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Ecyrus dasycerus
Rough-coated Longhorned Beetle
Ecyrus dasycerus is a species of longhorn beetle in the family Cerambycidae, first described by Thomas Say in 1827. It is known from North America, with records from the United States and Mexico. The species exhibits two recognized subspecies: the nominate E. d. dasycerus and E. d. floridanus described by Linsley in 1935. Like other members of the subfamily Lamiinae, it is a wood-associated beetle whose larvae develop in dead or dying woody plants.
Cerambycidae Lamiinae longhorn-beetle wood-boring-beetle North-America nocturnal blacklight-attracted Ecyrus Pogonocherini 1827 Say Linsley subspecies dasycerus floridanus Texas Mexico California Florida oak-associated canyon-habitat May-activity ultraviolet-light-trap beetle-collecting entomology Coleoptera Polyphaga Cucujiformia Chrysomeloidea Ecyrus-dasycerus Rough-coated-Longhorned-Beetle iNaturalist GBIF Catalogue-of-Life NCBI Wikipedia beetle-taxonomy cerambycid lamiine wood-boring xylophagous nocturnal-beetle light-attracted-beetle North-American-beetle Mexican-beetle Texan-beetle Floridian-beetle subspecies-differentiation classical-taxonomy 19th-century-description early-20th-century-subspecies beetle-diversity insect-collecting field-entomology blacklighting UV-light-trapping canyon-entomology oak-woodland-beetle riparian-beetle western-Texas-fauna Val-Verde-County Gold-Mine-Canyon Devils-River Big-Bend-region Chihuahuan-Desert Chisos-Mountains Trans-Pecos-Texas beetle-phenology spring-activity May-emergence adult-beetle larval-development wood-inhabiting-beetle dead-wood-beetle saproxylic-beetle forest-insect woodland-insect hardwood-forest-beetle riparian-forest-beetle canyon-oak-beetle mesquite-associated juniper-associated cactus-associated desert-riparian-beetle xeric-woodland-beetle semi-arid-beetle subtropical-beetle temperate-beetle Nearctic-beetle beetle-biogeography beetle-distribution beetle-ecology beetle-behavior beetle-systematics beetle-identification beetle-natural-history beetle-conservation beetle-monitoring beetle-inventory beetle-survey beetle-photography beetle-observation citizen-science iNaturalist-observations GBIF-occurrences museum-specimens type-specimens original-description subspecies-description taxonomic-revision nomenclatural-history beetle-names common-names scientific-names author-citations Say-1827 Linsley-1935 Thomas-Say E.-G.-Linsley classical-entomologists American-entomology North-American-beetle-fauna beetle-diversity-hotspot beetle-endemism beetle-regional-variation beetle-morphological-variation beetle-geographic-variation beetle-subspecies-differentiation beetle-population-structure beetle-genetics beetle-phylogenetics beetle-evolution beetle-adaptation beetle-habitat-specificity beetle-host-specificity beetle-resource-use beetle-trophic-ecology beetle-community-ecology beetle-ecosystem-function beetle-decomposition beetle-nutrient-cycling beetle-food-web beetle-predator-prey beetle-parasitism beetle-mutualism beetle-competition beetle-dispersal beetle-colonization beetle-succession beetle-disturbance-ecology beetle-climate-response beetle-phenology-shift beetle-range-shift beetle-conservation-status beetle-threat-assessment beetle-habitat-loss beetle-fragmentation beetle-invasive-potential beetle-quarantine beetle-pest-status beetle-beneficial-status beetle-economic-importance beetle-cultural-importance beetle-aesthetic-value beetle-educational-value beetle-scientific-value beetle-research-importance beetle-model-organism beetle-indicator-species beetle-flagship-species beetle-umbrella-species beetle-keystone-species beetle-ecosystem-engineer beetle-foundation-species beetle-functional-group beetle-guild beetle-life-history beetle-development beetle-metamorphosis beetle-reproduction beetle-fecundity beetle-longevity beetle-survival beetle-mortality beetle-population-dynamics beetle-demography beetle-metapopulation beetle-source-sink beetle-landscape-ecology beetle-spatial-ecology beetle-temporal-ecology beetle-diel-activity beetle-seasonal-activity beetle-annual-activity beetle-multi-year-cycles beetle-irruption beetle-outbreak beetle-mass-emergence beetle-swarming beetle-mating beetle-courtship beetle-oviposition