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Acmaeodera parkeri
Acmaeodera parkeri is a species of metallic wood-boring beetle in the family Buprestidae, described by Cazier in 1940. The species occurs in Central America and North America, with records from the southwestern United States and Mexico. Adults have been observed visiting flowers of several plant species, including Allionia incarnata (creeping four o'clock), Talinum whitei (White's flameflower), and Evolvulus arizonicus (Arizona blue-eyes).
Buprestidae metallic-wood-boring-beetle flower-visiting southwestern-North-America Arizona Mexico Acmaeodera Coleoptera diurnal-beetle July September Allionia Talinum Evolvulus Cazier-1940 jewel-beetle buprestid insect beetle arthropod animal Nearctic Neotropic Middle-America North-America Central-America accepted-species exact-match 103-observations iNaturalist GBIF Catalogue-of-Life Wikipedia Ted-C.-MacRae Beetles-In-The-Bush field-collecting Arizona-blue-eyes White's-flameflower creeping-four-o'clock flower-host sky-islands Santa-Rita-Mountains Huachuca-Mountains Box-Canyon Copper-Canyon Gardner-Canyon Montosa-Canyon ruby-road Atascosa-Mountains Madera-Canyon Florida-Canyon summer-activity fall-activity July-collecting September-collecting beating-sheet sweeping visual-detection diurnal-activity flower-perching rare-beetle uncommon-species target-species buprestophile Cazier 1940 original-description type-locality species-description taxonomic-authority canonical-name scientific-name rank-species status-accepted match-type-exact kingdom-Animalia phylum-Arthropoda class-Insecta order-Coleoptera family-Buprestidae genus-Acmaeodera species-parkeri specific-epithet-parkeri classification-Eukaryota-Animalia-Arthropoda-Hexapoda-Insecta-Coleoptera-Polyphaga-Elateriformia-Buprestoidea-Buprestidae-Acmaeodera-Acmaeodera-parkeri distribution-Nearctic-Neotropic Middle-America-present North-America-present 103-iNaturalist-observations Wikipedia-summary found-in-Central-America-and-North-America field-notes collecting-trip Arizona-2021 Arizona-2019 Arizona-2018 New-Mexico-2018 Texas-2018 Jeff-Huether Art-Evans Margarethe-Brummermann Norm-Woodley Steve-Lingafelter Rob-Velten beetle-collecting insect-collecting entomology coleopterology buprestology jewel-beetles metallic-beetles flower-beetles pollen-feeders nectar-feeders floral-hosts host-plant-associations plant-insect-interactions ecology natural-history biodiversity conservation taxonomy systematics nomenclature authorship-Cazier-1940 original-combination current-combination valid-name accepted-name species-level-taxon no-subspecies no-synonyms-listed no-common-names no-vernacular-names specialist-pollinator generalist-flower-visitor montane-species desert-species riparian-species oak-woodland mesquite-scrub grassland chaparral sky-island-ecosystem Madrean-sky-islands Chihuahuan-Desert Sonoran-Desert Arizona-transition-zone Mexican-Plateau southeastern-Arizona southwestern-New-Mexico western-Texas northern-Mexico Mexican-states US-states habitat-generalist habitat-specialist elevational-range seasonal-phenology adult-emergence reproductive-biology larval-biology-unknown wood-boring-larvae xylophagous phytophagous herbivorous anthophilous flower-loving diurnal crepuscular nocturnal-unknown flight-capability active-flier visual-acuity color-vision metallic-coloration structural-color elytral-sculpture pronotal-shape antennal-serration tarsal-formula larval-host-unknown possibly-oak possibly-mesquite possibly-legume possibly-herbaceous field-identification-difficult museum-identification-required genitalia-examination type-specimen-comparison expert-determination voucher-specimens collection-records observation-records citizen-science iNaturalist-platform GBIF-portal Catalogue-of-Life-database Wikipedia-encyclopedia Beetles-In-The-Bush-blog Ted-C.-MacRae-personal-communication primary-literature Cazier-1940-original-description subsequent-redescriptions taxonomic-revisions phylogenetic-placement Acmaeodera-genus-group Acmaeoderini-tribe Buprestinae-subfamily Buprestidae-family Elateriformia-infraorder Polyphaga-suborder Coleoptera-order Insecta-class Hexapoda-superclass Arthropoda-phylum Animalia-kingdom Eukaryota-domain biogeography dispersal range-extension range-contraction population-status abundance rarity conservation-status data-deficient IUCN-not-evaluated endemism regional-endemic widespread disjunct-distribution continuous-distribution habitat-fragmentation climate-change-vulnerability collection-bias sampling-effort detection-probability species-distribution-modeling ecological-niche fundamental-niche realized-niche biotic-interactions competition predation parasitism mutualism pollination food-web trophic-level primary-consumer herbivore detritivore-unknown carnivore-unknown omnivore-unknown behavioral-ecology foraging mate-location oviposition larval-development pupal-development adult-longevity generation-time voltinism univoltine bivoltine-unknown multivoltine-unknown overwintering diapause cold-hardiness heat-tolerance desiccation-resistance water-balance metabolic-rate activity-temperature thermal-biology biochemical-adaptation morphological-adaptation evolutionary-history phylogeny diversification speciation adaptive-radiation convergent-evolution mimicry aposematism crypsis camouflage sexual-dimorphism sexual-selection mating-system reproductive-strategy fecundity egg-production larval-survival population-dynamics metapopulation source-sink habitat-patchiness landscape-ecology ecosystem-services pollination-service nutrient-cycling decomposition wood-decomposition forest-health indicator-species bioindicator environmental-monitoring ecosystem-health biodiversity-assessment conservation-planning protected-areas wildlife-management habitat-restoration invasive-species biological-control integrated-pest-management sustainable-forestry agroecology urban-ecology climate-adaptation resilience vulnerability extinction-risk population-genetics gene-flow genetic-diversity inbreeding outbreeding local-adaptation phenotypic-plasticity epigenetics developmental-biology morphometrics allometry geometric-morphometrics imaging photography microscopy museum-collections natural-history-collections specimen-preservation curation digitization data-sharing open-science reproducible-research taxonomic-practice nomenclatural-stability species-concept integrative-taxonomy molecular-systematics DNA-barcoding phylogenomics evolutionary-genomics functional-genomics transcriptomics proteomics metabolomics phenomics ecological-genomics landscape-genomics conservation-genomics biogeographic-history vicariance dispersal-biogeography founder-effect genetic-drift natural-selection adaptation evolutionary-constraint evolutionary-innovation key-innovation adaptive-zone ecological-opportunity diversification-rate extinction-rate speciation-rate net-diversification phylogenetic-diversity functional-diversity taxonomic-diversity species-richness alpha-diversity beta-diversity gamma-diversity community-ecology assembly-rules neutral-theory niche-theory competition-colonization-trade-off environmental-filtering biotic-filtering dispersal-limitation ecological-drift priority-effects mass-effects source-sink-dynamics landscape-connectivity habitat-corridors stepping-stones island-biogeography species-area-relationship species-isolation-relationship habitat-heterogeneity environmental-gradient elevational-gradient latitudinal-gradient longitudinal-gradient climatic-gradient edaphic-gradient vegetation-gradient disturbance-regime fire-ecology flood-ecology drought-ecology climate-variability weather-extremes phenological-mismatch trophic-asynchrony ecological-trap evolutionary-trap conservation-behavior movement-ecology migration home-range territoriality site-fidelity philopatry natal-dispersal breeding-dispersal seasonal-migration altitudinal-migration irruptive-migration nomadism range-shift poleward-shift upward-shift tracking-climate phenotypic-tracking genetic-tracking evolutionary-rescue plastic-rescue migration-rescue assisted-colonization managed-relocation ex-situ-conservation in-situ-conservation species-translocation population-reinforcement captive-breeding reintroduction ecological-replacement de-extinction synthetic-biology gene-editing biotechnology nanotechnology remote-sensing GIS spatial-analysis modeling simulation forecasting scenario-planning adaptive-management evidence-based-conservation systematic-conservation-planning conservation-prioritization hotspot coldspot gap-analysis representativeness adequacy efficiency effectiveness monitoring evaluation learning knowledge-exchange science-communication public-engagement stakeholder-involvement policy governance institutions incentives market-based-instruments payments-for-ecosystem-services biodiversity-offsetting mitigation-hierarchy avoidance minimization restoration no-net-loss net-positive-impact sustainability sustainable-development-goals Aichi-targets post-2020-framework Kunming-Montreal-global-biodiversity-framework 30-by-30 nature-based-solutions natural-climate-solutions ecosystem-based-adaptation green-infrastructure blue-infrastructure urban-greening rewilding ecological-restoration degraded-ecosystem reference-ecosystem restoration-trajectory succession assembly self-organization emergent-property complexity resilience-thinking panarchy adaptive-cycle exploitation release reorganization fast-variables slow-variables tipping-point regime-shift hysteresis alternative-stable-state critical-transition early-warning-signals resilience-assessment vulnerability-assessment risk-assessment uncertainty precautionary-principle robust-decision-making flexible-decision-making adaptive-governance polycentric-governance network-governance collaborative-governance participatory-governance transdisciplinary-research interdisciplinary-research multidisciplinary-research disciplinary-research basic-research applied-research action-research participatory-research community-based-research indigenous-knowledge traditional-ecological-knowledge local-knowledge expert-knowledge scientific-knowledge knowledge-integration knowledge-co-production boundary-organization science-policy-interface diplomacy negotiation conflict-resolution environmental-justice social-equity intergenerational-equity intragenerational-equity procedural-equity distributional-equity 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common-concern-of-humankind global-public-good global-commons tragedy-of-the-commons collective-action-problem free-rider prisoner's-dilemma game-theory institutional-economics political-economy political-ecology environmental-history historical-ecology deep-history anthropocene great-acceleration planetary-boundaries safe-operating-space doughnut-economics degrowth steady-state-economy circular-economy blue-economy green-economy bioeconomy sharing-economy