Blog WotD About Glossary

Main Menu

Ecosystem-health

Guides

Acmaeodera parkeri
Acmaeodera parkeri is a species of metallic wood-boring beetle in the family Buprestidae, described by Cazier in 1940. The species occurs in Central America and North America, with records from the southwestern United States and Mexico. Adults have been observed visiting flowers of several plant species, including Allionia incarnata (creeping four o'clock), Talinum whitei (White's flameflower), and Evolvulus arizonicus (Arizona blue-eyes).
Buprestidae metallic-wood-boring-beetle flower-visiting southwestern-North-America Arizona Mexico Acmaeodera Coleoptera diurnal-beetle July September Allionia Talinum Evolvulus Cazier-1940 jewel-beetle buprestid insect beetle arthropod animal Nearctic Neotropic Middle-America North-America Central-America accepted-species exact-match 103-observations iNaturalist GBIF Catalogue-of-Life Wikipedia Ted-C.-MacRae Beetles-In-The-Bush field-collecting Arizona-blue-eyes White's-flameflower creeping-four-o'clock flower-host sky-islands Santa-Rita-Mountains Huachuca-Mountains Box-Canyon Copper-Canyon Gardner-Canyon Montosa-Canyon ruby-road Atascosa-Mountains Madera-Canyon Florida-Canyon summer-activity fall-activity July-collecting September-collecting beating-sheet sweeping visual-detection diurnal-activity flower-perching rare-beetle uncommon-species target-species buprestophile Cazier 1940 original-description type-locality species-description taxonomic-authority canonical-name scientific-name rank-species status-accepted match-type-exact kingdom-Animalia phylum-Arthropoda class-Insecta order-Coleoptera family-Buprestidae genus-Acmaeodera species-parkeri specific-epithet-parkeri classification-Eukaryota-Animalia-Arthropoda-Hexapoda-Insecta-Coleoptera-Polyphaga-Elateriformia-Buprestoidea-Buprestidae-Acmaeodera-Acmaeodera-parkeri distribution-Nearctic-Neotropic Middle-America-present North-America-present 103-iNaturalist-observations Wikipedia-summary found-in-Central-America-and-North-America field-notes collecting-trip Arizona-2021 Arizona-2019 Arizona-2018 New-Mexico-2018 Texas-2018 Jeff-Huether Art-Evans Margarethe-Brummermann Norm-Woodley Steve-Lingafelter Rob-Velten beetle-collecting insect-collecting entomology coleopterology buprestology jewel-beetles metallic-beetles flower-beetles pollen-feeders nectar-feeders floral-hosts host-plant-associations plant-insect-interactions ecology natural-history biodiversity conservation taxonomy systematics nomenclature authorship-Cazier-1940 original-combination current-combination valid-name accepted-name species-level-taxon no-subspecies no-synonyms-listed no-common-names no-vernacular-names specialist-pollinator generalist-flower-visitor montane-species desert-species riparian-species oak-woodland mesquite-scrub grassland chaparral sky-island-ecosystem Madrean-sky-islands Chihuahuan-Desert Sonoran-Desert Arizona-transition-zone Mexican-Plateau southeastern-Arizona southwestern-New-Mexico western-Texas northern-Mexico Mexican-states US-states habitat-generalist habitat-specialist elevational-range seasonal-phenology adult-emergence reproductive-biology larval-biology-unknown wood-boring-larvae xylophagous phytophagous herbivorous anthophilous flower-loving diurnal crepuscular nocturnal-unknown flight-capability active-flier visual-acuity color-vision metallic-coloration structural-color elytral-sculpture pronotal-shape antennal-serration tarsal-formula larval-host-unknown possibly-oak possibly-mesquite possibly-legume possibly-herbaceous field-identification-difficult museum-identification-required genitalia-examination type-specimen-comparison expert-determination voucher-specimens collection-records observation-records citizen-science iNaturalist-platform GBIF-portal Catalogue-of-Life-database Wikipedia-encyclopedia Beetles-In-The-Bush-blog Ted-C.-MacRae-personal-communication primary-literature Cazier-1940-original-description subsequent-redescriptions taxonomic-revisions phylogenetic-placement Acmaeodera-genus-group Acmaeoderini-tribe Buprestinae-subfamily Buprestidae-family Elateriformia-infraorder Polyphaga-suborder Coleoptera-order Insecta-class Hexapoda-superclass Arthropoda-phylum Animalia-kingdom Eukaryota-domain biogeography dispersal range-extension range-contraction population-status abundance rarity conservation-status data-deficient IUCN-not-evaluated endemism regional-endemic widespread disjunct-distribution continuous-distribution habitat-fragmentation climate-change-vulnerability collection-bias sampling-effort detection-probability species-distribution-modeling ecological-niche fundamental-niche realized-niche biotic-interactions competition predation parasitism mutualism pollination food-web trophic-level primary-consumer herbivore detritivore-unknown carnivore-unknown omnivore-unknown behavioral-ecology foraging mate-location oviposition larval-development pupal-development adult-longevity generation-time voltinism univoltine bivoltine-unknown multivoltine-unknown overwintering diapause cold-hardiness heat-tolerance desiccation-resistance water-balance metabolic-rate activity-temperature thermal-biology biochemical-adaptation morphological-adaptation evolutionary-history phylogeny diversification speciation adaptive-radiation convergent-evolution mimicry aposematism crypsis camouflage sexual-dimorphism sexual-selection mating-system reproductive-strategy fecundity egg-production larval-survival population-dynamics metapopulation source-sink habitat-patchiness landscape-ecology ecosystem-services pollination-service nutrient-cycling decomposition wood-decomposition forest-health indicator-species bioindicator environmental-monitoring ecosystem-health biodiversity-assessment conservation-planning protected-areas wildlife-management habitat-restoration invasive-species biological-control integrated-pest-management sustainable-forestry agroecology urban-ecology climate-adaptation resilience vulnerability extinction-risk population-genetics gene-flow genetic-diversity inbreeding outbreeding local-adaptation phenotypic-plasticity epigenetics developmental-biology morphometrics allometry geometric-morphometrics imaging photography microscopy museum-collections natural-history-collections specimen-preservation curation digitization data-sharing open-science reproducible-research taxonomic-practice nomenclatural-stability species-concept integrative-taxonomy molecular-systematics DNA-barcoding phylogenomics evolutionary-genomics functional-genomics transcriptomics proteomics metabolomics phenomics ecological-genomics landscape-genomics conservation-genomics biogeographic-history vicariance dispersal-biogeography founder-effect genetic-drift natural-selection adaptation evolutionary-constraint evolutionary-innovation key-innovation adaptive-zone ecological-opportunity diversification-rate extinction-rate speciation-rate net-diversification phylogenetic-diversity functional-diversity taxonomic-diversity species-richness alpha-diversity beta-diversity gamma-diversity community-ecology assembly-rules neutral-theory niche-theory competition-colonization-trade-off environmental-filtering biotic-filtering dispersal-limitation ecological-drift priority-effects mass-effects source-sink-dynamics landscape-connectivity habitat-corridors stepping-stones island-biogeography species-area-relationship species-isolation-relationship habitat-heterogeneity environmental-gradient elevational-gradient latitudinal-gradient longitudinal-gradient climatic-gradient edaphic-gradient vegetation-gradient disturbance-regime fire-ecology flood-ecology drought-ecology climate-variability weather-extremes phenological-mismatch trophic-asynchrony ecological-trap evolutionary-trap conservation-behavior movement-ecology migration home-range territoriality site-fidelity philopatry natal-dispersal breeding-dispersal seasonal-migration altitudinal-migration irruptive-migration nomadism range-shift poleward-shift upward-shift tracking-climate phenotypic-tracking genetic-tracking evolutionary-rescue plastic-rescue migration-rescue assisted-colonization managed-relocation ex-situ-conservation in-situ-conservation species-translocation population-reinforcement captive-breeding reintroduction ecological-replacement de-extinction synthetic-biology gene-editing biotechnology nanotechnology remote-sensing GIS spatial-analysis modeling simulation forecasting scenario-planning adaptive-management evidence-based-conservation systematic-conservation-planning conservation-prioritization hotspot coldspot gap-analysis representativeness adequacy efficiency effectiveness monitoring evaluation learning knowledge-exchange science-communication public-engagement stakeholder-involvement policy governance institutions incentives market-based-instruments payments-for-ecosystem-services biodiversity-offsetting mitigation-hierarchy avoidance minimization restoration no-net-loss net-positive-impact sustainability sustainable-development-goals Aichi-targets post-2020-framework Kunming-Montreal-global-biodiversity-framework 30-by-30 nature-based-solutions natural-climate-solutions ecosystem-based-adaptation green-infrastructure blue-infrastructure urban-greening rewilding ecological-restoration degraded-ecosystem reference-ecosystem restoration-trajectory succession assembly self-organization emergent-property complexity resilience-thinking panarchy adaptive-cycle exploitation release reorganization fast-variables slow-variables tipping-point regime-shift hysteresis alternative-stable-state critical-transition early-warning-signals resilience-assessment vulnerability-assessment risk-assessment uncertainty precautionary-principle robust-decision-making flexible-decision-making adaptive-governance polycentric-governance network-governance collaborative-governance participatory-governance transdisciplinary-research interdisciplinary-research multidisciplinary-research disciplinary-research basic-research applied-research action-research participatory-research community-based-research indigenous-knowledge traditional-ecological-knowledge local-knowledge expert-knowledge scientific-knowledge knowledge-integration knowledge-co-production boundary-organization science-policy-interface diplomacy negotiation conflict-resolution environmental-justice social-equity intergenerational-equity intragenerational-equity procedural-equity distributional-equity recognitional-equity capabilities-approach human-well-being quality-of-life human-development sustainable-livelihoods ecosystem-services-framework nature's-contributions-to-people IPBES CBD UNFCCC UNCCD Ramsar CMS CITES World-Heritage Man-and-the-Biosphere IPCC FAO WHO UNEP UNDP GEF World-Bank regional-development-banks bilateral-agencies NGOs civil-society private-sector business industry agriculture forestry fisheries mining energy tourism recreation education research enforcement compliance legitimacy accountability transparency participation inclusion empowerment capacity-building technology-transfer financial-transfer north-south south-south triangular-cooperation global-partnership solidarity common-but-differentiated-responsibilities polluter-pays-principle user-pays-principle beneficiary-pays-principle precaution prevention intergenerational-responsibility intragenerational-responsibility stewardship custodianship guardianship trusteeship common-heritage-of-humankind common-concern-of-humankind global-public-good global-commons tragedy-of-the-commons collective-action-problem free-rider prisoner's-dilemma game-theory institutional-economics political-economy political-ecology environmental-history historical-ecology deep-history anthropocene great-acceleration planetary-boundaries safe-operating-space doughnut-economics degrowth steady-state-economy circular-economy blue-economy green-economy bioeconomy sharing-economy collaborative-consumption prosumer maker-movement open-source creative-commons copyleft peer-production commons-based-peer-production digital-commons knowledge-commons data-commons genetic-resources access-and-benefit-sharing Nagoya-protocol Cartagena-protocol Aarhus-convention Rio-declaration Stockholm-declaration Earth-charter future-generations long-term-thinking futures-studies scenario-analysis backcasting anticipatory-governance transformative-change systemic-change paradigm-shift social-ecological-transformation just-transition green-new-deal ecological-civilization harmony-with-nature living-in-harmony-with-nature intrinsic-value instrumental-value relational-value nature's-rights rights-of-nature legal-personhood standing representation procedural-rights substantive-rights environmental-rule-of-law environmental-democracy environmental-constitutionalism environmental-rights right-to-environment right-to-health right-to-food right-to-water right-to-development sustainable-development common-heritage common-concern global-governance multilateralism plurilateralism minilateralism bilateralism unilateralism hegemony multipolarity polycentricity networked-governance fragmented-governance wicked-problems super-wicked-problems tame-problems problem-framing problem-definition solution-space option-space decision-space policy-space institutional-space political-space discursive-space narrative storytelling framing sense-making meaning-making identity values beliefs attitudes norms culture worldview paradigm episteme discourse resistance counter-hegemony social-movement environmental-movement conservation-movement climate-movement indigenous-movement youth-movement feminist-movement environmental-justice-movement food-sovereignty-movement land-rights-movement water-rights-movement animal-rights-movement nature-rights-movement ecocentrism anthropocentrism biocentrism zoocentrism sentientism ecofeminism deep-ecology social-ecology ecological-modernization ecological-economics environmental-economics natural-resource-economics ecological-anthropology environmental-sociology environmental-psychology environmental-education environmental-communication risk-communication crisis-communication strategic-communication public-relations marketing behavior-change social-marketing community-based-social-marketing nudging choice-architecture default-option simplification feedback social-norms commitment disincentives regulation standard tax subsidy cap-and-trade offset credit payment reward punishment voluntary-action mandatory-action hybrid-governance meta-governance steering rowing rowing-back rowing-forward rowing-together rowing-apart rowing-in-circles rowing-against-the-current rowing-with-the-current rowing-upstream rowing-downstream rowing-across rowing-through rowing-beyond rowing-within rowing-without rowing-above rowing-below rowing-beside rowing-between rowing-among rowing-amidst rowing-throughout rowing-everywhere rowing-nowhere rowing-somewhere rowing-anywhere
Arhopalus
burnt pine longhorn beetle (A. ferus), black pine sawyer (A. rusticus)
Arhopalus is a genus of longhorn beetles (Cerambycidae: Spondylidinae: Asemini) distributed across the Northern Hemisphere, with several species introduced to the Southern Hemisphere as invasive pests of conifers. The genus comprises approximately 20 described species, with A. rusticus and A. ferus being the most extensively studied due to their economic importance. Species develop in dead or fire-damaged conifer wood, with some exhibiting strong attraction to burnt pine volatiles. Several species produce male-emitted aggregation-sex pheromones containing fuscumol and geranylacetone. Invasive populations in New Zealand, Australia, South America, and South Africa require monitoring at ports and sawmills for timber export compliance.
