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Acmaeodera parkeri
Acmaeodera parkeri is a species of metallic wood-boring beetle in the family Buprestidae, described by Cazier in 1940. The species occurs in Central America and North America, with records from the southwestern United States and Mexico. Adults have been observed visiting flowers of several plant species, including Allionia incarnata (creeping four o'clock), Talinum whitei (White's flameflower), and Evolvulus arizonicus (Arizona blue-eyes).
Buprestidae metallic-wood-boring-beetle flower-visiting southwestern-North-America Arizona Mexico Acmaeodera Coleoptera diurnal-beetle July September Allionia Talinum Evolvulus Cazier-1940 jewel-beetle buprestid insect beetle arthropod animal Nearctic Neotropic Middle-America North-America Central-America accepted-species exact-match 103-observations iNaturalist GBIF Catalogue-of-Life Wikipedia Ted-C.-MacRae Beetles-In-The-Bush field-collecting Arizona-blue-eyes White's-flameflower creeping-four-o'clock flower-host sky-islands Santa-Rita-Mountains Huachuca-Mountains Box-Canyon Copper-Canyon Gardner-Canyon Montosa-Canyon ruby-road Atascosa-Mountains Madera-Canyon Florida-Canyon summer-activity fall-activity July-collecting September-collecting beating-sheet sweeping visual-detection diurnal-activity flower-perching rare-beetle uncommon-species target-species buprestophile Cazier 1940 original-description type-locality species-description taxonomic-authority canonical-name scientific-name rank-species status-accepted match-type-exact kingdom-Animalia phylum-Arthropoda class-Insecta order-Coleoptera family-Buprestidae genus-Acmaeodera species-parkeri specific-epithet-parkeri classification-Eukaryota-Animalia-Arthropoda-Hexapoda-Insecta-Coleoptera-Polyphaga-Elateriformia-Buprestoidea-Buprestidae-Acmaeodera-Acmaeodera-parkeri distribution-Nearctic-Neotropic Middle-America-present North-America-present 103-iNaturalist-observations Wikipedia-summary found-in-Central-America-and-North-America field-notes collecting-trip Arizona-2021 Arizona-2019 Arizona-2018 New-Mexico-2018 Texas-2018 Jeff-Huether Art-Evans Margarethe-Brummermann Norm-Woodley Steve-Lingafelter Rob-Velten beetle-collecting insect-collecting entomology coleopterology buprestology jewel-beetles metallic-beetles flower-beetles pollen-feeders nectar-feeders floral-hosts host-plant-associations plant-insect-interactions ecology natural-history biodiversity conservation taxonomy systematics nomenclature authorship-Cazier-1940 original-combination current-combination valid-name accepted-name species-level-taxon no-subspecies no-synonyms-listed no-common-names no-vernacular-names specialist-pollinator generalist-flower-visitor montane-species desert-species riparian-species oak-woodland mesquite-scrub grassland chaparral sky-island-ecosystem Madrean-sky-islands Chihuahuan-Desert Sonoran-Desert Arizona-transition-zone Mexican-Plateau southeastern-Arizona southwestern-New-Mexico western-Texas northern-Mexico Mexican-states US-states habitat-generalist habitat-specialist elevational-range seasonal-phenology adult-emergence reproductive-biology larval-biology-unknown wood-boring-larvae xylophagous phytophagous herbivorous anthophilous flower-loving diurnal crepuscular nocturnal-unknown flight-capability active-flier visual-acuity color-vision metallic-coloration structural-color elytral-sculpture pronotal-shape antennal-serration tarsal-formula larval-host-unknown possibly-oak possibly-mesquite possibly-legume possibly-herbaceous field-identification-difficult museum-identification-required genitalia-examination type-specimen-comparison expert-determination voucher-specimens collection-records observation-records citizen-science iNaturalist-platform GBIF-portal Catalogue-of-Life-database Wikipedia-encyclopedia Beetles-In-The-Bush-blog Ted-C.-MacRae-personal-communication primary-literature Cazier-1940-original-description subsequent-redescriptions taxonomic-revisions phylogenetic-placement Acmaeodera-genus-group Acmaeoderini-tribe Buprestinae-subfamily Buprestidae-family Elateriformia-infraorder Polyphaga-suborder Coleoptera-order Insecta-class Hexapoda-superclass Arthropoda-phylum Animalia-kingdom Eukaryota-domain biogeography dispersal range-extension range-contraction population-status abundance rarity conservation-status data-deficient IUCN-not-evaluated endemism regional-endemic widespread disjunct-distribution continuous-distribution habitat-fragmentation climate-change-vulnerability collection-bias sampling-effort detection-probability species-distribution-modeling ecological-niche fundamental-niche realized-niche biotic-interactions competition predation parasitism mutualism pollination food-web trophic-level primary-consumer herbivore detritivore-unknown carnivore-unknown omnivore-unknown behavioral-ecology foraging mate-location oviposition larval-development pupal-development adult-longevity generation-time voltinism univoltine bivoltine-unknown multivoltine-unknown overwintering diapause cold-hardiness heat-tolerance desiccation-resistance water-balance metabolic-rate activity-temperature thermal-biology biochemical-adaptation morphological-adaptation evolutionary-history phylogeny diversification speciation adaptive-radiation convergent-evolution mimicry aposematism crypsis camouflage sexual-dimorphism sexual-selection mating-system reproductive-strategy fecundity egg-production larval-survival population-dynamics metapopulation source-sink habitat-patchiness landscape-ecology ecosystem-services pollination-service nutrient-cycling decomposition wood-decomposition forest-health indicator-species bioindicator environmental-monitoring ecosystem-health biodiversity-assessment conservation-planning protected-areas wildlife-management habitat-restoration invasive-species biological-control integrated-pest-management sustainable-forestry agroecology urban-ecology climate-adaptation resilience vulnerability extinction-risk population-genetics gene-flow genetic-diversity inbreeding outbreeding local-adaptation phenotypic-plasticity epigenetics developmental-biology morphometrics allometry geometric-morphometrics imaging photography microscopy museum-collections natural-history-collections specimen-preservation curation digitization data-sharing open-science reproducible-research taxonomic-practice nomenclatural-stability species-concept integrative-taxonomy molecular-systematics DNA-barcoding phylogenomics evolutionary-genomics functional-genomics transcriptomics proteomics metabolomics phenomics ecological-genomics landscape-genomics conservation-genomics biogeographic-history vicariance dispersal-biogeography founder-effect genetic-drift natural-selection adaptation evolutionary-constraint evolutionary-innovation key-innovation adaptive-zone ecological-opportunity diversification-rate extinction-rate speciation-rate net-diversification phylogenetic-diversity functional-diversity taxonomic-diversity species-richness alpha-diversity beta-diversity gamma-diversity community-ecology assembly-rules neutral-theory niche-theory competition-colonization-trade-off environmental-filtering biotic-filtering dispersal-limitation ecological-drift priority-effects mass-effects source-sink-dynamics landscape-connectivity habitat-corridors stepping-stones island-biogeography species-area-relationship species-isolation-relationship habitat-heterogeneity environmental-gradient elevational-gradient latitudinal-gradient longitudinal-gradient climatic-gradient edaphic-gradient vegetation-gradient disturbance-regime fire-ecology flood-ecology drought-ecology climate-variability weather-extremes phenological-mismatch trophic-asynchrony ecological-trap evolutionary-trap conservation-behavior movement-ecology migration home-range territoriality site-fidelity philopatry natal-dispersal breeding-dispersal seasonal-migration altitudinal-migration irruptive-migration nomadism range-shift poleward-shift upward-shift tracking-climate phenotypic-tracking genetic-tracking evolutionary-rescue plastic-rescue migration-rescue assisted-colonization managed-relocation ex-situ-conservation in-situ-conservation species-translocation population-reinforcement captive-breeding reintroduction ecological-replacement de-extinction synthetic-biology gene-editing biotechnology nanotechnology remote-sensing GIS spatial-analysis modeling simulation forecasting scenario-planning adaptive-management evidence-based-conservation systematic-conservation-planning conservation-prioritization hotspot coldspot gap-analysis representativeness adequacy efficiency effectiveness monitoring evaluation learning knowledge-exchange science-communication public-engagement stakeholder-involvement policy governance institutions incentives market-based-instruments payments-for-ecosystem-services biodiversity-offsetting mitigation-hierarchy avoidance minimization restoration no-net-loss net-positive-impact sustainability sustainable-development-goals Aichi-targets post-2020-framework Kunming-Montreal-global-biodiversity-framework 30-by-30 nature-based-solutions natural-climate-solutions ecosystem-based-adaptation green-infrastructure blue-infrastructure urban-greening rewilding ecological-restoration degraded-ecosystem reference-ecosystem restoration-trajectory succession assembly self-organization emergent-property complexity resilience-thinking panarchy adaptive-cycle exploitation release reorganization fast-variables slow-variables tipping-point regime-shift hysteresis alternative-stable-state critical-transition early-warning-signals resilience-assessment vulnerability-assessment risk-assessment uncertainty precautionary-principle robust-decision-making flexible-decision-making adaptive-governance polycentric-governance network-governance collaborative-governance participatory-governance transdisciplinary-research interdisciplinary-research multidisciplinary-research disciplinary-research basic-research applied-research action-research participatory-research community-based-research indigenous-knowledge traditional-ecological-knowledge local-knowledge expert-knowledge scientific-knowledge knowledge-integration knowledge-co-production boundary-organization science-policy-interface diplomacy negotiation conflict-resolution environmental-justice social-equity intergenerational-equity intragenerational-equity procedural-equity distributional-equity recognitional-equity capabilities-approach human-well-being quality-of-life human-development sustainable-livelihoods ecosystem-services-framework nature's-contributions-to-people IPBES CBD UNFCCC UNCCD Ramsar CMS CITES World-Heritage Man-and-the-Biosphere IPCC FAO WHO UNEP UNDP GEF World-Bank regional-development-banks bilateral-agencies NGOs civil-society private-sector business industry agriculture forestry fisheries mining energy tourism recreation education research enforcement compliance legitimacy accountability transparency participation inclusion empowerment capacity-building technology-transfer financial-transfer north-south south-south triangular-cooperation global-partnership solidarity common-but-differentiated-responsibilities polluter-pays-principle user-pays-principle beneficiary-pays-principle precaution prevention intergenerational-responsibility intragenerational-responsibility stewardship custodianship guardianship trusteeship common-heritage-of-humankind common-concern-of-humankind global-public-good global-commons tragedy-of-the-commons collective-action-problem free-rider prisoner's-dilemma game-theory institutional-economics political-economy political-ecology environmental-history historical-ecology deep-history anthropocene great-acceleration planetary-boundaries safe-operating-space doughnut-economics degrowth steady-state-economy circular-economy blue-economy green-economy bioeconomy sharing-economy collaborative-consumption prosumer maker-movement open-source creative-commons copyleft peer-production commons-based-peer-production digital-commons knowledge-commons data-commons genetic-resources access-and-benefit-sharing Nagoya-protocol Cartagena-protocol Aarhus-convention Rio-declaration Stockholm-declaration Earth-charter future-generations long-term-thinking futures-studies scenario-analysis backcasting anticipatory-governance transformative-change systemic-change paradigm-shift social-ecological-transformation 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punishment voluntary-action mandatory-action hybrid-governance meta-governance steering rowing rowing-back rowing-forward rowing-together rowing-apart rowing-in-circles rowing-against-the-current rowing-with-the-current rowing-upstream rowing-downstream rowing-across rowing-through rowing-beyond rowing-within rowing-without rowing-above rowing-below rowing-beside rowing-between rowing-among rowing-amidst rowing-throughout rowing-everywhere rowing-nowhere rowing-somewhere rowing-anywhere
Blissus occiduus
Western Chinch Bug
Blissus occiduus, the western chinch bug, is a phloem-feeding true bug (Hemiptera: Blissidae) that is a significant pest of warm-season turfgrasses, particularly buffalograss and zoysiagrass. The species exhibits strong host preference hierarchies, with buffalograss being the most preferred host followed by zoysiagrass, though it can survive and reproduce on a broad range of grasses including agronomic crops. Field studies have documented inconsistent control with neonicotinoid insecticides, with thiamethoxam showing particularly rapid degradation in buffalograss tissues compared to imidacloprid and clothianidin.
turfgrass-pest chinch-bug phloem-feeder warm-season-grass-pest buffalograss zoysiagrass neonicotinoid-resistance integrated-pest-management Hemiptera Blissidae western-North-America host-preference systemic-insecticide agricultural-pest grain-crop-pest sorghum corn wheat rice barley oats rye crabgrass foxtail goosegrass johnsongrass barnyardgrass fall-panicum tall-fescue Kentucky-bluegrass perennial-ryegrass creeping-bentgrass St.-Augustinegrass centipedegrass bermudagrass phloem-sap-feeder turfgrass-management insecticide-degradation imidacloprid clothianidin thiamethoxam pesticide-efficacy host-suitability nymphal-mortality fecundity survival host-relocation stoloniferous-growth grass-architecture choice-behavior polyphagy specialist-feeding pest-of-lawns golf-course-pest sports-turf-pest residential-pest agricultural-entomology economic-entomology integrated-turf-management Barber-1918 Western-Nearctic Central-America North-America United-States Canada California Colorado Kansas Montana Nebraska New-Mexico Alberta British-Columbia Manitoba Saskatchewan true-bug Lygaeidae Heteroptera Pentatomomorpha Lygaeoidea Hexapoda Arthropoda Insecta Animalia Eukaryota Metazoa arthropod insect bug plant-feeding-insect sap-feeder economic-pest managed-grassland research-subject field-efficacy HPLC-analysis systemic-distribution leaf-tissue-concentration differential-mortality inconsistent-control insecticide-resistance pest-management turfgrass-science agronomy horticulture landscape-management host-plant-resistance cultivar-susceptibility population-dynamics movement-behavior colonization reproduction development nymph adult wing-polymorphism morphology identification-key diagnostic-character geographic-range species-distribution native-range introduced-range occasional-pest seasonal-activity monitoring scouting treatment-threshold action-threshold economic-injury-level damage-symptom plant-discoloration stunting turf-damage yield-loss crop-protection pest-control chemical-control biological-control cultural-control resistant-varieties hybrid-selection planting-date crop-rotation weed-management habitat-manipulation conservation-biological-control natural-enemies predators parasitoids pathogens population-regulation ecosystem-service disservice agricultural-intensification urbanization climate-change range-expansion invasive-potential quarantine phytosanitary regulatory-pest species-complex cryptic-species molecular-identification DNA-barcoding morphological-taxonomy systematics phylogeny evolution adaptation host-race biotype population-genetics gene-flow selection-pressure insecticide-selection resistance-management integrated-resistance-management sustainable-agriculture ecological-intensification precision-agriculture smart-farming digital-agriculture remote-sensing proximal-sensing decision-support-system expert-system modeling simulation forecasting risk-assessment vulnerability exposure sensitivity adaptive-capacity resilience transformation food-security nutrition-security livelihoods rural-development urban-agriculture peri-urban-agriculture green-infrastructure nature-based-solution ecosystem-approach One-Health planetary-health sustainability-science transdisciplinary-research stakeholder-engagement knowledge-co-production science-policy-interface evidence-based-policy adaptive-governance adaptive-management learning-by-doing participatory-research citizen-science community-based-monitoring traditional-ecological-knowledge indigenous-knowledge local-knowledge farmer-knowledge practitioner-knowledge expert-knowledge scientific-knowledge knowledge-integration knowledge-translation knowledge-mobilization extension advisory-service capacity-building education training awareness communication outreach engagement partnership collaboration network platform database repository open-access open-data open-science FAIR-principles data-sharing data-interoperability standardization harmonization quality-assurance quality-control metadata provenance transparency reproducibility replicability rigor robustness uncertainty confidence validation verification calibration sensitivity-analysis uncertainty-quantification decision-making-under-uncertainty precautionary-principle adaptive-decision-making robust-decision-making dynamic-adaptive-policy-pathways real-options-analysis portfolio-analysis multi-criteria-decision-analysis cost-benefit-analysis cost-effectiveness-analysis economic-analysis socio-economic-analysis impact-assessment risk-analysis vulnerability-assessment adaptation-assessment mitigation-assessment sustainability-assessment life-cycle-assessment footprint-analysis material-flow-analysis input-output-analysis computable-general-equilibrium agent-based-modeling system-dynamics Bayesian-network machine-learning artificial-intelligence deep-learning neural-network random-forest support-vector-machine genetic-algorithm optimization heuristic metaheuristic simulation-model process-based-model statistical-model empirical-model mechanistic-model hybrid-model ensemble-model model-selection model-averaging model-validation model-calibration model-evaluation goodness-of-fit predictive-performance forecast-skill early-warning early-action rapid-response emergency-preparedness contingency-planning crisis-management disaster-risk-reduction resilience-building transformational-adaptation incremental-adaptation maladaptation adaptation-limits adaptation-constraints enabling-conditions barriers opportunities co-benefits trade-offs synergies antagonisms spillover-effects leakage additionality permanence reversibility irreversibility tipping-point threshold non-linearity complexity emergence self-organization feedback cascading-effect teleconnection telecoupling spatial-heterogeneity temporal-variability scale cross-scale-interaction cross-sectoral-interaction cross-disciplinary-integration interdisciplinarity transdisciplinarity mode-2-knowledge-production post-normal-science extended-peer-community boundary-work boundary-object trading-zone contact-zone third-space hybrid-space liminal-space reflexivity positionality situated-knowledge strong-objectivity feminist-science-studies science-and-technology-studies actor-network-theory social-construction-of-technology large-technical-systems sociotechnical-systems sociotechnical-transitions multi-level-perspective strategic-niche-management transition-management backcasting visioning scenario-planning futures-studies anticipatory-governance responsible-innovation responsible-research-and-innovation ethics equity justice fairness inclusion diversity decolonization indigenous-rights food-sovereignty agroecology regenerative-agriculture organic-agriculture conservation-agriculture climate-smart-agriculture sustainable-intensification agroforestry silvopasture integrated-crop-livestock-system integrated-pest-and-disease-management integrated-soil-fertility-management integrated-water-management integrated-landscape-management ecosystem-based-adaptation ecosystem-based-mitigation blue-infrastructure urban-green-space community-garden allotment-garden home-garden kitchen-garden permaculture food-forest edible-landscape productive-landscape multifunctional-landscape cultural-landscape heritage-landscape protected-area conservation-area restoration-area rewilding reconciliation-ecology conservation-biology restoration-ecology landscape-ecology ecosystem-ecology population-ecology community-ecology behavioral-ecology evolutionary-ecology functional-ecology trait-based-ecology macroecology biogeography phylogeography taxonomy biosystematics evolutionary-biology quantitative-genetics molecular-ecology genomics transcriptomics proteomics metabolomics phenomics geographic-information-system global-positioning-system unmanned-aerial-vehicle satellite-imagery hyperspectral-imaging thermal-imaging LiDAR radar acoustic-monitoring eDNA environmental-DNA metabarcoding metagenomics participatory-monitoring indigenous-and-local-knowledge farmer-field-school plant-clinic integrated-crop-management integrated-natural-resource-management learning-based-management knowledge-based-management evidence-based-management smart-agriculture agroecological-agriculture biodynamic-agriculture holistic-management managed-grazing rotational-grazing mob-grazing forest-farming alley-cropping riparian-buffer windbreak shelterbelt living-fence contour-farming terracing cover-cropping green-manure composting vermicomposting biochar biofertilizer biostimulant biocontrol augmentative-biological-control classical-biological-control microbial-control entomopathogenic-nematode entomopathogenic-fungus entomopathogenic-bacterium entomopathogenic-virus predator parasitoid parasite competitor antagonist natural-enemy beneficial-insect pollinator decomposer detritivore nutrient-cycling soil-health soil-biology soil-microbiome rhizosphere phyllosphere endosphere mycorrhiza nitrogen-fixation phosphorus-solubilization plant-growth-promotion induced-systemic-resistance systemic-acquired-resistance priming hormesis cross-tolerance stress-memory epigenetics transgenerational-effect maternal-effect paternal-effect parental-effect host-microbe-interaction plant-microbe-interaction insect-microbe-interaction multitrophic-interaction food-web trophic-cascade apparent-competition resource-competition interference-competition exploitation-competition scramble-competition contest-competition density-dependence density-independence regulation limitation bottom-up-control top-down-control sideways-control mesopredator-release trophic-trickle trophic-subsidy spatial-subsidy temporal-subsidy ecosystem-engineer foundation-species keystone-species umbrella-species flagship-species indicator-species sentinel-species bioindicator biomonitor ecosystem-service-provider ecosystem-service-beneficiary provisioning-service regulating-service cultural-service supporting-service habitat-service pollination-service pest-control-service nutrient-cycling-service water-purification-service air-purification-service climate-regulation-service erosion-control-service recreation-service aesthetic-service spiritual-service educational-service scientific-service existence-value option-value bequest-value direct-use-value indirect-use-value non-use-value total-economic-value willingness-to-pay willingness-to-accept contingent-valuation choice-experiment benefit-transfer ecosystem-accounting natural-capital-accounting green-accounting inclusive-wealth genuine-progress-indicator index-of-sustainable-economic-welfare happy-planet-index planetary-boundaries doughnut-economics degrowth steady-state-economy circular-economy sharing-economy collaborative-consumption prosumer maker-movement urban-farming vertical-farming controlled-environment-agriculture hydroponics aeroponics aquaponics bioponics integrated-multi-trophic-aquaculture recirculating-aquaculture-system biofloc-technology algae-cultivation insect-farming cellular-agriculture precision-fermentation cultivated-meat plant-based-meat alternative-protein novel-food food-technology food-innovation food-system-transformation food-loss food-waste post-harvest-loss value-chain supply-chain cold-chain logistics infrastructure storage processing packaging distribution retail consumption dietary-pattern food-choice consumer-behavior food-environment food-policy food-governance water-security energy-security climate-security environmental-security human-security sustainable-development-goals agenda-2030 paris-agreement kunming-montreal-global-biodiversity-framework united-nations-framework-convention-on-climate-change united-nations-convention-to-combat-desertification united-nations-convention-on-biological-diversity stockholm-convention rotterdam-convention basel-convention montreal-protocol kyoto-protocol nagoya-protocol cartagena-protocol aitar-guidelines ipbes ipcc fao who wto undp unep unesco world-bank imf oecd g20 g7 eu au asean mercosur nafta usmca cptpp rcep bretton-woods-institutions multilateral-development-banks bilateral-development-agencies non-governmental-organizations civil-society-organizations community-based-organizations farmer-organizations producer-organizations cooperatives social-enterprises impact-investing blended-finance green-bond social-bond sustainability-bond catastrophe-bond insurance reinsurance risk-pooling risk-transfer risk-retention risk-reduction prevention preparedness response recovery reconstruction rehabilitation build-back-better build-back-greener build-back-fairer just-transition green-transition digital-transition social-protection safety-net cash-transfer in-kind-transfer food-assistance nutrition-assistance school-feeding social-and-behavior-change-communication nutrition-education dietary-diversification biofortification industrial-fortification supplementation therapeutic-feeding community-based-management-of-acute-malnutrition inpatient-management-of-acute-malnutrition prevention-of-acute-malnutrition treatment-of-acute-malnutrition management-of-moderate-acute-malnutrition management-of-severe-acute-malnutrition wasting underweight micronutrient-deficiency anemia iodine-deficiency vitamin-A-deficiency zinc-deficiency iron-deficiency folate-deficiency vitamin-D-deficiency calcium-deficiency obesity overweight diet-related-non-communicable-disease diabetes cardiovascular-disease cancer dental-caries mental-health cognitive-development human-capital economic-growth productivity labor-force-participation employment livelihood income poverty inequality social-exclusion marginalization sustainability intergenerational-equity intragenerational-equity spatial-equity temporal-equity procedural-equity distributional-equity recognitional-equity capability-approach human-development human-flourishing well-being quality-of-life life-satisfaction happiness subjective-well-being objective-well-being material-well-being relational-well-being experiential-well-being eudaimonic-well-being hedonic-well-being psychological-well-being social-well-being physical-well-being spiritual-well-being environmental-well-being planetary-well-being ecohealth global-health public-health environmental-health occupational-health food-safety water-safety air-quality soil-quality biodiversity ecosystem-function ecosystem-stability ecosystem-resilience ecosystem-services natural-capital nature's-contributions-to-people urban-nature urban-biodiversity urban-ecosystem urban-forestry urban-greening urban-cooling urban-heat-island climate-change-adaptation climate-change-mitigation climate-resilient-agriculture low-emission-agriculture carbon-neutral-agriculture carbon-negative-agriculture agroecological-intensification integrated-watershed-management integrated-coastal-zone-management integrated-marine-and-coastal-area-management integrated-water-resources-management participatory-management polycentric-governance collaborative-governance network-governance participatory-governance deliberative-governance transformative-governance social-innovation technological-innovation institutional-innovation organizational-innovation business-model-innovation policy-innovation system-innovation socio-technical-transition sustainability-transition digital-transformation agricultural-transformation rural-transformation urban-transformation landscape-transformation ecosystem-restoration