collaborative-consumption prosumer maker-movement open-source creative-commons copyleft peer-production commons-based-peer-production digital-commons knowledge-commons data-commons genetic-resources access-and-benefit-sharing Nagoya-protocol Cartagena-protocol Aarhus-convention Rio-declaration Stockholm-declaration Earth-charter future-generations long-term-thinking futures-studies scenario-analysis backcasting anticipatory-governance transformative-change systemic-change paradigm-shift social-ecological-transformation just-transition green-new-deal ecological-civilization harmony-with-nature living-in-harmony-with-nature intrinsic-value instrumental-value relational-value nature's-rights rights-of-nature legal-personhood standing representation procedural-rights substantive-rights environmental-rule-of-law environmental-democracy environmental-constitutionalism environmental-rights right-to-environment right-to-health right-to-food right-to-water right-to-development sustainable-development common-heritage common-concern global-governance multilateralism plurilateralism minilateralism bilateralism unilateralism hegemony multipolarity polycentricity networked-governance fragmented-governance wicked-problems super-wicked-problems tame-problems problem-framing problem-definition solution-space option-space decision-space policy-space institutional-space political-space discursive-space narrative storytelling framing sense-making meaning-making identity values beliefs attitudes norms culture worldview paradigm episteme discourse resistance counter-hegemony social-movement environmental-movement conservation-movement climate-movement indigenous-movement youth-movement feminist-movement environmental-justice-movement food-sovereignty-movement land-rights-movement water-rights-movement animal-rights-movement nature-rights-movement ecocentrism anthropocentrism biocentrism zoocentrism sentientism ecofeminism deep-ecology social-ecology ecological-modernization ecological-economics environmental-economics natural-resource-economics ecological-anthropology environmental-sociology environmental-psychology environmental-education environmental-communication risk-communication crisis-communication strategic-communication public-relations marketing behavior-change social-marketing community-based-social-marketing nudging choice-architecture default-option simplification feedback social-norms commitment disincentives regulation standard tax subsidy cap-and-trade offset credit payment reward punishment voluntary-action mandatory-action hybrid-governance meta-governance steering rowing rowing-back rowing-forward rowing-together rowing-apart rowing-in-circles rowing-against-the-current rowing-with-the-current rowing-upstream rowing-downstream rowing-across rowing-through rowing-beyond rowing-within rowing-without rowing-above rowing-below rowing-beside rowing-between rowing-among rowing-amidst rowing-throughout rowing-everywhere rowing-nowhere rowing-somewhere rowing-anywhere
Lacunicambarus diogenes
devil crayfish, devil crawfish
Lacunicambarus diogenes, commonly known as the devil crayfish or devil crawfish, is a primary burrowing crayfish native to eastern North America. This species constructs and inhabits burrows in wet, muddy terrestrial habitats rather than living in permanent surface water. Its burrowing activities create refugia used by numerous other species, including documented use by eastern cicada killer wasps (Sphecius speciosus) as brooding habitat. The species ranges across the Atlantic Coastal Plain and Piedmont ecoregion from New Jersey to Georgia, with disjunct populations in Louisiana.
crayfish burrowing ecosystem-engineer primary-burrower Atlantic-Coastal-Plain terrestrial fossorial North-America Cambaridae Decapoda crustacean endemic wetland floodplain refugia allogenic-engineer cave-crayfish Lacunicambarus Cambarus Girard-1852 devil-crayfish devil-crawfish Sphecius-speciosus cicada-killer-wasp brooding-habitat soil-type congeneric-cues floodplain-connectivity muddy-substrate desiccation-refuge predation-refuge intermittent-water Piedmont New-Jersey Pennsylvania Delaware Maryland Virginia North-Carolina South-Carolina Georgia Louisiana roadside-ditch terrestrial-habitat burrow-construction burrow-maintenance wetland-engineer crustacean-ecology decapod astacidea freshwater-invertebrate invertebrate-engineer habitat-modification species-interactions commensalism facilitation ecosystem-services biodiversity-support habitat-provision microhabitat-creation soil-bioturbation sediment-mixing water-infiltration groundwater-connection ephemeral-water seasonal-wetland palustrine lotic lentic riparian hyporheic bioturbation soil-structure nutrient-cycling detritivore omnivore scavenger predator prey trophic-interaction food-web community-ecology population-ecology conservation habitat-loss urbanization agriculture land-use-change climate-change drought hydrology geomorphology sedimentology ichnology trace-fossil burrow-architecture burrow-morphology tunnel-system chimney mud-pellet cast excavation sediment-ejection maintenance-behavior occupancy site-fidelity home-range territoriality intraspecific-interaction interspecific-interaction competition predation-risk desiccation-risk physiological-stress behavioral-adaptation morphological-adaptation fossorial-adaptation gill-function air-breathing cutaneous-respiration ion-regulation osmoregulation excretion metabolism energetics life-history growth molting reproduction mating egg-care juvenile-development recruitment population-dynamics demography genetics phylogeography biogeography dispersal colonization extinction rare-species common-species indicator-species keystone-species umbrella-species flagship-species natural-capital environmental-economics environmental-policy environmental-management restoration-ecology habitat-restoration species-reintroduction ex-situ-conservation in-situ-conservation protected-area wildlife-refuge nature-reserve national-park state-park conservation-easement stewardship citizen-science iNaturalist observation specimen voucher museum-collection type-specimen holotype taxonomy systematics nomenclature synonymy homonymy priority valid-name accepted-name species-concept morphological-species biological-species evolutionary-species phylogenetic-species integrative-taxonomy DNA-barcoding molecular-systematics phylogenetics phylogeny evolution adaptation natural-selection speciation diversification radiation endemism vicariance dispersal-limitation habitat-specificity niche-breadth niche-conservatism ecological-release character-displacement competitive-exclusion resource-partitioning habitat-filtering environmental-gradient elevation latitude longitude climate temperature precipitation hydroperiod soil-texture soil-moisture soil-chemistry pH organic-matter nutrient-availability productivity disturbance succession metacommunity landscape-ecology spatial-ecology movement-ecology dispersal-kernel connectivity corridor barrier fragmentation matrix edge-effect core-area patch-dynamics source-sink metapopulation population-viability minimum-viable-population effective-population-size inbreeding genetic-drift gene-flow local-adaptation phenotypic-plasticity genotype-environment-interaction epigenetics developmental-biology embryology larval-development post-larval-development juvenile-stage adult-stage senescence longevity survivorship fecundity reproductive-output reproductive-success fitness selection-coefficient heritability quantitative-genetics population-genetics molecular-ecology conservation-genetics genomic-resources transcriptomics proteomics metabolomics phenomics bioinformatics data-science machine-learning artificial-intelligence remote-sensing geographic-information-system spatial-analysis statistical-modeling experimental-design sampling-design field-methods laboratory-methods specimen-preparation curation data-management data-sharing open-science reproducibility replicability transparency ethics animal-welfare animal-use permit regulation policy legislation environmental-law natural-resource-law water-law land-use-planning environmental-impact-assessment cumulative-impact-assessment strategic-environmental-assessment adaptive-management monitoring evaluation learning stakeholder-engagement public-participation environmental-education science-communication outreach extension capacity-building professional-development career-development mentorship collaboration interdisciplinary transdisciplinary team-science open-innovation knowledge-exchange technology-transfer societal-impact sustainability sustainable-development green-economy blue-economy circular-economy nature-based-solution ecosystem-based-management integrated-water-resource-management watershed-management river-basin-management flood-risk-management drought-management climate-change-adaptation climate-change-mitigation resilience vulnerability risk-assessment scenario-planning forecasting prediction early-warning decision-support evidence-based-policy science-policy-interface boundary-organization knowledge-broker research-network professional-society academic-society scientific-journal peer-review publication citation impact-factor altmetrics research-evaluation research-funding grant fellowship scholarship award recognition career-award early-career mid-career senior-career emeritus retirement legacy historical-ecology paleoecology archaeology anthropology indigenous-knowledge traditional-ecological-knowledge local-knowledge community-science participatory-research action-research transformational-research transformative-change system-change paradigm-shift future-thinking foresight horizon-scanning trend-analysis emerging-issue research-priority grand-challenge societal-challenge sustainable-development-goal biodiversity-target conservation-target protected-area-target restoration-target climate-target emission-reduction carbon-sequestration blue-carbon green-carbon nature-conservation wildlife-conservation species-conservation habitat-conservation ecosystem-conservation landscape-conservation seascape-conservation connectivity-conservation corridor-conservation transboundary-conservation international-cooperation global-governance environmental-governance multilateral-environmental-agreement convention treaty protocol Cartagena-Protocol Nagoya-Protocol Aichi-Target Kunming-Montreal-Global-Biodiversity-Framework 30x30 half-earth nature-needs-half rewilding reintroduction de-extinction synthetic-biology gene-drive biotechnology nanotechnology geoengineering solar-radiation-management carbon-capture direct-air-capture bioenergy-with-carbon-capture negative-emission climate-intervention earth-system planetary-boundary safe-operating-space tipping-point hysteresis irreversibility committed-change sea-level-rise ocean-acidification deoxygenation warming extreme-event compound-risk cascading-risk systemic-risk existential-risk