Reflex-bleeding

Guides

• Ceraeochrysa claveri

  trash bug (larval stage)

  A green lacewing in the family Chrysopidae, Ceraeochrysa claveri is a polyphagous predator with established value in biological control of agricultural pests. Larvae are "trash carriers" that construct dorsal camouflage packets using assorted materials including lichen, plant trichomes, and notably the wax of mealybugs and other insects. The species has been documented incorporating live mealybug nymphs into these packets, representing the first recorded instance of active placement of live prey onto the larval packet. Adults feed on pollen and may contribute to pollination. The species shows high genetic diversity across agroecosystems and is compatible with certain biopesticides including Bacillus thuringiensis serovar aizawai.

  biological-controltrash-carriermealybug-predatorintegrated-pest-managementcamouflage-behaviorphoresyreflex-bleedingpollinatoragroecosystemgenetic-diversitybiopesticide-compatibilityChrysopidaeNeuroptera

• Coccinella septempunctata

  seven-spot ladybird, seven-spotted ladybug, seven-spotted lady beetle, C-7

  Coccinella septempunctata is a predatory lady beetle native to Europe, Asia, and North Africa, now widely established in North America following intentional introduction for biological control. Adults measure 6.5–7.8 mm with distinctive red elytra bearing seven black spots. The species is among the most extensively studied coccinellids due to its importance in aphid biocontrol and its invasive success in new ranges. It completes up to five generations annually with a six-week development cycle from egg to adult.

  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• Exapion ulicis

  Gorse Seed Weevil

  Exapion ulicis is a small seed-feeding weevil specialized on gorse (Ulex europaeus). Adults are light gray with a prominent snout roughly half the body length. The species is native to western Europe and has been introduced to New Zealand, California, Hawaii, and other regions as a biological control agent targeting invasive gorse populations. Larval feeding destroys seeds within pods, reducing plant spread, while adult feeding on stems and spines causes minor damage.

  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rational-extensive-Pareto-optimal-individually-rational-Nash-stable-setminimal-weakly-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-retentive-Pareto-optimal-individually-rational-Nash-stable-setminimal-weakly-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-weakly-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-setminimal-weakly-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-strongly-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-collectively-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-Nash-individually-rational-Pareto-optimal-individually-rational-Nash-stable-setminimal-individually-rational-individually-rational-Pareto-optimal-individually-rational-Nash-stable-set

• Harmonia

  Greater Lady Beetles

  Harmonia is a genus of lady beetles (family Coccinellidae) commonly known as Greater Lady Beetles. The genus includes the widely introduced Multicolored Asian Lady Beetle (Harmonia axyridis), which has become established across North America and other regions as a biological control agent for aphids and other soft-bodied pests. Members of this genus are predatory beetles with highly variable coloration and spot patterns. The genus is native to Asia, with some species now distributed globally due to intentional introductions for pest management.

  lady-beetleladybugbiological-controlinvasive-speciesaphid-predatorhome-invaderCoccinellidaearboreal-predatorautumn-aggregationreflex-bleeding

• Harmonia dimidiata

  Greater Asian Lady Beetle

  Harmonia dimidiata is a medium to large predaceous ladybird beetle (7.4–10 mm) native to southern Asia with introduced populations in North America. Adults display reddish-yellow elytra with black markings in a thirteen-spot pattern. The species is a voracious aphid predator with documented prey including Rhopalosiphum maidis, Aphis gossypii, Myzus persicae, and Aphis pomi. Temperature strongly influences its development, fecundity, and predatory capacity, with optimal performance at 24°C. Males produce yellow reflex blood containing harmonine as both chemical defense and nuptial gift during courtship.

  lady-beetleladybirdpredatoraphidbiocontrolagricultureintroduced-speciesnuptial-giftreflex-bleedingtemperature-dependent-development

• Lytta

  blister beetles

  Lytta is a genus of blister beetles in the family Meloidae, established by Fabricius in 1775. The genus comprises approximately 70 described species in North America and over 100 species worldwide, making it one of the more diverse genera in its tribe. Species are distributed across the Americas, Europe, Asia, and Africa. Members are known for producing cantharidin, a defensive compound that causes blistering upon contact with skin.