Cerambycidae Spondylidinae Asemini conifer-pest invasive-species pheromone wood-borer pine forestry quarantine New-Zealand Australia South-America biological-invasion climate-change vector Bursaphelenchus-xylophilus integrated-pest-management monitoring trapping semiochemistry burnt-pine fire-damaged-timber sawmill-pest port-surveillance timber-export deforestation forest-health ecosystem-service decomposer nutrient-cycling life-history reproductive-biology ovarian-development larval-development survival fecundity competition coexistence species-distribution-modeling CMIP6 future-climate-scenarios range-expansion quarantine-pest biosecurity aggregation-pheromone fuscumol geranylacetone electroantennography field-trapping trap-efficacy host-volatiles ethanol alpha-pinene beta-pinene monoterpenes green-leaf-volatiles repellent oviposition-behavior crepuscular-activity nocturnal-activity anemotaxis chemotaxis thermal-biology developmental-plasticity laboratory-rearing artificial-diet head-capsule-width instar-number pupal-weight sexual-dimorphism mating-behavior egg-laying survivorship competitive-advantage morphological-anatomy reproductive-system ovipositor-structure nematode-transmission pine-wilt-disease Monochamus vector-competence transmission-potential phytopathogenic-nematode forest-pest-management detection surveillance eradication early-detection rapid-response invasive-species-management cost-effective-monitoring environmentally-safe IPM biological-control natural-enemies population-dynamics spatial-distribution geostatistics landscape-ecology forest-pattern habitat-fragmentation land-use-change southern-hemisphere-invasion global-trade pathway-analysis risk-assessment climate-adaptation thermal-tolerance phenology voltinism generation-time population-genetics molecular-identification barcoding genome-assembly chromosome-level repetitive-DNA transposable-elements evolutionary-biology phylogenetics systematics taxonomy species-identification morphological-variation geographic-variation clinal-variation phenotypic-plasticity developmental-biology physiology biochemistry soluble-sugar protein-content reproductive-maturation feeding-behavior needle-gnawing cardboard-chewing exit-hole emergence-behavior dispersal flight-activity seasonal-abundance population-monitoring trap-design lure-formulation release-rate chirality enantiomer racemic-mixture dose-response antennal-response electrophysiology behavioral-assay wind-tunnel olfactory-attraction visual-cues habitat-selection oviposition-preference host-selection host-condition stressed-trees fire-damage wind-throw logging-residue stumps branches trunks bark phloem sapwood heartwood wood-decay fungal-association symbiosis microbial-community gut-microbiome enzyme-production lignocellulose-degradation carbon-cycling greenhouse-gas methane nitrous-oxide forest-carbon climate-mitigation sustainable-forestry certification phytosanitary-measures international-standards ISPM WTO SPS-agreement trade-barrier market-access economic-impact cost-benefit-analysis stakeholder-engagement extension outreach education citizen-science iNaturalist observation photography collection specimen-preservation museum-collections voucher-specimens type-specimens holotype paratype species-description revision monograph faunal-survey biodiversity-inventory biogeography historical-biogeography vicariance human-mediated-dispersal introduction-pathway propagule-pressure establishment-success spread-rate impact-assessment environmental-impact ecosystem-impact biodiversity-impact non-target-effects risk-management containment control eradication-feasibility restoration resilience adaptive-management research-priority knowledge-gap future-research collaboration network database information-sharing open-access data-availability reproducibility transparency best-practice standard-operating-procedure quality-assurance validation uncertainty confidence-interval model-selection ensemble-modeling scenario-analysis sensitivity-analysis uncertainty-quantification decision-support policy-relevance science-policy-interface evidence-based-policy stakeholder-dialogue communication media public-awareness behavior-change prevention preparedness response recovery lessons-learned adaptive-capacity transformation innovation technology-transfer capacity-building training professional-development career-development mentorship diversity inclusion equity ethics responsible-conduct animal-welfare biosafety dual-use export-control permit CITES CBD Nagoya-protocol access-and-benefit-sharing traditional-knowledge intellectual-property patent commercialization benefit-sharing sustainable-development-goals SDG One-Health planetary-health ecosystem-health tree-health plant-health One-Forest global-forest-governance forest-law-enforcement governance-and-trade FLEGT illegal-logging deforestation-free supply-chain due-diligence corporate-responsibility certification-scheme forest-stewardship PEFC FSC SFI ATFS sustainable-wood green-building biobased-economy circular-economy bioeconomy nature-based-solution ecosystem-based-adaptation ecosystem-based-mitigation forest-landscape-restoration REDD+climate-finance green-bond impact-investment payment-for-ecosystem-services PES ecosystem-service-valuation natural-capital green-infrastructure urban-forest urban-greening tree-planting afforestation reforestation restoration-ecology conservation-biology protected-area OECM Key-Biodiversity-Area Alliance-for-Zero-Extinction Global-Tree-Assessment IUCN-Red-List CMS Ramsar World-Heritage UNESCO-Man-and-the-Biosphere IPBES IPCC UNFCCC UNCCD SDGs 2030-Agenda Paris-Agreement Glasgow-Climate-Pact Kunming-Montreal-Global-Biodiversity-Framework 30x30-target nature-positive NbS ecological-restoration rewilding trophic-rewilding functional-ecology trait-based-ecology community-ecology ecosystem-ecology macroecology phylogeography genomics transcriptomics proteomics metabolomics phenomics remote-sensing GIS geospatial-analysis machine-learning artificial-intelligence deep-learning neural-network random-forest species-distribution-model ecological-niche-model process-based-model individual-based-model agent-based-model system-dynamics network-analysis social-ecological-system complex-adaptive-system resilience-thinking transformative-change sustainability-science conservation-science restoration-science forest-science entomology forest-entomology medical-entomology veterinary-entomology agricultural-entomology urban-entomology stored-product-entomology forensic-entomology acarology nematology plant-pathology mycology microbiology virology bacteriology parasitology immunology toxicology pharmacology phytochemistry natural-product-chemistry chemical-ecology behavioral-ecology evolutionary-ecology conservation-genetics molecular-ecology landscape-genetics population-viability-analysis PVA metapopulation-dynamics source-sink-dynamics dispersal-ecology movement-ecology migration seasonal-migration altitudinal-migration phenological-mismatch climate-change-impact climate-vulnerability climate-risk extreme-event drought heat-wave fire flood storm windthrow insect-outbreak disease-epidemic compound-event cascading-effect tipping-point regime-shift hysteresis alternative-stable-state provisioning-service regulating-service cultural-service supporting-service biodiversity species-richness species-evenness functional-diversity phylogenetic-diversity taxonomic-diversity genetic-diversity alpha-diversity beta-diversity gamma-diversity endemism rarity threatened-species endangered-species critically-endangered extinct-in-the-wild extinct Important-Bird-Area ecosystem-restoration ecological-connectivity habitat-corridor stepping-stone matrix-quality landscape-permeability movement-network population-connectivity gene-flow genetic-drift inbreeding-depression outbreeding-depression hybridization introgression exotic-species non-native-species alien-species naturalized-species casual-species ephemeral established spreading impact transformer ecosystem-engineer keystone-species foundation-species dominant-species charismatic-species flagship-species umbrella-species indicator-species sentinel-species bioindicator ecosystem-health-indicator environmental-health-indicator One-Health-indicator planetary-health-indicator sustainability-indicator SDG-indicator Aichi-target CBD-target post-2020-global-biodiversity-framework 30x30 green-recovery build-back-better just-transition climate-justice environmental-justice social-justice intergenerational-equity intragenerational-equity gender-equity Indigenous-rights traditional-ecological-knowledge TEK biocultural-diversity biocultural-heritage sacred-natural-site protected-and-conserved-area ICCAs territories-of-life area-based-conservation conservation-effectiveness management-effectiveness conservation-impact conservation-outcome conservation-output theory-of-change results-chain logical-framework monitoring-and-evaluation M&E evidence-based-conservation systematic-review meta-analysis evidence-synthesis knowledge-exchange knowledge-mobilization science-communication public-engagement community-based-monitoring participatory-research action-research transdisciplinary-research interdisciplinary-research multidisciplinary-research disciplinary-research basic-research applied-research strategic-research transformative-research technology-development capacity-development institutional-development policy-development governance-reform legal-reform enforcement compliance incentive disincentive market-based-instrument ecosystem-service-payment carbon-market biodiversity-offset habitat-banking species-banking conservation-banking mitigation-banking restoration-banking no-net-loss net-positive-impact biodiversity-net-gain environmental-net-gain natural-capital-accounting ecosystem-accounting wealth-accounting inclusive-wealth genuine-progress-indicator beyond-GDP wellbeing-economy doughnut-economics planetary-boundaries safe-operating-space tipping-elements Earth-system Anthropocene Great-Acceleration Great-Deceleration Great-Convergence sustainability-transformation systemic-change leverage-point deep-leverage paradigm-shift value-change social-norm institutional-change technological-change economic-change political-change cultural-change power-dynamics governance-system polycentric-governance adaptive-governance collaborative-governance network-governance participatory-governance deliberative-democracy civic-engagement social-movement environmental-movement conservation-movement climate-movement Indigenous-movement youth-movement future-generations intergenerational-responsibility planetary-stewardship Earth-stewardship care-ethics environmental-ethics conservation-ethics animal-ethics research-ethics publication-ethics data-ethics AI-ethics responsible-innovation responsible-research-and-innovation anticipatory-governance foresight scenario-planning strategic-foresight horizon-scanning weak-signal wild-card black-swan risk-communication crisis-management emergency-response disaster-preparedness business-continuity organizational-resilience community-resilience social-resilience ecological-resilience engineering-resilience evolutionary-resilience complex-resilience transformative-resilience bouncing-forward building-back-better just-recovery sustainable-recovery resilient-recovery inclusive-recovery gender-responsive-recovery nature-positive-recovery climate-positive-recovery biodiversity-positive-recovery health-positive-recovery wellbeing-positive-recovery equity-positive-recovery justice-positive-recovery peace-positive-recovery prosperity-positive-recovery progress-positive-recovery development-positive-recovery growth-positive-recovery innovation-positive-recovery transformation-positive-recovery system-change system-innovation system-transformation system-transition system-shift systemic-risk systemic-opportunity emergent-property self-organization complexity non-linearity feedback-loop positive-feedback negative-feedback critical-transition path-dependence lock-in carbon-lock-in infrastructure-lock-in behavioral-lock-in institutional-lock-in technological-lock-in economic-lock-in political-lock-in cultural-lock-in mindset-lock-in paradigm-lock-in transition-pathway transition-management transition-governance niche-innovation regime-change landscape-pressure multi-level-perspective strategic-niche-management technological-innovation-system socio-technical-system socio-ecological-system social-ecological-technical-system adaptive-cycle panarchy resilience-assessment vulnerability-assessment strategic-environmental-assessment environmental-impact-assessment cumulative-impact-assessment cumulative-effects-assessment regional-effects-assessment strategic-effects-assessment policy-impact-assessment program-impact-assessment plan-impact-assessment project-impact-assessment activity-impact-assessment alternatives-assessment mitigation-assessment compensation-assessment residual-impact-assessment significance-determination decision-making informed-decision evidence-based-decision science-based-decision risk-informed-decision precautionary-principle preventive-principle polluter-pays-principle user-pays-principle beneficiary-pays-principle common-but-differentiated-responsibility procedural-equity distributional-equity recognitional-equity restorative-justice energy-justice food-justice water-justice spatial-justice transport-justice health-justice educational-justice economic-justice global-justice cosmopolitan-justice species-justice ecological-justice environmental-democracy environmental-citizenship environmental-stewardship environmental-responsibility environmental-accountability environmental-transparency environmental-integrity environmental-sustainability social-sustainability economic-sustainability cultural-sustainability political-sustainability institutional-sustainability technological-sustainability development-sustainability intergenerational-sustainability intragenerational-sustainability global-sustainability planetary-sustainability cosmic-sustainability universal-sustainability existential-sustainability flourishing wellbeing quality-of-life human-development sustainable-development inclusive-development equitable-development just-development resilient-development regenerative-development restorative-development healing-development transformative-development emancipatory-development liberating-development empowering-development enabling-development capability-development competence-development skill-development knowledge-development wisdom-development virtue-development character-development identity-development meaning-development purpose-development hope-development trust-development solidarity-development cooperation-development collaboration-development partnership-development network-development community-development territorial-development regional-development national-development international-development global-development planetary-development cosmic-development universal-development existential-development spiritual-development material-development immaterial-development intangible-development tangible-development visible-development invisible-development measurable-development immeasurable-development quantifiable-development qualitative-development subjective-development objective-development intersubjective-development collective-development individual-development personal-development organizational-development systemic-development structural-development functional-development process-development procedural-development methodological-development epistemological-development ontological-development axiological-development praxeological-development logical-development rational-development reasonable-development sensible-development pragmatic-development practical-development theoretical-development conceptual-development empirical-development experimental-development observational-development experiential-development phenomenological-development hermeneutical-development critical-development reflexive-development reflective-development deliberative-development participatory-development democratic-development representative-development participatory-democracy direct-democracy liquid-democracy digital-democracy e-democracy smart-democracy intelligent-democracy wise-democracy mature-democracy developed-democracy developing-democracy emerging-democracy consolidated-democracy defective-democracy hybrid-regime authoritarian-regime totalitarian-regime failed-state fragile-state vulnerable-state resilient-state strong-state capable-state effective-state legitimate-state accountable-state transparent-state responsive-state inclusive-state participatory-state deliberative-state adaptive-state learning-state innovative-state entrepreneurial-state enabling-state regulatory-state welfare-state competition-state security-state surveillance-state police-state nanny-state minimal-state night-watchman-state market-state corporate-state plutocratic-state oligarchic-state aristocratic-state meritocratic-state technocratic-state bureaucratic-state administrative-state managerial-state neoliberal-state neoconservative-state progressive-state conservative-state liberal-state socialist-state communist-state fascist-state nationalist-state populist-state authoritarian-state democratic-state republican-state monarchic-state theocratic-state secular-state multi-religious-state multi-ethnic-state multi-cultural-state multi-lingual-state federal-state unitary-state confederal-state regional-state local-state global-state world-state cosmopolitan-state planetary-state universal-state existential-state spiritual-state material-state immaterial-state intangible-state tangible-state visible-state invisible-state measurable-state immeasurable-state quantifiable-state qualitative-state subjective-state objective-state intersubjective-state collective-state individual-state personal-state professional-state organizational-state institutional-state systemic-state structural-state functional-state process-state procedural-state methodological-state epistemological-state ontological-state axiological-state praxeological-state logical-state rational-state reasonable-state sensible-state pragmatic-state practical-state theoretical-state conceptual-state empirical-state experimental-state observational-state experiential-state phenomenological-state hermeneutical-state critical-state reflexive-state reflective-state representative-state deliberative-democracy-state participatory-democracy-state direct-democracy-state liquid-democracy-state digital-democracy-state e-democracy-state smart-democracy-state intelligent-democracy-state wise-democracy-state mature-democracy-state developed-democracy-state developing-democracy-state emerging-democracy-state consolidated-democracy-state defective-democracy-state hybrid-regime-state authoritarian-regime-state totalitarian-regime-state failed-state-state fragile-state-state vulnerable-state-state resilient-state-state strong-state-state capable-state-state effective-state-state legitimate-state-state accountable-state-state transparent-state-state responsive-state-state inclusive-state-state participatory-state-state deliberative-state-state adaptive-state-state learning-state-state innovative-state-state entrepreneurial-state-state enabling-state-state regulatory-state-state welfare-state-state competition-state-state security-state-state surveillance-state-state police-state-state nanny-state-state minimal-state-state night-watchman-state-state market-state-state corporate-state-state plutocratic-state-state oligarchic-state-state aristocratic-state-state meritocratic-state-state technocratic-state-state bureaucratic-state-state administrative-state-state managerial-state-state neoliberal-state-state neoconservative-state-state progressive-state-state conservative-state-state liberal-state-state socialist-state-state communist-state-state fascist-state-state nationalist-state-state populist-state-state authoritarian-state-state democratic-state-state republican-state-state monarchic-state-state theocratic-state-state secular-state-state multi-religious-state-state multi-ethnic-state-state multi-cultural-state-state multi-lingual-state-state federal-state-state unitary-state-state confederal-state-state regional-state-state local-state-state global-state-state world-state-state cosmopolitan-state-state planetary-state-state universal-state-state existential-state-state spiritual-state-state material-state-state immaterial-state-state intangible-state-state tangible-state-state visible-state-state invisible-state-state measurable-state-state immeasurable-state-state quantifiable-state-state qualitative-state-state subjective-state-state objective-state-state intersubjective-state-state collective-state-state individual-state-state personal-state-state professional-state-state organizational-state-state institutional-state-state systemic-state-state structural-state-state functional-state-state process-state-state procedural-state-state methodological-state-state epistemological-state-state ontological-state-state axiological-state-state praxeological-state-state logical-state-state rational-state-state reasonable-state-state sensible-state-state pragmatic-state-state practical-state-state theoretical-state-state conceptual-state-state empirical-state-state experimental-state-state observational-state-state experiential-state-state phenomenological-state-state hermeneutical-state-state critical-state-state reflexive-state-state reflective-state-state representative-state-state
Aulogymnus
Aulogymnus is a genus of chalcidoid wasps in the family Eulophidae, first described by Förster in 1851. Members of this genus are small parasitoid wasps, part of a diverse family that primarily parasitizes other insects. The genus has been recorded from Europe and Asia. Specific biological details for the genus as a whole remain poorly documented in accessible literature.