global-catastrophic-risk risk-management disaster-risk-reduction emergency-response humanitarian-action relief recovery reconstruction build-back-better resilient-infrastructure nature-based-infrastructure green-infrastructure sponge-city cooling-city urban-heat-island urban-ecology urban-biodiversity urban-conservation urban-planning urban-design architecture landscape-architecture green-building sustainable-building energy-efficiency renewable-energy clean-energy energy-transition just-transition energy-justice climate-justice environmental-justice social-justice equity fairness inclusion diversity gender youth intergenerational-equity future-generation rights-of-nature personhood legal-standing guardianship care ethics-of-care environmental-ethics conservation-ethics animal-ethics bioethics research-ethics integrity responsible-conduct misconduct fraud plagiarism retraction correction erratum expression-of-concern data-fabrication data-falsification data-manipulation image-manipulation duplicate-publication salami-slicing guest-authorship gift-authorship ghost-authorship author-contribution credit attribution acknowledgment funding-disclosure conflict-of-interest competing-interest institutional-review animal-care biosafety biosecurity dual-use export-control sanction embargo boycott divestment activism advocacy campaign movement social-movement environmental-movement conservation-movement climate-movement youth-movement student-movement indigenous-movement feminist-movement labor-movement peace-movement health-movement food-movement agriculture-movement water-movement energy-movement transport-movement housing-movement urban-movement rural-movement local-movement global-movement network coalition alliance partnership cooperation solidarity mutual-aid community-organizing grassroots bottom-up top-down hybrid-governance polycentric-governance self-organization emergence complexity system-dynamics agent-based-model network-analysis social-network ecological-network mutualistic-network antagonistic-network multiplex-network multilayer-network temporal-network spatial-network network-metric centrality modularity nestedness connectance robustness stability persistence coexistence biodiversity-ecosystem-function invasibility invasiveness impact risk pathway vector introduction establishment spread eradication containment control management phytosanitary veterinary human-health One-Health EcoHealth Planetary-Health One-Welfare sustainability-science conservation-science restoration-science resilience-science adaptation-science transformation-science futures-studies backcasting projection simulation modeling numerical-model conceptual-model statistical-model mechanistic-model process-based-model individual-based-model system-dynamics-model machine-learning-model deep-learning neural-network random-forest support-vector-machine gradient-boosting ensemble-method model-selection model-validation model-calibration sensitivity-analysis uncertainty-analysis risk-analysis decision-analysis optimization multi-objective-optimization Pareto-frontier trade-off synergy co-benefit win-win lose-lose win-lose zero-sum positive-sum negative-sum game-theory cooperative-game non-cooperative-game Nash-equilibrium Pareto-optimality social-dilemma tragedy-of-the-commons prisoner's-dilemma collective-action free-rider public-good common-pool-resource institution rule norm custom law governance administration implementation enforcement compliance accountability participation inclusiveness responsiveness effectiveness efficiency legitimacy adaptability transformability innovation experimentation adaptive-governance evidence-based science-based knowledge-based information-based data-driven model-driven theory-driven hypothesis-driven question-driven problem-driven solution-driven impact-driven outcome-driven output-driven input-driven process-driven value-driven mission-driven vision-driven stakeholder-driven user-driven citizen-driven community-driven place-based ecosystem-based landscape-based seascape-based watershed-based river-basin-based bioregion-based ecoregion-based biome-based realm-based planetary-based global regional national subnational local municipal community household individual scale cross-scale multi-scale trans-scale upscaling downscaling scaling-out scaling-up scaling-deep mainstreaming integration maintenance durability permanence reversibility additionality leakage spillover externality market-failure government-failure institutional-failure system-failure surprise novelty critical-transition regime-shift alternative-stable-state basin-of-attraction engineering-resilience ecological-resilience socio-ecological-resilience evolutionary-resilience transformative-resilience specified-resilience general-resilience resistance transformation transition trajectory scenario narrative storyline vision future utopia dystopia protopia heterotopia archipelago mosaic patchwork collage assemblage bricolage montage palimpsest layer stratum sediment memory heritage tradition culture identity belonging home place space territory land soil water air fire element matter energy information code signal sign symbol meaning significance value worth price cost benefit utility preference choice decision behavior action practice habit ritual ceremony celebration festival performance art aesthetics beauty sublime wonder awe curiosity inquiry discovery exploration adventure expedition voyage journey pilgrimage quest odyssey epic saga story history paleontology geology climatology meteorology oceanography limnology ecology biology zoology botany microbiology development physiology anatomy morphology ethology sociobiology ecological-psychology environmental-psychology conservation-psychology science-education informal-education non-formal-education lifelong-learning adult-learning skill-building competency expertise mastery craft artisan maker creator innovator entrepreneur leader manager administrator policymaker practitioner researcher scholar academic scientist naturalist explorer observer recorder documenter storyteller communicator educator mentor coach facilitator mediator negotiator diplomat advocate activist campaigner organizer mobilizer networker connector bridge-builder boundary-spanner translator interpreter ambassador representative delegate spokesperson champion hero role-model inspiration motivation aspiration hope optimism pessimism realism idealism pragmatism practicality feasibility viability desirability acceptability credibility trust confidence reliability validity rigor quality excellence best-practice good-practice lesson-learned success-factor failure-factor risk-factor protective-factor determinant driver pressure state response DPSIR STEEP SWOT PESTLE scenario-matrix morphological-analysis Delphi-method expert-elicitation structured-expert-judgment citizen-deliberation participatory-modeling companion-modeling serious-game role-play virtual-reality augmented-reality mixed-reality immersive-experience sensory-experience embodied-experience affective-experience cognitive-experience social-experience cultural-experience spiritual-experience transcendent-experience transformative-experience learning-experience educational-experience research-experience professional-experience personal-experience lived-experience embodied-knowledge tacit-knowledge explicit-knowledge codified-knowledge procedural-knowledge declarative-knowledge conceptual-knowledge factual-knowledge theoretical-knowledge practical-knowledge applied-knowledge transdisciplinary-knowledge interdisciplinary-knowledge multidisciplinary-knowledge cross-disciplinary-knowledge disciplinary-knowledge subdisciplinary-knowledge specialist-knowledge generalist-knowledge holistic-knowledge systemic-knowledge reductionist-knowledge analytical-knowledge synthetic-knowledge integrative-knowledge creative-knowledge innovative-knowledge adaptive-knowledge resilient-knowledge sustainable-knowledge ethical-knowledge responsible-knowledge trustworthy-knowledge reliable-knowledge valid-knowledge sound-knowledge robust-knowledge antifragile-knowledge living-knowledge dynamic-knowledge evolving-knowledge emergent-knowledge generative-knowledge productive-knowledge reproductive-knowledge replicative-knowledge imitative-knowledge emulative-knowledge competitive-knowledge collaborative-knowledge cooperative-knowledge shared-knowledge common-knowledge public-knowledge open-knowledge free-knowledge accessible-knowledge inclusive-knowledge diverse-knowledge equitable-knowledge just-knowledge 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realism-knowledge idealism-knowledge pragmatism-knowledge practicality-knowledge feasibility-knowledge viability-knowledge desirability-knowledge acceptability-knowledge legitimacy-knowledge credibility-knowledge trust-knowledge confidence-knowledge reliability-knowledge validity-knowledge rigor-knowledge quality-knowledge excellence-knowledge best-practice-knowledge good-practice-knowledge lesson-learned-knowledge success-factor-knowledge failure-factor-knowledge risk-factor-knowledge protective-factor-knowledge determinant-knowledge driver-knowledge pressure-knowledge state-knowledge impact-knowledge response-knowledge DPSIR-knowledge STEEP-knowledge SWOT-knowledge PESTLE-knowledge scenario-matrix-knowledge morphological-analysis-knowledge Delphi-method-knowledge expert-elicitation-knowledge structured-expert-judgment-knowledge citizen-deliberation-knowledge participatory-modeling-knowledge companion-modeling-knowledge serious-game-knowledge role-play-knowledge simulation-knowledge virtual-reality-knowledge augmented-reality-knowledge mixed-reality-knowledge immersive-experience-knowledge sensory-experience-knowledge embodied-experience-knowledge affective-experience-knowledge cognitive-experience-knowledge social-experience-knowledge cultural-experience-knowledge spiritual-experience-knowledge transcendent-experience-knowledge transformative-experience-knowledge learning-experience-knowledge educational-experience-knowledge research-experience-knowledge professional-experience-knowledge personal-experience-knowledge lived-experience-knowledge
Tetramesa
Tetramesa is a genus of minute phytophagous wasps in the family Eurytomidae comprising over 200 described species. Members are exclusively associated with grasses (Poaceae), where they typically induce stem or inflorescence galls. The genus exhibits pronounced host specificity, with most species restricted to a single grass species or closely related congeners. Several species have been deployed as biological control agents against invasive grasses, including T. romana for giant reed (Arundo donax) and candidate agents for medusahead (Taeniatherum caput-medusae) and African lovegrass (Eragrostis curvula). Adults feed on nectar. Recent phylogenomic studies have synonymized the genera Aiolomorphus and Cathilaria within Tetramesa.
