Natural-dispersal

Guides

• Leptoglossus

  leaf-footed bugs

  Leptoglossus is a genus of true bugs in the leaf-footed bug family Coreidae, tribe Anisoscelini. Species are characterized by leaflike dilations of the hind tibia, a diagnostic trait of the genus. The genus is distributed throughout the Americas, with some introduced populations in Europe and Asia. Several species are economically significant agricultural pests, notably L. occidentalis, which has become invasive in multiple continents.

  Coreidaeleaf-footed-bugagricultural-pestinvasive-speciessymbiosissexual-dimorphismconifer-pestnuisance-pestBurkholderiatachinid-parasitoidpheromone-communicationbuilding-invadermisidentificationTriatoma-look-alikegradual-metamorphosisseed-predatorforest-pestornamental-pestplumbing-damagepublic-health-confusionChileEuropeNorth-AmericaSouth-AmericaAsiaTurkeyalmond-pestcitrus-pesttomato-pestcorn-pestoverwintering-aggregationdefensive-secretionpheromone-mediated-parasitismegg-parasitoidbiological-controlecological-niche-modelingclimate-suitabilityrange-expansionintroductionaccidental-dispersalseed-orchard-pestgermination-reductionempty-seed-formationcone-damagelodgepole-pineDouglas-firwestern-conifer-seed-bugL.-occidentalisL.-zonatusL.-phyllopusL.-clypealisL.-australisL.-chilensisL.-oppositusmale-combatfemoral-weaponabdominal-glandpheromone-glandtibial-dilationleaf-like-hind-legtrue-bugHemipteraHeteropteraPentatomomorphaAnisosceliniphytophagousplant-feedingstylet-feedingsalivary-sheathoverwinteringdiapausebuilding-entrystructural-pestnuisance-odorflight-sounddroning-flightaggregation-behaviormale-pheromonefemale-choiceparasitoid-hostbiological-control-agentintegrated-pest-managementmonitoringearly-detectionpreventioninvasion-biologyalien-speciesnon-native-pestglobal-spreadclimate-changeniche-modelingsuitable-habitatestablishment-riskquarantinephytosanitaryforest-healthseed-productionreforestationafforestationconifer-forestrypine-seedseed-viabilityeconomic-impactcrop-lossyield-reductionkernel-damagefruit-damagenut-damagevector-of-diseaseChagas-diseasekissing-bugpublic-alarmmisinformationeducationidentification-toolshealth-system-burdenpesticide-overuseurban-ecologydomiciliaryperidomiciliaryAndean-regionPatagoniaMediterraneantemperatesubtropicaltropicalagroecosystemnatural-enemypredatorparasitepathogensymbiontgut-microbiomesoil-ingestionfitness-benefitreproductive-successsperm-morphologytesticular-morphologyaccessory-glandpheromone-blendspecies-specific-odorcherry-scentvanilla-scentcinnamon-scentrose-scentolfactory-communicationmate-locationmate-recognitionsexual-selectionmale-male-competitionweapon-morphologyallometrydevelopmental-plasticitynymphal-instaregg-barrelegg-arrangementlinear-ovipositionleaf-surfaceplant-tissuestylet-penetrationenzymatic-digestionfluid-feedingphloem-feedingxylem-feedingseed-feedingcone-feedingfruit-feedingkernel-feedingnut-feedingcrop-feedinghost-plant-rangepolyphagyoligophagyspecialistgeneralistpest-statusdamage-thresholdeconomic-injury-levelmanagement-strategycultural-controlphysical-controlchemical-controlresistancetolerancehost-plant-resistancemonitoring-traplight-trappopulation-densitydistribution-mapspread-ratecolonizationestablishmentpopulation-dynamicslife-tabledevelopmental-ratethermal-requirementsdegree-daysphenologyvoltinismunivoltinebivoltinemultivoltinegeneration-timeoverwintering-sitehibernaculumshelter-seekingcold-hardinessfreeze-tolerancesupercoolingwater-relationsdesiccation-resistancestarvation-resistancedispersal-abilityflight-capacitywalking-behaviorclimbing-behavioraggregation-pheromonealarm-pheromonestink-glandmetathoracic-glanddorsoabdominal-glandventral-abdominal-glandsexual-glandmorphological-defensechemical-defensemechanical-defenseautotomyleg-losspredation-riskparasitism-riskparasitoid-riskpathogen-riskcompetitionintraspecificinterspecificresource-competitionmating-competitionsperm-competitioncryptic-female-choicereproductive-isolationspeciationphylogenysystematicstaxonomymorphologyanatomyhistologyultrastructurespermatogenesisspermatozoasperm-lengthnuclear-lengthcyst-productionfollicle-numbertestis-structurereproductive-anatomygenitaliamating-behaviorcopulationinseminationovipositionegg-productionfecundityfertilityhatching-successnymphal-survivaladult-longevitysex-ratiooperational-sex-ratiopopulation-sex-ratioeffective-population-sizegenetic-diversitygene-flowpopulation-structurephylogeographybiogeographyhistorical-biogeographyvicariancedispersalrange-shiftrange-contractionaltitudinal-distributionlatitudinal-distributionlongitudinal-distributionisland-distributioncontinental-distributionendemismcosmopolitanismintroduced-rangenative-rangesource-populationfounder-populationinvasion-frontlag-phaseestablishment-phasespread-phaseequilibrium-phaseimpact-assessmentrisk-assessmenthorizon-scanningearly-warningrapid-responseeradicationcontainmentcontrolmanagementadaptationmitigationrestorationconservationbiodiversityecosystem-serviceecosystem-functionfood-webtrophic-levelprimary-consumerherbivorefruit-predatorplant-animal-interactionmutualismantagonismpredationparasitismcommensalismamensalismfacilitationindirect-interactiontrait-mediated-interactiondensity-mediated-interactionbehavioral-ecologyevolutionary-ecologyfunctional-ecologyphysiological-ecologypopulation-ecologycommunity-ecologyecosystem-ecologylandscape-ecologymacroecologyglobal-ecologyapplied-ecologyagricultural-ecologyforest-ecologyconservation-biologyinvasion-ecologyrestoration-ecologypest-managementconservation-biological-controlaugmentative-biological-controlclassical-biological-controlparasitoidnematodefungusbacteriumvirusentomopathogenmicrobial-controlsterile-insect-techniquegenetic-controlmating-disruptionattract-and-killpush-pulltrap-cropborder-croprefugehabitat-managementcultural-practicecrop-rotationtillageirrigationfertilizationpruningharvest-timingsanitationresidue-managementweed-managementcover-cropcompanion-plantingintercroppingagroforestrysilvicultureforest-managementseed-orchard-managementcone-collectionseed-extractionseed-testinggermination-testingseed-qualityseed-vigorempty-seedinsect-damageinsect-injuryfeeding-scarpuncturestylet-sheathsalivary-flangesymptomsigndiagnosissamplingeconomic-thresholddecision-supportexpert-systemmodelingsimulationforecastingclimate-modelniche-modelspecies-distribution-modelhabitat-suitabilityrisk-mappinginvasion-pathwayintroduction-vectortradetransporttourismmigrationnatural-dispersalhuman-mediated-dispersalaccidental-introductionintentional-introductionreleaseescapecultivationornamentalforestryagriculturehorticultureurbanizationglobalizationland-use-changehabitat-fragmentationhabitat-lossdegradationpollutionpesticidefertilizernutrient-enrichmenteutrophicationacidificationwarmingdroughtextreme-eventdisturbancesuccessionrecoveryresiliencestabilityvariabilityuncertaintysustainable-developmentgreen-economycircular-economybioeconomyecosystem-approachone-healthplanetary-healthenvironmental-healthpublic-healthveterinary-healthplant-healthfood-securitynutritionlivelihoodpovertyinequalitygenderindigenous-knowledgetraditional-knowledgelocal-knowledgecitizen-sciencestakeholder-engagementpolicygovernanceregulationlegislationinternational-cooperationcapacity-buildingawarenesscommunicationoutreachextensionadvisory-serviceresearchinnovationtechnology-transferknowledge-exchangenetworkingpartnershipcollaborationevaluationevidence-based-policyscience-policy-interfacediplomacyadvocacyactivismsocial-movementenvironmental-justiceenvironmental-ethicsenvironmental-philosophyenvironmental-humanitiesenvironmental-historyenvironmental-literatureenvironmental-artenvironmental-educationenvironmental-communicationscience-communicationrisk-communicationcrisis-communicationhealth-communicationbehavior-changepro-environmental-behaviorsustainable-behaviorconsumptionproductionwasterecyclingreusereductionefficiencyrenewablecleangreenbluenaturalorganicagroecologypermacultureregenerativerestorativehealingtransformativetransdisciplinaryinterdisciplinarymultidisciplinarycross-disciplinarydisciplinarydisciplinary