Alternative-stable-state

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Acmaeodera parkeri
Acmaeodera parkeri is a species of metallic wood-boring beetle in the family Buprestidae, described by Cazier in 1940. The species occurs in Central America and North America, with records from the southwestern United States and Mexico. Adults have been observed visiting flowers of several plant species, including Allionia incarnata (creeping four o'clock), Talinum whitei (White's flameflower), and Evolvulus arizonicus (Arizona blue-eyes).
Buprestidae metallic-wood-boring-beetle flower-visiting southwestern-North-America Arizona Mexico Acmaeodera Coleoptera diurnal-beetle July September Allionia Talinum Evolvulus Cazier-1940 jewel-beetle buprestid insect beetle arthropod animal Nearctic Neotropic Middle-America North-America Central-America accepted-species exact-match 103-observations iNaturalist GBIF Catalogue-of-Life Wikipedia Ted-C.-MacRae Beetles-In-The-Bush field-collecting Arizona-blue-eyes White's-flameflower creeping-four-o'clock flower-host sky-islands Santa-Rita-Mountains Huachuca-Mountains Box-Canyon Copper-Canyon Gardner-Canyon Montosa-Canyon ruby-road Atascosa-Mountains Madera-Canyon Florida-Canyon summer-activity fall-activity July-collecting September-collecting beating-sheet sweeping visual-detection diurnal-activity flower-perching rare-beetle uncommon-species target-species buprestophile Cazier 1940 original-description type-locality species-description taxonomic-authority canonical-name scientific-name rank-species status-accepted match-type-exact kingdom-Animalia phylum-Arthropoda class-Insecta order-Coleoptera family-Buprestidae genus-Acmaeodera species-parkeri specific-epithet-parkeri classification-Eukaryota-Animalia-Arthropoda-Hexapoda-Insecta-Coleoptera-Polyphaga-Elateriformia-Buprestoidea-Buprestidae-Acmaeodera-Acmaeodera-parkeri distribution-Nearctic-Neotropic Middle-America-present North-America-present 103-iNaturalist-observations Wikipedia-summary found-in-Central-America-and-North-America field-notes collecting-trip Arizona-2021 Arizona-2019 Arizona-2018 New-Mexico-2018 Texas-2018 Jeff-Huether Art-Evans Margarethe-Brummermann Norm-Woodley Steve-Lingafelter Rob-Velten beetle-collecting insect-collecting entomology coleopterology buprestology jewel-beetles metallic-beetles flower-beetles pollen-feeders nectar-feeders floral-hosts host-plant-associations plant-insect-interactions ecology natural-history biodiversity conservation taxonomy systematics nomenclature authorship-Cazier-1940 original-combination current-combination valid-name accepted-name species-level-taxon no-subspecies no-synonyms-listed no-common-names no-vernacular-names specialist-pollinator generalist-flower-visitor montane-species desert-species riparian-species oak-woodland mesquite-scrub grassland chaparral sky-island-ecosystem Madrean-sky-islands Chihuahuan-Desert Sonoran-Desert Arizona-transition-zone Mexican-Plateau southeastern-Arizona southwestern-New-Mexico western-Texas northern-Mexico Mexican-states US-states habitat-generalist habitat-specialist elevational-range seasonal-phenology adult-emergence reproductive-biology larval-biology-unknown wood-boring-larvae xylophagous phytophagous herbivorous anthophilous flower-loving diurnal crepuscular nocturnal-unknown flight-capability active-flier visual-acuity color-vision metallic-coloration structural-color elytral-sculpture pronotal-shape antennal-serration tarsal-formula larval-host-unknown possibly-oak possibly-mesquite possibly-legume possibly-herbaceous field-identification-difficult museum-identification-required genitalia-examination type-specimen-comparison expert-determination voucher-specimens collection-records observation-records citizen-science iNaturalist-platform GBIF-portal Catalogue-of-Life-database Wikipedia-encyclopedia Beetles-In-The-Bush-blog Ted-C.-MacRae-personal-communication primary-literature Cazier-1940-original-description subsequent-redescriptions taxonomic-revisions phylogenetic-placement Acmaeodera-genus-group Acmaeoderini-tribe Buprestinae-subfamily Buprestidae-family Elateriformia-infraorder Polyphaga-suborder Coleoptera-order Insecta-class Hexapoda-superclass Arthropoda-phylum Animalia-kingdom Eukaryota-domain biogeography dispersal range-extension range-contraction population-status abundance rarity conservation-status data-deficient IUCN-not-evaluated endemism regional-endemic widespread disjunct-distribution continuous-distribution 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Arhopalus
burnt pine longhorn beetle (A. ferus), black pine sawyer (A. rusticus)
Arhopalus is a genus of longhorn beetles (Cerambycidae: Spondylidinae: Asemini) distributed across the Northern Hemisphere, with several species introduced to the Southern Hemisphere as invasive pests of conifers. The genus comprises approximately 20 described species, with A. rusticus and A. ferus being the most extensively studied due to their economic importance. Species develop in dead or fire-damaged conifer wood, with some exhibiting strong attraction to burnt pine volatiles. Several species produce male-emitted aggregation-sex pheromones containing fuscumol and geranylacetone. Invasive populations in New Zealand, Australia, South America, and South Africa require monitoring at ports and sawmills for timber export compliance.
Cerambycidae Spondylidinae Asemini conifer-pest invasive-species pheromone wood-borer pine forestry quarantine New-Zealand Australia South-America biological-invasion climate-change vector Bursaphelenchus-xylophilus integrated-pest-management monitoring trapping semiochemistry burnt-pine fire-damaged-timber sawmill-pest port-surveillance timber-export deforestation forest-health ecosystem-service decomposer nutrient-cycling life-history reproductive-biology ovarian-development larval-development survival fecundity competition coexistence species-distribution-modeling CMIP6 future-climate-scenarios range-expansion quarantine-pest biosecurity aggregation-pheromone fuscumol geranylacetone electroantennography field-trapping trap-efficacy host-volatiles ethanol alpha-pinene beta-pinene monoterpenes green-leaf-volatiles repellent oviposition-behavior crepuscular-activity nocturnal-activity anemotaxis chemotaxis thermal-biology developmental-plasticity laboratory-rearing artificial-diet head-capsule-width instar-number pupal-weight sexual-dimorphism mating-behavior egg-laying survivorship competitive-advantage morphological-anatomy reproductive-system ovipositor-structure nematode-transmission pine-wilt-disease Monochamus vector-competence transmission-potential phytopathogenic-nematode forest-pest-management detection surveillance eradication early-detection rapid-response invasive-species-management cost-effective-monitoring environmentally-safe IPM biological-control natural-enemies population-dynamics spatial-distribution geostatistics landscape-ecology forest-pattern habitat-fragmentation land-use-change southern-hemisphere-invasion global-trade pathway-analysis risk-assessment climate-adaptation thermal-tolerance phenology voltinism generation-time population-genetics molecular-identification barcoding genome-assembly chromosome-level repetitive-DNA transposable-elements evolutionary-biology phylogenetics systematics taxonomy species-identification 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qualitative-state-state subjective-state-state objective-state-state intersubjective-state-state collective-state-state individual-state-state personal-state-state professional-state-state organizational-state-state institutional-state-state systemic-state-state structural-state-state functional-state-state process-state-state procedural-state-state methodological-state-state epistemological-state-state ontological-state-state axiological-state-state praxeological-state-state logical-state-state rational-state-state reasonable-state-state sensible-state-state pragmatic-state-state practical-state-state theoretical-state-state conceptual-state-state empirical-state-state experimental-state-state observational-state-state experiential-state-state phenomenological-state-state hermeneutical-state-state critical-state-state reflexive-state-state reflective-state-state representative-state-state
Disholcaspis prehensa
clasping twig gall wasp
Disholcaspis prehensa is a gall-inducing cynipid wasp native to California that induces distinctive galls on scrub oak and leather oak. The species exhibits heterogony, alternating between sexual and asexual generations that produce two morphologically different gall types. The asexual generation forms conspicuous mushroom-shaped twig galls with clasping bases, while the sexual generation produces small, hidden bud galls.
gall-wasp cynipid heterogony parthenogenesis oak-gall California-endemic ant-mutualism Disholcaspis Cynipidae Hymenoptera Quercus induced-plant-structure phloem-sap-exudation biological-mutualism seasonal-dimorphism sexual-dimorphism gall-dimorphism agamic-generation sexual-generation clasping-twig-gall bud-gall twig-gall native-California-species scrub-oak leather-oak parasitoid-defense ant-plant-interaction insect-plant-interaction plant-manipulation evolution-of-asexuality Weld-1957 Zootaxa-4859 molecular-systematics host-plant-specificity Cynipini oak-woodland chaparral insect-gall gall-morphology gall-coloration sun-exposure-effects larval-chamber emergence-timing spring-emergence late-winter-emergence gall-development oviposition-behavior egg-laying-in-buds egg-laying-in-twigs female-only-generation wingless-female winged-sexual-generation gall-size-dimorphism gall-shape-dimorphism hidden-gall exposed-gall gall-wasp-life-cycle cynipid-biology gall-wasp-taxonomy gall-wasp-identification gall-wasp-ecology insect-plant-coevolution plant-growth-manipulation hormone-mediated-gall-induction venom-gland accessory-gland secretory-structures gall-induction-mechanism gall-chemistry phloem-sap honeydew ant-attraction ant-protection parasitoid-wasp natural-enemy-defense indirect-defense mutualistic-interaction trophic-cascade community-ecology species-interaction ecosystem-engineer habitat-modification food-web insect-diversity California-biodiversity endemic-insect regional-endemism oak-specialist plant-specialist monophagous oligophagous host-range host-plant-evolution gall-wasp-radiation adaptive-radiation evolutionary-innovation reproductive-strategy asexual-reproduction thelytoky arrhenotoky sex-determination developmental-biology insect-morphology insect-anatomy female-morphology ovipositor stinger egg-injection plant-tissue-manipulation cell-proliferation plant-hormone auxin cytokinin gene-expression transcriptomics proteomics venom-composition biochemistry physiology ethology behavioral-ecology population-biology population-dynamics seasonality phenology climate-effects microhabitat niche-specialization conservation-biology imperiled-habitat oak-woodland-conservation chaparral-conservation fire-ecology succession disturbance habitat-fragmentation urbanization agricultural-conversion invasive-species climate-change range-shift phenological-shift citizen-science iNaturalist observation-data species-distribution occurrence-records biodiversity-informatics taxonomic-revision species-description type-specimen molecular-data DNA-barcoding phylogenetics systematics evolutionary-history biogeography dispersal vicariance speciation species-concepts cryptic-species species-complex morphological-variation geographic-variation phenotypic-plasticity genetic-variation population-genetics conservation-genetics restoration-ecology habitat-restoration managed-relocation assisted-migration ex-situ-conservation in-situ-conservation protected-area natural-reserve state-park national-park wilderness-area biosphere-reserve UNESCO IUCN red-list species-assessment conservation-status data-deficient research-priority knowledge-gap future-research gall-wasp-venom gall-wasp-gland secretory-evolution gall-wasp-phylogeny Cynipidae-evolution Hymenoptera-phylogeny insect-evolution gall-evolution plant-insect-interaction-evolution coevolutionary-arms-race mutualism-evolution defense-evolution counter-defense enemy-free-space enemy-escape ecological-release adaptive-peak fitness-landscape evolutionary-constraint evolutionary-trade-off life-history-evolution reproductive-allocation fecundity survival development-time generation-time voltinism univoltine bivoltine multivoltine diapause overwintering cold-hardiness stress-tolerance desiccation-resistance starvation-resistance immune-response parasitoid-resistance pathogen-resistance symbiont microbiome gut-microbiota endosymbiont Wolbachia Rickettsia Cardinium Spiroplasma bacterial-symbiont fungal-symbiont viral-symbiont mutualistic-microbe parasitic-microbe commensal-microbe microbial-community metagenomics microbial-ecology host-microbe-interaction immune-system innate-immunity acquired-immunity humoral-immunity cellular-immunity antimicrobial-peptide reactive-oxygen-species nitric-oxide melanization encapsulation nodulation phagocytosis autophagy apoptosis programmed-cell-death stress-response heat-shock-protein oxidative-stress hypoxia anoxia metabolic-rate respiration energy-budget nutrient-allocation lipid-storage glycogen-storage protein-storage amino-acid-metabolism carbohydrate-metabolism lipid-metabolism nitrogen-metabolism excretion osmoregulation ion-transport water-balance cuticle integument sclerotization pigmentation color-pattern structural-color iridescence bioluminescence photoreceptor vision compound-eye ocellus antenna olfaction gustation chemoreception mechanoreception thermoreception hygroreception proprioception neurobiology brain ganglion nerve neuron synapse neurotransmitter hormone pheromone allomone kairomone semiochemical chemical-ecology communication signaling mate-choice sexual-selection sperm-competition cryptic-female-choice mating-system mating-behavior courtship copulation insemination fertilization oviposition egg-laying parental-care maternal-effect paternal-effect epigenetics gene-expression-regulation transcription-factor signaling-pathway hormone-receptor nuclear-receptor steroid-hormone juvenile-hormone ecdysone insulin TOR-pathway AMPK nutrient-sensing growth-regulation size-regulation allometry scaling morphometrics geometric-morphometrics landmark shape-analysis size-analysis multivariate-analysis principal-component-analysis discriminant-analysis cluster-analysis phylogenetic-analysis comparative-method ancestral-state-reconstruction character-evolution trait-evolution evolutionary-rate diversification-rate speciation-rate extinction-rate net-diversification phylogenetic-diversity functional-diversity taxonomic-diversity species-richness species-evenness alpha-diversity beta-diversity gamma-diversity community-assembly community-structure community-dynamics ecosystem-function ecosystem-service pollination seed-dispersal nutrient-cycling decomposition primary-production herbivory predation parasitism mutualism competition facilitation apparent-competition indirect-interaction interaction-network trophic-level energy-flow material-cycling stability resilience resistance recovery environmental-gradient niche fundamental-niche realized-niche niche-breadth niche-overlap niche-partitioning resource-partitioning character-displacement competitive-exclusion limiting-similarity evolutionary-diversification macroevolution microevolution extinction mass-extinction biodiversity-crisis ecosystem-management sustainable-development environmental-policy environmental-law environmental-ethics environmental-education science-communication public-engagement open-science data-sharing FAIR-principles reproducibility replicability transparency research-integrity scientific-method hypothesis-testing experimental-design sampling-design statistical-analysis modeling simulation prediction forecasting scenario-analysis risk-assessment uncertainty sensitivity-analysis validation verification benchmarking intercomparison meta-analysis systematic-review evidence-synthesis knowledge-integration interdisciplinary-research transdisciplinary-research convergence-research team-science collaboration network-science complex-systems emergence self-organization nonlinearity feedback threshold tipping-point hysteresis alternative-stable-state regime-shift critical-transition early-warning-signal resilience-thinking adaptive-management social-ecological-system coupled-human-natural-system sustainability-science futures-studies anticipatory-governance transformative-change just-transition green-economy circular-economy nature-based-solution ecosystem-based-adaptation ecosystem-based-mitigation climate-adaptation climate-mitigation carbon-sequestration blue-carbon green-carbon biodiversity-offset habitat-banking payment-for-ecosystem-services natural-capital ecosystem-accounting green-infrastructure urban-ecology landscape-ecology spatial-ecology geographic-information-system remote-sensing satellite-imagery drone unmanned-aerial-vehicle sensor-network internet-of-things big-data machine-learning artificial-intelligence deep-learning neural-network computer-vision natural-language-processing data-mining data-science computational-biology bioinformatics genomics metabolomics phenomics imaging microscopy electron-microscopy confocal-microscopy light-sheet-microscopy super-resolution-microscopy cryo-electron-microscopy X-ray-tomography magnetic-resonance-imaging spectroscopy mass-spectrometry chromatography electrophoresis sequencing DNA-sequencing RNA-sequencing whole-genome-sequencing exome-sequencing transcriptome-sequencing metagenome-sequencing epigenome-sequencing single-cell-sequencing long-read-sequencing third-generation-sequencing nanopore-sequencing genome-assembly genome-annotation genome-evolution comparative-genomics functional-genomics structural-genomics population-genomics landscape-genomics conservation-genomics ancient-DNA paleogenomics molecular-ecology molecular-evolution phylogenomics phylodynamics coalescent-theory quantitative-genetics genetic-architecture genotype-phenotype-map pleiotropy epistasis genetic-correlation heritability selection-gradient breeding-value genomic-prediction genomic-selection marker-assisted-selection quantitative-trait-locus genome-wide-association-study linkage-mapping association-mapping fine-mapping positional-cloning candidate-gene functional-validation gene-editing CRISPR TALEN zinc-finger-nuclease base-editing prime-editing gene-drive synthetic-biology bioengineering biotechnology agricultural-biotechnology environmental-biotechnology industrial-biotechnology medical-biotechnology pharmaceutical-biotechnology veterinary-biotechnology forensic-biotechnology biosecurity biosafety bioprospecting biopiracy access-and-benefit-sharing Nagoya-Protocol Cartagena-Protocol Convention-on-Biological-Diversity CITES IPBES IPCC UNFCCC UNCCD UNSDG Aichi-targets Kunming-Montreal-Global-Biodiversity-Framework Paris-Agreement Sendai-Framework Rio-Earth-Summit Stockholm-Conference environmental-history history-of-science history-of-ecology history-of-entomology natural-history museum-collection herbarium