  blister-beetlescantharidinSpanish-flyMeloidaedefensive-chemistryagricultural-pestreflex-bleedingchemosensory-genestriungulin-larvaebee-parasitismlocust-egg-predation

• Meloe impressus

  blister beetle, oil beetle

  Meloe impressus is a species of blister beetle in the family Meloidae, native to North America. It belongs to a genus commonly known as oil beetles, characterized by their defensive secretion of cantharidin, a potent blistering agent. The species is recorded across western and central Canada and the United States. Like other Meloe species, it likely exhibits the complex life history involving parasitic larvae that exploit solitary bees as hosts, though specific details for this species remain poorly documented.

  blister-beetleoil-beetlebee-parasitecantharidinground-nesting-beesMeloidaeNorth-Americatriungulinreflex-bleedingsolitary-bee-parasite

• Stictocephala stimulea

  Stictocephala stimulea is a treehopper species in the family Membracidae, first described by Van Duzee in 1914. Treehoppers in this genus are characterized by their enlarged, often ornate pronotum that extends over the body like a helmet or thorn. The specific epithet 'stimulea' suggests possible resemblance to or association with stinging organisms, though this refers to appearance rather than actual sting. Like other membracids, this species likely feeds on plant phloem sap using piercing-sucking mouthparts.

  MembracidaetreehopperAuchenorrhynchaCicadomorphaStictocephalaCeresiniSmiliinaeHemipterasap-feederpronotumhelmetthorn-mimicryplant-feeding-insectphloem-feedercryptic-colorationcamouflageVan-Duzee-1914NearcticNorth-AmericaFloridaNorth-CarolinaSouth-Dakotaunderstudied-speciesdata-deficienttaxonomyentomologyinsectarthropodtrue-bugauchenorrhynchancicadomorphleafhopper-relativeplant-hopperornate-pronotumpronotal-extensiondorsal-helmetinsect-morphologyinsect-systematicsbiodiversitynatural-historyconservationinsect-ecologyinsect-behaviorinsect-developmentinsect-distributionbiogeographyspecies-descriptiontype-specimenoriginal-descriptionhistorical-taxonomymuseum-specimencollection-recordoccurrence-dataGBIFiNaturalistcitizen-scienceobservation-recordrange-mapdistribution-datageographic-rangeendemicnative-speciesregional-faunalocal-biodiversityhabitat-specificityhost-plantfeeding-guildphytophagousherbivoreplant-insect-interactioncoevolutionmimicrydefensecrypsisevolutionary-adaptationmorphological-adaptationexoskeleton-modificationstructural-colorationinsect-identificationfield-guidediagnostic-charactertaxonomic-keyspecies-complexcryptic-speciesmorphological-variationgeographic-variationphenotypic-plasticitylife-historyreproductionovipositionnymphinstarmetamorphosishemimetabolousincomplete-metamorphosispopulation-dynamicscommunity-ecologyfood-webtrophic-levelprimary-consumerecosystem-functionnutrient-cyclingenergy-flowbiological-controlpest-statusagricultural-relevanceforestryurban-ecologyinvasive-potentialclimate-changehabitat-lossconservation-statusdata-gapresearch-needtaxonomic-revisionphylogeneticsmolecular-systematicsDNA-barcodingintegrative-taxonomynatural-history-collectionspecimen-digitizationopen-dataFAIR-databiodiversity-informaticsbiological-databasetaxonomic-authoritynomenclaturezoological-nomenclatureInternational-Code-of-Zoological-Nomenclaturetype-localitytype-specimen-depositionauthor-citationoriginal-combinationcurrent-combinationsynonymyhomonymynomenclatural-acttaxonomic-historyclassification-systemphylogenetic-classificationevolutionary-relationshipsmonophylycladelineagestem-groupcrown-groupdivergence-timemolecular-clockfossil-recordpaleontologyextantextinctliving-fossilevolutionary-historybiogeographic-historydispersalvicariancerange-expansionrange-contractionrefugiumglacial-relictpostglacial-colonizationanthropogenic-impacthuman-disturbanceland-use-changeurbanizationagricultural-intensificationpesticide-usehabitat-fragmentationconnectivitycorridorprotected-areaconservation-planningsystematic-conservation-planningpriority-areahotspotendemic-areacenter-of-origincenter-of-diversityevolutionary-significant-unitmanagement-unitpopulationmetapopulationsource-sink-dynamicsdemographyvital-ratessurvivalfecunditygeneration-timepopulation-growth-ratecarrying-capacitydensity-dependenceregulationlimiting-factorenvironmental-stochasticitydemographic-stochasticitygenetic-stochasticityinbreeding-depressionoutbreeding-depressiongenetic-diversityheterozygosityallelic-richnesseffective-population-sizegenetic-driftgene-flowmigrationisolation-by-distanceisolation-by-resistancelandscape-geneticspopulation-genomicsphylogeographyhistorical-demographycoalescent-theoryapproximate-Bayesian-computationspecies-distribution-modelecological-niche-modelhabitat-suitabilitybioclimatic-envelopeclimate-envelopeniche-conservatismniche-evolutionrealized-nichefundamental-nichebiotic-interactioncompetitionpredationparasitismmutualismcommensalismamensalismfacilitationapparent-competitionindirect-interactiontrophic-cascadetop-down-controlbottom-up-controlmesopredator-releasekeystone-speciesecosystem-engineerfoundation-speciesdominant-speciescharacteristic-speciesindicator-speciesumbrella-speciesflagship-speciessurrogate-speciesfocal-speciestarget-speciespest-speciesinvasive-speciesnaturalized-speciesalien-speciesnon-native-speciesintroduced-speciescryptogenic-speciesrange-shiftpoleward-shiftelevational-shiftphenological-shifttemporal-mismatchtrophic-mismatchphenologyseasonalityvoltinismunivoltinebivoltinemultivoltinepartial-generationprolonged-diapauseaestivationhibernationoverwinteringoverwintering-strategycold-hardinessfreeze-tolerancefreeze-avoidancesupercoolingcryoprotectantthermal-biologythermoregulationbehavioral-thermoregulationmicroclimate-selectionhabitat-selectionoviposition-site-selectionhost-selectionmate-choicesexual-selectionsperm-competitioncryptic-female-choicereproductive-isolationprezygotic-isolationpostzygotic-isolationspeciationallopatric-speciationsympatric-speciationparapatric-speciationperipatric-speciationreinforcementhybridizationintrogressionhybrid-zoneclineecotonecontact-zonesecondary-contactexogenous-selectionendogenous-selectionecological-speciationmutation-order-speciationsexual-conflictsexual-dimorphismsexual-size-dimorphismsexual-dichromatismpolymorphismcolor-polymorphismmorphphasegregarious-phasesolitary-phasedensity-dependent-polyphenismreaction-normgenotype-environment-interactionplasticitycanalizationgene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• Timarcha