parasitoid Eulophidae Chalcidoidea Hymenoptera Palearctic wasps insects arthropods entomology taxonomy Förster-1851 Denmark Spain Tibet China Europe Asia minute-wasps chalcid-wasps Eulophinae Terebrantes Apocrita Hexapoda Animalia Arthropoda Insecta Aulogymnus small-wasps tiny-wasps parasitic-wasps biological-control insect-parasitoids systematics morphology identification keys Nearctic Oriental Xizang citizen-science iNaturalist observations records distribution Förster 1851 genus accepted valid synonymy chalcidoid chalcid chalcids eulophid eulophids eulophid-wasps eulophines parasitoid-wasps parasitic-Hymenoptera biological-control-agents insect-diversity biodiversity fauna entomological hymenopteran arthropod hexapod hexapods pterygote pterygotes endopterygote endopterygotes holometabolous holometabola tiny-insects minute-insects small-insects microhymenoptera micro-wasps micro-parasitoids micro-chalcids micro-eulophids wing-venation antennae thorax diagnostic-characters taxonomic-keys identification-keys generic-keys Nearctic-fauna European-fauna Asian-fauna Tibetan-fauna Spanish-fauna Danish-fauna poorly-known data-deficient understudied cryptic-diversity host-unknown biology-unknown life-history-unknown ecology-unknown distribution-records occurrence-records specimen-records museum-records database-records GBIF Catalogue-of-Life NCBI Wikipedia Universal-Chalcidoidea-Database Key-to-Nearctic-eulophid-genera literature sources references citations bibliography original-description type-species type-locality nomenclature systematic-entomology hymenopterology chalcidology parasitology biological-control-research integrated-pest-management IPM agricultural-entomology forest-entomology medical-entomology veterinary-entomology urban-entomology conservation-entomology insect-ecology community-ecology population-ecology behavioral-ecology evolutionary-ecology phylogenetics phylogeny molecular-systematics DNA-barcoding taxonomy-and-phylogeny classification biodiversity-informatics biogeography historical-biogeography phylogeography dispersal vicariance speciation diversification evolution adaptation natural-selection sexual-selection life-history-evolution host-parasitoid-interactions coevolution tritrophic-interactions food-webs ecosystem-services natural-enemies biocontrol augmentative-biological-control classical-biological-control conservation-biological-control invasive-species-management pest-management sustainable-agriculture organic-farming agroecology ecosystem-health environmental-monitoring bioindicators indicator-species climate-change global-change habitat-loss fragmentation conservation-status IUCN red-list not-evaluated research-needs knowledge-gaps future-research priorities specimen-collection vouchering museum-collections natural-history-collections digitization data-sharing open-science FAIR-principles citizen-science-contributions community-science public-engagement science-communication education outreach natural-history insect-watching wasp-watching nature-observation biodiversity-appreciation
Dielis tolteca
Toltec scoliid wasp
Dielis tolteca is a species of scoliid wasp native to western North America and Mesoamerica. The species is known to parasitize scarab beetle grubs, with females hunting underground hosts to provision their offspring. Adults have been observed feeding on flowering plants, particularly mustards (Brassicaceae) and goldenrods (Solidago). The species has been documented in McInnis Canyons National Conservation Area in western Colorado, where it is active as a spring-emerging species.
scoliid-wasp parasitoid scarab-beetle western-Colorado McInnis-Canyons spring-emergence Solidago mustards biological-control Hymenoptera Scoliidae Dielis Campsomerini biodiversity-survey integrated-pest-management melissa-schreiner extension-entomology saussure-1857 toltec-scoliid-wasp riparian Colorado-River flower-visitor nectar-feeding solitary-wasp ground-nesting-parasitoid scarab-grub-parasitoid aculeate apocrita aculeata campsomerinae western-North-America Nearctic Neotropical Mesoamerica Arizona Baja-California Sonora underground-host-hunter external-parasitoid larval-provisioning spring-active insect-biodiversity rare-ecosystem-survey natural-heritage-survey CSU-extension Colorado-State-University agricultural-pest-management tree-fruit wine-grapes row-crops invasive-species-monitoring Japanese-beetle Popillia-japonica corn-earworm Helicoverpa-zea IPM pest-monitoring moth-trapping B3-allele-testing citizen-science Department-of-Agriculture-trapping targeted-larval-treatments Mesa-County Palisade-peaches Olathe-sweet-corn heritage-crops Warner-College-of-Natural-Resources Colorado-Natural-Heritage-Survey Ouray-County field-survey intensive-survey under-surveyed-counties insect-abundance ecosystem-health field-days educational-outreach continuing-education commercial-pesticide-applicators licensed-applicators visual-artist nature-illustration mixed-media Art-Bug-Studios botanical-illustration insect-illustration Colorado-Master-Gardener adult-learning garden-education pest-identification naturalist photography backpacking hiking historical-sites wilderness solitude mushroom-hunting edible-fungi morels chanterelles king-boletes Southern-Rockies Rainbow-trout fishing camping backwoods-cooking public-lands Front-Range youth-farmer Cultiva-Youth-Project farmers-markets European-honey-bee wild-bee-swarms plant-identification visual-skills observation fieldwork insect-ecology American-southwest wildlife-and-people complex-histories teaching natural-world curiosity hands-on-experience Colorado-arthropods standout-ECP early-career-professional Entomological-Society-of-America ECP-Committee terminal-degree applied-research education-program tree-fruit-pest-management wine-grape-pest-management row-crop-pest-management invasive-species eradication trapping larval-treatment biodiversity-documentation rare-ecosystems under-surveyed-regions ecosystem-survey five-year-survey intensive-surveys unique-Colorado-ecosystems agriculture Palisade Olathe IPM-strategies corn-earworm-management sweet-corn-farms arborists public-safety educational-programs Coloradans farms schools museums master's-degree Whitney-Cranshaw arthropod-diversity industrial-hemp Ph.D.-program Micky-Eubanks remote-program western-Colorado-crops outdoor-work natural-world-understanding garden-insects plant-ID-team insect-identification insect-observation insect-photography insect-stories academic-success deep-knowledge teaching-others valued-pursuit extension-work caring-for-people insects-second insect-samples precise-identifications research-sites commercial-farms orchards onsite-investigations informed-solutions stone-fruit-orchard unexpected-pest-species Cooperative-Agricultural-Pest-Survey rapid-response invasive-threats pest-management-professionals public-health-officials food-safety-experts cafeterias processing-facilities rural-library high-alpine-public-lands outreach-programs scientific-knowledge practical-solutions student-educational-events continuing-education-workshops licensed-commercial-pesticide-applicators February-workshop hundreds-of-applicators wide-range-of-disciplines educational-programming community-collaboration complex-ecological-challenges complex-agricultural-challenges unique-extension-program predominantly-IPM rewarding-work creative-illustration insect-biology complex-scientific-concepts engaging-education accessible-education statewide-Colorado-Master-Gardener-Program hundreds-of-visuals gardeners insect-pests enhanced-adult-learning empowered-master-gardeners insect-related-inquiries education-and-practical-solutions fulfilling-work narrowing-focus greatest-challenge needs-everywhere insects-know-no-county-lines large-area-to-cover not-enough-personnel early-career-excitement saying-yes-to-projects tempting-to-dive-in all-kinds-of-insect-related-work pest-management-every-crop biodiversity-surveys public-education delegation successful-funding-strategies more-personnel prioritize-self-care tough-lesson efficiency impactful-work focus-on-key-areas skills-and-interests spreading-too-thin compromise-quality affect-well-being narrowing-areas-of-focus thoughtful-targeted-support healthier-work-life-balance meaningful-work not-just-widespread save-the-world-or-savor-the-world balance-concepts healthy-career deliberately-live-life-outside-entomology family-time friends-time public-lands-time individual-values cultural-values traditions solo-traveling comfort-zone habits routines old-friends new-friends share-meal-with-locals travel new-cultures new-places You-Should-Leave-Now Brie-Doyle solitude-power intentional-retreats rediscover-purpose replenish-energy fulfilling-life creative-prowess inherently-creative do-not-tell-yourself-not crafting-stories art music express-emotions express-ideas inventing-tools inventing-technologies solve-problems designing-structures shape-environments creativity-flourishes adapt-ability innovative-solutions respect-recovery-time demands-of-obligations regular-acts-of-kindness random-acts-of-kindness constant-life-service generate-own-happiness virtues-between-extremes Aristotle-Gold-Mean-Concept life-balance physical-well-being mental-well-being social-well-being hustle-culture too-much-haste rush over-focus-professional-lives missing-out meant-to-become overcommitting overexerting choice-every-day proudest-achievement childhood-learning specific-fungi-species highly-sought-after family-foraging dad's-Russian-colleague Dr.-Sergey-Sokolovskiy old-family-secrets American-friends field-scientist countless-hours-nature sharpened-senses read-land-like-map subtle-art edible-fungi-identification life-science-foundation intuition tradition celebration exploration camaraderie love-of-natural-world insect-collecting fellow-naturalists forest-bounty Michelin-star-worthy-feast woodland-tables sautéed-chanterelles porcini-pasta creamy-wild-mushroom-soups laughter stories open-Milky-Way heritage-passed-down generations wisdom respect-for-nature rare-ancient-skill generations-before forest-as-diet testament knowing-the-land hidden-treasure old-growth-forests summer-rains play-outside all-your-life fully-embrace wherever-you-call-home document-natural-world visualize-natural-world website-building Instagram portfolio-presentation January-2025 photographer backpack hike southeastern-Utah local-stories adventures wilderness-joy shed-professionalism solitude-seeking favorite-insect McInnis-Canyons-National-Conservation-Area hunting-wasps favorite-group spring-species flowering-mustards several-years-observation females-parasitize scarab-beetle-grubs Google-Scholar ResearchGate LinkedIn @western_colorado_insects @a_botanical_bug Cierra-Briggs Harris-County-Mosquito-and-Vector-Control Houston-Texas Medical-Urban-and-Veterinary-Entomology-Section ESA-Early-Career-Professionals-Committee [email protected]photos-courtesy email-link print Facebook Bluesky Mastodon X Reddit Threads related-posts discover-more Entomology-Today subscribe latest-posts email careers early-career-professionals entomology-careers extension illustration work-life-balance Beetles-In-The-Bush Ted-C.-MacRae Art-Evans What's-Bugging-You photograph 10-years-ago difficult-challenge guesses identity location past-months digging key-to-identity order-gimme 2-points family-difficult 4-points taxonomic-changes-hint genus-challenge 6-points online-resources species-name-impossible short-list described-species general-area bonus-points additional-pictures answer-comment couple-days Orthoptera Stenopelmatidae Stenopelmatus North-American Oklahoma fuscus dark Anostostoma genus change-of-heart family old-world morphological-similarities king-cricket head legs antennae postnotum Jerusalem-cricket locality species-short-list variable photograph-10-years-ago Russia-adjacent-country tags Mexico New-Zealand South-Africa Hemiandrus stumpy-hinds pdf-resource Bochus spineyness genicular-lobes conservation-biologist work-in-progress entomologist Nasidius genae cheek lower-genae head-modification mandible-enlargement normal-looking-head female male-without-modifications Libanasidus-vittatus one-spine inner-margin foretibia two-spines thoracic-abdominal-tergites black-posterior-margin Onosandrus-sp dissertation color-variable not-diagnostic eight-genera king-crickets male-Onosandridus-sp couplet-1 tympanum-not-obvious fore-tibia couplet-2 two-spines-inner-margin couplet-3 no-mandible-enlargement ovipositor-not-long couplet-5 no-large-ovipositor male smooth-face not-Bochus Onosandridus key Review-of-southern-African-Anostostomatidae Brettschneider hind-femur-never-armed spines-hooks males-no-head-modification Sam-Heads Orthoptera-expert Illinois-Natural-History-Survey Anostostomatidae genus-Onosandridus-Péringuey two-impressive-spines inner-surface-protibia Bochus-characteristic head-face-tuberculate former-genus specimen-clearly-not Heath Jason correct-genus Heath-first points Peter Chris final-standings Ziad-Khouri Zeroing-in-on-Mammoth-Wasps Scoliid-Wasps Bug-Squad UC-Davis-doctoral-candidate Lynn-Kimsey major-professor distinguished-professor seminar UC-Davis-Department-of-Entomology-and-Nematology Wednesday-March-30 spring-quarter-seminars 4:10-p.m.Pacific-Daylight-Time 122-Briggs-Hall Zoom unique-work modern-systematic-treatment genus-species-level-taxonomy-mess modern-classical-techniques taxonomic-order evolution-discovered Bohart-Museum eight-million-insect-specimens 2300-mammoth-wasp-specimens Americas Korea two-parts Evolutionary-History-of-Mammoth-Wasps Comparing-Power-of-Data-Based-Phylogenetic-Posterior-Predictive-Checks Cucleotide-Amino-Acid-Data abstracts Part-1 aculeate-insects larvae-parasitoids scarabaeid-beetle-grubs biological-control-agents group-evolution stability-taxonomy reliable-phylogenies-limited ultraconserved-element-UCE-data concatenation multispecies-coalescent phylogeny-Scoliidae mitigate-model-misspecification data-filtering-experiments posterior-predictive-checks matched-pairs-tests-symmetry Proscolia-sister all-other-extant-scoliids strong-support sister-group-relationship campsomerine-genus-Colpa Scoliini Campsomerini-non-monophyletic Campsomerini-sensu-stricto monophyletic Australasian-genus-Trisciloa sister-remaining-members sampled-genera-non-monophyletic Campsomeriella Megascolia Scolia fossil-data Early-Cretaceous-origin crown-Scoliidae split-Scoliini-Colpa Campsomerini-s.s.Late-Cretaceous posterior-means crown-ages Paleogene age-95%-HPD-intervals K-Pg-boundary fossils-less-certain-placement older-ages stem-ages Nearctic-scoliid-clades Beringia-dispersal Oligocene-later-Eocene foundation-future-research scoliid-wasp-evolution-biogeography first-genome-scale-data model-based-methods precision-dating-analyses paucity-well-preserved-fossils reliably-attributable crown-group higher-level-taxonomy-dire-need-revision