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Disholcaspis prehensa
clasping twig gall wasp
Disholcaspis prehensa is a gall-inducing cynipid wasp native to California that induces distinctive galls on scrub oak and leather oak. The species exhibits heterogony, alternating between sexual and asexual generations that produce two morphologically different gall types. The asexual generation forms conspicuous mushroom-shaped twig galls with clasping bases, while the sexual generation produces small, hidden bud galls.
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ulicis
Gorse Seed Weevil
Exapion ulicis is a small seed-feeding weevil specialized on gorse (Ulex europaeus). Adults are light gray with a prominent snout roughly half the body length. The species is native to western Europe and has been introduced to New Zealand, California, Hawaii, and other regions as a biological control agent targeting invasive gorse populations. Larval feeding destroys seeds within pods, reducing plant spread, while adult feeding on stems and spines causes minor damage.
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-collectively-retentive-Pareto-optimal-setminimal-individually-retentive-Pareto-optimal-setminimal-Nash-retentive-Pareto-optimal-setminimal-individually-rational-retentive-Pareto-optimal-setminimal-weakly-stable-Pareto-optimal-setminimal-strongly-stable-Pareto-optimal-setminimal-collectively-stable-Pareto-optimal-setminimal-individually-stable-Pareto-optimal-setminimal-Nash-stable-Pareto-optimal-setminimal-individually-rational-stable-Pareto-optimal-setminimal-weakly-Nash-stable-Pareto-optimal-setminimal-strongly-Nash-stable-Pareto-optimal-setminimal-collectively-Nash-stable-Pareto-optimal-setminimal-individually-Nash-stable-Pareto-optimal-setminimal-Nash-Nash-stable-Pareto-optimal-setminimal-individually-rational-Nash-stable-Pareto-optimal-setminimal-weakly-individually-rational-Pareto-optimal-setminimal-strongly-individually-rational-Pareto-optimal-setminimal-collectively-individually-rational-Pareto-optimal-setminimal-individually-individually-rational-Pareto-optimal-setminimal-Nash-individually-rational-Pareto-optimal-setminimal-individually-rational-individually-rational-Pareto-optimal-setminimal-weakly-Pareto-optimal-Nash-stable-setminimal-strongly-Pareto-optimal-Nash-stable-setminimal-collectively-Pareto-optimal-Nash-stable-setminimal-individually-Pareto-optimal-Nash-stable-setminimal-Nash-Pareto-optimal-Nash-stable-setminimal-individually-rational-Pareto-optimal-Nash-stable-setminimal-weakly-comprehensive-Pareto-optimal-Nash-stable-setminimal-strongly-comprehensive-Pareto-optimal-Nash-stable-setminimal-collectively-comprehensive-Pareto-optimal-Nash-stable-setminimal-individually-comprehensive-Pareto-optimal-Nash-stable-setminimal-Nash-comprehensive-Pareto-optimal-Nash-stable-setminimal-individually-rational-comprehensive-Pareto-optimal-Nash-stable-setminimal-weakly-extensive-Pareto-optimal-Nash-stable-setminimal-strongly-extensive-Pareto-optimal-Nash-stable-setminimal-collectively-extensive-Pareto-optimal-Nash-stable-setminimal-individually-extensive-Pareto-optimal-Nash-stable-setminimal-Nash-extensive-Pareto-optimal-Nash-stable-setminimal-individually-rational-extensive-Pareto-optimal-Nash-stable-setminimal-weakly-retentive-Pareto-optimal-Nash-stable-setminimal-strongly-retentive-Pareto-optimal-Nash-stable-setminimal-collectively-retentive-Pareto-optimal-Nash-stable-setminimal-individually-retentive-Pareto-optimal-Nash-stable-setminimal-Nash-retentive-Pareto-optimal-Nash-stable-setminimal-individually-rational-retentive-Pareto-optimal-Nash-stable-setminimal-weakly-stable-Pareto-optimal-Nash-stable-setminimal-strongly-stable-Pareto-optimal-Nash-stable-setminimal-collectively-stable-Pareto-optimal-Nash-stable-setminimal-individually-stable-Pareto-optimal-Nash-stable-setminimal-Nash-stable-Pareto-optimal-Nash-stable-setminimal-individually-rational-stable-Pareto-optimal-Nash-stable-setminimal-weakly-Nash-stable-Pareto-optimal-Nash-stable-setminimal-strongly-Nash-stable-Pareto-optimal-Nash-stable-setminimal-collectively-Nash-stable-Pareto-optimal-Nash-stable-setminimal-individually-Nash-stable-Pareto-optimal-Nash-stable-setminimal-Nash-Nash-stable-Pareto-optimal-Nash-stable-setminimal-individually-rational-Nash-stable-Pareto-optimal-Nash-stable-setminimal-weakly-individually-rational-Pareto-optimal-Nash-stable-setminimal-strongly-individually-rational-Pareto-optimal-Nash-stable-setminimal-collectively-individually-rational-Pareto-optimal-Nash-stable-setminimal-individually-individually-rational-Pareto-optimal-Nash-stable-setminimal-Nash-individually-rational-Pareto-optimal-Nash-stable-setminimal-individually-rational-individually-rational-Pareto-optimal-Nash-stable-setminimal-weakly-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-weakly-comprehensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-comprehensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-comprehensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-comprehensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-comprehensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-comprehensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-weakly-extensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-extensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-extensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-extensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-extensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-extensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-weakly-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-weakly-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-weakly-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-weakly-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setTortopus
Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.