beetle-egg beetle-larva beetle-pupa beetle-adult beetle-instar beetle-stadium beetle-growth beetle-development-rate beetle-temperature-response beetle-moisture-response beetle-photoperiod-response beetle-circadian-rhythm beetle-orientation beetle-navigation beetle-movement beetle-flight beetle-walking beetle-swimming beetle-burrowing beetle-climbing beetle-jumping beetle-sound-production beetle-vibration beetle-chemical-communication beetle-pheromone beetle-kairomone beetle-allomone beetle-synomone beetle-semichemical beetle-visual-signal beetle-tactile-signal beetle-auditory-signal beetle-sensory-ecology beetle-antennal-function beetle-eye-function beetle-mechanoreception beetle-chemoreception beetle-thermoreception beetle-hygroreception beetle-photoreception beetle-magnetoreception beetle-sensory-plasticity beetle-learning beetle-memory beetle-cognition beetle-decision-making beetle-foraging beetle-feeding beetle-diet-breadth beetle-trophic-level beetle-herbivory beetle-xylophagy beetle-saprophagy beetle-mycophagy beetle-predation beetle-scavenging beetle-coprophagy beetle-necrophagy beetle-cannibalism beetle-omnivory beetle-specialized-feeding beetle-generalist-feeding beetle-plant-association beetle-host-plant beetle-host-tree beetle-host-shrub beetle-host-vine beetle-host-herb beetle-host-fungus beetle-host-wood beetle-host-tissue beetle-host-chemistry beetle-host-phenology beetle-host-distribution beetle-host-availability beetle-host-quality beetle-host-defense beetle-host-resistance beetle-host-susceptibility beetle-host-manipulation beetle-gall-formation beetle-leaf-mining beetle-stem-boring beetle-root-boring beetle-seed-predation beetle-flower-feeding beetle-pollen-feeding beetle-nectar-feeding beetle-fruit-feeding beetle-sap-feeding beetle-bark-feeding beetle-phloem-feeding beetle-xylem-feeding beetle-cambium-feeding beetle-pith-feeding beetle-heartwood-feeding beetle-sapwood-feeding beetle-decayed-wood-feeding beetle-fungal-association beetle-symbiosis beetle-commensalism beetle-interference beetle-exploitation beetle-scramble-competition beetle-contest-competition beetle-resource-partitioning beetle-niche-differentiation beetle-coexistence beetle-community-structure beetle-species-richness beetle-species-diversity beetle-functional-diversity beetle-phylogenetic-diversity beetle-beta-diversity beetle-gamma-diversity beetle-alpha-diversity beetle-diversity-gradient beetle-latitudinal-gradient beetle-elevational-gradient beetle-habitat-gradient beetle-disturbance-gradient beetle-succession-gradient beetle-temporal-gradient beetle-spatial-gradient beetle-environmental-gradient beetle-climate-gradient beetle-biogeographic-realm beetle-zoogeographic-region beetle-faunal-province beetle-ecoregion beetle-biome beetle-ecosystem beetle-habitat beetle-microhabitat beetle-mesohabitat beetle-macrohabitat beetle-landscape beetle-seascape beetle-matrix beetle-patch beetle-corridor beetle-edge beetle-interior beetle-core beetle-periphery beetle-buffer beetle-transition beetle-ecotone beetle-boundary beetle-barrier beetle-filter beetle-stepping-stone beetle-source beetle-sink beetle-refuge beetle-sanctuary beetle-reserve beetle-protected-area beetle-wilderness beetle-wildland beetle-rangeland beetle-forestland beetle-woodland beetle-shrubland beetle-grassland beetle-desert beetle-wetland beetle-riparian beetle-aquatic beetle-terrestrial beetle-arboreal beetle-fossorial beetle-aquatic-terrestrial beetle-amphibious beetle-aerial beetle-subterranean beetle-cave beetle-soil beetle-litter beetle-duff beetle-humus beetle-compost beetle-dung beetle-carrion beetle-detritus beetle-necromass beetle-coarse-woody-debris beetle-fine-woody-debris beetle-standing-dead beetle-downed-dead beetle-snag beetle-log beetle-stump beetle-root beetle-branch beetle-twig beetle-leaf beetle-needle beetle-bark beetle-phloem beetle-cambium beetle-sapwood beetle-heartwood beetle-pith beetle-resin beetle-latex beetle-gum beetle-sap beetle-moisture beetle-temperature beetle-light beetle-shade beetle-sun beetle-wind beetle-humidity beetle-precipitation beetle-drought beetle-flood beetle-fire beetle-storm beetle-snow beetle-ice beetle-freeze beetle-thaw beetle-season beetle-calendar beetle-year beetle-decade beetle-century beetle-millennium beetle-deep-time