  bloody-nosed beetles

  Timarcha is a genus of flightless leaf beetles comprising over 100 species across three subgenera, with a disjunct distribution spanning the Mediterranean region and western North America. The genus exhibits several unusual traits for Chrysomelidae, including complete apterism (winglessness), fused elytra, and archaic genital morphology. All species are uniformly black and herbivorous, with host plant associations concentrated in Rubiaceae and Plumbaginaceae. The most familiar species is T. tenebricosa, commonly known as the bloody-nosed beetle, named for its defensive reflex bleeding behavior.

  apterousflightlessreflex-bleedingMediterraneanHolarctic-disjunctionherbivoryChrysomelidaeleaf-beetleancient-lineageRubiaceaePlumbaginaceaedefense-behaviorthanatosismonovoltinediapauseparasite-mediated-selectionmate-choicecryptic-species-complexphylogeographymolecular-systematics

• Tritoma

  pleasing fungus beetles

  Tritoma is a genus of pleasing fungus beetles (family Erotylidae) comprising over 100 species distributed worldwide, with greatest diversity in the Old World. Members are associated with fungi, with some species feeding on euagaric mushrooms and mycorrhizae. The genus is currently considered paraphyletic based on molecular evidence and may require taxonomic revision into two separate genera. Tritoma bipustulata, a common European species with distinctive black-and-red spotted coloration, has been studied for its chemical defensive system.

  Erotylidaefungivorechemical-defenseparaphyletic-genusreflex-bleedingOld-World-distribution