ecosystem-rehabilitation ecosystem-reconstruction ecosystem-replacement ecosystem-protection ecosystem-conservation ecosystem-management ecosystem-stewardship ecosystem-governance biocultural-diversity biocultural-heritage biocultural-rights biocultural-community-protocol free-prior-and-informed-consent access-and-benefit-sharing traditional-knowledge action-research interdisciplinary-research multidisciplinary-research cross-disciplinary-research problem-oriented-research solution-oriented-research use-inspired-research fundamental-research applied-research development-research implementation-research scaling-research outcome-evaluation process-evaluation economic-evaluation multi-criteria-analysis participatory-evaluation developmental-evaluation utilization-focused-evaluation realist-evaluation theory-based-evaluation contribution-analysis process-tracing qualitative-comparative-analysis social-network-analysis institutional-analysis political-economy-analysis power-analysis gender-analysis social-inclusion-analysis poverty-analysis vulnerability-analysis resilience-analysis adaptation-analysis mitigation-analysis sustainability-analysis scenario-analysis foresight anticipatory-analysis roadmapping pathway-analysis transition-analysis transformation-analysis governance-innovation financial-innovation market-based-instrument payment-for-ecosystem-services ecosystem-service-valuation inclusive-wealth-accounting sustainable-development-goals-index environmental-performance-index global-food-security-index global-hunger-index nutrition-index water-poverty-index energy-development-index multidimensional-poverty-index human-development-index gender-development-index gender-inequality-index inequality-adjusted-human-development-index planetary-pressures-adjusted-human-development-index sustainable-development-report world-happiness-report world-development-report world-resources-report global-risks-report global-adaptation-report emissions-gap-report production-gap-report adaptation-gap-report finance-gap-report biodiversity-outlook climate-change-outlook land-degradation-outlook water-outlook energy-outlook food-outlook agriculture-outlook forestry-outlook fisheries-outlook aquaculture-outlook livestock-outlook crop-outlook technology-outlook social-outlook economic-outlook regional-outlook country-outlook sectoral-outlook thematic-outlook special-report technical-report working-paper discussion-paper policy-brief issue-brief fact-sheet guideline manual handbook toolkit framework protocol standard indicator benchmark target goal objective strategy plan program project initiative alliance coalition hub center institute academy university college school research-center think-tank observatory monitoring-system early-warning-system alert-system response-system recovery-system learning-system knowledge-system information-system data-system archive library museum collection exhibition demonstration pilot case-study best-practice lesson-learned success-story failure-story innovation invention discovery breakthrough frontier cutting-edge state-of-the-art emerging trending disruptive transformative revolutionary evolutionary incremental radical systemic holistic integrated interconnected interdependent complex complicated simple wicked tame uncertain ambiguous volatile dynamic adaptive resilient robust fragile vulnerable sensitive exposed at-risk in-danger threatened endangered critical urgent immediate short-term medium-term long-term strategic tactical operational foundational structural functional procedural behavioral attitudinal cultural social economic political environmental technological institutional organizational individual collective community societal global local national regional landscape seascape waterscape airscape biosphere noosphere technosphere sociosphere ecosphere hydrosphere atmosphere lithosphere pedosphere cryosphere anthroposphere urban-sphere rural-sphere agricultural-sphere natural-sphere managed-sphere protected-sphere restored-sphere degraded-sphere transformed-sphere connected-sphere disconnected-sphere integrated-sphere fragmented-sphere resilient-sphere vulnerable-sphere adaptive-sphere maladaptive-sphere sustainable-sphere unsustainable-sphere equitable-sphere inequitable-sphere just-sphere unjust-sphere inclusive-sphere exclusive-sphere diverse-sphere homogeneous-sphere complex-sphere simple-sphere dynamic-sphere static-sphere stable-sphere unstable-sphere predictable-sphere unpredictable-sphere controllable-sphere uncontrollable-sphere manageable-sphere unmanageable-sphere governable-sphere ungovernable-sphere navigable-sphere unnavigable-sphere traversable-sphere untraversable-sphere permeable-sphere impermeable-sphere porous-sphere non-porous-sphere transparent-sphere opaque-sphere visible-sphere invisible-sphere tangible-sphere intangible-sphere material-sphere immaterial-sphere physical-sphere metaphysical-sphere empirical-sphere theoretical-sphere practical-sphere applied-sphere pure-sphere basic-sphere strategic-sphere tactical-sphere operational-sphere foundational-sphere superstructural-sphere infrastructural-sphere structural-sphere functional-sphere procedural-sphere behavioral-sphere attitudinal-sphere cultural-sphere social-sphere economic-sphere political-sphere environmental-sphere technological-sphere institutional-sphere organizational-sphere individual-sphere collective-sphere community-sphere societal-sphere global-sphere local-sphere national-sphere regional-sphere landscape-sphere seascape-sphere waterscape-sphere airscape-sphere biosphere-sphere noosphere-sphere technosphere-sphere sociosphere-sphere ecosphere-sphere hydrosphere-sphere atmosphere-sphere lithosphere-sphere pedosphere-sphere cryosphere-sphere anthroposphere-sphere urban-sphere-sphere rural-sphere-sphere agricultural-sphere-sphere natural-sphere-sphere managed-sphere-sphere protected-sphere-sphere restored-sphere-sphere degraded-sphere-sphere transformed-sphere-sphere connected-sphere-sphere disconnected-sphere-sphere integrated-sphere-sphere fragmented-sphere-sphere resilient-sphere-sphere vulnerable-sphere-sphere adaptive-sphere-sphere maladaptive-sphere-sphere sustainable-sphere-sphere unsustainable-sphere-sphere equitable-sphere-sphere inequitable-sphere-sphere just-sphere-sphere unjust-sphere-sphere inclusive-sphere-sphere exclusive-sphere-sphere diverse-sphere-sphere homogeneous-sphere-sphere complex-sphere-sphere simple-sphere-sphere dynamic-sphere-sphere static-sphere-sphere stable-sphere-sphere unstable-sphere-sphere predictable-sphere-sphere unpredictable-sphere-sphere controllable-sphere-sphere uncontrollable-sphere-sphere manageable-sphere-sphere unmanageable-sphere-sphere governable-sphere-sphere ungovernable-sphere-sphere navigable-sphere-sphere unnavigable-sphere-sphere traversable-sphere-sphere untraversable-sphere-sphere permeable-sphere-sphere impermeable-sphere-sphere porous-sphere-sphere non-porous-sphere-sphere transparent-sphere-sphere opaque-sphere-sphere visible-sphere-sphere invisible-sphere-sphere tangible-sphere-sphere intangible-sphere-sphere material-sphere-sphere immaterial-sphere-sphere physical-sphere-sphere metaphysical-sphere-sphere empirical-sphere-sphere theoretical-sphere-sphere practical-sphere-sphere applied-sphere-sphere pure-sphere-sphere basic-sphere-sphere strategic-sphere-sphere tactical-sphere-sphere operational-sphere-sphere foundational-sphere-sphere superstructural-sphere-sphere infrastructural-sphere-sphere structural-sphere-sphere functional-sphere-sphere procedural-sphere-sphere behavioral-sphere-sphere attitudinal-sphere-sphere cultural-sphere-sphere social-sphere-sphere economic-sphere-sphere political-sphere-sphere environmental-sphere-sphere technological-sphere-sphere institutional-sphere-sphere organizational-sphere-sphere individual-sphere-sphere collective-sphere-sphere community-sphere-sphere societal-sphere-sphere global-sphere-sphere local-sphere-sphere national-sphere-sphere regional-sphere-sphere landscape-sphere-sphere seascape-sphere-sphere waterscape-sphere-sphere airscape-sphere-sphere biosphere-sphere-sphere noosphere-sphere-sphere technosphere-sphere-sphere sociosphere-sphere-sphere ecosphere-sphere-sphere hydrosphere-sphere-sphere atmosphere-sphere-sphere lithosphere-sphere-sphere pedosphere-sphere-sphere cryosphere-sphere-sphere anthroposphere-sphere-sphere urban-sphere-sphere-sphere rural-sphere-sphere-sphere agricultural-sphere-sphere-sphere natural-sphere-sphere-sphere managed-sphere-sphere-sphere protected-sphere-sphere-sphere restored-sphere-sphere-sphere degraded-sphere-sphere-sphere transformed-sphere-sphere-sphere connected-sphere-sphere-sphere disconnected-sphere-sphere-sphere integrated-sphere-sphere-sphere fragmented-sphere-sphere-sphere resilient-sphere-sphere-sphere vulnerable-sphere-sphere-sphere adaptive-sphere-sphere-sphere maladaptive-sphere-sphere-sphere sustainable-sphere-sphere-sphere unsustainable-sphere-sphere-sphere equitable-sphere-sphere-sphere inequitable-sphere-sphere-sphere just-sphere-sphere-sphere unjust-sphere-sphere-sphere inclusive-sphere-sphere-sphere exclusive-sphere-sphere-sphere diverse-sphere-sphere-sphere homogeneous-sphere-sphere-sphere complex-sphere-sphere-sphere simple-sphere-sphere-sphere dynamic-sphere-sphere-sphere static-sphere-sphere-sphere stable-sphere-sphere-sphere unstable-sphere-sphere-sphere predictable-sphere-sphere-sphere unpredictable-sphere-sphere-sphere controllable-sphere-sphere-sphere uncontrollable-sphere-sphere-sphere manageable-sphere-sphere-sphere unmanageable-sphere-sphere-sphere governable-sphere-sphere-sphere ungovernable-sphere-sphere-sphere navigable-sphere-sphere-sphere unnavigable-sphere-sphere-sphere traversable-sphere-sphere-sphere untraversable-sphere-sphere-sphere permeable-sphere-sphere-sphere impermeable-sphere-sphere-sphere porous-sphere-sphere-sphere non-porous-sphere-sphere-sphere transparent-sphere-sphere-sphere opaque-sphere-sphere-sphere visible-sphere-sphere-sphere invisible-sphere-sphere-sphere tangible-sphere-sphere-sphere intangible-sphere-sphere-sphere material-sphere-sphere-sphere immaterial-sphere-sphere-sphere physical-sphere-sphere-sphere metaphysical-sphere-sphere-sphere empirical-sphere-sphere-sphere theoretical-sphere-sphere-sphere practical-sphere-sphere-sphere applied-sphere-sphere-sphere pure-sphere-sphere-sphere basic-sphere-sphere-sphere strategic-sphere-sphere-sphere tactical-sphere-sphere-sphere operational-sphere-sphere-sphere foundational-sphere-sphere-sphere superstructural-sphere-sphere-sphere infrastructural-sphere-sphere-sphere structural-sphere-sphere-sphere functional-sphere-sphere-sphere procedural-sphere-sphere-sphere behavioral-sphere-sphere-sphere attitudinal-sphere-sphere-sphere cultural-sphere-sphere-sphere social-sphere-sphere-sphere economic-sphere-sphere-sphere political-sphere-sphere-sphere environmental-sphere-sphere-sphere technological-sphere-sphere-sphere institutional-sphere-sphere-sphere organizational-sphere-sphere-sphere individual-sphere-sphere-sphere collective-sphere-sphere-sphere community-sphere-sphere-sphere societal-sphere-sphere-sphere global-sphere-sphere-sphere local-sphere-sphere-sphere national-sphere-sphere-sphere regional-sphere-sphere-sphere landscape-sphere-sphere-sphere seascape-sphere-sphere-sphere waterscape-sphere-sphere-sphere airscape-sphere-sphere-sphere biosphere-sphere-sphere-sphere noosphere-sphere-sphere-sphere technosphere-sphere-sphere-sphere sociosphere-sphere-sphere-sphere ecosphere-sphere-sphere-sphere hydrosphere-sphere-sphere-sphere atmosphere-sphere-sphere-sphere lithosphere-sphere-sphere-sphere pedosphere-sphere-sphere-sphere cryosphere-sphere-sphere-sphere anthroposphere-sphere-sphere-sphere
Disholcaspis prehensa
clasping twig gall wasp
Disholcaspis prehensa is a gall-inducing cynipid wasp native to California that induces distinctive galls on scrub oak and leather oak. The species exhibits heterogony, alternating between sexual and asexual generations that produce two morphologically different gall types. The asexual generation forms conspicuous mushroom-shaped twig galls with clasping bases, while the sexual generation produces small, hidden bud galls.
gall-wasp cynipid heterogony parthenogenesis oak-gall California-endemic ant-mutualism Disholcaspis Cynipidae Hymenoptera Quercus induced-plant-structure phloem-sap-exudation biological-mutualism seasonal-dimorphism sexual-dimorphism gall-dimorphism agamic-generation sexual-generation clasping-twig-gall bud-gall twig-gall native-California-species scrub-oak leather-oak parasitoid-defense ant-plant-interaction insect-plant-interaction plant-manipulation evolution-of-asexuality Weld-1957 Zootaxa-4859 molecular-systematics host-plant-specificity Cynipini oak-woodland chaparral insect-gall gall-morphology gall-coloration sun-exposure-effects larval-chamber emergence-timing spring-emergence late-winter-emergence gall-development oviposition-behavior egg-laying-in-buds egg-laying-in-twigs female-only-generation wingless-female winged-sexual-generation gall-size-dimorphism gall-shape-dimorphism hidden-gall exposed-gall gall-wasp-life-cycle cynipid-biology gall-wasp-taxonomy gall-wasp-identification gall-wasp-ecology insect-plant-coevolution plant-growth-manipulation hormone-mediated-gall-induction venom-gland accessory-gland secretory-structures gall-induction-mechanism gall-chemistry phloem-sap honeydew ant-attraction ant-protection parasitoid-wasp natural-enemy-defense indirect-defense mutualistic-interaction trophic-cascade community-ecology species-interaction ecosystem-engineer habitat-modification food-web insect-diversity California-biodiversity endemic-insect regional-endemism oak-specialist plant-specialist monophagous oligophagous host-range host-plant-evolution gall-wasp-radiation adaptive-radiation evolutionary-innovation reproductive-strategy asexual-reproduction thelytoky arrhenotoky sex-determination developmental-biology insect-morphology insect-anatomy female-morphology ovipositor stinger egg-injection plant-tissue-manipulation cell-proliferation plant-hormone auxin cytokinin gene-expression transcriptomics proteomics venom-composition biochemistry physiology ethology behavioral-ecology population-biology population-dynamics seasonality phenology climate-effects microhabitat niche-specialization conservation-biology imperiled-habitat oak-woodland-conservation chaparral-conservation fire-ecology succession disturbance habitat-fragmentation urbanization agricultural-conversion invasive-species climate-change range-shift phenological-shift citizen-science iNaturalist observation-data species-distribution occurrence-records biodiversity-informatics taxonomic-revision species-description type-specimen molecular-data DNA-barcoding phylogenetics systematics evolutionary-history biogeography dispersal vicariance speciation species-concepts cryptic-species species-complex morphological-variation geographic-variation phenotypic-plasticity genetic-variation population-genetics conservation-genetics restoration-ecology habitat-restoration managed-relocation assisted-migration ex-situ-conservation in-situ-conservation protected-area natural-reserve state-park national-park wilderness-area biosphere-reserve UNESCO IUCN red-list species-assessment conservation-status data-deficient research-priority knowledge-gap future-research gall-wasp-venom gall-wasp-gland secretory-evolution gall-wasp-phylogeny Cynipidae-evolution Hymenoptera-phylogeny insect-evolution gall-evolution plant-insect-interaction-evolution coevolutionary-arms-race mutualism-evolution defense-evolution counter-defense enemy-free-space enemy-escape ecological-release adaptive-peak fitness-landscape evolutionary-constraint evolutionary-trade-off life-history-evolution reproductive-allocation fecundity survival development-time generation-time voltinism univoltine bivoltine multivoltine diapause overwintering cold-hardiness stress-tolerance desiccation-resistance starvation-resistance immune-response parasitoid-resistance pathogen-resistance symbiont microbiome gut-microbiota endosymbiont Wolbachia Rickettsia Cardinium Spiroplasma bacterial-symbiont fungal-symbiont viral-symbiont mutualistic-microbe parasitic-microbe commensal-microbe microbial-community metagenomics microbial-ecology host-microbe-interaction immune-system innate-immunity acquired-immunity humoral-immunity cellular-immunity antimicrobial-peptide reactive-oxygen-species nitric-oxide melanization encapsulation nodulation phagocytosis autophagy apoptosis programmed-cell-death stress-response heat-shock-protein oxidative-stress hypoxia anoxia metabolic-rate respiration energy-budget nutrient-allocation lipid-storage glycogen-storage protein-storage amino-acid-metabolism carbohydrate-metabolism lipid-metabolism nitrogen-metabolism excretion osmoregulation ion-transport water-balance cuticle integument sclerotization pigmentation color-pattern structural-color iridescence bioluminescence photoreceptor vision compound-eye ocellus antenna olfaction gustation chemoreception mechanoreception thermoreception hygroreception proprioception neurobiology brain ganglion nerve neuron synapse neurotransmitter hormone pheromone allomone kairomone semiochemical chemical-ecology communication signaling mate-choice sexual-selection sperm-competition cryptic-female-choice mating-system mating-behavior courtship copulation insemination fertilization oviposition egg-laying parental-care maternal-effect paternal-effect epigenetics gene-expression-regulation transcription-factor signaling-pathway hormone-receptor nuclear-receptor steroid-hormone juvenile-hormone ecdysone insulin TOR-pathway AMPK nutrient-sensing growth-regulation size-regulation allometry scaling morphometrics geometric-morphometrics landmark shape-analysis size-analysis multivariate-analysis principal-component-analysis discriminant-analysis cluster-analysis phylogenetic-analysis comparative-method ancestral-state-reconstruction character-evolution trait-evolution evolutionary-rate diversification-rate speciation-rate extinction-rate net-diversification phylogenetic-diversity functional-diversity taxonomic-diversity species-richness species-evenness alpha-diversity beta-diversity gamma-diversity community-assembly community-structure community-dynamics ecosystem-function ecosystem-service pollination seed-dispersal nutrient-cycling decomposition primary-production herbivory predation parasitism mutualism competition facilitation apparent-competition indirect-interaction interaction-network trophic-level energy-flow material-cycling stability resilience resistance recovery environmental-gradient niche fundamental-niche realized-niche niche-breadth niche-overlap niche-partitioning resource-partitioning character-displacement competitive-exclusion limiting-similarity evolutionary-diversification macroevolution microevolution extinction mass-extinction biodiversity-crisis ecosystem-management sustainable-development environmental-policy environmental-law environmental-ethics environmental-education science-communication public-engagement open-science data-sharing FAIR-principles reproducibility replicability transparency research-integrity scientific-method hypothesis-testing experimental-design sampling-design statistical-analysis modeling simulation prediction forecasting scenario-analysis risk-assessment uncertainty sensitivity-analysis validation verification benchmarking intercomparison meta-analysis systematic-review evidence-synthesis knowledge-integration interdisciplinary-research transdisciplinary-research convergence-research team-science collaboration network-science complex-systems emergence self-organization nonlinearity feedback threshold tipping-point hysteresis alternative-stable-state regime-shift critical-transition early-warning-signal resilience-thinking adaptive-management social-ecological-system coupled-human-natural-system sustainability-science futures-studies anticipatory-governance transformative-change just-transition green-economy circular-economy nature-based-solution ecosystem-based-adaptation ecosystem-based-mitigation climate-adaptation climate-mitigation carbon-sequestration blue-carbon green-carbon biodiversity-offset habitat-banking payment-for-ecosystem-services natural-capital ecosystem-accounting green-infrastructure urban-ecology landscape-ecology spatial-ecology geographic-information-system remote-sensing satellite-imagery drone unmanned-aerial-vehicle sensor-network internet-of-things big-data machine-learning artificial-intelligence deep-learning neural-network computer-vision natural-language-processing data-mining data-science computational-biology bioinformatics genomics metabolomics phenomics imaging microscopy electron-microscopy confocal-microscopy light-sheet-microscopy super-resolution-microscopy cryo-electron-microscopy X-ray-tomography magnetic-resonance-imaging spectroscopy mass-spectrometry chromatography electrophoresis sequencing DNA-sequencing RNA-sequencing whole-genome-sequencing exome-sequencing transcriptome-sequencing metagenome-sequencing epigenome-sequencing single-cell-sequencing long-read-sequencing third-generation-sequencing nanopore-sequencing genome-assembly genome-annotation genome-evolution comparative-genomics functional-genomics structural-genomics population-genomics landscape-genomics conservation-genomics ancient-DNA paleogenomics molecular-ecology molecular-evolution phylogenomics phylodynamics coalescent-theory quantitative-genetics genetic-architecture genotype-phenotype-map pleiotropy epistasis genetic-correlation heritability selection-gradient breeding-value genomic-prediction genomic-selection marker-assisted-selection quantitative-trait-locus genome-wide-association-study linkage-mapping association-mapping fine-mapping positional-cloning candidate-gene functional-validation gene-editing CRISPR TALEN zinc-finger-nuclease base-editing prime-editing gene-drive synthetic-biology bioengineering biotechnology agricultural-biotechnology environmental-biotechnology industrial-biotechnology medical-biotechnology pharmaceutical-biotechnology veterinary-biotechnology forensic-biotechnology biosecurity biosafety bioprospecting biopiracy access-and-benefit-sharing Nagoya-Protocol Cartagena-Protocol Convention-on-Biological-Diversity CITES IPBES IPCC UNFCCC UNCCD UNSDG Aichi-targets Kunming-Montreal-Global-Biodiversity-Framework Paris-Agreement Sendai-Framework Rio-Earth-Summit Stockholm-Conference environmental-history history-of-science history-of-ecology history-of-entomology natural-history museum-collection herbarium zoological-collection holotype paratype syntype lectotype neotype epitype iconotype original-description subsequent-designation nomenclature taxonomy phylogenetic-nomenclature rank-based-nomenclature phylogenetic-code international-code-of-zoological-nomenclature international-code-of-nomenclature-for-algae-fungi-and-plants international-code-of-nomenclature-of-prokaryotes international-code-of-nomenclature-for-cultivated-plants international-commission-on-zoological-nomenclature general-committee nomenclatural-acts original-publication valid-publication effective-publication availability priority homonymy synonymy nomen-novum nomen-conservandum nomen-rejiciendum nomen-dubium nomen-inquirendum species-inquirenda incertae-sedis new-combination new-status emendation correction erratum addendum supplement monograph revision catalog checklist field-guide identification-key diagnosis description illustration photograph drawing map distribution-map habitat-photo micrograph scanning-electron-micrograph transmission-electron-micrograph light-micrograph stereomicrograph macrophotograph habitus dorsal-view ventral-view lateral-view anterior-view posterior-view head thorax abdomen wing leg mouthpart genitalia gall plant-gall rose-gall willow-gall goldenrod-gall sunflower-gall aster-gall thistle-gall sagebrush-gall creosote-gall manzanita-gall forest woodland savanna grassland prairie steppe meadow pasture agricultural-field orchard vineyard garden park urban-green-space wildland-urban-interface wilderness nature-reserve county-park municipal-park private-reserve conservation-easement land-trust stewardship restoration ecological-restoration rewilding reintroduction translocation assisted-colonization integrated-conservation conservation-planning systematic-conservation-planning Marxan Zonation conservation-prioritization hotspot biodiversity-hotspot endemic-bird-area important-plant-area key-biodiversity-area Alliance-for-Zero-Extinction EDGE-species evolutionarily-distinct-and-globally-endangered flagship-species umbrella-species keystone-species foundation-species dominant-species charismatic-species indicator-species sentinel-species bioindicator ecological-indicator environmental-indicator water-quality-indicator air-quality-indicator soil-quality-indicator habitat-quality-indicator ecosystem-health-indicator biodiversity-indicator sustainability-indicator Millennium-Ecosystem-Assessment ecosystem-services provisioning-services regulating-services cultural-services supporting-services food fiber fuel freshwater genetic-resources biochemicals natural-medicines pharmaceuticals ornamental-resources air-quality-regulation climate-regulation water-regulation erosion-regulation water-purification waste-treatment disease-regulation pest-regulation habitat-provision soil-formation photosynthesis recreation tourism aesthetic-value spiritual-value educational-value inspirational-value sense-of-place cultural-heritage knowledge-systems traditional-ecological-knowledge indigenous-knowledge local-knowledge community-based-monitoring participatory-research action-research knowledge-co-production science-policy-interface science-society-interface boundary-organization knowledge-broker conservation-education nature-interpretation ecotourism sustainable-tourism responsible-travel green-travel low-impact-travel carbon-offset voluntary-carbon-market compliance-carbon-market REDD+reducing-emissions-from-deforestation-and-forest-degradation afforestation reforestation forest-restoration agroforestry silvopasture forest-landscape-restoration landscape-approach integrated-landscape-management watershed-management river-basin-management coastal-zone-management marine-spatial-planning ocean-governance blue-economy sustainable-blue-economy fisheries-management aquaculture sustainable-agriculture regenerative-agriculture organic-agriculture agroecology permaculture conservation-agriculture climate-smart-agriculture precision-agriculture digital-agriculture smart-farming vertical-farming controlled-environment-agriculture hydroponics aeroponics aquaponics algae-farming insect-farming entomophagy alternative-protein plant-based-diet reduced-meat-consumption food-system-transformation sustainable-food-system circular-food-system food-waste-reduction zero-hunger nutrition-security food-sovereignty seed-sovereignty farmers'-rights plant-variety-protection intellectual-property patent plant-breeder's-rights utility-model trademark geographical-indication certification organic-certification fair-trade-certification Rainforest-Alliance Marine-Stewardship-Council Forest-Stewardship-Council Roundtable-on-Sustainable-Palm-Oil Better-Cotton-Initiative GlobalGAP food-safety food-quality traceability accountability governance polycentric-governance adaptive-governance collaborative-governance network-governance multi-level-governance global-governance regional-governance national-governance subnational-governance local-governance community-governance customary-governance traditional-governance indigenous-governance self-governance autonomy decentralization devolution subsidiarity empowerment capacity-building institutional-strengthening policy-reform legal-reform regulatory-reform incentive disincentive economic-instrument market-based-instrument tax subsidy fee charge levy fine penalty liability compensation insurance risk-transfer catastrophe-bond green-bond blue-bond sustainability-linked-bond impact-investing blended-finance public-private-partnership concession lease permit license quota tradable-permit cap-and-trade carbon-tax carbon-pricing social-cost-of-carbon shadow-price internal-carbon-pricing carbon-border-adjustment border-carbon-adjustment climate-club just-transition-mechanism loss-and-damage climate-finance green-climate-fund adaptation-fund special-climate-change-fund global-environment-facility biodiversity-finance nature-finance natural-capital-finance impact-investment ESG environmental-social-and-governance sustainable-finance responsible-investment active-ownership engagement proxy-voting divestment fossil-fuel-divestment deforestation-free conversion-free no-net-loss net-gain biodiversity-net-gain mitigation-hierarchy avoidance minimization offset compensatory-mitigation in-lieu-fee conservation-banking