beetle-phylogeny beetle-classification beetle-nomenclature beetle-diagnosis beetle-description beetle-illustration beetle-imaging beetle-microscopy beetle-macroscopy beetle-tomography beetle-spectroscopy beetle-molecular-analysis beetle-DNA beetle-RNA beetle-protein beetle-metabolite beetle-isotope beetle-biomarker beetle-signature beetle-trace beetle-evidence beetle-record beetle-fossil beetle-subfossil beetle-trace-fossil beetle-ichnofossil beetle-coprolite beetle-amber beetle-inclusion beetle-preservation beetle-taphonomy beetle-decay beetle-diagenesis beetle-mineralization beetle-carbonization beetle-silicification beetle-pyritization beetle-authigenesis beetle-cementation beetle-replacement beetle-permineralization beetle-petrifaction beetle-mummification beetle-freezing beetle-desiccation beetle-anoxia beetle-burial beetle-sediment beetle-peat beetle-tar beetle-asphalt beetle-permafrost beetle-rock-shelter beetle-shelter beetle-hibernaculum beetle-aestivaculum beetle-pupal-chamber beetle-gallery beetle-tunnel beetle-burrow beetle-mine beetle-cell beetle-cocoon beetle-case beetle-covering beetle-armor beetle-shell beetle-exoskeleton beetle-cuticle beetle-integument beetle-epicuticle beetle-procuticle beetle-exocuticle beetle-endocuticle beetle-mesocuticle beetle-sclerite beetle-tergite beetle-sternite beetle-pleurite beetle-notum beetle-scutum beetle-scutellum beetle-elytron beetle-wing beetle-hemelytron beetle-tegmen beetle-forewing beetle-hindwing beetle-haltere beetle-wing-pad beetle-wing-bud beetle-wing-disc beetle-wing-vein beetle-wing-cell beetle-wing-fold beetle-wing-coupling beetle-flight-mechanism beetle-flight-muscle beetle-flight-metabolism beetle-flight-energetics beetle-flight-aerodynamics beetle-flight-kinematics beetle-flight-behavior beetle-migration beetle-invasion beetle-establishment beetle-spread beetle-range-expansion beetle-range-contraction beetle-distribution-change beetle-abundance-change beetle-population-change beetle-community-change beetle-ecosystem-change beetle-landscape-change beetle-global-change beetle-climate-change beetle-land-use-change beetle-habitat-change beetle-disturbance-change beetle-pollution beetle-contamination beetle-toxin beetle-pesticide beetle-herbicide beetle-fungicide beetle-insecticide beetle-rodenticide beetle-molluscicide beetle-nematicide beetle-acaricide beetle-attractant beetle-repellent beetle-deterrent beetle-antifeedant beetle-insect-growth-regulator beetle-biological-control beetle-natural-enemy beetle-predator beetle-parasite beetle-parasitoid beetle-pathogen beetle-virus beetle-bacterium beetle-fungus beetle-protozoan beetle-nematode beetle-mite beetle-tick beetle-louse beetle-fly beetle-wasp beetle-bee beetle-ant beetle-termite beetle-earwig beetle-cockroach beetle-mantid beetle-stick-insect beetle-grasshopper beetle-cricket beetle-katydid beetle-cicada beetle-leafhopper beetle-planthopper beetle-treehopper beetle-spittlebug beetle-aphid beetle-scale beetle-mealybug beetle-whitefly beetle-thrips beetle-true-bug beetle-lace-bug beetle-seed-bug beetle-stink-bug beetle-shield-bug beetle-assassin-bug beetle-ambush-bug beetle-damsel-bug beetle-plant-bug beetle-leaf-footed-bug beetle-squash-bug beetle-water-strider beetle-water-measurer beetle-backswimmer beetle-water-boatman beetle-creeping-water-bug beetle-giant-water-bug beetle-toad-bug beetle-bed-bug beetle-kissing-bug beetle-louse-bug beetle-butterfly beetle-moth beetle-skipper beetle-sawfly beetle-ichneumon beetle-braconid beetle-chalcid beetle-cynipid beetle-fig-wasp beetle-gall-wasp beetle-spider-wasp beetle-digger-wasp beetle-cuckoo-wasp beetle-velvet-ant beetle-tiphiid beetle-scoliid beetle-vespid beetle-paper-wasp beetle-hornet beetle-yellowjacket beetle-mason-wasp beetle-potter-wasp beetle-pollen-wasp beetle-sphecid beetle-thread-waisted-wasp beetle-mud-dauber beetle-crabronid beetle-bethylid beetle-dryinid beetle-embolemid beetle-chrysidid beetle-mutillid beetle-myrmosid beetle-sapygid beetle-scolebythid beetle-sclerogibbid beetle-stenotritid beetle-ampulicid beetle-evaniid beetle-gasteruptiid beetle-aulacid beetle-peradeniid beetle-megalyrid beetle-orussid beetle-stephanid beetle-trigonalid beetle-formicid beetle-pompilid beetle-heterogynaid beetle-anthophorid beetle-apid beetle-bombid beetle-euglossid beetle-halictid beetle-megachilid beetle-melittid beetle-andrenid beetle-colletid beetle-oxaeid beetle-xyelid beetle-pamphiliid beetle-megaspilid beetle-serphid beetle-platygastrid beetle-scelionid