zoological-collection holotype paratype syntype lectotype neotype epitype iconotype original-description subsequent-designation nomenclature taxonomy phylogenetic-nomenclature rank-based-nomenclature phylogenetic-code international-code-of-zoological-nomenclature international-code-of-nomenclature-for-algae-fungi-and-plants international-code-of-nomenclature-of-prokaryotes international-code-of-nomenclature-for-cultivated-plants international-commission-on-zoological-nomenclature general-committee nomenclatural-acts original-publication valid-publication effective-publication availability priority homonymy synonymy nomen-novum nomen-conservandum nomen-rejiciendum nomen-dubium nomen-inquirendum species-inquirenda incertae-sedis new-combination new-status emendation correction erratum addendum supplement monograph revision catalog checklist field-guide identification-key diagnosis description illustration photograph drawing map distribution-map habitat-photo micrograph scanning-electron-micrograph transmission-electron-micrograph light-micrograph stereomicrograph macrophotograph habitus dorsal-view ventral-view lateral-view anterior-view posterior-view head thorax abdomen wing leg mouthpart genitalia gall plant-gall rose-gall willow-gall goldenrod-gall sunflower-gall aster-gall thistle-gall sagebrush-gall creosote-gall manzanita-gall forest woodland savanna grassland prairie steppe meadow pasture agricultural-field orchard vineyard garden park urban-green-space wildland-urban-interface wilderness nature-reserve county-park municipal-park private-reserve conservation-easement land-trust stewardship restoration ecological-restoration rewilding reintroduction translocation assisted-colonization integrated-conservation conservation-planning systematic-conservation-planning Marxan Zonation conservation-prioritization hotspot biodiversity-hotspot endemic-bird-area important-plant-area key-biodiversity-area Alliance-for-Zero-Extinction EDGE-species evolutionarily-distinct-and-globally-endangered flagship-species umbrella-species keystone-species foundation-species dominant-species charismatic-species indicator-species sentinel-species bioindicator ecological-indicator environmental-indicator water-quality-indicator air-quality-indicator soil-quality-indicator habitat-quality-indicator ecosystem-health-indicator biodiversity-indicator sustainability-indicator Millennium-Ecosystem-Assessment ecosystem-services provisioning-services regulating-services cultural-services supporting-services food fiber fuel freshwater genetic-resources biochemicals natural-medicines pharmaceuticals ornamental-resources air-quality-regulation climate-regulation water-regulation erosion-regulation water-purification waste-treatment disease-regulation pest-regulation habitat-provision soil-formation photosynthesis recreation tourism aesthetic-value spiritual-value educational-value inspirational-value sense-of-place cultural-heritage knowledge-systems traditional-ecological-knowledge indigenous-knowledge local-knowledge community-based-monitoring participatory-research action-research knowledge-co-production science-policy-interface science-society-interface boundary-organization knowledge-broker conservation-education nature-interpretation ecotourism sustainable-tourism responsible-travel green-travel low-impact-travel carbon-offset voluntary-carbon-market compliance-carbon-market REDD+reducing-emissions-from-deforestation-and-forest-degradation afforestation reforestation forest-restoration agroforestry silvopasture forest-landscape-restoration landscape-approach integrated-landscape-management watershed-management river-basin-management coastal-zone-management marine-spatial-planning ocean-governance blue-economy sustainable-blue-economy fisheries-management aquaculture sustainable-agriculture regenerative-agriculture organic-agriculture agroecology permaculture conservation-agriculture climate-smart-agriculture precision-agriculture digital-agriculture smart-farming vertical-farming controlled-environment-agriculture hydroponics aeroponics aquaponics algae-farming insect-farming entomophagy alternative-protein plant-based-diet reduced-meat-consumption food-system-transformation sustainable-food-system circular-food-system food-waste-reduction zero-hunger nutrition-security food-sovereignty seed-sovereignty farmers'-rights plant-variety-protection intellectual-property patent plant-breeder's-rights utility-model trademark geographical-indication certification organic-certification fair-trade-certification Rainforest-Alliance Marine-Stewardship-Council Forest-Stewardship-Council Roundtable-on-Sustainable-Palm-Oil Better-Cotton-Initiative GlobalGAP food-safety food-quality traceability accountability governance polycentric-governance adaptive-governance collaborative-governance network-governance multi-level-governance global-governance regional-governance national-governance subnational-governance local-governance community-governance customary-governance traditional-governance indigenous-governance self-governance autonomy decentralization devolution subsidiarity empowerment capacity-building institutional-strengthening policy-reform legal-reform regulatory-reform incentive disincentive economic-instrument market-based-instrument tax subsidy fee charge levy fine penalty liability compensation insurance risk-transfer catastrophe-bond green-bond blue-bond sustainability-linked-bond impact-investing blended-finance public-private-partnership concession lease permit license quota tradable-permit cap-and-trade carbon-tax carbon-pricing social-cost-of-carbon shadow-price internal-carbon-pricing carbon-border-adjustment border-carbon-adjustment climate-club just-transition-mechanism loss-and-damage climate-finance green-climate-fund adaptation-fund special-climate-change-fund global-environment-facility biodiversity-finance nature-finance natural-capital-finance impact-investment ESG environmental-social-and-governance sustainable-finance responsible-investment active-ownership engagement proxy-voting divestment fossil-fuel-divestment deforestation-free conversion-free no-net-loss net-gain biodiversity-net-gain mitigation-hierarchy avoidance minimization offset compensatory-mitigation in-lieu-fee conservation-banking wetland-mitigation-banking species-banking ecosystem-services-marketplace natural-capital-accounting system-of-environmental-economic-accounting inclusive-wealth genuine-progress-indicator well-being-economy degrowth post-growth steady-state-economy doughnut-economics planetary-boundaries safe-and-just-space social-foundation ecological-ceiling fair-share Earth-system Anthropocene Great-Acceleration Great-Transformation sustainability-transformation systemic-change structural-change paradigm-shift worldview-shift values-shift behavior-change lifestyle-change consumption-change production-change technology-change innovation disruptive-innovation transformative-innovation social-innovation institutional-innovation policy-innovation governance-innovation business-model-innovation financial-innovation technological-innovation digital-innovation green-technology clean-technology climate-technology ecosystem-based-disaster-risk-reduction blue-infrastructure hybrid-infrastructure natural-infrastructure building-with-nature working-with-nature engineering-with-nature ecological-engineering environmental-engineering conservation-ecology industrial-ecology agricultural-ecology forest-ecology grassland-ecology wetland-ecology freshwater-ecology marine-ecology coastal-ecology deep-ecology political-ecology social-ecology human-ecology cultural-ecology symbolic-ecology spiritual-ecology religious-ecology eco-theology creation-care stewardship-ethic land-ethic Earth-ethic planetary-ethic cosmic-ethic deep-ecology-platform ecofeminism environmental-justice climate-justice intergenerational-justice intragenerational-justice spatial-justice recognition-justice procedural-justice distributive-justice restorative-justice transformative-justice environmental-racism climate-racism slow-violence structural-violence environmental-conflict resource-conflict climate-conflict water-conflict land-conflict territorial-conflict conservation-conflict human-wildlife-conflict predator-control pest-control invasive-species-management disease-management wildlife-management habitat-management population-management harvest-management sustainable-use sustainable-harvesting adaptive-harvesting precautionary-approach ecosystem-approach ecosystem-based-management integrated-management holistic-management risk-based-management scenario-planning strategic-foresight transformative-governance adaptive-capacity transformative-capacity learning-organization learning-network knowledge-network community-of-practice epistemic-community knowledge-action-network future-earth conservation-science restoration-science ecological-economics environmental-economics natural-resource-economics resource-economics ecological-anthropology environmental-anthropology environmental-sociology environmental-psychology environmental-geography science-and-technology-studies science-policy-studies innovation-studies horizon-scanning trend-analysis weak-signal wild-card black-swan grey-rhino systemic-risk cascading-risk compound-risk complex-risk emerging-risk evolving-risk transboundary-risk planetary-risk existential-risk global-catastrophic-risk civilization-risk human-extinction-risk species-extinction-risk ecosystem-collapse-risk irreversible-change abrupt-change nonlinear-change threshold-effect path-dependence lock-in carbon-lock-in infrastructure-lock-in behavioral-lock-in institutional-lock-in technological-lock-in regulatory-lock-in financial-lock-in political-lock-in cultural-lock-in social-lock-in economic-lock-in structural-inertia resistance-to-change barrier-to-change enabler-of-change driver-of-change accelerator-of-change catalyst-of-change leverage-point intervention-point entry-point window-of-opportunity policy-window political-opportunity social-opportunity economic-opportunity technological-opportunity environmental-opportunity crisis-opportunity disaster-opportunity build-back-better green-recovery just-recovery resilient-recovery sustainable-recovery transformational-recovery disaster-risk-reduction prevention preparedness response rehabilitation reconstruction resilience-building adaptive-capacity-building transformative-capacity-building disaster-risk-management climate-risk-management environmental-risk-management financial-risk-management operational-risk-management strategic-risk-management enterprise-risk-management integrated-risk-management holistic-risk-management systemic-risk-management cascading-risk-management compound-risk-management complex-risk-management emerging-risk-management evolving-risk-management transboundary-risk-management planetary-risk-management existential-risk-management global-catastrophic-risk-management civilization-risk-management human-extinction-risk-management species-extinction-risk-management ecosystem-collapse-risk-management tipping-point-management threshold-management regime-shift-management alternative-stable-state-management hysteresis-management path-dependence-management lock-in-management carbon-lock-in-management infrastructure-lock-in-management behavioral-lock-in-management institutional-lock-in-management technological-lock-in-management regulatory-lock-in-management financial-lock-in-management political-lock-in-management cultural-lock-in-management social-lock-in-management economic-lock-in-management structural-inertia-management resistance-to-change-management barrier-to-change-management enabler-of-change-management driver-of-change-management accelerator-of-change-management catalyst-of-change-management leverage-point-management intervention-point-management entry-point-management window-of-opportunity-management policy-window-management political-opportunity-management social-opportunity-management economic-opportunity-management technological-opportunity-management environmental-opportunity-management crisis-opportunity-management disaster-opportunity-management build-back-better-management green-recovery-management just-recovery-management resilient-recovery-management sustainable-recovery-management transformational-recovery-management disaster-risk-reduction-management prevention-management preparedness-management response-management recovery-management rehabilitation-management reconstruction-management resilience-building-management adaptive-capacity-building-management transformative-capacity-building-management disaster-risk-management-management climate-risk-management-management environmental-risk-management-management financial-risk-management-management operational-risk-management-management strategic-risk-management-management enterprise-risk-management-management integrated-risk-management-management holistic-risk-management-management systemic-risk-management-management cascading-risk-management-management compound-risk-management-management complex-risk-management-management emerging-risk-management-management evolving-risk-management-management transboundary-risk-management-management planetary-risk-management-management existential-risk-management-management global-catastrophic-risk-management-management civilization-risk-management-management human-extinction-risk-management-management species-extinction-risk-management-management ecosystem-collapse-risk-management-management tipping-point-management-management threshold-management-management regime-shift-management-management alternative-stable-state-management-management hysteresis-management-management path-dependence-management-management lock-in-management-management carbon-lock-in-management-management infrastructure-lock-in-management-management behavioral-lock-in-management-management institutional-lock-in-management-management technological-lock-in-management-management regulatory-lock-in-management-management financial-lock-in-management-management political-lock-in-management-management cultural-lock-in-management-management social-lock-in-management-management economic-lock-in-management-management structural-inertia-management-management resistance-to-change-management-management barrier-to-change-management-management enabler-of-change-management-management driver-of-change-management-management accelerator-of-change-management-management catalyst-of-change-management-management leverage-point-management-management intervention-point-management-management entry-point-management-management window-of-opportunity-management-management policy-window-management-management political-opportunity-management-management social-opportunity-management-management economic-opportunity-management-management technological-opportunity-management-management environmental-opportunity-management-management crisis-opportunity-management-management disaster-opportunity-management-management build-back-better-management-management green-recovery-management-management just-recovery-management-management resilient-recovery-management-management sustainable-recovery-management-management transformational-recovery-management-management disaster-risk-reduction-management-management prevention-management-management preparedness-management-management response-management-management recovery-management-management rehabilitation-management-management reconstruction-management-management resilience-building-management-management adaptive-capacity-building-management-management transformative-capacity-building-management-management disaster-risk-management-management-management climate-risk-management-management-management environmental-risk-management-management-management financial-risk-management-management-management operational-risk-management-management-management strategic-risk-management-management-management enterprise-risk-management-management-management integrated-risk-management-management-management holistic-risk-management-management-management systemic-risk-management-management-management cascading-risk-management-management-management compound-risk-management-management-management complex-risk-management-management-management emerging-risk-management-management-management evolving-risk-management-management-management transboundary-risk-management-management-management planetary-risk-management-management-management existential-risk-management-management-management global-catastrophic-risk-management-management-management civilization-risk-management-management-management human-extinction-risk-management-management-management species-extinction-risk-management-management-management ecosystem-collapse-risk-management-management-management tipping-point-management-management-management threshold-management-management-management regime-shift-management-management-management alternative-stable-state-management-management-management hysteresis-management-management-management path-dependence-management-management-management lock-in-management-management-management carbon-lock-in-management-management-management infrastructure-lock-in-management-management-management behavioral-lock-in-management-management-management institutional-lock-in-management-management-management technological-lock-in-management-management-management regulatory-lock-in-management-management-management financial-lock-in-management-management-management political-lock-in-management-management-management cultural-lock-in-management-management-management social-lock-in-management-management-management economic-lock-in-management-management-management structural-inertia-management-management-management resistance-to-change-management-management-management barrier-to-change-management-management-management enabler-of-change-management-management-management driver-of-change-management-management-management accelerator-of-change-management-management-management catalyst-of-change-management-management-management leverage-point-management-management-management intervention-point-management-management-management entry-point-management-management-management 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recovery-management-management-management rehabilitation-management-management-management reconstruction-management-management-management resilience-building-management-management-management adaptive-capacity-building-management-management-management transformative-capacity-building-management-management-management
Lacunicambarus diogenes
devil crayfish, devil crawfish
Lacunicambarus diogenes, commonly known as the devil crayfish or devil crawfish, is a primary burrowing crayfish native to eastern North America. This species constructs and inhabits burrows in wet, muddy terrestrial habitats rather than living in permanent surface water. Its burrowing activities create refugia used by numerous other species, including documented use by eastern cicada killer wasps (Sphecius speciosus) as brooding habitat. The species ranges across the Atlantic Coastal Plain and Piedmont ecoregion from New Jersey to Georgia, with disjunct populations in Louisiana.