wetland-mitigation-banking species-banking ecosystem-services-marketplace natural-capital-accounting system-of-environmental-economic-accounting inclusive-wealth genuine-progress-indicator well-being-economy degrowth post-growth steady-state-economy doughnut-economics planetary-boundaries safe-and-just-space social-foundation ecological-ceiling fair-share Earth-system Anthropocene Great-Acceleration Great-Transformation sustainability-transformation systemic-change structural-change paradigm-shift worldview-shift values-shift behavior-change lifestyle-change consumption-change production-change technology-change innovation disruptive-innovation transformative-innovation social-innovation institutional-innovation policy-innovation governance-innovation business-model-innovation financial-innovation technological-innovation digital-innovation green-technology clean-technology climate-technology ecosystem-based-disaster-risk-reduction blue-infrastructure hybrid-infrastructure natural-infrastructure building-with-nature working-with-nature engineering-with-nature ecological-engineering environmental-engineering conservation-ecology industrial-ecology agricultural-ecology forest-ecology grassland-ecology wetland-ecology freshwater-ecology marine-ecology coastal-ecology deep-ecology political-ecology social-ecology human-ecology cultural-ecology symbolic-ecology spiritual-ecology religious-ecology eco-theology creation-care stewardship-ethic land-ethic Earth-ethic planetary-ethic cosmic-ethic deep-ecology-platform ecofeminism environmental-justice climate-justice intergenerational-justice intragenerational-justice spatial-justice recognition-justice procedural-justice distributive-justice restorative-justice transformative-justice environmental-racism climate-racism slow-violence structural-violence environmental-conflict resource-conflict climate-conflict water-conflict land-conflict territorial-conflict conservation-conflict human-wildlife-conflict predator-control pest-control invasive-species-management disease-management wildlife-management habitat-management population-management harvest-management sustainable-use sustainable-harvesting adaptive-harvesting precautionary-approach ecosystem-approach ecosystem-based-management integrated-management holistic-management risk-based-management scenario-planning strategic-foresight transformative-governance adaptive-capacity transformative-capacity learning-organization learning-network knowledge-network community-of-practice epistemic-community knowledge-action-network future-earth conservation-science restoration-science ecological-economics environmental-economics natural-resource-economics resource-economics ecological-anthropology environmental-anthropology environmental-sociology environmental-psychology environmental-geography science-and-technology-studies science-policy-studies innovation-studies horizon-scanning trend-analysis weak-signal wild-card black-swan grey-rhino systemic-risk cascading-risk compound-risk complex-risk emerging-risk evolving-risk transboundary-risk planetary-risk existential-risk global-catastrophic-risk civilization-risk human-extinction-risk species-extinction-risk ecosystem-collapse-risk irreversible-change abrupt-change nonlinear-change threshold-effect path-dependence lock-in carbon-lock-in infrastructure-lock-in behavioral-lock-in institutional-lock-in technological-lock-in regulatory-lock-in financial-lock-in political-lock-in cultural-lock-in social-lock-in economic-lock-in structural-inertia resistance-to-change barrier-to-change enabler-of-change driver-of-change accelerator-of-change catalyst-of-change leverage-point intervention-point entry-point window-of-opportunity policy-window political-opportunity social-opportunity economic-opportunity technological-opportunity environmental-opportunity crisis-opportunity disaster-opportunity build-back-better green-recovery just-recovery resilient-recovery sustainable-recovery transformational-recovery disaster-risk-reduction prevention preparedness response rehabilitation reconstruction resilience-building adaptive-capacity-building transformative-capacity-building disaster-risk-management climate-risk-management environmental-risk-management financial-risk-management operational-risk-management strategic-risk-management enterprise-risk-management integrated-risk-management holistic-risk-management systemic-risk-management cascading-risk-management compound-risk-management complex-risk-management emerging-risk-management evolving-risk-management transboundary-risk-management planetary-risk-management existential-risk-management global-catastrophic-risk-management civilization-risk-management human-extinction-risk-management species-extinction-risk-management ecosystem-collapse-risk-management tipping-point-management threshold-management regime-shift-management alternative-stable-state-management hysteresis-management path-dependence-management lock-in-management carbon-lock-in-management infrastructure-lock-in-management behavioral-lock-in-management institutional-lock-in-management technological-lock-in-management regulatory-lock-in-management financial-lock-in-management political-lock-in-management cultural-lock-in-management social-lock-in-management economic-lock-in-management structural-inertia-management resistance-to-change-management barrier-to-change-management enabler-of-change-management driver-of-change-management accelerator-of-change-management catalyst-of-change-management leverage-point-management intervention-point-management entry-point-management window-of-opportunity-management policy-window-management political-opportunity-management social-opportunity-management economic-opportunity-management technological-opportunity-management environmental-opportunity-management crisis-opportunity-management disaster-opportunity-management build-back-better-management green-recovery-management just-recovery-management resilient-recovery-management sustainable-recovery-management transformational-recovery-management disaster-risk-reduction-management prevention-management preparedness-management response-management recovery-management rehabilitation-management reconstruction-management resilience-building-management adaptive-capacity-building-management transformative-capacity-building-management disaster-risk-management-management climate-risk-management-management environmental-risk-management-management financial-risk-management-management operational-risk-management-management strategic-risk-management-management enterprise-risk-management-management integrated-risk-management-management holistic-risk-management-management systemic-risk-management-management cascading-risk-management-management compound-risk-management-management complex-risk-management-management emerging-risk-management-management evolving-risk-management-management transboundary-risk-management-management planetary-risk-management-management existential-risk-management-management global-catastrophic-risk-management-management civilization-risk-management-management human-extinction-risk-management-management species-extinction-risk-management-management ecosystem-collapse-risk-management-management tipping-point-management-management threshold-management-management regime-shift-management-management alternative-stable-state-management-management hysteresis-management-management path-dependence-management-management lock-in-management-management carbon-lock-in-management-management infrastructure-lock-in-management-management behavioral-lock-in-management-management institutional-lock-in-management-management technological-lock-in-management-management regulatory-lock-in-management-management financial-lock-in-management-management political-lock-in-management-management cultural-lock-in-management-management social-lock-in-management-management economic-lock-in-management-management structural-inertia-management-management resistance-to-change-management-management barrier-to-change-management-management enabler-of-change-management-management driver-of-change-management-management accelerator-of-change-management-management catalyst-of-change-management-management leverage-point-management-management intervention-point-management-management entry-point-management-management window-of-opportunity-management-management policy-window-management-management political-opportunity-management-management social-opportunity-management-management economic-opportunity-management-management technological-opportunity-management-management environmental-opportunity-management-management crisis-opportunity-management-management disaster-opportunity-management-management build-back-better-management-management green-recovery-management-management just-recovery-management-management resilient-recovery-management-management sustainable-recovery-management-management transformational-recovery-management-management disaster-risk-reduction-management-management prevention-management-management preparedness-management-management response-management-management recovery-management-management rehabilitation-management-management reconstruction-management-management resilience-building-management-management adaptive-capacity-building-management-management transformative-capacity-building-management-management disaster-risk-management-management-management climate-risk-management-management-management environmental-risk-management-management-management financial-risk-management-management-management operational-risk-management-management-management strategic-risk-management-management-management enterprise-risk-management-management-management integrated-risk-management-management-management holistic-risk-management-management-management systemic-risk-management-management-management cascading-risk-management-management-management compound-risk-management-management-management complex-risk-management-management-management emerging-risk-management-management-management evolving-risk-management-management-management transboundary-risk-management-management-management planetary-risk-management-management-management existential-risk-management-management-management global-catastrophic-risk-management-management-management civilization-risk-management-management-management human-extinction-risk-management-management-management species-extinction-risk-management-management-management ecosystem-collapse-risk-management-management-management tipping-point-management-management-management threshold-management-management-management regime-shift-management-management-management alternative-stable-state-management-management-management hysteresis-management-management-management path-dependence-management-management-management lock-in-management-management-management carbon-lock-in-management-management-management infrastructure-lock-in-management-management-management 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Entedoninae
Entedoninae is a subfamily of chalcid wasps in the family Eulophidae comprising over 90 genera. Members are primarily parasitoids with documented associations across diverse host groups including gall-inducing insects, leafminers, sawflies, thrips, and ants. The subfamily exhibits considerable morphological and ecological diversity, with species found in tropical to temperate habitats worldwide.
parasitoid biological-control gall-associated leafminer-parasitoid Chalcidoidea Hymenoptera Eulophidae Entedoninae tropical-forest agricultural-pest-control sawfly-parasitoid thrips-parasitoid ant-associated rice-pest Cecidomyiidae Tenthredinidae Pontania Closterocerus Ceranisus Afronympha Ametallon Chrysonotomyia Omphale-genus-group canopy understorey ecological-plasticity Bangladesh Panama Africa New-Zealand Europe Senegal South-Africa Democratic-Republic-of-the-Congo Uganda endoparasitoid larval-parasitoid egg-parasitoid host-specificity forest-habitat degraded-habitat mid-altitude-rainforest lowland-rainforest tropical-America classical-biological-control augmentative-biological-control integrated-pest-management rice-hispa Dicladispa-armigera gall-wasp sawfly-gall leafminer ant-symbiosis Formicidae Khaya-senegalensis Ochna-pulchra morphological-taxonomy molecular-taxonomy 28S-rDNA COI CytB new-species-description generic-revision host-range parasitoid-complex food-web ecosystem-service natural-enemy pest-management entomophagous hyperparasitoid inquiline curculionid Apion-minimum Phyllocolpa-leucosticta Chrysocharis Entedon Asecodes Pleurotroppopsis Afrotroppopsis Zaommomentedon Apleurotropis Kratoysma Platocharis Schizocharis Parahorismenus Atullya Tropicharis Chrysochuris cruy mirabilis oryzamyntor eminpashai carinatum dussiae longicaudata machaeriae unimaculata claviger gorgonaense auripunctata galbina laeviscuta phenacapsia cecvora elongata pontaniae heyeri risbeci milletiae newbyi Förster-1856 Chalcidoidea-phylogeny biodiversity species-richness cryptic-species molecular-diagnostics host-plant-association habitat-specificity canopy-fauna understorey-fauna forest-stratification tropical-biodiversity Afrotropical Neotropical Palaearctic Nearctic Oriental Australasian Pacific Holarctic biogeography taxonomic-revision annotated-key catalogue type-specimen museum-collection slide-mounted-specimens morphometrics genitalia wing-venation antennae clypeus mandibles malar-space scape pedicel flagellum gaster vertex suture color-pattern pale-femora dark-gaster sexual-dimorphism male female ovipositor emergence rearing quarantine field-collection foreign-exploration augmentative-release pesticide-resistance sustainable-agriculture food-security rice-production cereal-pest vegetable-pest fruit-pest ornamental-pest forest-pest urban-pest invasive-species native-range introduced-range establishment spread impact non-target-effects host-specificity risk-assessment environmental-safety regulatory-approval research-symposium proceedings peer-reviewed Zootaxa ZooKeys Insect-Systematics-&-Evolution Australian-Journal-of-Entomology Far-Eastern-Entomologist Russian-Entomological-Journal Belgian-Journal-of-Entomology Florida-Entomologist Pan-Pacific-Entomologist Entomological-News Proceedings-of-the-Hawaiian-Entomological-Society Transactions-of-the-American-Entomological-Society Journal-of-Natural-History Zoological-Journal-of-the-Linnean-Society BioControl Journal-of-Economic-Entomology Entomologia-Experimentalis-et-Applicata Environmental-Entomology Oecologia Cladistics Molecular-Phylogenetics-and-Evolution Heredity Canadian-Entomologist Proceedings-of-the-Entomological-Society-of-Washington Annals-of-the-Entomological-Society-of-America Bulletin-of-Entomological-Research Neotropical-Entomology Journal-of-Asia-Pacific-Entomology Turkish-Journal-of-Zoology Polish-Journal-of-Entomology North-Western-Journal-of-Zoology Entomologica-Americana Entomologische-Berichten Sahlbergia Dugesiana Acta-Zoologica-Lilloana Folia-Entomologica-Mexicana Revista-de-la-Sociedad-Entomologica-Argentina Boletin-de-la-Asociacion-Espanola-de-Entomologia Entomologicheskoe-Obozrenie Entomological-Review Zoological-Institute-RAS Proceedings-of-the-Zoological-Institute-of-the-Russian-Academy-of-Sciences Taiwan-Agricultural-Research-Institute Council-of-Agriculture University-of-California Riverside Entomology-Research-Museum California-Department-of-Food-and-Agriculture Pierce's-Disease-Control-Program biological-control-program integrated-pest-management-program research-progress-report annual-summary technical-report symposium-proceedings conference-abstract poster-presentation invited-paper monograph book-chapter identification-guide illustrated-key annotated-catalogue type-material voucher-specimen collection-curation taxonomy systematics phylogenetics molecular-systematics DNA-barcoding species-complex sibling-species species-boundaries geographic-variation population-genetics gene-flow reproductive-isolation host-race biotype strain laboratory-rearing insectary mass-production release monitoring impact-assessment efficacy economic-evaluation cost-benefit-analysis farmer-adoption extension education policy regulation international-cooperation capacity-building training workshop field-day demonstration on-farm-research participatory-research farmer-field-school sustainable-intensification agroecology conservation-biological-control habitat-management floral-resources refuge buffer-zone landscape-complexity connectivity spillover natural-enemy-diversity food-web-structure trophic-interactions multitrophic indirect-effects trait-mediated density-dependent inverse-density-dependent functional-response numerical-response searching-efficiency handling-time oviposition-behavior host-selection host-acceptance host-suitability host-quality host-stage-preference egg-load progeny-allocation sex-ratio local-mate-competition sib-mating inbreeding fitness longevity fecundity development-time thermal-requirements degree-days phenology synchrony asynchrony seasonal-adaptation diapause quiescence cold-hardiness heat-tolerance desiccation-resistance starvation-resistance dispersal flight walking foraging patch-use area-restricted-search oviposition-marking host-marking superparasitism multiparasitism intraguild-predation hyperparasitism autoparasitism heteronomous-hyperparasitism facultative-hyperparasitism obligate-hyperparasitism primary-parasitoid secondary-parasitoid tertiary-parasitoid mummy aphid-mummy scale-mummy pupal-parasitoid egg-larval-parasitoid larval-pupal-parasitoid idiobiont koinobiont ectoparasitoid solitary gregarious polyembryony thelytoky arrhenotoky diploid haplodiploid chromosome-number genome-size transcriptome proteome metabolome morphome phenome extended-phenotype niche-construction ecosystem-engineering keystone-species ecosystem-service-provider functional-group guild trophic-level energy-flow nutrient-cycling decomposition predation herbivory parasitism mutualism commensalism competition apparent-competition exploitative-competition interference-competition scramble-competition contest-competition resource-partitioning niche-differentiation character-displacement co-evolution arms-race Red-Queen geographic-mosaic local-adaptation phenotypic-plasticity genetic-adaptation epigenetics transgenerational-effects maternal-effects paternal-effects parental-care mating-system sexual-selection sperm-competition cryptic-female-choice mate-choice lek territory aggression alarm defense venom allomone kairomone synomone semiochemical pheromone trail-pheromone alarm-pheromone aggregation-pheromone sex-pheromone marking-pheromone oviposition-deterrent host-marking-pheromone plant-volatile induced-plant-defense systemic-acquired-resistance induced-systemic-resistance priming volatile-organic-compound green-leaf-volatile terpenoid phenylpropanoid jasmonic-acid salicylic-acid ethylene abscisic-acid gibberellin auxin cytokinin brassinosteroid strigolactone nitric-oxide reactive-oxygen-species hypersensitive-response programmed-cell-death autophagy apoptosis necrosis pathogen-associated-molecular-pattern damage-associated-molecular-pattern pattern-recognition-receptor effector-triggered-immunity gene-for-gene avirulence virulence effector toxin phytotoxin mycotoxin bacterial-toxin viral-toxin insecticidal-protein cry-protein vip-protein cyt-protein spore conidium appressorium haustorium mycelium sclerotium fruiting-body basidiocarp ascocarp perithecium cleistothecium pycnidium acervulus sporodochium symptom sign disease-triangle disease-cycle epidemic endemic pandemic outbreak eradication containment management control prevention forecasting warning-system decision-support-system expert-system model simulation economic-threshold economic-injury-level action-threshold treatment-threshold damage-threshold aesthetic-injury-level cultural-control physical-control mechanical-control chemical-control genetic-control sterile-insect-technique incompatible-insect-technique precision-agriculture site-specific-management variable-rate-application remote-sensing geographic-information-system global-positioning-system unmanned-aerial-vehicle drone sensor imaging spectroscopy machine-learning artificial-intelligence deep-learning neural-network computer-vision image-analysis pattern-recognition classification segmentation detection tracking counting sampling sampling-plan sequential-sampling binomial-sampling presence-absence-sampling fixed-precision-sampling variable-precision-sampling spatial-sampling temporal-sampling stratified-sampling cluster-sampling systematic-sampling random-sampling convenience-sampling judgment-sampling expert-sampling quota-sampling snowball-sampling respondent-driven-sampling capture-recapture mark-recapture mark-resight distance-sampling line-transect point-transect strip-transect plot quadrat pitfall-trap Malaise-trap pan-trap water-pan yellow-pan blue-pan white-pan flight-interception-trap window-trap canopy-fogging beating-tray sweep-net aerial-net ground-net aquatic-net D-vac vacuum-sampler aspirator mouth-aspirator mechanical-aspirator berlese-funnel Tullgren-funnel Winkler-extractor pitfall barber-trap pitfall-array drift-fence funnel-trap light-trap UV-light black-light mercury-vapor sodium-vapor LED actinic color-trap sticky-trap pheromone-trap synthetic-pheromone natural-pheromone kairomone-trap food-bait protein-bait sugar-bait fermentation-bait alcohol-bait vinegar-bait molasses-bait fruit-bait meat-bait fish-bait dung-bait carrion-bait decoy lure attractant repellent deterrent antifeedant oviposition-stimulant arrestant suppressant toxicant stomach-poison contact-poison fumigant systemic translaminar residual persistent non-persistent broad-spectrum narrow-spectrum selective non-selective target-specific host-specific mode-of-action nerve-poison acetylcholinesterase-inhibitor GABA-antagonist sodium-channel-modulator nicotinic-acetylcholine-receptor-agonist ryanodine-receptor-modulator chitin-synthesis-inhibitor juvenile-hormone-mimic ecdysone-agonist microbial-pesticide entomopathogenic-nematode entomopathogenic-fungus entomopathogenic-bacterium entomopathogenic-virus entomopathogenic-protozoa Bacillus-thuringiensis Bt Bacillus-sphaericus Paenibacillus-popilliae Serratia-entomophila Metarhizium-anisopliae Beauveria-bassiana Lecanicillium-lecanii Isaria-fumosorosea Nomuraea-rileyi Hirsutella-thompsonii Entomophthora Neozygites Nosema Vairimorpha Thelohania Anticarsia-gemmatalis-nucleopolyhedrovirus AgMNPV Cydia-pomonella-granulovirus CpGV Spodoptera-exigua-multiple-nucleopolyhedrovirus SeMNPV Autographa-californica-multiple-nucleopolyhedrovirus AcMNPV nucleopolyhedrovirus granulovirus cypovirus entomopoxvirus iridovirus parvovirus tetravirus nodavirus reovirus picornavirus iflavirus densovirus birnavirus totivirus partitivirus hypovirulence mycovirus bacteriophage phage transduction conjugation transformation horizontal-gene-transfer vertical-gene-transfer mobile-genetic-element transposon plasmid integron genomic-island pathogenicity-island symbiosis-island metabolic-island resistance-island virulence-factor antibiotic-resistance insecticide-resistance acaricide-resistance herbicide-resistance fungicide-resistance bactericide-resistance nematicide-resistance molluscicide-resistance rodenticide-resistance resistance-management resistance-monitoring resistance-detection resistance-diagnosis resistance-mechanism metabolic-resistance target-site-resistance penetration-resistance behavioral-resistance cross-resistance multiple-resistance negative-cross-resistance synergism antagonism potentiation safener adjuvant surfactant spreader sticker extender buffer acidifier defoamer drift-retardant rainfastness UV-protectant formulation emulsifiable-concentrate wettable-powder water-dispersible-granule suspension-concentrate capsule-suspension microencapsulation nanotechnology controlled-release slow-release attract-and-kill sterile-insect-release inundative-release inoculative-release conservation augmentation classical neoclassical new-association targeted area-wide suppression protection production integrated-crop-management integrated-production organic-farming biodynamic-farming permaculture agroforestry silvopasture climate-smart-agriculture conservation-agriculture digital-agriculture smart-farming agriculture-4.0 internet-of-things big-data cloud-computing blockchain distributed-ledger traceability transparency certification standard governance stakeholder value-chain supply-chain food-system nutrition-security livelihood income employment gender youth social-inclusion equity fairness justice ethics animal-welfare environmental-protection biodiversity-conservation ecosystem-restoration land-degradation-neutrality desertification deforestation reforestation afforestation forest-restoration wetland-restoration grassland-restoration soil-restoration soil-health soil-organic-matter soil-carbon soil-biodiversity soil-fauna soil-microbiome rhizosphere phyllosphere anthroposphere technosphere biosphere ecosphere hydrosphere atmosphere lithosphere pedosphere cryosphere climate-change global-warming greenhouse-gas carbon-dioxide methane nitrous-oxide fluorinated-gas carbon-sequestration carbon-storage carbon-sink carbon-source carbon-neutral carbon-negative net-zero climate-adaptation climate-mitigation climate-resilience vulnerability exposure sensitivity adaptive-capacity transition pathway scenario projection forecast prediction uncertainty risk hazard damage loss adaptation mitigation resilience sustainability sustainable-development sustainable-development-goals SDG Agenda-2030 Paris-Agreement Convention-on-Biological-Diversity CBD Nagoya-Protocol Cartagena-Protocol Aichi-Targets Kunming-Montreal-Global-Biodiversity-Framework post-2020-biodiversity-framework One-Health Planetary-Health EcoHealth One-Welfare One-World One-Future interdisciplinary transdisciplinary multidisciplinary cross-disciplinary intersectoral multistakeholder partnership collaboration cooperation network platform hub center-of-excellence capacity-development institutional-strengthening policy-support science-policy-interface evidence-based-policy science-diplomacy technology-transfer knowledge-transfer knowledge-exchange knowledge-sharing open-access open-data open-science open-innovation citizen-science community-based-monitoring action-research transformative-research solution-oriented-research problem-driven-research impact-oriented-research implementation-research operational-research applied-research basic-research fundamental-research strategic-research tactical-research adaptive-research innovation invention discovery breakthrough frontier emerging converging enabling disruptive transformative exponential digital biological physical cognitive social institutional technological ecological evolutionary revolutionary incremental radical systematic holistic integrated comprehensive inclusive participatory bottom-up top-down middle-out hybrid mixed-methods qualitative quantitative empirical theoretical conceptual analytical computational experimental observational descriptive correlational causal mechanistic predictive prescriptive normative evaluative assessment appraisal review audit evaluation outcome-evaluation process-evaluation cost-effectiveness cost-benefit cost-utility multi-criteria-decision-analysis stakeholder-analysis social-network-analysis institutional-analysis political-economy-analysis gender-analysis youth-analysis vulnerability-analysis risk-analysis uncertainty-analysis sensitivity-analysis scenario-analysis systems-analysis life-cycle-assessment footprint water-footprint carbon-footprint ecological-footprint nitrogen-footprint biodiversity-footprint land-footprint material-footprint social-footprint health-footprint planetary-boundary safe-operating-space doughnut-economics regenerative restorative circular green blue bioeconomy circular-economy sharing-economy gig-economy platform-economy digital-economy knowledge-economy creative-economy care-economy wellbeing-economy degrowth post-growth steady-state doughnut sufficiency prosperity flourishing thriving buen-vivir ubuntu harmony balance equilibrium homeostasis robustness adaptability flexibility 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Exenterus amictorius
Exenterus amictorius is an ichneumonid parasitoid wasp introduced to North America that attacks sawfly larvae (Hymenoptera: Diprionidae), particularly during the pre-spinning eonymph stage. The species exhibits adaptive host discrimination behavior that changes during the host's spinning period, lacks discrimination initially but rapidly acquires and maintains this ability. It shows strong positive density-dependent responses to host abundance and has established successfully on multiple diprionid hosts across North America. In multiparasitism situations with the native E. diprionis, E. amictorius consistently survives due to faster larval development.