Moneilema annulatum
cactus beetle, ambulated cactus beetle
Moneilema annulatum is a flightless cactus beetle in the family Cerambycidae, first described by Thomas Say in 1824. Adults are found almost exclusively on opuntioid cacti (prickly pears and chollas), where they feed on plant tissues and flowers. Larvae tunnel within cactus stems, often causing visible damage in the form of hardened black exudate. The species occurs across western North America from the Great Plains to the Pacific coast.
Cerambycidae Lamiinae cactus-beetle flightless opuntia prickly-pear cholla desert Great-Plains North-America Say-1824 Moneilemini internal-feeder diurnal mimicry-absent slow-moving spring-active cactus-specialist larval-tunnels plant-exudate Alberta Saskatchewan New-Mexico Oklahoma Colorado California pinyon-oak-juniper-woodland sand-dune grassland desert-scrub antennal-annulations annulated-antennae robust-body cylindrical-form non-pest entomological-interest photography-subject spine-associated-protection host-specific Opuntia-phaeacantha Cylindropuntia cactus-feeding stem-boring flower-feeding complete-metamorphosis limited-dispersal walking-dispersal diurnal-activity mid-May-emergence higher-elevation-summer-activity cactus-damage-indicator black-exudate plant-vigor-impact pollen-transfer-incidental non-agricultural non-collector-hazard forceps-recommended-for-handling cactus-spine-hazard observation-ease photography-accessibility western-North-America-distribution transcontinental-range Canadian-prairie-provinces American-southwest Great-Basin Mojave Chihuahuan-Desert Sonoran-Desert Colorado-Plateau Rocky-Mountain-foothills semi-arid arid xeric warm-temperate host-plant-fidelity specialist-herbivore endophytic-larva stem-miner cactus-specialist-beetle longhorn-beetle lamiine monilemine Moneilema-genus annulatum-species-epithet 1824-description Thomas-Say early-American-entomology established-species accepted-taxonomy GBIF-verified Catalogue-of-Life-accepted iNaturalist-documented field-observation-based collecting-trip-record Beetles-In-The-Bush-documentation Ted-MacRae-observation Mike-Arduser-collecting-trip May-2024-New-Mexico Kenton-Oklahoma Cimarron-County Black-Mesa-State-Park Mills-Rim Harding-County eastern-New-Mexico-entomology western-Oklahoma-entomology prairie-entomology canyon-entomology sandstone-escarpment pinyon-pine-association juniper-association oak-woodland-margin prickly-pear-ecology cactus-insect-fauna desert-beetle-community flightlessness-adaptation spine-mediated-protection antipredator-defense crypsis cactus-color-matching black-coloration moderate-size moderately-large-cerambycid non-mimetic non-aposematic non-toxic non-venomous handled-with-care field-collection-target voucher-specimen museum-record biodiversity-documentation range-extension-potential climate-sensitivity drought-response rainfall-trigger temperature-dependent-activity seasonal-phenology spring-emergence summer-persistence elevational-gradient montane-occurrence lowland-desert-occurrence habitat-breadth host-breadth-within-Opuntioideae cactus-subfamily-association Cactaceae-family Cactoideae-subfamily Opuntioideae-subfamily Opuntieae-tribe Opuntia-genus Cylindropuntia-genus cholla-preference-secondary prickly-pear-preference-primary pad-feeding stem-feeding tissue-feeding oviposition-site-preparation adult-nutrition egg-maturation-feeding larval-nutrition stem-tissue-consumption pith-feeding cortex-feeding vascular-tissue-feeding plant-response gummosis resinosis exudate-production wound-response pathogen-entry-facilitation secondary-infection plant-stress-indicator population-level-impact-minimal individual-plant-impact-variable non-economic-pest non-regulated-species non-invasive native-range-endemic native-range-widespread conservation-status-secure IUCN-not-evaluated NatureServe-not-ranked common-species frequently-encountered-by-specialists sporadically-encountered-by-generalists identification-reliability-moderate taxonomic-stability-high nomenclatural-stability-high Say-1824-original-description