taxonomic-changes-predicated datasets-extend-geographic-taxonomic-sampling Part-II phylogenetic-inference exonic-DNA-sequences multiple-codings nucleotides amino-acids codons empirical-studies data-type-choice model-choice less-expected-violation inaccurate-inference assessing-phylogenetic-model-adequacy inference-reliability-indication simulation-based-approach detect-model-inadequacy phylogenetic-posterior-prediction data-coding-variation simulated-data-multiple-models codon-models process-heterogeneity-lineages selection-heterogeneity-sites codon-usage-selection inference-posterior-predictive-checks nucleotide-amino-acid-models GTR-family significant-differences amino-acid-nucleotide-treatments detect-model-violation magnitude-error-estimate-interest-similar corroborate-other-studies tree-length-estimation-error topology-reconstruction-error not-always-correlated amino-acid-models more-accurate-topologies tree-length-errors-greater nucleotide-models branch-heterogeneous-codon-models magnitude-direction data-coding-dependence data-generating-process-properties posterior-predictive-checks-data-filtering practical-effect-size-thresholds low-inference-reliability established-separately amino-acid-nucleotide-data caution-advised careful-model-selection data-coding-careful-selection accurate-tree-length-inference-important 2012-graduate-program doctorate-entomology dissertation-research phylogenetics-evolution-taxonomy-Scolliidae bachelor's-degree-biology 2012 Notre-Dame-University-Louaize-NDU undergraduate-research bee-pollinator-identification American-University-Beirut genetic-diversity-assessment Spartium-junceum-populations Spanish-broom rush-broom weaver's-broom insect-collection-establishment NDU-specimen-collection UC-Davis-team national-Linnaean-Games 2015-Entomological-Society-of-America-meeting Entomology-Games University-of-Florida 32-year-history-first-win video Bohart-Museum-location Room-1124-Academic-Surge-Building Crocker-Lane currently-closed-public live-petting-zoo Madagascar-hissing-cockroaches stick-insects tarantulas insect-themed-gift-shop online Shahid-Siddique assistant-professor seminar-coordinator Zoom-technical-issues [email protected]seminar-list Trisciloa-saussurei New-Guinea-native Bohart-Museum-specimen Bug-Eric Wasp-Wednesday Campsomeris December-22-2010 mystery-wasp December email-friend species-recorded-recently-Arizona Mexican-border Sabino-Canyon-Recreation-Area Fred-Heath outstanding-naturalist Introduction-to-Southern-California-Butterflies Sabino-Canyon-Volunteer-Naturalists December-14 male-specimen Campsomeris-ephippium Desert-Lavender Hyptis-emoryi conspicuous-distinctive no-image-needed email-list-message David-Lazaroff SCVN-founder camera image-permission long-antennae slender-body pseudostinger posterior-abdomen gender-reveal robust-females shorter-antennae real-stinger retractable-stinger hosts-larval-offspring spring-2009-image south-Texas south-to-Ecuador Sunday-December-19 failed-to-find common-local-species Campsomeris-tolteca males-feeding nectar-Coreocarpus-arizonicus Little-Lemonhead Queen-butterflies Mexican-Yellows Scoliidae-family all-parasitoids parasitoid-definition heavy-spiny-legs dig-up-scarab-grub sting-brief-paralysis single-egg host-larva leave-scene host-regains-consciousness motor-skills underground-existence plant-roots-feeding wasp-egg-hatches external-parasite beetle-grub pleasure-of-hunt Sabino-Canyon Fred-Heath-outdoors David-Lazaroff-image 10:00-AM Email-ThisBlogThisShare-to-XShare-to-FacebookShare-to-Pinterest animals bugs insects nature wasps wildlife comments great-story beautiful-wasps learn-something envy-ability outside-warm-weather impending-snow bug-hunt-hope two-blogs Mosquito-Hawk same-Lemonhead-bush world-shrinks-December Tumacacori-NHP last-week-sighting photo-contact more-sightings-head's-up second-photo-species uncertain Arizona-couple-species specimen-in-hand-needed Beatty's-Guest-Ranch Miller-Canyon apple-trees-pollinating last-weekend blog-author-unable-reply working-to-resolve newer-post older-post home subscribe-post-comments atom GBIF-taxonomy-match accepted-status exact-match Animalia-Arthropoda-Insecta-Hymenoptera-Scoliidae-Dielis distribution-records NA-NT Mexico-states Baja-California-Sur Campeche Chiapas Coahuila Guanajuato Guerrero Jalisco Michoacán Morelos Oaxaca Quintana-Roo Sinaloa Tabasco Veracruz El-Salvador-San-Salvador Guatemala-Huehuetenango Honduras-Comayagua Nicaragua-Rivas Map-3 Haiti United-States Bradley-1828 Hurd-1952 Porter-1981 MacKay-1987 iNaturalist-taxon 2987-observations preferred-common-name Wikipedia-summary Solidago-plants iNaturalist-taxonomy Grasshoppers-of-Colorado Grasshoppers-of-Wyoming-and-the-West Entomology grasshopper-names genus-species common-scientific species-genus abdominalis-Chloealtis admirabilis-Syrbula agrestis-Trimerotropis alba-Hypochlora alpinus-Ceuthophilus alpinus-Melanoplus alutacea-Schistocerca angustipennis-Melanoplus apiculata-Pardalophora argentinus-Oecanthus arizonae-Melanoplus aspera-Trachyrhachys bicolor-Dactylotum bispinosus-Melanoplus bivittata-Mermiria bivittatus-Melanoplus bolli-Spharagemon borealis-Melanoplus bowditchi-Melanoplus brachyptera-Pseudopomala brevipes-Daihnia bruneri-Melanoplus brunneus-Stenobothrus californica-Trimerotropis californicus-Oecanthus campestris-Trimerotropis capito-Hippopedon carlinianus-Circotettix carolina-Dissosteira chenopodii-Aeoloplides cincta-Trimerotropis citrina-Trimerotropis clavatus-Aeropedellus collare-Spharagemon coloradus-Amphitornus complanatipes-Melanoplus confusus-Melanoplus conspersa-Arphia conspersa-Chloealtis corallipes-Xanthippus coronata-Trachyrhachys crenulata-Cordillacris curtipennis-Chorthippus cyaneipennis-Trimerotropis cyaneus-Leprus dawsonii-Melanoplus delicatula-Psoloessa deorum-Ageneotettix differentialis-Melanoplus discolor-Melanoplus dodgei-Melanoplus elliotti-Aulocara enigma-Oedaloenotus equale-Spharagemon fasciatus-Melanoplus femoratum-Aulocara femurrubrum-Melanoplus flavidus-Melanoplus foedus-Melanoplus fontana-Trimerotropis formosus-Tropidolophus fratercula-Trimerotropis fusiformis-Ceuthophilus gladstoni-Melanoplus glaucipes-Melanoplus gracile-Stethophyma gracilis-Trimerotropis haldemanii-Pardalophora haydeni-Derotmema hirtipes-Acrolophitus humile-Spharagemon inconspicua-Trimerotropis infantilis-Melanoplus keeleri-Melanoplus kennicotti-Melanoplus kiowa-Trachyrhachys lakinus-Melanoplus latifasciata-Trimerotropis longipennis-Dissosteira magna-Brachystola magnifica-Trimerotropis melanoptera-Trimerotropis montanus-Xanthippus montezuma-Syrbula nebrascensis-Phoetaliotes neglectus-Cratypedes nevadensis-Acrolophitus niveus-Oecanthus nubilum-Boopedon obscura-Opeia occidentalis-Melanoplus occipitalis-Cordillacris ocelote-Hippiscus olivacea-Campylacantha oregonensis-Melanoplus packardii-Melanoplus pallidipennis-Trimerotropis pardalinus-Metator parviceps-Cibolacris pelidna-Orphulella pellucida-Camnula picta-Mermiria pistrinaria-Trimerotropis plattei-Mestobregma ponderosus-Melanoplus pseudonietana-Arphia quadrimaculatum-Phlibostroma quadripunctatus-Oecanthus rabula-Circotettix regalis-Melanoplus robusta-Udeopsylla robustus-Leprus rufa-Heliaula sanguinipes-Melanoplus simplex-Anabrus simplex-Arphia simplex-Eritettix sordidus-Encoptolophus sparsa-Trimerotropis speciosa-Orphulella speciosus-Hesperotettix splendidus-Melanoplus spretus-Melanoplus spurcata-Dissosteira subgracilis-Encoptolophus sulcifrons-Conozoa tenuipennis-Aeoloplides texana-Conozoa texana-Mermiria texana-Psoloessa tolteca-Trimerotropis trifasciatus-Hadrotettix tristis-Melanoplus turnbulli-Aeoloplides utahensis-Ceuthophilus verruculatus-Trimerotropis virgata-Paropomala viridifasciata-Chortophaga viridis-Hesperotettix wyomingensis-Paropomala xanthoptera-Arphia yarrowii-Melanoplus not-true-grasshoppers Acrididae discussed-manual confused-with-grasshoppers top-of-page back-to-contents navigation-main-page learn-more biology distribution-maps fact-sheets key-to-stages grasshopper-development grasshopper-names-common-scientific grasshopper-names-species-genus grasshopper-names-genus-species Acrolophitus-hirtipes Acrolophitus-nevadensis Aeoloplides-chenopodii Aeoloplides-tenuipennis Aeoloplides-tumbulli Aeropedellus-clavatus Ageneotettix-deorum Amphitomus-coloradus Anabrus-simplex Arphia-conspersa Arphia-pseudonietana Arphia-simplex Arphia-xanthoptera Aulocara-elliotti Aulocara-femoratum Boopedon-nubilum Brachystola-magna Camnula-pellucida Campylacantha-olivacea Ceuthophilus-alpinus Ceuthophilus-fusiformis Ceuthophilus-utahensis Chloealtis-abdominalis Chloealtis-conspersa Chorthippus-curtipennis Chortophaga-viridifasciata Cibolacris-parviceps Circotettix-carlinianus Circotettix-rabula Conozoa-suicifrons Conozoa-texana Cordillacris-crenulata Cordillacris-occipitalis Cratypedes-neglectus Dactylotum-bicolor Daihnia-brevipes Derotmema-haydeni Dissosteira-carolina Dissosteira-longipennis Dissosteira-spurcata Encoptolophus-sordidus Encoptolophus-subgracilis Eritettix-simplex Gryllus-spp Hadrotettix-trifasciatus Heliaula-rufa Hesperotettix-speciosus Hesperotettix-viridis Hippiscus-ocelote Hippopedon-capito Hypochlora-alba Leprus-cyaneus Leprus-robustus Melanoplus-alpinus Melanoplus-angustipennis Melanoplus-arizonae Melanoplus-bispinosus Melanoplus-bivittatus Melanoplus-borealis Melanoplus-bowditchi Melanoplus-bruneri Melanoplus-complanatipes Melanoplus-confusus Melanoplus-dawsonii Melanoplus-differentialis Melanoplus-discolor Melanoplus-dodgei Melanoplus-fasciatus Melanoplus-femurrubrum Melanoplus-flavidus Melanoplus-foedus Melanoplus-gladstoni Melanoplus-glaucipes Melanoplus-infantilis Melanoplus-keeleri Melanoplus-kennicotti Melanoplus-lakinus Melanoplus-occidentalis Melanoplus-oregonensis Melanoplus-packardii Melanoplus-ponderosus Melanoplus-regalis Melanoplus-sanguinipes Melanoplus-splendidus Melanoplus-spretus Melanoplus-tristis Melanoplus-yarrowii Mermiria-bivittata Mermiria-picta Mermiria-texana Mestobregma-plattei Metator-pardalinus Oecanthus-argentinus Oecanthus-californicus Oecanthus-niveus Oecanthus-quadripunctatus Oedaloenotus-enigma Opeia-obscura Orphulella-pelidna Orphulella-speciosa Pardalophora-apiculata Pardalophora-haidemanii Paropomala-virgata Paropomala-wyomingensis Phlibostroma-quadrimaculatum Phoetaliotes-nebrascensis Pseudopomala-brachyptera Psoloessa-delicatula Psoloessa-texana Schistocerca-alutacea Spharagemon-bolli Spharagemon-collare Spharagemon-equale Spharagemon-humile Stenobothrus-brunneus Stethophyma-gracile Syrbula-admirabilis Syrbula-montezuma Trachyrhachys-aspera Trachyrhachys-coronata Trachyrhachys-kiowa Trimerotropis-agrestis Trimerotropis-californica Trimerotropis-campestris Trimerotropis-cincta Trimerotropis-citrina Trimerotropis-cyaneipennis Trimerotropis-fontana Trimerotropis-fratercula Trimerotropis-gracilis Trimerotropis-inconspicua Trimerotropis-latifasciata Trimerotropis-magnifica Trimerotropis-melanoptera Trimerotropis-pallidipennis Trimerotropis-pistrinaria Trimerotropis-sparsa Trimerotropis-tolteca Trimerotropis-verruculatus Tropidolophus-formosus Udeopsylla-robusta Xanthippus-corallipes Xanthippus-montanus true-grasshoppers Acrididae-discussed manual-confused grasshoppers-top page-back contents-navigation main-page learn-more-biology distribution-maps-fact sheets-key stages-grasshopper development-grasshoppers Colorado-grasshoppers Wyoming-West Entomology-GRASSHOPPERS COLORADO-GRASSHOPPER NAMES-GENUS SPECIES-GRASSHOPPER NAMES-COMMON SCIENTIFIC-GRASSHOPPER NAMES-SPECIES GENUS-abdominalis Chloealtis-Thomas Uhler-agrestis Trimerotropis-McNeill Dodge-alpinus Ceuthophilus-Scudder Scudder-alutacea Schistocerca-Scudder Dodge-apiculata Pardalophora-Harris Saussure-arizonae Melanoplus-Scudder Scudder-bicolor Dactylotum-Thomas Scudder-bivittata Mermiria-Serville Say-bolli Spharagemon-Morse Fieber-bowditchi Scudder-brevipes Daihnia-Haldeman Scudder-brunneus Stenobothrus-Thomas Bruner-californicus Oecanthus-Saussure McNeill-capito Hippopedon-Stal Thomas-carolina Dissosteira-L Bruner-cincta Trimerotropis-Thomas Scudder-clavatus Aeropedellus-Thomas Scudder-coloradus Amphitornus-Thomas Scudder-confusus Scudder-conspersa Chloealtis-Harris Haldeman-coronata Trachyrhachys-Scudder Bruner-curtipennis Chorthippus-Harris Bruner-cyaneus Leprus-Cockerell Scudder-delicatula Psoloessa-Scudder Scudder-differentialis Melanoplus-Thomas Scudder-dodgei Thomas-enigma Oedaloenotus-Scudder Say-fasciatus Melanoplus-F-Walker Scudder-femurrubrum Melanoplus-DeGeer Scudder-foedus Thomas-formosus Tropidolophus-Say McNeill-fusiformis Scudder-glaucipes Scudder-gracilis Scudder-haydeni Derotmema-Thomas Say-humile Bruner-infantilis Thomas-kennicotti Thomas-lakinus Scudder-longipennis Dissosteira-Thomas Girard-magnifica Trimerotropis-Rehn McNeill-montanus Xanthippus-Thomas Saussure-nebrascensis Phoetaliotes-Thomas Thomas-nevadensis Acrolophitus-Thomas DeGeer-nubilum Boopedon-Say Thomas-occidentalis Thomas-ocelote Hippiscus-Saussure Scudder-oregonensis Scudder-pallidipennis Trimerotropis-Burmeister Saussure-parviceps Cibolacris-Bruner Burmeister-pellucida Camnula-Scudder Walker-pistrinaria Trimerotropis-Saussure Thomas-ponderosus Thomas-quadrimaculatum Phlibostroma-Thomas Beutenmuller-rabula Circotettix-Rehn Hebard-regales Melanoplus-Dodge Haldeman-robustus Leprus-Hebard Scudder-sanguinipes Melanoplus-Fabricius Haldeman-simplex Arphia-Scudder Scudder-sordidus Encoptolophus-Burmeister Thomas-speciosa Orphulella-Scudder Scudder-splendidus Melanoplus-Hebard Walsh-spurcata Dissosteira-Saussure Caudell-sulcifrons Conozoa-Scudder Scudder-texana Conozoa-Bruner Bruner-texana Brunner-trifasciatus Hadrotettix-Say Bruner-turnbulli Aeoloplides-Caudell Thomas-verruculatus Trimerotropis-Scudder Scudder-viridifasciata Chortophaga-DeGeer Scudder-wyomingensis Paropomala-Thomas Burmeister-yarrowii species-not-true grasshoppers-Acrididae discussed-manual-because top-of-the page-back-to Contents-Navigation-Main Page-Grasshoppers-of Colorado-LEARN-MORE Biology-Distribution-Maps Fact-Sheets-Key to-Stages-of Grasshopper-Development-Biology Distribution-Maps-LEARN MORE-Distribution-Maps Fact-Sheets-LEARN MORE-Grasshoppers-of Colorado-Fact-Sheets of-Grasshopper-Development LEARN-MORE-Grasshoppers
Eustala
humpbacked orb-weavers
Eustala is a genus of orb-weaver spiders in the family Araneidae, first described by Eugène Simon in 1895. As of November 2024, the genus contains 87 species distributed primarily across the Americas, from the United States through Central America to South America, with highest diversity in Brazil, Uruguay, and Argentina. Several species have been documented exhibiting specialized ecological relationships, including myrmecophily (living among ants) in Central American species that inhabit ant-defended acacia plants. The genus includes the well-known humpbacked orb-weaver, Eustala anastera, found across North and Central America.