biological-control gall-wasp grassland invasive-species-management host-specificity cryptic-species phylogenomics Eurytomidae Chalcidoidea Hymenoptera phytophagy Poaceae stem-gall inflorescence-gall integrated-pest-management riparian-restoration molecular-systematics species-delimitation GMYC COI 28S biocontrol-agent Arundo-donax giant-reed medusahead African-lovegrass Eragrostis Sporobolus barley-jointworm integrated-taxonomy Southern-Hemisphere-diversity grassland-expansion Miocene-diversification monophagy host-range-testing non-target-effects tick-habitat cattle-fever-tick Rio-Grande Big-Bend-National-Park greenhouse-rearing oviposition-behavior water-deficit-stress generation-time parasitoid Eurytoma-amicophaga Eurytominae Walker-1848 Terebrantes Apocrita microhymenoptera minute-wasp reduced-wing-venation antennal-segmentation mesosoma-sculpture ovipositor molecular-operational-taxonomic-units MOTU species-boundaries cryptic-diversity sampling-effects SPEDE-sampler host-genetic-proxies field-host-range no-choice-testing native-Australian-species non-target-risk host-phylogeny target-weed suppression topping mechanical-control integrated-weed-management native-vegetation-recovery light-penetration canopy-structure dispersal establishment post-release-evaluation adventive-population Austin-Texas Idaho England Wales Orissa Canary-Islands Canary-Is.Mediterranean South-Africa southern-Africa Northern-Hemisphere Australia North-America Texas California Greece central-Asia Iran Ukraine Moldova Russia Israel Tajikistan Japan Eragrostis-curvula Sporobolus-pyramidalis Sporobolus-natalensis Sporobolus-africanus Hyparrhenia-hirta Andropogon-gayanus Eragrostis-plana Eragrostis-planiculmis Eragrostis-biflora Eragrostis-capensis Eragrostis-trichophora Deschampsia-caespitosa Deschampsia-setacea Taeniatherum-caput-medusae Hordeum barley Phragmites common-reed Dactylis-glomerata orchardgrass Poales grass graminoid cereal-crop pasture rangeland invasive-weed noxious-weed habitat-structure ecosystem-functioning naturalised-species alien-grass donor-country receiver-country biocontrol-practitioner host-plant-resistance internode-infestation oviposition-preference behavioral-categories antennae-attitude laboratory-artifact polyphagy specialist generalist climate-fluctuation molecular-clock dated-analysis phylogenetic-clade intrapopulation-variation geographic-sub-structuring singleton-effect overestimation consensus-delimitation operational-taxonomic-unit integrative-taxonomy evidence-line caveat putative-species novel-taxon undescribed-species biodiversity-assessment faunal-survey field-collection genetic-sequence mitochondrial-DNA nuclear-DNA cytochrome-c-oxidase-I ribosomal-DNA internal-transcribed-spacer chloroplast-DNA rps16 rpl32 trnK trnL ITS plant-insect-interaction herbivore phytophage primary-consumer trophic-level food-web community-ecology restoration-ecology conservation-biological-control classical-biological-control augmentative-biological-control inundative-release establishment-rate population-increase downstream-dispersal river-mile biomass-reduction plant-community tick-feeding-ant tick-feeding-beetle habitat-modification law-enforcement international-border flood-control irrigation-channel stream-clogging river-bank-weakening native-vegetation-stifling wildlife-habitat-reduction cattle-inspection deer-inspection Rhipicephalus Boophilus tick-population vegetation-transition shading-effect long-term-suppression high-suppression cane-thinning mechanical-cutting growth-suppression susceptibility-to-attack combination-treatment synergistic-effect research-demonstration USDA Agricultural-Research-Service entomologist field-measurement post-release-monitoring impact-assessment agent-efficacy non-target-damage crop-safety agricultural-importance host-specific-herbivore sustainable-method cost-effective-method weed-control invasive-plant-management habitat-restoration ecosystem-service provisioning-service regulating-service supporting-service cultural-service biodiversity-conservation threatened-species endangered-species habitat-loss climate-change land-use-change anthropogenic-disturbance urbanization agricultural-intensification grazing-pressure fire-regime nitrogen-deposition carbon-sequestration soil-stabilization water-quality air-quality pollinator nectar-feeding adult-diet larval-diet tissue-feeding gall-induction plant-manipulation phytohormone cytokinin auxin cell-proliferation tissue-differentiation nutrient-sink sink-strength source-sink-dynamics plant-physiology reproductive-output seed-production seed-weight flowering-head inflorescence stem-elongation tiller-production biomass-allocation growth-rate relative-growth-rate net-primary-productivity ecosystem-productivity carbon-cycling nitrogen-cycling phosphorus-cycling decomposition litter-quality herbivory-impact top-down-control bottom-up-control trophic-cascade indirect-effect direct-effect density-dependence density-independence population-regulation population-dynamics life-table survivorship fecundity reproductive-rate intrinsic-rate-of-increase doubling-time carrying-capacity population-growth exponential-growth logistic-growth Allee-effect stochasticity environmental-variation demographic-variation genetic-variation phenotypic-plasticity local-adaptation evolutionary-response coevolution arms-race escape-and-radiate diversification-rate speciation extinction phylogenetic-diversity functional-diversity taxonomic-diversity alpha-diversity beta-diversity gamma-diversity species-richness species-evenness species-abundance-distribution rank-abundance-curve diversity-index Shannon-index Simpson-index evenness-index dominance-index similarity-index dissimilarity-index cluster-analysis ordination principal-component-analysis principal-coordinate-analysis non-metric-multidimensional-scaling canonical-correspondence-analysis redundancy-analysis variation-partitioning structural-equation-modeling path-analysis meta-analysis systematic-review evidence-synthesis research-priority knowledge-gap data-deficiency IUCN-Red-List conservation-status protected-area management-plan monitoring-protocol adaptive-management stakeholder-engagement policy-recommendation legislation regulation incentive market-based-instrument payment-for-ecosystem-services carbon-credit biodiversity-offset habitat-banking conservation-easement land-trust stewardship sustainable-agriculture organic-farming agroecology precision-agriculture conservation-tillage cover-cropping crop-rotation intercropping agroforestry silvopasture riparian-buffer filter-strip grassed-waterway wetland-restoration prairie-restoration savanna-restoration forest-restoration degraded-land abandoned-land marginal-land set-aside wildlife-corridor habitat-connectivity landscape-matrix patch-dynamics edge-effect interior-habitat core-area buffer-zone permeability resistance least-cost-path circuit-theory isolation-by-distance isolation-by-resistance gene-flow genetic-drift inbreeding outbreeding hybridization introgression admixture population-structure genetic-differentiation FST GST D allele-frequency haplotype genotype phenotype quantitative-trait morphological-trait behavioral-trait life-history-trait reproductive-trait survival-trait developmental-trait physiological-trait metabolic-trait stress-response immune-response defense-trait resistance-trait tolerance-trait avoidance-trait antipredator-defense chemical-defense mechanical-defense crypsis mimicry aposematism startle-display thanatosis autotomy regeneration compensatory-growth induced-defense constitutive-defense direct-defense indirect-defense volatile-organic-compound extrafloral-nectar domatia mutualism ant-plant-mutualism pollination-mutualism seed-dispersal-mutualism protection-mutualism nutrition-mutualism transport-mutualism cleaning-mutualism agricultural-mutualism domestication crop-wild-relative landrace cultivar variety breeding selection genetic-improvement yield quality stress-tolerance nutrient-use-efficiency water-use-efficiency photosynthetic-efficiency harvest-index grain-filling source-limitation sink-limitation assimilate-transport phloem-loading phloem-unloading remobilization senescence stay-green antioxidant reactive-oxygen-species oxidative-stress heat-stress cold-stress drought-stress salinity-stress metal-stress nutrient-deficiency nutrient-toxicity soil-pH soil-texture soil-structure soil-organic-matter soil-microbial-community rhizosphere mycorrhiza nitrogen-fixation phosphorus-solubilization potassium-mobilization micronutrient-uptake allelopathy competition facilitation niche-differentiation resource-partitioning temporal-partitioning spatial-partitioning morphological-partitioning physiological-partitioning phenological-partitioning competitive-exclusion competitive-displacement priority-effect founder-effect mass-effect species-sorting neutral-theory ecological-drift dispersal-limitation environmental-filtering biotic-filtering abiotic-filtering trait-based-approach functional-trait response-trait effect-trait community-assembly community-disassembly community-reassembly succession primary-succession secondary-succession pioneer-species climax-species keystone-species ecosystem-engineer foundation-species dominant-species subordinate-species transient-species resident-species migrant-species nomadic-species irruptive-species invasive-species naturalized-species casual-species established-species spreading-species widespread-species restricted-species endemic-species cosmopolitan-species disjunct-distribution relict-distribution refugium glacial-refugium interglacial-refugium postglacial-colonization range-shift range-expansion range-contraction range-fragmentation source-population sink-population metapopulation population-viability minimum-viable-population effective-population-size genetic-rescue demographic-rescue assisted-migration managed-relocation ex-situ-conservation in-situ-conservation cryopreservation seed-bank gene-bank