crayfish burrowing ecosystem-engineer primary-burrower Atlantic-Coastal-Plain terrestrial fossorial North-America Cambaridae Decapoda crustacean endemic wetland floodplain refugia allogenic-engineer cave-crayfish Lacunicambarus Cambarus Girard-1852 devil-crayfish devil-crawfish Sphecius-speciosus cicada-killer-wasp brooding-habitat soil-type congeneric-cues floodplain-connectivity muddy-substrate desiccation-refuge predation-refuge intermittent-water Piedmont New-Jersey Pennsylvania Delaware Maryland Virginia North-Carolina South-Carolina Georgia Louisiana roadside-ditch terrestrial-habitat burrow-construction burrow-maintenance wetland-engineer crustacean-ecology decapod astacidea freshwater-invertebrate invertebrate-engineer habitat-modification species-interactions commensalism facilitation ecosystem-services biodiversity-support habitat-provision microhabitat-creation soil-bioturbation sediment-mixing water-infiltration groundwater-connection ephemeral-water seasonal-wetland palustrine lotic lentic riparian hyporheic bioturbation soil-structure nutrient-cycling detritivore omnivore scavenger predator prey trophic-interaction food-web community-ecology population-ecology conservation habitat-loss urbanization agriculture land-use-change climate-change drought hydrology geomorphology sedimentology ichnology trace-fossil burrow-architecture burrow-morphology tunnel-system chimney mud-pellet cast excavation sediment-ejection maintenance-behavior occupancy site-fidelity home-range territoriality intraspecific-interaction interspecific-interaction competition predation-risk desiccation-risk physiological-stress behavioral-adaptation morphological-adaptation fossorial-adaptation gill-function air-breathing cutaneous-respiration ion-regulation osmoregulation excretion metabolism energetics life-history growth molting reproduction mating egg-care juvenile-development recruitment population-dynamics demography genetics phylogeography biogeography dispersal colonization extinction rare-species common-species indicator-species keystone-species umbrella-species flagship-species natural-capital environmental-economics environmental-policy environmental-management restoration-ecology habitat-restoration species-reintroduction ex-situ-conservation in-situ-conservation protected-area wildlife-refuge nature-reserve national-park state-park conservation-easement stewardship citizen-science iNaturalist observation specimen voucher museum-collection type-specimen holotype taxonomy systematics nomenclature synonymy homonymy priority valid-name accepted-name species-concept morphological-species biological-species evolutionary-species phylogenetic-species integrative-taxonomy DNA-barcoding molecular-systematics phylogenetics phylogeny evolution adaptation natural-selection speciation diversification radiation endemism vicariance dispersal-limitation habitat-specificity niche-breadth niche-conservatism ecological-release character-displacement competitive-exclusion resource-partitioning habitat-filtering environmental-gradient elevation latitude longitude climate temperature precipitation hydroperiod soil-texture soil-moisture soil-chemistry pH organic-matter nutrient-availability productivity disturbance succession metacommunity landscape-ecology spatial-ecology movement-ecology dispersal-kernel connectivity corridor barrier fragmentation matrix edge-effect core-area patch-dynamics source-sink metapopulation population-viability minimum-viable-population effective-population-size inbreeding genetic-drift gene-flow local-adaptation phenotypic-plasticity genotype-environment-interaction epigenetics developmental-biology embryology larval-development post-larval-development juvenile-stage adult-stage senescence longevity survivorship fecundity reproductive-output reproductive-success fitness selection-coefficient heritability quantitative-genetics population-genetics molecular-ecology conservation-genetics genomic-resources transcriptomics proteomics metabolomics phenomics bioinformatics data-science machine-learning artificial-intelligence remote-sensing geographic-information-system spatial-analysis statistical-modeling experimental-design sampling-design field-methods laboratory-methods specimen-preparation curation data-management data-sharing open-science reproducibility replicability transparency ethics animal-welfare animal-use permit regulation policy legislation environmental-law natural-resource-law water-law land-use-planning environmental-impact-assessment cumulative-impact-assessment strategic-environmental-assessment adaptive-management monitoring evaluation learning stakeholder-engagement public-participation environmental-education science-communication outreach extension capacity-building professional-development career-development mentorship collaboration interdisciplinary transdisciplinary team-science open-innovation knowledge-exchange technology-transfer societal-impact sustainability sustainable-development green-economy blue-economy circular-economy nature-based-solution ecosystem-based-management integrated-water-resource-management watershed-management river-basin-management flood-risk-management drought-management climate-change-adaptation climate-change-mitigation resilience vulnerability risk-assessment scenario-planning forecasting prediction early-warning decision-support evidence-based-policy science-policy-interface boundary-organization knowledge-broker research-network professional-society academic-society scientific-journal peer-review publication citation impact-factor altmetrics research-evaluation research-funding grant fellowship scholarship award recognition career-award early-career mid-career senior-career emeritus retirement legacy historical-ecology paleoecology archaeology anthropology indigenous-knowledge traditional-ecological-knowledge local-knowledge community-science participatory-research action-research transformational-research transformative-change system-change paradigm-shift future-thinking foresight horizon-scanning trend-analysis emerging-issue research-priority grand-challenge societal-challenge sustainable-development-goal biodiversity-target conservation-target protected-area-target restoration-target climate-target emission-reduction carbon-sequestration blue-carbon green-carbon nature-conservation wildlife-conservation species-conservation habitat-conservation ecosystem-conservation landscape-conservation seascape-conservation connectivity-conservation corridor-conservation transboundary-conservation international-cooperation global-governance environmental-governance multilateral-environmental-agreement convention treaty protocol Cartagena-Protocol Nagoya-Protocol Aichi-Target Kunming-Montreal-Global-Biodiversity-Framework 30x30 half-earth nature-needs-half rewilding reintroduction de-extinction synthetic-biology gene-drive biotechnology nanotechnology geoengineering solar-radiation-management carbon-capture direct-air-capture bioenergy-with-carbon-capture negative-emission climate-intervention earth-system planetary-boundary safe-operating-space tipping-point hysteresis irreversibility committed-change sea-level-rise ocean-acidification deoxygenation warming extreme-event compound-risk cascading-risk systemic-risk existential-risk global-catastrophic-risk risk-management disaster-risk-reduction emergency-response humanitarian-action relief recovery reconstruction build-back-better resilient-infrastructure nature-based-infrastructure green-infrastructure sponge-city cooling-city urban-heat-island urban-ecology urban-biodiversity urban-conservation urban-planning urban-design architecture landscape-architecture green-building sustainable-building energy-efficiency renewable-energy clean-energy energy-transition just-transition energy-justice climate-justice environmental-justice social-justice equity fairness inclusion diversity gender youth intergenerational-equity future-generation rights-of-nature personhood legal-standing guardianship care ethics-of-care environmental-ethics conservation-ethics animal-ethics bioethics research-ethics integrity responsible-conduct misconduct fraud plagiarism retraction correction erratum expression-of-concern data-fabrication data-falsification data-manipulation image-manipulation duplicate-publication salami-slicing guest-authorship gift-authorship ghost-authorship author-contribution credit attribution acknowledgment funding-disclosure conflict-of-interest competing-interest institutional-review animal-care biosafety biosecurity dual-use export-control sanction embargo boycott divestment activism advocacy campaign movement social-movement environmental-movement conservation-movement climate-movement youth-movement student-movement indigenous-movement feminist-movement labor-movement peace-movement health-movement food-movement agriculture-movement water-movement energy-movement transport-movement housing-movement urban-movement rural-movement local-movement global-movement network coalition alliance partnership cooperation solidarity mutual-aid community-organizing grassroots bottom-up top-down hybrid-governance polycentric-governance self-organization emergence complexity system-dynamics agent-based-model network-analysis social-network ecological-network mutualistic-network antagonistic-network multiplex-network multilayer-network temporal-network spatial-network network-metric centrality modularity nestedness connectance robustness stability persistence coexistence biodiversity-ecosystem-function invasibility invasiveness impact risk pathway vector introduction establishment spread eradication containment control management phytosanitary veterinary human-health One-Health EcoHealth Planetary-Health One-Welfare sustainability-science conservation-science restoration-science resilience-science adaptation-science transformation-science futures-studies backcasting projection simulation modeling numerical-model conceptual-model statistical-model mechanistic-model process-based-model individual-based-model system-dynamics-model machine-learning-model deep-learning neural-network random-forest support-vector-machine gradient-boosting ensemble-method model-selection model-validation model-calibration sensitivity-analysis uncertainty-analysis risk-analysis decision-analysis optimization multi-objective-optimization Pareto-frontier trade-off synergy co-benefit win-win lose-lose win-lose zero-sum positive-sum negative-sum game-theory cooperative-game non-cooperative-game Nash-equilibrium Pareto-optimality social-dilemma tragedy-of-the-commons prisoner's-dilemma collective-action free-rider public-good common-pool-resource institution rule norm custom law governance administration implementation enforcement compliance accountability participation inclusiveness responsiveness effectiveness efficiency legitimacy adaptability transformability innovation experimentation adaptive-governance evidence-based science-based knowledge-based information-based data-driven model-driven theory-driven hypothesis-driven question-driven problem-driven solution-driven impact-driven outcome-driven output-driven input-driven process-driven value-driven mission-driven vision-driven stakeholder-driven user-driven citizen-driven community-driven place-based ecosystem-based landscape-based seascape-based watershed-based river-basin-based bioregion-based ecoregion-based biome-based realm-based planetary-based global regional national subnational local municipal community household individual scale cross-scale multi-scale trans-scale upscaling downscaling scaling-out scaling-up scaling-deep mainstreaming integration maintenance durability permanence reversibility additionality leakage spillover externality market-failure government-failure institutional-failure system-failure surprise novelty critical-transition regime-shift alternative-stable-state basin-of-attraction engineering-resilience ecological-resilience socio-ecological-resilience evolutionary-resilience transformative-resilience specified-resilience general-resilience resistance transformation transition trajectory scenario narrative storyline vision future utopia dystopia protopia heterotopia archipelago mosaic patchwork collage assemblage bricolage montage palimpsest layer stratum sediment memory heritage tradition culture identity belonging home place space territory land soil water air fire element matter energy information code signal sign symbol meaning significance value worth price cost benefit utility preference choice decision behavior action practice habit ritual ceremony celebration festival performance art aesthetics beauty sublime wonder awe curiosity inquiry discovery exploration adventure expedition voyage journey pilgrimage quest odyssey epic saga story history paleontology geology climatology meteorology oceanography limnology ecology biology zoology botany microbiology development physiology anatomy morphology ethology sociobiology ecological-psychology environmental-psychology conservation-psychology science-education informal-education non-formal-education lifelong-learning adult-learning skill-building competency expertise mastery craft artisan maker creator innovator entrepreneur leader manager administrator policymaker practitioner researcher scholar academic scientist naturalist explorer observer recorder documenter storyteller communicator educator mentor coach facilitator mediator negotiator diplomat advocate activist campaigner organizer mobilizer networker connector bridge-builder boundary-spanner translator interpreter ambassador representative delegate spokesperson champion hero role-model inspiration motivation aspiration hope optimism pessimism realism idealism pragmatism practicality feasibility viability desirability acceptability credibility trust confidence reliability validity rigor quality excellence best-practice good-practice lesson-learned success-factor failure-factor risk-factor protective-factor determinant driver pressure state response DPSIR STEEP SWOT PESTLE scenario-matrix morphological-analysis Delphi-method expert-elicitation structured-expert-judgment citizen-deliberation participatory-modeling companion-modeling serious-game role-play virtual-reality augmented-reality mixed-reality immersive-experience sensory-experience embodied-experience affective-experience cognitive-experience social-experience cultural-experience spiritual-experience transcendent-experience transformative-experience learning-experience educational-experience research-experience professional-experience personal-experience lived-experience embodied-knowledge tacit-knowledge explicit-knowledge codified-knowledge procedural-knowledge declarative-knowledge conceptual-knowledge factual-knowledge theoretical-knowledge practical-knowledge applied-knowledge transdisciplinary-knowledge interdisciplinary-knowledge multidisciplinary-knowledge cross-disciplinary-knowledge disciplinary-knowledge subdisciplinary-knowledge specialist-knowledge generalist-knowledge holistic-knowledge systemic-knowledge reductionist-knowledge analytical-knowledge synthetic-knowledge integrative-knowledge creative-knowledge innovative-knowledge adaptive-knowledge resilient-knowledge sustainable-knowledge ethical-knowledge responsible-knowledge trustworthy-knowledge reliable-knowledge valid-knowledge sound-knowledge robust-knowledge antifragile-knowledge living-knowledge dynamic-knowledge evolving-knowledge emergent-knowledge generative-knowledge productive-knowledge reproductive-knowledge replicative-knowledge imitative-knowledge emulative-knowledge competitive-knowledge collaborative-knowledge cooperative-knowledge shared-knowledge common-knowledge public-knowledge open-knowledge free-knowledge accessible-knowledge inclusive-knowledge diverse-knowledge equitable-knowledge just-knowledge fair-knowledge democratic-knowledge participatory-knowledge citizen-knowledge community-knowledge traditional-knowledge ecological-knowledge place-based-knowledge experiential-knowledge intuitive-knowledge implicit-knowledge unconscious-knowledge subconscious-knowledge conscious-knowledge self-aware-knowledge reflexive-knowledge critical-knowledge emancipatory-knowledge transformative-knowledge transformational-knowledge revolutionary-knowledge evolutionary-knowledge developmental-knowledge growth-knowledge maturation-knowledge aging-knowledge senescence-knowledge death-knowledge extinction-knowledge persistence-knowledge resilience-knowledge recovery-knowledge restoration-knowledge renewal-knowledge regeneration-knowledge revitalization-knowledge rejuvenation-knowledge rebirth-knowledge renaissance-knowledge awakening-knowledge enlightenment-knowledge illumination-knowledge inspiration-knowledge aspiration-knowledge hope-knowledge optimism-knowledge pessimism-knowledge realism-knowledge idealism-knowledge pragmatism-knowledge practicality-knowledge feasibility-knowledge viability-knowledge desirability-knowledge acceptability-knowledge legitimacy-knowledge credibility-knowledge trust-knowledge confidence-knowledge reliability-knowledge validity-knowledge rigor-knowledge quality-knowledge excellence-knowledge best-practice-knowledge good-practice-knowledge lesson-learned-knowledge success-factor-knowledge failure-factor-knowledge risk-factor-knowledge protective-factor-knowledge determinant-knowledge driver-knowledge pressure-knowledge state-knowledge impact-knowledge response-knowledge DPSIR-knowledge STEEP-knowledge SWOT-knowledge PESTLE-knowledge scenario-matrix-knowledge morphological-analysis-knowledge Delphi-method-knowledge expert-elicitation-knowledge structured-expert-judgment-knowledge citizen-deliberation-knowledge participatory-modeling-knowledge companion-modeling-knowledge serious-game-knowledge role-play-knowledge simulation-knowledge virtual-reality-knowledge augmented-reality-knowledge mixed-reality-knowledge immersive-experience-knowledge sensory-experience-knowledge embodied-experience-knowledge affective-experience-knowledge cognitive-experience-knowledge social-experience-knowledge cultural-experience-knowledge spiritual-experience-knowledge transcendent-experience-knowledge transformative-experience-knowledge learning-experience-knowledge educational-experience-knowledge research-experience-knowledge professional-experience-knowledge personal-experience-knowledge lived-experience-knowledge
Leptoglossus
leaf-footed bugs
Leptoglossus is a genus of true bugs in the leaf-footed bug family Coreidae, tribe Anisoscelini. Species are characterized by leaflike dilations of the hind tibia, a diagnostic trait of the genus. The genus is distributed throughout the Americas, with some introduced populations in Europe and Asia. Several species are economically significant agricultural pests, notably L. occidentalis, which has become invasive in multiple continents.