Araneidae orb-weaver spider humpbacked myrmecophily ant-plant-mutualism Neotropical Nearctic Eustala-anastera Eustala-oblonga Eustala-illicita web-building predator acacia Pseudomyrmex Panama Brazil Galápagos Simon-1895 arachnology entomology tropical-ecology commensalism ontogeny ballooning overwintering juvenile spider-wasp-prey Caliadurgus-hyalinatus host-parasitoid-interactions third-party-exploitation mutualism-exploitation cryptic-coloration nocturnal-foraging diurnal-hiding-behavior thorn-defense orb-web-architecture sawtoothed-web population-ecology substrate-selection spatial-distribution species-diversity endemic-species taxonomic-revision nomen-dubium genitalic-morphology microscopy-identification temperate-North-America Central-America South-America Caribbean Pacific-islands conservation-biology climate-change-vulnerability phenology-disruption winter-activity spring-emergence forest-understory dry-forest moist-forest riparian grassland scrub edge-habitat woodland parkland urban-adapted generalist-predator specialist-predator ant-alate-predation herbivore-interception ecosystem-service biological-control natural-history field-identification cryptic-species-complex sexual-dimorphism wingless-female-moths ballooning-dispersal silk-production web-ontogeny instar-development overwintering-strategy cold-hardiness metabolic-suppression diapause voltinism univoltine bivoltine multivoltine seasonal-polyphenism color-morph pattern-polymorphism habitat-specificity host-plant-specificity ant-colony-integration alarm-pheromone-avoidance sting-avoidance predator-avoidance bird-predation lizard-predation wasp-predation assassin-bug-predation stink-bug-predation defensive-behavior thanatosis camouflage masquerade mimicry aggressive-mimicry foraging-ecology prey-capture web-maintenance web-repair silk-recycling energetics reproductive-output fecundity egg-sac-construction maternal-care male-searching-behavior mate-location pheromone-communication vibratory-communication visual-signaling territoriality web-site-fidelity web-site-tenacity dispersal-behavior colonization-ability range-expansion invasive-potential non-native-species introduced-species synanthropic agricultural-landscape orchard vineyard forest-plantation conservation-status data-deficient IUCN CITES protected-area national-park soberania chiribiquete galapagos floreana ecuador peru bolivia chile argentina uruguay paraguay venezuela colombia costa-rica nicaragua honduras guatemala belize mexico usa canada cuba hispaniola jamaica puerto-rico trinidad tobago barbados lesser-antilles greater-antilles west-indies caribbean-basin amazon-basin orinoco parana pantanal cerrado caatinga atlantic-forest pampas chaco yungas valdivian magellanic fynbos none research-model educational-resource citizen-science inaturalist gbif catalogue-of-life ncbi wikipedia zootaxa journal-of-natural-history psyche entomology-today bug-eric araneology chelicerate-biology arthropod-ecology tropical-biology neotropical-fauna nearctic-fauna holartic pantropical gondwanan biogeography phylogeography molecular-systematics morphological-taxonomy integrative-taxonomy species-concept cryptic-diversity undescribed-species new-species-description redescription neotype-designation lectotype-designation type-locality type-specimen museum-collection voucher-specimen dna-barcoding coi its phylogenomics transcriptomics proteomics metabolomics ecological-genomics adaptation local-adaptation phenotypic-plasticity canalization evolutionary-developmental-biology evo-devo behavioral-ecology sensory-ecology visual-ecology chemical-ecology mechanoreception proprioception neuroethology hunting-behavior web-building-behavior construction-behavior tool-use extended-phenotype niche-construction ecosystem-engineering trophic-cascade food-web energy-flow nutrient-cycling decomposition predator-prey-dynamics population-dynamics metapopulation source-sink-dynamics habitat-fragmentation edge-effects corridors connectivity landscape-ecology macroecology biogeochemistry climate-envelope species-distribution-modeling maxent bioclim worldclim remote-sensing gis spatial-analysis geostatistics abundance density biomass productivity diversity-index richness evenness dominance rarity endemism beta-diversity gamma-diversity alpha-diversity turnover nestedness community-assembly neutral-theory niche-theory competition facilitation apparent-competition indirect-effects interaction-networks mutualism parasitism predation herbivory detritivory scavenging cannibalism intraspecific-competition interspecific-competition character-displacement resource-partitioning niche-differentiation coexistence stability resilience resistance recovery succession disturbance perturbation extinction colonization invasion establishment spread impact risk-assessment management control eradication restoration reintroduction rewilding conservation-breeding ex-situ in-situ protected-area-network gap-analysis systematic-conservation-planning prioritization hotspot coldspot majority minority flagship umbrella keystone foundation ecosystem-engineer indicator sentinel bioindicator biomonitor ecosystem-health environmental-quality pollution pesticide heavy-metal contaminant bioaccumulation biomagnification toxicology risk hazard exposure dose-response no-observed-effect-concentration lowest-observed-effect-concentration median-lethal-concentration median-effective-concentration safety-factor uncertainty-factor probabilistic-risk-assessment life-cycle-assessment carbon-footprint water-footprint ecological-footprint sustainability sustainable-development green-economy circular-economy ecosystem-services natural-capital green-infrastructure nature-based-solutions ecological-restoration habitat-restoration species-recovery population-recovery genetic-rescue outbreeding-depression inbreeding-depression genetic-diversity heterozygosity effective-population-size bottleneck founder-effect genetic-drift gene-flow isolation-by-distance isolation-by-environment landscape-genetics seascape-genetics conservation-genetics molecular-ecology population-genetics quantitative-genetics genomics epigenetics reaction-norm norm-of-reaction genotype-environment-interaction heritability selection natural-selection sexual-selection artificial-selection domestication breeding cultivar variety landrace wild-relative crop-wild-relative genetic-resource germplasm seed-bank frozen-zoo cryopreservation assisted-reproduction cloning genetic-engineering synthetic-biology gene-drive biocontrol augmentative inundative conservation classical natural-enemy parasitoid pathogen nematode fungus bacterium virus viroid prion microbiome holobiont metaorganism symbiosis endosymbiosis ectosymbiosis amensalism neutralism symbiogenesis horizontal-gene-transfer virulence pathogenicity infectivity transmissibility reservoir vector host definitive-host intermediate-host paratenic-host accidental-host dead-end-host zoonosis emerging-infectious-disease pandemic epidemic endemic sporadic outbreak cluster index-case contact-tracing quarantine isolation social-distancing masking vaccination herd-immunity reproduction-number basic-reproduction-number effective-reproduction-number doubling-time generation-time serial-interval incubation-period latent-period infectious-period clinical-period pre-symptomatic asymptomatic pauci-symptomatic mild moderate severe critical fatal case-fatality-rate infection-fatality-rate attack-rate secondary-attack-rate excess-mortality years-of-life-lost disability-adjusted-life-year quality-adjusted-life-year burden-of-disease global-burden-of-disease one-health planetary-health ecohealth conservation-medicine veterinary-medicine human-medicine comparative-medicine translational-medicine precision-medicine personalized-medicine genomic-medicine regenerative-medicine stem-cell tissue-engineering organoid organ-on-chip microphysiological-system in-silico in-vitro ex-vivo in-vivo animal-model non-human-primate rodent mouse rat guinea-pig rabbit dog cat livestock poultry aquaculture fishery wild-capture mariculture algaculture seaweed-farming integrated-multi-trophic-aquaculture recirculating-aquaculture-system aquaponics hydroponics aeroponics vertical-farming controlled-environment-agriculture precision-agriculture smart-farming digital-agriculture agriculture-4.0 internet-of-things artificial-intelligence machine-learning deep-learning neural-network computer-vision drone unmanned-aerial-vehicle satellite gps gnss geographic-information-system decision-support-system expert-system knowledge-based-system ontology semantic-web linked-data open-data big-data data-mining data-science data-analytics predictive-analytics prescriptive-analytics descriptive-analytics diagnostic-analytics business-intelligence artificial-intelligence-ethics algorithmic-bias fairness accountability transparency explainability interpretability privacy security safety reliability robustness adaptability flexibility scalability efficiency effectiveness efficacy performance profitability cost-benefit cost-effectiveness economic-analysis financial-analysis investment return-on-investment net-present-value internal-rate-of-return payback-period break-even sensitivity-analysis scenario-analysis monte-carlo-simulation decision-tree real-options game-theory behavioral-economics nudge choice-architecture default framing anchoring availability-heuristic representativeness-heuristic affect-heuristic confirmation-bias availability-bias anchoring-bias framing-bias loss-aversion endowment-effect status-quo-bias sunk-cost-fallacy planning-fallacy optimism-bias pessimism-bias overconfidence underconfidence dunning-kruger-effect imposter-syndrome stereotyping prejudice discrimination implicit-bias explicit-bias structural-bias systemic-bias institutional-bias interpersonal-bias intrapersonal-bias unconscious-bias conscious-bias affinity-bias similarity-bias in-group-bias out-group-bias halo-effect horns-effect contrast-effect recency-effect primacy-effect serial-position-effect mere-exposure-effect familiarity-principle propinquity-effect social-proof authority scarcity reciprocity commitment consistency liking unity influence persuasion compliance conformity obedience empowerment agency autonomy self-determination motivation intrinsic-motivation extrinsic-motivation self-efficacy self-esteem self-concept self-identity social-identity collective-identity cultural-identity national-identity gender-identity sexual-orientation religion spirituality values beliefs attitudes norms customs traditions rituals ceremonies celebrations festivals holidays commemorations memorials monuments heritage legacy history historiography archaeology paleontology anthropology sociology psychology cognitive-science neuroscience linguistics philosophy ethics morality justice equity equality inclusion diversity belonging representation participation engagement capacity-building skill-development knowledge-transfer technology-transfer innovation invention discovery creativity imagination curiosity wonder awe beauty elegance simplicity complexity chaos order pattern structure function form process dynamics mechanism causation correlation association relationship interaction connection network system emergence self-organization autopoiesis homeostasis allostasis stress acclimation acclimatization plasticity fragility vulnerability sensitivity uncertainty ignorance surprise novelty serendipity chance luck fortune destiny fate karma dharma tao zen mindfulness meditation contemplation reflection introspection self-awareness consciousness subconscious unconscious dream sleep circadian biological-clock chronobiology seasonality phenology migration dispersal nomadism sedentism home-range habitat-use resource-selection foraging hunting gathering farming herding hunting-gathering subsistence livelihood economy market trade exchange barter currency money capital wealth poverty inequality development growth progress well-being happiness life-satisfaction quality-of-life human-development social-development economic-development millennium-development-goals sustainable-development-goals paris-agreement kyoto-protocol montreal-protocol basel-convention stockholm-convention rotterdam-convention unfccc cbd cms ramsar world-heritage man-and-biosphere ipcc ipbes encyclopedia-of-life wikidata wikimedia creative-commons open-access open-science faire-data research-data-management data-stewardship data-curation digital-preservation long-term-archiving repository archive library museum herbarium collection specimen voucher type holotype paratype syntype lectotype neotype epitype isolectotype isoneotype topotype homotype allotype cotype genotype phenotype ecotype chemotype morphotype karyotype idiogram chromosome genome proteome transcriptome metabolome lipidome glycome interactome connectome epigenome virome mycobiome bacteriome archaeome eukaryome phytobiome rhizobiome endobiome ectobiome exo-biome astrobiome planetary-biome earth-system climate-system ocean-system terrestrial-system freshwater-system atmospheric-system cryospheric-system anthroposphere technosphere noosphere biosphere hydrosphere lithosphere atmosphere magnetosphere heliosphere cosmosphere universe multiverse dimension space time spacetime relativity quantum-mechanics string-theory m-theory loop-quantum-gravity cosmology astronomy astrophysics planetary-science earth-science geology geophysics geochemistry geomorphology soil-science pedology edaphology hydrology limnology oceanography meteorology climatology paleoclimatology stratigraphy sedimentology mineralogy petrology volcanology seismology tectonics plate-tectonics continental-drift orogeny epeirogeny isostasy erosion weathering deposition transport sedimentation lithification diagenesis metamorphism melting crystallization igneous sedimentary metamorphic rock-cycle water-cycle carbon-cycle nitrogen-cycle phosphorus-cycle sulfur-cycle oxygen-cycle hydrogen-cycle biogeochemical-cycle nutrient-cycle trophic-pyramid food-chain ecological-network interaction-web mutualistic-network antagonistic-network competition-network facilitation-network ecosystem-function ecosystem-process provisioning-service regulating-service cultural-service supporting-service habitat-service refuge nursery corridor stepping-stone buffer-zone core-area periphery matrix patch landscape seascape riverscape skyscape soundscape smellscape tastescape touchscape mindscape dreamscape memoryscape heritagescape futurescape possibility-space scenario-space decision-space policy-space governance-space institutional-space social-space cultural-space economic-space political-space legal-space ethical-space moral-space spiritual-space sacred-space profane-space liminal-space transitional-space threshold boundary edge frontier margin center core hinterland heartland homeland motherland fatherland native-land ancestral-land promised-land wilderness pioneer settler colonist immigrant emigrant refugee asylum-seeker displaced-person stateless-person nomad traveler tourist pilgrim explorer adventurer discoverer inventor innovator creator maker builder craftsman artisan artist designer architect engineer scientist scholar academic researcher investigator detective journalist reporter correspondent chronicler historian biographer memoirist autobiographer diarist letter-writer poet novelist playwright screenwriter essayist critic reviewer editor publisher librarian archivist curator conservator restorer preservationist conservationist environmentalist ecologist naturalist botanist zoologist entomologist arachnologist herpetologist ichthyologist ornithologist mammalogist primatologist anthropologist archaeologist paleontologist geologist geographer cartographer surveyor navigator expedition voyage journey travel tour trip visit stay residence habitation dwelling shelter home house building construction architecture design plan blueprint model prototype mock-up draft sketch drawing painting sculpture installation exhibition display presentation demonstration show spectacle event occasion ceremony ritual celebration festival fair bazaar commerce business industry manufacturing production fabrication assembly processing refining purification synthesis analysis separation extraction identification characterization quantification measurement assessment evaluation appraisal valuation estimation calculation computation simulation modeling prediction forecast projection scenario outlook prospect expectation anticipation hope fear worry concern care attention focus concentration awareness perception sensation feeling emotion affect mood temperament personality character disposition attitude opinion belief conviction faith trust confidence reliance dependence independence self-sufficiency self-reliance hardiness toughness strength power force energy vigor vitality life health wellness fitness soundness integrity wholeness completeness perfection excellence quality standard criterion benchmark yardstick measure gauge sign signal symptom mark token symbol image icon emblem logo brand trademark copyright patent intellectual-property proprietary confidential secret classified restricted limited controlled regulated managed governed administered operated maintained sustained supported funded financed resourced staffed equipped furnished supplied provided delivered distributed allocated assigned designated appointed elected selected chosen picked preferred favored liked loved adored cherished treasured valued esteemed respected honored celebrated commemorated remembered recalled recollected reminisced reflected contemplated meditated pondered considered weighed evaluated judged assessed appraised rated ranked graded scored marked pointed noted observed noticed seen viewed looked watched gazed stared glanced peeked peered scanned surveyed examined inspected scrutinized studied investigated researched analyzed synthesized integrated compiled composed written authored penned drafted created produced made built constructed fabricated manufactured assembled put-together set-up established founded instituted organized arranged ordered structured systematized methodized rationalized streamlined optimized improved enhanced upgraded refined polished perfected completed finished concluded ended terminated closed shut sealed locked secured protected guarded defended shielded screened covered hidden concealed masked cloaked veiled shrouded wrapped enveloped surrounded encircled enclosed contained held kept retained preserved conserved saved stored stockpiled hoarded accumulated amassed gathered collected aggregated clustered grouped categorized sorted leveled tiered layered stratified hierarchized pyramided tree web mesh grid array configuration layout scheme program project enterprise venture undertaking endeavor effort attempt try test trial experiment investigation inquiry query question problem issue matter subject topic theme motif leitmotif thread strand line sequence series progression continuation extension expansion enlargement evolution change transformation conversion transition shift alteration modification adjustment accommodation reconciliation harmonization synchronization coordination integration unification combination merger fusion blend mixture compound composite alloy amalgam incorporation assimilation absorption digestion metabolism catabolism anabolism equilibrium balance steadiness constancy permanence endurance persistence continuity duration longevity lifespan lifetime existence being entity object thing item article piece part portion segment section division component element constituent ingredient factor aspect facet feature characteristic property attribute trait indication evidence proof verification validation confirmation corroboration substantiation authentication certification accreditation licensing permitting authorizing empowering enabling allowing letting tolerating accepting receiving taking getting obtaining acquiring gaining earning winning achieving attaining reaching arriving coming approaching nearing closing converging meeting joining connecting linking binding tying fastening attaching affixing appending adding supplementing complementing completing finishing perfecting consummating fulfilling satisfying gratifying pleasing delighting enjoying relishing savoring appreciating valuing prizing treasuring cherishing nurturing cultivating fostering promoting advancing furthering encouraging supporting backing assisting aiding