botanic-garden arboretum zoo aquarium microbial-culture-collection DNA-bank tissue-bank sperm-bank oocyte-bank embryo-bank reintroduction translocation reinforcement supplementation headstarting captive-breeding propagation nucleus-population release-program soft-release hard-release acclimatization habitat-preparation predator-control competitor-control disease-control parasite-control health-screening quarantine veterinary-care nutrition-management environmental-enrichment behavioral-training imprinting fostering cross-fostering hand-rearing parent-rearing semi-natural-enclosure pre-release-conditioning survival-rate reproduction-rate dispersal-rate site-fidelity homeward-orientation migration navigation orientation magnetic-sense olfactory-sense visual-sense auditory-sense mechanosensory-sense thermosensory-sense hygrosensory-sense electrosensory-sense polarized-light celestial-cues landmark-cues odor-plume wind-direction sun-compass star-compass moon-compass magnetic-compass inertial-navigation path-integration cognitive-map spatial-memory temporal-memory episodic-memory procedural-memory associative-learning operant-conditioning classical-conditioning habituation sensitization song-learning tool-use problem-solving innovation social-learning cultural-transmission teaching cooperation altruism kin-selection reciprocal-altruism group-selection multilevel-selection selfish-gene inclusive-fitness reproductive-skew dominance-hierarchy territoriality home-range activity-center den-site nest-site roost-site singing-post display-site lek arena scrape burrow cavity crevice hole tunnel gallery chamber cell comb paper-nest mud-nest silk-nest spun-cocoon pupal-case ootheca egg-mass clutch brood litter covey pack herd flock colony society superorganism eusociality cooperative-breeding communal-breeding polygyny polyandry promiscuity monogamy serial-monogamy genetic-monogamy social-monogamy extra-pair-copulation sperm-competition cryptic-female-choice sexual-selection intersexual-selection intrasexual-selection mate-choice mate-guarding mate-switching divorce widowhood mate-replacement breeding-season breeding-cycle breeding-attempt breeding-success fledging-success recruitment juvenile-survival adult-survival longevity life-expectancy maximum-lifespan aging telomere oxidative-damage mitochondrial-dysfunction cellular-senescence apoptosis necrosis autophagy proteostasis DNA-repair epigenetic-modification transgenerational-effect maternal-effect paternal-effect parental-care maternal-care paternal-care biparental-care alloparental-care brood-care egg-care larval-care pupal-care food-provisioning nest-defense territory-defense predator-defense disease-defense parasite-defense hygiene-behavior grooming nest-cleaning cannibalism siblicide infanticide filial-cannibalism sexual-cannibalism cannibalistic-oophagy trophic-egg regurgitation trophallaxis food-sharing information-sharing alarm-call mobbing distraction-display broken-wing-display tailing persecution attack escape flight fight freeze faint death-feigning tonic-immobility startle-response withdrawal avoidance dishabituation latent-inhibition blocking overshadowing relative-validity conditioned-inhibition spontaneous-recovery renewal reinstatement rapid-reacquisition latent-learning insight-learning observational-learning vicarious-learning emulation imitation social-facilitation social-inhibition audience-effect bystander-effect conformity obedience compliance commitment consistency foot-in-the-door door-in-the-face low-ball that's-not-all sunk-cost endowment-effect loss-aversion risk-aversion risk-seeking probability-weighting hyperbolic-discounting temporal-discounting intertemporal-choice optimal-foraging marginal-value-theorem prey-choice patch-choice diet-breadth central-place-foraging load-size travel-time handling-time search-time encounter-rate prey-density prey-profitability prey-handling prey-detection prey-pursuit prey-capture prey-consumption prey-rejection ingestion-rate digestion-rate assimilation-efficiency production-efficiency ecological-efficiency trophic-transfer-efficiency Lindeman-efficiency ten-percent-rule pyramid-of-numbers pyramid-of-biomass pyramid-of-energy food-chain trophic-network energy-flow material-cycling biogeochemical-cycle carbon-cycle nitrogen-cycle phosphorus-cycle sulfur-cycle water-cycle hydrological-cycle atmospheric-circulation ocean-circulation thermohaline-circulation El-Niño La-Niña ENSO NAO AO PDO AMO climate-oscillation climate-anomaly weather-extreme drought flood heat-wave cold-snap storm hurricane typhoon cyclone tornado thunderstorm lightning wildfire volcanic-eruption earthquake tsunami landslide avalanche meteor-impact mass-extinction background-extinction speciation-rate extinction-rate net-diversification turnover standing-diversity fossil-record taphonomy preservation-bias sampling-bias collection-bias publication-bias citation-bias language-bias geographic-bias temporal-bias taxonomic-bias size-bias abundance-bias habitat-bias facies-bias Lagerstätte Konservat-Lagerstätte Konzentrat-Lagerstätte trace-fossil body-fossil chemical-fossil molecular-fossil biomarker isotope-signature paleoclimate-proxy paleoecology paleobiogeography paleoecosystem paleocommunity paleoenvironment paleogeography paleolatitude paleolongitude continental-drift plate-tectonics seafloor-spreading subduction orogeny mountain-building basin-formation sedimentation erosion weathering soil-formation pedogenesis horizon-development profile-development chronosequence toposequence biosequence climosequence landform geomorphology topography relief elevation slope aspect curvature roughness drainage watershed catchment basin subbasin reach segment zone riffle pool run glide cascade waterfall wetland marsh swamp bog fen mire peatland tundra taiga boreal-forest temperate-forest temperate-rainforest temperate-grassland tropical-forest tropical-rainforest tropical-dry-forest tropical-savanna tropical-grassland desert hot-desert cold-desert semiarid steppe prairie pampas veld savanna woodland shrubland chaparral maquis garigue fynbos kwongan mallee mulga mogote karst cave sinkhole doline polje uvala ponor spring seep oasis wadi arroyo gully ravine canyon gorge valley dale glen strath rift graben horst fault fold anticline syncline monocline dome plateau mesa butte escarpment bluff cliff crag tor inselberg bornhardt kopje hill knoll mound ridge spur saddle pass col gap notch cleft fissure joint bedding-plane foliation lineation cleavage schistosity gneissic-banding magma lava igneous-rock sedimentary-rock metamorphic-rock mineral crystal grain texture structure fabric composition chemistry mineralogy petrology geochemistry geophysics geochronology radiometric-dating isotopic-dating dendrochronology varve-chronology lichenometry archaeomagnetism paleomagnetism thermoluminescence optically-stimulated-luminescence electron-spin-resonance cosmogenic-nuclide exposure-dating burial-dating terrestrial-cosmogenic-nuclide in-situ-cosmogenic-nuclide meteoric-cosmogenic-nuclide fallout-radionuclide lead-210 cesium-137 plutonium americium radiocarbon carbon-14 calibration-curve intcal marine-reservoir-effect terrestrial-reservoir-effect hard-water-effect contamination pretreatment combustion graphitization accelerator-mass-spectrometry liquid-scintillation-counting gas-proportional-counting beta-counting gamma-spectrometry alpha-spectrometry mass-spectrometry inductively-coupled-plasma-mass-spectrometry thermal-ionization-mass-spectrometry secondary-ion-mass-spectrometry laser-ablation-inductively-coupled-plasma-mass-spectrometry electron-microprobe scanning-electron-microscopy transmission-electron-microscopy light-microscopy confocal-microscopy fluorescence-microscopy polarizing-microscopy phase-contrast-microscopy differential-interference-contrast-microscopy dark-field-microscopy bright-field-microscopy stereomicroscopy dissecting-microscopy microtomy sectioning staining mounting embedding fixation preservation collection curation vouchering repository herbarium museum archive database registry catalogue checklist inventory survey monitoring assessment evaluation appraisal audit review synthesis rapid-review scoping-review narrative-review critical-review state-of-the-art-review literature-review bibliometric-analysis citation-analysis co-citation-analysis bibliographic-coupling co-authorship-analysis collaboration-network knowledge-map concept-map ontology taxonomy classification nomenclature typification designation description diagnosis illustration measurement morphometry meristics count ratio index score rank grade scale level category class group set subset cluster clade lineage branch stem crown root node tip terminal internal basal derived ancestral plesiomorphic apomorphic synapomorphic autapomorphic homologous homoplastic convergent parallel reversal gain loss duplication deletion insertion inversion fusion fission polyploidy aneuploidy chromosome-number genome-size gene-number transcriptome proteome metabolome lipidome glycome interactome phenome connectome epigenome microbiome virome mycobiome bacteriome phytobiome rhizobiome endobiome ectobiome parasitome pathobiome diseasome toxome exposome foodome nutriome pharmacome toxicome adverse-outcome-pathway mode-of-action mechanism-of-action target-site receptor-binding enzyme-inhibition ion-channel-blockade membrane-disruption DNA-intercalation inflammation cell-differentiation cell-migration cell-adhesion cell-signaling signal-transduction gene-expression transcription translation post-translational-modification protein-folding protein-degradation protein-trafficking vesicle-transport exocytosis endocytosis phagocytosis pinocytosis mitophagy pexophagy ribophagy ER-phagy lysophagy lipophagy glycophagy ferritinophagy aggrephagy xenophagy entosis cytosis syncytium plasmodesmata gap-junction tight-junction adherens-junction desmosome hemidesmosome focal-adhesion cell-polarity cell-asymmetry cell-division mitosis meiosis cytokinesis karyokinesis chromosome-segregation spindle-assembly checkpoint DNA-replication DNA-recombination transposition retrotransposition horizontal-gene-transfer vertical-gene-transfer endosymbiosis symbiogenesis holobiont hologenome extended-phenotype niche-construction ecosystem-engineering keystone-process ecological-function ecosystem-function ecosystem-good natural-capital natural-resource renewable-resource non-renewable-resource common-pool-resource public-good private-good club-good externality market-failure government-failure policy-failure institutional-failure tragedy-of-the-commons free-rider collective-action social-dilemma prisoner's-dilemma chicken-game assurance-game coordination-game zero-sum-game positive-sum-game negative-sum-game win-win lose-lose win-lose Pareto-optimal Pareto-improvement Kaldor-Hicks-improvement cost-benefit-analysis cost-effectiveness-analysis multi-criteria-decision-analysis decision-support-system expert-system knowledge-based-system rule-based-system case-based-reasoning machine-learning deep-learning neural-network random-forest support-vector-machine decision-tree regression-tree classification-tree clustering-algorithm dimensionality-reduction feature-selection feature-extraction pattern-recognition image-analysis signal-processing time-series-analysis spatial-analysis geographic-information-system remote-sensing satellite-imagery aerial-photography drone-imagery LiDAR radar sonar seismic geophysical geochemical biogeochemical ecoinformatics bioinformatics computational-biology systems-biology synthetic-biology biotechnology nanotechnology genetic-engineering genome-editing CRISPR TALEN zinc-finger-nuclease transgenic cisgenic intragenic subgenic agrobacterium-mediated-transformation biolistics electroporation microinjection protoplast-fusion somatic-hybridization cybrid asymmetric-hybrid synthetic-polyploid doubled-haploid haploid-induction anther-culture microspore-culture ovule-culture embryo-culture endosperm-culture protoplast-culture cell-culture tissue-culture organ-culture shoot-culture root-culture callus-culture suspension-culture bioreactor hairy-root-culture transformation hardening micropropagation clonal-propagation meristem-culture shoot-tip-culture axillary-bud-culture nodal-culture leaf-culture petiole-culture stem-culture internode-culture rhizome-culture bulb-culture corm-culture tuber-culture seed-culture zygotic-embryo-culture somatic-embryo-culture somatic-embryogenesis organogenesis caulogenesis rhizogenesis adventitious-shoot-formation direct-regeneration indirect-regeneration primary-metabolism secondary-metabolism biosynthesis biotransformation bioconversion biodegradation bioremediation phytoremediation mycoremediation zooremediation bioaugmentation biostimulation natural-attenuation monitored-natural-attenuation engineered-natural-attenuation permeable-reactive-barrier pump-and-treat soil-vapor-extraction air-sparging thermal-desorption incineration landfilling containment capping slurry-wall grout-curtain sheet-piling clay-liner geomembrane geotextile geogrid geocomposite geosynthetic geofabric geonet geocell geopipe geostrip geofoam geobag geotube geocontainer geomat geoblanket geospacer geodrain geoinjection geogrout geopier geocolumn geopile geowall geoberm geotrench geocut geofill geolift geoslab geopanel geoblock geobrick geotile geopaver geoweb geomattress geocushion
Tortopus
Tortopus is a genus of burrowing mayflies in the family Polymitarcyidae, restricted to six species based on cladistic revision: T. igaranus, T. circumfluus, T. harrisi, T. zottai, T. bellus, and T. arenales. The genus is characterized by distinctive morphological synapomorphies including female parastyli receptors with long anterior furrows, entirely flattened penes, and nymphs bearing two subapical tubercles on mandibular tusks. Nymphs construct U-shaped tunnels in clay banks, a behavior that renders them difficult to sample with standard aquatic survey methods.
Ephemeroptera Polymitarcyidae burrowing-mayflies Neotropical clay-bank-habitat U-shaped-tunnels mandibular-tusks Tortopsis Campsurinae Panamerican-distribution cladistic-revision synapomorphy parastyli-receptors flattened-penes furcasternal-protuberances subapical-tubercles limnological-sampling-bias nymph-morphology adult-morphology type-species-T.-igaranus new-species-T.-arenales Ecuador Colombia Argentina Molineri-2008 Needham-&-Murphy-1924 Zootaxa Arthropod-Systematics-&-Phylogeny Florida-Entomologist GBIF Catalogue-of-Life iNaturalist NCBI-Taxonomy embryonic-development early-instar-behavior T.-harrisi T.-bellus T.-circumfluus T.-zottai T.-igaranus T.-incertus T.-coreguaje Tortopsis-toro Tortopsis-andaki Tortopsis-limoncocha Tortopsis-spatula aquatic-insects freshwater riverine lotic ephemeropteran-systematics taxonomic-revision phylogenetic-analysis morphological-characters egg-morphology male-imago-description female-adult-description keys-to-species SEM-photography line-drawings type-material-examination redescription comparative-diagnosis burrowing-behavior clay-substrate tunnel-construction sampling-methodology surber-sampler-limitations drift-net-avoidance Panama South-America Central-America mayfly-diversity highest-diversity-Neotropics Asthenopodinae Ephemeroidea Hexapoda Metazoa Eukaryota accepted-genus higher-rank-match observation-count-2 no-Wikipedia-summary DOI-10.11646/zootaxa.2481.1.1 DOI-10.11646/zootaxa.2436.1.4 DOI-10.3897/asp.79.e62735 DOI-10.2307/3493493 Scott-et-al.-1959 Navás Ulmer Traver Lugo-Ortiz-&-McCafferty Domínguez Banks McDunnough Pictet Needham-&-Murphy freshwater-ecology aquatic-entomology insect-development nymphal-stages imago-description alate-adults wing-venation R-sector gonopore-structure parastyli-length pedestal-morphology lateral-view-curvature mandibular-morphology tusk-tubercles mesosternal-structure abdominal-sternum-notch furcasternum protuberance-configuration receptor-socket-orientation IR-vein R2-vein claw-like-structure gonopore-association penis-base-separation female-fore-wing additional-veins comparative-morphology type-specimen holotype allotype paratype neotropical-species new-species-description species-key identification-key male-adult-key female-adult-key nymph-key generic-key Polymitarcyidae-genera systematics phylogeny cladistics monophyly sister-group character-state transformation-series parsimony-analysis morphological-data-set external-morphology egg-chorion micropyle egg-burster hatching emergence behavioral-study early-instar embryology oviposition substrate-preference habitat-specificity endemic regional-diversity conservation biomonitoring water-quality-indicator macroinvertebrate benthic infaunal interstitial hyporheic river-bank stream-bank erosion sediment particle-size clay-content bank-stability root-mat vegetation-association flow-regime current-velocity depth width stream-order catchment basin ecoregion biogeography dispersal colonization vicariance speciation adaptive-radiation morphological-divergence convergent-evolution homoplasy character-conflict bootstrap-support Bremer-support consensus-tree strict-consensus majority-rule taxonomic-stability nomenclatural-act original-description subsequent-designation type-locality type-depository museum-collection specimen-preservation alcohol-preservation slide-mounting SEM-preparation imaging-techniques scientific-illustration photography measurement morphometrics ratios proportions allometry sexual-dimorphism geographic-variation intraspecific-variation interspecific-variation diagnostic-characters identification-confidence taxonomic-expertise literature-review bibliographic-search database-query online-repository open-access paywall abstract-only full-text-unavailable citation-analysis reference-list source-quality evidence-strength knowledge-gap research-need future-study collaboration field-work collection-trip sampling-effort specimen-availability type-examination revision-scope taxonomic-concept circumscription inclusion exclusion species-group species-complex cryptic-species morphospecies phylogenetic-species evolutionary-significant-unit operational-taxonomic-unit DNA-barcoding molecular-systematics integrative-taxonomy traditional-taxonomy alpha-taxonomy beta-taxonomy gamma-taxonomy classification-hierarchy Linnaean-system rank-based-nomenclature phylogenetic-nomenclature clade-name crown-group stem-group total-evidence congruence incongruence data-partition character-weighting implied-weighting equal-weighting ordered-characters unordered-characters multistate-characters binary-characters continuous-characters meristic-characters qualitative-characters quantitative-characters morphological-matrix data-matrix character-matrix taxon-sampling character-sampling missing-data inapplicable-data polymorphism intraspecific-polymorphism fixed-difference diagnostic-difference unique-derived-character autapomorphy shared-derived-character symplesiomorphy convergent-character parallel-evolution reversal character-optimization ACCTRAN DELTRAN unambiguous-optimization ambiguous-optimization most-parsimonious-tree tree-length consistency-index retention-index rescaled-consistency-index homoplasy-index character-fit character-congruence character-support branch-support node-support clade-stability taxonomic-stability-index phylogenetic-signal noise saturation substitution-model rate-heterogeneity among-site-rate-variation gamma-distribution invariant-sites model-selection Akaike-information-criterion Bayesian-information-criterion likelihood-ratio-test Bayesian-inference Markov-chain-Monte-Carlo posterior-probability prior-probability hyperprior convergence effective-sample-size burn-in stationarity trace-plot parameter-estimate credible-interval highest-posterior-density tree-topology tree-shape lineage-through-time diversification-rate extinction-rate speciation-rate net-diversification birth-death-model Yule-model coalescent-model population-size effective-population-size