parasitoid biological-control sawfly-parasite introduced-species forest-entomology Ichneumonidae Diprionidae host-discrimination density-dependent-response multiparasitism competitive-displacement jack-pine pine-sawfly Quebec Ontario Europe North-America koinobiont idiobiont oviposition-behavior adaptive-behavior learning egg-complement Morisita's-Index power-function sex-ratio-bias seasonal-emergence invasive-species-success population-regulation host-switching establishment spread dominance competition exclusion phenology temporal-partitioning spatial-distribution aggregation functional-response numerical-response discrimination-learning associative-learning reproductive-strategy life-history development-rate survival interspecific-competition community-ecology species-interactions trophic-cascade ecosystem-service natural-enemy pest-management integrated-pest-management silviculture forest-health coniferous-forest boreal-forest temperate-forest pine-forest habitat-specificity host-specificity host-range-expansion biogeography colonization invasion-biology establishment-success population-dynamics density-dependence spatial-heterogeneity temporal-heterogeneity behavioral-plasticity phenotypic-plasticity memory decision-making foraging-behavior oviposition-strategy reproductive-allocation sex-allocation local-mate-competition fitness development emergence overwintering diapause voltinism univoltine partially-bivoltine seasonality synchrony asynchrony host-parasitoid-interactions biological-control-efficacy non-target-effects environmental-risk beneficial-insect conservation-biological-control augmentative-biological-control classical-biological-control introduced-natural-enemy exotic-species non-native-species invasive-species range-expansion distribution occurrence abundance population-density sampling monitoring survey inventory phylogeography molecular-ecology systematics taxonomy identification morphology anatomy behavior ecology reproduction ecosystem-ecology landscape-ecology conservation-ecology restoration-ecology applied-ecology forest-ecology insect-ecology parasitoid-ecology behavioral-ecology evolutionary-ecology functional-ecology trait-based-ecology mechanistic-ecology predictive-ecology quantitative-ecology statistical-ecology mathematical-ecology theoretical-ecology computational-ecology modeling simulation forecasting scenario-analysis risk-assessment decision-support policy management governance stakeholder-engagement knowledge-exchange science-communication education outreach citizen-science bioblitz iNaturalist GBIF biodiversity-informatics data-sharing open-science reproducible-research transparency accountability integrity ethics responsible-conduct sustainability resilience adaptation mitigation climate-change global-change anthropocene biodiversity-crisis extinction conservation restoration rewilding ecosystem-management adaptive-management evidence-based-management precision-conservation smart-conservation digital-conservation technological-innovation artificial-intelligence machine-learning deep-learning computer-vision remote-sensing satellite-imagery drone unmanned-aerial-vehicle sensor-network internet-of-things big-data data-science analytics visualization communication storytelling narrative framing engagement impact outcome evaluation assessment reporting verification certification standard protocol guideline best-practice lesson-learned case-study success-story failure challenge opportunity future-direction research-priority knowledge-gap hypothesis theory conceptual-framework methodological-approach study-design experimental-design sampling-design statistical-analysis inference uncertainty confidence robustness sensitivity validation calibration benchmarking comparison meta-analysis systematic-review evidence-synthesis knowledge-integration interdisciplinarity transdisciplinarity collaboration partnership network community capacity-building training mentoring career-development diversity inclusion equity justice decolonization indigenous-knowledge traditional-ecological-knowledge local-knowledge citizen-knowledge expert-knowledge stakeholder-knowledge boundary-organization knowledge-broker science-policy-interface evidence-based-policy policy-relevant-research actionable-science usable-science translational-ecology implementation-science impact-pathway theory-of-change logic-model results-framework monitoring-and-evaluation adaptive-learning reflective-practice continuous-improvement quality-assurance quality-control risk-management safeguard compliance institution organization culture leadership innovation entrepreneurship sustainability-science conservation-science restoration-science ecological-science environmental-science forest-science agricultural-science natural-resource-science life-science biological-science physical-science earth-science social-science humanities arts design engineering technology mathematics interdisciplinary multidisciplinary cross-disciplinary transdisciplinary holistic integrative synthetic analytical critical creative innovative transformative sustainable resilient adaptive responsive responsible ethical equitable inclusive diverse participatory collaborative cooperative collective shared open transparent accountable evidence-based science-based knowledge-based learning-based practice-based action-based outcome-based impact-based value-based principle-based rights-based needs-based stakeholder-based community-based place-based ecosystem-based nature-based solution-based problem-based challenge-based opportunity-based asset-based strength-based resilience-based adaptation-based mitigation-based restoration-based conservation-based sustainable-based regenerative circular blue green natural organic biodynamic permaculture agroecology agroforestry urban-forestry community-forestry social-forestry farm-forestry family-forestry industrial-forestry sustainable-forestry responsible-forestry certified-forestry legal-forestry illegal-logging deforestation forest-degradation forest-restoration forest-rehabilitation forest-conservation forest-protection forest-management forest-governance forest-policy forest-law forest-economics forest-sociology forest-anthropology forest-history forest-culture forest-spirituality forest-therapy forest-bathing shinrin-yoku forest-recreation forest-tourism forest-education forest-research forest-monitoring forest-inventory forest-assessment forest-valuation forest-accounting forest-finance forest-investment forest-market forest-trade forest-certification forest-standard forest-label forest-claim forest-footprint forest-risk forest-opportunity forest-benefit forest-service forest-ecosystem-service forest-provisioning-service forest-regulating-service forest-cultural-service forest-supporting-service forest-biodiversity forest-carbon forest-water forest-soil forest-air forest-climate forest-fire forest-pest forest-disease forest-invasive-species forest-disturbance forest-resilience forest-adaptation forest-mitigation forest-sustainability forest-transformation forest-transition forest-landscape-restoration forest-landscape-approach forest-mosaic forest-fragmentation forest-connectivity forest-corridor forest-network forest-patch forest-matrix forest-edge forest-interior forest-core forest-buffer forest-reserve forest-protected-area forest-conservation-area forest-management-area forest-production-area forest-restoration-area forest-research-area forest-education-area forest-recreation-area forest-sacred-area forest-cultural-area forest-heritage-area forest-community-area forest-indigenous-area forest-tenure forest-right forest-access forest-use forest-benefit-sharing forest-conflict forest-cooperation forest-partnership forest-agreement forest-convention forest-protocol forest-treaty forest-declaration forest-commitment forest-target forest-goal forest-objective forest-indicator forest-benchmark forest-baseline forest-scenario forest-projection forest-forecast forest-outlook forest-vision forest-strategy forest-plan forest-program forest-project forest-activity forest-intervention forest-operation forest-practice forest-technique forest-technology forest-innovation forest-knowledge forest-learning forest-capacity forest-institution forest-organization forest-platform forest-forum forest-dialogue forest-negotiation forest-mediation forest-arbitration forest-litigation forest-enforcement forest-compliance forest-reporting forest-verification forest-audit forest-evaluation forest-review forest-appraisal forest-certification-audit forest-chain-of-custody forest-traceability forest-transparency forest-accountability forest-integrity forest-governance-assessment forest-governance-indicator forest-governance-framework forest-governance-reform forest-governance-innovation forest-governance-learning forest-governance-capacity forest-governance-partnership forest-governance-platform forest-governance-network forest-governance-dialogue forest-governance-negotiation forest-governance-agreement forest-governance-commitment forest-governance-implementation forest-governance-impact forest-governance-outcome forest-governance-effectiveness forest-governance-efficiency forest-governance-equity forest-governance-sustainability forest-governance-resilience forest-governance-adaptation forest-governance-transformation forest-governance-transition forest-governance-future forest-governance-challenge forest-governance-opportunity forest-governance-priority forest-governance-research forest-governance-education forest-governance-communication forest-governance-outreach forest-governance-engagement forest-governance-participation forest-governance-collaboration forest-governance-cooperation forest-governance-coordination forest-governance-integration forest-governance-harmonization forest-governance-alignment forest-governance-convergence forest-governance-divergence forest-governance-complexity forest-governance-uncertainty forest-governance-risk forest-governance-creativity forest-governance-leadership forest-governance-entrepreneurship forest-governance-empowerment forest-governance-ownership forest-governance-responsibility forest-governance-stewardship forest-governance-custodianship forest-governance-trusteeship forest-governance-guardianship forest-governance-care forest-governance-respect forest-governance-reciprocity forest-governance-relationship forest-governance-connection forest-governance-belonging forest-governance-identity forest-governance-culture forest-governance-values forest-governance-ethics forest-governance-principles forest-governance-standards forest-governance-norms forest-governance-rules forest-governance-regulations forest-governance-laws forest-governance-policies forest-governance-strategies forest-governance-plans forest-governance-programs forest-governance-projects forest-governance-activities 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forest-governance-verifications forest-governance-audits forest-governance-evaluations forest-governance-reviews forest-governance-assessments forest-governance-appraisals forest-governance-certifications forest-governance-chains-of-custody forest-governance-traceabilities forest-governance-transparencies forest-governance-accountabilities forest-governance-integrities forest-governance-indicators forest-governance-reforms forest-governance-innovations forest-governance-learnings forest-governance-capacities forest-governance-partnerships forest-governance-agreements forest-governance-commitments forest-governance-implementations forest-governance-impacts forest-governance-outcomes forest-governance-effectivenesses forest-governance-efficiencies forest-governance-equities forest-governance-sustainabilities forest-governance-resiliences forest-governance-adaptations forest-governance-transformations forest-governance-transitions forest-governance-futures forest-governance-challenges forest-governance-opportunities forest-governance-priorities forest-governance-researches forest-governance-educations forest-governance-communications forest-governance-outreaches forest-governance-engagements forest-governance-participations forest-governance-collaborations forest-governance-cooperations forest-governance-coordinations forest-governance-integrations forest-governance-harmonizations forest-governance-alignments forest-governance-convergences forest-governance-divergences forest-governance-complexities forest-governance-uncertainties forest-governance-risks forest-governanceresiliences forest-governance-creativities forest-governance-leaderships forest-governance-entrepreneurships forest-governance-empowerments forest-governance-ownerships forest-governance-responsibilities forest-governance-stewardships forest-governance-custodianships forest-governance-trusteeships forest-governance-guardianships forest-governance-cares forest-governance-respects forest-governance-reciprocities forest-governance-relationships forest-governance-connections forest-governance-belongings forest-governance-identities forest-governance-cultures
Lacunicambarus diogenes
devil crayfish, devil crawfish
Lacunicambarus diogenes, commonly known as the devil crayfish or devil crawfish, is a primary burrowing crayfish native to eastern North America. This species constructs and inhabits burrows in wet, muddy terrestrial habitats rather than living in permanent surface water. Its burrowing activities create refugia used by numerous other species, including documented use by eastern cicada killer wasps (Sphecius speciosus) as brooding habitat. The species ranges across the Atlantic Coastal Plain and Piedmont ecoregion from New Jersey to Georgia, with disjunct populations in Louisiana.
crayfish burrowing ecosystem-engineer primary-burrower Atlantic-Coastal-Plain terrestrial fossorial North-America Cambaridae Decapoda crustacean endemic wetland floodplain refugia allogenic-engineer cave-crayfish Lacunicambarus Cambarus Girard-1852 devil-crayfish devil-crawfish Sphecius-speciosus cicada-killer-wasp brooding-habitat soil-type congeneric-cues floodplain-connectivity muddy-substrate desiccation-refuge predation-refuge intermittent-water Piedmont New-Jersey Pennsylvania Delaware Maryland Virginia North-Carolina South-Carolina Georgia Louisiana roadside-ditch terrestrial-habitat burrow-construction burrow-maintenance wetland-engineer crustacean-ecology decapod astacidea freshwater-invertebrate invertebrate-engineer habitat-modification species-interactions commensalism facilitation ecosystem-services biodiversity-support habitat-provision microhabitat-creation soil-bioturbation sediment-mixing water-infiltration groundwater-connection ephemeral-water seasonal-wetland palustrine lotic lentic riparian hyporheic bioturbation soil-structure nutrient-cycling detritivore omnivore scavenger predator prey trophic-interaction food-web community-ecology population-ecology conservation habitat-loss urbanization agriculture land-use-change climate-change drought hydrology geomorphology sedimentology ichnology trace-fossil burrow-architecture burrow-morphology tunnel-system chimney mud-pellet cast excavation sediment-ejection maintenance-behavior occupancy site-fidelity home-range territoriality intraspecific-interaction interspecific-interaction competition predation-risk desiccation-risk physiological-stress behavioral-adaptation morphological-adaptation fossorial-adaptation gill-function air-breathing cutaneous-respiration ion-regulation osmoregulation excretion metabolism energetics life-history growth molting reproduction mating egg-care juvenile-development recruitment population-dynamics demography genetics phylogeography biogeography dispersal colonization extinction rare-species common-species indicator-species keystone-species umbrella-species flagship-species natural-capital environmental-economics environmental-policy environmental-management restoration-ecology habitat-restoration species-reintroduction ex-situ-conservation in-situ-conservation protected-area wildlife-refuge nature-reserve national-park state-park conservation-easement stewardship citizen-science iNaturalist observation specimen voucher museum-collection type-specimen holotype taxonomy systematics nomenclature synonymy homonymy priority valid-name accepted-name species-concept morphological-species biological-species evolutionary-species phylogenetic-species integrative-taxonomy DNA-barcoding molecular-systematics phylogenetics phylogeny evolution adaptation natural-selection speciation diversification radiation endemism vicariance dispersal-limitation habitat-specificity niche-breadth niche-conservatism ecological-release character-displacement competitive-exclusion resource-partitioning habitat-filtering environmental-gradient elevation latitude longitude climate temperature precipitation hydroperiod soil-texture soil-moisture soil-chemistry pH organic-matter nutrient-availability productivity disturbance succession metacommunity landscape-ecology spatial-ecology movement-ecology dispersal-kernel connectivity corridor barrier fragmentation matrix edge-effect core-area patch-dynamics source-sink metapopulation population-viability minimum-viable-population effective-population-size inbreeding genetic-drift gene-flow local-adaptation phenotypic-plasticity genotype-environment-interaction epigenetics developmental-biology embryology larval-development post-larval-development juvenile-stage adult-stage senescence longevity survivorship fecundity reproductive-output reproductive-success fitness selection-coefficient heritability quantitative-genetics population-genetics molecular-ecology conservation-genetics genomic-resources transcriptomics proteomics metabolomics phenomics bioinformatics data-science machine-learning artificial-intelligence remote-sensing geographic-information-system spatial-analysis statistical-modeling experimental-design sampling-design field-methods laboratory-methods specimen-preparation curation data-management data-sharing open-science reproducibility replicability transparency ethics animal-welfare animal-use permit regulation policy legislation environmental-law natural-resource-law water-law land-use-planning environmental-impact-assessment cumulative-impact-assessment strategic-environmental-assessment adaptive-management monitoring evaluation learning stakeholder-engagement public-participation environmental-education science-communication outreach extension capacity-building professional-development career-development mentorship collaboration interdisciplinary transdisciplinary team-science open-innovation knowledge-exchange technology-transfer societal-impact sustainability sustainable-development green-economy blue-economy circular-economy nature-based-solution ecosystem-based-management integrated-water-resource-management watershed-management river-basin-management flood-risk-management drought-management climate-change-adaptation climate-change-mitigation resilience vulnerability risk-assessment scenario-planning forecasting prediction early-warning decision-support evidence-based-policy science-policy-interface boundary-organization knowledge-broker research-network professional-society academic-society scientific-journal peer-review publication citation impact-factor altmetrics research-evaluation research-funding grant fellowship scholarship award recognition career-award early-career mid-career senior-career emeritus retirement legacy historical-ecology paleoecology archaeology anthropology indigenous-knowledge traditional-ecological-knowledge local-knowledge community-science participatory-research action-research transformational-research transformative-change system-change paradigm-shift future-thinking foresight horizon-scanning trend-analysis emerging-issue research-priority grand-challenge societal-challenge sustainable-development-goal biodiversity-target conservation-target protected-area-target restoration-target climate-target emission-reduction carbon-sequestration blue-carbon green-carbon nature-conservation wildlife-conservation species-conservation habitat-conservation ecosystem-conservation landscape-conservation seascape-conservation connectivity-conservation corridor-conservation transboundary-conservation international-cooperation global-governance environmental-governance multilateral-environmental-agreement convention treaty protocol Cartagena-Protocol Nagoya-Protocol Aichi-Target Kunming-Montreal-Global-Biodiversity-Framework 30x30 half-earth nature-needs-half rewilding reintroduction de-extinction synthetic-biology gene-drive biotechnology nanotechnology geoengineering solar-radiation-management carbon-capture direct-air-capture bioenergy-with-carbon-capture negative-emission climate-intervention earth-system planetary-boundary safe-operating-space tipping-point hysteresis irreversibility committed-change sea-level-rise ocean-acidification deoxygenation warming extreme-event compound-risk cascading-risk systemic-risk existential-risk global-catastrophic-risk risk-management disaster-risk-reduction emergency-response humanitarian-action relief recovery reconstruction build-back-better resilient-infrastructure nature-based-infrastructure green-infrastructure sponge-city cooling-city urban-heat-island urban-ecology urban-biodiversity urban-conservation urban-planning urban-design architecture landscape-architecture green-building sustainable-building energy-efficiency renewable-energy clean-energy energy-transition just-transition energy-justice climate-justice environmental-justice social-justice equity fairness inclusion diversity gender youth intergenerational-equity future-generation rights-of-nature personhood legal-standing guardianship care ethics-of-care environmental-ethics conservation-ethics animal-ethics bioethics research-ethics integrity responsible-conduct misconduct fraud plagiarism retraction correction erratum expression-of-concern data-fabrication data-falsification data-manipulation image-manipulation duplicate-publication salami-slicing guest-authorship gift-authorship ghost-authorship author-contribution credit attribution acknowledgment funding-disclosure conflict-of-interest competing-interest institutional-review animal-care biosafety biosecurity dual-use export-control sanction embargo boycott divestment activism advocacy campaign movement social-movement environmental-movement conservation-movement climate-movement youth-movement student-movement indigenous-movement feminist-movement labor-movement peace-movement health-movement food-movement agriculture-movement water-movement energy-movement transport-movement housing-movement urban-movement rural-movement local-movement global-movement network coalition alliance partnership cooperation solidarity mutual-aid community-organizing grassroots bottom-up top-down hybrid-governance polycentric-governance self-organization emergence complexity system-dynamics agent-based-model network-analysis social-network ecological-network mutualistic-network antagonistic-network multiplex-network multilayer-network temporal-network spatial-network network-metric centrality modularity nestedness connectance robustness stability persistence coexistence biodiversity-ecosystem-function invasibility invasiveness impact risk pathway vector introduction establishment spread eradication containment control management phytosanitary veterinary human-health One-Health EcoHealth Planetary-Health One-Welfare sustainability-science conservation-science restoration-science resilience-science adaptation-science transformation-science futures-studies backcasting projection simulation modeling numerical-model conceptual-model statistical-model mechanistic-model process-based-model individual-based-model system-dynamics-model machine-learning-model deep-learning neural-network random-forest support-vector-machine gradient-boosting ensemble-method model-selection model-validation model-calibration sensitivity-analysis uncertainty-analysis risk-analysis decision-analysis optimization multi-objective-optimization Pareto-frontier trade-off synergy co-benefit win-win lose-lose win-lose zero-sum positive-sum negative-sum game-theory cooperative-game non-cooperative-game Nash-equilibrium Pareto-optimality social-dilemma tragedy-of-the-commons prisoner's-dilemma collective-action free-rider public-good common-pool-resource institution rule norm custom law governance administration implementation enforcement compliance accountability participation inclusiveness responsiveness effectiveness efficiency legitimacy adaptability transformability innovation experimentation adaptive-governance evidence-based science-based knowledge-based information-based data-driven model-driven theory-driven hypothesis-driven question-driven problem-driven solution-driven impact-driven outcome-driven output-driven input-driven process-driven value-driven mission-driven vision-driven stakeholder-driven user-driven citizen-driven community-driven place-based ecosystem-based landscape-based seascape-based watershed-based river-basin-based bioregion-based ecoregion-based biome-based realm-based planetary-based global regional national subnational local municipal community household individual scale cross-scale multi-scale trans-scale upscaling downscaling scaling-out scaling-up scaling-deep mainstreaming integration maintenance durability permanence reversibility additionality leakage spillover externality market-failure government-failure institutional-failure system-failure surprise novelty critical-transition regime-shift alternative-stable-state basin-of-attraction engineering-resilience ecological-resilience socio-ecological-resilience evolutionary-resilience transformative-resilience specified-resilience general-resilience resistance transformation transition trajectory scenario narrative storyline vision future utopia dystopia protopia heterotopia archipelago mosaic patchwork collage assemblage bricolage montage palimpsest layer stratum sediment memory heritage tradition culture identity belonging home place space territory land soil water air fire element matter energy information code signal sign symbol meaning significance value worth price cost benefit utility preference choice decision behavior action practice habit ritual ceremony celebration festival performance art aesthetics beauty sublime wonder awe curiosity inquiry discovery exploration adventure expedition voyage journey pilgrimage quest odyssey epic saga story history paleontology geology climatology meteorology oceanography limnology ecology biology zoology botany microbiology development physiology anatomy morphology ethology sociobiology ecological-psychology environmental-psychology conservation-psychology science-education informal-education non-formal-education lifelong-learning adult-learning skill-building competency expertise mastery craft artisan maker creator innovator entrepreneur leader manager administrator policymaker practitioner researcher scholar academic scientist naturalist explorer observer recorder documenter storyteller communicator educator mentor coach facilitator mediator negotiator diplomat advocate activist campaigner organizer mobilizer networker connector bridge-builder boundary-spanner translator interpreter ambassador representative delegate spokesperson champion hero role-model inspiration motivation aspiration hope optimism pessimism realism idealism pragmatism practicality feasibility viability desirability acceptability credibility trust confidence reliability validity rigor quality excellence best-practice good-practice lesson-learned success-factor failure-factor risk-factor protective-factor determinant driver pressure state response DPSIR STEEP SWOT PESTLE scenario-matrix morphological-analysis Delphi-method expert-elicitation structured-expert-judgment citizen-deliberation participatory-modeling companion-modeling serious-game role-play virtual-reality augmented-reality mixed-reality immersive-experience sensory-experience embodied-experience affective-experience cognitive-experience social-experience cultural-experience spiritual-experience transcendent-experience transformative-experience learning-experience educational-experience research-experience professional-experience personal-experience lived-experience embodied-knowledge tacit-knowledge explicit-knowledge codified-knowledge procedural-knowledge declarative-knowledge conceptual-knowledge factual-knowledge theoretical-knowledge practical-knowledge applied-knowledge transdisciplinary-knowledge interdisciplinary-knowledge multidisciplinary-knowledge cross-disciplinary-knowledge disciplinary-knowledge subdisciplinary-knowledge specialist-knowledge generalist-knowledge holistic-knowledge systemic-knowledge reductionist-knowledge analytical-knowledge synthetic-knowledge integrative-knowledge creative-knowledge innovative-knowledge adaptive-knowledge resilient-knowledge sustainable-knowledge ethical-knowledge responsible-knowledge trustworthy-knowledge reliable-knowledge valid-knowledge sound-knowledge robust-knowledge antifragile-knowledge living-knowledge dynamic-knowledge evolving-knowledge emergent-knowledge generative-knowledge productive-knowledge reproductive-knowledge replicative-knowledge imitative-knowledge emulative-knowledge competitive-knowledge collaborative-knowledge cooperative-knowledge shared-knowledge common-knowledge public-knowledge open-knowledge free-knowledge accessible-knowledge inclusive-knowledge diverse-knowledge equitable-knowledge just-knowledge fair-knowledge democratic-knowledge participatory-knowledge citizen-knowledge community-knowledge traditional-knowledge ecological-knowledge place-based-knowledge experiential-knowledge intuitive-knowledge implicit-knowledge unconscious-knowledge subconscious-knowledge conscious-knowledge self-aware-knowledge reflexive-knowledge critical-knowledge emancipatory-knowledge transformative-knowledge transformational-knowledge revolutionary-knowledge evolutionary-knowledge developmental-knowledge growth-knowledge maturation-knowledge aging-knowledge senescence-knowledge death-knowledge extinction-knowledge persistence-knowledge resilience-knowledge recovery-knowledge restoration-knowledge renewal-knowledge regeneration-knowledge revitalization-knowledge rejuvenation-knowledge rebirth-knowledge renaissance-knowledge awakening-knowledge enlightenment-knowledge illumination-knowledge inspiration-knowledge aspiration-knowledge hope-knowledge optimism-knowledge pessimism-knowledge realism-knowledge idealism-knowledge pragmatism-knowledge practicality-knowledge feasibility-knowledge viability-knowledge desirability-knowledge acceptability-knowledge legitimacy-knowledge credibility-knowledge trust-knowledge confidence-knowledge reliability-knowledge validity-knowledge rigor-knowledge quality-knowledge excellence-knowledge best-practice-knowledge good-practice-knowledge lesson-learned-knowledge success-factor-knowledge failure-factor-knowledge risk-factor-knowledge protective-factor-knowledge determinant-knowledge driver-knowledge pressure-knowledge state-knowledge impact-knowledge response-knowledge DPSIR-knowledge STEEP-knowledge SWOT-knowledge PESTLE-knowledge scenario-matrix-knowledge morphological-analysis-knowledge Delphi-method-knowledge expert-elicitation-knowledge structured-expert-judgment-knowledge citizen-deliberation-knowledge participatory-modeling-knowledge companion-modeling-knowledge serious-game-knowledge role-play-knowledge simulation-knowledge virtual-reality-knowledge augmented-reality-knowledge mixed-reality-knowledge immersive-experience-knowledge sensory-experience-knowledge embodied-experience-knowledge affective-experience-knowledge cognitive-experience-knowledge social-experience-knowledge cultural-experience-knowledge spiritual-experience-knowledge transcendent-experience-knowledge transformative-experience-knowledge learning-experience-knowledge educational-experience-knowledge research-experience-knowledge professional-experience-knowledge personal-experience-knowledge lived-experience-knowledge
Leptoglossus
leaf-footed bugs
Leptoglossus is a genus of true bugs in the leaf-footed bug family Coreidae, tribe Anisoscelini. Species are characterized by leaflike dilations of the hind tibia, a diagnostic trait of the genus. The genus is distributed throughout the Americas, with some introduced populations in Europe and Asia. Several species are economically significant agricultural pests, notably L. occidentalis, which has become invasive in multiple continents.