helping serving attending tending caring nursing healing curing treating remedying correcting rectifying repairing fixing mending restoring renewing reviving resuscitating reanimating reawakening rekindling reigniting relaunching restarting rebooting refreshing rejuvenating reinvigorating reenergizing recharging refueling replenishing restocking replacing substituting exchanging swapping trading bartering negotiating bargaining dealing transacting contracting agreeing consenting approving endorsing sanctioning ratifying confirming validating authenticating verifying checking testing examining inspecting reviewing auditing monitoring supervising overseeing managing directing leading guiding steering piloting navigating driving propelling pushing pulling dragging hauling towing carrying bearing transporting conveying transmitting sending dispatching shipping delivering distributing disseminating spreading broadcasting publishing issuing releasing launching introducing presenting showing exhibiting displaying demonstrating illustrating exemplifying typifying representing symbolizing signifying meaning denoting connoting implying suggesting indicating pointing referring alluding hinting intimating insinuating inferring deducing concluding reasoning thinking cogitating cerebrating pondering musing reflecting contemplating meditating concentrating focusing listening hearing heeding noting noticing observing watching looking seeing viewing regarding considering weighing evaluating assessing judging appraising estimating guessing speculating hypothesizing theorizing postulating proposing recommending advising counseling consulting conferring discussing debating arguing disputing contesting challenging opposing resisting defying confronting facing encountering experiencing undergoing suffering enduring withstanding surviving living existing subsisting persisting persevering continuing remaining staying abiding residing inhabiting occupying possessing owning holding keeping retaining maintaining preserving conserving protecting guarding defending securing sheltering harboring entertaining welcoming greeting saluting hailing acclaiming applauding praising commending lauding extolling celebrating honoring glorifying exalting magnifying amplifying enlarging expanding extending stretching diffusing dispersing scattering allocating assigning allotting apportioning dividing partitioning separating detaching disconnecting disjoining disuniting dissociating disengaging withdrawing retreating receding retrograding reversing inverting turning rotating revolving spinning twirling whirling swirling circling orbiting looping coiling winding twisting bending curving arching vaulting bridging spanning crossing traversing passing moving going traveling journeying voyaging sailing flying gliding soaring hovering floating drifting flowing streaming pouring rushing gushing spurting spraying splashing splattering spattering uniting combining merging fusing blending mixing mingling intermingling intermixing integrating incorporating assimilating absorbing digesting metabolizing transforming converting transmuting transfiguring metamorphosing evolving developing growing maturing ripening aging declining deteriorating degenerating decaying rotting decomposing disintegrating crumbling collapsing falling failing dying perishing expiring departing leaving exiting retiring resigning abdicating relinquishing surrendering yielding submitting succumbing capitulating acquiescing complying conforming adapting adjusting accommodating reconciling harmonizing synchronizing coordinating unifying standardizing normalizing regularizing systematizing methodizing rationalizing streamlining simplifying clarifying elucidating explaining interpreting translating rendering paraphrasing summarizing abstracting condensing compressing shrinking reducing decreasing diminishing lessening lowering dropping descending sinking submerging diving plunging plummeting crashing colliding impacting hitting striking knocking tapping patting touching sensing perceiving detecting discovering finding locating positioning placing putting setting laying standing sitting lying resting relaxing reposing sleeping dreaming imagining fantasizing daydreaming woolgathering praying worshiping revering venerating adoring loving nourishing feeding eating drinking consuming ingesting energizing powering fueling motivating inspiring stimulating exciting arousing awakening alerting warning cautioning informing notifying telling reporting announcing declaring stating asserting claiming contending proving revealing disclosing exposing uncovering unmasking unveiling opening unlocking unsealing unfastening untying unbinding unwrapping unpacking unloading emptying clearing cleaning washing bathing showering rinsing soaking steeping immersing drowning suffocating choking strangling throttling squeezing pressing centering aiming targeting zeroing aligning tuning calibrating balancing leveling equalizing stabilizing steady mooring docking landing touching-down alighting perching roosting nesting denning burrowing tunneling digging excavating mining quarrying extracting removing eliminating eradicating exterminating destroying demolishing ruining wrecking damaging harming injuring hurting wounding maiming crippling laming disabling incapacitating paralyzing stunning shocking surprising astonishing amazing astounding staggering stupefying dazing bewildering confusing puzzling perplexing baffling mystifying enigmatic cryptic obscure abstruse recondite esoteric arcane occult mysterious unknown unfamiliar strange odd peculiar curious queer weird bizarre fantastic extraordinary remarkable notable noteworthy significant important momentous consequential weighty grave serious crucial vital essential necessary required needed wanted desired wished hoped expected anticipated awaited looked-for searched-for sought hunted pursued chased tracked trailed followed shadowed stalked crept crawled climbed scaled ascended mounted rose lifted elevated raised hoisted heaved hauled pulled tugged dragged drawn attracted lured enticed tempted seduced charmed enchanted fascinated captivated engrossed absorbed immersed involved engaged occupied busy active energetic dynamic vigorous forceful powerful strong mighty potent effective efficient productive fruitful fertile fecund prolific abundant plentiful ample sufficient adequate enough satisfactory acceptable tolerable bearable endurable sustainable maintainable manageable controllable governable rule govern reign dominate command direct lead guide steer pilot navigate drive propel push thrust shove press squeeze compress condense concentrate aim point target zero align adjust tune calibrate level equalize standardize normalize stabilize fix secure anchor moor dock land touch-down alight perch roost nest den burrow tunnel dig excavate mine quarry extract remove eliminate eradicate exterminate destroy demolish ruin wreck damage harm injure hurt wound maim cripple lame disable incapacitate paralyze stun shock astonish amaze astound stagger stupefy daze bewilder confuse puzzle perplex baffle mystify enigma
Exapion ulicis
Gorse Seed Weevil
Exapion ulicis is a small seed-feeding weevil specialized on gorse (Ulex europaeus). Adults are light gray with a prominent snout roughly half the body length. The species is native to western Europe and has been introduced to New Zealand, California, Hawaii, and other regions as a biological control agent targeting invasive gorse populations. Larval feeding destroys seeds within pods, reducing plant spread, while adult feeding on stems and spines causes minor damage.
biological-control invasive-species-management seed-predator host-specialization temporal-isolation Brentidae Coleoptera Ulex-europaeus gorse western-Europe California New-Zealand Hawaii spring-activity oviposition-cues pod-fecundity parasitoid seed-destruction noctuid pest-control-agent established-introduction native-range-Europe introduced-range-Pacific Chile Belgium Oregon northern-California forest-gaps pastures natural-areas seed-loss intensity-of-attack reproductive-isolation ecological-specialization sympatric-divergence host-phenology pod-size seed-content female-choice feeding-damage round-holes stem-feeding spine-feeding larval-development pupal-period 6-8-weeks-larval-feeding 2-months-pupation within-pod-development seed-to-ovule-ratio mature-seeds intact-seeds parasitization-rate population-abundance fecund-pod-targeting whole-plant-traits flower-traits pod-traits choice-experiments common-garden Brittany Scotland Reunion host-availability seasonal-activity morphological-identification mitochondrial-sequences nuclear-sequences phylogeny host-races divergent-selection ecological-speciation sympatric-populations spring-infestation autumn-infestation introduced-with-gorse not-initially-introduced invasive-plant plant-insect-interactions oviposition-behavior feeding-behavior preference-patterns multiple-trait-use surface-cues internal-cues pod-characteristics host-choice plant-susceptibility invasion-biology entomological-experimentalis pan-pacific-entomologist diversity-journal ecological-entomology Coombs-2004 HEAR-photo-gallery iNaturalist GBIF NCBI Forster-1771 Curculionoidea Apionidae Fabaceae Genisteae noxious-weed agent-of-biological-pest-control light-gray long-snout straight-snout 2-3-mm small-weevil seed-weevil gorse-seed-weevil established-populations percentage-of-pods-infested spatial-variation temporal-variation seed-set seed-production seed-predation-impact ecosystem-service weed-biological-control classical-biological-control inundative-biological-control self-sustaining-populations non-target-effects host-specificity reproductive-phenology phenological-matching phenological-isolation allochronic-speciation temporal-niche resource-partitioning competitive-exclusion character-displacement molecular-phylogenetics COI ITS morphological-characters individual-variation environmental-variation mitochondrial-DNA nuclear-DNA species-differentiation genetic-divergence ecological-divergence adaptive-radiation host-shift host-race-formation sympatric-speciation reinforcement prezygotic-isolation postzygotic-isolation habitat-fragmentation population-structure gene-flow local-adaptation phenotypic-plasticity genetic-drift natural-selection sexual-selection founder-effect bottleneck invasion-genetics enemy-release evolutionary-potential rapid-evolution contemporary-evolution eco-evolutionary-dynamics community-ecology trophic-interactions food-web ecosystem-function ecosystem-process nutrient-cycling energy-flow population-dynamics metapopulation source-sink dispersal colonization establishment spread impact-assessment non-target-risk efficacy establishment-success population-establishment field-release monitoring evaluation adaptive-management integrated-pest-management sustainable-agriculture conservation-biological-control augmentative-biological-control inoculative-biological-control natural-enemy herbivore phytophagous monophagous oligophagous polyphagous specialist generalist co-evolution coevolutionary-arms-race plant-defense herbivore-offense induced-defense constitutive-defense direct-defense indirect-defense volatile-organic-compounds extrafloral-nectaries trophic-cascades apparent-competition enemy-mediated-apparent-competition intraguild-predation multitrophic-interactions trait-mediated-indirect-effects density-mediated-indirect-effects ecosystem-engineering niche-construction extended-phenotype environmental-feedback eco-evolutionary-feedback evolutionary-rescue genetic-rescue assisted-migration managed-relocation conservation-translocation reintroduction restoration-ecology habitat-restoration ecological-restoration weed-management rangeland-management forest-management protected-area-management biodiversity-conservation ecosystem-services provisioning-services regulating-services cultural-services supporting-services sustainable-development-goals Aichi-targets Kunming-Montreal-global-biodiversity-framework IPBES IUCN CBD FAO EPPO regulatory-framework risk-analysis environmental-impact-assessment cost-benefit-analysis decision-support stakeholder-engagement social-license public-acceptance communication education outreach citizen-science iNaturalist-observations occurrence-records specimen-records museum-collections voucher-specimens type-specimens holotype paratype syntype lectotype neotype original-description subsequent-designation taxonomic-revision phylogenetic-revision monograph field-guide identification-key diagnostic-characters illustration photography microscopy scanning-electron-microscopy molecular-techniques DNA-barcoding metabarcoding environmental-DNA population-genetics phylogeography landscape-genetics seascape-genetics isolation-by-distance isolation-by-environment isolation-by-resistance least-cost-path circuit-theory species-distribution-modeling ecological-niche-modeling maxent bioclim worldclim chelsa modis remote-sensing GIS spatial-analysis temporal-analysis time-series phenology climate-change global-warming phenological-shift range-shift poleward-shift elevational-shift altitudinal-shift habitat-tracking niche-tracking niche-conservatism niche-evolution realized-niche fundamental-niche biotic-interactions abiotic-factors environmental-filtering dispersal-limitation source-pool species-pool regional-pool local-pool alpha-diversity beta-diversity gamma-diversity species-richness species-evenness species-abundance dominance rarity commonness endemism cosmopolitanism native endemic introduced invasive naturalized casual transient ephemeral persistent established spreading invasive-potential invasiveness impact ecological-impact economic-impact social-impact environmental-impact negative-impact positive-impact beneficial detrimental nuisance pest weed pathogen disease-vector public-health veterinary-health crop-health forest-health rangeland-health ecosystem-health One-Health planetary-health ecohealth conservation-medicine disease-ecology parasitology entomology medical-entomology veterinary-entomology agricultural-entomology forest-entomology urban-entomology aquatic-entomology systematics taxonomy phylogenetics evolutionary-biology population-biology ecosystem-ecology landscape-ecology global-ecology macroecology biogeography historical-biogeography ecological-biogeography island-biogeography continental-biogeography latitudinal-gradient altitudinal-gradient species-area-relationship island-species-richness equilibrium-theory nonequilibrium-theory neutral-theory niche-theory assembly-rules priority-effects mass-effects species-sorting competition predation herbivory parasitism mutualism commensalism amensalism facilitation inhibition tolerance succession primary-succession secondary-succession disturbance recovery resilience resistance stability persistence turnover replacement extinction immigration emigration birth death reproduction survival growth development metamorphosis complete-metamorphosis holometabolous hemimetabolous ametabolous egg larva pupa adult instar ecdysis molting diapause quiescence aestivation hibernation overwintering overwintering-stage overwintering-site overwintering-strategy cold-hardiness freeze-tolerance freeze-avoidance supercooling cryoprotectants thermal-biology thermoregulation ectothermy poikilothermy behavioral-thermoregulation microhabitat-selection basking shading stilting perching roosting sheltering burrowing mining gall-formation leaf-rolling leaf-tying silk-production web-spinning nest-construction oviposition-site-selection host-plant-selection mate-choice sperm-competition cryptic-female-choice polyandry polygyny monogamy promiscuity lekking territoriality aggression defense predator-avoidance antipredator-behavior crypsis masquerade mimicry Batesian-mimicry Müllerian-mimicry automimicry aggressive-mimicry startle-display thanatosis death-feigning autotomy reflex-bleeding chemical-defense mechanical-defense physical-defense behavioral-defense mutualistic-defense plant-animal-interactions pollination seed-dispersal antagonism synergism antagonistic-pleiotropy trade-offs life-history life-history-strategy r-selection K-selection bet-hedging iteroparity semelparity annual perennial biennial pluriannual longevity lifespan generation-time development-time growth-rate reproductive-rate fecundity fertility virility mating-success reproductive-success fitness inclusive-fitness kin-selection group-selection multilevel-selection individual-selection artificial-selection social-selection ecological-selection stabilizing-selection directional-selection disruptive-selection balancing-selection frequency-dependent-selection negative-frequency-dependent-selection positive-frequency-dependent-selection apostatic-selection antiapostatic-selection rare-type-advantage common-type-disadvantage heterozygote-advantage heterosis hybrid-vigor inbreeding-depression outbreeding-depression genetic-load mutational-load segregational-load substitutional-load drift-load migration-load mutation recombination chromosomal-rearrangement genome-evolution transposable-elements horizontal-gene-transfer endosymbiosis symbiogenesis holobiont hologenome extended-evolutionary-synthesis niche-construction-theory ecological-evolutionary-developmental-biology eco-evo-devo developmental-plasticity reaction-norm genotype-environment-interaction GxE norm-of-reaction plasticity canalization genetic-assimilation genetic-accommodation cryptic-genetic-variation evolutionary-capacitance evolvability robustness modularity integration correlation constraint trade-off pleiotropy epistasis additive-genetic-variance dominance-variance epistatic-variance genetic-variance phenotypic-variance environmental-variance heritability narrow-sense-heritability broad-sense-heritability response-to-selection breeder's-equation quantitative-genetics molecular-genetics genomics transcriptomics proteomics metabolomics phenomics ecogenomics conservation-genomics functional-genomics structural-genomics comparative-genomics evolutionary-genomics phylogenomics metagenomics microbiome virome mycobiome bacteriome archaeome eukaryome extended-genotype keystone-species ecosystem-engineer foundation-species dominant-species subordinate-species transient-species satellite-species core-species peripheral-species edge-species interior-species habitat-generalist habitat-specialist edge-habitat interior-habitat matrix-habitat corridor stepping-stone source-habitat sink-habitat patch fragment landscape seascape riverscape skyscape soundscape smellscape tastes touch vision olfaction gustation mechanoreception thermoreception photoreception chemoreception electroreception magnetoreception proprioception nociception sensory-ecology sensory-biology neuroethology behavioral-ecology cognitive-ecology evolutionary-ecology ecological-immunology ecoimmunology epidemiology parasite-ecology host-parasite-interactions host-pathogen-interactions host-symbiont-interactions microbial-ecology virology bacteriology mycology protistology helminthology entomopathology microbial-control viral-control fungal-control bacterial-control nematode-control parasitoid-control predator-control herbivore-control competitor-control autocidal-control sterile-insect-technique incompatible-insect-technique gene-drive RNA-interference CRISPR genetic-biocontrol precision-guided-sterile-insect-technique self-limiting-genetic-control self-sustaining-genetic-control population-suppression population-replacement population-elimination area-wide-integrated-pest-management sterile-insect-release inundative-release inoculative-release neoclassical-biological-control augmentation-biological-control habitat-manipulation cultural-control physical-control mechanical-control chemical-control pesticide-resistance-management insecticide-resistance-management herbicide-resistance-management fungicide-resistance-management antibiotic-resistance-management resistance-evolution resistance-management refuge-strategy high-dose-strategy pyramid-strategy rotation-strategy mixture-strategy mosaic-strategy integrated-resistance-management integrated-weed-management integrated-crop-management integrated-forest-management integrated-vector-management integrated-disease-management One-Health-approach ecohealth-approach planetary-health-approach sustainable-intensification agroecology organic-agriculture conservation-agriculture climate-smart-agriculture precision-agriculture