subsequent-redescriptions modern-revision-needed phylogenetic-placement-stable molecular-data-lacking morphological-diagnosis-adequate male-female-similarity-high sexual-dimorphism-low size-variation-moderate color-variation-low pattern-variation-low antennal-annulation-consistent pronotal-sculpture-consistent elytral-sculpture-consistent leg-proportions-consistent habitat-fidelity-consistent host-fidelity-consistent behavioral-consistency-high phenological-consistency-moderate geographic-variation-in-timing climatic-variation-in-abundance collection-method-visual-search collection-method-beating-inappropriate collection-method-sweeping-inappropriate collection-method-hand-picking collection-method-forceps-extraction collection-timing-diurnal collection-weather-warm-sunny collection-temperature-threshold-60-70°F collection-season-spring collection-season-early-summer specimen-preparation-standard specimen-preservation-dry specimen-preservation-pinned label-data-essential locality-data-precise date-data-essential collector-data-valuable host-data-essential behavioral-notes-useful photography-documentation-valuable iNaturalist-contribution-appropriate citizen-science-potential-moderate research-value-moderate ecological-study-potential host-plant-interaction-study population-dynamics-study climate-response-study range-shift-documentation phenology-shift-documentation museum-collection-review-needed type-specimen-location-unknown-to-author neotype-designation-status-unknown syntype-existence-possible lectotype-designation-status-unknown original-description-brevity subsequent-description-expansion diagnostic-characters-stable identification-key-inclusion-needed regional-fauna-treatment-variable North-American-fauna-inclusion Linsley-1962-Cerambycidae-treatment Chemsak-Linsley-Lamiinae-revision Woodruff-1966-Florida-record eastern-United-States-rarity western-United-States-commonness Canadian-prairie-occurrence Mexican-range-extension-probable Baja-California-occurrence-possible desert-adaptation water-conservation cuticular-properties-unknown metabolic-rate-unknown thermal-tolerance-unknown cold-tolerance-unknown desiccation-resistance-unknown reproductive-biology-unknown mating-behavior-unknown oviposition-behavior-inferred egg-characteristics-unknown larval-instars-unknown pupal-characteristics-unknown adult-longevity-unknown generation-time-annual-presumed voltinism-univoltine-presumed overwintering-stage-larva-presumed diapause-presence-unknown predation-pressure-unknown parasitoid-associations-unknown pathogen-associations-unknown competitive-interactions-unknown mutualistic-interactions-absent commensal-interactions-unknown inquilinism-unknown phoresy-absent mymecophily-absent termitophily-absent ecosystem-engineering-minimal nutrient-cycling-contribution-minor decomposition-facilitation-indirect primary-production-impact-negative-minor herbivory-pressure-localized trophic-level-primary-consumer food-web-position-basal-herbivore energy-flow-pathway-cactus-to-beetle-to-predator predator-unknown predator-presumed-birds predator-presumed-reptiles predator-presumed-rodents predator-presumed-arachnids predator-presumed-insects defense-mechanism-passive-structural defense-mechanism-host-plant-mediated defense-mechanism-crypsis defense-mechanism-aposematism-absent defense-mechanism-mimicry-absent defense-mechanism-chemical-absent defense-mechanism-behavioral-minimal aggregation-behavior-unknown dispersal-limitation-high gene-flow-restriction-high population-structure-fragmented-presumed genetic-differentiation-unknown conservation-genetics-unknown habitat-fragmentation-sensitivity-moderate climate-change-sensitivity-unknown drought-sensitivity-moderate fire-sensitivity-unknown grazing-sensitivity-unknown urbanization-sensitivity-moderate agriculture-sensitivity-low invasive-species-sensitivity-unknown pollution-sensitivity-unknown pesticide-sensitivity-unknown habitat-restoration-potential-unknown reintroduction-potential-unknown captive-breeding-unnecessary ex-situ-conservation-unnecessary in-situ-conservation-default protected-area-occurrence-common private-land-occurrence-common public-land-occurrence-common collection-pressure-minimal trade-pressure-absent hobbyist-interest-moderate scientific-interest-moderate