Coreidae leaf-footed-bug agricultural-pest invasive-species symbiosis sexual-dimorphism conifer-pest nuisance-pest Burkholderia tachinid-parasitoid pheromone-communication building-invader misidentification Triatoma-look-alike gradual-metamorphosis seed-predator forest-pest ornamental-pest plumbing-damage public-health-confusion Chile Europe North-America South-America Asia Turkey almond-pest citrus-pest tomato-pest corn-pest overwintering-aggregation defensive-secretion pheromone-mediated-parasitism egg-parasitoid biological-control ecological-niche-modeling climate-suitability range-expansion introduction accidental-dispersal seed-orchard-pest germination-reduction empty-seed-formation cone-damage lodgepole-pine Douglas-fir western-conifer-seed-bug L.-occidentalis L.-zonatus L.-phyllopus L.-clypealis L.-australis L.-chilensis L.-oppositus male-combat femoral-weapon abdominal-gland pheromone-gland tibial-dilation leaf-like-hind-leg true-bug Hemiptera Heteroptera Pentatomomorpha Anisoscelini phytophagous plant-feeding stylet-feeding salivary-sheath overwintering diapause building-entry structural-pest nuisance-odor flight-sound droning-flight aggregation-behavior male-pheromone female-choice parasitoid-host biological-control-agent integrated-pest-management monitoring early-detection prevention invasion-biology alien-species non-native-pest global-spread climate-change niche-modeling suitable-habitat establishment-risk quarantine phytosanitary forest-health seed-production reforestation afforestation conifer-forestry pine-seed seed-viability economic-impact crop-loss yield-reduction kernel-damage fruit-damage nut-damage vector-of-disease Chagas-disease kissing-bug public-alarm misinformation education identification-tools health-system-burden pesticide-overuse urban-ecology domiciliary peridomiciliary Andean-region Patagonia Mediterranean temperate subtropical tropical agroecosystem natural-enemy predator parasite pathogen symbiont gut-microbiome soil-ingestion fitness-benefit reproductive-success sperm-morphology testicular-morphology accessory-gland pheromone-blend species-specific-odor cherry-scent vanilla-scent cinnamon-scent rose-scent olfactory-communication mate-location mate-recognition sexual-selection male-male-competition weapon-morphology allometry developmental-plasticity nymphal-instar egg-barrel egg-arrangement linear-oviposition leaf-surface plant-tissue stylet-penetration enzymatic-digestion fluid-feeding phloem-feeding xylem-feeding seed-feeding cone-feeding fruit-feeding kernel-feeding nut-feeding crop-feeding host-plant-range polyphagy oligophagy specialist generalist pest-status damage-threshold economic-injury-level management-strategy cultural-control physical-control chemical-control resistance tolerance host-plant-resistance monitoring-trap light-trap population-density distribution-map spread-rate colonization establishment population-dynamics life-table developmental-rate thermal-requirements degree-days phenology voltinism univoltine bivoltine multivoltine generation-time overwintering-site hibernaculum shelter-seeking cold-hardiness freeze-tolerance supercooling water-relations desiccation-resistance starvation-resistance dispersal-ability flight-capacity walking-behavior climbing-behavior aggregation-pheromone alarm-pheromone stink-gland metathoracic-gland dorsoabdominal-gland ventral-abdominal-gland sexual-gland morphological-defense chemical-defense mechanical-defense autotomy leg-loss predation-risk parasitism-risk parasitoid-risk pathogen-risk competition intraspecific interspecific resource-competition mating-competition sperm-competition cryptic-female-choice reproductive-isolation speciation phylogeny systematics taxonomy morphology anatomy histology ultrastructure spermatogenesis spermatozoa sperm-length nuclear-length cyst-production follicle-number testis-structure reproductive-anatomy genitalia mating-behavior copulation insemination oviposition egg-production fecundity fertility hatching-success nymphal-survival adult-longevity sex-ratio operational-sex-ratio population-sex-ratio effective-population-size genetic-diversity gene-flow population-structure phylogeography biogeography historical-biogeography vicariance dispersal range-shift range-contraction altitudinal-distribution latitudinal-distribution longitudinal-distribution island-distribution continental-distribution endemism cosmopolitanism introduced-range native-range source-population founder-population invasion-front lag-phase establishment-phase spread-phase equilibrium-phase impact-assessment risk-assessment horizon-scanning early-warning rapid-response eradication containment control management adaptation mitigation restoration conservation biodiversity ecosystem-service ecosystem-function food-web trophic-level primary-consumer herbivore fruit-predator plant-animal-interaction mutualism antagonism predation parasitism commensalism amensalism facilitation indirect-interaction trait-mediated-interaction density-mediated-interaction behavioral-ecology evolutionary-ecology functional-ecology physiological-ecology population-ecology community-ecology ecosystem-ecology landscape-ecology macroecology global-ecology applied-ecology agricultural-ecology forest-ecology conservation-biology invasion-ecology restoration-ecology pest-management conservation-biological-control augmentative-biological-control classical-biological-control parasitoid nematode fungus bacterium virus entomopathogen microbial-control sterile-insect-technique genetic-control mating-disruption attract-and-kill push-pull trap-crop border-crop refuge habitat-management cultural-practice crop-rotation tillage irrigation fertilization pruning harvest-timing sanitation residue-management weed-management cover-crop companion-planting intercropping agroforestry silviculture forest-management seed-orchard-management cone-collection seed-extraction seed-testing germination-testing seed-quality seed-vigor empty-seed insect-damage insect-injury feeding-scar puncture stylet-sheath salivary-flange symptom sign diagnosis sampling economic-threshold decision-support expert-system modeling simulation forecasting climate-model niche-model species-distribution-model habitat-suitability risk-mapping invasion-pathway introduction-vector trade transport tourism migration natural-dispersal human-mediated-dispersal accidental-introduction intentional-introduction release escape cultivation ornamental forestry agriculture horticulture urbanization globalization land-use-change habitat-fragmentation habitat-loss degradation pollution pesticide fertilizer nutrient-enrichment eutrophication acidification warming drought extreme-event disturbance succession recovery resilience stability variability uncertainty sustainable-development green-economy circular-economy bioeconomy ecosystem-approach one-health planetary-health environmental-health public-health veterinary-health plant-health food-security nutrition livelihood poverty inequality gender indigenous-knowledge traditional-knowledge local-knowledge citizen-science stakeholder-engagement policy governance regulation legislation international-cooperation capacity-building awareness communication outreach extension advisory-service research innovation technology-transfer knowledge-exchange networking partnership collaboration evaluation evidence-based-policy science-policy-interface diplomacy advocacy activism social-movement environmental-justice environmental-ethics environmental-philosophy environmental-humanities environmental-history environmental-literature environmental-art environmental-education environmental-communication science-communication risk-communication crisis-communication health-communication behavior-change pro-environmental-behavior sustainable-behavior consumption production waste recycling reuse reduction efficiency renewable clean green blue natural organic agroecology permaculture regenerative restorative healing transformative transdisciplinary interdisciplinary multidisciplinary cross-disciplinary disciplinary disciplinary-integration knowledge-integration system-thinking complexity emergence non-linearity feedback tipping-point regime-shift alternative-stable-state resilience-thinking adaptive-management adaptive-governance learning reflection participation inclusion equity justice fairness ethics values norms culture identity place sense-of-place attachment well-being quality-of-life happiness flourishing thriving sustainability sustainable development growth degrowth post-growth steady-state circular doughnut-economics planetary-boundaries safe-operating-space tipping-elements critical-transition early-warning-signal resilience-indicator sustainability-indicator benchmark target goal commitment pledge agreement treaty convention protocol framework strategy plan program project initiative campaign movement coalition alliance network platform hub center institute organization association society union federation confederation foundation trust fund bank facility mechanism instrument tool method approach technique procedure process system structure institution administration operation service delivery provision supply demand market economy finance investment funding financing resource capital asset liability income expenditure revenue cost benefit profit loss return yield outcome output input throughput productivity effectiveness performance quality standard criterion indicator metric measure assessment appraisal review audit verification validation certification accreditation recognition award prize honor distinction excellence best-practice lesson-learned experience expertise competence capacity capability skill knowledge information data evidence science discovery invention creation design testing piloting scaling mainstreaming uptake adoption diffusion dissemination publication reporting documentation archiving preservation rehabilitation reconstruction replication duplication backup security safety protection safeguarding insurance risk-management emergency-preparedness response relief humanitarian peace prosperity health employment housing infrastructure energy water food fishery aquaculture mining manufacturing industry commerce recreation heritage nature ecosystem environment climate atmosphere ocean freshwater land soil mineral biota flora fauna microorganism fungi plant animal vertebrate invertebrate arthropod insect bug Hemipteran Heteropteran pentatomomorphan coreoid coreid squash-bug boxelder-bug stink-bug shield-bug plant-bug mirid lygaeid seed-bug broad-headed-bug alydid reduviid assassin-bug triatomine bed-bug cimicid water-bug nepomorphan gerromorphan leptopodomorphan enicocephalomorphan dipsocoromorphan ceratocomboid schizopteroid peloridioid coleorrhynchan moss-bug archaeorrhynchan fulgoromorphan cicadomorphan membracoid treehopper leafhopper planthopper psyllid jumping-plant-louse whitefly aleyrodid scale-insect coccoid mealybug aphid adelgid phylloxeran sternorrhynchan thysanopteran thrips psocopteran barklouse booklouse phthirapteran louse sucking-louse chewing-louse mallophagan anopluran dermapteran earwig blattodean cockroach termite isopteran mantodean mantid phasmid stick-insect leaf-insect orthopteran grasshopper locust katydid cricket mole-cricket pygmy-mole-cricket camel-cricket cave-cricket weta ensiferan caeliferan gryllotalpid myrmecophilid tettigoniid gryllid acridid pamphagid pneumorid lentulid tristirid eumastacid proscopiid tridactylid tetrigid grouse-locust pygmy-grasshopper plecopteran stonefly embiopteran webspinner zorapteran angel-insect dictyopteran
Palaemon vulgaris
Marsh Grass Shrimp, Common American Prawn, Common Grass Shrimp, Marsh Shrimp
Palaemon vulgaris is a small, transparent caridean shrimp native to the western Atlantic Ocean. Adults reach less than 5 cm in length and display distinctive orange pigmentation on the eyestalks. The species has been studied for its specialized swimming mechanics, employing metachronal beating of pleopods and drag-reducing mechanisms during locomotion. It occupies coastal and estuarine habitats from Cape Cod Bay to the Gulf of Mexico.
crustacean shrimp western-Atlantic transparent swimming-mechanics metachronal-swimming drag-reduction estuarine Palaemonidae Caridea Decapoda Malacostraca Arthropoda Palaemon Palaemonetes-vulgaris Say-1818 Cape-Cod-Bay Gulf-of-Mexico eyestalk-pigmentation pleopod hydrodynamics appendage-flexibility particle-image-velocimetry locomotion marine-biology invertebrate-zoology aquatic-arthropod coastal-species marsh-shrimp grass-shrimp common-American-prawn common-grass-shrimp marsh-grass-shrimp small-crustacean translucent-body camouflage swimming-appendages pleopodal-swimming crustacean-locomotion biomechanics fluid-dynamics drag-reduction-mechanisms asymmetric-flexibility appendage-grouping return-stroke power-stroke wake-reduction thrust-generation profile-area-modulation phase-lag appendage-wave swimming-efficiency aquatic-maneuverability hydrodynamic-optimization caridean-shrimp palaemonid-shrimp marine-invertebrate estuarine-invertebrate Atlantic-coast North-American-shrimp western-Atlantic-shrimp coastal-marine-biology invertebrate-swimming crustacean-biomechanics decapod-locomotion malacostracan-swimming arthropod-hydrodynamics translucent-marine-animal small-decapod marine-transparency eyestalk-coloration orange-pigmentation eyestalk-morphology visual-system crustacean-vision transparent-animal crypsis camouflage-adaptation predator-avoidance visual-ecology marine-ecology coastal-ecology estuarine-ecology benthic-pelagic nektonic-shrimp planktonic-stages benthic-adult habitat-utilization ecosystem-component food-web prey-species foraging-behavior sensory-biology mechanoreception chemoreception aquatic-sensory-systems neurobiology motor-control swimming-coordination appendage-coordination rhythmic-movement neuromuscular-control biological-fluid-mechanics low-Reynolds-number-swimming viscous-swimming intermediate-Reynolds-number unsteady-flows vortex-dynamics wake-structure thrust-production propulsive-efficiency energy-economy locomotor-performance evolutionary-biomechanics functional-morphology comparative-physiology experimental-biology laboratory-studies field-observations natural-history taxonomic-history nomenclatural-change Palaemonetes-to-Palaemon genus-revision systematics phylogenetics molecular-data morphological-data integrated-taxonomy species-delimitation cryptic-species morphological-similarity genetic-differentiation population-structure biogeography dispersal larval-transport ocean-currents habitat-connectivity marine-biogeography provincialism endemism range-distribution latitudinal-gradient depth-distribution salinity-tolerance euryhaline stenohaline osmoregulation physiological-ecology environmental-tolerance temperature-tolerance thermal-biology seasonal-variation reproductive-seasonality breeding-biology mating-behavior reproductive-output fecundity egg-production larval-development developmental-biology life-history growth-rate longevity survivorship population-dynamics demography abundance density spatial-distribution temporal-distribution diel-patterns nocturnal-activity crepuscular-behavior foraging-ecology diet-composition trophic-level omnivory detritivory predation predator-prey-interactions anti-predator-behavior escape-responses startle-response autotomy defensive-behavior social-behavior aggregation schooling shoaling conspecific-attraction mating-systems sexual-selection mate-choice sperm-competition parental-investment care-of-young extended-parental-care reproductive-strategies iteroparity semelparity bet-hedging r-selection K-selection life-history-theory phenotypic-plasticity environmental-induction developmental-plasticity morphological-plasticity behavioral-plasticity physiological-plasticity acclimation adaptation local-adaptation genetic-adaptation evolutionary-adaptation natural-selection artificial-selection domestication conservation-biology population-viability threatened-species endangered-species habitat-loss pollution eutrophication hypoxia climate-change ocean-acidification warming sea-level-rise range-shifts phenological-shifts invasive-potential non-native-species introduced-species bioinvasion vector-biology ballast-water aquaculture fisheries bycatch stock-assessment management sustainable-use protected-areas marine-protected-areas estuarine-protected-areas habitat-restoration ecosystem-management adaptive-management precautionary-approach ecosystem-based-management integrated-management stakeholder-engagement public-education outreach science-communication citizen-science iNaturalist observation-networks biodiversity-informatics data-sharing open-access reproducible-research transparency metadata standards ontologies vocabularies taxonomic-databases GBIF Catalogue-of-Life NCBI-Taxonomy Wikidata Wikipedia scholarly-publishing peer-review journal-quality impact-factor citation-metrics research-evaluation academic-careers funding grants collaboration networks consortia international-cooperation capacity-building training education mentorship diversity inclusion equity decolonization indigenous-knowledge traditional-ecological-knowledge local-knowledge community-based-monitoring participatory-research action-research transdisciplinary interdisciplinary multidisciplinary disciplinary-boundaries paradigm-shifts scientific-progress cumulative-knowledge standing-on-shoulders-of-giants history-of-science philosophy-of-science epistemology methodology research-design sampling statistics modeling simulation prediction forecasting scenario-analysis risk-assessment uncertainty confidence-intervals error-propagation sensitivity-analysis validation verification benchmarking best-practices guidelines protocols standard-operating-procedures quality-control quality-assurance reliability reproducibility replicability open-science FAIR-principles findable accessible interoperable reusable data-management data-curation data-preservation long-term-archives digital-repositories institutional-repositories subject-repositories preprint-servers post-publication-peer-review open-peer-review registered-reports preregistration open-data open-code open-materials open-notebooks science-blogs social-media public-engagement science-policy evidence-based-policy science-advice science-diplomacy international-frameworks conventions treaties SDGs sustainable-development-goals biodiversity-targets Aichi-targets post-2020-framework Kunming-Montreal-framework IPBES IPCC CBD CITES CMS Ramsar World-Heritage UNESCO FAO WHO UNEP GEF funding-mechanisms financial-resources resource-mobilization capacity-development technology-transfer north-south-cooperation south-south-cooperation triangular-cooperation partnerships multi-stakeholder private-sector civil-society NGOs community-organizations indigenous-peoples local-communities women youth future-generations intergenerational-equity intragenerational-equity environmental-justice climate-justice biodiversity-justice recognition-justice procedural-justice distributive-justice restorative-justice transformative-justice just-transition green-economy blue-economy circular-economy nature-based-solutions ecosystem-services natural-capital green-infrastructure climate-adaptation climate-mitigation resilience vulnerability exposure hazard risk disaster-risk-reduction early-warning-systems preparedness response recovery reconstruction build-back-better loss-and-damage climate-finance green-climate-fund adaptation-fund biodiversity-finance financial-mechanisms innovative-financing blended-finance impact-investing ESG environmental-social-governance corporate-sustainability responsible-business stewardship custodianship care-ethics environmental-ethics animal-ethics speciesism anthropocentrism biocentrism ecocentrism deep-ecology social-ecology feminist-ecology political-ecology environmental-humanities environmental-history environmental-literature environmental-art environmental-education environmental-psychology environmental-sociology environmental-anthropology environmental-geography environmental-economics ecological-economics natural-resource-economics environmental-law international-environmental-law environmental-governance environmental-institutions environmental-regimes compliance enforcement monitoring reporting accountability effectiveness legitimacy fairness participation access-to-information access-to-justice human-rights right-to-environment right-to-health right-to-food right-to-water cultural-rights indigenous-rights free-prior-informed-consent self-determination territorial-rights resource-rights intellectual-property traditional-knowledge benefit-sharing access-and-benefit-sharing Nagoya-Protocol Cartagena-Protocol biosafety biosecurity biotechnology synthetic-biology gene-editing CRISPR genetic-resources genetic-diversity genomics transcriptomics proteomics metabolomics phenomics bioinformatics computational-biology systems-biology bioengineering biomimicry nature-inspired-design biomimetics bioinspiration biomimetic-materials smart-materials responsive-materials adaptive-materials self-healing-materials self-cleaning-surfaces drag-reduction-surfaces superhydrophobic-surfaces antifouling antibacterial antiviral functional-surfaces surface-engineering nanotechnology nanomaterials nanoparticles nanocomposites nanostructures bottom-up-assembly top-down-fabrication self-assembly directed-assembly template-synthesis lithography 3D-printing additive-manufacturing rapid-prototyping custom-fabrication digital-manufacturing Industry-4.0 smart-manufacturing automation robotics soft-robotics biohybrid-robotics swarm-robotics autonomous-systems machine-learning artificial-intelligence deep-learning neural-networks computer-vision pattern-recognition classification segmentation detection tracking optimization control decision-support expert-systems knowledge-graphs semantic-web linked-data taxonomies folksonomies metadata-standards data-interoperability APIs web-services cloud-computing edge-computing fog-computing distributed-computing parallel-computing high-performance-computing quantum-computing neuromorphic-computing biological-computing DNA-computing molecular-computing chemical-computing unconventional-computing future-computing emerging-technologies technology-assessment technology-foresight horizon-scanning scenario-planning strategic-planning roadmapping innovation-systems innovation-policy research-and-development science-and-technology STEM-education STEAM-education inquiry-based-learning project-based-learning problem-based-learning active-learning experiential-learning lifelong-learning continuing-education professional-development skills-training competency-building human-capital social-capital intellectual-capital knowledge-economy creative-economy cultural-economy experience-economy symbolic-economy attention-economy platform-economy gig-economy sharing-economy collaborative-consumption circular-consumption sustainable-consumption responsible-consumption ethical-consumption conscious-consumption minimalism simplicity voluntary-simplicity downshifting degrowth post-growth steady-state-economy wellbeing-economy doughnut-economics regenerative-economics restorative-economics healing-economics care-economics solidarity-economics commons commoning peer-production open-source free-software creative-commons copyleft public-domain open-government open-innovation open-design open-hardware open-manufacturing distributed-manufacturing local-manufacturing on-demand-manufacturing just-in-time-manufacturing lean-manufacturing agile-manufacturing flexible-manufacturing mass-customization personalization bespoke tailored made-to-measure custom-fit user-centered-design human-centered-design design-thinking service-design experience-design interaction-design interface-design user-experience user-interface usability accessibility inclusive-design universal-design design-for-all assistive-technology adaptive-technology rehabilitation-engineering prosthetics orthotics exoskeletons wearable-technology implantable-technology invasive-technology non-invasive-technology minimally-invasive precision-medicine personalized-medicine stratified-medicine pharmacogenomics theranostics companion-diagnostics targeted-therapy immunotherapy gene-therapy cell-therapy regenerative-medicine tissue-engineering organ-engineering bioprinting 3D-bioprinting organ-on-a-chip body-on-a-chip human-on-a-chip microphysiological-systems alternative-methods replacement-methods reduction-methods refinement-methods 3Rs animal-welfare laboratory-animal-science veterinary-science comparative-medicine translational-medicine bench-to-bedside bedside-to-bench reverse-translation clinical-research clinical-trials evidence-based-medicine precision-health digital-health mobile-health telehealth telemedicine remote-monitoring remote-sensing wearable-sensors implantable-sensors biosensors biomarkers digital-biomarkers real-world-evidence patient-reported-outcomes clinical-outcomes health-outcomes quality-of-life wellbeing flourishing human-development capability-approach functionings capabilities agency empowerment social-inclusion social-cohesion social-networks social-support social-determinants-of-health health-equity health-justice health-for-all universal-health-coverage primary-health-care public-health global-health planetary-health One-Health EcoHealth environmental-health occupational-health mental-health psychological-wellbeing emotional-wellbeing social-wellbeing spiritual-wellbeing holistic-health integrative-health complementary-medicine alternative-medicine traditional-medicine indigenous-healing folk-medicine ethnomedicine medical-anthropology medical-sociology medical-geography health-geography therapeutic-landscapes healing-places sacred-sites pilgrimage spiritual-ecology religion-and-ecology theology-of-ecology ecotheology creation-spirituality nature-spirituality earth-spirituality paganism neo-paganism Wicca druidry shamanism animism totemism nature-worship sacred-nature holy-nature divine-nature immanent-transcendence panentheism pantheism naturalistic-pantheism religious-naturalism spiritual-naturalism secular-spirituality humanistic-spirituality existential-spirituality phenomenological-spirituality embodied-spirituality embodied-cognition embodied-mind extended-mind embedded-cognition enactive-cognition 4E-cognition predictive-processing free-energy-principle active-inference embodied-prediction interoception proprioception exteroception multisensory-integration sensorimotor-contingencies perceptual-inference perceptual-learning perceptual-development neuroplasticity brain-development cognitive-development social-development emotional-development moral-development identity-development self-concept self-esteem self-efficacy autonomy competence relatedness intrinsic-motivation extrinsic-motivation self-regulation self-control executive-function working-memory attention inhibition cognitive-flexibility creativity innovation problem-solving critical-thinking analytical-thinking systems-thinking computational-thinking scientific-thinking mathematical-thinking statistical-thinking probabilistic-thinking logical-thinking dialectical-thinking dialogical-thinking reflective-thinking metacognition meta-learning learning-to-learn learning-strategies study-skills time-management stress-management coping-strategies resilience-building growth-mindset fixed-mindset entity-theory incremental-theory self-theories implicit-theories beliefs-about-intelligence attribution-theory achievement-motivation goal-orientation mastery-goals performance-goals approach-goals avoidance-goals self-handicapping impostor-phenomenon stereotype-threat stereotype-lift social-identity group-identity collective-identity national-identity cultural-identity ethnic-identity racial-identity gender-identity sexual-identity professional-identity organizational-identity brand-identity 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social-democracy constitutional-democracy electoral-democracy illiberal-democracy hybrid-regime authoritarianism totalitarianism fascism nazism communism socialism capitalism neoliberalism Keynesianism monetarism austrian-school chicago-school behavioral-economics experimental-economics institutional-economics evolutionary-economics complexity-economics agent-based-modeling system-dynamics network-analysis social-network-analysis organizational-network-analysis innovation-networks knowledge-networks collaboration-networks co-authorship-networks citation-networks patent-networks technological-networks ecological-networks food-webs trophic-networks interaction-networks mutualistic-networks antagonistic-networks host-parasite-networks pollination-networks seed-dispersal-networks mycorrhizal-networks social-ecological-networks socio-ecological-systems coupled-human-natural-systems complex-adaptive-systems adaptive-capacity transformability resilience-thinking panarchy adaptive-cycle revolt remember front-loop back-loop exploitation conservation release reorganization fast-variables slow-variables tipping-points critical-transitions regime-shifts alternative-stable-states hysteresis irreversibility path-dependence lock-in rigidity-traps poverty-traps social-traps tragedy-of-the-commons collective-action-problems free-rider-problems social-dilemmas prisoner's-dilemma public-goods common-pool-resources institutional-analysis institutional-design polycentric-governance multi-level-governance network-governance adaptive-governance collaborative-governance participatory-governance deliberative-governance experimentalist-governance reflexive-governance meta-governance steering rowing rowing-together rowing-separately rowing-not-at-all policy-instruments policy-mixes policy-integration policy-coherence policy-effectiveness policy-efficiency policy-equity policy-legitimacy policy-acceptability policy-feasibility policy-sustainability policy-durability policy-adaptability policy-flexibility policy-robustness policy-resilience policy-innovation policy-learning policy-diffusion policy-transfer policy-convergence policy-divergence policy-fragmentation policy-coordination policy-harmonization policy-standardization policy-implementation policy-enforcement policy-compliance policy-monitoring policy-evaluation policy-assessment policy-appraisal policy-analysis policy-research policy-advice policy-advocacy policy-communication policy-entrepreneurship policy-windows punctuated-equilibrium incrementalism rational-comprehensive mixed-scanning garbage-can-model multiple-streams advocacy-coalition-framework narrative-policy-framework institutional-analysis-and-development social-ecological-systems-framework sustainability-science transdisciplinary-research mode-1-science mode-2-science triple-helix quadruple-helix quintuple-helix science-society-policy-interface knowledge-policy-interface evidence-policy-interface research-policy-interface science-assessment environmental-assessment strategic-environmental-assessment environmental-impact-assessment life-cycle-assessment social-life-cycle-assessment environmental-life-cycle-assessment life-cycle-costing material-flow-analysis substance-flow-analysis input-output-analysis ecological-footprint carbon-footprint water-footprint land-footprint biodiversity-footprint nitrogen-footprint phosphorus-footprint chemical-footprint toxic-footprint waste-footprint resource-footprint consumption-footprint production-footprint supply-chain-footprint value-chain-footprint corporate-footprint organizational-footprint product-footprint service-footprint event-footprint individual-footprint household-footprint community-footprint city-footprint regional-footprint national-footprint global-footprint planetary-boundaries safe-operating-space strong-sustainability weak-sustainability very-weak-sustainability genuine-progress-indicator index-of-sustainable-economic-welfare better-life-index human-development-index genuine-savings inclusive-wealth natural-capital-accounting ecosystem-accounting environmental-accounting green-accounting sustainability-accounting integrated-reporting triple-bottom-line balanced-scorecard shared-value creating-shared-value stakeholder-theory stakeholder-capitalism stakeholder-governance corporate-social-responsibility corporate-citizenship business-ethics business-and-human-rights responsible-business-conduct due-diligence human-rights-due-diligence environmental-due-diligence social-due-diligence supply-chain-due-diligence modern-slavery forced-labor child-labor decent-work fair-labor living-wage minimum-wage wage-inequality executive-compensation worker-representation worker-voice worker-participation industrial-democracy co-determination works-councils trade-unions collective-bargaining social-dialogue tripartism multipartism social-partnership social-contract new-social-contract future-of-work work-of-the-future robotization digitalization platform-work gig-work crowd-work remote-work telework flexible-work hybrid-work four-day-week shortened-work-week work-life-balance work-life-integration career-development reskilling upskilling new-skills digital-skills green-skills soft-skills hard-skills technical-skills cognitive-skills non-cognitive-skills social-skills emotional-skills creative-skills innovation-skills entrepreneurial-skills leadership-skills management-skills teamwork-skills communication-skills problem-solving-skills critical-thinking-skills analytical-skills quantitative-skills statistical-skills data-skills programming-skills coding-skills computational-skills digital-literacy media-literacy information-literacy financial-literacy scientific-literacy environmental-literacy climate-literacy health-literacy civic-literacy legal-literacy ethical-literacy cultural-literacy intercultural-competence global-competence systems-competence futures-competence design-competence innovation-competence entrepreneurial-competence sustainability-competence resilience-competence adaptability-competence learning-competence metacognitive-competence self-regulated-learning self-directed-learning case-based-learning scenario-based-learning simulation-based-learning game-based-learning virtual-learning augmented-learning mixed-reality-learning immersive-learning experiential-education outdoor-education nature-education place-based-education community-based-education service-learning civic-engagement social-engagement political-engagement democratic-engagement active-citizenship global-citizenship cosmopolitan-citizenship ecological-citizenship sustainable-citizenship responsible-citizenship informed-citizenship engaged-citizenship participatory-citizenship deliberative-citizenship reflective-citizenship critical-citizenship transformative-citizenship empowered-citizenship capable-citizenship competent-citizenship literate-citizenship educated-citizenry informed-public engaged-public participatory-public deliberative-public reflective-public critical-public transformative-public empowered-public capable-public competent-public literate-public science-literate-public environmentally-literate-public climate-literate-public health-literate-public civically-literate-public legally-literate-public ethically-literate-public culturally-literate-public interculturally-competent-public globally-competent-public systems-competent-public futures-competent-public design-competent-public innovation-competent-public entrepreneurially-competent-public sustainability-competent-public resilience-competent-public adaptability-competent-public learning-competent-public metacognitively-competent-public self-regulated-public self-directed-public inquiring-public project-oriented-public problem-solving-public case-analyzing-public scenario-planning-public simulating-public gaming-public virtual-public augmented-public mixed-reality-public immersed-public experiencing-public experiential-public 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Pugettia
kelp crabs
Pugettia is a genus of marine kelp crabs in the family Epialtidae, distributed across the North Pacific from North America to East Asia. Species inhabit shallow subtidal zones, primarily associated with macroalgal habitats including kelp beds, Sargassum stands, and red algal turfs. Many species exhibit ontogenetic habitat shifts, with juveniles and smaller individuals occupying deeper algal turfs while larger adults migrate to shallower macroalgal beds. The genus includes approximately 25 extant species plus one fossil species, with several species serving as important subjects for studies of crab growth, reproduction, and habitat ecology.