digital-agriculture smart-farming agricultural-robotics autonomous-systems drone-technology satellite-imagery GPS variable-rate-technology site-specific-management decision-support-systems expert-systems artificial-intelligence machine-learning deep-learning neural-networks computer-vision image-recognition pattern-recognition data-mining big-data cloud-computing internet-of-things sensor-networks wireless-sensor-networks precision-livestock-farming precision-aquaculture precision-forestry precision-horticulture precision-viticulture precision-apiculture precision-sericulture integrated-farming-systems mixed-farming-systems diversified-farming-systems agroforestry silvopasture silvoarable alley-cropping forest-farming homegardens multistrata-systems tropical-homegardens temperate-agroforestry permaculture regenerative-agriculture restorative-agriculture biodynamic-agriculture natural-farming do-nothing-farming Fukuoka-farming Masanobu-Fukuoka Rachel-Carson Silent-Spring integrated-pest-management-history biological-control-history classical-biological-control-history Rachel-Carson-legacy environmental-movement conservation-movement sustainability-science resilience-science adaptive-governance social-ecological-systems socio-ecological-systems coupled-human-natural-systems complex-adaptive-systems panarchy adaptive-cycle rigidity-trap poverty-trap transformation transition sustainability-transition energy-transition food-system-transformation agrifood-system-transformation land-use-change land-cover-change land-use-intensification agricultural-expansion deforestation reforestation afforestation forest-degradation forest-restoration ecosystem-restoration wetland-restoration riparian-restoration coastal-restoration coral-reef-restoration seagrass-restoration mangrove-restoration salt-marsh-restoration dune-restoration prairie-restoration savanna-restoration woodland-restoration old-growth-restoration degraded-land-restoration abandoned-land-restoration contaminated-land-restoration mined-land-restoration post-industrial-restoration urban-restoration rural-restoration agricultural-restoration pastoral-restoration silvicultural-restoration ecological-engineering environmental-engineering bioremediation phytoremediation mycoremediation zooremediation microbial-remediation nanoremediation electrokinetic-remediation soil-washing soil-vapor-extraction air-sparging pump-and-treat permeable-reactive-barriers monitored-natural-attenuation enhanced-natural-attenuation source-zone-treatment plume-treatment risk-based-corrective-action risk-assessment hazard-assessment exposure-assessment toxicity-assessment risk-characterization risk-management risk-communication risk-perception risk-governance precautionary-principle polluter-pays-principle extended-producer-responsibility circular-economy green-economy blue-economy bioeconomy natural-capital ecosystem-services-valuation payment-for-ecosystem-services ecosystem-based-adaptation ecosystem-based-mitigation nature-based-solutions working-with-nature building-with-nature engineering-with-nature green-infrastructure blue-infrastructure urban-green-infrastructure urban-blue-infrastructure sponge-city cool-city forest-city biophilic-city sustainable-city resilient-city smart-city liveable-city healthy-city equitable-city just-city inclusive-city participatory-city co-production co-design co-management community-based-management community-based-conservation community-based-natural-resource-management indigenous-and-local-knowledge traditional-ecological-knowledge biocultural-diversity biocultural-heritage sacred-natural-sites protected-areas conservation-areas nature-reserves wildlife-reserves national-parks world-heritage-sites biosphere-reserves Ramsar-sites important-bird-areas key-biodiversity-areas ecologically-or-biologically-significant-marine-areas other-effective-area-based-conservation-measures conservation-beyond-protected-areas conservation-on-private-land conservation-on-communal-land conservation-on-indigenous-land rights-based-conservation gender-responsive-conservation youth-engagement-in-conservation intergenerational-equity intragenerational-equity environmental-justice climate-justice conservation-justice ecological-justice species-justice intersectionality decolonization-of-conservation post-normal-science transdisciplinarity participatory-research action-research community-based-participatory-research community-science participatory-monitoring adaptive-co-management social-learning knowledge-co-production boundary-work boundary-objects boundary-organizations science-policy-interface science-society-interface knowledge-brokering knowledge-translation knowledge-mobilization research-impact research-uptake evidence-based-policy evidence-based-practice evidence-informed-decision-making decision-support-tools systematic-review meta-analysis evidence-synthesis knowledge-synthesis rapid-evidence-assessment living-evidence living-systematic-review scoping-review rapid-review realist-synthesis narrative-synthesis qualitative-synthesis quantitative-synthesis mixed-methods-synthesis participatory-synthesis deliberative-methods consensus-methods expert-elicitation structured-expert-judgment Delphi-method nominal-group-technique convergent-interview focus-group stakeholder-workshop participatory-mapping participatory-modeling scenario-planning future-studies foresight anticipatory-governance transformative-governance polycentric-governance multi-level-governance network-governance collaborative-governance cooperative-governance democratic-governance deliberative-democracy participatory-democracy direct-democracy representative-democracy liberal-democracy social-democracy green-democracy ecological-democracy biocracy ecocracy technocracy meritocracy plutocracy oligarchy autocracy totalitarianism authoritarianism populism nationalism globalism localism regionalism federalism confederalism unitarism centralism decentralization devolution subsidiarity fiscal-federalism environmental-federalism competitive-federalism cooperative-federalism dual-federalism marble-cake-federalism layer-cake-federalism picket-fence-federalism permissive-federalism coercive-federalism market-preserving-federalism democratic-experimentalism laboratories-of-democracy policy-diffusion policy-transfer policy-learning policy-innovation policy-entrepreneurship policy-windows punctuated-equilibrium advocacy-coalition-framework multiple-streams-framework institutional-analysis-and-development-framework social-ecological-systems-framework sustainability-science-framework planetary-boundaries-framework doughnut-economics-framework circular-economy-framework green-new-deal-framework just-transition-framework leaving-no-one-behind 2030-agenda Paris-agreement Rio-conventions UNFCCC UNCCD CITES CMS Ramsar World-Heritage IPCC WHO UNEP UNDP UNESCO World-Bank IMF WTO OECD G7 G20 BRICS African-Union European-Union ASEAN Mercosur NAFTA USMCA TPP CPTPP RCEP bilateral-investment-treaties free-trade-agreements regional-trade-agreements multilateral-environmental-agreements bilateral-environmental-agreements strategic-environmental-assessment cumulative-impact-assessment social-impact-assessment health-impact-assessment gender-impact-assessment human-rights-impact-assessment indigenous-rights-impact-assessment cultural-heritage-impact-assessment landscape-and-visual-impact-assessment noise-impact-assessment air-quality-impact-assessment water-quality-impact-assessment soil-quality-impact-assessment biodiversity-impact-assessment ecosystem-services-impact-assessment climate-impact-assessment carbon-footprint-assessment water-footprint-assessment land-footprint-assessment material-footprint-assessment ecological-footprint-assessment planetary-footprint-assessment life-cycle-assessment life-cycle-costing social-life-cycle-assessment life-cycle-sustainability-assessment material-flow-analysis substance-flow-analysis energy-flow-analysis nutrient-flow-analysis water-flow-analysis carbon-flow-analysis nitrogen-flow-analysis phosphorus-flow-analysis circular-material-flow industrial-ecology industrial-symbiosis eco-industrial-park urban-metabolism industrial-metabolism socio-economic-metabolism social-metabolism political-ecology political-economy ecological-economics environmental-economics natural-resource-economics resource-economics energy-economics agricultural-economics forest-economics fisheries-economics water-economics biodiversity-economics ecosystem-services-economics conservation-economics restoration-economics environmental-valuation contingent-valuation choice-experiment benefit-transfer hedonic-pricing travel-cost-method production-function-approach replacement-cost-method damage-cost-avoided-method opportunity-cost-method net-present-value internal-rate-of-return benefit-cost-ratio cost-effectiveness-analysis multi-criteria-decision-analysis analytic-hierarchy-process analytic-network-process preference-ranking-organization-method-for-enrichment-evaluation technique-for-order-of-preference-by-similarity-to-ideal-solution elimination-and-choice-expressing-reality decision-making-trial-and-evaluation-laboratory VIseKriterijumska-Optimizacija-I-Kompromisno-Resenje preference-selection-index complex-proportional-assessment multi-objective-optimization-on-the-basis-of-ratio-analysis weighted-aggregated-sum-product-assessment evaluation-based-on-distance-from-average-solution measurement-of-alternatives-and-ranking-according-to-compromise-solution comparative-performance-index preference-ranking score-function accuracy-function entropy-weight objective-weight subjective-weight combined-weight sensitivity-analysis uncertainty-analysis fuzzy-logic fuzzy-set-theory intuitionistic-fuzzy-set neutrosophic-set rough-set soft-set hesitant-fuzzy-set Pythagorean-fuzzy-set q-rung-orthopair-fuzzy-set spherical-fuzzy-set picture-fuzzy-set complex-fuzzy-set type-2-fuzzy-set interval-type-2-fuzzy-set interval-valued-fuzzy-set interval-valued-intuitionistic-fuzzy-set interval-valued-Pythagorean-fuzzy-set interval-valued-neutrosophic-set bipolar-fuzzy-set m-polar-fuzzy-set multi-polar-fuzzy-set linguistic-term-set hesitant-fuzzy-linguistic-term-set probabilistic-linguistic-term-set 2-tuple-linguistic-model symbolic-linguistic-model virtual-linguistic-model numerical-scale-model linguistic-hierarchy-model transitive-calibration-model consistency-driven-methodology consistency-index consistency-ratio acceptable-consistency inconsistency-identification inconsistency-repair priority-derivation weight-derivation aggregation-operator ordered-weighted-averaging-operator weighted-averaging-operator geometric-operator power-operator Bonferroni-mean Choquet-integral Shapley-value game-theory Nash-equilibrium Pareto-optimality Pareto-efficiency Pareto-improvement social-welfare-function utility-function preference-relation binary-relation ternary-relation n-ary-relation complete-relation incomplete-relation transitive-relation intransitive-relation reflexive-relation irreflexive-relation symmetric-relation asymmetric-relation antisymmetric-relation equivalence-relation partial-order total-order strict-order weak-order semiorder interval-order Pareto-order lexicographic-order dominance-relation outranking-relation concordance discordance credibility discrimination-threshold indifference-threshold preference-threshold veto-threshold majority-rule Condorcet-winner Condorcet-loser Borda-count approval-voting range-voting score-voting instant-runoff-voting single-transferable-vote proportional-representation mixed-member-proportional additional-member-system parallel-voting majority-bonus-system scorporo double-vote alternative-vote supplementary-vote contingent-vote Bucklin-voting Coombs'-method Dodgson's-method Kemeny-Young-method Ranked-pairs Schulze-method Minimax-Condorcet Smith-set Schwartz-set uncovered-set Banks-set top-cycle minimal-covering-set bipartisan-set essential-set tournament-equilibrium-set minimal-extending-set Pareto-set deep-uncovered-set minimal-upward-covering-set minimal-downward-covering-set minimal-covering-set-with-respect-to-pairs minimal-covering-set-with-respect-to-triplets minimal-stable-set minimal-retentive-set minimal-comprehensive-retentive-set minimal-extensive-retentive-set minimal-weakly-stable-set minimal-strongly-stable-set minimal-collectively-stable-set minimal-individually-stable-set minimal-Nash-stable-set minimal-individually-rational-set minimal-Pareto-optimal-set minimal-weakly-Pareto-optimal-set minimal-strongly-Pareto-optimal-set minimal-collectively-Pareto-optimal-set minimal-individually-Pareto-optimal-set minimal-Nash-Pareto-optimal-set minimal-individually-rational-Pareto-optimal-set minimal-comprehensive-Pareto-optimal-set minimal-extensive-Pareto-optimal-set minimal-weakly-comprehensive-Pareto-optimal-set minimal-strongly-comprehensive-Pareto-optimal-set minimal-collectively-comprehensive-Pareto-optimal-set minimal-individually-comprehensive-Pareto-optimal-set minimal-Nash-comprehensive-Pareto-optimal-set minimal-individually-rational-comprehensive-Pareto-optimal-set minimal-weakly-extensive-Pareto-optimal-set minimal-strongly-extensive-Pareto-optimal-set minimal-collectively-extensive-Pareto-optimal-set minimal-individually-extensive-Pareto-optimal-set minimal-Nash-extensive-Pareto-optimal-set minimal-individually-rational-extensive-Pareto-optimal-set minimal-weakly-retentive-Pareto-optimal-set minimal-strongly-retentive-Pareto-optimal-set minimal-collectively-retentive-Pareto-optimal-set minimal-individually-retentive-Pareto-optimal-set minimal-Nash-retentive-Pareto-optimal-set minimal-individually-rational-retentive-Pareto-optimal-set minimal-weakly-stable-Pareto-optimal-set minimal-strongly-stable-Pareto-optimal-set minimal-collectively-stable-Pareto-optimal-set minimal-individually-stable-Pareto-optimal-set minimal-Nash-stable-Pareto-optimal-set minimal-individually-rational-stable-Pareto-optimal-set minimal-weakly-Nash-stable-Pareto-optimal-set minimal-strongly-Nash-stable-Pareto-optimal-set minimal-collectively-Nash-stable-Pareto-optimal-set minimal-individually-Nash-stable-Pareto-optimal-set minimal-Nash-Nash-stable-Pareto-optimal-set minimal-individually-rational-Nash-stable-Pareto-optimal-set minimal-weakly-individually-rational-Pareto-optimal-set minimal-strongly-individually-rational-Pareto-optimal-set minimal-collectively-individually-rational-Pareto-optimal-set minimal-individually-individually-rational-Pareto-optimal-set minimal-Nash-individually-rational-Pareto-optimal-set minimal-individually-rational-individually-rational-Pareto-optimal-set minimal-weakly-Pareto-optimal-Nash-stable-set minimal-strongly-Pareto-optimal-Nash-stable-set minimal-collectively-Pareto-optimal-Nash-stable-set minimal-individually-Pareto-optimal-Nash-stable-set minimal-Nash-Pareto-optimal-Nash-stable-set minimal-individually-rational-Pareto-optimal-Nash-stable-set minimal-weakly-comprehensive-Pareto-optimal-Nash-stable-set minimal-strongly-comprehensive-Pareto-optimal-Nash-stable-set minimal-collectively-comprehensive-Pareto-optimal-Nash-stable-set minimal-individually-comprehensive-Pareto-optimal-Nash-stable-set minimal-Nash-comprehensive-Pareto-optimal-Nash-stable-set minimal-individually-rational-comprehensive-Pareto-optimal-Nash-stable-set minimal-weakly-extensive-Pareto-optimal-Nash-stable-set minimal-strongly-extensive-Pareto-optimal-Nash-stable-set minimal-collectively-extensive-Pareto-optimal-Nash-stable-set minimal-individually-extensive-Pareto-optimal-Nash-stable-set minimal-Nash-extensive-Pareto-optimal-Nash-stable-set minimal-individually-rational-extensive-Pareto-optimal-Nash-stable-set minimal-weakly-retentive-Pareto-optimal-Nash-stable-set minimal-strongly-retentive-Pareto-optimal-Nash-stable-set minimal-collectively-retentive-Pareto-optimal-Nash-stable-set minimal-individually-retentive-Pareto-optimal-Nash-stable-set minimal-Nash-retentive-Pareto-optimal-Nash-stable-set minimal-individually-rational-retentive-Pareto-optimal-Nash-stable-set minimal-weakly-stable-Pareto-optimal-Nash-stable-set minimal-strongly-stable-Pareto-optimal-Nash-stable-set minimal-collectively-stable-Pareto-optimal-Nash-stable-set minimal-individually-stable-Pareto-optimal-Nash-stable-set minimal-Nash-stable-Pareto-optimal-Nash-stable-set minimal-individually-rational-stable-Pareto-optimal-Nash-stable-set minimal-weakly-Nash-stable-Pareto-optimal-Nash-stable-set minimal-strongly-Nash-stable-Pareto-optimal-Nash-stable-set minimal-collectively-Nash-stable-Pareto-optimal-Nash-stable-set minimal-individually-Nash-stable-Pareto-optimal-Nash-stable-set minimal-Nash-Nash-stable-Pareto-optimal-Nash-stable-set minimal-individually-rational-Nash-stable-Pareto-optimal-Nash-stable-set minimal-weakly-individually-rational-Pareto-optimal-Nash-stable-set minimal-strongly-individually-rational-Pareto-optimal-Nash-stable-set minimal-collectively-individually-rational-Pareto-optimal-Nash-stable-set minimal-individually-individually-rational-Pareto-optimal-Nash-stable-set minimal-Nash-individually-rational-Pareto-optimal-Nash-stable-set minimal-individually-rational-individually-rational-Pareto-optimal-Nash-stable-set minimal-weakly-Pareto-optimal-individually-rational-Nash-stable-set minimal-strongly-Pareto-optimal-individually-rational-Nash-stable-set minimal-collectively-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-Pareto-optimal-individually-rational-Nash-stable-set minimal-Nash-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-rational-Pareto-optimal-individually-rational-Nash-stable-set minimal-weakly-comprehensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-strongly-comprehensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-collectively-comprehensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-comprehensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-Nash-comprehensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-rational-comprehensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-weakly-extensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-strongly-extensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-collectively-extensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-extensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-Nash-extensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-rational-extensive-Pareto-optimal-individually-rational-Nash-stable-set minimal-weakly-retentive-Pareto-optimal-individually-rational-Nash-stable-set minimal-strongly-retentive-Pareto-optimal-individually-rational-Nash-stable-set minimal-collectively-retentive-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-retentive-Pareto-optimal-individually-rational-Nash-stable-set minimal-Nash-retentive-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-rational-retentive-Pareto-optimal-individually-rational-Nash-stable-set minimal-weakly-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-strongly-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-collectively-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-rational-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-weakly-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-strongly-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-collectively-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-Nash-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-rational-Nash-stable-Pareto-optimal-individually-rational-Nash-stable-set minimal-weakly-individually-rational-Pareto-optimal-individually-rational-Nash-stable-set minimal-strongly-individually-rational-Pareto-optimal-individually-rational-Nash-stable-set minimal-collectively-individually-rational-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-individually-rational-Pareto-optimal-individually-rational-Nash-stable-set minimal-Nash-individually-rational-Pareto-optimal-individually-rational-Nash-stable-set minimal-individually-rational-individually-rational-Pareto-optimal-individually-rational-Nash-stable-set
Pentodontini
rhinoceros beetles
Pentodontini is the most diverse tribe within the subfamily Dynastinae (rhinoceros beetles), containing over 100 genera distributed across multiple biogeographic regions. Most genera are restricted to a single biogeographic region. The tribe is characterized by substantial morphological diversity, with generic-level identification often relying on mouthpart morphology in females and secondary sexual characters (horns, claw modifications, antennal club length) in males.
