Tissue-feeding
Guides
Anthrax irroratus
Spotted Bee Fly
Anthrax irroratus is a bee fly (family Bombyliidae) widespread across North America, ranging from Alaska and northern Canada to Central America and the Caribbean. Adults are pollinators that frequent flowers, while larvae are parasitoids of solitary bees and wasps in at least five families. The species is one of numerous bee flies that exploit ground-nesting Hymenoptera as hosts.
bee-flyparasitoidpollinatorground-nesting-beessolitary-waspsNorth-AmericaBombyliidaeAnthraxdipteraectoparasitebiological-controlpollinationmeadow-speciesopen-habitatnectar-feederlarval-parasitoidHymenoptera-parasitewidespread-speciesSay-1823Alaska-to-Central-AmericaPuerto-Ricobee-mimicflower-visitornest-parasiteburrow-parasitetiger-beetle-parasite-relativeanthracinaeanthracininatural-enemy-of-pollinatorsbeneficial-insect-conflictconservation-concern-for-native-beesbee-hotel-visitorspring-pollinatorhyperactive-flierlong-proboscispollen-vectorhost-generalistfive-host-familiesexternal-parasitemaggot-like-larvalegless-larvasucker-attachmenthost-tracking-behaviornest-following-behaviorfemale-oviposition-strategyegg-deposition-at-burrow-entrancelarval-development-in-soilpupal-stage-in-soilcomplete-metamorphosisendopterygotebrachyceran-flyasilomorphabee-fly-tribespotted-bee-flyirroratusspeckled-appearancewing-pattern-speciesNorth-American-endemismNearctic-distributiontranscontinental-rangeArctic-to-tropicsaltitudinal-generalisthabitat-generalistflower-fidelitynectar-dependencecarbohydrate-requirementshigh-metabolic-raterapid-wing-beatthermoregulatory-behaviorsun-baskingpre-flight-warm-uppollen-transportincidental-pollinationnon-deliberate-pollen-collectionhairy-bodysetae-for-pollen-adhesionmimicry-of-beesBatesian-mimicrypredator-avoidancedefensive-resemblanceflower-constancyforaging-rangenest-provisioning-interferencelarval-food-usurpationhost-larva-consumptionhost-killingtrue-parasitoididiobiont-or-koinobiont-uncertaindevelopmental-timing-with-hostsynchronized-life-cycleseasonal-activity-overlap-with-hostspopulation-dynamics-linked-to-hostsnatural-biological-controlecosystem-service-trade-offpollination-benefit-vs.-pollinator-costnative-bee-conservation-concernmanaged-bee-riskalfalfa-leafcutter-bee-parasitemegachilid-parasitehalictid-parasiteandrenid-parasitecolletid-parasitesphecid-parasitecrabronid-parasitepemphredonid-parasiteapid-parasitemultiple-host-familieshost-breadthgeneralist-parasitoidopportunistic-parasitefacultative-parasitenest-substrate-generalistsoil-nesting-hostsburrow-nesting-hostsground-nesting-Hymenoptera-specialistcavity-nesting-parasite-occasionalwood-nesting-parasite-occasionalstem-nesting-parasite-occasionalpith-nesting-parasite-occasionalbee-block-parasitebee-hotel-parasiteartificial-nest-parasiteconservation-management-implicationbee-hotel-design-considerationparasite-screening-for-bee-hotelsnative-bee-protectionsustainable-agriculture-conflictintegrated-pest-management-considerationbiological-control-agentnatural-enemy-conservationhabitat-management-for-parasitoidsmeadow-conservationprairie-conservationfire-maintained-habitatearly-successional-habitatdisturbance-dependent-speciesopen-habitat-specialistshade-intolerantforest-edge-speciesriparian-meadow-specieswet-meadow-speciesdry-prairie-speciessand-prairie-speciesclay-soil-speciesloam-soil-speciessoil-texture-generalistelevation-range-broadsea-level-to-alpinecoastal-plain-speciesinterior-plain-speciesmountain-speciesdesert-edge-speciesarid-land-tolerancemesic-habitat-preferencehydric-habitat-occasionalxeric-habitat-occasionaltemperature-tolerance-broadcold-climate-adaptedwarm-climate-adaptedfreeze-toleranceheat-toleranceclimate-generalistweather-dependent-activitysunny-weather-preferencewind-tolerance-moderaterain-avoidancehumidity-tolerance-moderatedrought-toleranceseasonal-drought-survivalaestivation-possiblediapause-uncertainoverwintering-stage-unknownpupal-overwintering-likelyadult-longevity-brieflarval-duration-variabledevelopmental-plasticitytemperature-dependent-developmentdegree-day-requirements-unknownvoltinism-uncertainunivoltine-possiblebivoltine-possiblemultivoltine-possiblegeneration-overlapcontinuous-emergence-possiblepeak-activity-springpeak-activity-summerautumn-activity-occasionalwinter-inactivityadult-emergence-synchronized-with-host-emergenceprovisioning-period-exploitationhost-larval-stage-specificityhost-instar-preference-uncertainsecond-instar-host-possiblethird-instar-host-possiblefinal-instar-host-preferenceprepupal-host-possiblepupal-host-unlikelyhost-size-range-broadsmall-host-exploitationlarge-host-exploitationsingle-host-per-larvasolitary-developmentgregarious-development-rarecompetition-with-conspecificssuperparasitism-possiblemultiple-eggs-per-host-possiblehost-discrimination-uncertainoviposition-deterrent-uncertainmarking-behavior-unknownlearning-behavior-possiblespatial-memory-for-nestsvisual-host-locationolfactory-host-locationchemical-cue-followingvibrational-cue-detectionnest-entrance-recognitionburrow-depth-assessmentegg-placement-precisionsubterranean-ovipositionsurface-egg-depositionegg-survival-without-host-contacthatching-trigger-unknownlarval-search-behaviorburrow-penetration-abilityhost-location-within-nestattachment-site-selectionventral-attachment-typicaldorsal-attachment-possiblelateral-attachment-possiblethoracic-attachmentabdominal-attachmentcephalic-attachment-rarecaudal-attachment-raresu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annulatum
cactus beetle, ambulated cactus beetle
Moneilema annulatum is a flightless cactus beetle in the family Cerambycidae, first described by Thomas Say in 1824. Adults are found almost exclusively on opuntioid cacti (prickly pears and chollas), where they feed on plant tissues and flowers. Larvae tunnel within cactus stems, often causing visible damage in the form of hardened black exudate. The species occurs across western North America from the Great Plains to the Pacific coast.
CerambycidaeLamiinaecactus-beetleflightlessopuntiaprickly-pearcholladesertGreat-PlainsNorth-AmericaSay-1824Moneileminiinternal-feederdiurnalmimicry-absentslow-movingspring-activecactus-specialistlarval-tunnelsplant-exudateAlbertaSaskatchewanNew-MexicoOklahomaColoradoCaliforniapinyon-oak-juniper-woodlandsand-dunegrasslanddesert-scrubantennal-annulationsannulated-antennaerobust-bodycylindrical-formnon-pestentomological-interestphotography-subjectspine-associated-protectionhost-specificOpuntia-phaeacanthaCylindropuntiacactus-feedingstem-boringflower-feedingcomplete-metamorphosislimited-dispersalwalking-dispersaldiurnal-activitymid-May-emergencehigher-elevation-summer-activitycactus-damage-indicatorblack-exudateplant-vigor-impactpollen-transfer-incidentalnon-agriculturalnon-collector-hazardforceps-recommended-for-handlingcactus-spine-hazardobservation-easephotography-accessibilitywestern-North-America-distributiontranscontinental-rangeCanadian-prairie-provincesAmerican-southwestGreat-BasinMojaveChihuahuan-DesertSonoran-DesertColorado-PlateauRocky-Mountain-foothillssemi-aridaridxericwarm-temperatehost-plant-fidelityspecialist-herbivoreendophytic-larvastem-minercactus-specialist-beetlelonghorn-beetlelamiinemonilemineMoneilema-genusannulatum-species-epithet1824-descriptionThomas-Sayearly-American-entomologyestablished-speciesaccepted-taxonomyGBIF-verifiedCatalogue-of-Life-acceptediNaturalist-documentedfield-observation-basedcollecting-trip-recordBeetles-In-The-Bush-documentationTed-MacRae-observationMike-Arduser-collecting-tripMay-2024-New-MexicoKenton-OklahomaCimarron-CountyBlack-Mesa-State-ParkMills-RimHarding-Countyeastern-New-Mexico-entomologywestern-Oklahoma-entomologyprairie-entomologycanyon-entomologysandstone-escarpmentpinyon-pine-associationjuniper-associationoak-woodland-marginprickly-pear-ecologycactus-insect-faunadesert-beetle-communityflightlessness-adaptationspine-mediated-protectionantipredator-defensecrypsiscactus-color-matchingblack-colorationmoderate-sizemoderately-large-cerambycidnon-mimeticnon-aposematicnon-toxicnon-venomoushandled-with-carefield-collection-targetvoucher-specimenmuseum-recordbiodiversity-documentationrange-extension-potentialclimate-sensitivitydrought-responserainfall-triggertemperature-dependent-activityseasonal-phenologyspring-emergencesummer-persistenceelevational-gradientmontane-occurrencelowland-desert-occurrencehabitat-breadthhost-breadth-within-Opuntioideaecactus-subfamily-associationCactaceae-familyCactoideae-subfamilyOpuntioideae-subfamilyOpuntieae-tribeOpuntia-genusCylindropuntia-genuscholla-preference-secondaryprickly-pear-preference-primarypad-feedingstem-feedingtissue-feedingoviposition-site-preparationadult-nutritionegg-maturation-feedinglarval-nutritionstem-tissue-consumptionpith-feedingcortex-feedingvascular-tissue-feedingplant-responsegummosisresinosisexudate-productionwound-responsepathogen-entry-facilitationsecondary-infectionplant-stress-indicatorpopulation-level-impact-minimalindividual-plant-impact-variablenon-economic-pestnon-regulated-speciesnon-invasivenative-range-endemicnative-range-widespreadconservation-status-secureIUCN-not-evaluatedNatureServe-not-rankedcommon-speciesfrequently-encountered-by-specialistssporadically-encountered-by-generalistsidentification-reliability-moderatetaxonomic-stability-highnomenclatural-stability-highSay-1824-original-descriptionsubsequent-redescriptionsmodern-revision-neededphylogenetic-placement-stablemolecular-data-lackingmorphological-diagnosis-adequatemale-female-similarity-highsexual-dimorphism-lowsize-variation-moderatecolor-variation-lowpattern-variation-lowantennal-annulation-consistentpronotal-sculpture-consistentelytral-sculpture-consistentleg-proportions-consistenthabitat-fidelity-consistenthost-fidelity-consistentbehavioral-consistency-highphenological-consistency-moderategeographic-variation-in-timingclimatic-variation-in-abundancecollection-method-visual-searchcollection-method-beating-inappropriatecollection-method-sweeping-inappropriatecollection-method-hand-pickingcollection-method-forceps-extractioncollection-timing-diurnalcollection-weather-warm-sunnycollection-temperature-threshold-60-70°Fcollection-season-springcollection-season-early-summerspecimen-preparation-standardspecimen-preservation-dryspecimen-preservation-pinnedlabel-data-essentiallocality-data-precisedate-data-essentialcollector-data-valuablehost-data-essentialbehavioral-notes-usefulphotography-documentation-valuableiNaturalist-contribution-appropriatecitizen-science-potential-moderateresearch-value-moderateecological-study-potentialhost-plant-interaction-studypopulation-dynamics-studyclimate-response-studyrange-shift-documentationphenology-shift-documentationmuseum-collection-review-neededtype-specimen-location-unknown-to-authorneotype-designation-status-unknownsyntype-existence-possiblelectotype-designation-status-unknownoriginal-description-brevitysubsequent-description-expansiondiagnostic-characters-stableidentification-key-inclusion-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Tetramesa is a genus of minute phytophagous wasps in the family Eurytomidae comprising over 200 described species. Members are exclusively associated with grasses (Poaceae), where they typically induce stem or inflorescence galls. The genus exhibits pronounced host specificity, with most species restricted to a single grass species or closely related congeners. Several species have been deployed as biological control agents against invasive grasses, including T. romana for giant reed (Arundo donax) and candidate agents for medusahead (Taeniatherum caput-medusae) and African lovegrass (Eragrostis curvula). Adults feed on nectar. Recent phylogenomic studies have synonymized the genera Aiolomorphus and Cathilaria within Tetramesa.
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