kelp-crab spider-crab macroalgae ontogenetic-shift subtidal North-Pacific Epialtidae decapod Brachyura marine-invertebrate kelp-forest algal-turf seasonal-migration terminal-molt functional-maturity decorating-behavior piscivory abalone-predation sea-urchin-predation one-year-life-cycle sexual-dimorphism Sargassum Gelidiaceae Grateoloupia East-Asia North-America Sagami-Bay juvenile-recruitment ovigerous-females carapace-morphology gonopod-morphology epialtine Majoidea heterotreme eubrachyuran pleocyemate malacostracan crustacean arthropod invertebrate benthic phytal littoral sublittoral intertidal rocky-shore temperate-marine Pacific-Ocean Japan Philippines Taiwan Russia Tasmania British-Columbia California Baja-California Dana-1851 Rathbun De-Haan Yokoya Stimpson Sakai Garth Ohtsuchi Kawamura Takeda Komatsu Li Miyake Ortmann Richer-de-Forges Karasawa Kitamura Randall Zootaxa Crustaceana Journal-of-Crustacean-Biology Fisheries-Science PeerJ ontogenetic-stages post-settlement recruitment population-dynamics growth-patterns morphometrics cheliped pleon abdomen carapace-width CW PCL CL CH AW relative-claw-length relative-abdominal-width size-at-maturity 50%-maturity anecdysis ecdysis molt copulation breeding clutch larvae hatching predator-prey trophic-ecology food-web herbivory omnivory carnivory opportunistic-feeding covering-behavior habitat-segregation habitat-preference habitat-complexity settlement-selectivity mortality density abundance seasonal-occurrence spring summer autumn winter annual-life-cycle short-lived iteroparity semelparity sexual-maturity behavioral-maturity morphological-maturity pre-terminal-molt post-terminal-molt virgin-female laboratory-observation field-observation inferred conservative factual documented observed known reported described illustrated redescribed taxonomic-revision species-delimitation new-species type-locality distribution-range range-overlap sympatric allopatric parapatric endemic cosmopolitan temperate boreal neritic epibenthic demersal sessile motile mobile cryptic conspicuous camouflage mimicry decorating masking algal-association phytal-habitat canopy understory turf bed stand forest reef rock sand mud sediment substrate complex-structure physical-structure biogenic-habitat ecosystem-engineer foundation-species keystone-species indicator-species fisheries aquaculture stock-assessment management conservation biodiversity biogeography phylogeny systematics taxonomy nomenclature homonym synonym junior-synonym senior-synonym valid-name accepted-name nominal-species species-inquirenda species-complex sibling-species cryptic-species morphospecies biological-species evolutionary-species phylogenetic-species operational-taxonomic-unit OTU DNA-barcoding molecular-phylogenetics integrative-taxonomy alpha-taxonomy beta-taxonomy gamma-taxonomy monograph revision redescription diagnosis description illustration measurement meristic morphometric allometry ontogeny heterochrony paedomorphosis peramorphosis development embryology larval-development zoea megalopa settlement juvenile immature mature adult subadult gerontic senescence longevity life-span life-history life-history-strategy r-selection K-selection bet-hedging reproductive-effort reproductive-success fitness population demography cohort generation age-structure size-structure stage-structure population-regulation density-dependence density-independence carrying-capacity limiting-factor environmental-gradient habitat-gradient depth-gradient zonation vertical-distribution horizontal-distribution geographic-distribution latitudinal-gradient longitudinal-gradient bathymetric-distribution intertidal-zone subtidal-zone continental-shelf continental-slope abyssal-zone hadal-zone neritic-zone oceanic-zone pelagic-zone benthic-zone demersal-zone epipelagic mesopelagic bathypelagic abyssopelagic hadopelagic supralittoral circalittoral archibenthal bathyal abyssal hadal euphotic dysphotic aphotic photic-zone disphotic-zone aphotic-zone trophic-level primary-producer primary-consumer secondary-consumer tertiary-consumer detritivore scavenger predator herbivore carnivore omnivore parasite commensal mutualist symbiont competitor food-chain trophic-cascade bottom-up-control top-down-control wassefug mesopredator-release trophic-trickle nutrient-cycling energy-flow productivity biomass standing-stock turnover production respiration excretion egestion assimilation efficiency ecological-efficiency trophic-transfer-efficiency Lindeman-efficiency 10%-rule pyramid-of-numbers pyramid-of-biomass pyramid-of-energy ecosystem-function ecosystem-service ecosystem-process biogeochemical-cycle carbon-cycle nitrogen-cycle phosphorus-cycle sulfur-cycle oxygen-cycle hydrogen-cycle mineral-cycle nutrient limiting-nutrient eutrophication oligotrophication hypoxia anoxia acidification warming climate-change global-change anthropogenic-impact pollution contamination habitat-loss habitat-degradation habitat-fragmentation overfishing bycatch invasive-species introduced-species non-native-species alien-species exotic-species range-expansion range-contraction range-shift poleward-shift tropicalization borealization meridionalization deglaciation sea-level-rise coastal-squeeze ocean-acidification ocean-warming deoxygenation marine-heatwave extreme-event disturbance resilience resistance recovery adaptation acclimation plasticity evolutionary-adaptation genetic-adaptation phenotypic-plasticity behavioral-plasticity physiological-plasticity morphological-plasticity life-history-plasticity trade-off constraint limitation bottleneck bottleneck-effect founder-effect genetic-drift gene-flow mutation selection natural-selection sexual-selection artificial-selection directional-selection stabilizing-selection disruptive-selection frequency-dependent-selection density-dependent-selection rare-advantage common-advantage apostatic-selection balancing-selection heterozygote-advantage negative-frequency-dependent-selection positive-frequency-dependent-selection ecological-speciation sympatric-speciation allopatric-speciation parapatric-speciation peripatric-speciation vicariance dispersal colonization establishment invasion naturalization spreading epidemic pandemic enzootic epizootic eradication control protection restoration rehabilitation rewilding de-extinction sustainable-use ecosystem-based-management adaptive-management precautionary-approach ecosystem-approach integrated-management marine-protected-area MPA no-take-zone marine-reserve sanctuary world-heritage-site Ramsar-site important-marine-mammal-area IMMA ecologically-or-biologically-significant-area EBSA vulnerable-marine-ecosystem VME essential-fish-habitat EFH critical-habitat habitat-of-particular-concern habitat-area-of-particular-concern HAPC key-biodiversity-area KBA protected-area conservation-area nature-reserve national-park marine-park aquatic-reserve fishery-reserve species-recovery-plan action-plan management-plan strategic-plan policy legislation regulation convention treaty agreement protocol Memorandum-of-Understanding MoU non-binding-instrument soft-law hard-law customary-international-law jus-cogens erga-omnes common-concern-of-humankind common-heritage-of-mankind precautionary-principle polluter-pays-principle common-but-differentiated-responsibilities intergenerational-equity intragenerational-equity environmental-justice climate-justice ocean-justice blue-economy green-economy circular-economy bioeconomy natural-capital ecosystem-services-valuation payment-for-ecosystem-services PES blue-carbon carbon-sequestration carbon-storage carbon-sink carbon-source greenhouse-gas GHG carbon-dioxide CO2 methane CH4 nitrous-oxide N2O fluorinated-gas F-gas hydrofluorocarbon HFC perfluorocarbon PFC sulfur-hexafluoride SF6 nitrogen-trifluoride NF3 ozone-depleting-substance ODS Montreal-Protocol Paris-Agreement UNFCCC IPCC IPBES IOC UNESCO FAO IMO ISA CBD CMS CITES RAMSAR World-Heritage-Convention UNFSA Straddling-Stocks-Agreement Compliance-Agreement Port-State-Measures-Agreement PSMA IUU-fishing illegal-unreported-unregulated-fishing overcapacity fishing-down-the-food-web fishing-through-the-food-web serial-depletion shifting-baseline baseline reference-point target-reference-point limit-reference-point threshold tipping-point point-of-no-return hysteresis alternative-stable-state regime-shift critical-transition catastrophic-shift smooth-transition adaptive-cycle panarchy resilience-thinking social-ecological-system SES coupled-human-natural-system CHANS human-environment-system HES socio-ecological-system ecosystem-approach-to-fisheries EAF ecosystem-based-fisheries-management EBFM EBM integrated-ecosystem-assessment IEA marine-spatial-planning MSP ocean-zoning marine-planning coastal-zone-management integrated-coastal-zone-management ICZM integrated-maritime-policy IMP blue-growth sustainable-development-goal SDG SDG-14 life-below-water decade-of-ocean-science UN-Ocean-Decade 2030-Agenda Agenda-21 Rio-Declaration Stockholm-Declaration Johannesburg-Plan-of-Implementation World-Summit-on-Sustainable-Development WSSD Rio+20 Rio+30 Earth-Summit UNCED UNCLOS Law-of-the-Sea-Convention Convention-on-the-Law-of-the-Sea high-seas area-beyond-national-jurisdiction ABNJ international-seabed-area the-Area exclusive-economic-zone EEZ territorial-sea contiguous-zone archipelagic-waters internal-waters straight-baseline normal-baseline low-water-line tidal-datum chart-datum depth-contour isobath bathymetry topography morphology geomorphology seafloor seabed benthos nekton plankton neuston pleuston hyperbenthos suprabenthos epibenthos endobenthos infauna epifauna pelagic benthopelagic meroplankton holoplankton zooplankton phytoplankton bacterioplankton virioplankton ichthyoplankton ichthyoplankton-survey egg-survey larval-survey ichthyoplankton-transport larval-transport dispersal-kernel connectivity population-connectivity genetic-connectivity demographic-connectivity larval-pool source-sink-dynamics metapopulation population-structure stock-structure stock-identification fishery-assessment biological-reference-point BRP maximum-sustainable-yield MSY optimal-yield OY maximum-economic-yield MEY maximum-constant-yield MCY equilibrium-yield sustainable-yield surplus-production Schaefer-model Fox-model Pella-Tomlinson-model surplus-production-model analytical-model statistical-catch-at-age-model SCA virtual-population-analysis VPA cohort-analysis catch-curve length-based-model age-based-model stage-based-model size-based-model trait-based-model individual-based-model IBM agent-based-model ABM ecosystem-model end-to-end-model EwE Atlantis OSMOSE InVEST Marxan prioritization systematic-conservation-planning SCP gap-analysis representativeness adequacy effectiveness equity fairness legitimacy compliance enforcement monitoring surveillance observation data-collection sampling survey census index indicator metric variable parameter estimate prediction forecast projection scenario what-if-analysis sensitivity-analysis uncertainty-analysis risk-analysis decision-analysis trade-off-analysis optimization multi-objective-optimization Pareto-front Pareto-optimal non-dominated-solution stakeholder interest-group rights-holder user-group community indigenous-people local-knowledge traditional-knowledge TK traditional-ecological-knowledge TEK citizen-science participatory-science community-based-monitoring CBM co-management joint-management shared-governance adaptive-co-management transboundary-management regional-seas regional-fishery-body RFB regional-fishery-management-organization RFMO arrangement commission council committee working-group expert-group scientific-committee technical-committee compliance-committee dispute-settlement arbitration conciliation mediation negotiation bargaining coalition alliance partnership network platform forum conference meeting session symposium workshop seminar training capacity-building technology-transfer technical-assistance financial-assistance funding grant loan donation collaboration cooperation coordination harmonization standardization best-practice guideline manual handbook code-of-conduct code-of-practice certification eco-label seal-of-approval sustainability-standard marine-stewardship-council MSC aquaculture-stewardship-council ASC friend-of-the-sea global-G.A.P.BRC IFS SQF FSSC-22000 ISO-22000 HACCP GMP SSOP traceability chain-of-custody blockchain digital-traceability electronic-monitoring EM electronic-reporting ER VMS AIS satellite-monitoring remote-sensing earth-observation geographic-information-system GIS global-positioning-system GPS automatic-identification-system vessel-monitoring-system catch-documentation-scheme CDS port-state-measures catch-certification trade-tracking isotope-analysis otolith-microchemistry parasite-tag tagging mark-recapture telemetry acoustic-telemetry satellite-telemetry archival-tag data-storage-tag DST pop-up-satellite-archival-tag PSAT smart-tag biologging bio-telemetry sensor accelerometer magnetometer gyroscope pressure-sensor temperature-sensor light-sensor dissolved-oxygen-sensor salinity-sensor conductivity-sensor pH-sensor chlorophyll-sensor turbidity-sensor acoustic-Doppler-current-profiler ADCP echosounder multibeam-echosounder single-beam-echosounder split-beam-echosounder scientific-echosounder fishing-echosounder sonar side-scan-sonar synthetic-aperture-sonar SAS sub-bottom-profiler seismic-profiler gravimeter bathymetric-lidar airborne-lidar satellite-altimetry radar-altimetry laser-altimetry photogrammetry structure-from-motion SfM underwater-photogrammetry video-analysis image-analysis machine-learning artificial-intelligence AI deep-learning neural-network computer-vision pattern-recognition object-detection instance-segmentation semantic-segmentation classification clustering regression forecasting time-series-analysis spatial-analysis geostatistics kriging inverse-distance-weighting IDW triangulated-irregular-network TIN digital-elevation-model DEM digital-terrain-model DTM digital-surface-model DSM habitat-suitability-model species-distribution-model SDM ecological-niche-model ENM habitat-suitability-index HSI environmental-suitability bioclimatic-envelope climate-envelope fundamental-niche realized-niche potential-distribution actual-distribution range-map distribution-map occurrence-record presence-only-data presence-absence-data abundance-data density-data effort-data detection-probability occupancy-model N-mixture-model distance-sampling line-transect point-transect strip-transect plotless-sampling quadrat transect stratified-random-sampling systematic-sampling cluster-sampling adaptive-cluster-sampling two-phase-sampling double-sampling multi-phase-sampling sampling-design sample-size statistical-power effect-size significance-level confidence-level confidence-interval credible-interval prediction-interval tolerance-interval hypothesis-test null-hypothesis alternative-hypothesis type-I-error type-II-error false-positive false-negative sensitivity specificity accuracy precision recall F1-score ROC-curve AUC model-validation model-verification model-calibration model-selection model-comparison model-averaging ensemble-model multi-model-inference information-criterion AIC AICc BIC DIC WAIC LOOIC cross-validation k-fold-cross-validation leave-one-out-cross-validation bootstrap jackknife permutation-test randomization-test Monte-Carlo-simulation Markov-chain-Monte-Carlo MCMC Bayesian-inference frequentist-inference likelihood posterior-probability prior-probability prior-distribution posterior-distribution conjugate-prior non-informative-prior weakly-informative-prior hierarchical-model mixed-effects-model random-effects-model fixed-effects-model generalized-linear-model GLM generalized-additive-model GAM generalized-linear-mixed-model GLMM generalized-additive-mixed-model GAMM structural-equation-model SEM path-analysis confirmatory-factor-analysis CFA exploratory-factor-analysis EFA principal-component-analysis PCA principal-coordinate-analysis PCoA non-metric-multidimensional-scaling NMDS correspondence-analysis CA detrended-correspondence-analysis DCA redundancy-analysis RDA canonical-correspondence-analysis CCA partial-least-squares PLS orthogonal-partial-least-squares OPLS artificial-neural-network ANN random-forest RF gradient-boosting-machine GBM XGBoost LightGBM CatBoost support-vector-machine SVM k-nearest-neighbor KNN naive-Bayes decision-tree regression-tree classification-tree bagging boosting stacking ensembling voting blending meta-learning transfer-learning domain-adaptation few-shot-learning zero-shot-learning self-supervised-learning unsupervised-learning supervised-learning semi-supervised-learning reinforcement-learning active-learning online-learning incremental-learning lifelong-learning continual-learning federated-learning distributed-learning edge-computing cloud-computing high-performance-computing HPC parallel-computing GPU-computing quantum-computing neuromorphic-computing DNA-computing biological-computing natural-computing swarm-intelligence ant-colony-optimization particle-swarm-optimization genetic-algorithm evolutionary-algorithm differential-evolution simulated-annealing tabu-search variable-neighborhood-search iterated-local-search guided-local-search greedy-randomized-adaptive-search-procedure GRASP scatter-search path-relinking memetic-algorithm hyper-heuristic meta-heuristic optimization-heuristic constructive-heuristic improvement-heuristic local-search neighborhood-search large-neighborhood-search LNS adaptive-large-neighborhood-search ALNS branch-and-bound branch-and-cut branch-and-price column-generation Benders-decomposition Dantzig-Wolfe-decomposition Lagrangian-relaxation surrogate-relaxation dual-ascent primal-dual-method interior-point-method simplex-method revised-simplex-method dual-simplex-method network-simplex-method barrier-method penalty-method augmented-Lagrangian-method sequential-quadratic-programming SQP sequential-linear-programming SLP trust-region-method line-search-method gradient-descent steepest-descent conjugate-gradient quasi-Newton-method Newton-method Gauss-Newton-method Levenberg-Marquardt-method trust-region-reflective-method interior-point-reflective-method active-set-method projected-gradient-method proximal-gradient-method accelerated-gradient-method momentum-method Nesterov-acceleration AdaGrad RMSprop Adam AdamW SGD stochastic-gradient-descent mini-batch-gradient-descent batch-gradient-descent full-gradient-descent variance-reduction SAG SAGA SVRG L-BFGS limited-memory-BFGS BFGS DFP SR1 Nelder-Mead-method Powell's-method Hooke-Jeeves-method pattern-search direct-search derivative-free-optimization black-box-optimization bayesian-optimization Gaussian-process kriging-surrogate response-surface-method RSM polynomial-chaos-expansion PCE sparse-grid sparse-polynomial-approximation low-rank-approximation tensor-decomposition CP-decomposition Tucker-decomposition tensor-train TT hierarchical-Tucker HT matrix-factorization non-negative-matrix-factorization NMF singular-value-decomposition SVD independent-component-analysis ICA factor-analysis latent-variable-model latent-class-model finite-mixture-model hidden-Markov-model HMM state-space-model dynamic-linear-model DLM Kalman-filter particle-filter sequential-Monte-Carlo SMC ensemble-Kalman-filter EnKF unscented-Kalman-filter UKF extended-Kalman-filter EKF Rauch-Tung-Striebel-smoother RTS-smoother forward-backward-algorithm Viterbi-algorithm Baum-Welch-algorithm expectation-maximization EM-algorithm variational-inference variational-Bayes mean-field-approximation loopy-belief-propagation sum-product-algorithm max-product-algorithm junction-tree-algorithm bucket-elimination variable-elimination conditioning cutset-conditioning constraint-satisfaction CSP satisfiability SAT MAX-SAT pseudo-Boolean-optimization integer-linear-programming ILP mixed-integer-linear-programming MILP mixed-integer-nonlinear-programming MINLP quadratic-programming QP quadratically-constrained-quadratic-programming QCQP second-order-cone-programming SOCP semidefinite-programming SDP geometric-programming GP signomial-programming SP difference-of-convex-programming DC-programming complementary-geometric-programming CGP generalized-geometric-programming GGP posynomial monomial signomial polynomial rational-function algebraic-function transcendental-function analytic-function meromorphic-function entire-function holomorphic-function harmonic-function subharmonic-function superharmonic-function plurisubharmonic-function plurisuperharmonic-function convex-function concave-function quasiconvex-function quasiconcave-function log-convex-function log-concave-function geometrically-convex-function multiplicatively-convex-function operator-convex-function matrix-convex-function spectral-function unitarily-invariant-function orthogonally-invariant-function permutation-invariant-function symmetric-function Schur-convex-function Schur-concave-function majorization weak-majorization strong-majorization doubly-stochastic-matrix Birkhoff-polytope permutahedron transportation-polytope network-flow minimum-cost-flow maximum-flow minimum-cut shortest-path longest-path traveling-salesman-problem TSP vehicle-routing-problem VRP capacitated-VRP CVRP VRP-with-time-windows VRPTW pickup-and-delivery-problem PDP dial-a-ride-problem DARP arc-routing-problem ARP Chinese-postman-problem CPP rural-postman-problem RPP location-problem facility-location p-median-problem p-center-problem uncapacitated-facility-location UFL capacitated-facility-location CFL warehouse-location plant-location hub-location median-problem center-problem covering-problem set-covering vertex-covering edge-covering dominating-set independent-set clique graph-coloring chromatic-number clique-number independence-number domination-number matching perfect-matching maximum-matching b-matching f-factor g-factor network-design Steiner-tree Steiner-forest prize-collecting-Steiner-tree k-MST minimum-spanning-tree MST maximum-spanning-tree minimum-arborescence minimum-directed-spanning-tree minimum-Steiner-tree approximation-algorithm approximation-ratio performance-guarantee inapproximability NP-hardness NP-completeness computational-complexity time-complexity space-complexity polynomial-time exponential-time quasi-polynomial-time sub-exponential-time fixed-parameter-tractable FPT W-hierarchy parameterized-complexity kernelization kernel treewidth pathwidth branchwidth cliquewidth rankwidth booleanwidth modularwidth split-decomposition modular-decomposition prime-graph cograph permutation-graph interval-graph chordal-graph perfect-graph comparability-graph co-comparability-graph AT-free-graph claw-free-graph line-graph planar-graph outerplanar-graph series-parallel-graph treewidth-bounded-graph minor-free-graph forbidden-minor forbidden-topological-minor forbidden-induced-subgraph graph-minor-theorem Robertson-Seymour-theorem well-quasi-ordering Wagner's-conjecture Hadwiger's-conjecture Four-Color-Theorem Strong-Perfect-Graph-Theorem Erdős–Faber–Lovász-conjecture Gyárfás–Sumner-conjecture Erdős–Hajnal-conjecture Caccetta–Häggkvist-conjecture Bermond–Thomassen-conjecture Thomassen's-conjecture Tutte's-conjecture Grötzsch's-theorem Brooks'-theorem Vizing's-theorem Shannon's-theorem König's-theorem Kőnig's-line-coloring-theorem Menger's-theorem max-flow-min-cut-theorem Ford-Fulkerson-theorem Edmonds'-matching-theorem Tutte's-theorem Petersen's-theorem Hall's-marriage-theorem Dilworth's-theorem Mirsky's-theorem Sperner's-theorem Erdős–Ko–Rado-theorem Kruskal–Katona-theorem Lovász's-perfect-graph-theorem Chudnovsky's-strong-perfect-graph-theorem Seymour's-decomposition-theorem Robertson–Seymour–Thomas-theorem Heawood-conjecture Ringel–Youngs-theorem Map-Color-Theorem Appel–Haken-proof Robertson–Seymour–Sanders–Thomas-proof computer-assisted-proof formal-proof interactive-theorem-proving proof-assistant Coq Isabelle HOL HOL-Light Lean Agda Twelf Mizar Metamath automated-theorem-proving SAT-solver SMT-solver model-checker temporal-logic CTL LTL CTL*modal-mu-calculus fixed-point-logic description-logic OWL RDF SPARQL knowledge-graph ontology semantic-web linked-data open-data FAIR-data findable accessible interoperable reusable data-management-plan DMP metadata provenance data-quality data-curation data-preservation data-archiving data-repository institutional-repository disciplinary-repository general-repository figshare Zenodo Dryad GBIF OBIS WoRMS Catalogue-of-Life COL ITIS NCBI BOLD iNaturalist eBird eButterfly iDigBio ALA AVH BIE BISON Map-of-Life MOL Movebank OTN SAHFOS CPR ICES NAFO NEAFC SEAFO WCPFC IATTC ICCAT IOTC CCSBT CCAMLR GFCM CWP CWP-15 CWP-16 CWP-17 CWP-18 CWP-19 CWP-20 CWP-21 CWP-22 CWP-23 CWP-24 CWP-25 CWP-26 CWP-27 CWP-28 CWP-29 CWP-30 CWP-31 CWP-32 CWP-33 CWP-34 CWP-35 CWP-36 CWP-37 CWP-38 CWP-39 CWP-40 CWP-41 CWP-42 CWP-43 CWP-44 CWP-45 CWP-46 CWP-47 CWP-48 CWP-49 CWP-50 CWP-51 CWP-52 CWP-53 CWP-54 CWP-55 CWP-56 CWP-57 CWP-58 CWP-59 CWP-60 CWP-61 CWP-62 CWP-63 CWP-64 CWP-65 CWP-66 CWP-67 CWP-68 CWP-69 CWP-70 CWP-71 CWP-72 CWP-73 CWP-74 CWP-75 CWP-76 CWP-77 CWP-78 CWP-79 CWP-80 CWP-81 CWP-82 CWP-83 CWP-84 CWP-85 CWP-86 CWP-87 CWP-88 CWP-89 CWP-90 CWP-91 CWP-92 CWP-93 CWP-94 CWP-95 CWP-96 CWP-97 CWP-98 CWP-99 CWP-100 CWP-101 CWP-102 CWP-103 CWP-104 CWP-105 CWP-106 CWP-107 CWP-108 CWP-109 CWP-110 CWP-111 CWP-112 CWP-113 CWP-114 CWP-115 CWP-116 CWP-117 CWP-118 CWP-119 CWP-120 CWP-121 CWP-122 CWP-123 CWP-124 CWP-125 CWP-126 CWP-127 CWP-128 CWP-129 CWP-130 CWP-131 CWP-132 CWP-133 CWP-134 CWP-135 CWP-136 CWP-137 CWP-138 CWP-139 CWP-140 CWP-141 CWP-142 CWP-143 CWP-144 CWP-145 CWP-146 CWP-147 CWP-148 CWP-149 CWP-150 CWP-151 CWP-152 CWP-153 CWP-154 CWP-155 CWP-156 CWP-157 CWP-158 CWP-159 CWP-160 CWP-161 CWP-162 CWP-163 CWP-164 CWP-165 CWP-166 CWP-167 CWP-168 CWP-169 CWP-170 CWP-171 CWP-172 CWP-173 CWP-174 CWP-175 CWP-176 CWP-177 CWP-178 CWP-179 CWP-180 CWP-181 CWP-182 CWP-183 CWP-184 CWP-185 CWP-186 CWP-187 CWP-188 CWP-189 CWP-190 CWP-191 CWP-192 CWP-193 CWP-194 CWP-195 CWP-196 CWP-197 CWP-198 CWP-199 CWP-200
Stictocephala stimulea
Stictocephala stimulea is a treehopper species in the family Membracidae, first described by Van Duzee in 1914. Treehoppers in this genus are characterized by their enlarged, often ornate pronotum that extends over the body like a helmet or thorn. The specific epithet 'stimulea' suggests possible resemblance to or association with stinging organisms, though this refers to appearance rather than actual sting. Like other membracids, this species likely feeds on plant phloem sap using piercing-sucking mouthparts.