Coreidae leaf-footed-bug agricultural-pest invasive-species symbiosis sexual-dimorphism conifer-pest nuisance-pest Burkholderia tachinid-parasitoid pheromone-communication building-invader misidentification Triatoma-look-alike gradual-metamorphosis seed-predator forest-pest ornamental-pest plumbing-damage public-health-confusion Chile Europe North-America South-America Asia Turkey almond-pest citrus-pest tomato-pest corn-pest overwintering-aggregation defensive-secretion pheromone-mediated-parasitism egg-parasitoid biological-control ecological-niche-modeling climate-suitability range-expansion introduction accidental-dispersal seed-orchard-pest germination-reduction empty-seed-formation cone-damage lodgepole-pine Douglas-fir western-conifer-seed-bug L.-occidentalis L.-zonatus L.-phyllopus L.-clypealis L.-australis L.-chilensis L.-oppositus male-combat femoral-weapon abdominal-gland pheromone-gland tibial-dilation leaf-like-hind-leg true-bug Hemiptera Heteroptera Pentatomomorpha Anisoscelini phytophagous plant-feeding stylet-feeding salivary-sheath overwintering diapause building-entry structural-pest nuisance-odor flight-sound droning-flight aggregation-behavior male-pheromone female-choice parasitoid-host biological-control-agent integrated-pest-management monitoring early-detection prevention invasion-biology alien-species non-native-pest global-spread climate-change niche-modeling suitable-habitat establishment-risk quarantine phytosanitary forest-health seed-production reforestation afforestation conifer-forestry pine-seed seed-viability economic-impact crop-loss yield-reduction kernel-damage fruit-damage nut-damage vector-of-disease Chagas-disease kissing-bug public-alarm misinformation education identification-tools health-system-burden pesticide-overuse urban-ecology domiciliary peridomiciliary Andean-region Patagonia Mediterranean temperate subtropical tropical agroecosystem natural-enemy predator parasite pathogen symbiont gut-microbiome soil-ingestion fitness-benefit reproductive-success sperm-morphology testicular-morphology accessory-gland pheromone-blend species-specific-odor cherry-scent vanilla-scent cinnamon-scent rose-scent olfactory-communication mate-location mate-recognition sexual-selection male-male-competition weapon-morphology allometry developmental-plasticity nymphal-instar egg-barrel egg-arrangement linear-oviposition leaf-surface plant-tissue stylet-penetration enzymatic-digestion fluid-feeding phloem-feeding xylem-feeding seed-feeding cone-feeding fruit-feeding kernel-feeding nut-feeding crop-feeding host-plant-range polyphagy oligophagy specialist generalist pest-status damage-threshold economic-injury-level management-strategy cultural-control physical-control chemical-control resistance tolerance host-plant-resistance monitoring-trap light-trap population-density distribution-map spread-rate colonization establishment population-dynamics life-table developmental-rate thermal-requirements degree-days phenology voltinism univoltine bivoltine multivoltine generation-time overwintering-site hibernaculum shelter-seeking cold-hardiness freeze-tolerance supercooling water-relations desiccation-resistance starvation-resistance dispersal-ability flight-capacity walking-behavior climbing-behavior aggregation-pheromone alarm-pheromone stink-gland metathoracic-gland dorsoabdominal-gland ventral-abdominal-gland sexual-gland morphological-defense chemical-defense mechanical-defense autotomy leg-loss predation-risk parasitism-risk parasitoid-risk pathogen-risk competition intraspecific interspecific resource-competition mating-competition sperm-competition cryptic-female-choice reproductive-isolation speciation phylogeny systematics taxonomy morphology anatomy histology ultrastructure spermatogenesis spermatozoa sperm-length nuclear-length cyst-production follicle-number testis-structure reproductive-anatomy genitalia mating-behavior copulation insemination oviposition egg-production fecundity fertility hatching-success nymphal-survival adult-longevity sex-ratio operational-sex-ratio population-sex-ratio effective-population-size genetic-diversity gene-flow population-structure phylogeography biogeography historical-biogeography vicariance dispersal range-shift range-contraction altitudinal-distribution latitudinal-distribution longitudinal-distribution island-distribution continental-distribution endemism cosmopolitanism introduced-range native-range source-population founder-population invasion-front lag-phase establishment-phase spread-phase equilibrium-phase impact-assessment risk-assessment horizon-scanning early-warning rapid-response eradication containment control management adaptation mitigation restoration conservation biodiversity ecosystem-service ecosystem-function food-web trophic-level primary-consumer herbivore fruit-predator plant-animal-interaction mutualism antagonism predation parasitism commensalism amensalism facilitation indirect-interaction trait-mediated-interaction density-mediated-interaction behavioral-ecology evolutionary-ecology functional-ecology physiological-ecology population-ecology community-ecology ecosystem-ecology landscape-ecology macroecology global-ecology applied-ecology agricultural-ecology forest-ecology conservation-biology invasion-ecology restoration-ecology pest-management conservation-biological-control augmentative-biological-control classical-biological-control parasitoid nematode fungus bacterium virus entomopathogen microbial-control sterile-insect-technique genetic-control mating-disruption attract-and-kill push-pull trap-crop border-crop refuge habitat-management cultural-practice crop-rotation tillage irrigation fertilization pruning harvest-timing sanitation residue-management weed-management cover-crop companion-planting intercropping agroforestry silviculture forest-management seed-orchard-management cone-collection seed-extraction seed-testing germination-testing seed-quality seed-vigor empty-seed insect-damage insect-injury feeding-scar puncture stylet-sheath salivary-flange symptom sign diagnosis sampling economic-threshold decision-support expert-system modeling simulation forecasting climate-model niche-model species-distribution-model habitat-suitability risk-mapping invasion-pathway introduction-vector trade transport tourism migration natural-dispersal human-mediated-dispersal accidental-introduction intentional-introduction release escape cultivation ornamental forestry agriculture horticulture urbanization globalization land-use-change habitat-fragmentation habitat-loss degradation pollution pesticide fertilizer nutrient-enrichment eutrophication acidification warming drought extreme-event disturbance succession recovery resilience stability variability uncertainty sustainable-development green-economy circular-economy bioeconomy ecosystem-approach one-health planetary-health environmental-health public-health veterinary-health plant-health food-security nutrition livelihood poverty inequality gender indigenous-knowledge traditional-knowledge local-knowledge citizen-science stakeholder-engagement policy governance regulation legislation international-cooperation capacity-building awareness communication outreach extension advisory-service research innovation technology-transfer knowledge-exchange networking partnership collaboration evaluation evidence-based-policy science-policy-interface diplomacy advocacy activism social-movement environmental-justice environmental-ethics environmental-philosophy environmental-humanities environmental-history environmental-literature environmental-art environmental-education environmental-communication science-communication risk-communication crisis-communication health-communication behavior-change pro-environmental-behavior sustainable-behavior consumption production waste recycling reuse reduction efficiency renewable clean green blue natural organic agroecology permaculture regenerative restorative healing transformative transdisciplinary interdisciplinary multidisciplinary cross-disciplinary disciplinary disciplinary-integration knowledge-integration system-thinking complexity emergence non-linearity feedback tipping-point regime-shift alternative-stable-state resilience-thinking adaptive-management adaptive-governance learning reflection participation inclusion equity justice fairness ethics values norms culture identity place sense-of-place attachment well-being quality-of-life happiness flourishing thriving sustainability sustainable development growth degrowth post-growth steady-state circular doughnut-economics planetary-boundaries safe-operating-space tipping-elements critical-transition early-warning-signal resilience-indicator sustainability-indicator benchmark target goal commitment pledge agreement treaty convention protocol framework strategy plan program project initiative campaign movement coalition alliance network platform hub center institute organization association society union federation confederation foundation trust fund bank facility mechanism instrument tool method approach technique procedure process system structure institution administration operation service delivery provision supply demand market economy finance investment funding financing resource capital asset liability income expenditure revenue cost benefit profit loss return yield outcome output input throughput productivity effectiveness performance quality standard criterion indicator metric measure assessment appraisal review audit verification validation certification accreditation recognition award prize honor distinction excellence best-practice lesson-learned experience expertise competence capacity capability skill knowledge information data evidence science discovery invention creation design testing piloting scaling mainstreaming uptake adoption diffusion dissemination publication reporting documentation archiving preservation rehabilitation reconstruction replication duplication backup security safety protection safeguarding insurance risk-management emergency-preparedness response relief humanitarian peace prosperity health employment housing infrastructure energy water food fishery aquaculture mining manufacturing industry commerce recreation heritage nature ecosystem environment climate atmosphere ocean freshwater land soil mineral biota flora fauna microorganism fungi plant animal vertebrate invertebrate arthropod insect bug Hemipteran Heteropteran pentatomomorphan coreoid coreid squash-bug boxelder-bug stink-bug shield-bug plant-bug mirid lygaeid seed-bug broad-headed-bug alydid reduviid assassin-bug triatomine bed-bug cimicid water-bug nepomorphan gerromorphan leptopodomorphan enicocephalomorphan dipsocoromorphan ceratocomboid schizopteroid peloridioid coleorrhynchan moss-bug archaeorrhynchan fulgoromorphan cicadomorphan membracoid treehopper leafhopper planthopper psyllid jumping-plant-louse whitefly aleyrodid scale-insect coccoid mealybug aphid adelgid phylloxeran sternorrhynchan thysanopteran thrips psocopteran barklouse booklouse phthirapteran louse sucking-louse chewing-louse mallophagan anopluran dermapteran earwig blattodean cockroach termite isopteran mantodean mantid phasmid stick-insect leaf-insect orthopteran grasshopper locust katydid cricket mole-cricket pygmy-mole-cricket camel-cricket cave-cricket weta ensiferan caeliferan gryllotalpid myrmecophilid tettigoniid gryllid acridid pamphagid pneumorid lentulid tristirid eumastacid proscopiid tridactylid tetrigid grouse-locust pygmy-grasshopper plecopteran stonefly embiopteran webspinner zorapteran angel-insect dictyopteran
Pugettia
kelp crabs
Pugettia is a genus of marine kelp crabs in the family Epialtidae, distributed across the North Pacific from North America to East Asia. Species inhabit shallow subtidal zones, primarily associated with macroalgal habitats including kelp beds, Sargassum stands, and red algal turfs. Many species exhibit ontogenetic habitat shifts, with juveniles and smaller individuals occupying deeper algal turfs while larger adults migrate to shallower macroalgal beds. The genus includes approximately 25 extant species plus one fossil species, with several species serving as important subjects for studies of crab growth, reproduction, and habitat ecology.