-integrationknowledge-integrationsystem-thinkingcomplexityemergencenon-linearityfeedbacktipping-pointregime-shiftalternative-stable-stateresilience-thinkingadaptive-managementadaptive-governancelearningreflectionparticipationinclusionequityjusticefairnessethicsvaluesnormscultureidentityplacesense-of-placeattachmentwell-beingquality-of-lifehappinessflourishingthrivingsustainabilitysustainabledevelopmentgrowthdegrowthpost-growthsteady-statecirculardoughnut-economicsplanetary-boundariessafe-operating-spacetipping-elementscritical-transitionearly-warning-signalresilience-indicatorsustainability-indicatorbenchmarktargetgoalcommitmentpledgeagreementtreatyconventionprotocolframeworkstrategyplanprogramprojectinitiativecampaignmovementcoalitionalliancenetworkplatformhubcenterinstituteorganizationassociationsocietyunionfederationconfederationfoundationtrustfundbankfacilitymechanisminstrumenttoolmethodapproachtechniqueprocedureprocesssystemstructureinstitutionadministrationoperationservicedeliveryprovisionsupplydemandmarketeconomyfinanceinvestmentfundingfinancingresourcecapitalassetliabilityincomeexpenditurerevenuecostbenefitprofitlossreturnyieldoutcomeoutputinputthroughputproductivityeffectivenessperformancequalitystandardcriterionindicatormetricmeasureassessmentappraisalreviewauditverificationvalidationcertificationaccreditationrecognitionawardprizehonordistinctionexcellencebest-practicelesson-learnedexperienceexpertisecompetencecapacitycapabilityskillknowledgeinformationdataevidencesciencediscoveryinventioncreationdesigntestingpilotingscalingmainstreaminguptakeadoptiondiffusiondisseminationpublicationreportingdocumentationarchivingpreservationrehabilitationreconstructionreplicationduplicationbackupsecuritysafetyprotectionsafeguardinginsurancerisk-managementemergency-preparednessresponsereliefhumanitarianpeaceprosperityhealthemploymenthousinginfrastructureenergywaterfoodfisheryaquacultureminingmanufacturingindustrycommercerecreationheritagenatureecosystemenvironmentclimateatmosphereoceanfreshwaterlandsoilmineralbiotaflorafaunamicroorganismfungiplantanimalvertebrateinvertebratearthropodinsectbugHemipteranHeteropteranpentatomomorphancoreoidcoreidsquash-bugboxelder-bugstink-bugshield-bugplant-bugmiridlygaeidseed-bugbroad-headed-bugalydidreduviidassassin-bugtriatominebed-bugcimicidwater-bugnepomorphangerromorphanleptopodomorphanenicocephalomorphandipsocoromorphanceratocomboidschizopteroidpeloridioidcoleorrhynchanmoss-bugarchaeorrhynchanfulgoromorphancicadomorphanmembracoidtreehopperleafhopperplanthopperpsyllidjumping-plant-lousewhiteflyaleyrodidscale-insectcoccoidmealybugaphidadelgidphylloxeransternorrhynchanthysanopteranthripspsocopteranbarklousebooklousephthirapteranlousesucking-lousechewing-lousemallophagananoplurandermapteranearwigblattodeancockroachtermiteisopteranmantodeanmantidphasmidstick-insectleaf-insectorthopterangrasshopperlocustkatydidcricketmole-cricketpygmy-mole-cricketcamel-cricketcave-cricketwetaensiferancaeliferangryllotalpidmyrmecophilidtettigoniidgryllidacrididpamphagidpneumoridlentulidtristirideumastacidproscopiidtridactylidtetrigidgrouse-locustpygmy-grasshopperplecopteranstoneflyembiopteranwebspinnerzorapteranangel-insectdictyopteran

• Trichonephila clavata

  Jorō spider, Joro Spider, Parachute spider

  Trichonephila clavata, commonly known as the Jorō spider, is a large orb-weaving spider native to East Asia that has become established as an invasive species in the southeastern United States since approximately 2010. First confirmed in Georgia in 2014, it has expanded rapidly across multiple states through a combination of ballooning dispersal and human-mediated transport. The species is notable for its substantial size, striking coloration, and extensive golden webs, but poses minimal risk to humans due to small fangs and docile behavior. Its physiological adaptations—including higher metabolic rate, faster heart rate, and greater cold tolerance than its congener Trichonephila clavipes—suggest potential for continued northward range expansion.