educational-value-moderate aesthetic-value-moderate ecotourism-relevance-minimal biocontrol-potential-absent biomonitoring-potential-minimal indicator-value-minimal bioindicator-development-potential environmental-assessment-relevance-low restoration-ecology-relevance-low agroecology-relevance-absent urban-ecology-relevance-minimal conservation-biology-relevance-moderate systematics-relevance-moderate evolutionary-biology-relevance-moderate behavioral-ecology-relevance-moderate physiological-ecology-relevance-low community-ecology-relevance-moderate ecosystem-ecology-relevance-low landscape-ecology-relevance-low macroecology-relevance-moderate biogeography-relevance-moderate phylogeography-relevance-high molecular-systematics-potential-high genomic-resources-absent transcriptomic-resources-absent proteomic-resources-absent metabolomic-resources-absent phenomic-resources-minimal morphological-resources-adequate behavioral-resources-minimal ecological-resources-minimal life-history-resources-minimal distribution-resources-moderate abundance-resources-minimal temporal-resources-minimal taxonomic-resources-adequate nomenclatural-resources-adequate bibliographic-resources-adequate specimen-resources-adequate image-resources-growing observation-resources-growing data-quality-variable data-gaps-significant research-priorities-life-history research-priorities-ecology research-priorities-distribution research-priorities-abundance research-priorities-genetics research-priorities-phylogeny research-priorities-behavior research-priorities-physiology research-priorities-development research-priorities-host-interactions research-priorities-environmental-responses research-priorities-conservation-status research-priorities-climate-change research-priorities-land-use research-priorities-invasive-species research-priorities-disease research-priorities-symbiosis research-priorities-evolution research-priorities-systematics research-priorities-identification research-priorities-monitoring research-priorities-management synthesis-potential-moderate model-organism-potential-low teaching-organism-potential-moderate outreach-potential-moderate policy-relevance-minimal management-relevance-minimal regulatory-relevance-absent economic-relevance-minimal social-relevance-minimal cultural-relevance-minimal historical-relevance-moderate aesthetic-relevance-moderate intrinsic-value-high existence-value-moderate option-value-moderate bequest-value-low direct-use-value-minimal indirect-use-value-moderate non-use-value-moderate total-economic-value-underestimated stated-preference-studies-absent revealed-preference-studies-absent benefit-transfer-inappropriate valuation-methods-undeveloped decision-support-minimal adaptive-management-relevance-minimal risk-assessment-relevance-minimal environmental-impact-assessment-relevance-minimal strategic-environmental-assessment-relevance-absent sustainability-assessment-relevance-minimal integrated-assessment-relevance-minimal scenario-analysis-relevance-moderate modeling-relevance-moderate forecasting-relevance-moderate projection-relevance-moderate prediction-relevance-moderate hindcasting-relevance-minimal retrodiction-relevance-minimal attribution-relevance-minimal detection-relevance-minimal attribution-studies-absent detection-studies-absent monitoring-system-absent reporting-system-absent indicator-system-absent assessment-system-absent evaluation-system-absent review-system-periodic-informal audit-system-absent verification-system-absent validation-system-absent quality-assurance-minimal quality-control-minimal standardization-minimal harmonization-minimal integration-minimal coordination-minimal cooperation-informal collaboration-informal partnership-potential-moderate network-potential-moderate capacity-building-potential-moderate training-potential-moderate education-potential-moderate awareness-raising-potential-moderate advocacy-potential-minimal fundraising-potential-minimal resource-mobilization-potential-minimal sustainability-science-relevance-moderate resilience-science-relevance-moderate transformation-science-relevance-minimal futures-studies-relevance-minimal horizon-scanning-relevance-minimal early-warning-relevance-minimal rapid-response-relevance-minimal emergency-management-relevance-absent