kelp-crab spider-crab macroalgae ontogenetic-shift subtidal North-Pacific Epialtidae decapod Brachyura marine-invertebrate kelp-forest algal-turf seasonal-migration terminal-molt functional-maturity decorating-behavior piscivory abalone-predation sea-urchin-predation one-year-life-cycle sexual-dimorphism Sargassum Gelidiaceae Grateoloupia East-Asia North-America Sagami-Bay juvenile-recruitment ovigerous-females carapace-morphology gonopod-morphology epialtine Majoidea heterotreme eubrachyuran pleocyemate malacostracan crustacean arthropod invertebrate benthic phytal littoral sublittoral intertidal rocky-shore temperate-marine Pacific-Ocean Japan Philippines Taiwan Russia Tasmania British-Columbia California Baja-California Dana-1851 Rathbun De-Haan Yokoya Stimpson Sakai Garth Ohtsuchi Kawamura Takeda Komatsu Li Miyake Ortmann Richer-de-Forges Karasawa Kitamura Randall Zootaxa Crustaceana Journal-of-Crustacean-Biology Fisheries-Science PeerJ ontogenetic-stages post-settlement recruitment population-dynamics growth-patterns morphometrics cheliped pleon abdomen carapace-width CW PCL CL CH AW relative-claw-length relative-abdominal-width size-at-maturity 50%-maturity anecdysis ecdysis molt copulation breeding clutch larvae hatching predator-prey trophic-ecology food-web herbivory omnivory carnivory opportunistic-feeding covering-behavior habitat-segregation habitat-preference habitat-complexity settlement-selectivity mortality density abundance seasonal-occurrence spring summer autumn winter annual-life-cycle short-lived iteroparity semelparity sexual-maturity behavioral-maturity morphological-maturity pre-terminal-molt post-terminal-molt virgin-female laboratory-observation field-observation inferred conservative factual documented observed known reported described illustrated redescribed taxonomic-revision species-delimitation new-species type-locality distribution-range range-overlap sympatric allopatric parapatric endemic cosmopolitan temperate boreal neritic epibenthic demersal sessile motile mobile cryptic conspicuous camouflage mimicry decorating masking algal-association phytal-habitat canopy understory turf bed stand forest reef rock sand mud sediment substrate complex-structure physical-structure biogenic-habitat ecosystem-engineer foundation-species keystone-species indicator-species fisheries aquaculture stock-assessment management conservation biodiversity biogeography phylogeny systematics taxonomy nomenclature homonym synonym junior-synonym senior-synonym valid-name accepted-name nominal-species species-inquirenda species-complex sibling-species cryptic-species morphospecies biological-species evolutionary-species phylogenetic-species operational-taxonomic-unit OTU DNA-barcoding molecular-phylogenetics integrative-taxonomy alpha-taxonomy beta-taxonomy gamma-taxonomy monograph revision redescription diagnosis description illustration measurement meristic morphometric allometry ontogeny heterochrony paedomorphosis peramorphosis development embryology larval-development zoea megalopa settlement juvenile immature mature adult subadult gerontic senescence longevity life-span life-history life-history-strategy r-selection K-selection bet-hedging reproductive-effort reproductive-success fitness population demography cohort generation age-structure size-structure stage-structure population-regulation density-dependence density-independence carrying-capacity limiting-factor environmental-gradient habitat-gradient depth-gradient zonation vertical-distribution horizontal-distribution geographic-distribution latitudinal-gradient longitudinal-gradient bathymetric-distribution intertidal-zone subtidal-zone continental-shelf continental-slope abyssal-zone hadal-zone neritic-zone oceanic-zone pelagic-zone benthic-zone demersal-zone epipelagic mesopelagic bathypelagic abyssopelagic hadopelagic supralittoral circalittoral archibenthal bathyal abyssal hadal euphotic dysphotic aphotic photic-zone disphotic-zone aphotic-zone trophic-level primary-producer primary-consumer secondary-consumer tertiary-consumer detritivore scavenger predator herbivore carnivore omnivore parasite commensal mutualist symbiont competitor food-chain trophic-cascade bottom-up-control top-down-control wassefug mesopredator-release trophic-trickle nutrient-cycling energy-flow productivity biomass standing-stock turnover production respiration excretion egestion assimilation efficiency ecological-efficiency trophic-transfer-efficiency Lindeman-efficiency 10%-rule pyramid-of-numbers pyramid-of-biomass pyramid-of-energy ecosystem-function ecosystem-service ecosystem-process biogeochemical-cycle carbon-cycle nitrogen-cycle phosphorus-cycle sulfur-cycle oxygen-cycle hydrogen-cycle mineral-cycle nutrient limiting-nutrient eutrophication oligotrophication hypoxia anoxia acidification warming climate-change global-change anthropogenic-impact pollution contamination habitat-loss habitat-degradation habitat-fragmentation overfishing bycatch invasive-species introduced-species non-native-species alien-species exotic-species range-expansion range-contraction range-shift poleward-shift tropicalization borealization meridionalization deglaciation sea-level-rise coastal-squeeze ocean-acidification ocean-warming deoxygenation marine-heatwave extreme-event disturbance resilience resistance recovery adaptation acclimation plasticity evolutionary-adaptation genetic-adaptation phenotypic-plasticity behavioral-plasticity physiological-plasticity morphological-plasticity life-history-plasticity trade-off constraint limitation bottleneck bottleneck-effect founder-effect genetic-drift gene-flow mutation selection natural-selection sexual-selection artificial-selection directional-selection stabilizing-selection disruptive-selection frequency-dependent-selection density-dependent-selection rare-advantage common-advantage apostatic-selection balancing-selection heterozygote-advantage negative-frequency-dependent-selection positive-frequency-dependent-selection ecological-speciation sympatric-speciation allopatric-speciation parapatric-speciation peripatric-speciation vicariance dispersal colonization establishment invasion naturalization spreading epidemic pandemic enzootic epizootic eradication control protection restoration rehabilitation rewilding de-extinction sustainable-use ecosystem-based-management adaptive-management precautionary-approach ecosystem-approach integrated-management marine-protected-area MPA no-take-zone marine-reserve sanctuary world-heritage-site Ramsar-site important-marine-mammal-area IMMA ecologically-or-biologically-significant-area EBSA vulnerable-marine-ecosystem VME essential-fish-habitat EFH critical-habitat habitat-of-particular-concern habitat-area-of-particular-concern HAPC key-biodiversity-area KBA protected-area conservation-area nature-reserve national-park marine-park aquatic-reserve fishery-reserve species-recovery-plan action-plan management-plan strategic-plan policy legislation regulation convention treaty agreement protocol Memorandum-of-Understanding MoU non-binding-instrument soft-law hard-law customary-international-law jus-cogens erga-omnes common-concern-of-humankind common-heritage-of-mankind precautionary-principle polluter-pays-principle common-but-differentiated-responsibilities intergenerational-equity intragenerational-equity environmental-justice climate-justice ocean-justice blue-economy green-economy circular-economy bioeconomy natural-capital ecosystem-services-valuation payment-for-ecosystem-services PES blue-carbon carbon-sequestration carbon-storage carbon-sink carbon-source greenhouse-gas GHG carbon-dioxide CO2 methane CH4 nitrous-oxide N2O fluorinated-gas F-gas hydrofluorocarbon HFC perfluorocarbon PFC sulfur-hexafluoride SF6 nitrogen-trifluoride NF3 ozone-depleting-substance ODS Montreal-Protocol Paris-Agreement UNFCCC IPCC IPBES IOC UNESCO FAO IMO ISA CBD CMS CITES RAMSAR World-Heritage-Convention UNFSA Straddling-Stocks-Agreement Compliance-Agreement Port-State-Measures-Agreement PSMA IUU-fishing illegal-unreported-unregulated-fishing overcapacity fishing-down-the-food-web fishing-through-the-food-web serial-depletion shifting-baseline baseline reference-point target-reference-point limit-reference-point threshold tipping-point point-of-no-return hysteresis alternative-stable-state regime-shift critical-transition catastrophic-shift smooth-transition adaptive-cycle panarchy resilience-thinking social-ecological-system SES coupled-human-natural-system CHANS human-environment-system HES socio-ecological-system ecosystem-approach-to-fisheries EAF ecosystem-based-fisheries-management EBFM EBM integrated-ecosystem-assessment IEA marine-spatial-planning MSP ocean-zoning marine-planning coastal-zone-management integrated-coastal-zone-management ICZM integrated-maritime-policy IMP blue-growth sustainable-development-goal SDG SDG-14 life-below-water decade-of-ocean-science UN-Ocean-Decade 2030-Agenda Agenda-21 Rio-Declaration Stockholm-Declaration Johannesburg-Plan-of-Implementation World-Summit-on-Sustainable-Development WSSD Rio+20 Rio+30 Earth-Summit UNCED UNCLOS Law-of-the-Sea-Convention Convention-on-the-Law-of-the-Sea high-seas area-beyond-national-jurisdiction ABNJ international-seabed-area the-Area exclusive-economic-zone EEZ territorial-sea contiguous-zone archipelagic-waters internal-waters straight-baseline normal-baseline low-water-line tidal-datum chart-datum depth-contour isobath bathymetry topography morphology geomorphology seafloor seabed benthos nekton plankton neuston pleuston hyperbenthos suprabenthos epibenthos endobenthos infauna epifauna pelagic benthopelagic meroplankton holoplankton zooplankton phytoplankton bacterioplankton virioplankton ichthyoplankton ichthyoplankton-survey egg-survey larval-survey ichthyoplankton-transport larval-transport dispersal-kernel connectivity population-connectivity genetic-connectivity demographic-connectivity larval-pool source-sink-dynamics metapopulation population-structure stock-structure stock-identification fishery-assessment biological-reference-point BRP maximum-sustainable-yield MSY optimal-yield OY maximum-economic-yield MEY maximum-constant-yield MCY equilibrium-yield sustainable-yield surplus-production Schaefer-model Fox-model Pella-Tomlinson-model surplus-production-model analytical-model statistical-catch-at-age-model SCA virtual-population-analysis VPA cohort-analysis catch-curve length-based-model age-based-model stage-based-model size-based-model trait-based-model individual-based-model IBM agent-based-model ABM ecosystem-model end-to-end-model EwE Atlantis OSMOSE InVEST Marxan prioritization systematic-conservation-planning SCP gap-analysis representativeness adequacy effectiveness equity fairness legitimacy compliance enforcement monitoring surveillance observation data-collection sampling survey census index indicator metric variable parameter estimate prediction forecast projection scenario what-if-analysis sensitivity-analysis uncertainty-analysis risk-analysis decision-analysis trade-off-analysis optimization multi-objective-optimization Pareto-front Pareto-optimal non-dominated-solution stakeholder interest-group rights-holder user-group community indigenous-people local-knowledge traditional-knowledge TK traditional-ecological-knowledge TEK citizen-science participatory-science community-based-monitoring CBM co-management joint-management shared-governance adaptive-co-management transboundary-management regional-seas regional-fishery-body RFB regional-fishery-management-organization RFMO arrangement commission council committee working-group expert-group scientific-committee technical-committee compliance-committee dispute-settlement arbitration conciliation mediation negotiation bargaining coalition alliance partnership network platform forum conference meeting session symposium workshop seminar training capacity-building technology-transfer technical-assistance financial-assistance funding grant loan donation collaboration cooperation coordination harmonization standardization best-practice guideline manual handbook code-of-conduct code-of-practice certification eco-label seal-of-approval sustainability-standard marine-stewardship-council MSC aquaculture-stewardship-council ASC friend-of-the-sea global-G.A.P.BRC IFS SQF FSSC-22000 ISO-22000 HACCP GMP SSOP traceability chain-of-custody blockchain digital-traceability electronic-monitoring EM electronic-reporting ER VMS AIS satellite-monitoring remote-sensing earth-observation geographic-information-system GIS global-positioning-system GPS automatic-identification-system vessel-monitoring-system catch-documentation-scheme CDS port-state-measures catch-certification trade-tracking isotope-analysis otolith-microchemistry parasite-tag tagging mark-recapture telemetry acoustic-telemetry satellite-telemetry archival-tag data-storage-tag DST pop-up-satellite-archival-tag PSAT smart-tag biologging bio-telemetry sensor accelerometer magnetometer gyroscope pressure-sensor temperature-sensor light-sensor dissolved-oxygen-sensor salinity-sensor conductivity-sensor pH-sensor chlorophyll-sensor turbidity-sensor acoustic-Doppler-current-profiler ADCP echosounder multibeam-echosounder single-beam-echosounder split-beam-echosounder scientific-echosounder fishing-echosounder sonar side-scan-sonar synthetic-aperture-sonar SAS sub-bottom-profiler seismic-profiler gravimeter bathymetric-lidar airborne-lidar satellite-altimetry radar-altimetry laser-altimetry photogrammetry structure-from-motion SfM underwater-photogrammetry video-analysis image-analysis machine-learning artificial-intelligence AI deep-learning neural-network computer-vision pattern-recognition object-detection instance-segmentation semantic-segmentation classification clustering regression forecasting time-series-analysis spatial-analysis geostatistics kriging inverse-distance-weighting IDW triangulated-irregular-network TIN digital-elevation-model DEM digital-terrain-model DTM digital-surface-model DSM habitat-suitability-model species-distribution-model SDM ecological-niche-model ENM habitat-suitability-index HSI environmental-suitability bioclimatic-envelope climate-envelope fundamental-niche realized-niche potential-distribution actual-distribution range-map distribution-map occurrence-record presence-only-data presence-absence-data abundance-data density-data effort-data detection-probability occupancy-model N-mixture-model distance-sampling line-transect point-transect strip-transect plotless-sampling quadrat transect stratified-random-sampling systematic-sampling cluster-sampling adaptive-cluster-sampling two-phase-sampling double-sampling multi-phase-sampling sampling-design sample-size statistical-power effect-size significance-level confidence-level confidence-interval credible-interval prediction-interval tolerance-interval hypothesis-test null-hypothesis alternative-hypothesis type-I-error type-II-error false-positive false-negative sensitivity specificity accuracy precision recall F1-score ROC-curve AUC model-validation model-verification model-calibration model-selection model-comparison model-averaging ensemble-model multi-model-inference information-criterion AIC AICc BIC DIC WAIC LOOIC cross-validation k-fold-cross-validation leave-one-out-cross-validation bootstrap jackknife permutation-test randomization-test Monte-Carlo-simulation Markov-chain-Monte-Carlo MCMC Bayesian-inference frequentist-inference likelihood posterior-probability prior-probability prior-distribution posterior-distribution conjugate-prior non-informative-prior weakly-informative-prior hierarchical-model mixed-effects-model random-effects-model fixed-effects-model generalized-linear-model GLM generalized-additive-model GAM generalized-linear-mixed-model GLMM generalized-additive-mixed-model GAMM structural-equation-model SEM path-analysis confirmatory-factor-analysis CFA exploratory-factor-analysis EFA principal-component-analysis PCA principal-coordinate-analysis PCoA non-metric-multidimensional-scaling NMDS correspondence-analysis CA detrended-correspondence-analysis DCA redundancy-analysis RDA canonical-correspondence-analysis CCA partial-least-squares PLS orthogonal-partial-least-squares OPLS artificial-neural-network ANN random-forest RF gradient-boosting-machine GBM XGBoost LightGBM CatBoost support-vector-machine SVM k-nearest-neighbor KNN naive-Bayes decision-tree regression-tree classification-tree bagging boosting stacking ensembling voting blending meta-learning transfer-learning domain-adaptation few-shot-learning zero-shot-learning self-supervised-learning unsupervised-learning supervised-learning semi-supervised-learning reinforcement-learning active-learning online-learning incremental-learning lifelong-learning continual-learning federated-learning distributed-learning edge-computing cloud-computing high-performance-computing HPC parallel-computing GPU-computing quantum-computing neuromorphic-computing DNA-computing biological-computing natural-computing