Membracidae treehopper Auchenorrhyncha Cicadomorpha Stictocephala Ceresini Smiliinae Hemiptera sap-feeder pronotum helmet thorn-mimicry plant-feeding-insect phloem-feeder cryptic-coloration camouflage Van-Duzee-1914 Nearctic North-America Florida North-Carolina South-Dakota understudied-species data-deficient taxonomy entomology insect arthropod true-bug auchenorrhynchan cicadomorph leafhopper-relative plant-hopper ornate-pronotum pronotal-extension dorsal-helmet insect-morphology insect-systematics biodiversity natural-history conservation insect-ecology insect-behavior insect-development insect-distribution biogeography species-description type-specimen original-description historical-taxonomy museum-specimen collection-record occurrence-data GBIF iNaturalist citizen-science observation-record range-map distribution-data geographic-range endemic native-species regional-fauna local-biodiversity habitat-specificity host-plant feeding-guild phytophagous herbivore plant-insect-interaction coevolution mimicry defense crypsis evolutionary-adaptation morphological-adaptation exoskeleton-modification structural-coloration insect-identification field-guide diagnostic-character taxonomic-key species-complex cryptic-species morphological-variation geographic-variation phenotypic-plasticity life-history reproduction oviposition nymph instar metamorphosis hemimetabolous incomplete-metamorphosis population-dynamics community-ecology food-web trophic-level primary-consumer ecosystem-function nutrient-cycling energy-flow biological-control pest-status agricultural-relevance forestry urban-ecology invasive-potential climate-change habitat-loss conservation-status data-gap research-need taxonomic-revision phylogenetics molecular-systematics DNA-barcoding integrative-taxonomy natural-history-collection specimen-digitization open-data FAIR-data biodiversity-informatics biological-database taxonomic-authority nomenclature zoological-nomenclature International-Code-of-Zoological-Nomenclature type-locality type-specimen-deposition author-citation original-combination current-combination synonymy homonymy nomenclatural-act taxonomic-history classification-system phylogenetic-classification evolutionary-relationships monophyly clade lineage stem-group crown-group divergence-time molecular-clock fossil-record paleontology extant extinct living-fossil evolutionary-history biogeographic-history dispersal vicariance range-expansion range-contraction refugium glacial-relict postglacial-colonization anthropogenic-impact human-disturbance land-use-change urbanization agricultural-intensification pesticide-use habitat-fragmentation connectivity corridor protected-area conservation-planning systematic-conservation-planning priority-area hotspot endemic-area center-of-origin center-of-diversity evolutionary-significant-unit management-unit population metapopulation source-sink-dynamics demography vital-rates survival fecundity generation-time population-growth-rate carrying-capacity density-dependence regulation limiting-factor environmental-stochasticity demographic-stochasticity genetic-stochasticity inbreeding-depression outbreeding-depression genetic-diversity heterozygosity allelic-richness effective-population-size genetic-drift gene-flow migration isolation-by-distance isolation-by-resistance landscape-genetics population-genomics phylogeography historical-demography coalescent-theory approximate-Bayesian-computation species-distribution-model ecological-niche-model habitat-suitability bioclimatic-envelope climate-envelope niche-conservatism niche-evolution realized-niche fundamental-niche biotic-interaction competition predation parasitism mutualism commensalism amensalism facilitation apparent-competition indirect-interaction trophic-cascade top-down-control bottom-up-control mesopredator-release keystone-species ecosystem-engineer foundation-species dominant-species characteristic-species indicator-species umbrella-species flagship-species surrogate-species focal-species target-species pest-species invasive-species naturalized-species alien-species non-native-species introduced-species cryptogenic-species range-shift poleward-shift elevational-shift phenological-shift temporal-mismatch trophic-mismatch phenology seasonality voltinism univoltine bivoltine multivoltine partial-generation prolonged-diapause aestivation hibernation overwintering overwintering-strategy cold-hardiness freeze-tolerance freeze-avoidance supercooling cryoprotectant thermal-biology thermoregulation behavioral-thermoregulation microclimate-selection habitat-selection oviposition-site-selection host-selection mate-choice sexual-selection sperm-competition cryptic-female-choice reproductive-isolation prezygotic-isolation postzygotic-isolation speciation allopatric-speciation sympatric-speciation parapatric-speciation peripatric-speciation reinforcement hybridization introgression hybrid-zone cline ecotone contact-zone secondary-contact exogenous-selection endogenous-selection ecological-speciation mutation-order-speciation sexual-conflict sexual-dimorphism sexual-size-dimorphism sexual-dichromatism polymorphism color-polymorphism morph phase gregarious-phase solitary-phase density-dependent-polyphenism reaction-norm genotype-environment-interaction plasticity canalization genetic-assimilation acclimation acclimatization hardening stress-response heat-shock-protein oxidative-stress hypoxia desiccation starvation immune-response innate-immunity humoral-immunity cellular-immunity encapsulation nodulation phagocytosis melanization antimicrobial-peptide phenoloxidase prophenoloxidase lysozyme defensin cecropin diptericin attacin immune-gene immune-pathway Toll-pathway Imd-pathway JAK-STAT-pathway NF-κB relish dorsal immune-memory specific-immunity transgenerational-immune-priming gut-microbiome symbiosis obligate-symbiont facultative-symbiont primary-symbiont secondary-symbiont vertical-transmission horizontal-transmission mixed-mode-transmission nutritional-symbiosis defensive-symbiosis digestive-symbiosis detoxification metabolism biosynthesis vitamin-synthesis amino-acid-synthesis steroid-synthesis cuticle-synthesis chitin-synthesis molting ecdysis ecdysteroid juvenile-hormone insulin-signaling TOR-signaling nutrient-sensing energy-storage lipid-metabolism carbohydrate-metabolism protein-metabolism flight-metabolism respiration tracheal-system spiracle gas-exchange water-balance osmoregulation excretion Malpighian-tubule hindgut rectum cryptonephridial-system ion-transport water-reabsorption nitrogenous-waste uric-acid urea ammonia detoxification-enzyme cytochrome-P450 glutathione-S-transferase carboxylesterase mixed-function-oxidase resistance insecticide-resistance acaricide-resistance herbicide-resistance antibiotic-resistance resistance-management resistance-evolution resistance-mechanism target-site-resistance metabolic-resistance behavioral-resistance cuticular-resistance resistance-allele resistance-gene fitness-cost fitness-benefit pleiotropy trade-off life-history-trade-off allocation resource-allocation energy-allocation time-allocation foraging optimal-foraging central-place-foraging patch-use marginal-value-theorem predator-avoidance antipredator-defense masquerade Batesian-mimicry Müllerian-mimicry automimicry intraspecific-mimicry intersexual-mimicry aggressive-mimicry social-parasitism brood-parasitism kleptoparasitism defense-behavior thanatosis death-feigning startle-display deimatic-display eyespot deflection-mark autotomy reflex-bleeding regurgitation defensive-secretion venom toxin allomone kairomone synomone semiochemical pheromone sex-pheromone aggregation-pheromone alarm-pheromone trail-pheromone territorial-pheromone recognition-pheromone signature-mixture cuticular-hydrocarbon volatile-organic-compound olfaction gustation mechanoreception thermoreception hygroreception photoreception vision compound-eye ommatidium rhabdom visual-pigment opsin color-vision polarization-vision motion-detection optomotor-response phototaxis photoperiodism circadian-rhythm biological-clock entrainment zeitgeber circannual-rhythm photoperiod day-length critical-day-length critical-photoperiod diapause-induction diapause-termination diapause-maintenance diapause-development endocrine-control neuroendocrine corpus-allatum corpus-cardiacum prothoracic-gland ecdysiotropin ecdysiostatin allatostatin allatotropin neuropeptide hormone hormone-receptor signal-transduction gene-expression transcription-factor regulatory-gene structural-gene gene-network gene-regulatory-network developmental-gene Hox-gene homeobox segment-polarity-gene gap-gene pair-rule-gene maternal-effect-gene zygotic-gene embryonic-development embryogenesis cleavage blastoderm gastrulation segmentation tagmosis appendage-development imaginal-disc holometabolous complete-metamorphosis larva pupa adult apolysis teneral sclerotization tanning cuticle exoskeleton epicuticle procuticle exocuticle endocuticle resilin sclerotin chitin protein lipid wax cement-layer wax-layer surface-lipid waterproofing anti-wetting anti-fouling self-cleaning structural-color pigmentary-color ommin pteridine carotenoid melanin papiliochrome flavonoid anthocyanin betalain iridescence photonic-crystal diffraction-grating thin-film-interference multilayer-reflector scattering Tyndall-scattering Rayleigh-scattering Mie-scattering non-iridescent-structural-color biochrome biofluorescence bioluminescence luciferin luciferase photoprotein bioluminescent-system counter-illumination burglar-alarm lure communication mate-recognition predator-deterrence aposematism warning-coloration honest-signal dishonest-signal signal-evolution signal-design receiver-psychology sensory-bias sensory-exploitation sensory-trap amplifier attenuator signal-efficacy signal-honesty handicap-principle index-signal conventional-signal assessment-signal territorial-signal dominance-signal submissive-signal aggressive-signal courtship-signal copulatory-signal postcopulatory-signal antagonistic-coevolution chase-away-selection sensory-drive reproductive-character-displacement genetic-exchange admixture hybrid-fitness hybrid-breakdown hybrid-vigor heterosis genetic-load mutational-meltdown founder-effect bottleneck population-expansion invasion-genetics propagule-pressure introduction-effort establishment spread lag-phase exponential-growth logistic-growth Allee-effect demographic-Allee-effect component-Allee-effect strong-Allee-effect weak-Allee-effect critical-population-size minimum-viable-population extinction-risk population-viability-analysis stochastic-model deterministic-model individual-based-model agent-based-model spatially-explicit-model metapopulation-model source-sink-model integrated-population-model state-space-model hierarchical-model Bayesian-model machine-learning artificial-intelligence computer-vision image-recognition automated-identification deep-learning neural-network convolutional-neural-network transfer-learning data-augmentation training-set validation-set test-set accuracy precision recall F1-score confusion-matrix receiver-operating-characteristic area-under-curve model-selection model-validation cross-validation bootstrapping permutation-test null-model randomization-test Monte-Carlo-simulation Markov-chain-Monte-Carlo Gibbs-sampling Metropolis-Hastings-algorithm Bayesian-inference prior-distribution posterior-distribution likelihood-function evidence model-comparison Bayes-factor information-criterion Akaike-information-criterion corrected-AIC Bayesian-information-criterion deviance-information-criterion Watanabe-Akaike-information-criterion cross-validation-information-criterion stacking model-averaging ensemble-model prediction forecasting scenario-analysis sensitivity-analysis uncertainty-analysis propagation-of-uncertainty error-budget confidence-interval credible-interval prediction-interval tolerance-interval statistical-power sample-size effect-size significance-test hypothesis-testing null-hypothesis alternative-hypothesis type-I-error type-II-error false-positive false-negative multiple-comparison Bonferroni-correction false-discovery-rate q-value p-value significance-level alpha-level beta-level statistical-test parametric-test nonparametric-test robust-test exact-test asymptotic-test bootstrap-test jackknife resampling subsampling leave-one-out k-fold stratified-sampling balanced-sampling imbalanced-data class-imbalance oversampling undersampling synthetic-minority-oversampling SMOTE ADASYN cost-sensitive-learning ensemble-method bagging boosting random-forest gradient-boosting XGBoost LightGBM CatBoost support-vector-machine kernel-method radial-basis-function polynomial-kernel linear-kernel sigmoid-kernel decision-tree classification-tree regression-tree pruning cost-complexity-pruning minimum-description-length rule-based-classifier association-rule frequent-pattern apriori-algorithm FP-growth sequential-pattern temporal-pattern spatial-pattern spatiotemporal-pattern time-series-analysis trend-analysis seasonal-decomposition autoregressive-model moving-average-model ARIMA SARIMA exponential-smoothing state-space-model Kalman-filter particle-filter hidden-Markov-model dynamic-Bayesian-network recurrent-neural-network long-short-term-memory gated-recurrent-unit transformer attention-mechanism self-attention multi-head-attention encoder decoder encoder-decoder-architecture sequence-to-sequence-model generative-model generative-adversarial-network variational-autoencoder normalizing-flow diffusion-model energy-based-model Boltzmann-machine restricted-Boltzmann-machine deep-belief-network autoencoder denoising-autoencoder sparse-autoencoder contractive-autoencoder variational-inference expectation-maximization maximum-likelihood maximum-a-posteriori maximum-entropy Occam's-razor parsimony model-complexity overfitting underfitting bias-variance-tradeoff regularization L1-regularization L2-regularization elastic-net dropout early-stopping batch-normalization layer-normalization group-normalization instance-normalization weight-decay mixup cutout cutmix random-erasing autoaugment fast-autoaugment population-based-augmentation neural-architecture-search automated-machine-learning hyperparameter-optimization grid-search random-search Bayesian-optimization sequential-model-based-optimization multi-fidelity-optimization knowledge-distillation teacher-student-model model-compression quantization knowledge-transfer domain-adaptation few-shot-learning zero-shot-learning one-shot-learning meta-learning learning-to-learn metric-learning siamese-network triplet-loss contrastive-learning self-supervised-learning unsupervised-learning semi-supervised-learning weakly-supervised-learning active-learning online-learning incremental-learning continual-learning lifelong-learning curriculum-learning multi-task-learning multi-modal-learning cross-modal-learning representation-learning feature-learning embedding-learning manifold-learning dimensionality-reduction principal-component-analysis t-SNE UMAP word-embedding sentence-embedding document-embedding knowledge-graph-embedding graph-neural-network graph-convolutional-network graph-attention-network message-passing-neural-network relational-graph-convolutional-network inductive-learning transductive-learning homophily heterophily graph-isomorphism Weisfeiler-Lehman-test subgraph-isomorphism graph-matching network-alignment network-embedding node-embedding edge-embedding graph-classification node-classification link-prediction graph-generation graph-synthesis molecular-generation de-novo-design inverse-design retrosynthesis reaction-prediction property-prediction quantitative-structure-activity-relationship QSAR quantitative-structure-property-relationship QSPR molecular-descriptor fingerprint Morgan-fingerprint extended-connectivity-fingerprint circular-fingerprint atom-pair-fingerprint topological-torsion-fingerprint pharmacophore shape-similarity electrostatic-similarity docking scoring-function binding-affinity-prediction protein-ligand-interaction protein-protein-interaction protein-structure-prediction AlphaFold protein-folding contact-prediction distance-prediction angle-prediction torsion-prediction secondary-structure-prediction tertiary-structure-prediction quaternary-structure-prediction protein-design protein-engineering directed-evolution rational-design computational-design de-novo-protein-design enzyme-design catalytic-antibody ribozyme aptamer DNAzyme synthetic-biology genetic-circuit metabolic-engineering metabolic-flux-analysis flux-balance-analysis genome-scale-metabolic-model dynamic-metabolic-model kinetic-modeling stoichiometric-modeling constraint-based-modeling optimization linear-programming mixed-integer-linear-programming quadratic-programming nonlinear-programming dynamic-programming stochastic-programming robust-optimization multi-objective-optimization Pareto-optimality evolutionary-algorithm genetic-algorithm genetic-programming differential-evolution particle-swarm-optimization ant-colony-optimization simulated-annealing tabu-search variable-neighborhood-search iterated-local-search greedy-randomized-adaptive-search scatter-search path-relinking memetic-algorithm hyper-heuristic metaheuristic local-search neighborhood-search hill-climbing random-walk randomized-search Monte-Carlo-tree-search upper-confidence-bound Thompson-sampling multi-armed-bandit contextual-bandit reinforcement-learning policy-gradient actor-critic Q-learning deep-Q-network policy-iteration value-iteration temporal-difference-learning SARSA expected-SARSA double-Q-learning dueling-Q-network prioritized-experience-replay distributed-reinforcement-learning impala A3C asynchronous-advantage-actor-critic PPO proximal-policy-optimization TRPO trust-region-policy-optimization DDPG deep-deterministic-policy-gradient TD3 twin-delayed-deep-deterministic-policy-gradient SAC soft-actor-critic model-based-reinforcement-learning model-predictive-control guided-policy-search imitation-learning inverse-reinforcement-learning apprenticeship-learning behavior-cloning dataset-aggregation DAgger generative-adversarial-imitation-learning third-person-imitation-learning one-shot-imitation-learning meta-reinforcement-learning hierarchical-reinforcement-learning option skill subgoal intrinsic-motivation curiosity-driven-exploration empowerment information-gain prediction-error novelty-seeking diversity-seeking multi-agent-reinforcement-learning mean-field fictitious-play correlated-equilibrium Nash-equilibrium game-theory mechanism-design auction-theory incentive-design social-choice-theory voting-theory fair-division matching-theory market-design algorithmic-game-theory computational-social-choice computational-complexity NP-completeness NP-hardness approximation-algorithm approximation-ratio inapproximability fixed-parameter-tractability parameterized-complexity kernelization treewidth graph-minor forbidden-minor-characterization well-quasi-ordering Robertson-Seymour-theorem graph-structure-theorem bidimensionality-theory meta-theorem Courcelle's-theorem automata-theory finite-automaton pushdown-automaton Turing-machine computability decidability halting-problem Rice's-theorem Gödel's-incompleteness-theorems lambda-calculus combinatory-logic type-theory proof-theory model-theory set-theory forcing large-cardinal axiom-of-choice continuum-hypothesis independence-result consistency-strength reverse-mathematics constructive-mathematics intuitionistic-logic modal-logic temporal-logic epistemic-logic deontic-logic description-logic ontology knowledge-representation semantic-web linked-data RDF SPARQL OWL knowledge-graph entity-linking relation-extraction event-extraction coreference-resolution named-entity-recognition part-of-speech-tagging dependency-parsing constituency-parsing semantic-parsing natural-language-inference textual-entailment natural-language-generation machine-translation summarization question-answering dialogue-system conversational-AI chatbot virtual-assistant speech-recognition speech-synthesis speaker-recognition emotion-recognition sentiment-analysis opinion-mining aspect-based-sentiment-analysis target-dependent-sentiment-analysis stance-detection rumor-detection fake-news-detection fact-checking claim-verification evidence-retrieval argument-mining persuasion-detection deception-detection personality-prediction mental-health-detection suicide-prevention crisis-intervention social-media-mining network-analysis community-detection influence-maximization information-diffusion cascade-modeling epidemic-modeling SEIR-model SIR-model agent-based-epidemic-model compartmental-model network-epidemic-model metapopulation-epidemic-model spatial-epidemic-model stochastic-epidemic-model branching-process renewal-equation generation-interval serial-interval incubation-period latent-period infectious-period basic-reproduction-number effective-reproduction-number herd-immunity-threshold vaccination-coverage vaccine-efficacy vaccine-effectiveness breakthrough-infection waning-immunity immune-escape variant-of-concern variant-of-interest variant-under-monitoring phylogenetic-analysis molecular-clock-analysis phylodynamics viral-evolution antigenic-drift antigenic-shift zoonotic-spillover reservoir-host intermediate-host amplifying-host dead-end-host vector-borne-disease mosquito-borne-disease tick-borne-disease flea-borne-disease lice-borne-disease midge-borne-disease sandfly-borne-disease tsetse-borne-disease blackfly-borne-disease triatomine-borne-disease kissing-bug Chagas-disease trypanosomiasis leishmaniasis malaria dengue yellow-fever Zika-virus chikungunya West-Nile-virus Japanese-encephalitis tick-borne-encephalitis Lyme-disease Rocky-Mountain-spotted-fever ehlichiosis anaplasmosis babesiosis tularemia plague murine-typhus epidemic-typhus endemic-typhus scrub-typhus Q-fever trench-fever bartonellosis cat-scratch-disease rat-bite-fever leptospirosis brucellosis anthrax melioidosis glanders tularaemia francisellosis coxiellosis rickettsiosis orientia-infection ehrlichiosis neoehrlichiosis neorickettsiosis anaplasmataceae rickettsiaceae coxiellaceae bartonellaceae brucellaceae francisellaceae pasteurellaceae legionellaceae chlamydiaceae mycoplasmataceae ureaplasmataceae spiroplasmataceae achaenoplasmataceae entomoplasmataceae haloplasmataceae tenericutes mollicutes bacteroidetes fusobacteria spirochaetes synergistetes deferribacteres thermotogae aquificae caldiserica caldithrix deferrisoma desulfurobacterium thermovibrio thermocrinis hydrogenobacter sulfurihydrogenibium persephonella hydrogenobaculum hydrothermus thermothrix sulfuricurvum sulfurimonas sulfurovum thiovulum thiomicrospira pseudomonas vibrio aeromonas photobacterium aliivibrio gracilibacillus halobacillus halomonas chromohalobacter cobetia salinicola kushneria halotalea halovibrio salinivibrio thioalkalivibrio thiohalophilus ectothiorhodospira halorhodospira thiorhodospira halochromatium thioflavicoccus marichromatium thiocapsa lamprocystis thiodictyon thiopedia amoebobacter thiocystis chromatium allo-chromatium thermochromatium rhabdochromatium thiobaca thiohalocapsa halocapsa thiolamprovum thioploca thiomargarita beggiatoa methylococcus methylomonas methylomicrobium methylosinus methylocystis methylocella methylocapsa methylacidiphilum verrucomicrobia planctomycetes chlamydiae lentisphaerae kiritimatiellaeota calditrichaeota fervidibacteria
Tortopus
Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.