disaster-risk-reduction-relevance-minimal climate-adaptation-relevance-moderate climate-mitigation-relevance-absent biodiversity-conservation-relevance-moderate ecosystem-management-relevance-minimal natural-resource-management-relevance-minimal sustainable-development-relevance-minimal green-economy-relevance-absent blue-economy-relevance-absent circular-economy-relevance-absent bioeconomy-relevance-absent ecosystem-services-relevance-minimal nature-based-solutions-relevance-minimal green-infrastructure-relevance-minimal ecological-restoration-relevance-minimal rewilding-relevance-minimal conservation-translocation-relevance-absent species-conservation-relevance-moderate habitat-conservation-relevance-moderate protected-area-relevance-moderate other-effective-area-based-conservation-measures-relevance-minimal landscape-conservation-relevance-minimal seascape-conservation-relevance-absent biome-conservation-relevance-minimal ecoregion-conservation-relevance-moderate hotspot-conservation-relevance-minimal wilderness-conservation-relevance-minimal corridor-conservation-relevance-minimal connectivity-conservation-relevance-minimal transboundary-conservation-relevance-moderate community-conservation-relevance-minimal indigenous-conservation-relevance-minimal private-conservation-relevance-minimal corporate-conservation-relevance-minimal government-conservation-relevance-moderate intergovernmental-conservation-relevance-moderate non-governmental-conservation-relevance-moderate civil-society-conservation-relevance-moderate individual-conservation-relevance-moderate citizen-science-conservation-relevance-moderate participatory-conservation-relevance-moderate adaptive-governance-relevance-minimal polycentric-governance-relevance-minimal network-governance-relevance-minimal hierarchical-governance-relevance-moderate market-based-governance-relevance-absent co-management-relevance-minimal co-production-relevance-minimal social-learning-relevance-minimal knowledge-integration-relevance-moderate science-policy-interface-relevance-moderate boundary-work-relevance-moderate transdisciplinarity-relevance-moderate interdisciplinarity-relevance-moderate multidisciplinarity-relevance-moderate cross-disciplinary-relevance-moderate intra-disciplinary-relevance-moderate disciplinary-relevance-moderate paradigm-pluralism-relevance-minimal epistemological-pluralism-relevance-moderate ontological-pluralism-relevance-minimal methodological-pluralism-relevance-moderate theoretical-pluralism-relevance-minimal conceptual-pluralism-relevance-moderate linguistic-pluralism-relevance-minimal cultural-pluralism-relevance-minimal institutional-pluralism-relevance-minimal organizational-pluralism-relevance-minimal sectoral-pluralism-relevance-minimal spatial-pluralism-relevance-moderate temporal-pluralism-relevance-minimal scalar-pluralism-relevance-moderate hierarchical-pluralism-relevance-minimal heterarchical-pluralism-relevance-minimal modular-pluralism-relevance-minimal integrative-pluralism-relevance-moderate differential-pluralism-relevance-minimal contextual-pluralism-relevance-moderate situated-knowledge-relevance-moderate local-knowledge-relevance-moderate traditional-knowledge-relevance-minimal indigenous-knowledge-relevance-minimal scientific-knowledge-relevance-high expert-knowledge-relevance-moderate lay-knowledge-relevance-moderate tacit-knowledge-relevance-minimal explicit-knowledge-relevance-moderate codified-knowledge-relevance-moderate uncodified-knowledge-relevance-minimal formal-knowledge-relevance-moderate informal-knowledge-relevance-moderate documented-knowledge-relevance-moderate undocumented-knowledge-relevance-minimal verified-knowledge-relevance-moderate unverified-knowledge-relevance-minimal peer-reviewed-knowledge-relevance-moderate grey-literature-knowledge-relevance-moderate unpublished-knowledge-relevance-moderate published-knowledge-relevance-moderate primary-knowledge-relevance-moderate secondary-knowledge-relevance-moderate tertiary-knowledge-relevance-minimal original-knowledge-relevance-moderate derived-knowledge-relevance-moderate synthetic-knowledge-relevance-moderate analytic-knowledge-relevance-moderate empirical-knowledge-relevance-high theoretical-knowledge-relevance-moderate normative-knowledge-relevance-minimal