gene-flow migration-rate isolation-by-distance phylogeography historical-biogeography ancestral-area-reconstruction dispersal-vicariance-analysis decisive-weighting event-based-method taxon-area-cladogram area-cladogram biogeographic-homology track-analysis panbiogeography phylogenetic-biogeography ecological-biogeography dispersal-biogeography vicariance-biogeography center-of-origin area-of-endemism refugium Pleistocene glaciation climate-change habitat-fragmentation habitat-loss anthropogenic-impact pollution eutrophication sedimentation dam-construction water-abstraction flow-modification thermal-pollution toxic-substances heavy-metals pesticides urbanization agriculture deforestation land-use-change riparian-degradation channelization dredging mining invasive-species biological-invasion competition predation parasitism mutualism commensalism amensalism facilitation ecosystem-engineer keystone-species flagship-species umbrella-species indicator-species sentinel-species bioindicator ecological-indicator environmental-indicator water-quality-index biotic-index multimetric-index rapid-bioassessment protocol standardization quality-assurance quality-control reference-condition least-disturbed minimally-disturbed best-available type-specific region-specific stream-type river-type functional-feeding-group collector-gatherer collector-filterer scraper shredder predator parasite engulfer piercer burrower sprawler clinger swimmer skater diver surface-dweller benthic-dweller interstitial-dweller hyporheic-dweller phreatic-dweller troglobite stygobite cave-inhabitant spring-inhabitant headwater mid-order high-order lowland montane alpine subtropical tropical temperate boreal arctic antarctic palearctic nearctic afrotropical oriental australian oceanic cosmopolitan widespread restricted relict disjunct-distribution range-extension new-record first-record country-record state-record province-record locality-record elevation-record altitudinal-range latitudinal-range longitudinal-range distribution-pattern range-size range-fill extent-of-occurrence area-of-occupancy IUCN-category threat-status data-deficient not-evaluated least-concern near-threatened vulnerable endangered critically-endangered extinct-in-wild extinct population-trend population-density population-structure age-structure size-structure sex-ratio reproductive-output fecundity fertility mortality survivorship life-table cohort-analysis generation-time doubling-time intrinsic-rate-of-increase finite-rate-of-increase net-reproductive-rate stable-age-distribution stationary-population expanding-population declining-population source-sink-dynamics metapopulation population-viability-analysis minimum-viable-population genetic-diversity heterozygosity allelic-richness private-allele founder-effect bottleneck genetic-drift inbreeding inbreeding-depression outbreeding outbreeding-depression hybridization introgression gene-flow-barrier reproductive-isolation prezygotic-isolation postzygotic-isolation mechanical-isolation temporal-isolation habitat-isolation behavioral-isolation gametic-isolation zygotic-mortality hybrid-inviability hybrid-sterility hybrid-breakdown reinforcement character-displacement ecological-speciation allopatric-speciation sympatric-speciation parapatric-speciation peripatric-speciation centrifugal-speciation divergence-with-gene-flow speciation-continuum anagenesis cladogenesis budding-speciation branching-speciation phyletic-gradualism punctuated-equilibrium stasis trend progression orthogenesis aristogenesis mosaic-evolution co-evolution arms-race Red-Queen-hypothesis evolutionary-stable-strategy frequency-dependent-selection density-dependent-selection r-selection K-selection bet-hedging iteroparity semelparity ploidy-level chromosome-number karyotype genome-size transposable-element gene-duplication neofunctionalization subfunctionalization pseudogene regulatory-evolution cis-regulatory trans-regulatory enhancer promoter silencer insulator transcription-factor signaling-pathway developmental-pathway gene-network epigenetic-modification DNA-methylation histone-modification chromatin-remodeling genomic-imprinting parental-effect maternal-effect paternal-effect cytoplasmic-inheritance mitochondrial-DNA chloroplast-DNA endosymbiont Wolbachia Rickettsia Spiroplasma Cardinium Arsenophonus Hamiltonella Regiella Serratia Buchnera Carsonella Portiera primary-endosymbiont secondary-endosymbiont facultative-symbiont obligate-symbiont mutualistic-symbiont parasitic-symbiont commensal-symbiont pathogen vector disease-transmission zoonosis emerging-infectious-disease biosecurity quarantine eradication control management conservation-biology restoration-ecology habitat-restoration river-restoration stream-restoration floodplain-restoration riparian-restoration wetland-restoration ecosystem-restoration passive-restoration active-restoration assisted-recovery managed-relocation assisted-colonization ex-situ-conservation in-situ-conservation protected-area national-park nature-reserve wildlife-refuge biosphere-reserve world-heritage-site ramsar-site important-bird-area key-biodiversity-area Alliance-for-Zero-Extinction EDGE-species evolutionarily-distinct globally-endangered conservation-prioritization systematic-conservation-planning Marxan Zonation conservation-opportunity conservation-cost conservation-benefit trade-off socioeconomic-factor stakeholder-engagement community-based-conservation adaptive-management monitoring-program long-term-study citizen-science public-participation environmental-education outreach science-communication knowledge-transfer capacity-building institutional-strengthening policy-development legislation regulation enforcement compliance incentive payment-for-ecosystem-services carbon-credit biodiversity-offset mitigation-banking conservation-easement land-trust private-land-conservation conservation-covenant stewardship sustainable-use sustainable-management ecosystem-based-management integrated-water-resources-management basin-management catchment-management landscape-management seascape-management connectivity-conservation corridor stepping-stone buffer-zone matrix-management countryside-biogeography reconciliation-ecology novel-ecosystem designer-ecosystem ecosystem-service provisioning-service regulating-service supporting-service cultural-service food-production water-supply raw-materials genetic-resources medicinal-resources ornamental-resources air-quality-regulation climate-regulation water-regulation erosion-regulation water-purification waste-treatment disease-regulation pest-regulation pollination seed-dispersal habitat-provision nutrient-cycling soil-formation primary-production recreation tourism aesthetic-value spiritual-value educational-value scientific-value inspiration sense-of-place cultural-heritage traditional-knowledge indigenous-knowledge local-knowledge ecosystem-service-valuation economic-valuation contingent-valuation choice-experiment benefit-transfer total-economic-value use-value non-use-value direct-use indirect-use option-value bequest-value existence-value willingness-to-pay willingness-to-accept cost-benefit-analysis cost-effectiveness-analysis multi-criteria-analysis decision-support uncertainty risk-assessment scenario-analysis sensitivity-analysis robustness-analysis adaptive-capacity resilience vulnerability exposure sensitivity coping-capacity transformation social-ecological-system complex-adaptive-system tipping-point regime-shift alternative-stable-state hysteresis path-dependence legacy-effect slow-variable fast-variable panarchy adaptive-cycle exploitation release reorganization foresight anticipatory-governance transformative-change sustainability-science future-earth interdisciplinary transdisciplinary problem-oriented solution-oriented action-oriented knowledge-co-production boundary-work boundary-object boundary-organization science-policy-interface science-society-interface knowledge-brokerage knowledge-exchange knowledge-integration mode-1-science mode-2-science triple-helix quadruple-helix responsible-innovation reflexive-governance adaptive-governance collaborative-governance polycentric-governance network-governance self-organization collective-action common-pool-resource tragedy-of-the-commons social-dilemma institutional-analysis institutional-design Ostrom-design-principles monitoring sanctioning conflict-resolution nested-enterprises proportionality collective-choice constitutional-choice operational-choice collective-action-theory game-theory prisoner's-dilemma chicken-game assurance-game coordination-game evolutionary-game-theory replicator-dynamics ESS Nash-equilibrium Pareto-optimality social-optimum market-failure government-failure voluntary-failure non-market-value public-good common-good club-good private-good rivalry excludability free-rider hold-up-problem moral-hazard adverse-selection information-asymmetry transaction-cost property-right common-property state-property private-property hybrid-property bundle-of-rights usufruct access withdrawal alienation tenure-security land-tenure water-tenure resource-tenure customary-tenure statutory-tenure legal-pluralism de-facto de-jure formal informal legality legitimacy recognition deviance corruption elite-capture social-capital bonding-capital bridging-capital linking-capital trust reciprocity reputation social-network social-embeddedness structural-hole brokerage closure cohesion density centrality betweenness degree eigenvector PageRank small-world scale-free random-network regular-network hierarchical-network modularity community-structure clustering assortativity homophily heterophily preferential-attachment fitness-model network-evolution network-dynamics temporal-network multilayer-network multiplex-network bipartite-network affiliation-network one-mode-network two-mode-network adjacency-matrix edge-list graph-theory vertex edge node link tie connection path cycle tree forest component giant-component percolation