  invasive-speciesorb-weaverballooningurban-wildlifebiological-controlcold-tolerancestartle-responsecannibalismkleptoparasitismJapanese-folkloreNephilidaeTrichonephilaspider-silkrange-expansionphysiological-adaptationurban-ecologyagricultural-pest-predatoriNaturalistGeorgiaMarylandEast-AsiaUnited-Statesspider-bitevenomdocilenon-aggressiveweb-architectureegg-sacoverwinteringsexual-dimorphismmetabolic-rateheart-ratefreeze-toleranceroad-ecologyprey-capturecompetitionnative-spider-impactspublic-perceptionmedia-sensationalismmanagementpesticide-alternativesbroom-removalornamentalstructural-pestornithologycardinalweb-strengthtrophic-interactionsDNA-metabarcodingdiet-analysisspecies-distribution-modelphysiological-entomologybehavioral-ecologyarachnologyballooning-dispersalhuman-mediated-dispersalhitchhikercargo-stowawayport-of-entryclimate-changewarmingnorthward-expansionDMVHoward-CountyElkridgeIlchesterWest-Elkridgebeech-forestsecondary-forestsuburbanroadsidetelephone-polestreetlampforest-edgedisturbance-toleranceprey-detectionvibrational-noiseauditory-disturbancemotionlessfreezingshynessHannibal-Lecterstink-buglanternflyevolutionary-historyancient-associationecosystem-servicebiocontrolnatural-enemyinvasive-pestbrown-marmorated-stink-bugspotted-lanternflyHalyomorpha-halysLycorma-delicatulacompetitive-exclusiondominant-orb-weaverabundantpopulation-growthrapid-spreadrange-span120,000-km2physiological-plasticitymetabolismtwice-as-high77%-higher-heart-rate74%-freeze-survivaldevelopment-rateseasonal-windowlife-cycle-completionthermal-limitationcongener-comparisonnatural-dispersalhuman-transportiNaturalist-recordcitizen-scienceconfirmed-sightingbreeding-populationmale-and-femalegravid-femaleegg-casestowawaycargo-containerport-introductionlocal-portvehicle-transporthitchhikingaccelerated-arrivalseveral-yearsnatural-meansprognosticationpredictionmodelsuitable-habitatabundant-suitable-habitateastern-North-Americawestern-North-Americapotential-rangeinvasive-rangepotential-impactenvironmental-impactecological-impactecosystem-impactmonitorcautiousevidence-basedreasonable-journalismsensationalismexaggerationuncritical-acceptanceharmlesscarefully-monitorednext-stepsresearch-opportunitycall-for-researchknowledge-gaplittle-knownnovel-ecosystemspreadingrapidly-expandinginvasive-predatorinvasive-orb-weavernon-nativeintroducedestablishedcolonizedthrivingdoing-finehappymerely-20-minutes-awayroad-tripvisitaccess-spreadbeech-leaf-diseaseBLDstate-forestremarkable-colonyamongst-beech-treesconfirmed-prognosticationDavis-and-Frickescape-relative-warmthexpand-range-northwardeastern-seaboardmerely-20-minutes-away-from-homeresist-opportunityamazing-predatorstales-of-arrivalprevious-episodes20222024reduce-angstlarge-non-native-spiderestablishing-in-DMVfacts-presentedpast-episodevenomous-terrible-painfulNahexpert-Rick-Hoebekerisks-smallpuny-fangsunlikely-pierce-skinvideocompletely-non-aggressiveeastern-Howard-Countyhaphazard-webslittered-with-remainsformer-victimsleavesshed-exoskeletonsmuch-larger-femaledwarfs-matepositioned-just-abovestrand-of-silkspinneretsunderside-of-abdomennear-red-moundrelative-to-handlarge-and-docilewait-and-seewhat-Jorō-means-to-ecosystemshelp-other-spidersput-a-beat-down-on-invasive-pestsstink-bugsspotted-lanternfliespassive-huntersenormous-webslarger-than-a-meter-in-diametercapture-prey-snared-in-silkarachnophobes-scaryarachnophiles-beautifulimportant-ecosystem-servicescrop-pestsnative-rangeAsiahaving-an-old-friend-over-for-dinnerreunitefirst-timelarge-spidersjuicy-prey-itemsfeathered-and-non-feathered-reptilesposes-no-known-threatdifficult-to-predict-impactnon-native-speciesexperts-suggestbeyond-scary-miengive-indigenous-large-orb-weavers-run-for-their-moneyblack-and-yellow-garden-spidermarbled-orb-weaverspotted-orb-weaverother-parts-of-worldmost-abundant-and-dominant-orb-weaverwhat-will-it-meanonly-time-will-tellfinal-tidbitJorō-shapeshifterJorō-gumobeautiful-womanseduces-menbinds-with-silkdevoursbad-dateacknowledgementsRick-Hoebekeidentifying-JorōinsightsDavid-CoyleBob-Bellingergreat-imagesknowledgestudiesVeni-vidi-viciNelsen-et-al.Physiological-evaluationNephila-clavata-newly-recordedHoebeke-et-al.Life-Cycle-Habitat-and-VariationMooreStartle-responsesDavis-and-AneraoHow-Urban-Tolerant-Are-TheyDavis-et-al.inspirationdetailsstarsDave-ClementMiri-TalabacMaddie-Potterhooked-up-with-colonydifferentiateresources.ipmcenters.orglinkJorō-guruDavid-Coyle's-takeYouTubecalm-your-fearsJournal-of-Economic-EntomologyJournal-of-Medical-Entomologychemical-management-strategieseffective-management-methodsspider-control-productssafe-and-effectiveunlikely-to-bitegenerally-harmlessbig-spiderscolorful-spidersinvasive-spidersin-your-face-spiderswebs-on-homeslandscape-plantsstructuresmiddle-of-where-people-live-work-and-playnorthern-Georgia-2014presumed-arrived-few-years-prior15-yearsincreased-dramaticallymuch-of-Southeastdisjunct-populationsnortheastern-U.S.firestorm-of-interestrapidly-expanding-rangesoutheastern-statesmany-countiesCarolinasTennesseeOklahomamore-than-a-centuryparts-of-Floridararely-Pennsylvaniatropicsthermal-limitationshigher-metabolismfaster-heart-ratebetter-ability-to-tolerate-freezingcomplete-life-cycle-rapidlychilly-temperaturessuite-of-adaptationsexpand-northwardwarm-loving-cousinfindings-support-potentialmuch-yet-to-be-learnedsuccessfully-survive-northern-wintersnation-and-world-warmsouthern-species-expandhigher-latitudes-and-altitudesrapidly-range-expansion-will-occurtypical-mode-of-dispersalaerial-dispersalballooning-on-strands-of-silkCharlotte's-babiesCharlotte's-Webspectacular-monikerparachute-spiderrain-down-from-airplaneslong-distance-transithuman-assistancegood-hitchhikersinseminated-gravid-femaleegg-case-stowawayarrival-in-DMVseveral-years-by-natural-meansnew-introduction-at-local-porthuman-assist-in-vehiclesaccelerate-arrivalterrible-and-painfulsmall-fangsvisited-cousingolden-silk-spiderCosta-Ricaexceeding-meter-diameterindigenous-large-orb-weaversrun-for-their-moneyminimal-direct-impactfirestormAndrew-DavisBenjamin-FrickUniversity-of-Georgiapast-weeknative-to-eastern-AsiaJapanChinaKoreaTaiwanknown-in-Georgia-since-2013spreading-rapidlyjoined-its-cousinTrichonephila-clavipesestablished-over-160-yearsbetter-tolerate-freezingsurvive-northern-wintersrapidly-range-expansiontypical-modelong-distancenew-introductionhuman-assistarachnophobesarachnophilesbrown-marmorated-stink-bugsdifficult-to-predictscary-mienindigenous-orb-weaversMary-NouriSarah-MorganWAMUrelatively-harmlessspider-seasonfallleaves-changing-colorcool-crisp-autumn-airpumpkin-spiceextension-entomologistSoutheastnot-just-spiders20-feet-widehomespeople-live-work-and-play15-years-or-soinflux-of-callspublicentomology-extension-programsmedia-frenzytwo-questionsget-rid-of-Jorō-spidersare-Jorō-spiders-dangerousClemson-UniversitySouthern-Adventist-UniversityWashington-CollegeUnion-Collegetwo-new-studiestest-methods-for-eliminatinghow-likely-to-biteharmscoured-internetrecommended-treatment-optionsspider-specific-management-recommendationsmedically-relevantwidow-spidersbrown-reclusesdo-not-enter-homesrelatively-newrecommended-productslabelled-spider-control-productstested-productsseveral-products-eliminatenot-legal-pesticidesmachine-lubricantnot-recommendedcommercially-availablestrongly-encouragelabelled-productseasiest-and-cheapestwithout-chemicalsbroom-or-stickwalked-into-spider-webrode-biketook-spider-to-facenot-great-feelingwonder-if-biteygive-bitten-superpowersunique-studiesevaluated-behaviorsreact-to-peoplerather-drop-off-webhang-around-and-bitereally-provokevolunteered-to-get-bittenyes-you-read-correctlyakin-to-mosquito-bitelittle-rednesslittle-swellinglargely-gone-next-dayno-acquired-superpowerstake-home-messagetwofoldsimplest-and-most-effectivewithout-pesticidesfeel-you-need-pesticidesproduct-labeled-for-spider-controltry-not-to-freak-outdoesn't-want-to-be-on-youodds-of-getting-bitten-extremely-lowenjoy-glorious-fall-weatherdrink-pumpkin-spiced-beveragewatch-where-you-walkKeep-calm-and-carry-onbites-unlikelycause-minimal-discomfortassociate-professorDepartment-of-Forestry-and-Environmental-Conservationsocialsemailextensionfeaturedpest-managementEntomology-Todaylatest-postsemail-subscriptionMarch-2022wonderedmake-its-way-to-DMVSeptember-2022two-observationsthree-years40-sightingshappy-and-doing-just-fineseveral-Howard-County-locationsweek-or-so-agoteam-of-scientistsUniversitymissionkilling-ultra-valuable-beech-treesthriving-amongst-beech-treesvisit-amazing-predatorsmature-femalesboth-Trichonephila-speciesthree-locally-common-orb-weaving-speciestime-spent-immobilemild-disturbancebrief-puff-of-airair-puff-response-datafive-other-North-American-speciespublished-literature453-observationsfreezing-behavior10-spider-speciesremained-immobile-under-a-minuteboth-Trichonephila-spidersover-an-hourunprecedentedshyest-ever-documentedtolerate-urban-environmentsremaining-motionlessfleeingnonsexual-cannibalismmating-activityfemales-cannibalize-malesfood-availabilityterritorial-aggressionSoutheastern-United-Statesexpanding-rangeshybehavioral-reactionsdescriptive-observationsphoto-documentationanecdotal-observationscontrolled-pairingslabnatural-websfieldone-female-biting-and-killing-anothershort-fightsimilar-sizecontainer25-trialsfights-ensued-40%different-sizes27-trialsfights-happened-18%larger-females-not-always-aggressor52-lab-trialssix-bouts9%direct-killingfield-trialsempty-web14-trialsone-fight7%aggressor-killing-and-wrappingsilkaway-from-any-webnot-necessarily-territorialityintraspecific-aggressionprovoked-or-stresseddeserves-more-studynewly-invasivebiologyphysiologynew-rangeclosely-related-speciessuccessfully-established160-yearsinvestigationcompareonline-recordsiNaturalist.orgseasonal-distributionstimingfemalesphysiological-traitsenvironmental-tolerancemetabolic-ratescardiac-functioningcold-exposuresurvivalbrief-below-freezing-temperatureshorter-seasoncomplete-lifecyclenarrow-periodsuitable-weatherinherently-higher-metabolismlow-temperaturesurvive-better74%-comp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