swarm-intelligence ant-colony-optimization particle-swarm-optimization genetic-algorithm evolutionary-algorithm differential-evolution simulated-annealing tabu-search variable-neighborhood-search iterated-local-search guided-local-search greedy-randomized-adaptive-search-procedure GRASP scatter-search path-relinking memetic-algorithm hyper-heuristic meta-heuristic optimization-heuristic constructive-heuristic improvement-heuristic local-search neighborhood-search large-neighborhood-search LNS adaptive-large-neighborhood-search ALNS branch-and-bound branch-and-cut branch-and-price column-generation Benders-decomposition Dantzig-Wolfe-decomposition Lagrangian-relaxation surrogate-relaxation dual-ascent primal-dual-method interior-point-method simplex-method revised-simplex-method dual-simplex-method network-simplex-method barrier-method penalty-method augmented-Lagrangian-method sequential-quadratic-programming SQP sequential-linear-programming SLP trust-region-method line-search-method gradient-descent steepest-descent conjugate-gradient quasi-Newton-method Newton-method Gauss-Newton-method Levenberg-Marquardt-method trust-region-reflective-method interior-point-reflective-method active-set-method projected-gradient-method proximal-gradient-method accelerated-gradient-method momentum-method Nesterov-acceleration AdaGrad RMSprop Adam AdamW SGD stochastic-gradient-descent mini-batch-gradient-descent batch-gradient-descent full-gradient-descent variance-reduction SAG SAGA SVRG L-BFGS limited-memory-BFGS BFGS DFP SR1 Nelder-Mead-method Powell's-method Hooke-Jeeves-method pattern-search direct-search derivative-free-optimization black-box-optimization bayesian-optimization Gaussian-process kriging-surrogate response-surface-method RSM polynomial-chaos-expansion PCE sparse-grid sparse-polynomial-approximation low-rank-approximation tensor-decomposition CP-decomposition Tucker-decomposition tensor-train TT hierarchical-Tucker HT matrix-factorization non-negative-matrix-factorization NMF singular-value-decomposition SVD independent-component-analysis ICA factor-analysis latent-variable-model latent-class-model finite-mixture-model hidden-Markov-model HMM state-space-model dynamic-linear-model DLM Kalman-filter particle-filter sequential-Monte-Carlo SMC ensemble-Kalman-filter EnKF unscented-Kalman-filter UKF extended-Kalman-filter EKF Rauch-Tung-Striebel-smoother RTS-smoother forward-backward-algorithm Viterbi-algorithm Baum-Welch-algorithm expectation-maximization EM-algorithm variational-inference variational-Bayes mean-field-approximation loopy-belief-propagation sum-product-algorithm max-product-algorithm junction-tree-algorithm bucket-elimination variable-elimination conditioning cutset-conditioning constraint-satisfaction CSP satisfiability SAT MAX-SAT pseudo-Boolean-optimization integer-linear-programming ILP mixed-integer-linear-programming MILP mixed-integer-nonlinear-programming MINLP quadratic-programming QP quadratically-constrained-quadratic-programming QCQP second-order-cone-programming SOCP semidefinite-programming SDP geometric-programming GP signomial-programming SP difference-of-convex-programming DC-programming complementary-geometric-programming CGP generalized-geometric-programming GGP posynomial monomial signomial polynomial rational-function algebraic-function transcendental-function analytic-function meromorphic-function entire-function holomorphic-function harmonic-function subharmonic-function superharmonic-function plurisubharmonic-function plurisuperharmonic-function convex-function concave-function quasiconvex-function quasiconcave-function log-convex-function log-concave-function geometrically-convex-function multiplicatively-convex-function operator-convex-function matrix-convex-function spectral-function unitarily-invariant-function orthogonally-invariant-function permutation-invariant-function symmetric-function Schur-convex-function Schur-concave-function majorization weak-majorization strong-majorization doubly-stochastic-matrix Birkhoff-polytope permutahedron transportation-polytope network-flow minimum-cost-flow maximum-flow minimum-cut shortest-path longest-path traveling-salesman-problem TSP vehicle-routing-problem VRP capacitated-VRP CVRP VRP-with-time-windows VRPTW pickup-and-delivery-problem PDP dial-a-ride-problem DARP arc-routing-problem ARP Chinese-postman-problem CPP rural-postman-problem RPP location-problem facility-location p-median-problem p-center-problem uncapacitated-facility-location UFL capacitated-facility-location CFL warehouse-location plant-location hub-location median-problem center-problem covering-problem set-covering vertex-covering edge-covering dominating-set independent-set clique graph-coloring chromatic-number clique-number independence-number domination-number matching perfect-matching maximum-matching b-matching f-factor g-factor network-design Steiner-tree Steiner-forest prize-collecting-Steiner-tree k-MST minimum-spanning-tree MST maximum-spanning-tree minimum-arborescence minimum-directed-spanning-tree minimum-Steiner-tree approximation-algorithm approximation-ratio performance-guarantee inapproximability NP-hardness NP-completeness computational-complexity time-complexity space-complexity polynomial-time exponential-time quasi-polynomial-time sub-exponential-time fixed-parameter-tractable FPT W-hierarchy parameterized-complexity kernelization kernel treewidth pathwidth branchwidth cliquewidth rankwidth booleanwidth modularwidth split-decomposition modular-decomposition prime-graph cograph permutation-graph interval-graph chordal-graph perfect-graph comparability-graph co-comparability-graph AT-free-graph claw-free-graph line-graph planar-graph outerplanar-graph series-parallel-graph treewidth-bounded-graph minor-free-graph forbidden-minor forbidden-topological-minor forbidden-induced-subgraph graph-minor-theorem Robertson-Seymour-theorem well-quasi-ordering Wagner's-conjecture Hadwiger's-conjecture Four-Color-Theorem Strong-Perfect-Graph-Theorem Erdős–Faber–Lovász-conjecture Gyárfás–Sumner-conjecture Erdős–Hajnal-conjecture Caccetta–Häggkvist-conjecture Bermond–Thomassen-conjecture Thomassen's-conjecture Tutte's-conjecture Grötzsch's-theorem Brooks'-theorem Vizing's-theorem Shannon's-theorem König's-theorem Kőnig's-line-coloring-theorem Menger's-theorem max-flow-min-cut-theorem Ford-Fulkerson-theorem Edmonds'-matching-theorem Tutte's-theorem Petersen's-theorem Hall's-marriage-theorem Dilworth's-theorem Mirsky's-theorem Sperner's-theorem Erdős–Ko–Rado-theorem Kruskal–Katona-theorem Lovász's-perfect-graph-theorem Chudnovsky's-strong-perfect-graph-theorem Seymour's-decomposition-theorem Robertson–Seymour–Thomas-theorem Heawood-conjecture Ringel–Youngs-theorem Map-Color-Theorem Appel–Haken-proof Robertson–Seymour–Sanders–Thomas-proof computer-assisted-proof formal-proof interactive-theorem-proving proof-assistant Coq Isabelle HOL HOL-Light Lean Agda Twelf Mizar Metamath automated-theorem-proving SAT-solver SMT-solver model-checker temporal-logic CTL LTL CTL*modal-mu-calculus fixed-point-logic description-logic OWL RDF SPARQL knowledge-graph ontology semantic-web linked-data open-data FAIR-data findable accessible interoperable reusable data-management-plan DMP metadata provenance data-quality data-curation data-preservation data-archiving data-repository institutional-repository disciplinary-repository general-repository figshare Zenodo Dryad GBIF OBIS WoRMS Catalogue-of-Life COL ITIS NCBI BOLD iNaturalist eBird eButterfly iDigBio ALA AVH BIE BISON Map-of-Life MOL Movebank OTN SAHFOS CPR ICES NAFO NEAFC SEAFO WCPFC IATTC ICCAT IOTC CCSBT CCAMLR GFCM CWP CWP-15 CWP-16 CWP-17 CWP-18 CWP-19 CWP-20 CWP-21 CWP-22 CWP-23 CWP-24 CWP-25 CWP-26 CWP-27 CWP-28 CWP-29 CWP-30 CWP-31 CWP-32 CWP-33 CWP-34 CWP-35 CWP-36 CWP-37 CWP-38 CWP-39 CWP-40 CWP-41 CWP-42 CWP-43 CWP-44 CWP-45 CWP-46 CWP-47 CWP-48 CWP-49 CWP-50 CWP-51 CWP-52 CWP-53 CWP-54 CWP-55 CWP-56 CWP-57 CWP-58 CWP-59 CWP-60 CWP-61 CWP-62 CWP-63 CWP-64 CWP-65 CWP-66 CWP-67 CWP-68 CWP-69 CWP-70 CWP-71 CWP-72 CWP-73 CWP-74 CWP-75 CWP-76 CWP-77 CWP-78 CWP-79 CWP-80 CWP-81 CWP-82 CWP-83 CWP-84 CWP-85 CWP-86 CWP-87 CWP-88 CWP-89 CWP-90 CWP-91 CWP-92 CWP-93 CWP-94 CWP-95 CWP-96 CWP-97 CWP-98 CWP-99 CWP-100 CWP-101 CWP-102 CWP-103 CWP-104 CWP-105 CWP-106 CWP-107 CWP-108 CWP-109 CWP-110 CWP-111 CWP-112 CWP-113 CWP-114 CWP-115 CWP-116 CWP-117 CWP-118 CWP-119 CWP-120 CWP-121 CWP-122 CWP-123 CWP-124 CWP-125 CWP-126 CWP-127 CWP-128 CWP-129 CWP-130 CWP-131 CWP-132 CWP-133 CWP-134 CWP-135 CWP-136 CWP-137 CWP-138 CWP-139 CWP-140 CWP-141 CWP-142 CWP-143 CWP-144 CWP-145 CWP-146 CWP-147 CWP-148 CWP-149 CWP-150 CWP-151 CWP-152 CWP-153 CWP-154 CWP-155 CWP-156 CWP-157 CWP-158 CWP-159 CWP-160 CWP-161 CWP-162 CWP-163 CWP-164 CWP-165 CWP-166 CWP-167 CWP-168 CWP-169 CWP-170 CWP-171 CWP-172 CWP-173 CWP-174 CWP-175 CWP-176 CWP-177 CWP-178 CWP-179 CWP-180 CWP-181 CWP-182 CWP-183 CWP-184 CWP-185 CWP-186 CWP-187 CWP-188 CWP-189 CWP-190 CWP-191 CWP-192 CWP-193 CWP-194 CWP-195 CWP-196 CWP-197 CWP-198 CWP-199 CWP-200
Rothschildia lebeau
Lebeau's Silk Moth, Forbes Moth
Rothschildia lebeau is a giant silkmoth in the family Saturniidae, distributed from the Lower Rio Grande Valley through eastern Mexico to Brazil. Adults are short-lived and do not feed, relying on fat reserves accumulated during larval development. The species exhibits a life history tradeoff between larval survival and adult body size based on host plant quality, with larger males showing higher mating success.
giant-silkmoth Saturniidae host-plant-tradeoff sexual-selection tropical-dry-forest Barro-Colorado-Island pheromone-communication non-feeding-adult complete-metamorphosis cultural-significance indigenous-use cocoon-harvesting mating-success body-size-dimorphism seasonal-variation microbial-associations protistan-diversity caterpillar-microbiome herbivore forest-ecology Neotropical Mexico Central-America South-America Panama Costa-Rica Brazil Texas Rio-Grande-Valley exhibit-species butterfly-house education conservation light-pollution suburban-sprawl introduced-parasites rearing cottage-industry vestigial-mouthparts fat-reserves short-lived-adult female-sedentary-behavior male-dispersal oviposition-site-selection fitness-consequences life-history-evolution quantitative-genetic-variation wet-season-phenology developmental-plasticity survival-size-tradeoff herbivore-performance plant-animal-interactions microbial-observatory molecular-characterization eukaryotic-microbiome saturniid-caterpillar Lepidoptera Bombycoidea Rothschildia Guérin-Méneville-1868 Attacus-lebeau Lebeau's-Silk-Moth Forbes-Moth Rothschildia-lebeau-forbesi windows transparent-wing-patches scale-less-wing-areas antennae sexual-dimorphism mate-choice sperm-competition reproductive-success larval-performance caterpillar-growth pupal-development adult-emergence longevity predation bat-predation nocturnal crepuscular moth silkmoth silkworm giant-moth forest-moth tropical-moth Neotropical-fauna biodiversity insect-collection museum-specimen Bohart-Museum American-Museum-of-Natural-History entomology lepidopterology systematics taxonomy natural-history field-study observational-study experimental-ecology evolutionary-ecology behavioral-ecology population-biology community-ecology ecosystem-function food-web trophic-interaction herbivory folivory tree-insect-interaction phenology climate-effects seasonality environmental-variation habitat-heterogeneity host-race ecotype local-adaptation phenotypic-plasticity genetic-variation quantitative-trait trade-off allocation reproductive-allocation parental-investment offspring-size offspring-number optimal-clutch-size mate-competition intrasexual-selection intersexual-selection ornament signal chemical-communication olfaction sensory-ecology foraging movement-ecology dispersal migration home-range activity-pattern diel-cycle circadian-rhythm photoperiod temperature humidity rainfall precipitation dry-season wet-season tropical-seasonality El-Niño climate-change global-warming habitat-disturbance forest-fragmentation deforestation agricultural-expansion urbanization conservation-biology threatened-species common-species population-decline extinction-risk protected-area national-park biological-reserve research-station Smithsonian-Tropical-Research-Institute Barro-Colorado-Island-Nature-Monument Gatun-Lake Panama-Canal Isthmus-of-Panama Central-American-isthmus land-bridge biogeography vicariance dispersal-limitation species-range range-expansion range-contraction range-shift poleward-shift elevational-shift altitudinal-gradient latitudinal-gradient thermal-tolerance desiccation-resistance water-balance physiological-ecology metabolic-rate energetics nutritional-ecology stoichiometry nitrogen phosphorus carbon plant-secondary-metabolite tannin alkaloid terpenoid defense induced-defense constitutive-defense herbivore-offense counter-adaptation coevolution evolutionary-arms-race phylogenetic-constraint phylogenetic-signal comparative-method phylogenetic-comparative-analysis trait-evolution reproductive-strategy iteroparity semelparity big-bang-reproduction suicidal-reproduction capital-breeder income-breeder resource-acquisition resource-allocation storage fat-body lipid glycogen protein amino-acid vitamin micronutrient digestion gut-microbiome gut-flora intestinal-microorganism symbiosis mutualism commensalism parasitism pathogen disease immunity immune-response defense-mechanism resistance tolerance recovery survival mortality fecundity fertility sterility hybridization introgression gene-flow population-structure genetic-differentiation FST isolation-by-distance isolation-by-environment landscape-genetics landscape-ecology spatial-ecology geographic-information-system remote-sensing satellite-imagery habitat-suitability species-distribution-model ecological-niche-model MaxEnt BIOMOD ensemble-forecast climate-envelope niche-conservatism niche-evolution realized-niche fundamental-niche biotic-interaction competition facilitation apparent-competition indirect-effect trophic-cascade keystone-species ecosystem-engineer foundation-species dominant-species rare-species functional-diversity functional-trait trait-based-ecology community-assembly filtering sorting neutral-theory niche-theory unified-theory metacommunity metapopulation source-sink-dynamics rescue-effect mass-effect patch-dynamics landscape-mosaic connectivity corridor stepping-stone matrix-quality edge-effect interior-forest core-area buffer-zone fragment-size isolation perimeter-area-ratio shape-index fractal-dimension habitat-amount threshold-effect extinction-debt relaxation-time lag-effect time-delay historical-legacy founder-effect bottleneck genetic-drift inbreeding inbreeding-depression outbreeding outbreeding-depression heterosis hybrid-vigor genetic-rescue assisted-migration managed-relocation translocation reintroduction restoration-ecology habitat-restoration ecological-restoration rewilding de-extinction synthetic-biology conservation-genetics conservation-genomics population-genomics landscape-genomics environmental-DNA eDNA metabarcoding metagenomics transcriptomics proteomics metabolomics phenomics imaging computer-vision machine-learning artificial-intelligence deep-learning neural-network convolutional-neural-network recurrent-neural-network generative-adversarial-network transfer-learning data-augmentation training-set validation-set test-set cross-validation bootstrap jackknife permutation randomization Monte-Carlo Markov-chain Bayesian-inference posterior-probability prior-probability likelihood model-selection information-criterion AIC BIC DIC WAIC LOO model-averaging multimodel-inference structural-equation-model path-analysis causal-inference directed-acyclic-graph potential-outcomes propensity-score instrumental-variable regression-discontinuity difference-in-differences synthetic-control panel-data longitudinal-data time-series cross-sectional-data spatial-data spatiotemporal-data point-pattern areal-data geostatistics kriging variogram spatial-autocorrelation Moran's-I Geary's-c Getis-Ord-Gi LISA hot-spot cold-spot cluster-detection scan-statistic network-analysis graph-theory small-world scale-free-network centrality betweenness closeness eigenvector PageRank community-detection modularity clustering-coefficient path-length diameter radius percolation robustness resilience vulnerability adaptive-capacity transformability social-ecological-system complex-adaptive-system emergent-property self-organization nonlinearity threshold tipping-point hysteresis alternative-stable-state regime-shift critical-transition early-warning-signal resilience-indicator panarchy adaptive-cycle exploitation release reorganization fast-variable slow-variable controlling-variable driver pressure state impact response DPSIR ecosystem-service provisioning-service regulating-service cultural-service supporting-service natural-capital green-infrastructure nature-based-solution ecosystem-based-adaptation ecosystem-based-mitigation