phase-transition criticality self-organized-criticality power-law heavy-tail Pareto-distribution lognormal-distribution exponential-distribution Poisson-distribution binomial-distribution normal-distribution Gaussian-distribution statistical-distribution probability-distribution cumulative-distribution probability-density-function cumulative-distribution-function survival-function hazard-function moment mean median mode variance standard-deviation coefficient-of-variation skewness kurtosis quantile percentile decile quartile interquartile-range range minimum maximum outlier robust-statistic parametric-test non-parametric-test hypothesis-testing null-hypothesis alternative-hypothesis significance-level p-value confidence-interval type-I-error type-II-error statistical-power effect-size sample-size sampling-design experimental-design observational-study quasi-experiment natural-experiment randomized-controlled-trial before-after-control-impact BACI impact-assessment environmental-impact-assessment strategic-environmental-assessment cumulative-impact-assessment social-impact-assessment health-impact-assessment life-cycle-assessment carbon-footprint water-footprint ecological-footprint material-flow-analysis substance-flow-analysis energy-analysis exergy-analysis emergy-analysis input-output-analysis environmentally-extended-input-output social-accounting-matrix computable-general-equilibrium agent-based-model system-dynamics-model Bayesian-network structural-equation-model path-analysis factor-analysis principal-component-analysis cluster-analysis discriminant-analysis canonical-correspondence-analysis redundancy-analysis non-metric-multidimensional-scaling ordination classification regression-analysis linear-regression nonlinear-regression generalized-linear-model generalized-additive-model mixed-effects-model multilevel-model hierarchical-model Bayesian-hierarchical-model state-space-model hidden-Markov-model time-series-analysis spatial-analysis geostatistics kriging variogram spatial-autocorrelation Moran's-I Geary's-c Getis-Ord-Gi local-indicator-of-spatial-association LISA hot-spot cold-spot spatial-cluster spatial-outlier edge-effect boundary-effect modifiable-areal-unit-problem MAUP ecological-fallacy atomistic-fallacy cross-scale-inference upscaling downscaling scaling-law allometric-scaling fractal fractal-dimension self-similarity multi-fractal lacunarity roughness texture pattern process pattern-process-relationship neutral-model null-model randomization-test permutation-test bootstrap jackknife cross-validation model-averaging model-comparison information-criterion Bayesian-model-averaging ensemble-modeling consensus-model prediction forecast projection scenario narrative-scenario quantitative-scenario storyline shared-socioeconomic-pathway representative-concentration-pathway emission-scenario climate-scenario impact-scenario adaptation-scenario mitigation-scenario transformation-scenario backcasting forecasting trend-extrapolation expert-judgment Delphi-method participatory-modeling stakeholder-analysis influence-diagram conceptual-model mental-model system-archetype leverage-point intervention-point policy-instrument economic-instrument regulatory-instrument voluntary-instrument information-instrument market-based-instrument command-and-control-instrument subsidy tax charge fee levy deposit-refund tradable-permit emission-trading carbon-tax biodiversity-banking conditional-cash-transfer community-based-natural-resource-management joint-forest-management co-management adaptive-co-management transboundary-management international-regime global-governance multilateral-environmental-agreement convention treaty cartagena-protocol nagoya-protocol Aichi-target sustainable-development-goal SDG post-2020-global-biodiversity-framework Kunming-Montreal-global-biodiversity-framework 30-by-30-target nature-based-solution NbS ecosystem-based-adaptation ecosystem-based-mitigation green-infrastructure blue-infrastructure natural-infrastructure hybrid-infrastructure climate-smart-agriculture climate-smart-landscape climate-smart-development low-carbon-development green-economy circular-economy bioeconomy blue-economy sustainable-consumption sustainable-production life-cycle-thinking cradle-to-cradle industrial-ecology industrial-symbiosis urban-metabolism socio-ecological-metabolism material-intensity energy-intensity carbon-intensity decoupling absolute-decoupling relative-decoupling resource-efficiency eco-efficiency eco-innovation green-technology clean-technology environmental-technology appropriate-technology participatory-technology technology-transfer technology-diffusion innovation-system national-innovation-system sectoral-innovation-system technological-innovation-system socio-technical-system socio-technical-transition multi-level-perspective niche-innovation regime landscape transition-management strategic-niche-management transition-experiment living-lab real-world-laboratory urban-living-lab rural-living-lab sustainability-transition fundamental-change system-innovation directionality reflexivity responsiveness anticipation learning-by-doing doing-by-learning social-learning organizational-learning policy-learning transformative-capacity capacity-development institutional-learning evolutionary-governance institutional-change path-dependency lock-in carbon-lock-in technological-lock-in institutional-lock-in behavioral-lock-in cultural-lock-in unlocking destabilization phase-out divestment stranded-asset just-transition transition-pathway transition-scenario deep-decarbonization net-zero carbon-neutral climate-neutral nature-positive biodiversity-positive restoration-positive regenerative circular doughnut-economics planetary-boundary safe-operating-space environmental-ceiling social-foundation intergenerational-equity intragenerational-equity distributive-justice procedural-justice recognition-justice environmental-justice climate-justice energy-justice water-justice food-justice spatial-justice territorial-justice ecological-justice species-justice intersectionality adaptation mitigation sustainability sustainable-development well-being human-development capability-approach human-security environmental-security climate-security water-security food-security energy-security livelihood-security disaster-risk-reduction climate-change-adaptation climate-change-mitigation loss-and-damage climate-finance green-climate-fund adaptation-fund global-environment-facility biodiversity-finance nature-finance blended-finance impact-investment ESG environmental-social-governance sustainable-finance green-bond social-bond sustainability-bond blue-bond nature-bond biodiversity-credit offset inset nature-based-credit ecosystem-service-credit payment-for-results results-based-payment conditional-payment performance-based-payment outcome-based-payment environmental-service-payment PSA PES REDD REDD+AFOLU LULUCF NAMA NAMC NDC NAP NBSAP national-adaptation-plan national-biodiversity-strategy-and-action-plan nationally-determined-contribution nationally-appropriate-mitigation-action intended-nationally-determined-contribution long-term-low-emission-development-strategy net-zero-target carbon-budget remaining-carbon-budget emission-gap adaptation-gap finance-gap implementation-gap ambition-loop ratchet-mechanism global-stocktake transparency-framework enhanced-transparency-framework common-but-differentiated-responsibilities respective-capabilities climate-equity climate-litigation climate-governance climate-diplomacy climate-negotiation UNFCCC COP CMP CMA SBSTA SBI IPCC IPBES UNCCD CBD CITES Ramsar World-Heritage Man-and-the-Biosphere MAB UNESCO FAO WHO ILO UNDP UNEP UN-Habitat World-Bank IMF WTO OECD GEF GCF AF multilateral-development-bank regional-development-bank national-development-bank development-finance-institution export-credit-agency sovereign-wealth-fund pension-fund insurance-company asset-manager asset-owner corporate-investor retail-investor impact-investor philanthropic-investor development-partner bilateral-donor multilateral-donor private-donor foundation NGO CSO civil-society-organization non-governmental-organization community-based-organization indigenous-peoples-organization women's-organization youth-organization faith-based-organization research-organization think-tank university academic-institution knowledge-institution capacity-building-institution training-institution extension-service advisory-service consulting-firm engineering-firm technology-provider equipment-supplier service-provider contractor operator maintainer utility municipality local-government provincial-government national-government federal-government ministry department agency authority commission 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seed-commons water-commons fisheries-commons forest-commons pasture-commons land-commons urban-commons atmospheric-commons global-commons common-heritage-of-mankind common-concern-of-humankind restorative-justice multispecies-justice more-than-human anthropocene capitalocene plantationocene chthulucene symbiocene good-anthropocene great-acceleration great-transformation great-decoupling great-convergence great-divergence great-fragmentation great-integration great-transition great-work long-now deep-time futures-thinking future-studies strategic-foresight scenario-planning trend-analysis weak-signal wild-card black-swan grey-rhino blue-sky green-swan systemic-change structural-change cultural-change behavioral-change individual-change collective-change social-change organizational-change technological-change economic-change political-change ecological-change planetary-change earth-system-change biodiversity-change ocean-change cryosphere-change atmospheric-change biogeochemical-change 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resolution-sphere-change dissolution-sphere-change absolution-sphere-change solution-sphere-change

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