesini, native to North and Central America. The genus contains at least five described species, with Metcalfa pruinosa (citrus flatid planthopper) being the most economically significant due to its invasive spread to Europe and Asia. Adults are small, ranging 5.5–8 mm in length with a waxy, often bluish or grayish appearance. The genus is characterized by forewings with parallel cross-veins along the costal margin, distinguishing it from related families.
Microterys
Microterys is a large genus of parasitoid wasps in the family Encyrtidae (Chalcidoidea), with its center of distribution in the northern parts of the Northern Hemisphere. Species are important natural enemies of various scale insects (Coccoidea), including soft scales (Coccidae), wax scales (Ceroplastes), and mealybugs (Pseudococcidae). The genus has been extensively studied for biological control applications, particularly for managing pest scale insects on citrus and other crops. Several species have been introduced to new regions as biocontrol agents, including Microterys flavus in California.
parasitoidbiological-controlscale-insectCoccoideaEncyrtidaeChalcidoideaHymenopteraovipositorintegrated-pest-managementcitrus-pestsoft-scalemealybughyperparasitoidhost-specificitybiotypeclassical-biological-controlsynchrotron-microtomographyoviposition-behaviornatural-enemyagricultural-entomologytaxonomyGiraultAustraliaChinaCaliforniaSyriaIndiaCzech-RepublicEuropeAsiaNorth-AmericaSouthern-HemisphereCeroplastesCoccusPhenacoccusKermesParthenolecaniumMetaceronemaChaetococcusKerriaLacciferGascardiaSphaerolecaniumCerococcusChloropulvinariapest-managementhost-selectionhost-penetrationenvenomationegg-depositionhost-feedingvalvifervalvulamusculoskeletal-system3D-morphologySEMSR-µCTinvasive-speciesbiocontrol-introductionhyperparasitismefficiency-reductionnatural-enemy-conservationcitrusCamelliaMagnoliafruit-cropsornamental-plantsforestrylac-insectwax-scaleplum-scalecottony-maple-scaleEuropean-fruit-lecaniumbrown-soft-scalecitricola-scalecitrus-snow-scaleJapanese-wax-scaletype-materialspecies-descriptionredescriptionidentification-keyintraspecific-variationdiagnostic-charactersNorthern-Hemispherecenter-of-distributioncosmopolitannew-speciesnew-recordfaunisticsmuseum-collectionUCRTimberlakeCompereGordhTriapitsynNoyesXuSugonjaevPrinslooRosenIshiiMercetDalmanHowardAshmeadWalkerFerrièreKerrichAnneckeMynhardtHayatSubba-RaoTrjapitzinViggianiDe-SantisMayrThomsonWestwoodGahanCrawfordBlanchardPerkinsRisbecBruesBennettErdösSilvestriDomenichiniDozierCockerellCoquilletFonscolombeRatzeburgMatsumuraVassiljevYoshimotoGhesquièreTachikawaLogvinovskayaSiscaroAgarwalAhmadGhaniBurksGirault-speciesAustralian-faunaChinese-faunaIndian-faunaSyrian-faunaEuropean-faunaCalifornian-faunaglobal-distributioneconomic-entomologyapplied-entomologysystematicsmorphologybehavioral-ecologyfunctional-morphologyevolutionary-successdiverse-groupunderstudiedminute-waspsmicro-Hymenopteraslide-collectionvoucher-specimenshost-recordsundetermined-specimensvalid-generasynonymymanuscript-namesnomina-nudaunpublished-namesre-organizationcuratorhistorical-notescollection-growthcollecting-activitiesidentificationsortingdrawerspointsslidesparatypesbackbonebiological-control-projectsCalifornia-entomologistsCitrus-Experiment-StationHawaiian-collectionbee-taxonomysystematistsstudentsstaffworldlargestimportantvaluemere-numbertaxa-representedreportedscientific-literatureconductedaugmentedcontributionscountriesrapidlygrowingcompletelistpresenteditalicizedmiscellaneousundeterminedfull-drawersunsortedspecimens-per-slidemore-than-500-slidesat-least-700-slidessometimes-more-than-one-specimen-per-slidesynchrotron-X-ray-phase-contrast-microtomographyscanning-electron-microscopyin-vivo-documentation3D-modelmicrostructurescuticular-elementsmotion-patternsphasesparasitizationworking-mechanismkey-traitadaptive-evolutionreproductive-successmechanical-aspectsfunctional-aspectsfully-understoodenormously-diversespecializedparasitisingplant-pestsecological-roleeconomic-roleimprove-understandingmechanicsmode-of-functiondetailedanalysisidentifiedconsistingrecently-discovereddescribedelucidatedpenetrationassessmentinternal-organsdevelopedunderlyingobservedcontributinghighly-diverse-groupchalcidoid-waspsFrontiers-in-ZoologyActa-Musei-SilesiaeArab-Journal-for-Plant-ProtectionAustralian-Journal-of-EntomologyInsect-ScienceOriental-InsectsJournal-of-Natural-HistoryPhytoparasiticaAnnals-of-the-Entomological-Society-of-AmericaEnvironmental-EntomologyGBIFCatalogue-of-LifeNCBI-TaxonomyiNaturalistWikipediaDOI10.1186/s12983-025-00575-110.2478/cszma-2021-000710.22268/AJPP-41.3.29230510.1111/j.1440-6055.1975.tb02058.x10.1111/j.1744-7917.2000.tb00345.x10.1080/00305316.2011.64682710.1111/j.1440-6055.1973.tb01670.x10.1080/00222933.2020.178660810.1007/s12600-019-00727-010.1007/s12600-018-0691-510.1093/aesa/54.2.22210.1093/ee/7.6.87420252021202319752000201119732020201920181950s1960s1876199920162017AprilJuneFebruaryNorth-MoraviaNE-Czech-RepublicMoravskoslezské-Beskydy-MtsMorávkaTurkmenistanCentral-AsiaLattakia-ProvinceAl-SanobarDabbaSyrian-coastDamascus-UniversityBiological-Control-Studies-and-Research-CenterBCSRCFaculty-of-AgricultureQueensland-MuseumZhejiangFujianYunnanGuangdongNortheast-ChinaWestern-AustraliaJilinfirst-recordnew-to-sciencefirst-timeadded-to-faunareviewednotesillustrationshabitusredescriptionsclose-relationshipdiscussedkey-providedfemalesmalestransferredextentvalidityevaluatedcorrectionbehaviorresponsehoneydewtaxonomy-matchhigher-rankacceptedcanonical-namerankstatusmatch-typeclassificationEukaryotaHexapodaApocritaTerebrantesEncyrtinaeobservations-countnoneevidencepaper-summaryconfidence-notesinferredexplicitly-statedvisible-textfull-paperaccessibleprovided-contentcited-referencesdetailed-biologytype-localitynative-rangegenerallimited-tobased-onno-information-providedabstractgenus-level-biologyfamily-level-biologydefinitivefull-text-requiredinferred-fromJaposhvili-1999Yasnosh-&-Japoshvili-1998high-hyperparasitoids-levelsreduced-efficiencynaturally-occurringmicroscopic-imagesmorphologicalmorpho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glaber
black household ant, black house ant, tramp ant, Copper-bellied Ant
Ochetellus glaber is a small ant species native to Australia, widely introduced globally through human commerce. Workers measure 2–3 mm, with queens reaching 5.2–5.5 mm. The species is recognized as a species complex with cryptic diversity, complicating identification and management decisions. It nests arboreally and in structures, forming conspicuous trails to exploit honeydew and food resources. Considered a household pest, it has established populations in Hawaii, Florida, New Zealand, and multiple Asian and Pacific regions.
Oncometopia
sharpshooters
Oncometopia is a genus of sharpshooters (leafhoppers in the family Cicadellidae) found in North and South America, erected by Carl Stål in 1869. Species range from 9.4 to 15.0 mm in length and are characterized by distinctive morphological features including a moderately produced head, coriaceous forewings with apical membranes, and specialized hindleg tarsomere proportions. Several species are economically important as vectors of plant pathogens, particularly Xylella fastidiosa, which causes diseases such as phony peach disease and citrus variegated chlorosis.
Oncometopia hamiltoni
Oncometopia hamiltoni is a species of sharpshooter leafhopper in the family Cicadellidae, described by Rakitov in 2016. It belongs to the tribe Proconiini, a group of large, active leafhoppers known for their xylem-feeding habits and distinctive "raining" behavior caused by forceful excretion of excess fluid. The species is part of a genus containing several economically significant sharpshooters that have been studied for their role in plant disease transmission and their interactions with egg parasitoids.
Oncometopia orbona
Broad-headed Sharpshooter
Oncometopia orbona, commonly known as the broad-headed sharpshooter, is a large leafhopper in the family Cicadellidae. It feeds on xylem sap from plants, processing large volumes of nutrient-poor fluid and excreting the excess as honeydew—a behavior that creates the phenomenon of 'raining trees.' The species possesses distinctive white chalky brochosomes on its sides, used to powder egg-laying sites. It is a strong flier with excellent eyesight and serves as a host for egg parasitoids in the family Mymaridae.
Panaphis juglandis
Walnut Aphid, Large Walnut Aphid
Panaphis juglandis, the large walnut aphid, is a phloem-feeding pest of walnut trees (Juglans regia). It colonizes exclusively the upper (adaxial) surface of walnut compound leaves, forming dense colonies along the main vascular veins of the outer leaflets. Native to the Old World (Central Asia and southeastern Europe), it has spread across Europe and was introduced to California in the early 20th century. The species reduces tree vigor, nut yield, and quality, and excretes honeydew that promotes sooty mold growth. It forms mutualistic relationships with ant species that tend and protect colonies from predators.
Petrusa
Petrusa is a genus of planthoppers in the family Flatidae, subfamily Flatinae. Members of this genus are found in the Neotropical region, with Petrusa epilepsis being a notable species associated with seagrape and other host plants in the Caribbean. The genus is characterized by the typical flatid morphology with wings held tent-like over the body and parallel cross-veins along the costal margin of the forewings.
Polistes dominula
European Paper Wasp
Polistes dominula is a highly successful invasive social wasp native to Eurasia that has established populations across North America, South America, New Zealand, South Africa, and other regions. First detected in North America near Boston in 1978, it has become one of the most abundant wasps on the continent. The species builds small, exposed paper nests in protected locations and preys primarily on live insects, particularly caterpillars. Unlike yellowjackets, it does not scavenge for meat or sugar. Its rapid spread has been attributed to ecological flexibility, superior competitive ability, and tolerance of human-altered environments.
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Publilia is a genus of treehoppers (Membracidae) native to North America, with species occurring primarily in the eastern and central United States. The genus is notable for forming dense aggregations on host plants, particularly goldenrods (Solidago spp.), where nymphs and adults are tended by ants in exchange for honeydew. This mutualistic relationship has made Publilia a model system for studying ant-hemipteran interactions and their ecological consequences for host plants. The genus is distinguished from related treehoppers by its relatively unadorned pronotum compared to the elaborate projections seen in many Neotropical membracids.
Publilia porrecta
Publilia porrecta is a Nearctic treehopper species in the family Membracidae, tribe Polyglyptini. The genus Publilia is known for ant-tended mutualistic associations, with multiple ant species frequently visiting aggregations. Like other members of its tribe, this species exhibits subsocial behaviors and is associated with herbaceous host plants. The species has been documented in western North America, including Utah and Mexico.
Pulvinaria floccifera
Cottony Camellia Scale, Cottony Yew Scale, Cottony Taxus Scale
Pulvinaria floccifera is a soft scale insect (Coccidae) known for producing conspicuous white, cottony ovisacs that cover eggs. It is a significant pest of woody ornamentals including camellia, holly, yew, and tea plants. The species has a broad global distribution spanning temperate and subtropical regions. Females are sessile and wingless, while males develop wings for mating. Heavy infestations produce honeydew that supports sooty mold growth, reducing plant vigor and aesthetic value.
Sarucallis
crape myrtle aphid
Sarucallis is a monotypic genus of aphids in the family Aphididae. The sole described species, Sarucallis kahawaluokalani, is commonly known as the crape myrtle aphid. Adults measure 1.2 to 1.8 millimeters in body length. The genus is native to Asia and has become widely distributed globally through the ornamental plant trade, tracking the spread of its host plant, crape myrtle (Lagerstroemia spp.).
Sternorrhyncha
plant-parasitic hemipterans
Sternorrhyncha is a suborder of Hemiptera containing aphids, whiteflies, scale insects, and psyllids—groups formerly classified in the obsolete order 'Homoptera'. The name refers to the rearward position of the mouthparts relative to the head. All members are obligate plant-feeders with piercing-sucking mouthparts adapted for extracting phloem or xylem sap. Many species exhibit highly modified life cycles including parthenogenesis, sexual dimorphism, flightless morphs, and eusociality. The group includes numerous economically important agricultural and horticultural pests.
Tapinoma
Tapinoma is a moderately diverse genus of ants in the subfamily Dolichoderinae, comprising approximately 87 described species (81 extant, 6 fossil) with a crown age estimated at 34–49 million years. Species occur worldwide in tropical and temperate regions, exhibiting generalized foraging behavior and flexible nesting habits. Several species, including T. sessile and T. melanocephalum, are notable as household pests and invasive species associated with human disturbance.
Toumeyella liriodendri
tuliptree scale
Toumeyella liriodendri, the tuliptree scale, is a soft scale insect (Coccidae) native to North America and specialized on Liriodendron tulipifera (tuliptree or yellow-poplar). This univoltine species overwinters as second-instar nymphs on twigs, matures to adults in spring, and produces eggs that hatch into crawlers in early summer. First-instar nymphs feed on leaf undersides before migrating back to twigs in autumn. Males are unknown; reproduction is presumed parthenogenetic. Heavy infestations cause branch dieback, reduced growth, and honeydew production that supports sooty mold growth. The species has been the subject of recent genomic research, with a 536Mb genome assembly revealing it as an early-diverging soft scale with 17 chromosomes.
Uroleucon taraxaci
Bronze-brown Dandelion Aphid
Uroleucon taraxaci is a species of aphid in the family Aphididae, commonly known as the Bronze-brown Dandelion Aphid. As a member of the genus Uroleucon, it shares characteristics with related species that exhibit parthenogenetic reproduction during spring and summer, forming all-female colonies. The species is associated with plants in the Asteraceae family, particularly dandelion (Taraxacum). Like other Uroleucon aphids, it produces honeydew that attracts predators and parasitoids. The species has been documented across parts of Europe and Asia.