payment-for-ecosystem-service REDD+biodiversity-offset habitat-banking mitigation-hierarchy avoidance minimization restoration compensation no-net-loss net-gain biodiversity-metric condition-metric landscape-metric connectivity-metric genetic-metric functional-metric irreplaceability threat opportunity conservation-planning systematic-conservation-planning Marxan Zonation prioritization efficiency effectiveness equity stakeholder governance institution policy law regulation incentive market-based-instrument command-and-control voluntary-agreement certification eco-label sustainable-use sustainable-management adaptive-management learning-by-doing monitoring evaluation indicator target goal vision scenario projection forecast uncertainty risk precautionary-principle safe-minimum-standard environmental-impact-assessment strategic-environmental-assessment cumulative-impact-assessment social-impact-assessment health-impact-assessment life-cycle-assessment carbon-footprint water-footprint ecological-footprint material-flow-analysis substance-flow-analysis input-output-analysis environmentally-extended-input-output-analysis multi-regional-input-output-analysis computable-general-equilibrium integrated-assessment-model earth-system-model global-circulation-model regional-climate-model downscaling bias-correction ensemble-simulation perturbed-physics emulator surrogate-model reduced-complexity-model simple-climate-model integrated-earth-system-model socioeconomic-scenario shared-socioeconomic-pathway representative-concentration-pathway climate-scenario emission-scenario mitigation-scenario adaptation-scenario impact-scenario vulnerability-scenario risk-scenario resilience-scenario transformation-scenario sustainability-scenario narrative storyline pathway trajectory transition transformation decarbonization electrification renewable-energy energy-efficiency circular-economy green-economy blue-economy bioeconomy nature-economy natural-capital-accounting ecosystem-accounting environmental-accounting green-accounting inclusive-wealth genuine-progress-indicator happy-planet-index human-development-index sustainable-development-goals Agenda-2030 Paris-Agreement Convention-on-Biological-Diversity Aichi-targets post-2020-global-biodiversity-framework Kunming-Montreal-Global-Biodiversity-Framework 30-by-30 half-earth nature-needs-half rewilding-Europe rewilding-Britain rewilding-North-America Pleistocene-rewilding trophic-rewilding ecological-replacement taxon-substitution revival-biology molecular-de-extinction proxy-species gene-drive CRISPR genome-editing genetic-modification biotechnology nanotechnology information-technology global-positioning-system unmanned-aerial-vehicle drone satellite sensor camera-trap acoustic-monitoring citizen-science community-science participatory-monitoring traditional-ecological-knowledge indigenous-knowledge local-knowledge farmer-knowledge fishers-knowledge co-production-of-knowledge transdisciplinarity interdisciplinarity multidisciplinarity post-normal-science mode-2-science triple-helix quadruple-helix quintuple-helix open-science open-data open-access open-source FAIR-data findable accessible interoperable reusable data-management-plan metadata provenance quality-assurance quality-control validation verification uncertainty-quantification error-propagation sensitivity-analysis scenario-analysis Monte-Carlo-simulation Bayesian-belief-network influence-diagram decision-tree multi-criteria-decision-analysis cost-benefit-analysis cost-effectiveness-analysis multi-attribute-utility-theory game-theory mechanism-design incentive-compatibility revelation-principle social-choice-theory voting-theory fair-division institutional-design policy-design governance-design architecture polycentric-governance adaptive-governance collaborative-governance network-governance hierarchical-governance market-governance community-governance hybrid-governance meta-governance steering rowing regulatory-state enabling-state partner-state entrepreneurial-state innovation-state strategic-state developmental-state welfare-state green-state sustainable-state resilient-state smart-state digital-state platform-state algorithmic-state surveillance-state authoritarian-state democratic-state deliberative-democracy participatory-democracy direct-democracy representative-democracy liberal-democracy social-democracy green-democracy ecological-democracy biocracy ecofascism deep-ecology social-ecology ecofeminism environmental-justice climate-justice intergenerational-justice intragenerational-justice spatial-justice territorial-justice recognition-justice procedural-justice distributive-justice restorative-justice transformative-justice reparative-justice transitional-justice ecological-justice species-justice multispecies-justice more-than-human-justice anthropocene capitalocene plantationocene chthulucene symbiocene good-anthropocene bad-anthropocene great-acceleration great-divergence great-convergence great-transformation great-decoupling great-disruption great-moderation great-recession great-depression great-war great-famine great-migration great-replacement great-reset great-refusal great-enrichment great-leveling great-compression
Tortopus
Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.
Ephemeroptera Polymitarcyidae burrowing-mayflies Neotropical clay-bank-habitat U-shaped-tunnels mandibular-tusks Tortopsis Campsurinae Panamerican-distribution cladistic-revision synapomorphy parastyli-receptors flattened-penes furcasternal-protuberances subapical-tubercles limnological-sampling-bias nymph-morphology adult-morphology type-species-T.-igaranus new-species-T.-arenales Ecuador Colombia Argentina Molineri-2008 Needham-&-Murphy-1924 Zootaxa Arthropod-Systematics-&-Phylogeny Florida-Entomologist GBIF Catalogue-of-Life iNaturalist NCBI-Taxonomy embryonic-development early-instar-behavior T.-harrisi T.-bellus T.-circumfluus T.-zottai T.-igaranus T.-incertus T.-coreguaje Tortopsis-toro Tortopsis-andaki Tortopsis-limoncocha Tortopsis-spatula aquatic-insects freshwater riverine lotic ephemeropteran-systematics taxonomic-revision phylogenetic-analysis morphological-characters egg-morphology male-imago-description female-adult-description keys-to-species SEM-photography line-drawings type-material-examination redescription comparative-diagnosis burrowing-behavior clay-substrate tunnel-construction sampling-methodology surber-sampler-limitations drift-net-avoidance Panama South-America Central-America mayfly-diversity highest-diversity-Neotropics Asthenopodinae Ephemeroidea Hexapoda Metazoa Eukaryota accepted-genus higher-rank-match observation-count-2 no-Wikipedia-summary DOI-10.11646/zootaxa.2481.1.1 DOI-10.11646/zootaxa.2436.1.4 DOI-10.3897/asp.79.e62735 DOI-10.2307/3493493 Scott-et-al.-1959 Navás Ulmer Traver Lugo-Ortiz-&-McCafferty Domínguez Banks McDunnough Pictet Needham-&-Murphy freshwater-ecology aquatic-entomology insect-development nymphal-stages imago-description alate-adults wing-venation R-sector gonopore-structure parastyli-length pedestal-morphology lateral-view-curvature mandibular-morphology tusk-tubercles mesosternal-structure abdominal-sternum-notch furcasternum protuberance-configuration receptor-socket-orientation IR-vein R2-vein claw-like-structure gonopore-association penis-base-separation female-fore-wing additional-veins comparative-morphology type-specimen holotype allotype paratype neotropical-species new-species-description species-key identification-key male-adult-key female-adult-key nymph-key generic-key Polymitarcyidae-genera systematics phylogeny cladistics monophyly sister-group character-state transformation-series parsimony-analysis morphological-data-set external-morphology egg-chorion micropyle egg-burster hatching emergence behavioral-study early-instar embryology oviposition substrate-preference habitat-specificity endemic regional-diversity conservation biomonitoring water-quality-indicator macroinvertebrate benthic infaunal interstitial hyporheic river-bank stream-bank erosion sediment particle-size clay-content bank-stability root-mat vegetation-association flow-regime current-velocity depth width stream-order catchment basin ecoregion biogeography dispersal colonization vicariance speciation adaptive-radiation morphological-divergence convergent-evolution homoplasy character-conflict bootstrap-support Bremer-support consensus-tree strict-consensus majority-rule taxonomic-stability nomenclatural-act original-description subsequent-designation type-locality type-depository museum-collection specimen-preservation alcohol-preservation slide-mounting SEM-preparation imaging-techniques scientific-illustration photography measurement morphometrics ratios proportions allometry sexual-dimorphism geographic-variation intraspecific-variation interspecific-variation diagnostic-characters identification-confidence taxonomic-expertise literature-review bibliographic-search database-query online-repository open-access paywall abstract-only full-text-unavailable citation-analysis reference-list source-quality evidence-strength knowledge-gap research-need future-study collaboration field-work collection-trip sampling-effort specimen-availability type-examination revision-scope taxonomic-concept circumscription inclusion exclusion species-group species-complex cryptic-species morphospecies phylogenetic-species evolutionary-significant-unit operational-taxonomic-unit DNA-barcoding molecular-systematics integrative-taxonomy traditional-taxonomy alpha-taxonomy beta-taxonomy gamma-taxonomy classification-hierarchy Linnaean-system rank-based-nomenclature phylogenetic-nomenclature clade-name crown-group stem-group total-evidence congruence incongruence data-partition character-weighting implied-weighting equal-weighting ordered-characters unordered-characters multistate-characters binary-characters continuous-characters meristic-characters qualitative-characters quantitative-characters morphological-matrix data-matrix character-matrix taxon-sampling character-sampling missing-data inapplicable-data polymorphism intraspecific-polymorphism fixed-difference diagnostic-difference unique-derived-character autapomorphy shared-derived-character symplesiomorphy convergent-character parallel-evolution reversal character-optimization ACCTRAN DELTRAN unambiguous-optimization ambiguous-optimization most-parsimonious-tree tree-length consistency-index retention-index rescaled-consistency-index homoplasy-index character-fit character-congruence character-support branch-support node-support clade-stability taxonomic-stability-index phylogenetic-signal noise saturation substitution-model rate-heterogeneity among-site-rate-variation gamma-distribution invariant-sites model-selection Akaike-information-criterion Bayesian-information-criterion likelihood-ratio-test Bayesian-inference Markov-chain-Monte-Carlo posterior-probability prior-probability hyperprior convergence effective-sample-size burn-in stationarity trace-plot parameter-estimate credible-interval highest-posterior-density tree-topology tree-shape lineage-through-time diversification-rate extinction-rate speciation-rate net-diversification birth-death-model Yule-model coalescent-model population-size effective-population-size gene-flow migration-rate isolation-by-distance phylogeography historical-biogeography ancestral-area-reconstruction dispersal-vicariance-analysis decisive-weighting event-based-method taxon-area-cladogram area-cladogram biogeographic-homology track-analysis panbiogeography phylogenetic-biogeography ecological-biogeography dispersal-biogeography vicariance-biogeography center-of-origin area-of-endemism refugium Pleistocene glaciation climate-change habitat-fragmentation habitat-loss anthropogenic-impact pollution eutrophication sedimentation dam-construction water-abstraction flow-modification thermal-pollution toxic-substances heavy-metals pesticides urbanization agriculture deforestation land-use-change riparian-degradation channelization dredging mining invasive-species biological-invasion competition predation parasitism mutualism commensalism amensalism facilitation ecosystem-engineer keystone-species flagship-species umbrella-species indicator-species sentinel-species bioindicator ecological-indicator environmental-indicator water-quality-index biotic-index multimetric-index rapid-bioassessment protocol standardization quality-assurance quality-control reference-condition least-disturbed minimally-disturbed best-available type-specific region-specific stream-type river-type functional-feeding-group collector-gatherer collector-filterer scraper shredder predator parasite engulfer piercer burrower sprawler clinger swimmer skater diver surface-dweller benthic-dweller interstitial-dweller hyporheic-dweller phreatic-dweller troglobite stygobite cave-inhabitant spring-inhabitant headwater mid-order high-order lowland montane alpine subtropical tropical temperate boreal arctic antarctic palearctic nearctic afrotropical oriental australian oceanic cosmopolitan widespread restricted relict disjunct-distribution range-extension new-record first-record country-record state-record province-record locality-record elevation-record altitudinal-range latitudinal-range longitudinal-range distribution-pattern range-size range-fill extent-of-occurrence area-of-occupancy IUCN-category threat-status data-deficient not-evaluated least-concern near-threatened vulnerable endangered critically-endangered extinct-in-wild extinct population-trend population-density population-structure age-structure size-structure sex-ratio reproductive-output fecundity fertility mortality survivorship life-table cohort-analysis generation-time doubling-time intrinsic-rate-of-increase finite-rate-of-increase net-reproductive-rate stable-age-distribution stationary-population expanding-population declining-population source-sink-dynamics metapopulation population-viability-analysis minimum-viable-population genetic-diversity heterozygosity allelic-richness private-allele founder-effect bottleneck genetic-drift inbreeding inbreeding-depression outbreeding outbreeding-depression hybridization introgression gene-flow-barrier reproductive-isolation prezygotic-isolation postzygotic-isolation mechanical-isolation temporal-isolation habitat-isolation behavioral-isolation gametic-isolation zygotic-mortality hybrid-inviability hybrid-sterility hybrid-breakdown reinforcement character-displacement ecological-speciation allopatric-speciation sympatric-speciation parapatric-speciation peripatric-speciation centrifugal-speciation divergence-with-gene-flow speciation-continuum anagenesis cladogenesis budding-speciation branching-speciation phyletic-gradualism punctuated-equilibrium stasis trend progression orthogenesis aristogenesis mosaic-evolution co-evolution arms-race Red-Queen-hypothesis evolutionary-stable-strategy frequency-dependent-selection density-dependent-selection r-selection K-selection bet-hedging iteroparity semelparity ploidy-level chromosome-number karyotype genome-size transposable-element gene-duplication neofunctionalization subfunctionalization pseudogene regulatory-evolution cis-regulatory trans-regulatory enhancer promoter silencer insulator transcription-factor signaling-pathway developmental-pathway gene-network epigenetic-modification DNA-methylation histone-modification chromatin-remodeling genomic-imprinting parental-effect maternal-effect paternal-effect cytoplasmic-inheritance mitochondrial-DNA chloroplast-DNA endosymbiont Wolbachia Rickettsia Spiroplasma Cardinium Arsenophonus Hamiltonella Regiella Serratia Buchnera Carsonella Portiera primary-endosymbiont secondary-endosymbiont facultative-symbiont obligate-symbiont mutualistic-symbiont parasitic-symbiont commensal-symbiont pathogen vector disease-transmission zoonosis emerging-infectious-disease biosecurity quarantine eradication control management conservation-biology restoration-ecology habitat-restoration river-restoration stream-restoration floodplain-restoration riparian-restoration wetland-restoration ecosystem-restoration passive-restoration active-restoration assisted-recovery managed-relocation assisted-colonization ex-situ-conservation in-situ-conservation protected-area national-park nature-reserve wildlife-refuge biosphere-reserve world-heritage-site ramsar-site important-bird-area key-biodiversity-area Alliance-for-Zero-Extinction EDGE-species evolutionarily-distinct globally-endangered conservation-prioritization systematic-conservation-planning Marxan Zonation conservation-opportunity conservation-cost conservation-benefit trade-off socioeconomic-factor stakeholder-engagement community-based-conservation adaptive-management monitoring-program long-term-study citizen-science public-participation environmental-education outreach science-communication knowledge-transfer capacity-building institutional-strengthening policy-development legislation regulation enforcement compliance incentive