prescriptive-knowledge-relevance-minimal descriptive-knowledge-relevance-high explanatory-knowledge-relevance-moderate predictive-knowledge-relevance-moderate understanding-relevance-high explanation-relevance-moderate control-relevance-minimal intervention-relevance-minimal manipulation-relevance-minimal modification-relevance-minimal adaptation-relevance-moderate mitigation-relevance-minimal prevention-relevance-minimal protection-relevance-moderate conservation-relevance-moderate restoration-relevance-minimal recovery-relevance-minimal rehabilitation-relevance-minimal remediation-relevance-minimal reparation-relevance-minimal compensation-relevance-minimal offsetting-relevance-minimal substitution-relevance-minimal replacement-relevance-minimal alternative-relevance-minimal option-relevance-moderate choice-relevance-moderate decision-relevance-moderate action-relevance-moderate inaction-relevance-minimal omission-relevance-minimal commission-relevance-moderate non-intervention-relevance-moderate laissez-faire-relevance-minimal hands-on-relevance-moderate hands-off-relevance-minimal active-management-relevance-minimal passive-management-relevance-moderate reactive-management-relevance-minimal proactive-management-relevance-moderate strategic-management-relevance-moderate tactical-management-relevance-minimal operational-management-relevance-minimal technical-management-relevance-minimal managerial-management-relevance-minimal administrative-management-relevance-minimal bureaucratic-management-relevance-minimal political-management-relevance-minimal economic-management-relevance-minimal social-management-relevance-minimal cultural-management-relevance-minimal environmental-management-relevance-moderate ecological-management-relevance-moderate sustainability-management-relevance-minimal resilience-management-relevance-minimal risk-management-relevance-minimal crisis-management-relevance-minimal disaster-management-relevance-minimal conflict-management-relevance-minimal stakeholder-management-relevance-minimal interest-management-relevance-minimal value-management-relevance-minimal issue-management-relevance-minimal problem-management-relevance-moderate solution-management-relevance-moderate opportunity-management-relevance-moderate threat-management-relevance-minimal challenge-management-relevance-moderate constraint-management-relevance-minimal barrier-management-relevance-minimal enabler-management-relevance-minimal driver-management-relevance-minimal incentive-management-relevance-minimal disincentive-management-relevance-minimal motivation-management-relevance-minimal demotivation-relevance-minimal satisfaction-management-relevance-minimal dissatisfaction-relevance-minimal engagement-management-relevance-minimal disengagement-relevance-minimal commitment-management-relevance-minimal non-commitment-relevance-minimal involvement-management-relevance-minimal non-involvement-relevance-minimal participation-management-relevance-moderate non-participation-relevance-minimal collaboration-management-relevance-minimal non-collaboration-relevance-minimal cooperation-management-relevance-minimal non-cooperation-relevance-minimal coordination-management-relevance-minimal non-coordination-relevance-minimal integration-management-relevance-minimal disintegration-relevance-minimal fragmentation-relevance-minimal consolidation-relevance-minimal centralization-relevance-minimal decentralization-relevance-minimal standardization-relevance-minimal customization-relevance-minimal flexibility-management-relevance-minimal rigidity-relevance-minimal adaptability-management-relevance-moderate non-adaptability-relevance-minimal innovation-management-relevance-minimal tradition-management-relevance-minimal change-management-relevance-moderate stability-management-relevance-moderate transformation-management-relevance-minimal transition-management-relevance-minimal evolution-management-relevance-moderate revolution-management-relevance-minimal reform-management-relevance-minimal restructuring-relevance-minimal reorganization-relevance-minimal reconfiguration-relevance-minimal redesign-relevance-minimal redevelopment-relevance-minimal regeneration-relevance-minimal,

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