Ephemeroptera Polymitarcyidae burrowing-mayflies Neotropical clay-bank-habitat U-shaped-tunnels mandibular-tusks Tortopsis Campsurinae Panamerican-distribution cladistic-revision synapomorphy parastyli-receptors flattened-penes furcasternal-protuberances subapical-tubercles limnological-sampling-bias nymph-morphology adult-morphology type-species-T.-igaranus new-species-T.-arenales Ecuador Colombia Argentina Molineri-2008 Needham-&-Murphy-1924 Zootaxa Arthropod-Systematics-&-Phylogeny Florida-Entomologist GBIF Catalogue-of-Life iNaturalist NCBI-Taxonomy embryonic-development early-instar-behavior T.-harrisi T.-bellus T.-circumfluus T.-zottai T.-igaranus T.-incertus T.-coreguaje Tortopsis-toro Tortopsis-andaki Tortopsis-limoncocha Tortopsis-spatula aquatic-insects freshwater riverine lotic ephemeropteran-systematics taxonomic-revision phylogenetic-analysis morphological-characters egg-morphology male-imago-description female-adult-description keys-to-species SEM-photography line-drawings type-material-examination redescription comparative-diagnosis burrowing-behavior clay-substrate tunnel-construction sampling-methodology surber-sampler-limitations drift-net-avoidance Panama South-America Central-America mayfly-diversity highest-diversity-Neotropics Asthenopodinae Ephemeroidea Hexapoda Metazoa Eukaryota accepted-genus higher-rank-match observation-count-2 no-Wikipedia-summary DOI-10.11646/zootaxa.2481.1.1 DOI-10.11646/zootaxa.2436.1.4 DOI-10.3897/asp.79.e62735 DOI-10.2307/3493493 Scott-et-al.-1959 Navás Ulmer Traver Lugo-Ortiz-&-McCafferty Domínguez Banks McDunnough Pictet Needham-&-Murphy freshwater-ecology aquatic-entomology insect-development nymphal-stages imago-description alate-adults wing-venation R-sector gonopore-structure parastyli-length pedestal-morphology lateral-view-curvature mandibular-morphology tusk-tubercles mesosternal-structure abdominal-sternum-notch furcasternum protuberance-configuration receptor-socket-orientation IR-vein R2-vein claw-like-structure gonopore-association penis-base-separation female-fore-wing additional-veins comparative-morphology type-specimen holotype allotype paratype neotropical-species new-species-description species-key identification-key male-adult-key female-adult-key nymph-key generic-key Polymitarcyidae-genera systematics phylogeny cladistics monophyly sister-group character-state transformation-series parsimony-analysis morphological-data-set external-morphology egg-chorion micropyle egg-burster hatching emergence behavioral-study early-instar embryology oviposition substrate-preference habitat-specificity endemic regional-diversity conservation biomonitoring water-quality-indicator macroinvertebrate benthic infaunal interstitial hyporheic river-bank stream-bank erosion sediment particle-size clay-content bank-stability root-mat vegetation-association flow-regime current-velocity depth width stream-order catchment basin ecoregion biogeography dispersal colonization vicariance speciation adaptive-radiation morphological-divergence convergent-evolution homoplasy character-conflict bootstrap-support Bremer-support consensus-tree strict-consensus majority-rule taxonomic-stability nomenclatural-act original-description subsequent-designation type-locality type-depository museum-collection specimen-preservation alcohol-preservation slide-mounting SEM-preparation imaging-techniques scientific-illustration photography measurement morphometrics ratios proportions allometry sexual-dimorphism geographic-variation intraspecific-variation interspecific-variation diagnostic-characters identification-confidence taxonomic-expertise literature-review bibliographic-search database-query online-repository open-access paywall abstract-only full-text-unavailable citation-analysis reference-list source-quality evidence-strength knowledge-gap research-need future-study collaboration field-work collection-trip sampling-effort specimen-availability type-examination revision-scope taxonomic-concept circumscription inclusion exclusion species-group species-complex cryptic-species morphospecies phylogenetic-species evolutionary-significant-unit operational-taxonomic-unit DNA-barcoding molecular-systematics integrative-taxonomy traditional-taxonomy alpha-taxonomy beta-taxonomy gamma-taxonomy classification-hierarchy Linnaean-system rank-based-nomenclature phylogenetic-nomenclature clade-name crown-group stem-group total-evidence congruence incongruence data-partition character-weighting implied-weighting equal-weighting ordered-characters unordered-characters multistate-characters binary-characters continuous-characters meristic-characters qualitative-characters quantitative-characters morphological-matrix data-matrix character-matrix taxon-sampling character-sampling missing-data inapplicable-data polymorphism intraspecific-polymorphism fixed-difference diagnostic-difference unique-derived-character autapomorphy shared-derived-character symplesiomorphy convergent-character parallel-evolution reversal character-optimization ACCTRAN DELTRAN unambiguous-optimization ambiguous-optimization most-parsimonious-tree tree-length consistency-index retention-index rescaled-consistency-index homoplasy-index character-fit character-congruence character-support branch-support node-support clade-stability taxonomic-stability-index phylogenetic-signal noise saturation substitution-model rate-heterogeneity among-site-rate-variation gamma-distribution invariant-sites model-selection Akaike-information-criterion Bayesian-information-criterion likelihood-ratio-test Bayesian-inference Markov-chain-Monte-Carlo posterior-probability prior-probability hyperprior convergence effective-sample-size burn-in stationarity trace-plot parameter-estimate credible-interval highest-posterior-density tree-topology tree-shape lineage-through-time diversification-rate extinction-rate speciation-rate net-diversification birth-death-model Yule-model coalescent-model population-size effective-population-size gene-flow migration-rate isolation-by-distance phylogeography historical-biogeography ancestral-area-reconstruction dispersal-vicariance-analysis decisive-weighting event-based-method taxon-area-cladogram area-cladogram biogeographic-homology track-analysis panbiogeography phylogenetic-biogeography ecological-biogeography dispersal-biogeography vicariance-biogeography center-of-origin area-of-endemism refugium Pleistocene glaciation climate-change habitat-fragmentation habitat-loss anthropogenic-impact pollution eutrophication sedimentation dam-construction water-abstraction flow-modification thermal-pollution toxic-substances heavy-metals pesticides urbanization agriculture deforestation land-use-change riparian-degradation channelization dredging mining invasive-species biological-invasion competition predation parasitism mutualism commensalism amensalism facilitation ecosystem-engineer keystone-species flagship-species umbrella-species indicator-species sentinel-species bioindicator ecological-indicator environmental-indicator water-quality-index biotic-index multimetric-index rapid-bioassessment protocol standardization quality-assurance quality-control reference-condition least-disturbed minimally-disturbed best-available type-specific region-specific stream-type river-type functional-feeding-group collector-gatherer collector-filterer scraper shredder predator parasite engulfer piercer burrower sprawler clinger swimmer skater diver surface-dweller benthic-dweller interstitial-dweller hyporheic-dweller phreatic-dweller troglobite stygobite cave-inhabitant spring-inhabitant headwater mid-order high-order lowland montane alpine subtropical tropical temperate boreal arctic antarctic palearctic nearctic afrotropical oriental australian oceanic cosmopolitan widespread restricted relict disjunct-distribution range-extension new-record first-record country-record state-record province-record locality-record elevation-record altitudinal-range latitudinal-range longitudinal-range distribution-pattern range-size range-fill extent-of-occurrence area-of-occupancy IUCN-category threat-status data-deficient not-evaluated least-concern near-threatened vulnerable endangered critically-endangered extinct-in-wild extinct population-trend population-density population-structure age-structure size-structure sex-ratio reproductive-output fecundity fertility mortality survivorship life-table cohort-analysis generation-time doubling-time intrinsic-rate-of-increase finite-rate-of-increase net-reproductive-rate stable-age-distribution stationary-population expanding-population declining-population source-sink-dynamics metapopulation population-viability-analysis minimum-viable-population genetic-diversity heterozygosity allelic-richness private-allele founder-effect bottleneck genetic-drift inbreeding inbreeding-depression outbreeding outbreeding-depression hybridization introgression gene-flow-barrier reproductive-isolation prezygotic-isolation postzygotic-isolation mechanical-isolation temporal-isolation habitat-isolation behavioral-isolation gametic-isolation zygotic-mortality hybrid-inviability hybrid-sterility hybrid-breakdown reinforcement character-displacement ecological-speciation allopatric-speciation sympatric-speciation parapatric-speciation peripatric-speciation centrifugal-speciation divergence-with-gene-flow speciation-continuum anagenesis cladogenesis budding-speciation branching-speciation phyletic-gradualism punctuated-equilibrium stasis trend progression orthogenesis aristogenesis mosaic-evolution co-evolution arms-race Red-Queen-hypothesis evolutionary-stable-strategy frequency-dependent-selection density-dependent-selection r-selection K-selection bet-hedging iteroparity semelparity ploidy-level chromosome-number karyotype genome-size transposable-element gene-duplication neofunctionalization subfunctionalization pseudogene regulatory-evolution cis-regulatory trans-regulatory enhancer promoter silencer insulator transcription-factor signaling-pathway developmental-pathway gene-network epigenetic-modification DNA-methylation histone-modification chromatin-remodeling genomic-imprinting parental-effect maternal-effect paternal-effect cytoplasmic-inheritance mitochondrial-DNA chloroplast-DNA endosymbiont Wolbachia Rickettsia Spiroplasma Cardinium Arsenophonus Hamiltonella Regiella Serratia Buchnera Carsonella Portiera primary-endosymbiont secondary-endosymbiont facultative-symbiont obligate-symbiont mutualistic-symbiont parasitic-symbiont commensal-symbiont pathogen vector disease-transmission zoonosis emerging-infectious-disease biosecurity quarantine eradication control management conservation-biology restoration-ecology habitat-restoration river-restoration stream-restoration floodplain-restoration riparian-restoration wetland-restoration ecosystem-restoration passive-restoration active-restoration assisted-recovery managed-relocation assisted-colonization ex-situ-conservation in-situ-conservation protected-area national-park nature-reserve wildlife-refuge biosphere-reserve world-heritage-site ramsar-site important-bird-area key-biodiversity-area Alliance-for-Zero-Extinction EDGE-species evolutionarily-distinct globally-endangered conservation-prioritization systematic-conservation-planning Marxan Zonation conservation-opportunity conservation-cost conservation-benefit trade-off socioeconomic-factor stakeholder-engagement community-based-conservation adaptive-management monitoring-program long-term-study citizen-science public-participation environmental-education outreach science-communication knowledge-transfer capacity-building institutional-strengthening policy-development legislation regulation enforcement compliance incentive payment-for-ecosystem-services carbon-credit biodiversity-offset mitigation-banking conservation-easement land-trust private-land-conservation conservation-covenant stewardship sustainable-use sustainable-management ecosystem-based-management integrated-water-resources-management basin-management catchment-management landscape-management seascape-management connectivity-conservation corridor stepping-stone buffer-zone matrix-management countryside-biogeography reconciliation-ecology novel-ecosystem designer-ecosystem ecosystem-service provisioning-service regulating-service supporting-service cultural-service food-production water-supply raw-materials genetic-resources medicinal-resources ornamental-resources air-quality-regulation climate-regulation water-regulation erosion-regulation water-purification waste-treatment disease-regulation pest-regulation pollination seed-dispersal habitat-provision nutrient-cycling soil-formation primary-production recreation tourism aesthetic-value spiritual-value educational-value scientific-value inspiration sense-of-place cultural-heritage traditional-knowledge indigenous-knowledge local-knowledge ecosystem-service-valuation economic-valuation contingent-valuation choice-experiment benefit-transfer total-economic-value use-value non-use-value direct-use indirect-use option-value bequest-value existence-value willingness-to-pay willingness-to-accept cost-benefit-analysis cost-effectiveness-analysis multi-criteria-analysis decision-support uncertainty risk-assessment scenario-analysis sensitivity-analysis robustness-analysis adaptive-capacity resilience vulnerability exposure sensitivity coping-capacity transformation social-ecological-system complex-adaptive-system tipping-point regime-shift alternative-stable-state hysteresis path-dependence legacy-effect slow-variable fast-variable panarchy adaptive-cycle exploitation release reorganization foresight anticipatory-governance transformative-change sustainability-science future-earth interdisciplinary transdisciplinary problem-oriented solution-oriented action-oriented knowledge-co-production boundary-work boundary-object boundary-organization science-policy-interface science-society-interface knowledge-brokerage knowledge-exchange knowledge-integration mode-1-science mode-2-science triple-helix quadruple-helix responsible-innovation reflexive-governance adaptive-governance collaborative-governance polycentric-governance network-governance self-organization collective-action common-pool-resource tragedy-of-the-commons social-dilemma institutional-analysis institutional-design Ostrom-design-principles monitoring sanctioning conflict-resolution nested-enterprises proportionality collective-choice constitutional-choice operational-choice collective-action-theory game-theory prisoner's-dilemma chicken-game assurance-game coordination-game evolutionary-game-theory replicator-dynamics ESS Nash-equilibrium Pareto-optimality social-optimum market-failure government-failure voluntary-failure non-market-value public-good common-good club-good private-good rivalry excludability free-rider hold-up-problem moral-hazard adverse-selection information-asymmetry transaction-cost property-right common-property state-property private-property hybrid-property bundle-of-rights usufruct access withdrawal alienation tenure-security land-tenure water-tenure resource-tenure customary-tenure statutory-tenure legal-pluralism de-facto de-jure formal informal legality legitimacy recognition deviance corruption elite-capture social-capital bonding-capital bridging-capital linking-capital trust reciprocity reputation social-network social-embeddedness structural-hole brokerage closure cohesion density centrality betweenness degree eigenvector PageRank small-world scale-free random-network regular-network hierarchical-network modularity community-structure clustering assortativity homophily heterophily preferential-attachment fitness-model network-evolution network-dynamics temporal-network multilayer-network multiplex-network bipartite-network affiliation-network one-mode-network two-mode-network adjacency-matrix edge-list graph-theory vertex edge node link tie connection path cycle tree forest component giant-component percolation phase-transition criticality self-organized-criticality power-law heavy-tail Pareto-distribution lognormal-distribution exponential-distribution Poisson-distribution binomial-distribution normal-distribution Gaussian-distribution statistical-distribution probability-distribution cumulative-distribution probability-density-function cumulative-distribution-function survival-function hazard-function moment mean median mode variance standard-deviation coefficient-of-variation skewness kurtosis quantile percentile decile quartile interquartile-range range minimum maximum outlier robust-statistic parametric-test non-parametric-test hypothesis-testing null-hypothesis alternative-hypothesis significance-level p-value confidence-interval type-I-error type-II-error statistical-power effect-size sample-size sampling-design experimental-design observational-study quasi-experiment natural-experiment randomized-controlled-trial before-after-control-impact BACI impact-assessment environmental-impact-assessment strategic-environmental-assessment cumulative-impact-assessment social-impact-assessment health-impact-assessment life-cycle-assessment carbon-footprint water-footprint ecological-footprint material-flow-analysis substance-flow-analysis energy-analysis exergy-analysis emergy-analysis input-output-analysis environmentally-extended-input-output social-accounting-matrix computable-general-equilibrium agent-based-model system-dynamics-model Bayesian-network structural-equation-model path-analysis factor-analysis principal-component-analysis cluster-analysis discriminant-analysis canonical-correspondence-analysis redundancy-analysis non-metric-multidimensional-scaling ordination classification regression-analysis linear-regression nonlinear-regression generalized-linear-model generalized-additive-model mixed-effects-model multilevel-model hierarchical-model Bayesian-hierarchical-model state-space-model hidden-Markov-model time-series-analysis spatial-analysis geostatistics kriging variogram spatial-autocorrelation Moran's-I Geary's-c Getis-Ord-Gi local-indicator-of-spatial-association LISA hot-spot cold-spot spatial-cluster spatial-outlier edge-effect boundary-effect modifiable-areal-unit-problem MAUP ecological-fallacy atomistic-fallacy cross-scale-inference upscaling downscaling scaling-law allometric-scaling fractal fractal-dimension self-similarity multi-fractal lacunarity roughness texture pattern process pattern-process-relationship neutral-model null-model randomization-test permutation-test bootstrap jackknife cross-validation model-averaging model-comparison information-criterion Bayesian-model-averaging ensemble-modeling consensus-model prediction forecast projection scenario narrative-scenario quantitative-scenario storyline shared-socioeconomic-pathway representative-concentration-pathway emission-scenario climate-scenario impact-scenario adaptation-scenario mitigation-scenario transformation-scenario backcasting forecasting trend-extrapolation expert-judgment Delphi-method participatory-modeling stakeholder-analysis influence-diagram conceptual-model mental-model system-archetype leverage-point intervention-point policy-instrument economic-instrument regulatory-instrument voluntary-instrument information-instrument market-based-instrument command-and-control-instrument subsidy tax charge fee levy deposit-refund tradable-permit emission-trading carbon-tax biodiversity-banking conditional-cash-transfer community-based-natural-resource-management joint-forest-management co-management adaptive-co-management transboundary-management international-regime global-governance multilateral-environmental-agreement convention treaty cartagena-protocol nagoya-protocol Aichi-target sustainable-development-goal SDG post-2020-global-biodiversity-framework Kunming-Montreal-global-biodiversity-framework 30-by-30-target nature-based-solution NbS ecosystem-based-adaptation ecosystem-based-mitigation green-infrastructure blue-infrastructure natural-infrastructure hybrid-infrastructure climate-smart-agriculture climate-smart-landscape climate-smart-development low-carbon-development green-economy circular-economy bioeconomy blue-economy sustainable-consumption sustainable-production life-cycle-thinking cradle-to-cradle industrial-ecology industrial-symbiosis urban-metabolism socio-ecological-metabolism material-intensity energy-intensity carbon-intensity decoupling absolute-decoupling relative-decoupling resource-efficiency eco-efficiency eco-innovation green-technology clean-technology environmental-technology appropriate-technology participatory-technology technology-transfer technology-diffusion innovation-system national-innovation-system sectoral-innovation-system technological-innovation-system socio-technical-system socio-technical-transition multi-level-perspective niche-innovation regime landscape transition-management strategic-niche-management transition-experiment living-lab real-world-laboratory urban-living-lab rural-living-lab sustainability-transition fundamental-change system-innovation directionality reflexivity responsiveness anticipation learning-by-doing doing-by-learning social-learning organizational-learning policy-learning transformative-capacity capacity-development institutional-learning evolutionary-governance institutional-change path-dependency lock-in carbon-lock-in technological-lock-in institutional-lock-in behavioral-lock-in cultural-lock-in unlocking destabilization phase-out divestment stranded-asset just-transition transition-pathway transition-scenario deep-decarbonization net-zero carbon-neutral climate-neutral nature-positive biodiversity-positive restoration-positive regenerative circular doughnut-economics planetary-boundary safe-operating-space environmental-ceiling social-foundation intergenerational-equity intragenerational-equity distributive-justice procedural-justice recognition-justice environmental-justice climate-justice energy-justice water-justice food-justice spatial-justice territorial-justice ecological-justice species-justice intersectionality adaptation mitigation sustainability sustainable-development well-being human-development capability-approach human-security environmental-security climate-security water-security food-security energy-security livelihood-security disaster-risk-reduction climate-change-adaptation climate-change-mitigation loss-and-damage climate-finance green-climate-fund adaptation-fund global-environment-facility biodiversity-finance nature-finance blended-finance impact-investment ESG environmental-social-governance sustainable-finance green-bond social-bond sustainability-bond blue-bond nature-bond biodiversity-credit offset inset nature-based-credit ecosystem-service-credit payment-for-results results-based-payment conditional-payment performance-based-payment outcome-based-payment environmental-service-payment PSA PES REDD REDD+AFOLU LULUCF NAMA NAMC NDC NAP NBSAP national-adaptation-plan national-biodiversity-strategy-and-action-plan nationally-determined-contribution nationally-appropriate-mitigation-action intended-nationally-determined-contribution long-term-low-emission-development-strategy net-zero-target carbon-budget remaining-carbon-budget emission-gap adaptation-gap finance-gap implementation-gap ambition-loop ratchet-mechanism global-stocktake transparency-framework enhanced-transparency-framework common-but-differentiated-responsibilities respective-capabilities climate-equity climate-litigation climate-governance climate-diplomacy climate-negotiation UNFCCC COP CMP CMA SBSTA SBI IPCC IPBES UNCCD CBD CITES Ramsar World-Heritage Man-and-the-Biosphere MAB UNESCO FAO WHO ILO UNDP UNEP UN-Habitat World-Bank IMF WTO OECD GEF GCF AF multilateral-development-bank regional-development-bank national-development-bank development-finance-institution export-credit-agency sovereign-wealth-fund pension-fund insurance-company asset-manager asset-owner corporate-investor retail-investor impact-investor philanthropic-investor development-partner bilateral-donor multilateral-donor private-donor foundation NGO CSO civil-society-organization non-governmental-organization community-based-organization indigenous-peoples-organization women's-organization youth-organization faith-based-organization research-organization think-tank university academic-institution knowledge-institution capacity-building-institution training-institution extension-service advisory-service consulting-firm engineering-firm technology-provider equipment-supplier service-provider contractor operator maintainer utility municipality local-government provincial-government national-government federal-government ministry department agency authority commission board council committee parliament congress legislature executive judiciary media journalist reporter editor publisher broadcaster platform social-media influencer celebrity activist advocate campaigner mobilizer organizer networker connector bridge-builder boundary-spanner knowledge-broker science-communicator extension-agent development-practitioner policy-maker decision-maker planner manager administrator leader champion entrepreneur innovator intrapreneur social-entrepreneur systems-entrepreneur institutional-entrepreneur policy-entrepreneur political-entrepreneur business-model business-model-innovation social-business inclusive-business base-of-pyramid fortune-at-bottom-of-pyramid shared-value creating-shared-value stakeholder-capitalism purpose-driven-business B-Corp benefit-corporation social-enterprise cooperative mutual association collective commons commoning peer-production open-source creative-commons knowledge-commons digital-commons data-commons genetic-commons seed-commons water-commons fisheries-commons forest-commons pasture-commons land-commons urban-commons atmospheric-commons global-commons common-heritage-of-mankind common-concern-of-humankind restorative-justice multispecies-justice more-than-human anthropocene capitalocene plantationocene chthulucene symbiocene good-anthropocene great-acceleration great-transformation great-decoupling great-convergence great-divergence great-fragmentation great-integration great-transition great-work long-now deep-time futures-thinking future-studies strategic-foresight scenario-planning trend-analysis weak-signal wild-card black-swan grey-rhino blue-sky green-swan systemic-change structural-change cultural-change behavioral-change individual-change collective-change social-change organizational-change technological-change economic-change political-change ecological-change planetary-change earth-system-change biodiversity-change ocean-change cryosphere-change atmospheric-change biogeochemical-change nitrogen-cycle-change phosphorus-cycle-change carbon-cycle-change water-cycle-change hydrological-change soil-change vegetation-change ecosystem-change landscape-change seascape-change biosphere-change noosphere-change technosphere-change anthroposphere-change sociosphere-change econosphere-change politosphere-change culturosphere-change semiosphere-change infosphere-change digisphere-change cybersphere-change urban-sphere-change agricultural-sphere-change industrial-sphere-change extractive-sphere-change manufacturing-sphere-change service-sphere-change financial-sphere-change knowledge-sphere-change innovation-sphere-change creativity-sphere-change culture-sphere-change identity-sphere-change meaning-sphere-change value-sphere-change purpose-sphere-change vision-sphere-change mission-sphere-change strategy-sphere-change tactic-sphere-change operation-sphere-change action-sphere-change practice-sphere-change habit-sphere-change routine-sphere-change ritual-sphere-change tradition-sphere-change custom-sphere-change convention-sphere-change norm-sphere-change rule-sphere-change law-sphere-change regulation-sphere-change standard-sphere-change protocol-sphere-change procedure-sphere-change process-sphere-change system-sphere-change structure-sphere-change function-sphere-change pattern-sphere-change dynamic-sphere-change state-sphere-change phase-sphere-change regime-sphere-change equilibrium-sphere-change stability-sphere-change instability-sphere-change chaos-sphere-change order-sphere-change complexity-sphere-change simplicity-sphere-change diversity-sphere-change uniformity-sphere-change heterogeneity-sphere-change homogeneity-sphere-change variation-sphere-change invariance-sphere-change change-sphere-change continuity-sphere-change discontinuity-sphere-change emergence-sphere-change submergence-sphere-change evolution-sphere-change devolution-sphere-change revolution-sphere-change involution-sphere-change convolution-sphere-change resolution-sphere-change dissolution-sphere-change absolution-sphere-change solution-sphere-change