payment-for-ecosystem-services carbon-credit biodiversity-offset mitigation-banking conservation-easement land-trust private-land-conservation conservation-covenant stewardship sustainable-use sustainable-management ecosystem-based-management integrated-water-resources-management basin-management catchment-management landscape-management seascape-management connectivity-conservation corridor stepping-stone buffer-zone matrix-management countryside-biogeography reconciliation-ecology novel-ecosystem designer-ecosystem ecosystem-service provisioning-service regulating-service supporting-service cultural-service food-production water-supply raw-materials genetic-resources medicinal-resources ornamental-resources air-quality-regulation climate-regulation water-regulation erosion-regulation water-purification waste-treatment disease-regulation pest-regulation pollination seed-dispersal habitat-provision nutrient-cycling soil-formation primary-production recreation tourism aesthetic-value spiritual-value educational-value scientific-value inspiration sense-of-place cultural-heritage traditional-knowledge indigenous-knowledge local-knowledge ecosystem-service-valuation economic-valuation contingent-valuation choice-experiment benefit-transfer total-economic-value use-value non-use-value direct-use indirect-use option-value bequest-value existence-value willingness-to-pay willingness-to-accept cost-benefit-analysis cost-effectiveness-analysis multi-criteria-analysis decision-support uncertainty risk-assessment scenario-analysis sensitivity-analysis robustness-analysis adaptive-capacity resilience vulnerability exposure sensitivity coping-capacity transformation social-ecological-system complex-adaptive-system tipping-point regime-shift alternative-stable-state hysteresis path-dependence legacy-effect slow-variable fast-variable panarchy adaptive-cycle exploitation release reorganization foresight anticipatory-governance transformative-change sustainability-science future-earth interdisciplinary transdisciplinary problem-oriented solution-oriented action-oriented knowledge-co-production boundary-work boundary-object boundary-organization science-policy-interface science-society-interface knowledge-brokerage knowledge-exchange knowledge-integration mode-1-science mode-2-science triple-helix quadruple-helix responsible-innovation reflexive-governance adaptive-governance collaborative-governance polycentric-governance network-governance self-organization collective-action common-pool-resource tragedy-of-the-commons social-dilemma institutional-analysis institutional-design Ostrom-design-principles monitoring sanctioning conflict-resolution nested-enterprises proportionality collective-choice constitutional-choice operational-choice collective-action-theory game-theory prisoner's-dilemma chicken-game assurance-game coordination-game evolutionary-game-theory replicator-dynamics ESS Nash-equilibrium Pareto-optimality social-optimum market-failure government-failure voluntary-failure non-market-value public-good common-good club-good private-good rivalry excludability free-rider hold-up-problem moral-hazard adverse-selection information-asymmetry transaction-cost property-right common-property state-property private-property hybrid-property bundle-of-rights usufruct access withdrawal alienation tenure-security land-tenure water-tenure resource-tenure customary-tenure statutory-tenure legal-pluralism de-facto de-jure formal informal legality legitimacy recognition deviance corruption elite-capture social-capital bonding-capital bridging-capital linking-capital trust reciprocity reputation social-network social-embeddedness structural-hole brokerage closure cohesion density centrality betweenness degree eigenvector PageRank small-world scale-free random-network regular-network hierarchical-network modularity community-structure clustering assortativity homophily heterophily preferential-attachment fitness-model network-evolution network-dynamics temporal-network multilayer-network multiplex-network bipartite-network affiliation-network one-mode-network two-mode-network adjacency-matrix edge-list graph-theory vertex edge node link tie connection path cycle tree forest component giant-component percolation phase-transition criticality self-organized-criticality power-law heavy-tail Pareto-distribution lognormal-distribution exponential-distribution Poisson-distribution binomial-distribution normal-distribution Gaussian-distribution statistical-distribution probability-distribution cumulative-distribution probability-density-function cumulative-distribution-function survival-function hazard-function moment mean median mode variance standard-deviation coefficient-of-variation skewness kurtosis quantile percentile decile quartile interquartile-range range minimum maximum outlier robust-statistic parametric-test non-parametric-test hypothesis-testing null-hypothesis alternative-hypothesis significance-level p-value confidence-interval type-I-error type-II-error statistical-power effect-size sample-size sampling-design experimental-design observational-study quasi-experiment natural-experiment randomized-controlled-trial before-after-control-impact BACI impact-assessment environmental-impact-assessment strategic-environmental-assessment cumulative-impact-assessment social-impact-assessment health-impact-assessment life-cycle-assessment carbon-footprint water-footprint ecological-footprint material-flow-analysis substance-flow-analysis energy-analysis exergy-analysis emergy-analysis input-output-analysis environmentally-extended-input-output social-accounting-matrix computable-general-equilibrium agent-based-model system-dynamics-model Bayesian-network structural-equation-model path-analysis factor-analysis principal-component-analysis cluster-analysis discriminant-analysis canonical-correspondence-analysis redundancy-analysis non-metric-multidimensional-scaling ordination classification regression-analysis linear-regression nonlinear-regression generalized-linear-model generalized-additive-model mixed-effects-model multilevel-model hierarchical-model Bayesian-hierarchical-model state-space-model hidden-Markov-model time-series-analysis spatial-analysis geostatistics kriging variogram spatial-autocorrelation Moran's-I Geary's-c Getis-Ord-Gi local-indicator-of-spatial-association LISA hot-spot cold-spot spatial-cluster spatial-outlier edge-effect boundary-effect modifiable-areal-unit-problem MAUP ecological-fallacy atomistic-fallacy cross-scale-inference upscaling downscaling scaling-law allometric-scaling fractal fractal-dimension self-similarity multi-fractal lacunarity roughness texture pattern process pattern-process-relationship neutral-model null-model randomization-test permutation-test bootstrap jackknife cross-validation model-averaging model-comparison information-criterion Bayesian-model-averaging ensemble-modeling consensus-model prediction forecast projection scenario narrative-scenario quantitative-scenario storyline shared-socioeconomic-pathway representative-concentration-pathway emission-scenario climate-scenario impact-scenario adaptation-scenario mitigation-scenario transformation-scenario backcasting forecasting trend-extrapolation expert-judgment Delphi-method participatory-modeling stakeholder-analysis influence-diagram conceptual-model mental-model system-archetype leverage-point intervention-point policy-instrument economic-instrument regulatory-instrument voluntary-instrument information-instrument market-based-instrument command-and-control-instrument subsidy tax charge fee levy deposit-refund tradable-permit emission-trading carbon-tax biodiversity-banking conditional-cash-transfer community-based-natural-resource-management joint-forest-management co-management adaptive-co-management transboundary-management international-regime global-governance multilateral-environmental-agreement convention treaty cartagena-protocol nagoya-protocol Aichi-target sustainable-development-goal SDG post-2020-global-biodiversity-framework Kunming-Montreal-global-biodiversity-framework 30-by-30-target nature-based-solution NbS ecosystem-based-adaptation ecosystem-based-mitigation green-infrastructure blue-infrastructure natural-infrastructure hybrid-infrastructure climate-smart-agriculture climate-smart-landscape climate-smart-development low-carbon-development green-economy circular-economy bioeconomy blue-economy sustainable-consumption sustainable-production life-cycle-thinking cradle-to-cradle industrial-ecology industrial-symbiosis urban-metabolism socio-ecological-metabolism material-intensity energy-intensity carbon-intensity decoupling absolute-decoupling relative-decoupling resource-efficiency eco-efficiency eco-innovation green-technology clean-technology environmental-technology appropriate-technology participatory-technology technology-transfer technology-diffusion innovation-system national-innovation-system sectoral-innovation-system technological-innovation-system socio-technical-system socio-technical-transition multi-level-perspective niche-innovation regime landscape transition-management strategic-niche-management transition-experiment living-lab real-world-laboratory urban-living-lab rural-living-lab sustainability-transition fundamental-change system-innovation directionality reflexivity responsiveness anticipation learning-by-doing doing-by-learning social-learning organizational-learning policy-learning transformative-capacity capacity-development institutional-learning evolutionary-governance institutional-change path-dependency lock-in carbon-lock-in technological-lock-in institutional-lock-in behavioral-lock-in cultural-lock-in unlocking destabilization phase-out divestment stranded-asset just-transition transition-pathway transition-scenario deep-decarbonization net-zero carbon-neutral climate-neutral nature-positive biodiversity-positive restoration-positive regenerative circular doughnut-economics planetary-boundary safe-operating-space environmental-ceiling social-foundation intergenerational-equity intragenerational-equity distributive-justice procedural-justice recognition-justice environmental-justice climate-justice energy-justice water-justice food-justice spatial-justice territorial-justice ecological-justice species-justice intersectionality adaptation mitigation sustainability sustainable-development well-being human-development capability-approach human-security environmental-security climate-security water-security food-security energy-security livelihood-security disaster-risk-reduction climate-change-adaptation climate-change-mitigation loss-and-damage climate-finance green-climate-fund adaptation-fund global-environment-facility biodiversity-finance nature-finance blended-finance impact-investment ESG environmental-social-governance sustainable-finance green-bond social-bond sustainability-bond blue-bond nature-bond biodiversity-credit offset inset nature-based-credit ecosystem-service-credit payment-for-results results-based-payment conditional-payment performance-based-payment outcome-based-payment environmental-service-payment PSA PES REDD REDD+AFOLU LULUCF NAMA NAMC NDC NAP NBSAP national-adaptation-plan national-biodiversity-strategy-and-action-plan nationally-determined-contribution nationally-appropriate-mitigation-action intended-nationally-determined-contribution long-term-low-emission-development-strategy net-zero-target carbon-budget remaining-carbon-budget emission-gap adaptation-gap finance-gap implementation-gap ambition-loop ratchet-mechanism global-stocktake transparency-framework enhanced-transparency-framework common-but-differentiated-responsibilities respective-capabilities climate-equity climate-litigation climate-governance climate-diplomacy climate-negotiation UNFCCC COP CMP CMA SBSTA SBI IPCC IPBES UNCCD CBD CITES Ramsar World-Heritage Man-and-the-Biosphere MAB UNESCO FAO WHO ILO UNDP UNEP UN-Habitat World-Bank IMF WTO OECD GEF GCF AF multilateral-development-bank regional-development-bank national-development-bank development-finance-institution export-credit-agency sovereign-wealth-fund pension-fund insurance-company asset-manager asset-owner corporate-investor retail-investor impact-investor philanthropic-investor development-partner bilateral-donor multilateral-donor private-donor foundation NGO CSO civil-society-organization non-governmental-organization community-based-organization indigenous-peoples-organization women's-organization youth-organization faith-based-organization research-organization think-tank university academic-institution knowledge-institution capacity-building-institution training-institution extension-service advisory-service consulting-firm engineering-firm technology-provider equipment-supplier service-provider contractor operator maintainer utility municipality local-government provincial-government national-government federal-government ministry department agency authority commission board council committee parliament congress legislature executive judiciary media journalist reporter editor publisher broadcaster platform social-media influencer celebrity activist advocate campaigner mobilizer organizer networker connector bridge-builder boundary-spanner knowledge-broker science-communicator extension-agent development-practitioner policy-maker decision-maker planner manager administrator leader champion entrepreneur innovator intrapreneur social-entrepreneur systems-entrepreneur institutional-entrepreneur policy-entrepreneur political-entrepreneur business-model business-model-innovation social-business inclusive-business base-of-pyramid fortune-at-bottom-of-pyramid shared-value creating-shared-value stakeholder-capitalism purpose-driven-business B-Corp benefit-corporation social-enterprise cooperative mutual association collective commons commoning peer-production open-source creative-commons knowledge-commons digital-commons data-commons genetic-commons seed-commons water-commons fisheries-commons forest-commons pasture-commons land-commons urban-commons atmospheric-commons global-commons common-heritage-of-mankind common-concern-of-humankind restorative-justice multispecies-justice more-than-human anthropocene capitalocene plantationocene chthulucene symbiocene good-anthropocene great-acceleration great-transformation great-decoupling great-convergence great-divergence great-fragmentation great-integration great-transition great-work long-now deep-time futures-thinking future-studies strategic-foresight scenario-planning trend-analysis weak-signal wild-card black-swan grey-rhino blue-sky green-swan systemic-change structural-change cultural-change behavioral-change individual-change collective-change social-change organizational-change technological-change economic-change political-change ecological-change planetary-change earth-system-change biodiversity-change ocean-change cryosphere-change atmospheric-change biogeochemical-change nitrogen-cycle-change phosphorus-cycle-change carbon-cycle-change water-cycle-change hydrological-change soil-change vegetation-change ecosystem-change landscape-change seascape-change biosphere-change noosphere-change technosphere-change anthroposphere-change sociosphere-change econosphere-change politosphere-change culturosphere-change semiosphere-change infosphere-change digisphere-change cybersphere-change urban-sphere-change agricultural-sphere-change industrial-sphere-change extractive-sphere-change manufacturing-sphere-change service-sphere-change financial-sphere-change knowledge-sphere-change innovation-sphere-change creativity-sphere-change culture-sphere-change identity-sphere-change meaning-sphere-change value-sphere-change purpose-sphere-change vision-sphere-change mission-sphere-change strategy-sphere-change tactic-sphere-change operation-sphere-change action-sphere-change practice-sphere-change habit-sphere-change routine-sphere-change ritual-sphere-change tradition-sphere-change custom-sphere-change convention-sphere-change norm-sphere-change rule-sphere-change law-sphere-change regulation-sphere-change standard-sphere-change protocol-sphere-change procedure-sphere-change process-sphere-change system-sphere-change structure-sphere-change function-sphere-change pattern-sphere-change dynamic-sphere-change state-sphere-change phase-sphere-change regime-sphere-change equilibrium-sphere-change stability-sphere-change instability-sphere-change chaos-sphere-change order-sphere-change complexity-sphere-change simplicity-sphere-change diversity-sphere-change uniformity-sphere-change heterogeneity-sphere-change homogeneity-sphere-change variation-sphere-change invariance-sphere-change change-sphere-change continuity-sphere-change discontinuity-sphere-change emergence-sphere-change submergence-sphere-change evolution-sphere-change devolution-sphere-change revolution-sphere-change involution-sphere-change convolution-sphere-change resolution-sphere-change dissolution-sphere-change absolution-sphere-change solution-sphere-change