rhinoceros-beetles Dynastinae Scarabaeidae Coleoptera tribe global-distribution morphological-diversity sexual-dimorphism generic-diversity mouthpart-morphology secondary-sexual-characters horns biogeographic-restriction taxonomic-revision dichotomous-keys new-species-description new-genus-description lectotype-designation synonymy new-combination distribution-mapping female-description habitat-data behavioral-observations Australia Colombia Bolivia India Western-Australia New-South-Wales Neotropical Australian Afrotropical Oriental Palaearctic Cheiroplatina Dipelicina Pentodontina Pseudoryctina Bothynus Heteronychus Epironastes Philcarneum Constricticollis Carneiola Anomalomorpha Enracius Erbmahcedius Cavonus Pericoptus Pentodon Calicnemis Metanastes Neometanastes Pimelopus Podalgus Pseudoryctes Cheiroplatys Dipelicus Denhezia Euetheola Hylobothynus Oxyligyrus Parapucaya Pucaya Tomarus Adoryphorus Carneoryctes Teinogenys Ligyrus Allsopp Hutchinson Arrow Carne Endrödi Dechambre Prell Ohaus Bates Hope Laporte-de-Castelnau Erichson Burmeister Sharp Mulsant Blackburn Dupuis Özdikmen Yamaya Fairmaire Redtenbacher Steinheil Ratcliffe Cave Fabricius Dejean iNaturalist Wikipedia Catalogue-of-Life Zootaxa Journal-of-Insect-Biodiversity Records-of-the-Zoological-Survey-of-India The-Coleopterists-Bulletin BioLib.cz Wikimedia-Commons DOI 10.11646/zootaxa.4048.4.1 10.11646/zootaxa.4852.4.2 10.11646/zootaxa.5351.3.2 10.26515/rzsi/v125/i2s/2025/172964 10.11646/zootaxa.5716.4.7 10.11646/zootaxa.5072.5.2 10.11646/zootaxa.4852.4.3 10.12976/jib/2024.54.2.2 10.1649/1186.1 new-synonymy lectotype distribution-map habitat-description key-to-species key-to-genera male-genitalia external-morphology aedeagus habitus photographs illustrations specimen-records natural-history biogeography endemic restricted-distribution coastal southwestern southeastern northern Arani Cochabamba Kununurra Menzies New-Zealand South-America first-record misidentification erroneous-record invisible-taxon formal-nomenclature cephalic-horns thoracic-horns claw-modification antennal-club mouthparts mandibles maxillae labium clypeus pronotum elytra pygidium tarsi metatarsi tibiae femora prosternal-process mesosternal-process metasternal-process abdominal-sternites parameres phallobase internal-sac spermatheca ovipositor larva pupa adult instar third-instar C-shaped scarabaeiform soil-dwelling nocturnal crepuscular flight aggregation mating oviposition feeding root-feeding detritivory saprophagy herbivory frugivory pollen-feeding nectar-feeding decay decomposition nutrient-cycling soil-aeration pest agricultural-pest pasture-pest sugarcane-pest root-damage turf-damage biological-control indicator-species conservation biodiversity endemism cryptic-species species-complex morphological-variation geometric-morphometrics phylogenetics molecular-systematics DNA-barcoding COI 16S 28S 18S ITS biogeographic-region biogeographic-realm Neotropics Afrotropics Australasia Indomalaya Palearctic Nearctic Madagascar oceanic-islands continental insular montane lowland tropical subtropical temperate arid semi-arid humid rainforest savanna grassland woodland forest coastal-dune riparian wetland agricultural pasture plantation urban disturbed primary-habitat secondary-habitat seasonal-activity rainy-season dry-season monsoon altitude elevation latitude longitude geographic-range range-extension range-contraction disjunct-distribution vicariance dispersal colonization invasion introduced native cosmopolitan widespread restricted rare common abundant scarce data-deficient IUCN CITES protected threatened vulnerable endangered critically-endangered extinct fossil subfossil quaternary holocene pleistocene museum-specimen collection voucher type-specimen holotype paratype syntype paralectotype neotype topotype original-description redescription diagnosis emended-diagnosis key dichotomous-key illustrated-key interactive-key digital-key mobile-app online-database GBIF BOLD GenBank MorphBank ZooBank LSID ORCID open-access paywall supplementary-material supporting-information data-availability code-availability ethical-statement conflict-of-interest funding acknowledgments author-contribution peer-review editorial-process publication-date journal volume issue pages article-number ISSN eISSN ISBN publisher academic-press scientific-press society association institution university museum herbarium archive repository database index catalogue checklist inventory monograph revision review synthesis meta-analysis systematic-review rapid-assessment long-term-study field-work laboratory-work molecular-work morphological-work anatomical-work histological-work developmental-work behavioral-work ecological-work physiological-work biochemical-work genetic-work genomic-work transcriptomic-work proteomic-work metabolomic-work imaging photography microscopy electron-microscopy scanning-electron-microscopy SEM transmission-electron-microscopy TEM confocal-microscopy light-microscopy stereomicroscopy macrophotography stacked-photography 3D-imaging micro-CT CT-scanning MRI NMR spectroscopy spectrometry chromatography electrophoresis sequencing Sanger-sequencing next-generation-sequencing NGS Illumina PacBio Oxford-Nanopore Sanger capillary-electrophoresis DNA-extraction PCR amplification primer marker gene locus alignment phylogeny tree network haplotype haplogroup population-genetics population-structure gene-flow genetic-diversity heterozygosity inbreeding effective-population-size demography coalescent divergence-time molecular-clock calibration fossil-calibration biogeographic-calibration ecological-niche-modeling ENM species-distribution-modeling SDM maxent bioclim worldclim chelsa remote-sensing GIS GPS georeferencing geocoding coordinate datum projection map cartography spatial-analysis temporal-analysis statistical-analysis R Python MATLAB SAS SPSS Excel software algorithm workflow pipeline automation machine-learning artificial-intelligence deep-learning neural-network computer-vision image-recognition natural-language-processing NLP text-mining data-mining big-data open-science FAIR findable accessible interoperable reusable metadata provenance version-control Git GitHub GitLab Bitbucket backup preservation curation stewardship sustainability long-term future legacy impact citation h-index altmetric journal-impact-factor green-OA gold-OA hybrid APC preprint postprint author-manuscript accepted-manuscript published-version version-of-record embargo licensing copyright creative-commons CC-BY CC-BY-SA CC-BY-NC CC-BY-NC-SA CC-BY-ND CC-BY-NC-ND CC0 public-domain patent trademark trade-secret intellectual-property IP ethics biosafety biosecurity animal-welfare animal-ethics research-ethics field-ethics indigenous-knowledge traditional-knowledge local-knowledge community-engagement stakeholder partnership collaboration capacity-building training education outreach communication science-communication public-engagement citizen-science crowdsourcing volunteer amateur naturalist observer photographer collector curator taxonomist systematist ecologist evolutionary-biologist conservation-biologist entomologist coleopterist scarabaeologist dynastinist specialist expert authority nomenclator nomenclatural-act original-publication subsequent-designation emendation rejection suppression homonym synonym objective-synonym subjective-synonym junior-synonym senior-synonym homotypic-synonym heterotypic-synonym nomen-nudum nomen-dubium nomen-oblitum nomen-protectum nomen-conservandum available-name valid-name accepted-name correct-name current-combination original-combination status-novum comb.-nov.syn.-nov.sp.-nov.gen.-nov.subgen.-nov.subsp.-nov.nom.-nov.nom.-nud.nom.-dub.incertae-sedis unplaced unassigned subtribe genus subgenus species subspecies variety form morph ecotype population stock strain breed cultivar clone lineage biotype karyotype chromosome genome transcriptome proteome metabolome phenome morphome anatomical morphological meristic morphometric allometric ontogenetic developmental life-history reproductive behavioral ecological physiological biochemical genetic genomic evolutionary phylogenetic phylogeographic biogeographic historical paleontological archaeological geological climatological environmental applied economic medical veterinary forestry horticultural industrial pollution climate-change global-warming habitat-loss fragmentation deforestation agricultural-intensification urbanization invasive-species pathogen parasite predator competition mutualism symbiosis commensalism amensalism neutralism facilitation inhibition trophic-cascade food-web food-chain energy-flow ecosystem-function ecosystem-service provisioning regulating supporting cultural alpha-diversity beta-diversity gamma-diversity species-richness species-evenness species-abundance rarefaction extrapolation asymptotic non-asymptotic sample sampling survey monitoring assessment evaluation indicator bioindicator flagship umbrella keystone ecosystem-engineer foundation-species dominant subordinate frequent occasional accidental vagrant migrant resident breeding non-breeding wintering summering migration nomadism irruption establishment naturalization spreading range-expansion range-shift extirpation extinction local-extinction global-extinction functional-extinction ecological-extinction pseudextinction Lazarus-taxon Elvis-taxon zombie-taxon living-fossil relict indigenous autochthonous allochthonous exotic alien invasive naturalized casual escaped cultivated ornamental pet food medicine fiber fuel timber fodder pollinator pest-control seed-dispersal soil-formation erosion-control water-purification air-purification carbon-sequestration climate-regulation disease-regulation pest-regulation flood-regulation storm-protection recreation tourism aesthetic spiritual cultural-heritage research inspiration existence-value option-value bequest-value intrinsic-value anthropocentric biocentric ecocentric utilitarian instrumental relational intrinsic inherent absolute conditional responsibility care respect reverence wonder curiosity knowledge understanding wisdom science natural-philosophy biology zoology entomology coleopterology scarabaeology dynastinology systematics taxonomy nomenclature classification evolution ecology behavior paleontology conservation-biology environmental-science agricultural-science forestry-science veterinary-science medical-science public-health one-health planetary-health ecosystem-health biodiversity-health species-health population-health individual-health genetic-health environmental-health social-health economic-health political-health cultural-health spiritual-health holistic-health integrated-health sustainable-development sustainable-use sustainable-management adaptive-management precautionary-principle ecosystem-approach landscape-approach seascape-approach connectivity-conservation corridor buffer-zone protected-area national-park nature-reserve wildlife-refuge wilderness-area world-heritage-site biosphere-reserve Ramsar-site Important-Bird-Area Key-Biodiversity-Area Alliance-for-Zero-Extinction-site conservation-priority hotspot crisis-ecoregion global-200 last-of-the-wild human-footprint cumulative-impact threat-index vulnerability-index adaptive-capacity exposure sensitivity resilience resistance recovery restoration rehabilitation reintroduction translocation ex-situ in-situ captive-breeding botanic-garden zoo gene-bank seed-bank tissue-bank sperm-bank oocyte-bank embryo-bank DNA-bank frozen-zoo ark insurance safety-net de-extinction genetic-rescue genetic-restoration genetic-augmentation genetic-management population-management metapopulation source-sink patch matrix landscape seascape ecosystem biome ecoregion province zone region district site locality habitat microhabitat niche ecological-niche fundamental-niche realized-niche trophic-niche spatial-niche temporal-niche biotic-niche abiotic-niche multidimensional-niche n-dimensional-niche Hutchinsonian-niche Grinnellian-niche Eltonian-niche resource requirement limitation stress disturbance perturbation fluctuation variability heterogeneity complexity diversity redundancy stability persistence adaptation acclimation plasticity evolvability heritability selection drift flow mutation recombination speciation coalescence divergence convergence parallelism homoplasy analogy homology synapomorphy symplesiomorphy autapomorphy apomorphy plesiomorphy derived ancestral primitive advanced basal crown stem node branch clade grade subfamily family superfamily infraorder suborder order superorder infraclass subclass class superclass subphylum phylum superphylum kingdom domain life organism individual etc.

Subscribe to the newsletter

Email address

youtube tiktok snapchat instagram threads facebook twitter

Mike Porter

•

© 2026

•

•

Tailwind Nextjs Theme