EphemeropteraPolymitarcyidaeburrowing-mayfliesNeotropicalclay-bank-habitatU-shaped-tunnelsmandibular-tusksTortopsisCampsurinaePanamerican-distributioncladistic-revisionsynapomorphyparastyli-receptorsflattened-penesfurcasternal-protuberancessubapical-tubercleslimnological-sampling-biasnymph-morphologyadult-morphologytype-species-T.-igaranusnew-species-T.-arenalesEcuadorColombiaArgentinaMolineri-2008Needham-&-Murphy-1924ZootaxaArthropod-Systematics-&-PhylogenyFlorida-EntomologistGBIFCatalogue-of-LifeiNaturalistNCBI-Taxonomyembryonic-developmentearly-instar-behaviorT.-harrisiT.-bellusT.-circumfluusT.-zottaiT.-igaranusT.-incertusT.-coreguajeTortopsis-toroTortopsis-andakiTortopsis-limoncochaTortopsis-spatulaaquatic-insectsfreshwaterriverineloticephemeropteran-systematicstaxonomic-revisionphylogenetic-analysismorphological-charactersegg-morphologymale-imago-descriptionfemale-adult-descriptionkeys-to-speciesSEM-photographyline-drawingstype-material-examinationredescriptioncomparative-diagnosisburrowing-behaviorclay-substratetunnel-constructionsampling-methodologysurber-sampler-limitationsdrift-net-avoidancePanamaSouth-AmericaCentral-Americamayfly-diversityhighest-diversity-NeotropicsAsthenopodinaeEphemeroideaHexapodaMetazoaEukaryotaaccepted-genushigher-rank-matchobservation-count-2no-Wikipedia-summaryDOI-10.11646/zootaxa.2481.1.1DOI-10.11646/zootaxa.2436.1.4DOI-10.3897/asp.79.e62735DOI-10.2307/3493493Scott-et-al.-1959NavásUlmerTraverLugo-Ortiz-&-McCaffertyDomínguezBanksMcDunnoughPictetNeedham-&-Murphyfreshwater-ecologyaquatic-entomologyinsect-developmentnymphal-stagesimago-descriptionalate-adultswing-venationR-sectorgonopore-structureparastyli-lengthpedestal-morphologylateral-view-curvaturemandibular-morphologytusk-tuberclesmesosternal-structureabdominal-sternum-notchfurcasternumprotuberance-configurationreceptor-socket-orientationIR-veinR2-veinclaw-like-structuregonopore-associationpenis-base-separationfemale-fore-wingadditional-veinscomparative-morphologytype-specimenholotypeallotypeparatypeneotropical-speciesnew-species-descriptionspecies-keyidentification-keymale-adult-keyfemale-adult-keynymph-keygeneric-keyPolymitarcyidae-generasystematicsphylogenycladisticsmonophylysister-groupcharacter-statetransformation-seriesparsimony-analysismorphological-data-setexternal-morphologyegg-chorionmicropyleegg-bursterhatchingemergencebehavioral-studyearly-instarembryologyovipositionsubstrate-preferencehabitat-specificityendemicregional-diversityconservationbiomonitoringwater-quality-indicatormacroinvertebratebenthicinfaunalinterstitialhyporheicriver-bankstream-bankerosionsedimentparticle-sizeclay-contentbank-stabilityroot-matvegetation-associationflow-regimecurrent-velocitydepthwidthstream-ordercatchmentbasinecoregionbiogeographydispersalcolonizationvicariancespeciationadaptive-radiationmorphological-divergenceconvergent-evolutionhomoplasycharacter-conflictbootstrap-supportBremer-supportconsensus-treestrict-consensusmajority-ruletaxonomic-stabilitynomenclatural-actoriginal-descriptionsubsequent-designationtype-localitytype-depositorymuseum-collectionspecimen-preservationalcohol-preservationslide-mountingSEM-preparationimaging-techniquesscientific-illustrationphotographymeasurementmorphometricsratiosproportionsallometrysexual-dimorphismgeographic-variationintraspecific-variationinterspecific-variationdiagnostic-charactersidentification-confidencetaxonomic-expertiseliterature-reviewbibliographic-searchdatabase-queryonline-repositoryopen-accesspaywallabstract-onlyfull-text-unavailablecitation-analysisreference-listsource-qualityevidence-strengthknowledge-gapresearch-needfuture-studycollaborationfield-workcollection-tripsampling-effortspecimen-availabilitytype-examinationrevision-scopetaxonomic-conceptcircumscriptioninclusionexclusionspecies-groupspecies-complexcryptic-speciesmorphospeciesphylogenetic-speciesevolutionary-significant-unitoperational-taxonomic-unitDNA-barcodingmolecular-systematicsintegrative-taxonomytraditional-taxonomyalpha-taxonomybeta-taxonomygamma-taxonomyclassification-hierarchyLinnaean-systemrank-based-nomenclaturephylogenetic-nomenclatureclade-namecrown-groupstem-grouptotal-evidencecongruenceincongruencedata-partitioncharacter-weightingimplied-weightingequal-weightingordered-charactersunordered-charactersmultistate-charactersbinary-characterscontinuous-charactersmeristic-charactersqualitative-charactersquantitative-charactersmorphological-matrixdata-matrixcharacter-matrixtaxon-samplingcharacter-samplingmissing-datainapplicable-datapolymorphismintraspecific-polymorphismfixed-differencediagnostic-differenceunique-derived-characterautapomorphyshared-derived-charactersymplesiomorphyconvergent-characterparallel-evolutionreversalcharacter-optimizationACCTRANDELTRANunambiguous-optimizationambiguous-optimizationmost-parsimonious-treetree-lengthconsistency-indexretention-indexrescaled-consistency-indexhomoplasy-indexcharacter-fitcharacter-congruencecharacter-supportbranch-supportnode-supportclade-stabilitytaxonomic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